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<document id="420401E27635284BD67167E9AF714F17" ID-CLB-Dataset="22814" ID-DOI="10.11646/zootaxa.4766.4.1" ID-GBIF-Dataset="1e2996e2-e31f-45e3-9149-60ebce934708" ID-ISSN="1175-5334" ID-Zenodo-Dep="3765652" ID-ZooBank="urn:lsid:zoobank.org:pub:2F8FEDC1-6891-46D1-B372-858CDCBE4051" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1587754498722" checkinUser="carolina" docAuthor="Kompier, Tom, Dow, Rory A. &amp; Steinhoff, Philip O. M." docDate="2020" docId="AF6A87D543686176FF6ADDAE4B6B3BD8" docLanguage="en" docName="zootaxa.4766.4.1.pdf.imf" docOrigin="Zootaxa 4766 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coeliccia curua Kompier &amp; Dow &amp; Steinhoff 2020, sp. nov." docType="treatment" docVersion="6" lastPageNumber="507" masterDocId="5353FFAD436C6170FFFDDB7048083A53" masterDocTitle="Five new species of Coeliccia Kirby, 1890 from Vietnam (Odonata: Platycnemididae), and information on several other species of the genus" masterLastPageNumber="538" masterPageNumber="501" pageNumber="505" updateTime="1698835840532" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="003BDCBFDD77F780D6908AC578A60C4D">Five new species of Coeliccia Kirby, 1890 from Vietnam (Odonata: Platycnemididae), and information on several other species of the genus</mods:title>
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<mods:namePart id="2E34D60CC4BBE4005CE031661E20824D">Kompier, Tom</mods:namePart>
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<mods:namePart id="B5081150BF1095980101335BB8DAEFBC">Dow, Rory A.</mods:namePart>
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<mods:namePart id="B8FCE0E1E9E283B0FDF15E65607141EA">Steinhoff, Philip O. M.</mods:namePart>
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<treatment id="AF6A87D543686176FF6ADDAE4B6B3BD8" ID-DOI="http://doi.org/10.5281/zenodo.3803422" ID-GBIF-Taxon="163739039" ID-Zenodo-Dep="3803422" LSID="urn:lsid:plazi:treatment:AF6A87D543686176FF6ADDAE4B6B3BD8" httpUri="http://treatment.plazi.org/id/AF6A87D543686176FF6ADDAE4B6B3BD8" lastPageId="6" lastPageNumber="507" pageId="4" pageNumber="505">
<subSubSection id="6FD9654843686174FF6ADDAE49D23CAA" box="[151,474,1758,1785]" pageId="4" pageNumber="505" type="nomenclature">
<paragraph id="277C36C343686174FF6ADDAE49D23CAA" blockId="4.[151,802,1758,1821]" box="[151,474,1758,1785]" pageId="4" pageNumber="505">
<heading id="7C3481AF43686174FF6ADDAE49D23CAA" bold="true" box="[151,474,1758,1785]" fontSize="10" level="2" pageId="4" pageNumber="505" reason="2">
<emphasis id="15B7EAD143686174FF6ADDAE49D23CAA" bold="true" box="[151,474,1758,1785]" pageId="4" pageNumber="505">
2.
<taxonomicName id="E0C34D4043686174FF48DDAE497C3CAA" ID-CoL="9X2BK" authority="Kompier &amp; Dow &amp; Steinhoff, 2020" authorityName="Kompier &amp; Dow &amp; Steinhoff" authorityYear="2020" box="[181,372,1758,1785]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="curua" status="sp. nov.">
<emphasis id="15B7EAD143686174FF48DDAE497C3CAA" bold="true" box="[181,372,1758,1785]" italics="true" pageId="4" pageNumber="505">Coeliccia curua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0E8457AA43686174FE86DDAE49D23CAA" box="[379,474,1758,1785]" pageId="4" pageNumber="505" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="6FD9654843686174FF6ADC724B2A3D4E" box="[151,802,1793,1821]" pageId="4" pageNumber="505" type="description">
<paragraph id="277C36C343686174FF6ADC724B2A3D4E" blockId="4.[151,802,1758,1821]" box="[151,802,1793,1821]" pageId="4" pageNumber="505">
(
<figureCitation id="BFF82A4643686174FF62DC7248FD3D4F" box="[159,245,1794,1820]" captionStart="FIGURE 1" captionStartId="3.[151,250,1981,2006]" captionTargetBox="[151,1436,993,1956]" captionTargetId="figure@3.[151,1436,993,1956]" captionTargetPageId="3" captionText="FIGURE 1. Head of male Coeliccia spp. in dorsolateral view. (a) C. caerulea sp. nov., holotype; (b) C. coronata sp. nov., holotype; (c) C. diomedea sp. nov., holotype; (d) C. curua sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.3765654" httpUri="https://zenodo.org/record/3765654/files/figure.png" pageId="4" pageNumber="505">Figs 1d</figureCitation>
,
<figureCitation id="BFF82A4643686174FEFFDC7249283D4F" box="[258,288,1794,1820]" captionStart="FIGURE 2" captionStartId="4.[151,250,1615,1640]" captionTargetBox="[151,1436,633,1590]" captionTargetId="figure@4.[151,1436,633,1590]" captionTargetPageId="4" captionText="FIGURE 2. Prothorax of male Coeliccia spp. in lateral view. (a) C. caerulea, holotype; (b) C. coronata, holotype; (c) C. diomedea, holotype; (d) C. curua, holotype." figureDoi="http://doi.org/10.5281/zenodo.3765656" httpUri="https://zenodo.org/record/3765656/files/figure.png" pageId="4" pageNumber="505">2d</figureCitation>
,
<figureCitation id="BFF82A4643686174FED1DC7249413D4F" box="[300,329,1794,1820]" captionStart="FIGURE 3" captionStartId="6.[151,250,1882,1907]" captionTargetBox="[151,1436,417,1857]" captionTargetId="figure@6.[151,1436,417,1857]" captionTargetPageId="6" captionText="FIGURE 3. Prothorax of Coeliccia spp. in dorsal view. [a-e] male, [f] female. (a) C. caerulea, holotype; (b) C. coronata, holotype; (c) C. diomedea, holotype; (d) C. curua, holotype; (e) C. pulchella sp. nov., holotype; (f) C. caerulea, paratype used for description." figureDoi="http://doi.org/10.5281/zenodo.3765658" httpUri="https://zenodo.org/record/3765658/files/figure.png" pageId="4" pageNumber="505">3d</figureCitation>
,
<figureCitation id="BFF82A4643686174FEABDC72497C3D4F" box="[342,372,1794,1820]" captionStart="FIGURE 4" captionStartId="7.[151,250,1090,1115]" captionTargetBox="[151,1437,181,1063]" captionTargetId="figure@7.[151,1437,181,1064]" captionTargetPageId="7" captionText="FIGURE 4. Thorax of male Coeliccia spp. in lateral view. (a) C. caerulea, holotype; (b) C. coronata, holotype; (c) C. diomedea, holotype; (d) C. curua, holotype." figureDoi="http://doi.org/10.5281/zenodo.3765660" httpUri="https://zenodo.org/record/3765660/files/figure.png" pageId="4" pageNumber="505">4d</figureCitation>
,
<figureCitation id="BFF82A4643686174FE7DDC7149963D4F" box="[384,414,1793,1820]" captionStart="FIGURE 5" captionStartId="8.[151,250,910,935]" captionTargetBox="[151,1436,181,885]" captionTargetId="figure@8.[151,1436,181,885]" captionTargetPageId="8" captionText="FIGURE 5. Thorax of male Coeliccia spp. in dorsal view. (a) C. caerulea, holotype; (b) C. coronata, holotype; (c) C. diomedea, holotype; (d) C. curua, holotype." figureDoi="http://doi.org/10.5281/zenodo.3765662" httpUri="https://zenodo.org/record/3765662/files/figure.png" pageId="4" pageNumber="505">5d</figureCitation>
,
<figureCitation id="BFF82A4643686174FE57DC7249BF3D4F" box="[426,439,1794,1820]" captionStart="FIGURE 7" captionStartId="9.[151,250,1813,1838]" captionTargetBox="[151,1436,181,1786]" captionTargetId="figure@9.[151,1436,181,1787]" captionTargetPageId="9" captionText="FIGURE 7. Abdomen tip of male Coeliccia spp. [a,e,g,i] lateral view, [c] dorsolateral view, [b,d,f,h,j] dorsal view. (a,b) C. caerulea, holotype; (c,d) C. pulchella, holotype; (e,f) C. curua, holotype; (g,h) C. coronata, holotype; (i,j) C. diomedea, holotype." figureDoi="http://doi.org/10.5281/zenodo.3765666" httpUri="https://zenodo.org/record/3765666/files/figure.png" pageId="4" pageNumber="505">7</figureCitation>
ef, 9, 10ab, 12, 25ad, 29cd)
</paragraph>
</subSubSection>
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<paragraph id="277C36C343686174FF6ADC3A4D133D37" blockId="4.[151,1436,1865,2037]" box="[151,1307,1865,1893]" pageId="4" pageNumber="505">
<materialsCitation id="97AB3C9E43686174FF6ADC3A4D133D37" ID-GBIF-Occurrence="2608670280" box="[151,1307,1865,1893]" collectingDate="2014-06-22" collectorName="Ba Be NP" elevation="250" latitude="22.413" location="Bac Kan Prov." longLatPrecision="75" longitude="105.61" pageId="4" pageNumber="505" specimenCount="1" specimenCount-male="1" stateProvince="Bac Kan Prov." typeStatus="holotype">
<emphasis id="15B7EAD143686174FF6ADC3A49033D37" bold="true" box="[151,267,1866,1892]" pageId="4" pageNumber="505">
<typeStatus id="F878886143686174FF6ADC3A490E3D37" box="[151,262,1866,1892]" pageId="4" pageNumber="505" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="31C5FD4A43686174FEEFDC3B49213D36" box="[274,297,1867,1893]" pageId="4" pageNumber="505" type="male"></specimenCount>
,
<collectorName id="8A36531543686174FEC8DC3B49AD3D36" box="[309,421,1867,1893]" pageId="4" pageNumber="505">Ba Be NP</collectorName>
,
<collectingRegion id="E507F82143686174FE4DDC3B4A5B3D36" box="[432,595,1867,1893]" country="Vietnam" name="Bac Kan" pageId="4" pageNumber="505">Bac Kan Prov.</collectingRegion>
(appr.
<geoCoordinate id="42F7500443686174FD5CDC3A4B0D3D37" box="[673,773,1866,1892]" degrees="22.413" direction="north" orientation="latitude" pageId="4" pageNumber="505" precision="55" value="22.413">22.413N</geoCoordinate>
,
<geoCoordinate id="42F7500443686174FCEDDC394B883D37" box="[784,896,1865,1892]" degrees="105.610" direction="east" orientation="longitude" pageId="4" pageNumber="505" precision="55" value="105.61">105.610E</geoCoordinate>
; alt.
<elevation id="ACEED1F043686174FC48DC394C2F3D36" box="[949,1063,1865,1893]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="505" unit="m" value="250.0">
<quantity id="E03B9B2643686174FC48DC394BF43D36" box="[949,1020,1865,1893]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="505" unit="m" value="250.0">250 m</quantity>
asl
</elevation>
.),
<date id="537D100343686174FBBDDC3A4CB33D37" box="[1088,1211,1866,1892]" pageId="4" pageNumber="505" value="2014-06-22">
<collectingDate id="4339E9EB43686174FBBDDC3A4CB33D37" box="[1088,1211,1866,1892]" pageId="4" pageNumber="505" value="2014-06-22">22 vi 2014</collectingDate>
</date>
, TK leg.
</materialsCitation>
</paragraph>
<paragraph id="277C36C343686174FF3ADC1E4AC53D83" blockId="4.[151,1436,1865,2037]" pageId="4" pageNumber="505">
<emphasis id="15B7EAD143686174FF3ADC1E494E3DDB" bold="true" box="[199,326,1902,1928]" pageId="4" pageNumber="505">
<typeStatus id="F878886143686174FF3ADC1E49493DDB" box="[199,321,1902,1928]" pageId="4" pageNumber="505" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="31C5FD4A43686174FEADDC1E49773DDA" box="[336,383,1902,1929]" pageId="4" pageNumber="505" type="male">1 ♂</specimenCount>
, same location and collector,
<date id="537D100343686174FD26DC1D4B6C3DDB" box="[731,868,1901,1928]" pageId="4" pageNumber="505" value="2013-07-25">25 vii 2013</date>
;
<specimenCount id="31C5FD4A43686174FC8EDC1E4BAB3DDA" box="[883,931,1902,1929]" pageId="4" pageNumber="505" type="male">1 ♂</specimenCount>
, Ba Be NP,
<collectingRegion id="E507F82143686174FBCBDC1F4CE83DDA" box="[1078,1248,1903,1929]" country="Vietnam" name="Bac Kan" pageId="4" pageNumber="505">Bac Kan Prov.</collectingRegion>
(appr.
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,
<geoCoordinate id="42F7500443686174FF6ADCE149003DFF" box="[151,264,1937,1964]" degrees="105.650" direction="east" orientation="longitude" pageId="4" pageNumber="505" precision="55" value="105.65">105.650E</geoCoordinate>
),
<date id="537D100343686174FEE1DCE2499B3DFF" box="[284,403,1938,1964]" pageId="4" pageNumber="505" value="2017-07-02">2 vii 2017</date>
, TK leg.;
<specimenCount id="31C5FD4A43686174FDF8DCE24A403DFE" box="[517,584,1938,1965]" pageId="4" pageNumber="505" type="male">2 ♂♂</specimenCount>
, Xuan Son NP,
<collectingRegion id="E507F82143686174FD00DCE24BA93DFE" box="[765,929,1938,1965]" country="Vietnam" name="Phu Tho" pageId="4" pageNumber="505">Phu Tho Prov.</collectingRegion>
(appr.
<geoCoordinate id="42F7500443686174FC0CDCE24C5D3DFF" box="[1009,1109,1938,1964]" degrees="21.141" direction="north" orientation="latitude" pageId="4" pageNumber="505" precision="55" value="21.141">21.141N</geoCoordinate>
,
<geoCoordinate id="42F7500443686174FB9FDCE24CDA3DFF" box="[1122,1234,1938,1964]" degrees="104.932" direction="east" orientation="longitude" pageId="4" pageNumber="505" precision="55" value="104.932">104.932E</geoCoordinate>
),
<date id="537D100343686174FB1BDCE24D633DFF" box="[1254,1387,1938,1964]" pageId="4" pageNumber="505" value="2017-07-23">23 vii 2017</date>
, TK leg.;
<specimenCount id="31C5FD4A43686174FF32DCC649183D82" box="[207,272,1974,2001]" pageId="4" pageNumber="505" type="male">2 ♂♂</specimenCount>
, same location and collector,
<date id="537D100343686174FDA1DCC64AC23D83" box="[604,714,1974,2000]" pageId="4" pageNumber="505" value="2018-06-09">9 vi 2018</date>
.
</paragraph>
</subSubSection>
<subSubSection id="6FD9654843686175FF3ADCAA4A553AA8" lastPageId="5" lastPageNumber="506" pageId="4" pageNumber="505" type="etymology">
<paragraph id="277C36C343686175FF3ADCAA4A553AA8" blockId="4.[151,1436,1865,2037]" lastBlockId="5.[151,1437,153,2015]" lastPageId="5" lastPageNumber="506" pageId="4" pageNumber="505">
<emphasis id="15B7EAD143686174FF3ADCAA49403DA7" bold="true" box="[199,328,2010,2036]" pageId="4" pageNumber="505">Etymology</emphasis>
. The term “
<emphasis id="15B7EAD143686174FE30DCAB4A293DA6" box="[461,545,2011,2037]" italics="true" pageId="4" pageNumber="505">Cụ Rùa</emphasis>
” (great grandfather turtle) was the name used by
<collectingRegion id="E507F82143686174FBBADCAA4C833DA7" box="[1095,1163,2010,2036]" country="Vietnam" name="Ha Noi" pageId="4" pageNumber="505">Hanoi</collectingRegion>
locals to refer to the last Giant Softshell Turtle
<taxonomicName id="E0C34D4043696175FE6DDBE94AEB3AE0" authority="(Gray, 1873)" baseAuthorityName="Gray" baseAuthorityYear="1873" box="[400,739,153,179]" class="Reptilia" family="Trionychidae" genus="Rafetus" kingdom="Animalia" order="Testudines" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="swinhoei">
<emphasis id="15B7EAD143696175FE6DDBE94A463AE0" box="[400,590,153,179]" italics="true" pageId="5" pageNumber="506">Rafetus swinhoei</emphasis>
(Gray, 1873)
</taxonomicName>
alive in the city. The last representative of the species in Hoan Kiem Lake was found dead on
<date id="537D100343696175FE00DBCD4ACB3A84" box="[509,707,189,215]" pageId="5" pageNumber="506" value="2016-01-19">January 19, 2016</date>
. Its demise serves as a stark reminder of the many threats to the Vietnamese fauna. A noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="6FD9654843696175FF3ADA7449AA3CC8" pageId="5" pageNumber="506" type="description">
<paragraph id="277C36C343696175FF3ADA74496A3848" blockId="5.[151,1437,153,2015]" pageId="5" pageNumber="506">
<emphasis id="15B7EAD143696175FF3ADA7449D53B4C" bold="true" box="[199,477,260,287]" pageId="5" pageNumber="506">
Description of
<typeStatus id="F878886143696175FE8BDA7549D53B4C" box="[374,477,261,287]" pageId="5" pageNumber="506" type="holotype">holotype</typeStatus>
</emphasis>
(
<figureCitation id="BFF82A4643696175FE16DA754A493B4C" box="[491,577,261,287]" captionStart="FIGURE 1" captionStartId="3.[151,250,1981,2006]" captionTargetBox="[151,1436,993,1956]" captionTargetId="figure@3.[151,1436,993,1956]" captionTargetPageId="3" captionText="FIGURE 1. Head of male Coeliccia spp. in dorsolateral view. (a) C. caerulea sp. nov., holotype; (b) C. coronata sp. nov., holotype; (c) C. diomedea sp. nov., holotype; (d) C. curua sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.3765654" httpUri="https://zenodo.org/record/3765654/files/figure.png" pageId="5" pageNumber="506">Figs 1d</figureCitation>
,
<figureCitation id="BFF82A4643696175FDB0DA754A633B4C" box="[589,619,261,287]" captionStart="FIGURE 2" captionStartId="4.[151,250,1615,1640]" captionTargetBox="[151,1436,633,1590]" captionTargetId="figure@4.[151,1436,633,1590]" captionTargetPageId="4" captionText="FIGURE 2. Prothorax of male Coeliccia spp. in lateral view. (a) C. caerulea, holotype; (b) C. coronata, holotype; (c) C. diomedea, holotype; (d) C. curua, holotype." figureDoi="http://doi.org/10.5281/zenodo.3765656" httpUri="https://zenodo.org/record/3765656/files/figure.png" pageId="5" pageNumber="506">2d</figureCitation>
,
<figureCitation id="BFF82A4643696175FD8ADA754A9C3B4C" box="[631,660,261,287]" captionStart="FIGURE 3" captionStartId="6.[151,250,1882,1907]" captionTargetBox="[151,1436,417,1857]" captionTargetId="figure@6.[151,1436,417,1857]" captionTargetPageId="6" captionText="FIGURE 3. Prothorax of Coeliccia spp. in dorsal view. [a-e] male, [f] female. (a) C. caerulea, holotype; (b) C. coronata, holotype; (c) C. diomedea, holotype; (d) C. curua, holotype; (e) C. pulchella sp. nov., holotype; (f) C. caerulea, paratype used for description." figureDoi="http://doi.org/10.5281/zenodo.3765658" httpUri="https://zenodo.org/record/3765658/files/figure.png" pageId="5" pageNumber="506">3d</figureCitation>
,
<figureCitation id="BFF82A4643696175FD5DDA754AB63B4C" box="[672,702,261,287]" captionStart="FIGURE 4" captionStartId="7.[151,250,1090,1115]" captionTargetBox="[151,1437,181,1063]" captionTargetId="figure@7.[151,1437,181,1064]" captionTargetPageId="7" captionText="FIGURE 4. Thorax of male Coeliccia spp. in lateral view. (a) C. caerulea, holotype; (b) C. coronata, holotype; (c) C. diomedea, holotype; (d) C. curua, holotype." figureDoi="http://doi.org/10.5281/zenodo.3765660" httpUri="https://zenodo.org/record/3765660/files/figure.png" pageId="5" pageNumber="506">4d</figureCitation>
,
<figureCitation id="BFF82A4643696175FD37DA744AE03B4C" box="[714,744,260,287]" captionStart="FIGURE 5" captionStartId="8.[151,250,910,935]" captionTargetBox="[151,1436,181,885]" captionTargetId="figure@8.[151,1436,181,885]" captionTargetPageId="8" captionText="FIGURE 5. Thorax of male Coeliccia spp. in dorsal view. (a) C. caerulea, holotype; (b) C. coronata, holotype; (c) C. diomedea, holotype; (d) C. curua, holotype." figureDoi="http://doi.org/10.5281/zenodo.3765662" httpUri="https://zenodo.org/record/3765662/files/figure.png" pageId="5" pageNumber="506">5d</figureCitation>
,
<figureCitation id="BFF82A4643696175FD0EDA754B083B4C" box="[755,768,261,287]" captionStart="FIGURE 7" captionStartId="9.[151,250,1813,1838]" captionTargetBox="[151,1436,181,1786]" captionTargetId="figure@9.[151,1436,181,1787]" captionTargetPageId="9" captionText="FIGURE 7. Abdomen tip of male Coeliccia spp. [a,e,g,i] lateral view, [c] dorsolateral view, [b,d,f,h,j] dorsal view. (a,b) C. caerulea, holotype; (c,d) C. pulchella, holotype; (e,f) C. curua, holotype; (g,h) C. coronata, holotype; (i,j) C. diomedea, holotype." figureDoi="http://doi.org/10.5281/zenodo.3765666" httpUri="https://zenodo.org/record/3765666/files/figure.png" pageId="5" pageNumber="506">7</figureCitation>
ef, 10ab, 12ef, 25cd).
<emphasis id="15B7EAD143696175FBD0DA754C623B4C" box="[1069,1130,261,287]" italics="true" pageId="5" pageNumber="506">Head</emphasis>
(
<figureCitation id="BFF82A4643696175FB84DA754CC23B73" box="[1145,1226,261,288]" captionStart="FIGURE 1" captionStartId="3.[151,250,1981,2006]" captionTargetBox="[151,1436,993,1956]" captionTargetId="figure@3.[151,1436,993,1956]" captionTargetPageId="3" captionText="FIGURE 1. Head of male Coeliccia spp. in dorsolateral view. (a) C. caerulea sp. nov., holotype; (b) C. coronata sp. nov., holotype; (c) C. diomedea sp. nov., holotype; (d) C. curua sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.3765654" httpUri="https://zenodo.org/record/3765654/files/figure.png" pageId="5" pageNumber="506">Fig. 1d</figureCitation>
). Labrum blackish brown, chestnut brown at free margin and basally. Anteclypeus white, with two darker areas left and right on lower half. Postclypeus shiny black. Base of mandibles black, anteroventral corner white. Genae white, the white narrowly extending over the antefrons above the postclypeus to about one-fourth in, remainder of frons matte black. Dorsal surface of head matte black, apart from two small bluish white spots between base of antennae and lateral ocelli and two oblong bluish white postocular spots. Antennae black, top of first segment white. Compound eyes three-colored in life, black or very dark chestnut above, green below, but posterior border between green lower half and blackish upper half is blue.
</paragraph>
<paragraph id="277C36C343696175FF3AD9554A64390C" blockId="5.[151,1437,153,2015]" pageId="5" pageNumber="506">
<emphasis id="15B7EAD143696175FF3AD9554928386C" box="[199,288,549,575]" italics="true" pageId="5" pageNumber="506">Thorax.</emphasis>
Prothorax (
<figureCitation id="BFF82A4643696175FE5ED95549F1386C" box="[419,505,549,575]" captionStart="FIGURE 2" captionStartId="4.[151,250,1615,1640]" captionTargetBox="[151,1436,633,1590]" captionTargetId="figure@4.[151,1436,633,1590]" captionTargetPageId="4" captionText="FIGURE 2. Prothorax of male Coeliccia spp. in lateral view. (a) C. caerulea, holotype; (b) C. coronata, holotype; (c) C. diomedea, holotype; (d) C. curua, holotype." figureDoi="http://doi.org/10.5281/zenodo.3765656" httpUri="https://zenodo.org/record/3765656/files/figure.png" pageId="5" pageNumber="506">Figs 2d</figureCitation>
,
<figureCitation id="BFF82A4643696175FDF8D9554A2C3813" box="[517,548,549,576]" captionStart="FIGURE 3" captionStartId="6.[151,250,1882,1907]" captionTargetBox="[151,1436,417,1857]" captionTargetId="figure@6.[151,1436,417,1857]" captionTargetPageId="6" captionText="FIGURE 3. Prothorax of Coeliccia spp. in dorsal view. [a-e] male, [f] female. (a) C. caerulea, holotype; (b) C. coronata, holotype; (c) C. diomedea, holotype; (d) C. curua, holotype; (e) C. pulchella sp. nov., holotype; (f) C. caerulea, paratype used for description." figureDoi="http://doi.org/10.5281/zenodo.3765658" httpUri="https://zenodo.org/record/3765658/files/figure.png" pageId="5" pageNumber="506">3d</figureCitation>
): matte black, propleuron dark brown with pale edges. Synthorax (
<figureCitation id="BFF82A4643696175FAF0D9554D6C386C" box="[1293,1380,549,575]" captionStart="FIGURE 4" captionStartId="7.[151,250,1090,1115]" captionTargetBox="[151,1437,181,1063]" captionTargetId="figure@7.[151,1437,181,1064]" captionTargetPageId="7" captionText="FIGURE 4. Thorax of male Coeliccia spp. in lateral view. (a) C. caerulea, holotype; (b) C. coronata, holotype; (c) C. diomedea, holotype; (d) C. curua, holotype." figureDoi="http://doi.org/10.5281/zenodo.3765660" httpUri="https://zenodo.org/record/3765660/files/figure.png" pageId="5" pageNumber="506">Figs 4d</figureCitation>
,
<figureCitation id="BFF82A4643696175FA92D9544D87386C" box="[1391,1423,548,575]" captionStart="FIGURE 5" captionStartId="8.[151,250,910,935]" captionTargetBox="[151,1436,181,885]" captionTargetId="figure@8.[151,1436,181,885]" captionTargetPageId="8" captionText="FIGURE 5. Thorax of male Coeliccia spp. in dorsal view. (a) C. caerulea, holotype; (b) C. coronata, holotype; (c) C. diomedea, holotype; (d) C. curua, holotype." figureDoi="http://doi.org/10.5281/zenodo.3765662" httpUri="https://zenodo.org/record/3765662/files/figure.png" pageId="5" pageNumber="506">5d</figureCitation>
): mesepisternum black, with a large oval pale blue antehumeral shield, the inner edge of which is somewhat irregular and the outer edge of which follows the mesopleural suture. Extent of this blue shield is about two-thirds of the width, and three-fifths of length of mesepisternum. Mesepimeron and mesinfraepisternum black. Metepisternum black with large rectangular pale blue area, which forward reaches proximity of mesocoxa. This blue area with a small rounded excision along its upper margin near the wing base, and with black margin along wing base connecting the black of mesepimeron to the well-defined black line over metapleural suture. Metepimeron bluish white with brown, irregular smudge near coxae, metinfraepisternum black. Legs: coxae and trochanters white. Femur grayish with black stripes. Spines and claws dark.
</paragraph>
<paragraph id="277C36C343696175FF3AD81A4C1D39D0" blockId="5.[151,1437,153,2015]" box="[199,1045,872,899]" pageId="5" pageNumber="506">
<emphasis id="15B7EAD143696175FF3AD81A491B39D0" box="[199,275,874,899]" italics="true" pageId="5" pageNumber="506">Wings.</emphasis>
Hyaline, 25 Px in FW,
<quantity id="E03B9B2643696175FDE7D8194A5B39D0" box="[538,595,873,899]" metricMagnitude="-1" metricUnit="m" metricValue="5.842" pageId="5" pageNumber="506" unit="in" value="23.0">23 in</quantity>
HW. Pt dark brown, covering two cells.
</paragraph>
<paragraph id="277C36C343696175FF3AD8FD4BFF39BC" blockId="5.[151,1437,153,2015]" pageId="5" pageNumber="506">
<emphasis id="15B7EAD143696175FF3AD8FD493139F4" box="[199,313,909,935]" italics="true" pageId="5" pageNumber="506">Abdomen.</emphasis>
S1 narrowly black dorsally, remainder bluish white; S2 upper half black and lower half white; S36 black above and white below, and with prominent subapical white annulus; S7 virtually black, but with large subapical white spot; S8 black with posteroventral corner yellow; S910 all yellow.
</paragraph>
<paragraph id="277C36C343696175FF3AD8894B243EF0" blockId="5.[151,1437,153,2015]" pageId="5" pageNumber="506">
<emphasis id="15B7EAD143696175FF3AD88949813E40" box="[199,393,1017,1043]" italics="true" pageId="5" pageNumber="506">Anal appendages</emphasis>
(
<figureCitation id="BFF82A4643696175FE65D88949E03E47" box="[408,488,1017,1044]" captionStart="FIGURE 10" captionStartId="12.[151,250,1788,1813]" captionTargetBox="[151,1436,345,1763]" captionTargetId="figure@12.[151,1436,345,1763]" captionTargetPageId="12" captionText="FIGURE 10. S10 and caudal appendages of male Coeliccia spp. [a,c,e] lateral view, [b,d,f] dorsal view. (a,b) C. curua, holotype; (c,d) C. curua, paratype, 2 vii 2017; (e,f) C.uenoi, Xuan Son NP, Phu Tho Prov., 9 vi 2018." figureDoi="http://doi.org/10.5281/zenodo.3765672" httpUri="https://zenodo.org/record/3765672/files/figure.png" pageId="5" pageNumber="506">Fig. 10</figureCitation>
ab). Yellow (Color in
<figureCitation id="BFF82A4643696175FD14D8894B3E3E40" box="[745,822,1017,1043]" captionStart="FIGURE 10" captionStartId="12.[151,250,1788,1813]" captionTargetBox="[151,1436,345,1763]" captionTargetId="figure@12.[151,1436,345,1763]" captionTargetPageId="12" captionText="FIGURE 10. S10 and caudal appendages of male Coeliccia spp. [a,c,e] lateral view, [b,d,f] dorsal view. (a,b) C. curua, holotype; (c,d) C. curua, paratype, 2 vii 2017; (e,f) C.uenoi, Xuan Son NP, Phu Tho Prov., 9 vi 2018." figureDoi="http://doi.org/10.5281/zenodo.3765672" httpUri="https://zenodo.org/record/3765672/files/figure.png" pageId="5" pageNumber="506">Fig. 10</figureCitation>
ab result of post-mortem changes). In lateral view the cerci twice as thick and a little shorter compared to paraprocts, apex curving downward, forming a small but distinct drooping point, and with subapical ventral expansion, tipped with black tooth. In dorsal view a small tooth at onethird of length of cerci, from where a dorsal ridge and the subapical ventral expansion start. The paraprocts of usual
<typeStatus id="F878886143696175FF6ADFF948C23EF0" box="[151,202,1161,1187]" pageId="5" pageNumber="506">type</typeStatus>
, apically curved inwards and with black tooth at apex.
</paragraph>
<paragraph id="277C36C343696175FF3ADFDD49E83F43" blockId="5.[151,1437,153,2015]" pageId="5" pageNumber="506">
<emphasis id="15B7EAD143696175FF3ADFDD496B3E94" box="[199,355,1197,1223]" italics="true" pageId="5" pageNumber="506">Genital ligula</emphasis>
(
<figureCitation id="BFF82A4643696175FE8EDFDD49CA3E9B" box="[371,450,1197,1224]" captionStart="FIGURE 12" captionStartId="17.[151,250,1512,1537]" captionTargetBox="[171,1416,181,1487]" captionTargetId="figure@17.[170,1416,181,1487]" captionTargetPageId="17" captionText="FIGURE 12. Genital ligula of Coeliccia curua. [a,c,e] lateral view, [b,d,f] ventral view, [a-c] Xuan Son NP, [d-f] Ba Vi NP. (a,b) paratype, 9 vi 2018; (c) second paratype, 9 vi 2018; (d) paratype, 25 vii 2013; (e-f) holotype." figureDoi="http://doi.org/10.5281/zenodo.3765676" httpUri="https://zenodo.org/record/3765676/files/figure.png" pageId="5" pageNumber="506">Fig. 12</figureCitation>
ef). The apical segment with apical flap and two robust flagella that curve back to base of apical segment and from there extend a little sideways, appearing to taper to a point (but actually ribbon-like and curled up under microscope).
</paragraph>
<paragraph id="277C36C343696175FF3ADE684D9F3C7C" blockId="5.[151,1437,153,2015]" pageId="5" pageNumber="506">
<emphasis id="15B7EAD143696175FF3ADE6849D83F60" bold="true" box="[199,464,1304,1331]" pageId="5" pageNumber="506">
Variation in
<typeStatus id="F878886143696175FEA7DE6949D83F60" box="[346,464,1305,1331]" pageId="5" pageNumber="506" type="paratype">paratypes</typeStatus>
</emphasis>
(
<figureCitation id="BFF82A4643696175FE23DE694A2E3F60" box="[478,550,1305,1331]" captionStart="FIGURE 9" captionStartId="11.[151,250,1555,1580]" captionTargetBox="[151,1436,181,1529]" captionTargetId="figure@11.[151,1436,181,1529]" captionTargetPageId="11" captionText="FIGURE 9. S10 and caudal appendages of male Coeliccia curua from Xuan Son NP. [a,c,e] lateral view, [b,d,f] dorsal view. (a,b) paratype, 23 vii 2017; (c,d) paratype, 9 vi 2018; (e,f) second paratype, 23 vii 2017." figureDoi="http://doi.org/10.5281/zenodo.3765670" httpUri="https://zenodo.org/record/3765670/files/figure.png" pageId="5" pageNumber="506">Figs 9</figureCitation>
,
<figureCitation id="BFF82A4643696175FDCDDE694A433F60" box="[560,587,1305,1331]" captionStart="FIGURE 12" captionStartId="17.[151,250,1512,1537]" captionTargetBox="[171,1416,181,1487]" captionTargetId="figure@17.[170,1416,181,1487]" captionTargetPageId="17" captionText="FIGURE 12. Genital ligula of Coeliccia curua. [a,c,e] lateral view, [b,d,f] ventral view, [a-c] Xuan Son NP, [d-f] Ba Vi NP. (a,b) paratype, 9 vi 2018; (c) second paratype, 9 vi 2018; (d) paratype, 25 vii 2013; (e-f) holotype." figureDoi="http://doi.org/10.5281/zenodo.3765676" httpUri="https://zenodo.org/record/3765676/files/figure.png" pageId="5" pageNumber="506">12</figureCitation>
ac, 25ab). Propleuron with pale central area and less obvious pale edges or completely black. In Xuan Son specimens the antehumeral shields are somewhat larger (
<figureCitation id="BFF82A4643696175FB8CDE4D4CCA3F04" box="[1137,1218,1340,1367]" captionStart="FIGURE 25" captionStartId="30.[151,250,1870,1895]" captionTargetBox="[151,1436,561,1845]" captionTargetId="figure@30.[151,1436,561,1845]" captionTargetPageId="30" captionText="FIGURE 25. Thorax and head of alive male Coeliccia spp. [a-d] C. curua, [e,f] C.uenoi. (a,b) paratype in dorsolateral (a) and lateral (b) view, Xuan Son NP, Phu Tho Prov., 23 vii 2017; (c) holotype in dorsolateral view; (d) holotype in lateral view; (e) dorsolateral view, Xuan Son NP, Phu Tho Prov., 9 vi 2018; (f) lateral view, Cuc Phuong NP, Ninh Binh Prov., 3 v 2014." figureDoi="http://doi.org/10.5281/zenodo.3765702" httpUri="https://zenodo.org/record/3765702/files/figure.png" pageId="5" pageNumber="506">Fig. 25</figureCitation>
ab). Blue stripe on metepisternum without black excision along upper margin or continuing over spiracle almost to reach mesinfraepisternum. Metepimeron without brown smudge near coxae. S34 with more white on proximal part of venter, S9 with antero-ventral corner dark brown in one
<typeStatus id="F878886143696175FD73DED94AE63F90" box="[654,750,1449,1475]" pageId="5" pageNumber="506" type="paratype">paratype</typeStatus>
or S10 almost completely yellow. Anal appendages deformed in one mature specimen. In Xuan Son specimens the paraprocts appear more slender and a little longer relative to the cerci (
<figureCitation id="BFF82A4643696175FF23DE81492B3C58" box="[222,291,1521,1547]" captionStart="FIGURE 9" captionStartId="11.[151,250,1555,1580]" captionTargetBox="[151,1436,181,1529]" captionTargetId="figure@11.[151,1436,181,1529]" captionTargetPageId="11" captionText="FIGURE 9. S10 and caudal appendages of male Coeliccia curua from Xuan Son NP. [a,c,e] lateral view, [b,d,f] dorsal view. (a,b) paratype, 23 vii 2017; (c,d) paratype, 9 vi 2018; (e,f) second paratype, 23 vii 2017." figureDoi="http://doi.org/10.5281/zenodo.3765670" httpUri="https://zenodo.org/record/3765670/files/figure.png" pageId="5" pageNumber="506">Fig. 9</figureCitation>
). Apico-lateral flagella of Xuan Son specimens shorter, not extending as obviously laterally in apical third (
<figureCitation id="BFF82A4643696175FF26DD6549243C7C" box="[219,300,1557,1583]" captionStart="FIGURE 12" captionStartId="17.[151,250,1512,1537]" captionTargetBox="[171,1416,181,1487]" captionTargetId="figure@17.[170,1416,181,1487]" captionTargetPageId="17" captionText="FIGURE 12. Genital ligula of Coeliccia curua. [a,c,e] lateral view, [b,d,f] ventral view, [a-c] Xuan Son NP, [d-f] Ba Vi NP. (a,b) paratype, 9 vi 2018; (c) second paratype, 9 vi 2018; (d) paratype, 25 vii 2013; (e-f) holotype." figureDoi="http://doi.org/10.5281/zenodo.3765676" httpUri="https://zenodo.org/record/3765676/files/figure.png" pageId="5" pageNumber="506">Fig. 12</figureCitation>
ac). 24 Px in left FW. Pt covering 1.5 cells (Xuan Son). 1921 Px in FW,
<quantity id="E03B9B2643696175FB92DD654CDA3C7C" box="[1135,1234,1557,1583]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" metricValueMax="5.08" metricValueMin="4.064" pageId="5" pageNumber="506" unit="in" value="18.0" valueMax="20.0" valueMin="16.0">1620 in</quantity>
HW (Xuan Son).
</paragraph>
<paragraph id="277C36C343696175FF3ADD4949AE3C24" blockId="5.[151,1437,153,2015]" pageId="5" pageNumber="506">Measurements (in mm). Ba Be: HW 3335; abdomen incl. appendages 5152; Xuan Son: HW 2627; abdomen incl. appendages 3739.</paragraph>
<paragraph id="277C36C343696175FF3ADDF149AA3CC8" blockId="5.[151,1437,153,2015]" box="[199,418,1665,1691]" pageId="5" pageNumber="506">
<emphasis id="15B7EAD143696175FF3ADDF1492D3CC8" bold="true" box="[199,293,1665,1691]" pageId="5" pageNumber="506">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="6FD9654843696176FF3ADDD44C933A84" lastPageId="6" lastPageNumber="507" pageId="5" pageNumber="506" type="diagnosis">
<paragraph id="277C36C343696176FF3ADDD44C933A84" blockId="5.[151,1437,153,2015]" lastBlockId="6.[151,1437,153,395]" lastPageId="6" lastPageNumber="507" pageId="5" pageNumber="506">
<emphasis id="15B7EAD143696175FF3ADDD449C63CEC" bold="true" box="[199,462,1700,1727]" pageId="5" pageNumber="506">Differential diagnosis.</emphasis>
<taxonomicName id="E0C34D4043696175FE2BDDD54A813CEC" authorityName="Kompier &amp; Dow &amp; Steinhoff" authorityYear="2020" box="[470,649,1701,1727]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="curua">
<emphasis id="15B7EAD143696175FE2BDDD54A813CEC" box="[470,649,1701,1727]" italics="true" pageId="5" pageNumber="506">Coeliccia curua</emphasis>
</taxonomicName>
has a typical genital ligula of the
<taxonomicName id="E0C34D4043696175FBF4DDD54C763CEC" authorityName="Laidlaw" authorityYear="1932" box="[1033,1150,1701,1727]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="pyriformis">
<emphasis id="15B7EAD143696175FBF4DDD54C763CEC" box="[1033,1150,1701,1727]" italics="true" pageId="5" pageNumber="506">pyriformis</emphasis>
</taxonomicName>
-group. Within this group it shares the combination of large blue antehumeral shields, bold black lines over mesopleural and metapleural sutures and yellow abdominal tip with
<taxonomicName id="E0C34D4043696175FDCDDD9D4A863D54" authorityName="Asahina" authorityYear="1997" box="[560,654,1773,1799]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="uenoi">
<emphasis id="15B7EAD143696175FDCDDD9D4A863D54" box="[560,654,1773,1799]" italics="true" pageId="5" pageNumber="506">C. uenoi</emphasis>
</taxonomicName>
(
<figureCitation id="BFF82A4643696175FD60DD9D4AE43D54" box="[669,748,1772,1799]" captionStart="FIGURE 25" captionStartId="30.[151,250,1870,1895]" captionTargetBox="[151,1436,561,1845]" captionTargetId="figure@30.[151,1436,561,1845]" captionTargetPageId="30" captionText="FIGURE 25. Thorax and head of alive male Coeliccia spp. [a-d] C. curua, [e,f] C.uenoi. (a,b) paratype in dorsolateral (a) and lateral (b) view, Xuan Son NP, Phu Tho Prov., 23 vii 2017; (c) holotype in dorsolateral view; (d) holotype in lateral view; (e) dorsolateral view, Xuan Son NP, Phu Tho Prov., 9 vi 2018; (f) lateral view, Cuc Phuong NP, Ninh Binh Prov., 3 v 2014." figureDoi="http://doi.org/10.5281/zenodo.3765702" httpUri="https://zenodo.org/record/3765702/files/figure.png" pageId="5" pageNumber="506">Fig. 25</figureCitation>
ef, 29ab) and
<taxonomicName id="E0C34D4043696175FC5FDD9D4C2C3D54" authorityName="Phan &amp; Tran" authorityYear="2018" box="[930,1060,1773,1799]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="phamiha">
<emphasis id="15B7EAD143696175FC5FDD9D4C2C3D54" box="[930,1060,1773,1799]" italics="true" pageId="5" pageNumber="506">C. phamiha</emphasis>
</taxonomicName>
. In
<taxonomicName id="E0C34D4043696175FBADDD9D4CDA3D54" authorityName="Phan &amp; Tran" authorityYear="2018" box="[1104,1234,1773,1799]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="phamiha">
<emphasis id="15B7EAD143696175FBADDD9D4CDA3D54" box="[1104,1234,1773,1799]" italics="true" pageId="5" pageNumber="506">C. phamiha</emphasis>
</taxonomicName>
the flagella of the genital ligula are directed laterally and posteriorly, before curving inward apically (
<bibRefCitation id="43524B3243696175FBC1DC614D183D78" author="Phan, Q. T. &amp; Tran, T. T. T." box="[1084,1296,1809,1835]" pageId="5" pageNumber="506" pagination="45 - 53" refId="ref19287" refString="Phan, Q. T. &amp; Tran, T. T. T. (2018) Description of Coeliccia phamiha sp. n. from central Vietnam (Odonata: Platycnemididae). International Journal of Odonatology, 21 (1), 45 - 53. https: // doi. org / 10.1080 / 13887890.2017.1419885" type="journal article" year="2018">Phan &amp; Tran 2018</bibRefCitation>
), whereas in
<taxonomicName id="E0C34D4043696175FF6ADC4548FE3D1C" authorityName="Asahina" authorityYear="1997" box="[151,246,1845,1871]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="uenoi">
<emphasis id="15B7EAD143696175FF6ADC4548FE3D1C" box="[151,246,1845,1871]" italics="true" pageId="5" pageNumber="506">C. uenoi</emphasis>
</taxonomicName>
they are robust and directed first laterally and then anteriorly (
<figureCitation id="BFF82A4643696175FC49DC454C0F3D1C" box="[948,1031,1845,1871]" captionStart="FIGURE 13" captionStartId="18.[151,250,1619,1644]" captionTargetBox="[151,1436,633,1594]" captionTargetId="figure@18.[151,1436,633,1594]" captionTargetPageId="18" captionText="FIGURE 13. Genital ligula of Coeliccia uenoi. [a,c] ventral view, [b,d] lateral view. (a,b) Xuan Son NP, Phu Tho Prov., 9 vi 2018; (c,d) Cuc Phuong NP, Ninh Binh Prov., 18 ix 2016." figureDoi="http://doi.org/10.5281/zenodo.3765678" httpUri="https://zenodo.org/record/3765678/files/figure.png" pageId="5" pageNumber="506">Fig. 13</figureCitation>
). In
<taxonomicName id="E0C34D4043696175FBC7DC454C943D1C" authorityName="Kompier &amp; Dow &amp; Steinhoff" authorityYear="2020" box="[1082,1180,1845,1871]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="curua">
<emphasis id="15B7EAD143696175FBC7DC454C943D1C" box="[1082,1180,1845,1871]" italics="true" pageId="5" pageNumber="506">C. curua</emphasis>
</taxonomicName>
the flagella are curved posteriorly, hidden in ventral view under terminal segment, but apical third directed laterally and visible in ventral view (although limited so or not at all in Xuan Son specimens) (
<figureCitation id="BFF82A4643696175FCA4DC0D4BA33DC4" box="[857,939,1917,1943]" captionStart="FIGURE 12" captionStartId="17.[151,250,1512,1537]" captionTargetBox="[171,1416,181,1487]" captionTargetId="figure@17.[170,1416,181,1487]" captionTargetPageId="17" captionText="FIGURE 12. Genital ligula of Coeliccia curua. [a,c,e] lateral view, [b,d,f] ventral view, [a-c] Xuan Son NP, [d-f] Ba Vi NP. (a,b) paratype, 9 vi 2018; (c) second paratype, 9 vi 2018; (d) paratype, 25 vii 2013; (e-f) holotype." figureDoi="http://doi.org/10.5281/zenodo.3765676" httpUri="https://zenodo.org/record/3765676/files/figure.png" pageId="5" pageNumber="506">Fig. 12</figureCitation>
). The antehumeral shields of
<taxonomicName id="E0C34D4043696175FB0CDC0D4D473DC4" authorityName="Asahina" authorityYear="1997" box="[1265,1359,1917,1943]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="uenoi">
<emphasis id="15B7EAD143696175FB0CDC0D4D473DC4" box="[1265,1359,1917,1943]" italics="true" pageId="5" pageNumber="506">C. uenoi</emphasis>
</taxonomicName>
have a characteristic step on the inner margin and are not oval (
<figureCitation id="BFF82A4643696175FCF9DCD14B5E3DE8" box="[772,854,1952,1979]" captionStart="FIGURE 25" captionStartId="30.[151,250,1870,1895]" captionTargetBox="[151,1436,561,1845]" captionTargetId="figure@30.[151,1436,561,1845]" captionTargetPageId="30" captionText="FIGURE 25. Thorax and head of alive male Coeliccia spp. [a-d] C. curua, [e,f] C.uenoi. (a,b) paratype in dorsolateral (a) and lateral (b) view, Xuan Son NP, Phu Tho Prov., 23 vii 2017; (c) holotype in dorsolateral view; (d) holotype in lateral view; (e) dorsolateral view, Xuan Son NP, Phu Tho Prov., 9 vi 2018; (f) lateral view, Cuc Phuong NP, Ninh Binh Prov., 3 v 2014." figureDoi="http://doi.org/10.5281/zenodo.3765702" httpUri="https://zenodo.org/record/3765702/files/figure.png" pageId="5" pageNumber="506">Figs 25</figureCitation>
ef, 29ab), and the black lines over the metapleural and mesopleural sutures are narrower.
<taxonomicName id="E0C34D4043696175FDB2DCB54B2D3D8C" authorityName="Phan &amp; Tran" authorityYear="2018" box="[591,805,1989,2015]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="phamiha">
<emphasis id="15B7EAD143696175FDB2DCB54B2D3D8C" box="[591,805,1989,2015]" italics="true" pageId="5" pageNumber="506">Coeliccia phamiha</emphasis>
</taxonomicName>
has even larger antehumeral shields that obviously and suddenly narrow at their center, and has black S910, besides structurally different and whitish cerci (
<bibRefCitation id="43524B32436A6176FAF6DBE948DC3A84" author="Phan, Q. T. &amp; Tran, T. T. T." pageId="6" pageNumber="507" pagination="45 - 53" refId="ref19287" refString="Phan, Q. T. &amp; Tran, T. T. T. (2018) Description of Coeliccia phamiha sp. n. from central Vietnam (Odonata: Platycnemididae). International Journal of Odonatology, 21 (1), 45 - 53. https: // doi. org / 10.1080 / 13887890.2017.1419885" type="journal article" year="2018">Phan &amp; Tran 2018</bibRefCitation>
). The caudal appendages of
<taxonomicName id="E0C34D40436A6176FDECDBCD4A7A3A84" authorityName="Kompier &amp; Dow &amp; Steinhoff" authorityYear="2020" box="[529,626,189,215]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="507" phylum="Arthropoda" rank="species" species="curua">
<emphasis id="15B7EAD1436A6176FDECDBCD4A7A3A84" box="[529,626,189,215]" italics="true" pageId="6" pageNumber="507">C. curua</emphasis>
</taxonomicName>
and
<taxonomicName id="E0C34D40436A6176FD54DBCD4B003A84" authorityName="Asahina" authorityYear="1997" box="[681,776,189,215]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="507" phylum="Arthropoda" rank="species" species="uenoi">
<emphasis id="15B7EAD1436A6176FD54DBCD4B003A84" box="[681,776,189,215]" italics="true" pageId="6" pageNumber="507">C. uenoi</emphasis>
</taxonomicName>
are similar in structure (
<figureCitation id="BFF82A46436A6176FBE7DBCD4C6B3A84" box="[1050,1123,189,215]" captionStart="FIGURE 9" captionStartId="11.[151,250,1555,1580]" captionTargetBox="[151,1436,181,1529]" captionTargetId="figure@11.[151,1436,181,1529]" captionTargetPageId="11" captionText="FIGURE 9. S10 and caudal appendages of male Coeliccia curua from Xuan Son NP. [a,c,e] lateral view, [b,d,f] dorsal view. (a,b) paratype, 23 vii 2017; (c,d) paratype, 9 vi 2018; (e,f) second paratype, 23 vii 2017." figureDoi="http://doi.org/10.5281/zenodo.3765670" httpUri="https://zenodo.org/record/3765670/files/figure.png" pageId="6" pageNumber="507">Figs 9</figureCitation>
,
<figureCitation id="BFF82A46436A6176FB92DBCD4C863A84" box="[1135,1166,189,215]" captionStart="FIGURE 10" captionStartId="12.[151,250,1788,1813]" captionTargetBox="[151,1436,345,1763]" captionTargetId="figure@12.[151,1436,345,1763]" captionTargetPageId="12" captionText="FIGURE 10. S10 and caudal appendages of male Coeliccia spp. [a,c,e] lateral view, [b,d,f] dorsal view. (a,b) C. curua, holotype; (c,d) C. curua, paratype, 2 vii 2017; (e,f) C.uenoi, Xuan Son NP, Phu Tho Prov., 9 vi 2018." figureDoi="http://doi.org/10.5281/zenodo.3765672" httpUri="https://zenodo.org/record/3765672/files/figure.png" pageId="6" pageNumber="507">10</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="6FD96548436A6176FF3ADB904B6B3BD8" pageId="6" pageNumber="507" type="biology_ecology">
<paragraph id="277C36C3436A6176FF3ADB904B6B3BD8" blockId="6.[151,1437,153,395]" pageId="6" pageNumber="507">
<emphasis id="15B7EAD1436A6176FF3ADB9049B23AA8" bold="true" box="[199,442,224,251]" pageId="6" pageNumber="507">Habitat and ecology.</emphasis>
In Ba Be NP
<taxonomicName id="E0C34D40436A6176FDACDB914B0B3AA8" authority="Kompier &amp; Dow &amp; Steinhoff, 2020" authorityName="Kompier &amp; Dow &amp; Steinhoff" authorityYear="2020" box="[593,771,225,251]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="507" phylum="Arthropoda" rank="species" species="curua" status="sp. nov.">
<emphasis id="15B7EAD1436A6176FDACDB914B0B3AA8" box="[593,771,225,251]" italics="true" pageId="6" pageNumber="507">Coeliccia curua</emphasis>
</taxonomicName>
<emphasis id="15B7EAD1436A6176FCF4DB914B6A3AA8" bold="true" box="[777,866,225,251]" pageId="6" pageNumber="507">
<taxonomicNameLabel id="0E8457AA436A6176FCF4DB914B6A3AA8" box="[777,866,225,251]" pageId="6" pageNumber="507" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was found at two different locations in karst mountains with seeps over rocks and forest floor, where it occurred with
<taxonomicName id="E0C34D40436A6176FC67DA754C393B4C" authorityName="Laidlaw" authorityYear="1932" box="[922,1073,261,287]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="507" phylum="Arthropoda" rank="species" species="pyriformis">
<emphasis id="15B7EAD1436A6176FC67DA754C393B4C" box="[922,1073,261,287]" italics="true" pageId="6" pageNumber="507">C. pyriformis</emphasis>
</taxonomicName>
and
<taxonomicName id="E0C34D40436A6176FB97DA7648DB3B10" authority="Kompier, 2016" authorityName="Kompier" authorityYear="2016" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="507" phylum="Arthropoda" rank="species" species="nigra">
<emphasis id="15B7EAD1436A6176FB97DA764D233B4C" box="[1130,1323,262,287]" italics="true" pageId="6" pageNumber="507">Protosticta nigra</emphasis>
Kompier, 2016
</taxonomicName>
. In Xuan Son NP it was found at two small and very shallow streams at appr.
<quantity id="E03B9B26436A6176FBBEDA594C8B3B10" box="[1091,1155,297,323]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="507" unit="m" value="900.0">900m</quantity>
asl. in degraded primary forest, similarly on karst mountains. Xuan Son NP boasts at least 7 other
<taxonomicName id="E0C34D40436A6176FC38DA3D4C383B34" authorityName="Kirby" authorityYear="1890" box="[965,1072,333,359]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="507" phylum="Arthropoda" rank="genus">
<emphasis id="15B7EAD1436A6176FC38DA3D4C383B34" box="[965,1072,333,359]" italics="true" pageId="6" pageNumber="507">Coeliccia</emphasis>
</taxonomicName>
species, including the similar
<emphasis id="15B7EAD1436A6176FA7EDA3D48D43BD8" italics="true" pageId="6" pageNumber="507">
<taxonomicName id="E0C34D40436A6176FA7EDA3D48D03BD8" authorityName="Asahina" authorityYear="1997" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="507" phylum="Arthropoda" rank="species" species="uenoi">C. uenoi</taxonomicName>
,
</emphasis>
but only
<taxonomicName id="E0C34D40436A6176FEBADA0149D53BD8" authorityName="Laidlaw" authorityYear="1932" box="[327,477,369,395]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="507" phylum="Arthropoda" rank="species" species="pyriformis">
<emphasis id="15B7EAD1436A6176FEBADA0149D53BD8" box="[327,477,369,395]" italics="true" pageId="6" pageNumber="507">C. pyriformis</emphasis>
</taxonomicName>
was found together with
<taxonomicName id="E0C34D40436A6176FD07DA014B533BD8" authorityName="Kompier &amp; Dow &amp; Steinhoff" authorityYear="2020" box="[762,859,369,395]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="507" phylum="Arthropoda" rank="species" species="curua">
<emphasis id="15B7EAD1436A6176FD07DA014B533BD8" box="[762,859,369,395]" italics="true" pageId="6" pageNumber="507">C. curua</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>