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<mods:title id="F590C7AA91B87DEB66B931F3D540EFA0">Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)</mods:title>
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<paragraph id="C40A82035DE7AEEB6875B50C72C57FF1" pageId="240" pageNumber="241">Taxon classification Animalia Coleoptera Dytiscidae</paragraph>
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<taxonomicName id="832910BEBC76D2ADC21E3FF834F87126" ID-CoL="3RM77" authority="Regimbart, 1889" authorityName="Regimbart" authorityYear="1889" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus flavopictus" order="Coleoptera" pageId="240" pageNumber="241" phylum="Arthropoda" rank="species" species="flavopictus">
Laccophilus flavopictus
<normalizedToken id="FA4C9B541794403F96CBFA0061F8E58A" originalValue="Régimbart">Regimbart</normalizedToken>
, 1889
</taxonomicName>
Figs 202-203, 371-372. 523, 577
</paragraph>
</subSubSection>
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<paragraph id="4375439030919365F79237A4EA606B00" pageId="240" pageNumber="241">
<taxonomicName id="DAEEB8B919CFD9144FD8359BA9F549F1" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus flavopictus" order="Coleoptera" pageId="240" pageNumber="241" phylum="Arthropoda" rank="species" species="flavopictus">Laccophilus flavopictus</taxonomicName>
<bibRefCitation id="7DDFD04CE87D5D9A5C20B4A1CE95EFFD" author="Regimbart, M" journalOrPublisher="Notes from the Leyden Museum" pageId="261" pageNumber="262" pagination="51 - 63" title="Dytiscidae et Gyrinidae nouveaux ou rares de la collection du Musee Royal de Leyde." volume="11" year="1889">
<normalizedToken id="964E75B6F064F713F92CD05A7E98B061" originalValue="Régimbart">Regimbart</normalizedToken>
1889
</bibRefCitation>
: 53 (original description, faunistics);
<bibRefCitation id="4375EB644391A80317CB0CF4249FABE2" author="Regimbart, M" journalOrPublisher="Memoires de la Societe entomologique de Belgique" pageId="262" pageNumber="263" pagination="1 - 244" title="Revision des Dytiscidae et Gyrinidae d'Afrique, Madagascar et iles voisines. En contribution a la faune entomologique du Congo." volume="4" year="1895">
<normalizedToken id="A931F26EBB82268983E8EE95CB620FB1" originalValue="Régimbart">Regimbart</normalizedToken>
1895
</bibRefCitation>
: 146 (description, faunistics);
<bibRefCitation id="85C14E0FD6BBB16094054FD16884C872" author="Zimmermann, A" journalOrPublisher="Junk, Berlin" pageId="263" pageNumber="264" title="Dytiscidae, Haliplidae, Hygrobiidae, Amphizoidae. In: Schenkling S (Ed.) Coleopterorum Catalogus. Vol. 4 pars 71. W." year="1920 a">Zimmermann 1920a</bibRefCitation>
:18 (catalogue, faunistics);
<bibRefCitation id="1B4C774149E49C3A219FB8D467E05141" author="Guignot, F" journalOrPublisher="Memoires du Museum National d'Histoire Naturelle Paris" pageId="256" pageNumber="257" pagination="215 - 322" title="Mission scientifique de l'Omo 6 (58). ColeopteraDytiscidae (2 e partie)." volume="19" year="1946 c">Guignot 1946c</bibRefCitation>
: 262, 312 (discussion);
<bibRefCitation id="32EF434D2C205B1656359CDA791FFECB" author="Guignot, F" journalOrPublisher="Exploration du Parc National Albert Mission H. Damas (1935 - 1936)" pageId="256" pageNumber="257" pagination="1 - 44" title="Dytiscidae et Gyrinidae." volume="16" year="1948">Guignot 1948</bibRefCitation>
: 14 (faunistics);
<bibRefCitation id="2B8E94713D0ABAE81510E0B99EAC78C8" author="Guignot, F" journalOrPublisher="Revue de Zoologie et Botanique Africaines" pageId="258" pageNumber="259" pagination="317 - 323" title="Hydrocanthares captures a la lumiere au Congo belge." volume="53" year="1956 c">Guignot 1956c</bibRefCitation>
: 321 (discussion);
<bibRefCitation id="095E650819063E88D41BDC3A137013F1" author="Legros, C" journalOrPublisher="Memoires de l'Institut Francais d'Afrique Noire" pageId="259" pageNumber="260" pagination="209 - 218" title="IX. Coleopteres Hydrocanthares (Mission M. Lamotte et Roy, juillet-decembre 1951)." volume="53" year="1958">Legros 1958</bibRefCitation>
: 211 (faunistics);
<bibRefCitation id="15CE187BA54AF00C49EB1E3B3DAFD03E" author="Guignot, F" journalOrPublisher="Annales du Musee Royal du Congo Belge (Serie 8 vo) Sciences Zoologiques" pageId="258" pageNumber="259" pagination="323 - 648" title="Revision des hydrocanthares d'Afrique (ColeopteraDytiscoidea). 2." volume="78" year="1959 a">Guignot 1959a</bibRefCitation>
: 537, 543 (description, faunistics, discussion);
<bibRefCitation id="76F11FE1D8BA8D24F1E1C709452D9C93" author="Nilsson, AN" journalOrPublisher="Entomologica Fennica" pageId="260" pageNumber="261" pagination="57 - 94" title="Taxonomy, distribution and habitats of the Dytiscidae (Coleoptera) of Ethiopia." volume="4" year="1993">Nilsson and Persson 1993</bibRefCitation>
: 79, 94 (faunistics, biology);
<bibRefCitation id="CB8B474EBEA7B35E135B6DF1C9B124E3" author="Nilsson, AN" journalOrPublisher="World catalogue of Insects" pageId="260" pageNumber="261" pagination="1 - 395" title="Dytiscidae." volume="3" year="2001">Nilsson 2001</bibRefCitation>
: 243 (catalogue, faunistics);
<bibRefCitation id="18AF40E3D04EF16738F4C70076D4AFCB" author="Nilsson, AN" journalOrPublisher="Apollo books, Stenstrup" pageId="260" pageNumber="261" title="A World Catalogue of the Family Dytiscidae, or the Diving Beetles (Coleoptera, Adephaga). Version 1. I. 2015" url="http://www2.emg.umu.se/projects/biginst/andersn/" year="2015">Nilsson 2015</bibRefCitation>
: 211 (catalogue, faunistics).
</paragraph>
<paragraph id="95E94284023BE23744E67DFB722CD09B" pageId="240" pageNumber="241">
<taxonomicName id="2505CB4C2277B7408F74FB89E0C2F2D2" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus bergeri" order="Coleoptera" pageId="240" pageNumber="241" phylum="Arthropoda" rank="species" species="bergeri">Laccophilus bergeri</taxonomicName>
<bibRefCitation id="B4A08FAAC8F33A87710FC17A9E966E03" author="Guignot, F" journalOrPublisher="Revue de Zoologie et Botanique Africaines" pageId="257" pageNumber="258" pagination="233 - 237" title="Hydrocanthares captures a la lumiere dans le Haut-Katanga." volume="47" year="1953 b">Guignot 1953b</bibRefCitation>
: 234, 236 (original description, faunistics);
<bibRefCitation id="D662760BBFA99C4C82B7C5949FF1F498" author="Guignot, F" journalOrPublisher="Annales du Musee Royal du Congo Belge (Serie 8 vo) Sciences Zoologiques" pageId="258" pageNumber="259" pagination="323 - 648" title="Revision des hydrocanthares d'Afrique (ColeopteraDytiscoidea). 2." volume="78" year="1959 a">Guignot 1959a</bibRefCitation>
: 534, 537, 542 (description, faunistics);
<bibRefCitation id="1CFF58D429E01A12874941B6EC69BF2C" author="Omer-Cooper, Joyce" journalOrPublisher="South African Animal Life" pageId="261" pageNumber="262" pagination="59 - 214" title="Chapter 2. Coleoptera: Dytiscidae." volume="11" year="1965">Omer-Cooper 1965</bibRefCitation>
: 76 (description, faunistics);
<bibRefCitation id="A930A5F99CF4BC6FF4A089A7B93EC33D" author="Omer-Cooper, Joyce" journalOrPublisher="Journal of the Entomological Society of Southern Africa" pageId="261" pageNumber="262" pagination="48 - 63" title="Dytiscidae (Coleoptera) of Nyasaland and Southern Rhodesia. X. Cybisterini and addenda." volume="29" year="1967">Omer-Cooper 1967</bibRefCitation>
: 58 (discussion, synonymy
<taxonomicName id="147C3667F647B022E9D093EAF2B08BA9" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus segmentatus" order="Coleoptera" pageId="240" pageNumber="241" phylum="Arthropoda" rank="species" species="segmentatus">Laccophilus segmentatus</taxonomicName>
and
<taxonomicName id="A548136B649D4307E055B48BA7DC07EB" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus bergeri" order="Coleoptera" pageId="240" pageNumber="241" phylum="Arthropoda" rank="species" species="bergeri">Laccophilus bergeri</taxonomicName>
);
<bibRefCitation id="C1D39F737D9F5D15BCB5659C94989402" author="Nilsson, AN" journalOrPublisher="World catalogue of Insects" pageId="260" pageNumber="261" pagination="1 - 395" title="Dytiscidae." volume="3" year="2001">Nilsson 2001</bibRefCitation>
: 240 (catalogue, faunistics);
<bibRefCitation id="1162E0AE5C5446B55911864E14A481A2" author="Nilsson, AN" journalOrPublisher="Apollo books, Stenstrup" pageId="260" pageNumber="261" title="A World Catalogue of the Family Dytiscidae, or the Diving Beetles (Coleoptera, Adephaga). Version 1. I. 2015" url="http://www2.emg.umu.se/projects/biginst/andersn/" year="2015">Nilsson 2015</bibRefCitation>
: 209 (catalogue, faunistics). New synonym.
</paragraph>
<paragraph id="FCDD4EC5C2924C1DFAEFA2F61503F97E" lastPageId="241" lastPageNumber="242" pageId="240" pageNumber="241">
<taxonomicName id="65D36198D7EE478EF937AA8C8B1ADB9E" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus segmentatus" order="Coleoptera" pageId="240" pageNumber="241" phylum="Arthropoda" rank="species" species="segmentatus">Laccophilus segmentatus</taxonomicName>
<bibRefCitation id="1B4EDD5AB77024D30405CA44108634E0" author="Omer-Cooper, Joyce" journalOrPublisher="Memoirs of the Entomological Society of Southern Africa" pageId="260" pageNumber="261" pagination="1 - 90" title="Dytiscidae (Coleoptera) collected in the Transvaal (Rhodes University Expedition 1948)." volume="5" year="1957">Omer-Cooper 1957</bibRefCitation>
: 12, 90 (original description, faunistics);
<bibRefCitation id="C7DFB294F732096C54BE10357C886167" author="Omer-Cooper, Joyce" journalOrPublisher="The Entomologist's Monthly Magazine" pageId="260" pageNumber="261" pagination="57 - 60" title="Some Dytiscidae (Col.) from Swaziland, Southern Africa, with the description of a new species." volume="94" year="1958 a">Omer-Cooper 1958a</bibRefCitation>
: 59 (faunistics);
<bibRefCitation id="0A5F3B1E09DCB111B98D71F00023B2E0" author="Omer-Cooper, Joyce" journalOrPublisher="Journal of the Entomological Society of Southern Africa" pageId="260" pageNumber="261" pagination="36 - 55" title="Dytiscidae from Nyasaland and Southern Rhodesia. III. Laccophilinae." volume="21" year="1958 b">Omer-Cooper 1958b</bibRefCitation>
: 37, 38, 39, 40, 42
<pageBreakToken id="2EB7F9D492A7CD2C92230B7050FD550C" pageId="241" pageNumber="242" start="start">(</pageBreakToken>
description, faunistics, biology, discussion);
<bibRefCitation id="353A71EDB5389A0119A7986AD4CDB162" author="Omer-Cooper, Joyce" journalOrPublisher="South African Animal Life" pageId="261" pageNumber="262" pagination="59 - 214" title="Chapter 2. Coleoptera: Dytiscidae." volume="11" year="1965">Omer-Cooper 1965</bibRefCitation>
: 80 (list synonym
<taxonomicName id="9C45414D03B72BBA33084A5ED2B2396B" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus bergeri" order="Coleoptera" pageId="241" pageNumber="242" phylum="Arthropoda" rank="species" species="bergeri">Laccophilus bergeri</taxonomicName>
Guignot);
<bibRefCitation id="E9834A7AF87CF6542CA4146432E23686" author="Omer-Cooper, Joyce" journalOrPublisher="Journal of the Entomological Society of Southern Africa" pageId="261" pageNumber="262" pagination="48 - 63" title="Dytiscidae (Coleoptera) of Nyasaland and Southern Rhodesia. X. Cybisterini and addenda." volume="29" year="1967">Omer-Cooper 1967</bibRefCitation>
: 58 (discussion, list, synonymy);
<bibRefCitation id="38F923CD4D097420AD109FBC9A539E6C" author="Nilsson, AN" journalOrPublisher="World catalogue of Insects" pageId="260" pageNumber="261" pagination="1 - 395" title="Dytiscidae." volume="3" year="2001">Nilsson 2001</bibRefCitation>
: 240 (catalogue, faunistics, list, synonymy);
<bibRefCitation id="F54E143BDE23A6672897E1F50B01C973" author="Nilsson, AN" journalOrPublisher="Apollo books, Stenstrup" pageId="260" pageNumber="261" title="A World Catalogue of the Family Dytiscidae, or the Diving Beetles (Coleoptera, Adephaga). Version 1. I. 2015" url="http://www2.emg.umu.se/projects/biginst/andersn/" year="2015">Nilsson 2015</bibRefCitation>
: 209 (catalogue, faunistics, list, synonymy,
<taxonomicName id="C9F5C7AF90BB3D774D83EF7D81553ABF" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus bergeri" order="Coleoptera" pageId="241" pageNumber="242" phylum="Arthropoda" rank="species" species="bergeri">Laccophilus bergeri</taxonomicName>
). New synonym.
</paragraph>
</subSubSection>
<subSubSection id="3404CA1CBB1F0C1B057677C4EBBE8682" pageId="241" pageNumber="242" type="type localities">
<paragraph id="A8FAD85B8A760A3A25F0F1237D130D32" pageId="241" pageNumber="242">Type localities.</paragraph>
<paragraph id="4D139AF4758236B31743E1A615D9F107" pageId="241" pageNumber="242">
<taxonomicName id="9E8F0154740C27EBFEB8F24BA4392757" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus flavopictus" order="Coleoptera" pageId="241" pageNumber="242" phylum="Arthropoda" rank="species" species="flavopictus">Laccophilus flavopictus</taxonomicName>
: Angola: Humpata.
</paragraph>
<paragraph id="203AF61E2DE8DF6507A434779BD2ED76" pageId="241" pageNumber="242">
<taxonomicName id="B487E93E5262B0784C806C022D0B1D69" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus bergeri" order="Coleoptera" pageId="241" pageNumber="242" phylum="Arthropoda" rank="species" species="bergeri">Laccophilus bergeri</taxonomicName>
: Zaire (DRC): Elisabethville.
</paragraph>
<paragraph id="E3FE4E3A9F91930A855BFAB48447EB2B" pageId="241" pageNumber="242">
<taxonomicName id="DB7B9136854CD3ED20D85C7C1AEA4F49" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus segmentatus" order="Coleoptera" pageId="241" pageNumber="242" phylum="Arthropoda" rank="species" species="segmentatus">Laccophilus segmentatus</taxonomicName>
: South Africa: Transvaal, Ermelo.
</paragraph>
</subSubSection>
<subSubSection id="48DC2009700F9B1AD3D7766952C42937" pageId="241" pageNumber="242" type="type material">
<paragraph id="160987980165994BD46C210BDB2773DA" pageId="241" pageNumber="242">Type material studied</paragraph>
<paragraph id="D8E137F46FA57A3F3E3FF61DEF2A59A7" pageId="241" pageNumber="242">
(6 exs.).
<taxonomicName id="796E1F29731556F5EFD67A1D780F5210" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus flavopictus" order="Coleoptera" pageId="241" pageNumber="242" phylum="Arthropoda" rank="species" species="flavopictus">Laccophilus flavopictus</taxonomicName>
: Holotype (monotypy): male: &quot;P.Y. v d Kellen Humpata Afr. trop /
<taxonomicName id="B6132B08684719B97EFDECFC4A73B149" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus flavopictus" order="Coleoptera" pageId="241" pageNumber="242" phylum="Arthropoda" rank="species" species="flavopictus">Laccophilus flavopictus</taxonomicName>
sp. n. type / type /
<taxonomicName id="3125E56040947B17AFB106475C9880C2" genus="Algae" lsidName="Algae flavopictus" pageId="241" pageNumber="242" rank="species" species="flavopictus">flavopictus</taxonomicName>
<normalizedToken id="F5BE013D51602982ABC7FEBEF704F444" originalValue="Régimbart">Regimbart</normalizedToken>
sp. n.&quot; (RMNH).
</paragraph>
<paragraph id="E6F17B8408280FB70267A45614381CED" pageId="241" pageNumber="242">
<taxonomicName id="6980D3333AE70C89C7F86FB0C3597E85" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus bergeri" order="Coleoptera" pageId="241" pageNumber="242" phylum="Arthropoda" rank="species" species="bergeri">Laccophilus bergeri</taxonomicName>
: Holotype: male: &quot;Holotypus / Coll. Mus. Congo Elisabethville
<normalizedToken id="42C6BDE3FCF249C719327ECBBB87D5EE" originalValue="lumière">lumiere</normalizedToken>
XI-50/VI-51 Ch. Seydel / Type / R. Det. J. 6182 / Guignot det. 1953
<taxonomicName id="BDFCA2645EE09A7E63D04BFC4B3A31DE" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus bergeri" order="Coleoptera" pageId="241" pageNumber="242" phylum="Arthropoda" rank="species" species="bergeri">Laccophilus bergeri</taxonomicName>
Guign. Type, male&quot; (MRAC; habitus in Fig. 523). - Paratypes: Same data as holotype, but: &quot;Paratypus female / R. Det. L. 6182&quot; (1 ex. MRAC); same as holotype, but: &quot;Allotypus female / (A la
<normalizedToken id="D923A2E4A17412EA845427F0F06D3A57" originalValue="lumière">lumiere</normalizedToken>
) X/XI. 1950/R. Det. K&quot; (1 ex. MRAC); same as holotype, but: &quot;Paratypus male / (A la
<normalizedToken id="B91C63133C900EB8E146C50095EDF807" originalValue="lumière">lumiere</normalizedToken>
) X/XI. 1950&quot; (1 ex. MRAC).
</paragraph>
<paragraph id="005ABDE3FA35914FAD7E4AC3D3067AC1" pageId="241" pageNumber="242">
<taxonomicName id="8687FFC468C2D2E9E7FB355D9FA8C3EC" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus segmentatus" order="Coleoptera" pageId="241" pageNumber="242" phylum="Arthropoda" rank="species" species="segmentatus">Laccophilus segmentatus</taxonomicName>
: Holotype: male: &quot;Type, male-symbol / Type / Transvaal, sluggish stream nr Ermelo 7. 12. 1948 J. O-C / Brit. Mus. 1957-660 /
<taxonomicName id="BC41312AEE89656F9F98820413B26EFF" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus" order="Coleoptera" pageId="241" pageNumber="242" phylum="Arthropoda" rank="genus">Laccophilus</taxonomicName>
sp. n. J. Balfour-Browne det. /
<taxonomicName id="FA174DE5678FDB399A4878081545650A" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus segmentatus" order="Coleoptera" pageId="241" pageNumber="242" phylum="Arthropoda" rank="species" species="segmentatus">Laccophilus segmentatus</taxonomicName>
O-C.&quot; (BMNH).
</paragraph>
</subSubSection>
<subSubSection id="A245972D6DDD350F6B517A604D58A8EF" pageId="241" pageNumber="242" type="materials_examined">
<paragraph id="308DE5FDF27EDDB108C9DBCA48846AED" pageId="241" pageNumber="242">Additional material studied</paragraph>
<paragraph id="5FF66A6F40393F44DEB9760D28DF35E0" pageId="241" pageNumber="242">
(25 exs.). Zaire: &quot;Elisabethville (
<normalizedToken id="60DB829BCDDFAA1006D17F1080DDB3C3" originalValue="à">a</normalizedToken>
la
<normalizedToken id="FC48D75DFA82E92BE93019A279D737B2" originalValue="lumière">lumiere</normalizedToken>
) 1957-1958 Seydel&quot; (1 ex. MRAC). - Zambia: &quot;Mwinilunga District, Ikelenge, nr Kalene, Zambezi Rapids / Pinhey 3.5. 1963 M.V. light trap&quot; (4 exs. BMNH, 1 ex. MZH); &quot;NC, Mkushi env. E 16-18.12. 2004 Snizek leg.&quot; (1 ex. NHPC); &quot;Northern Province 15 km E Luwingu 3-4.12. 2007 1400 m Kudrna Jr. leg. /
<taxonomicName id="ABFB14B4D1D1B9FEFC54B9E5AA4ADE7C" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus bergeri" order="Coleoptera" pageId="241" pageNumber="242" phylum="Arthropoda" rank="species" species="bergeri">Laccophilus bergeri</taxonomicName>
Guign. det. Pederzani&quot; (2 exs. CFP); &quot;100 km SW Serenje 7.12. 2002 Kantner&quot; (1 ex. NHMB). - Malawi: &quot;Dedza Sept. 1948 O-C. / small stream&quot; (1 ex. AMGS); &quot;Dedza Sept. 1948 O-C.&quot;(4 exs. AMGS); &quot;Dedza dam, lower Lilongwe rd 29.IX. 1948&quot; (1 ex. AMGS); &quot;Dedza Hotel, dam 29.IX. 1948&quot; (1 ex. AMGS). - Zimbabwe: &quot;Inyanga N. 1948 JOC. /
<taxonomicName id="CA60F97BB12410AF11C981DBC606BB96" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus segmentatus" order="Coleoptera" pageId="241" pageNumber="242" phylum="Arthropoda" rank="species" species="segmentatus">Laccophilus segmentatus</taxonomicName>
O-C. =
<taxonomicName id="51A228BCE0CA55FC993D3B53C1DD4C26" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus bergeri" order="Coleoptera" pageId="241" pageNumber="242" phylum="Arthropoda" rank="species" species="bergeri">Laccophilus bergeri</taxonomicName>
Guign. Det. J. Omer-Cooper&quot; (1 ex. AMGS); &quot;Stream Rusapi 13.XI. (?) 1948 /
<taxonomicName id="6A0A3422614400BBDB4AA075098D8DCA" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus segmentatus" order="Coleoptera" pageId="241" pageNumber="242" phylum="Arthropoda" rank="species" species="segmentatus">Laccophilus segmentatus</taxonomicName>
O-C. det. J. Omer-Cooper&quot; (1 ex. AMGS); &quot;Marandellas 2.11. 1948 J. Omer-Cooper&quot; (3 exs. MNHN). - Swaziland: &quot;Mbabane 5.XII. 1948 J.O.C. /
<taxonomicName id="BA92E095CDA5A1541F8434D7338C0DE5" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus segmentatus" order="Coleoptera" pageId="241" pageNumber="242" phylum="Arthropoda" rank="species" species="segmentatus">Laccophilus segmentatus</taxonomicName>
O-C. =
<taxonomicName id="99E25899C44E1432B15709EDDD7C00F2" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus bergeri" order="Coleoptera" pageId="241" pageNumber="242" phylum="Arthropoda" rank="species" species="bergeri">Laccophilus bergeri</taxonomicName>
Guign. Det. J. Omer-Cooper&quot; (1 ex. AMGS). - South Africa: &quot;Natal Durban Umgeni Tragardh&quot; (1 ex. AMGS); &quot;
<taxonomicName id="14C873428859FE0874DE1686A3D88A41" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus" order="Coleoptera" pageId="241" pageNumber="242" phylum="Arthropoda" rank="genus">Laccophilus</taxonomicName>
sp. n. J. Balfour-Browne / Paratype / Vlei nr
<normalizedToken id="2D5FA5E875C7FB4FDE25B0448B5B5F15" originalValue="Wasserman’s">Wasserman's</normalizedToken>
Beacon 6.12. 1948 /
<taxonomicName id="A64F0E90589FF1C9F9B906E588DD4058" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus segmentatus" order="Coleoptera" pageId="241" pageNumber="242" phylum="Arthropoda" rank="species" species="segmentatus">Laccophilus segmentatus</taxonomicName>
O-C.&quot; (1 ex. AMGS).
</paragraph>
</subSubSection>
<subSubSection id="4975A4C7239DF4513014BE8F0B472E93" pageId="241" pageNumber="242" type="comments on synonymy">
<paragraph id="5446FD6E4A17991C4186B6FFB6618F4F" pageId="241" pageNumber="242">Comments on synonymy.</paragraph>
<paragraph id="28313DB91B14BA291ED2D23C61DCFBA0" pageId="241" pageNumber="242">
Male holotypes of the three involved taxa,
<taxonomicName id="20B9B105D5CAF10FDEB100B7E4E78BA8" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus flavopictus" order="Coleoptera" pageId="241" pageNumber="242" phylum="Arthropoda" rank="species" species="flavopictus">Laccophilus flavopictus</taxonomicName>
,
<taxonomicName id="14DD80714F424A09ECFD083B8DCA82EB" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus bergeri" order="Coleoptera" pageId="241" pageNumber="242" phylum="Arthropoda" rank="species" species="bergeri">Laccophilus bergeri</taxonomicName>
, and
<taxonomicName id="162826A160D0E6F474FE48A18E17D010" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus segmentatus" order="Coleoptera" pageId="241" pageNumber="242" phylum="Arthropoda" rank="species" species="segmentatus">Laccophilus segmentatus</taxonomicName>
, have been examined and found to be conspecific. Because
<taxonomicName id="4D1D6F2B7F5BE9CC0DE4A7B74B366EB6" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus flavopictus" order="Coleoptera" pageId="241" pageNumber="242" phylum="Arthropoda" rank="species" species="flavopictus">Laccophilus flavopictus</taxonomicName>
is the oldest available name, it is also the valid name of the species.
</paragraph>
</subSubSection>
<subSubSection id="049967C4D3FA71480A8E4E459429BE48" lastPageId="242" lastPageNumber="243" pageId="241" pageNumber="242" type="diagnosis">
<paragraph id="793597C389385FA14D371A5F445C71BE" pageId="241" pageNumber="242">Diagnosis.</paragraph>
<paragraph id="8D76E793FEB65EA01FC5BCAA69F7F869" lastPageId="242" lastPageNumber="243" pageId="241" pageNumber="242">
<taxonomicName id="DC742A0127D180231B0196D04C201134" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus flavopictus" order="Coleoptera" pageId="241" pageNumber="242" phylum="Arthropoda" rank="species" species="flavopictus">Laccophilus flavopictus</taxonomicName>
is characterized by the following combination of features: Elytra almost black to blackish ferrugineous; basally provided with a quite narrow but distinct, transverse, pale ferrugineous marking and by non-hooked penis apex.
<pageBreakToken id="138ECB996A8D6039FFD7E177D23BDE73" pageId="242" pageNumber="243" start="start">Externally</pageBreakToken>
it resembles of some species not located in this species group, eg.
<taxonomicName id="E7C2F3971B0E294017F55049310C226F" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus luctuosus" order="Coleoptera" pageId="242" pageNumber="243" phylum="Arthropoda" rank="species" species="luctuosus">Laccophilus luctuosus</taxonomicName>
and
<taxonomicName id="94CBBE18DE440CEE569161CC3EE95079" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus secundus" order="Coleoptera" pageId="242" pageNumber="243" phylum="Arthropoda" rank="species" species="secundus">Laccophilus secundus</taxonomicName>
. Presence of a stridulation apparatus and shape of male genitalia, however, shows that
<taxonomicName id="F010DEA0DDE9AAFD6C5F219D8AFBE779" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus flavopictus" order="Coleoptera" pageId="242" pageNumber="243" phylum="Arthropoda" rank="species" species="flavopictus">Laccophilus flavopictus</taxonomicName>
belongs to another species group than these two species.
</paragraph>
</subSubSection>
<subSubSection id="443156CED012443D084FA40513088EC5" pageId="242" pageNumber="243" type="description">
<paragraph id="95BD89C77428DA3F139D1091320A47D4" pageId="242" pageNumber="243">Description.</paragraph>
<paragraph id="7D16F87BC188E2619AEA71ACE7E84691" pageId="242" pageNumber="243">Body length 3.4-3.7 mm, width 1.9-2.0 mm. Colour pattern generally distinct and exhibits only minor variation; sometimes in part, dark areas with somewhat vague delimitation (Fig. 523).</paragraph>
<paragraph id="2F35D0D063189084B583CD622BE0EF5A" pageId="242" pageNumber="243">Head: Pale ferrugineous to ferrugineous; posteriorly at pronotum darker; dark ferrugineous. Slightly mat, finely microsculptured; reticulation double (in part indistinct). Large meshes when discernible, contain 2-5 small meshes. At eyes and in a narrow, transverse area between eyes with very fine, somewhat irregular punctures; medially punctures indistinct and hardly visible.</paragraph>
<paragraph id="66C606C93F27225AE5D8649FA4972E92" pageId="242" pageNumber="243">Pronotum: Pale ferrugineous, frontally and basally with a medial, dark marking (Fig. 518). Submat to rather shiny, fienly microsculptured; reticulation in part double (partially indistinct). Large meshes generally contain 2-5 small meshes. At margins with fine, in part indistinct and sparse punctures.</paragraph>
<paragraph id="168C659B1A45CF2A268697FD66377DC0" pageId="242" pageNumber="243">Elytra: Blackish ferrugineous to dark ferrugineous, with distinct pale ferrugineous markings (Fig. 523). Submat to rather shiny, finely microsculptured. Reticulation in part double but double reticulation somewhat indistinct; large meshes, when discernible, contain 2-5 small meshes. Punctation very fine and form three rather irregular, longitudinal areas.</paragraph>
<paragraph id="BC55456FCFDF738C9680D3BEACDEAF24" pageId="242" pageNumber="243">Ventral aspect: Pale ferrugineous, abdominal ventrites at least in part darker, ferrugineous to dark ferrugineous. Stridulatory apparatus consists of about 30 fine ridges. Submat, finely microsculptured. Abdomen provided with fine, slightly sparse, curved striae. Apical ventrite provided with a sharp knob on one side (other side lacks knob) (Fig. 202). Prosternal process narrow, posteriorly slightly extended and apically pointed.</paragraph>
<paragraph id="A18E7ADB3DBDDE16B13BB803505E04BD" pageId="242" pageNumber="243">Legs: Pro- and mesotarsus slender, somewhat extended and provided with suckers.</paragraph>
<paragraph id="9658550E3F4FE1FB4002F22EEE5757C4" pageId="242" pageNumber="243">Male genitalia: Penis comparatively long, slightly angled; apical outline rounded to almost straight (Figs 371-372).</paragraph>
<paragraph id="F445BFB155CC94A1714A7C8007A0559C" pageId="242" pageNumber="243">Female: Externally as male but apical ventrite almost symmetric, lacking sharp knob (Fig. 203), Pro- and mesotarsus slender.</paragraph>
</subSubSection>
<subSubSection id="F59BDD7BDA275B2FC32D38E48E071737" pageId="242" pageNumber="243" type="distribution">
<paragraph id="AE32B079D5D8AD02ADE81BF49F6197EA" pageId="242" pageNumber="243">Distribution.</paragraph>
<paragraph id="32E1A50ADF79B0253B7B5D2C6AF5D0BA" pageId="242" pageNumber="243">Zaire, Angola, Zambia, Malawi, Zimbabwe, South Africa, Swaziland (Fig. 577). Only verified records are used in the map.</paragraph>
</subSubSection>
<subSubSection id="BC861491983614B4F953615B7E347D1B" pageId="242" pageNumber="243" type="collecting circumstances">
<paragraph id="D39B14D8A716D435F10D5438F5C8CE24" pageId="242" pageNumber="243">Collecting circumstances.</paragraph>
<paragraph id="D16129284F0CB9E5954BA63B4FF89136" pageId="242" pageNumber="243">
<bibRefCitation id="43A2AA14FCEBD8FBCF9FE3A2DD5E8F17" author="Omer-Cooper, Joyce" journalOrPublisher="Journal of the Entomological Society of Southern Africa" pageId="260" pageNumber="261" pagination="36 - 55" title="Dytiscidae from Nyasaland and Southern Rhodesia. III. Laccophilinae." volume="21" year="1958 b">Omer-Cooper (1958b)</bibRefCitation>
reports the species from different kinds of water bodies, as ponds, pools, streams, bogs and an artificial dam. Often in association with some vegetation as water lilies, rushes, weeds etc. Collected at light.
</paragraph>
</subSubSection>
</treatment>
</document>
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