283 lines
35 KiB
XML
283 lines
35 KiB
XML
<document id="53DE09B96231994ED72C493B4B72D576" ID-DOI="10.5281/zenodo.273937" ID-GBIF-Dataset="90a1585e-4f87-47d1-938d-e1663c2311b1" ID-ISSN="1175-5326" ID-Zenodo-Dep="273937" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459926661040" checkinUser="plazi" docAuthor="Pawson, David L. & Vance, Doris J." docDate="2007" docId="8C5BDF31EF2FFFDFFF3BF91A935BF87F" docLanguage="en" docName="zt01608p039.pdf" docOrigin="Zootaxa 1608" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Heteromolpadia joyceae Pawson & Vance, 2007, new species" docType="treatment" docVersion="8" lastPageNumber="38" masterDocId="7062A749EF2EFFDCFFACFFF59717FFC5" masterDocTitle="Heteromolpadia joyceae, a new species of sea cucumber from New Zealand, with a key to New Zealand Molpadiidae (Echinodermata: Holothuroidea: Molpadiida)" masterLastPageNumber="39" masterPageNumber="35" pageNumber="36" updateTime="1698229595558" updateUser="plazi">
|
||
<mods:mods id="35D6735744BAE04492C1795BC877B531" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="3788ED136ACDB055269C9BB64182E072">
|
||
<mods:title id="5E961B72C57D7C2F73C3AE4FCC6BEA32">Heteromolpadia joyceae, a new species of sea cucumber from New Zealand, with a key to New Zealand Molpadiidae (Echinodermata: Holothuroidea: Molpadiida)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="D1133B131326562B89F05C4CAD0B643B" type="personal">
|
||
<mods:role id="9383F6833591A2A63BB47D21F6CDAD91">
|
||
<mods:roleTerm id="1714225D2CB604168B1CFD6CE920A760">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="C7EB9B7ADA8D3E04024E186BC7EC7DC1">Pawson, David L.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="2C9C6B5217908E5B40D2F53C484457CE" type="personal">
|
||
<mods:role id="712906DA2AE2C694A38D2F57A8E6F0CE">
|
||
<mods:roleTerm id="6926F4716C3F40BDF2A78E013553D273">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="2DD965A7B11AA6666DC2329A41882677">Vance, Doris J.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="7E635A0729A361E5645ED201A39EDC08">text</mods:typeOfResource>
|
||
<mods:relatedItem id="F16DB0127D8A7C8BC541217F4FE82BE4" type="host">
|
||
<mods:titleInfo id="D8CB22214BBE9EAEB5387BEB03415BFB">
|
||
<mods:title id="710E8CEAAB7EE0AF4A9EE7F4D6B30208">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="980F05BEF0761E54006114CA2EA5EA7F">
|
||
<mods:date id="CED136D8BECE881E50869696B72C37A3">2007</mods:date>
|
||
<mods:detail id="BC5730486F4840842B69A870063F42C3" type="volume">
|
||
<mods:number id="C0425C53DEF1ED24AE22FDC80BE047E2">1608</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="65588E5208F2465A8FDDB971AF7CEA45" unit="page">
|
||
<mods:start id="AD4533653482D9DB3B1574C8D9102EB2">35</mods:start>
|
||
<mods:end id="403D8495141367346CE85D99ABE19E55">39</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="84483A27EE337752820BD5CBE7C9E3A6">journal article</mods:classification>
|
||
<mods:identifier id="8164A83CCBB912754E0BABEC3D6758AE" type="DOI">10.5281/zenodo.273937</mods:identifier>
|
||
<mods:identifier id="40C85D0EB23BC22776DEF821E0BBAC90" type="GBIF-Dataset">90a1585e-4f87-47d1-938d-e1663c2311b1</mods:identifier>
|
||
<mods:identifier id="06ED516224E75F2AD3F2E525CCFFFC3E" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="612E7BF1813485723D4C314FFBB778B0" type="Zenodo-Dep">273937</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="8C5BDF31EF2FFFDFFF3BF91A935BF87F" ID-DOI="http://doi.org/10.5281/zenodo.6238122" ID-GBIF-Taxon="119374385" ID-Zenodo-Dep="6238122" LSID="urn:lsid:plazi:treatment:8C5BDF31EF2FFFDFFF3BF91A935BF87F" httpUri="http://treatment.plazi.org/id/8C5BDF31EF2FFFDFFF3BF91A935BF87F" lastPageId="3" lastPageNumber="38" pageId="1" pageNumber="36">
|
||
<subSubSection id="4CE83DACEF2FFFDDFF3BF91A9639F8E9" pageId="1" pageNumber="36" type="nomenclature">
|
||
<paragraph id="044D6E27EF2FFFDDFF3BF91A954EF8CC" blockId="1.[151,601,1775,1836]" box="[151,601,1775,1801]" pageId="1" pageNumber="36">
|
||
<heading id="5F05D94BEF2FFFDDFF3BF91A954EF8CC" bold="true" box="[151,601,1775,1801]" fontSize="11" level="1" pageId="1" pageNumber="36" reason="1">
|
||
<emphasis id="3686B235EF2FFFDDFF3BF91A954EF8CC" bold="true" box="[151,601,1775,1801]" pageId="1" pageNumber="36">
|
||
<taxonomicName id="C3F215A4EF2FFFDDFF3BF91A96ACF8CC" ID-CoL="6LT8G" box="[151,443,1775,1801]" class="Holothuroidea" family="Molpadiidae" genus="Heteromolpadia" kingdom="Animalia" order="Molpadida" pageId="1" pageNumber="36" phylum="Echinodermata" rank="species" species="joyceae" status="sp. nov.">
|
||
<emphasis id="3686B235EF2FFFDDFF3BF91A96ACF8CC" bold="true" box="[151,443,1775,1801]" italics="true" pageId="1" pageNumber="36">Heteromolpadia joyceae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="2DB50F4EEF2FFFDDFE66F91A954EF8CC" box="[458,601,1775,1801]" pageId="1" pageNumber="36" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="044D6E27EF2FFFDDFF3BF8E79639F8E9" blockId="1.[151,601,1775,1836]" box="[151,302,1810,1836]" pageId="1" pageNumber="36">
|
||
<figureCitation id="9CC972A2EF2FFFDDFF3BF8E797E4F8E9" box="[151,243,1810,1836]" captionStart="FIGURE 1" captionStartId="2.[151,255,1866,1890]" captionTargetBox="[173,1409,320,1832]" captionTargetId="figure@2.[169,1417,317,1840]" captionTargetPageId="2" captionText="FIGURE 1. Heteromolpadia joyceae new species. A, Holotype, whole animal, total length 86 mm. B, scattered phosphatic deposits derived from dissolution of skin in liquid household bleach. C, Remnants of a spired table with three larger perforations and three smaller. D, Fragments of three-armed anchor plate. E, Spired tables from tail. F, spired tables from tail, higher magnification. Scale line in Figures 1 B-F = 100 Μm." httpUri="https://zenodo.org/record/273938/files/figure.png" pageId="1" pageNumber="36">Figure 1</figureCitation>
|
||
A–F
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4CE83DACEF2FFFDFFF3BF8AF935BF87F" lastPageId="3" lastPageNumber="38" pageId="1" pageNumber="36" type="description">
|
||
<paragraph id="044D6E27EF2FFFDDFF3BF8AF9590F8B4" blockId="1.[151,647,1882,1905]" box="[151,647,1882,1905]" pageId="1" pageNumber="36">
|
||
<treatmentCitationGroup id="24E24909EF2FFFDDFF3BF8AF9590F8B4" box="[151,647,1882,1905]" pageId="1" pageNumber="36">
|
||
<taxonomicName id="C3F215A4EF2FFFDDFF3BF8AF97E8F8B4" ID-CoL="7PKB8" box="[151,255,1882,1905]" class="Holothuroidea" family="Molpadiidae" genus="Molpadia" kingdom="Animalia" order="Molpadida" pageId="1" pageNumber="36" phylum="Echinodermata" rank="genus" status="sp. nov.">
|
||
<emphasis id="3686B235EF2FFFDDFF3BF8AF97E8F8B4" box="[151,255,1882,1905]" italics="true" pageId="1" pageNumber="36">Molpadia</emphasis>
|
||
</taxonomicName>
|
||
?
|
||
<taxonomicNameLabel id="2DB50F4EEF2FFFDDFEBDF8AF9653F8B4" box="[273,324,1882,1905]" pageId="1" pageNumber="36" rank="species">n.sp.</taxonomicNameLabel>
|
||
<treatmentCitation id="85534836EF2FFFDDFEE6F8AF9501F8B4" author="Pawson" box="[330,534,1882,1905]" page="13" pageId="1" pageNumber="36" year="1965">
|
||
<bibRefCitation id="606313D6EF2FFFDDFEE6F8AF96E7F8B4" author="Pawson" box="[330,496,1882,1905]" pageId="1" pageNumber="39" refString="Pawson, D. L. (1965 a) The bathyal holothurians of the New Zealand region. Zoology Publications from Victoria University of Wellington 39, 1 - 33." type="journal article" year="1965" yearSuffix="a">Pawson, 1965a</bibRefCitation>
|
||
: 13
|
||
</treatmentCitation>
|
||
; 1977: 98.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="044D6E27EF2FFFDEFF3BF86F97F9FEC7" blockId="1.[151,1436,1946,2012]" lastBlockId="2.[151,1436,152,258]" lastPageId="2" lastPageNumber="37" pageId="1" pageNumber="36">
|
||
Diagnosis: Body gray, typically with very numerous reddish spots approximately
|
||
<quantity id="C30AC3C2EF2FFFDDFBDFF86F93ADF871" box="[1139,1210,1946,1972]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="36" unit="mm" value="1.0">1 mm</quantity>
|
||
in diameter. Body wall ossicles in young specimens three-armed anchor plates, anchors, and three-pillared tables 100µm in average diameter with three larger perforations and often three additional smaller perforations. Body wall ossicles almost completely disappear as individuals grow. Tables in tail approximately 300Μm long, with 15–20 perforations.
|
||
</paragraph>
|
||
<caption id="508D3EAFEF2CFFDEFF3BF8BF94ADF809" httpUri="https://zenodo.org/record/273938/files/figure.png" pageId="2" pageNumber="37" targetBox="[173,1409,320,1832]" targetPageId="2">
|
||
<paragraph id="044D6E27EF2CFFDEFF3BF8BF94ADF809" blockId="2.[151,1436,1866,1996]" pageId="2" pageNumber="37">
|
||
<emphasis id="3686B235EF2CFFDEFF3BF8BF960CF8A7" bold="true" box="[151,283,1866,1890]" pageId="2" pageNumber="37">FIGURE 1.</emphasis>
|
||
<taxonomicName id="C3F215A4EF2CFFDEFE88F8BF953BF8A4" box="[292,556,1866,1889]" class="Holothuroidea" family="Molpadiidae" genus="Heteromolpadia" kingdom="Animalia" order="Molpadida" pageId="2" pageNumber="37" phylum="Echinodermata" rank="species" species="joyceae" status="sp. nov.">
|
||
<emphasis id="3686B235EF2CFFDEFE88F8BF953BF8A4" box="[292,556,1866,1889]" italics="true" pageId="2" pageNumber="37">Heteromolpadia joyceae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="2DB50F4EEF2CFFDEFD99F8BF95ABF8A4" box="[565,700,1866,1889]" pageId="2" pageNumber="37" rank="species">new species</taxonomicNameLabel>
|
||
. A, Holotype, whole animal, total length 86mm. B, scattered phosphatic deposits derived from dissolution of skin in liquid household bleach. C, Remnants of a spired table with three larger perforations and three smaller. D, Fragments of three-armed anchor plate. E, Spired tables from tail. F, spired tables from tail, higher magnification. Scale line in Figures 1B-F = 100Μm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="044D6E27EF2DFFDFFF6AFF6D94ACFE67" blockId="3.[151,1437,152,1978]" pageId="3" pageNumber="38">
|
||
Material Examined:
|
||
<typeStatus id="DB49D085EF2DFFDFFE10FF6D954DFF77" box="[444,602,152,178]" pageId="3" pageNumber="38" type="holotype">HOLOTYPE</typeStatus>
|
||
, Bay of Plenty, North Island,
|
||
<collectingCountry id="7CE52EB7EF2DFFDFFC68FF6D9370FF77" box="[964,1127,152,178]" name="New Zealand" pageId="3" pageNumber="38">New Zealand</collectingCountry>
|
||
, NZOI Station Z9005,
|
||
<date id="704C48E7EF2DFFDFFAD3FF6D928BFF77" box="[1407,1436,152,178]" pageId="3" pageNumber="38" value="1998-01-21">21</date>
|
||
January 1998,
|
||
<geoCoordinate id="61C608E0EF2DFFDFFEEEFF359677FF1F" box="[322,352,192,218]" direction="south" orientation="latitude" pageId="3" pageNumber="38" precision="9" value="-37.616665">37</geoCoordinate>
|
||
°37.00’S–
|
||
<geoCoordinate id="61C608E0EF2DFFDFFE74FF35954AFF1F" box="[472,605,192,218]" direction="south" orientation="latitude" pageId="3" pageNumber="38" precision="9" value="-37.65">37°39.00’S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="61C608E0EF2DFFDFFDC5FF3595E8FF1F" box="[617,767,192,218]" direction="east" orientation="longitude" pageId="3" pageNumber="38" precision="9" value="176.80817">176°48.49’E</geoCoordinate>
|
||
–
|
||
<geoCoordinate id="61C608E0EF2DFFDFFCA3FF3594B3FF1F" box="[783,932,192,218]" direction="east" orientation="longitude" pageId="3" pageNumber="38" precision="9" value="176.7455">176°44.73’E</geoCoordinate>
|
||
,
|
||
<quantity id="C30AC3C2EF2DFFDFFC1CFF359326FF1F" box="[944,1073,192,218]" metricMagnitude="2" metricUnit="m" metricValue="3.635" metricValueMax="3.67" metricValueMin="3.6" pageId="3" pageNumber="38" unit="m" value="363.5" valueMax="367.0" valueMin="360.0">360–367m</quantity>
|
||
,
|
||
<specimenCount id="12F4A5AEEF2DFFDFFB90FF3593D2FF1F" box="[1084,1221,192,218]" pageId="3" pageNumber="38" type="generic">1 specimen</specimenCount>
|
||
.
|
||
<typeStatus id="DB49D085EF2DFFDFFB62FF359264FF1F" box="[1230,1395,192,218]" pageId="3" pageNumber="38" type="paratype">PARATYPES</typeStatus>
|
||
,
|
||
<specimenCount id="12F4A5AEEF2DFFDFFAD3FF359601FEC7" pageId="3" pageNumber="38" type="generic">17 specimens</specimenCount>
|
||
: NZOI Station Z8999,
|
||
<date id="704C48E7EF2DFFDFFD8EFF1D9528FEC7" box="[546,575,232,258]" pageId="3" pageNumber="38" value="1998-01-20">20</date>
|
||
January 1998,
|
||
<geoCoordinate id="61C608E0EF2DFFDFFD5EFF1D9407FEC7" box="[754,784,232,258]" direction="south" orientation="latitude" pageId="3" pageNumber="38" precision="9" value="-37.613167">37</geoCoordinate>
|
||
°36.79’S–
|
||
<geoCoordinate id="61C608E0EF2DFFDFFC2BFF1D931AFEC7" box="[903,1037,232,258]" direction="south" orientation="latitude" pageId="3" pageNumber="38" precision="9" value="-38.783333">38°47.00’S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="61C608E0EF2DFFDFFBB6FF1D93B8FEC7" box="[1050,1199,232,258]" direction="east" orientation="longitude" pageId="3" pageNumber="38" precision="9" value="177.20166">177°12.10’E</geoCoordinate>
|
||
–
|
||
<geoCoordinate id="61C608E0EF2DFFDFFB6CFF1D9242FEC7" box="[1216,1365,232,258]" direction="east" orientation="longitude" pageId="3" pageNumber="38" precision="9" value="177.215">177°12.90’E</geoCoordinate>
|
||
,
|
||
<quantity id="C30AC3C2EF2DFFDFFACDFF1D97CBFEEF" metricMagnitude="2" metricUnit="m" metricValue="4.635" metricValueMax="4.67" metricValueMin="4.6" pageId="3" pageNumber="38" unit="m" value="463.5" valueMax="467.0" valueMin="460.0">460– 467m</quantity>
|
||
,
|
||
<specimenCount id="12F4A5AEEF2DFFDFFF58FEE59683FEEF" box="[244,404,272,298]" pageId="3" pageNumber="38" type="generic">5 specimens</specimenCount>
|
||
. NZOI Station Z9008,
|
||
<date id="704C48E7EF2DFFDFFD7DFEE595F9FEEF" box="[721,750,272,298]" pageId="3" pageNumber="38" value="1998-01-22">22</date>
|
||
January 1998,
|
||
<geoCoordinate id="61C608E0EF2DFFDFFC69FEE594F3FEEF" box="[965,996,272,298]" direction="south" orientation="latitude" pageId="3" pageNumber="38" precision="9" value="-37.1645">37</geoCoordinate>
|
||
°09.87’S–
|
||
<geoCoordinate id="61C608E0EF2DFFDFFBF0FEE593F7FEEF" box="[1116,1248,272,298]" direction="south" orientation="latitude" pageId="3" pageNumber="38" precision="9" value="-37.211166">37°12.67’S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="61C608E0EF2DFFDFFB54FEE5929CFEEF" box="[1272,1419,272,298]" direction="east" orientation="longitude" pageId="3" pageNumber="38" precision="9" value="176.36217">176°21.73’E</geoCoordinate>
|
||
–
|
||
<geoCoordinate id="61C608E0EF2DFFDFFF3BFECD9639FE97" box="[151,302,312,338]" direction="east" orientation="longitude" pageId="3" pageNumber="38" precision="9" value="176.38266">176°22.96’E</geoCoordinate>
|
||
,
|
||
<quantity id="C30AC3C2EF2DFFDFFE92FECD96A8FE97" box="[318,447,312,338]" metricMagnitude="2" metricUnit="m" metricValue="5.27" metricValueMax="5.36" metricValueMin="5.18" pageId="3" pageNumber="38" unit="m" value="527.0" valueMax="536.0" valueMin="518.0">518–536m</quantity>
|
||
,
|
||
<specimenCount id="12F4A5AEEF2DFFDFFE62FECD9571FE97" box="[462,614,312,338]" pageId="3" pageNumber="38" type="generic">9 specimens</specimenCount>
|
||
. NMNZ Station BS754,
|
||
<date id="704C48E7EF2DFFDFFC3AFECD94A4FE97" box="[918,947,312,338]" pageId="3" pageNumber="38" value="1979-01-23">23</date>
|
||
January 1979,
|
||
<geoCoordinate id="61C608E0EF2DFFDFFBDEFECD9386FE97" box="[1138,1169,312,338]" direction="south" orientation="latitude" pageId="3" pageNumber="38" precision="92" value="-37.146667">37</geoCoordinate>
|
||
°08.8’S
|
||
<geoCoordinate id="61C608E0EF2DFFDFFB5FFECD926DFE97" box="[1267,1402,312,338]" direction="east" orientation="longitude" pageId="3" pageNumber="38" precision="92" value="176.36333">176°21.8’E</geoCoordinate>
|
||
to
|
||
<geoCoordinate id="61C608E0EF2DFFDFFF3BFE95961AFEBF" box="[151,269,352,378]" direction="south" orientation="latitude" pageId="3" pageNumber="38" precision="92" value="-37.156666">37°09.4’S</geoCoordinate>
|
||
<geoCoordinate id="61C608E0EF2DFFDFFEB5FE9596B5FEBF" box="[281,418,352,378]" direction="east" orientation="longitude" pageId="3" pageNumber="38" precision="92" value="176.39667">176°23.8’E</geoCoordinate>
|
||
,
|
||
<quantity id="C30AC3C2EF2DFFDFFE1EFE959524FEBF" box="[434,563,352,378]" metricMagnitude="2" metricUnit="m" metricValue="5.72" metricValueMax="6.32" metricValueMin="5.12" pageId="3" pageNumber="38" unit="m" value="572.0" valueMax="632.0" valueMin="512.0">512–632m</quantity>
|
||
, mud,
|
||
<specimenCount id="12F4A5AEEF2DFFDFFD26FE959400FEBF" box="[650,791,352,378]" pageId="3" pageNumber="38" type="generic">1 specimen</specimenCount>
|
||
. NMNZ Station BS772,
|
||
<date id="704C48E7EF2DFFDFFBE4FE959372FEBF" box="[1096,1125,352,378]" pageId="3" pageNumber="38" value="1979-01-27">27</date>
|
||
January 1979,
|
||
<geoCoordinate id="61C608E0EF2DFFDFFA8AFE959251FEBF" box="[1318,1350,352,378]" direction="south" orientation="latitude" pageId="3" pageNumber="38" precision="92" value="-39.286667">39</geoCoordinate>
|
||
°17.2’S
|
||
<geoCoordinate id="61C608E0EF2DFFDFFF3BFE7D960AFE67" box="[151,285,392,418]" direction="east" orientation="longitude" pageId="3" pageNumber="38" precision="92" value="178.21167">178°12.7’E</geoCoordinate>
|
||
to
|
||
<geoCoordinate id="61C608E0EF2DFFDFFEEFFE7D96AEFE67" box="[323,441,392,418]" direction="south" orientation="latitude" pageId="3" pageNumber="38" precision="92" value="-39.321667">39°19.3’S</geoCoordinate>
|
||
<geoCoordinate id="61C608E0EF2DFFDFFE6CFE7D955FFE67" box="[448,584,392,418]" direction="east" orientation="longitude" pageId="3" pageNumber="38" precision="92" value="178.18834">178°11.3’E</geoCoordinate>
|
||
,
|
||
<quantity id="C30AC3C2EF2DFFDFFDF9FE7D95C2FE67" box="[597,725,392,418]" metricMagnitude="2" metricUnit="m" metricValue="2.82" metricValueMax="3.06" metricValueMin="2.58" pageId="3" pageNumber="38" unit="m" value="282.0" valueMax="306.0" valueMin="258.0">258–306m</quantity>
|
||
, mud,
|
||
<specimenCount id="12F4A5AEEF2DFFDFFC8FFE7D94A0FE67" box="[803,951,392,418]" pageId="3" pageNumber="38" type="generic">2 specimens</specimenCount>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="044D6E27EF2DFFDFFF6AFE459463FE0F" blockId="3.[151,1437,152,1978]" box="[198,884,432,458]" pageId="3" pageNumber="38">
|
||
Additional material:
|
||
<specimenCount id="12F4A5AEEF2DFFDFFE16FE45954EFE0F" box="[442,601,432,458]" pageId="3" pageNumber="38" type="generic">67 specimens</specimenCount>
|
||
from 16 NZOI stations.
|
||
</paragraph>
|
||
<paragraph id="044D6E27EF2DFFDFFF6AFE2D93B8FD27" blockId="3.[151,1437,152,1978]" pageId="3" pageNumber="38">
|
||
Description: Specimens typical of the Family
|
||
<taxonomicName id="C3F215A4EF2DFFDFFD4FFE2D946CFE37" box="[739,891,472,498]" class="Holothuroidea" family="Molpadiidae" kingdom="Animalia" order="Molpadida" pageId="3" pageNumber="38" phylum="Echinodermata" rank="family">Molpadiidae</taxonomicName>
|
||
, although the tail is relatively long, occupying 17–30% of the total length of the body.
|
||
<typeStatus id="DB49D085EF2DFFDFFDCBFDF595C3FDDF" box="[615,724,512,538]" pageId="3" pageNumber="38" type="holotype">Holotype</typeStatus>
|
||
(
|
||
<figureCitation id="9CC972A2EF2DFFDFFD4FFDF59454FDDF" box="[739,835,512,538]" captionStart="FIGURE 1" captionStartId="2.[151,255,1866,1890]" captionTargetBox="[173,1409,320,1832]" captionTargetId="figure@2.[169,1417,317,1840]" captionTargetPageId="2" captionText="FIGURE 1. Heteromolpadia joyceae new species. A, Holotype, whole animal, total length 86 mm. B, scattered phosphatic deposits derived from dissolution of skin in liquid household bleach. C, Remnants of a spired table with three larger perforations and three smaller. D, Fragments of three-armed anchor plate. E, Spired tables from tail. F, spired tables from tail, higher magnification. Scale line in Figures 1 B-F = 100 Μm." httpUri="https://zenodo.org/record/273938/files/figure.png" pageId="3" pageNumber="38">Figure 1</figureCitation>
|
||
A)
|
||
<quantity id="C30AC3C2EF2DFFDFFCC1FDF594AFFDDF" box="[877,952,512,538]" metricMagnitude="-2" metricUnit="m" metricValue="8.6" pageId="3" pageNumber="38" unit="mm" value="86.0">86mm</quantity>
|
||
in total length, of which the tail occupies
|
||
<quantity id="C30AC3C2EF2DFFDFFF3BFDDD97F1FD87" box="[151,230,552,578]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="38" unit="mm" value="16.0">16mm</quantity>
|
||
. In all specimens, total length is
|
||
<quantity id="C30AC3C2EF2DFFDFFDCBFDDD95EEFD87" box="[615,761,552,578]" metricMagnitude="-2" metricUnit="m" metricValue="8.4" metricValueMax="12.0" metricValueMin="4.8" pageId="3" pageNumber="38" unit="mm" value="84.0" valueMax="120.0" valueMin="48.0">48–120 mm</quantity>
|
||
. Body wall smooth to touch, thin, but tough and leathery. Color in alcohol gray to dirty white, with very numerous light to dark red irregularly shaped spots (
|
||
<figureCitation id="9CC972A2EF2DFFDFFAE2FDA597B5FD57" captionStart="FIGURE 1" captionStartId="2.[151,255,1866,1890]" captionTargetBox="[173,1409,320,1832]" captionTargetId="figure@2.[169,1417,317,1840]" captionTargetPageId="2" captionText="FIGURE 1. Heteromolpadia joyceae new species. A, Holotype, whole animal, total length 86 mm. B, scattered phosphatic deposits derived from dissolution of skin in liquid household bleach. C, Remnants of a spired table with three larger perforations and three smaller. D, Fragments of three-armed anchor plate. E, Spired tables from tail. F, spired tables from tail, higher magnification. Scale line in Figures 1 B-F = 100 Μm." httpUri="https://zenodo.org/record/273938/files/figure.png" pageId="3" pageNumber="38">Figure 1</figureCitation>
|
||
A), which are aggregations of phosphatic deposits, each spot approximately
|
||
<quantity id="C30AC3C2EF2DFFDFFB85FD8D9372FD57" box="[1065,1125,632,658]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="38" unit="mm" value="1.0">1mm</quantity>
|
||
in diameter. Spots closely aggregated towards anterior end of body, more scattered posteriorly. Tail grayish-white, lacking red spots. Oral disk typical, with 15 tentacles, the disk diameter approximately 10% of body length.
|
||
</paragraph>
|
||
<paragraph id="044D6E27EF2DFFDFFF6AFD059623FBE7" blockId="3.[151,1437,152,1978]" pageId="3" pageNumber="38">
|
||
In a
|
||
<quantity id="C30AC3C2EF2DFFDFFF51FD05965FFCCF" box="[253,328,752,778]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="38" unit="mm" value="90.0">90mm</quantity>
|
||
long
|
||
<typeStatus id="DB49D085EF2DFFDFFE22FD0596EFFCCF" box="[398,504,752,778]" pageId="3" pageNumber="38" type="paratype">Paratype</typeStatus>
|
||
, calcareous ring with interradial pieces
|
||
<quantity id="C30AC3C2EF2DFFDFFC7EFD059318FCCF" box="[978,1039,752,778]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="38" unit="mm" value="5.0">5mm</quantity>
|
||
high, radial pieces
|
||
<quantity id="C30AC3C2EF2DFFDFFB56FD059221FCCF" box="[1274,1334,752,778]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="38" unit="mm" value="7.0">7mm</quantity>
|
||
high, of which relatively short posterior projections occupy
|
||
<quantity id="C30AC3C2EF2DFFDFFCA0FCED945BFCF7" box="[780,844,792,818]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="38" unit="mm" value="2.0">2mm</quantity>
|
||
. Posterior projections with narrow notch 0.5mm deep. Polian vesicle single. Stone canal in mid-dorsal interradius whitish, approximately
|
||
<quantity id="C30AC3C2EF2DFFDFFAA9FCB59246FC9F" box="[1285,1361,832,858]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="38" unit="mm" value="7.0">7 mm</quantity>
|
||
long, madreporite approximately spherical, 0.5mm in diameter. Pore canal
|
||
<quantity id="C30AC3C2EF2DFFDFFC67FC9D9310FC47" box="[971,1031,872,898]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="38" unit="mm" value="1.0">1mm</quantity>
|
||
long extends from madreporite to attach to mid-dorsal body wall. Pore canal opens to exterior in an area distinguished by a bare gray patch approximately
|
||
<quantity id="C30AC3C2EF2DFFDFFEE5FC4D9683FC17" box="[329,404,952,978]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="38" unit="mm" value="10.0">10mm</quantity>
|
||
posterior to oral disk. Gonad with few branches, genital duct opens to exterior in middorsal interradius,
|
||
<quantity id="C30AC3C2EF2DFFDFFEDBFC1596A3FC3F" box="[375,436,992,1018]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="38" unit="mm" value="1.0">1mm</quantity>
|
||
posterior to oral disk. Gonad without ossicles. Course of intestine, and respiratory trees, typical.
|
||
</paragraph>
|
||
<paragraph id="044D6E27EF2DFFDFFF6AFBC593F1FAFF" blockId="3.[151,1437,152,1978]" pageId="3" pageNumber="38">
|
||
Body wall in all specimens contains primarily clusters of phosphatic deposits (
|
||
<figureCitation id="9CC972A2EF2DFFDFFBC3FBC593C5FB8F" box="[1135,1234,1072,1098]" captionStart="FIGURE 1" captionStartId="2.[151,255,1866,1890]" captionTargetBox="[173,1409,320,1832]" captionTargetId="figure@2.[169,1417,317,1840]" captionTargetPageId="2" captionText="FIGURE 1. Heteromolpadia joyceae new species. A, Holotype, whole animal, total length 86 mm. B, scattered phosphatic deposits derived from dissolution of skin in liquid household bleach. C, Remnants of a spired table with three larger perforations and three smaller. D, Fragments of three-armed anchor plate. E, Spired tables from tail. F, spired tables from tail, higher magnification. Scale line in Figures 1 B-F = 100 Μm." httpUri="https://zenodo.org/record/273938/files/figure.png" pageId="3" pageNumber="38">Figure 1</figureCitation>
|
||
B), derived from dissolution of calcareous ossicles. Rare remaining fragments of ossicles (
|
||
<figureCitation id="9CC972A2EF2DFFDFFC51FBAD937DFBB7" box="[1021,1130,1112,1138]" captionStart="FIGURE 1" captionStartId="2.[151,255,1866,1890]" captionTargetBox="[173,1409,320,1832]" captionTargetId="figure@2.[169,1417,317,1840]" captionTargetPageId="2" captionText="FIGURE 1. Heteromolpadia joyceae new species. A, Holotype, whole animal, total length 86 mm. B, scattered phosphatic deposits derived from dissolution of skin in liquid household bleach. C, Remnants of a spired table with three larger perforations and three smaller. D, Fragments of three-armed anchor plate. E, Spired tables from tail. F, spired tables from tail, higher magnification. Scale line in Figures 1 B-F = 100 Μm." httpUri="https://zenodo.org/record/273938/files/figure.png" pageId="3" pageNumber="38">Figures 1</figureCitation>
|
||
C–D) indicate that young specimens of this species carry in their body wall single anchor-plates that are apparently three-armed, along with typical molpadiid anchors. Fragments show that three-pillared tables of 100µm average diameter with three major perforations and often three smaller perforations are scattered in the body wall. Tail contains spired tables (
|
||
<figureCitation id="9CC972A2EF2DFFDFFE97FB0D968BFAD7" box="[315,412,1272,1298]" captionStart="FIGURE 1" captionStartId="2.[151,255,1866,1890]" captionTargetBox="[173,1409,320,1832]" captionTargetId="figure@2.[169,1417,317,1840]" captionTargetPageId="2" captionText="FIGURE 1. Heteromolpadia joyceae new species. A, Holotype, whole animal, total length 86 mm. B, scattered phosphatic deposits derived from dissolution of skin in liquid household bleach. C, Remnants of a spired table with three larger perforations and three smaller. D, Fragments of three-armed anchor plate. E, Spired tables from tail. F, spired tables from tail, higher magnification. Scale line in Figures 1 B-F = 100 Μm." httpUri="https://zenodo.org/record/273938/files/figure.png" pageId="3" pageNumber="38">Figure 1</figureCitation>
|
||
E–F) of average length 300µm, with an oval central area with 15–20 perforations, and two rod-like extensions, usually imperforate. The three-pillared spire averages 60ìm in height.
|
||
</paragraph>
|
||
<paragraph id="044D6E27EF2DFFDFFF6AFABD93ADFA4F" blockId="3.[151,1437,152,1978]" pageId="3" pageNumber="38">Distribution: Off the east coast of the North Island, and the northeast coast of the South Island, and the Chatham Rise, in 258–632 meters. Apparently this species is confined to the bathyal zone.</paragraph>
|
||
<paragraph id="044D6E27EF2DFFDFFF6AFA6D97C4FA1F" blockId="3.[151,1437,152,1978]" pageId="3" pageNumber="38">Ecology: Like all other molpadiids, prefers mud substrates. In all specimens, the intestine is packed with mud.</paragraph>
|
||
<paragraph id="044D6E27EF2DFFDFFF6AFA1D95D9F997" blockId="3.[151,1437,152,1978]" pageId="3" pageNumber="38">
|
||
Etymology: It is our pleasure to name this species for Mrs. Joyce McCullough of
|
||
<collectingCountry id="7CE52EB7EF2DFFDFFB00FA1D93E5F9C7" box="[1196,1266,1512,1538]" name="China" pageId="3" pageNumber="38">China</collectingCountry>
|
||
Grove, North Carolina, as a small token of appreciation of her untiring, endlessly cheerful, volunteer assistance in the field and in the laboratory over the past several years.
|
||
</paragraph>
|
||
<paragraph id="044D6E27EF2DFFDFFF6AF995935BF87F" blockId="3.[151,1437,152,1978]" pageId="3" pageNumber="38">
|
||
Remarks:
|
||
<bibRefCitation id="606313D6EF2DFFDFFE93F9959510F9BF" author="Pawson" box="[319,519,1632,1658]" pageId="3" pageNumber="39" refString="Pawson, D. L. (1965 a) The bathyal holothurians of the New Zealand region. Zoology Publications from Victoria University of Wellington 39, 1 - 33." type="journal article" year="1965" yearSuffix="a">Pawson (1965a)</bibRefCitation>
|
||
suspected that the two specimens he named “
|
||
<taxonomicName id="C3F215A4EF2DFFDFFB87F995938AF9BC" box="[1067,1181,1632,1657]" class="Holothuroidea" family="Molpadiidae" genus="Molpadia" kingdom="Animalia" order="Molpadida" pageId="3" pageNumber="38" phylum="Echinodermata" rank="genus" status="sp. nov.">
|
||
<emphasis id="3686B235EF2DFFDFFB87F995938AF9BC" box="[1067,1181,1632,1657]" italics="true" pageId="3" pageNumber="38">Molpadia</emphasis>
|
||
</taxonomicName>
|
||
?
|
||
<taxonomicNameLabel id="2DB50F4EEF2DFFDFFB1EF99593FDF9BF" box="[1202,1258,1632,1658]" pageId="3" pageNumber="38" rank="species">n.sp.</taxonomicNameLabel>
|
||
” represented a new species, but noted that more material was needed in order to properly characterize the species. The abundant material now available confirms the validity of this new taxon. In having single three-armed anchor plates in the body wall, this species falls into the genus
|
||
<taxonomicName id="C3F215A4EF2DFFDFFC80F92D938CF937" authority="Pawson, 1963" authorityName="Pawson" authorityYear="1963" box="[812,1179,1752,1778]" class="Holothuroidea" family="Molpadiidae" genus="Heteromolpadia" kingdom="Animalia" order="Molpadida" pageId="3" pageNumber="38" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="3686B235EF2DFFDFFC80F92D94FCF934" box="[812,1003,1752,1777]" italics="true" pageId="3" pageNumber="38">Heteromolpadia</emphasis>
|
||
<bibRefCitation id="606313D6EF2DFFDFFC5FF92D938CF937" author="Pawson" box="[1011,1179,1752,1778]" pageId="3" pageNumber="39" refString="Pawson, D. L. (1963) The holothurian fauna of Cook Strait, New Zealand. Zoology Publications from Victoria University of Wellington 36, 1 - 38." type="journal article" year="1963">Pawson, 1963</bibRefCitation>
|
||
</taxonomicName>
|
||
. From the Indonesian species
|
||
<taxonomicName id="C3F215A4EF2DFFDFFF5EF8F5950DF8DF" authority="Sluiter, 1901" authorityName="Sluiter" authorityYear="1901" box="[242,538,1792,1818]" class="Holothuroidea" family="Molpadiidae" genus="Heteromolpadia" kingdom="Animalia" order="Molpadida" pageId="3" pageNumber="38" phylum="Echinodermata" rank="species" species="tridens">
|
||
<emphasis id="3686B235EF2DFFDFFF5EF8F59670F8DC" box="[242,359,1792,1817]" italics="true" pageId="3" pageNumber="38">H. tridens</emphasis>
|
||
(
|
||
<bibRefCitation id="606313D6EF2DFFDFFEDAF8F59506F8DF" author="Sluiter" box="[374,529,1792,1818]" pageId="3" pageNumber="39" refString="Sluiter, C. P. (1901) Die Holothurien der Siboga-Expedition. Siboga Report 1, 1 - 142." type="book chapter" year="1901">Sluiter, 1901</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="C3F215A4EF2DFFDFFD89F8F595B5F8DC" box="[549,674,1792,1817]" class="Holothuroidea" family="Molpadiidae" genus="Heteromolpadia" kingdom="Animalia" order="Molpadida" pageId="3" pageNumber="38" phylum="Echinodermata" rank="species" species="joyceae">
|
||
<emphasis id="3686B235EF2DFFDFFD89F8F595B5F8DC" box="[549,674,1792,1817]" italics="true" pageId="3" pageNumber="38">H. joyceae</emphasis>
|
||
</taxonomicName>
|
||
differs in possessing spired tables in the body wall. Furthermore, in
|
||
<taxonomicName id="C3F215A4EF2DFFDFFF19F8DD963EF884" box="[181,297,1832,1857]" class="Holothuroidea" family="Molpadiidae" genus="Heteromolpadia" kingdom="Animalia" order="Molpadida" pageId="3" pageNumber="38" phylum="Echinodermata" rank="species" species="tridens">
|
||
<emphasis id="3686B235EF2DFFDFFF19F8DD963EF884" box="[181,297,1832,1857]" italics="true" pageId="3" pageNumber="38">H. tridens</emphasis>
|
||
</taxonomicName>
|
||
the three-armed ossicles and anchors are retained to some extent as the animal grows, and apparently do not undergo extensive dissolution.
|
||
<taxonomicName id="C3F215A4EF2DFFDFFD1CF8A59426F8AC" box="[688,817,1872,1897]" class="Holothuroidea" family="Molpadiidae" genus="Heteromolpadia" kingdom="Animalia" order="Molpadida" pageId="3" pageNumber="38" phylum="Echinodermata" rank="species" species="joyceae">
|
||
<emphasis id="3686B235EF2DFFDFFD1CF8A59426F8AC" box="[688,817,1872,1897]" italics="true" pageId="3" pageNumber="38">H. joyceae</emphasis>
|
||
</taxonomicName>
|
||
differs from the other
|
||
<collectingCountry id="7CE52EB7EF2DFFDFFBFCF8A593E6F8AF" box="[1104,1265,1872,1898]" name="New Zealand" pageId="3" pageNumber="38">New Zealand</collectingCountry>
|
||
congeners
|
||
<taxonomicName id="C3F215A4EF2DFFDFFAD3F8A596C7F857" authority="Theel, 1886" authorityName="Theel" authorityYear="1886" class="Holothuroidea" family="Molpadiidae" genus="Heteromolpadia" kingdom="Animalia" order="Molpadida" pageId="3" pageNumber="38" phylum="Echinodermata" rank="species" species="marenzelleri">
|
||
<emphasis id="3686B235EF2DFFDFFAD3F8A5963DF854" italics="true" pageId="3" pageNumber="38">H. marenzelleri</emphasis>
|
||
(
|
||
<bibRefCitation id="606313D6EF2DFFDFFE95F88D96DFF857" author="Theel" box="[313,456,1912,1938]" pageId="3" pageNumber="39" refString="Theel, H. (1886) Report on the Holothurioidea. 2. Challenger Scientific Results. Zoology, 14, 1 - 290." type="journal article" year="1886">Théel, 1886</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="C3F215A4EF2DFFDFFDA7F88D9401F857" authority="Pawson, 1965" authorityName="Pawson" authorityYear="1965" box="[523,790,1912,1938]" class="Holothuroidea" family="Molpadiidae" genus="Heteromolpadia" kingdom="Animalia" order="Molpadida" pageId="3" pageNumber="38" phylum="Echinodermata" rank="species" species="pikei">
|
||
<emphasis id="3686B235EF2DFFDFFDA7F88D957FF854" box="[523,616,1912,1937]" italics="true" pageId="3" pageNumber="38">H. pikei</emphasis>
|
||
<bibRefCitation id="606313D6EF2DFFDFFDC3F88D9401F857" author="Pawson" box="[623,790,1912,1938]" pageId="3" pageNumber="39" refString="Pawson, D. L. (1965 b) New sea-cucumbers (Echinodermata: Holothuroidea) from New Zealand waters. Records of the Dominion Museum Wellington 5 (11), 75 - 82." type="journal article" year="1965" yearSuffix="b">Pawson, 1965</bibRefCitation>
|
||
</taxonomicName>
|
||
b, in the nature of the ossicles in the tail; these are generally larger in
|
||
<taxonomicName id="C3F215A4EF2DFFDFFEE6F85596DFF87C" box="[330,456,1952,1977]" class="Holothuroidea" family="Molpadiidae" genus="Heteromolpadia" kingdom="Animalia" order="Molpadida" pageId="3" pageNumber="38" phylum="Echinodermata" rank="species" species="joyceae">
|
||
<emphasis id="3686B235EF2DFFDFFEE6F85596DFF87C" box="[330,456,1952,1977]" italics="true" pageId="3" pageNumber="38">H. joyceae</emphasis>
|
||
</taxonomicName>
|
||
, with far more numerous perforations (see key above).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |