320 lines
46 KiB
XML
320 lines
46 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.4813018" ID-GBIF-Dataset="a114ac18-a46f-456b-9052-1bbd63fa252d" ID-GBIF-Taxon="182398467" ID-Zenodo-Dep="4813018" approvedBy="tatiana" checkinTime="1622040502204" checkinUser="tatiana" docAuthor="Wang, Zhen, Olmi, Massimo, Shih, Chungkun, Ren, Dong & Gao, Taiping" docDate="2021" docId="03FB3512C47951108A14F8B006A55A8F" docLanguage="en" docName="cretaceousResearch.125.104861.pdf" docOrigin="Cretaceous Research 125, No. 104861" docTitle="Hybristodryinus viriosus Wang, Olmi and Gao 2021, sp. nov." docType="treatment" docVersion="3" lastPageId="3" lastPageNumber="4" masterDocId="FFC24D6AC47851138A70FFA203755F60" masterDocTitle="New species of Hybristodryinus from mid-Cretaceous Myanmar amber (Hymenoptera, Dryinidae)" masterLastPageNumber="6" masterPageNumber="1" pageId="1" pageNumber="2" updateTime="1643437438995" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New species of Hybristodryinus from mid-Cretaceous Myanmar amber (Hymenoptera, Dryinidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wang, Zhen</mods:namePart>
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<mods:affiliation>College of Life Sciences and Academy for Multidisciplinary Studies, Capital Normal University, 105 Xisanhuanbeilu, Haidian District, Beijing, 100048, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Olmi, Massimo</mods:namePart>
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<mods:affiliation>Tropical Entomology Research Center, Via De Gasperi 10, I- 01100 Viterbo, Italy</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Shih, Chungkun</mods:namePart>
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<mods:affiliation>College of Life Sciences and Academy for Multidisciplinary Studies, Capital Normal University, 105 Xisanhuanbeilu, Haidian District, Beijing, 100048, China & Department of Paleobiology, National Museum of Natural History, Smithsonian Institution, Washington, DC 20013 - 7012, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ren, Dong</mods:namePart>
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<mods:affiliation>College of Life Sciences and Academy for Multidisciplinary Studies, Capital Normal University, 105 Xisanhuanbeilu, Haidian District, Beijing, 100048, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Gao, Taiping</mods:namePart>
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<mods:affiliation>College of Life Sciences and Academy for Multidisciplinary Studies, Capital Normal University, 105 Xisanhuanbeilu, Haidian District, Beijing, 100048, China</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Cretaceous Research</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-04-23</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>125</mods:number>
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</mods:detail>
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<mods:detail type="numero">
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<mods:number>104861</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>6</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1016/j.cretres.2021.104861</mods:identifier>
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<mods:identifier type="GBIF-Dataset">a114ac18-a46f-456b-9052-1bbd63fa252d</mods:identifier>
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<mods:identifier type="Zenodo-Dep">4812995</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.4813018" ID-GBIF-Taxon="182398467" ID-Zenodo-Dep="4813018" LSID="urn:lsid:plazi:treatment:03FB3512C47951108A14F8B006A55A8F" httpUri="http://treatment.plazi.org/id/03FB3512C47951108A14F8B006A55A8F" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2">
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<subSubSection box="[100,662,1810,1830]" pageId="1" pageNumber="2" type="nomenclature">
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<paragraph blockId="1.[100,770,1810,1886]" box="[100,662,1810,1830]" pageId="1" pageNumber="2">
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<taxonomicName authority="Wang, Olmi and Gao" authorityName="Wang, Olmi and Gao" authorityYear="2021" box="[100,575,1810,1830]" class="Insecta" family="Dryinidae" genus="Hybristodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="viriosus" status="sp. nov.">
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<emphasis bold="true" box="[100,357,1810,1830]" italics="true" pageId="1" pageNumber="2">Hybristodryinus viriosus</emphasis>
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Wang, Olmi and Gao
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</taxonomicName>
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,
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<taxonomicNameLabel box="[585,662,1811,1830]" pageId="1" pageNumber="2" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="2" type="description">
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<paragraph blockId="1.[100,770,1810,1886]" pageId="1" pageNumber="2">LSID: urn:lsid:zoobank.org:act:79A30209-7D6E-4BAE-BA8B-31F7F 457D301</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="2" type="etymology">
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<paragraph blockId="1.[100,770,1908,1984]" pageId="1" pageNumber="2">
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<emphasis box="[100,207,1908,1928]" italics="true" pageId="1" pageNumber="2">Etymology.</emphasis>
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The species is named after the Latin adjective
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<taxonomicName authorityName="Wang, Olmi and Gao" authorityYear="2021" box="[695,770,1908,1928]" class="Insecta" family="Dryinidae" genus="Hybristodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="viriosus">
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<emphasis box="[695,770,1908,1928]" italics="true" pageId="1" pageNumber="2">viriosus</emphasis>
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</taxonomicName>
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(meaning, “strong”).
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</paragraph>
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</subSubSection>
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<subSubSection box="[100,658,1963,1984]" pageId="1" pageNumber="2" type="materials_examined">
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<paragraph blockId="1.[100,770,1908,1984]" box="[100,658,1963,1984]" pageId="1" pageNumber="2">
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<emphasis box="[100,188,1963,1983]" italics="true" pageId="1" pageNumber="2">Material.</emphasis>
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<materialsCitation ID-GBIF-Occurrence="3118973301" box="[194,658,1964,1984]" country="Afghanistan" pageId="1" pageNumber="2" specimenCode="CNU-HYM-MA2015109" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
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<typeStatus box="[194,291,1964,1983]" pageId="1" pageNumber="2">Holotype</typeStatus>
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:
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<specimenCount box="[300,370,1964,1983]" pageId="1" pageNumber="2" type="female">female</specimenCount>
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. No.
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<specimenCode box="[420,654,1964,1983]" pageId="1" pageNumber="2">CNU-HYM-MA2015109</specimenCode>
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.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="2" type="diagnosis">
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<paragraph blockId="1.[818,1488,177,700]" pageId="1" pageNumber="2">
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<emphasis box="[818,917,177,197]" italics="true" pageId="1" pageNumber="2">Diagnosis.</emphasis>
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Female of
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<taxonomicName authorityName="Engel" authorityYear="2005" box="[1020,1175,177,197]" class="Insecta" family="Dryinidae" genus="Hybristodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
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<emphasis box="[1020,1175,177,197]" italics="true" pageId="1" pageNumber="2">Hybristodryinus</emphasis>
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</taxonomicName>
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with occipital carina complete; antenna very slender, more than six times as long as head, reaching about second segment of metasoma; first flagellomere 19 times as long as broad; maxillary palpus very slender, with palpomere 4 about 20 times as long as broad.
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</paragraph>
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<paragraph blockId="1.[818,1488,177,700]" pageId="1" pageNumber="2">
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<emphasis box="[818,1040,317,337]" italics="true" pageId="1" pageNumber="2">
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<typeStatus box="[818,864,317,337]" pageId="1" pageNumber="2">Type</typeStatus>
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locality and horiz
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</emphasis>
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on.
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<materialsCitation ID-GBIF-Occurrence="3118973302" country="Myanmar" location="Hukawng Valley" pageId="1" pageNumber="2" specimenCount="1" stateProvince="Kachin">
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The amber specimen was collected from
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<collectingRegion box="[818,887,346,365]" pageId="1" pageNumber="2">Kachin</collectingRegion>
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(
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<location LSID="urn:lsid:plazi:treatment:03FB3512C47951108A14F8B006A55A8F:8E8DD2DFC479511289F4FEF8075B5E0D" box="[900,1070,346,365]" country="Myanmar" name="Hukawng Valley" pageId="1" pageNumber="2" stateProvince="Kachin">Hukawng Valley</location>
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) of northern
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<collectingCountry box="[1206,1303,346,365]" name="Myanmar" pageId="1" pageNumber="2" value="Myanmar">Myanmar</collectingCountry>
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, which was dated at 98.79 ± 0.62 Ma, earliest Cenomanian (
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<bibRefCitation author="Cruikshank, R. D. & Ko, K." box="[1230,1479,374,393]" journalOrPublisher="Journal of Asian Earth Sciences" pageId="1" pageNumber="2" pagination="441 ‾ 455" part="21" refId="ref4915" refString="Cruikshank, R. D., Ko, K., 2003. Geology of an amber locality in the Hukawng Valley, northern Myanmar. Journal of Asian Earth Sciences 21, 441 ‾ 455. https: // doi. org / 10.1016 / S 1367 - 9120 (02) 00044 - 5." title="Geology of an amber locality in the Hukawng Valley, northern Myanmar" type="journal article" year="2003">Cruikshank and Ko, 2003</bibRefCitation>
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;
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<bibRefCitation author="Shi, G. & Grimaldi, D. A. & Harlow, G. E. & Wang, J. & Wang, J. & Yang, M. & Lei, W. & Li, Q. & Li, X." box="[818,965,401,421]" journalOrPublisher="Cretaceous Research" pageId="1" pageNumber="2" pagination="155 ‾ 163" part="37" refId="ref6777" refString="Shi, G., Grimaldi, D. A., Harlow, G. E., Wang, J., Wang, J., Yang, M., Lei, W., Li, Q., Li, X., 2012. Age constraint on Burmese amber based on UePb dating of zircons. Cretaceous Research 37, 155 ‾ 163. https: // doi. org / 10.1016 / j. cretres. 2012.03.014." title="Age constraint on Burmese amber based on UePb dating of zircons" type="journal article" year="2012">Shi et al., 2012</bibRefCitation>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2" type="description">
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<paragraph blockId="1.[818,1488,177,700]" pageId="1" pageNumber="2">
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<emphasis box="[818,933,428,448]" italics="true" pageId="1" pageNumber="2">Description.</emphasis>
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Female (
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<figureCitation box="[1040,1110,429,449]" captionStart="Fig" captionStartId="2.[100,125,907,921]" captionTargetBox="[110,1469,172,874]" captionTargetId="figure-0@2.[106,1482,166,878]" captionTargetPageId="2" captionText="Fig. 1. Photomicrographs and line drawing of Hybristodryinus viriosus Wang, Olmi and Gao, sp. nov., holotype ♀ CNU-HYM-MA2015109 preserved in Cenomanian-aged Myanmar (Burmese) amber. A. Ventral view. B. Dorsal view. C. Line drawing in ventral view. Antennae and legs are artificially extended from the body for easy recognition and comparison. Scale bars: 1.0 mm for A, B and C." figureDoi="http://doi.org/10.5281/zenodo.4812997" httpUri="https://zenodo.org/record/4812997/files/figure.png" pageId="1" pageNumber="2">Figs. 1</figureCitation>
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<emphasis bold="true" box="[1110,1125,429,448]" italics="true" pageId="1" pageNumber="2">̅</emphasis>
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<figureCitation box="[1125,1138,430,449]" captionStart="Fig" captionStartId="3.[100,125,919,933]" captionTargetBox="[110,1478,181,887]" captionTargetId="figure-512@3.[106,1482,178,890]" captionTargetPageId="3" captionText="Fig. 3. Photomicrographs and line drawings of Hybristodryinus viriosus Wang, Olmi and Gao, sp. nov., holotype ♀ CNU-HYM-MA2015109.A. Ventral view of mesosoma and fore leg. B. Right chela. C. Line drawing of right chela. D. Forewing venation. E. Line drawing of forewing venation. Abbreviations: a, arolium; ec, enlarged claw; rc, rudimentary claw; T4, protarsomere 4; T5, protarsomere 5; Pt, Pterostigma. Scale bars: 0.4 mm for A, Band C; 1.0 mm for Eand F." figureDoi="http://doi.org/10.5281/zenodo.4813001" httpUri="https://zenodo.org/record/4813001/files/figure.png" pageId="1" pageNumber="2">3</figureCitation>
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). Fully winged; length
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<quantity box="[1402,1487,429,449]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="1" pageNumber="2" unit="mm" value="6.7">6.7 mm</quantity>
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(
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<figureCitation box="[826,986,457,476]" captionStart="Fig" captionStartId="2.[100,125,907,921]" captionTargetBox="[110,1469,172,874]" captionTargetId="figure-0@2.[106,1482,166,878]" captionTargetPageId="2" captionText="Fig. 1. Photomicrographs and line drawing of Hybristodryinus viriosus Wang, Olmi and Gao, sp. nov., holotype ♀ CNU-HYM-MA2015109 preserved in Cenomanian-aged Myanmar (Burmese) amber. A. Ventral view. B. Dorsal view. C. Line drawing in ventral view. Antennae and legs are artificially extended from the body for easy recognition and comparison. Scale bars: 1.0 mm for A, B and C." figureDoi="http://doi.org/10.5281/zenodo.4812997" httpUri="https://zenodo.org/record/4812997/files/figure.png" pageId="1" pageNumber="2">Figs. 1A, 1B, 1C</figureCitation>
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,
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<figureCitation box="[1001,1031,457,476]" captionStart="Fig" captionStartId="2.[100,125,1940,1954]" captionTargetBox="[110,1479,1011,1907]" captionTargetId="figure-62@2.[106,1482,1007,1910]" captionTargetPageId="2" captionText="Fig. 2. Photomicrographs of Hybristodryinus viriosus Wang, Olmi and Gao, sp. nov., holotype ♀ CNU-HYM-MA2015109. A. Lateral view. B. Lateral view of head and mesosoma. C. Ventral view of head.D. Dorsal view of head and mesosoma. E. Anterior view of head. Abbreviations: mp, maxillary palpus; oc, occipital carina. Scale bars:1.0 mm for A; 0.4 mm for B, C, D and E." figureDoi="http://doi.org/10.5281/zenodo.4812999" httpUri="https://zenodo.org/record/4812999/files/figure.png" pageId="1" pageNumber="2">2A</figureCitation>
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). Head black, except clypeus and mandible testaceous; antenna testaceous; mesosoma black; metasoma brown; legs testaceous. Antenna 10-segmented, clavate, very slender, apparently hairless; lengths of antennomeres in following proportions: 20:8:46:36:29:19:14:13:10:12; antenna more than six times as long as head (200:33), reaching about second segment of metasoma. ADO's not visible. Flagellomeres much longer than wide; antennomere 9 about five times as long as broad (10:2); antennomere 3 about 19 times as long as broad (46:2.4).
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</paragraph>
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<paragraph blockId="1.[818,1488,736,1983]" pageId="1" pageNumber="2">
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Head slightly convex, with sculpture not distinct; clypeus with anterior margin rounded (
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<figureCitation box="[1094,1263,764,783]" captionStart="Fig" captionStartId="2.[100,125,1940,1954]" captionTargetBox="[110,1479,1011,1907]" captionTargetId="figure-62@2.[106,1482,1007,1910]" captionTargetPageId="2" captionText="Fig. 2. Photomicrographs of Hybristodryinus viriosus Wang, Olmi and Gao, sp. nov., holotype ♀ CNU-HYM-MA2015109. A. Lateral view. B. Lateral view of head and mesosoma. C. Ventral view of head.D. Dorsal view of head and mesosoma. E. Anterior view of head. Abbreviations: mp, maxillary palpus; oc, occipital carina. Scale bars:1.0 mm for A; 0.4 mm for B, C, D and E." figureDoi="http://doi.org/10.5281/zenodo.4812999" httpUri="https://zenodo.org/record/4812999/files/figure.png" pageId="1" pageNumber="2">Figs. 2C, 2D, 2E</figureCitation>
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); mandible 4-dentate (progressively larger from dorsal to ventral tooth); frons slightly convex, not longitudinally excavated medially, with transverse fold situated half way, not reaching eyes laterally; occipital carina complete, bordered by furrow, not reaching eyes laterally; vertex without longitudinal keels connecting lateral ocelli to occipital carina; occiput deeply excavated; eye bulging, bare, much shorter than head (22:33); POL = 13; OL = 5 (but head partly crushed, so this measure is the average between right and left OL); OOL = 15; TL = 8; OPL = 4; greatest breadth of lateral ocelli much shorter than POL (4:13); frontal line absent. Maxillary palpus with six palpomeres, number of labial palpomeres not observable, palpal formula probably 6/?. Maxillary palpus very slender, with palpomere 4 about 20 times as long as broad (10:0.5).
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</paragraph>
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<paragraph blockId="1.[818,1488,736,1983]" pageId="1" pageNumber="2">
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Pronotum with deep anterior transverse impression between anterior collar and disc; disc humped, trapezoidal, not isosceles triangle shaped (
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<figureCitation box="[989,1108,1210,1230]" captionStart="Fig" captionStartId="2.[100,125,1940,1954]" captionTargetBox="[110,1479,1011,1907]" captionTargetId="figure-62@2.[106,1482,1007,1910]" captionTargetPageId="2" captionText="Fig. 2. Photomicrographs of Hybristodryinus viriosus Wang, Olmi and Gao, sp. nov., holotype ♀ CNU-HYM-MA2015109. A. Lateral view. B. Lateral view of head and mesosoma. C. Ventral view of head.D. Dorsal view of head and mesosoma. E. Anterior view of head. Abbreviations: mp, maxillary palpus; oc, occipital carina. Scale bars:1.0 mm for A; 0.4 mm for B, C, D and E." figureDoi="http://doi.org/10.5281/zenodo.4812999" httpUri="https://zenodo.org/record/4812999/files/figure.png" pageId="1" pageNumber="2">Figs. 2C, 2D</figureCitation>
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), not deeply longitudinally excavated medially; posterior collar absent; anterior surface of pronotum without longitudinal striae; sculpture of disc indistinct; pronotum longer than head (43:33); pronotal tubercle reaching tegula. Epicnemium present. Mesoscutum smooth, slightly shorter than pronotum (38:43). Notauli almost reaching posterior margin of mesoscutum (
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<figureCitation box="[960,1032,1378,1397]" captionStart="Fig" captionStartId="2.[100,125,1940,1954]" captionTargetBox="[110,1479,1011,1907]" captionTargetId="figure-62@2.[106,1482,1007,1910]" captionTargetPageId="2" captionText="Fig. 2. Photomicrographs of Hybristodryinus viriosus Wang, Olmi and Gao, sp. nov., holotype ♀ CNU-HYM-MA2015109. A. Lateral view. B. Lateral view of head and mesosoma. C. Ventral view of head.D. Dorsal view of head and mesosoma. E. Anterior view of head. Abbreviations: mp, maxillary palpus; oc, occipital carina. Scale bars:1.0 mm for A; 0.4 mm for B, C, D and E." figureDoi="http://doi.org/10.5281/zenodo.4812999" httpUri="https://zenodo.org/record/4812999/files/figure.png" pageId="1" pageNumber="2">Fig. 2D</figureCitation>
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); minimum distance between notauli slightly shorter than POL (10:13). Mesoscutellum with sculpture indistinct, shorter than mesoscutum (17:38). Metanotum shorter than scutellum (9:17), with sculpture indistinct. Metapectal-propodeal complex shorter than mesoscutum (30:38), reticulate rugose, areolae very wide, disc with posterior corners strongly projected posteriorly (
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<figureCitation box="[943,1020,1545,1564]" captionStart="Fig" captionStartId="2.[100,125,907,921]" captionTargetBox="[110,1469,172,874]" captionTargetId="figure-0@2.[106,1482,166,878]" captionTargetPageId="2" captionText="Fig. 1. Photomicrographs and line drawing of Hybristodryinus viriosus Wang, Olmi and Gao, sp. nov., holotype ♀ CNU-HYM-MA2015109 preserved in Cenomanian-aged Myanmar (Burmese) amber. A. Ventral view. B. Dorsal view. C. Line drawing in ventral view. Antennae and legs are artificially extended from the body for easy recognition and comparison. Scale bars: 1.0 mm for A, B and C." figureDoi="http://doi.org/10.5281/zenodo.4812997" httpUri="https://zenodo.org/record/4812997/files/figure.png" pageId="1" pageNumber="2">Figs. 1B</figureCitation>
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,
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<figureCitation box="[1032,1063,1545,1564]" captionStart="Fig" captionStartId="2.[100,125,1940,1954]" captionTargetBox="[110,1479,1011,1907]" captionTargetId="figure-62@2.[106,1482,1007,1910]" captionTargetPageId="2" captionText="Fig. 2. Photomicrographs of Hybristodryinus viriosus Wang, Olmi and Gao, sp. nov., holotype ♀ CNU-HYM-MA2015109. A. Lateral view. B. Lateral view of head and mesosoma. C. Ventral view of head.D. Dorsal view of head and mesosoma. E. Anterior view of head. Abbreviations: mp, maxillary palpus; oc, occipital carina. Scale bars:1.0 mm for A; 0.4 mm for B, C, D and E." figureDoi="http://doi.org/10.5281/zenodo.4812999" httpUri="https://zenodo.org/record/4812999/files/figure.png" pageId="1" pageNumber="2">2A</figureCitation>
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); propodeal declivity shorter than disc of metapectal-propodeal complex (10:20), indistinct, because hidden behind disc.
|
|
</paragraph>
|
|
<paragraph blockId="1.[818,1488,736,1983]" lastBlockId="3.[100,771,1053,1602]" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2">
|
|
Fore leg ratio: 38 (coxa): 41 (trochanter): 60 (femur): 70 (tibia): 38 (tarsomere 1): 5 (tarsomere 2): 8 (tarsomere 3): 40 (tarsomere 4): 51 (tarsomere 5); enlarged claw shorter than protarsomere 5 (37:51) (
|
|
<figureCitation box="[1078,1199,1713,1732]" captionStart="Fig" captionStartId="2.[100,125,907,921]" captionTargetBox="[110,1469,172,874]" captionTargetId="figure-0@2.[106,1482,166,878]" captionTargetPageId="2" captionText="Fig. 1. Photomicrographs and line drawing of Hybristodryinus viriosus Wang, Olmi and Gao, sp. nov., holotype ♀ CNU-HYM-MA2015109 preserved in Cenomanian-aged Myanmar (Burmese) amber. A. Ventral view. B. Dorsal view. C. Line drawing in ventral view. Antennae and legs are artificially extended from the body for easy recognition and comparison. Scale bars: 1.0 mm for A, B and C." figureDoi="http://doi.org/10.5281/zenodo.4812997" httpUri="https://zenodo.org/record/4812997/files/figure.png" pageId="1" pageNumber="2">Figs. 1A, 1C</figureCitation>
|
|
,
|
|
<figureCitation box="[1213,1326,1713,1732]" captionStart="Fig" captionStartId="3.[100,125,919,933]" captionTargetBox="[110,1478,181,887]" captionTargetId="figure-512@3.[106,1482,178,890]" captionTargetPageId="3" captionText="Fig. 3. Photomicrographs and line drawings of Hybristodryinus viriosus Wang, Olmi and Gao, sp. nov., holotype ♀ CNU-HYM-MA2015109.A. Ventral view of mesosoma and fore leg. B. Right chela. C. Line drawing of right chela. D. Forewing venation. E. Line drawing of forewing venation. Abbreviations: a, arolium; ec, enlarged claw; rc, rudimentary claw; T4, protarsomere 4; T5, protarsomere 5; Pt, Pterostigma. Scale bars: 0.4 mm for A, B and C; 1.0 mm for E and F." figureDoi="http://doi.org/10.5281/zenodo.4813001" httpUri="https://zenodo.org/record/4813001/files/figure.png" pageId="1" pageNumber="2">3A, 3B, 3C</figureCitation>
|
|
). Protrochanter very slender and long, with long and slender proximal stalk, broadened after half-way; protrochanter about 13 times as long as broad (41:3.2); protarsomere 3 produced into hook; rudimentary claw present, much shorter than enlarged claw (7:37); arolium much shorter than enlarged claw (6:37); enlarged claw with distal apex sharp, with two subapical teeth and one row of small lamellae (number of lamellae not distinct). Protarsomere 5 with two rows composed of many lamellae, one row formed by longer lamellae (number of lamellae indistinct); distal apex with group of lamellae (number of lamellae indistinct). Mid leg ratio (
|
|
<figureCitation captionStart="Fig" captionStartId="2.[100,125,907,921]" captionTargetBox="[110,1469,172,874]" captionTargetId="figure-0@2.[106,1482,166,878]" captionTargetPageId="2" captionText="Fig. 1. Photomicrographs and line drawing of Hybristodryinus viriosus Wang, Olmi and Gao, sp. nov., holotype ♀ CNU-HYM-MA2015109 preserved in Cenomanian-aged Myanmar (Burmese) amber. A. Ventral view. B. Dorsal view. C. Line drawing in ventral view. Antennae and legs are artificially extended from the body for easy recognition and comparison. Scale bars: 1.0 mm for A, B and C." figureDoi="http://doi.org/10.5281/zenodo.4812997" httpUri="https://zenodo.org/record/4812997/files/figure.png" pageId="3" pageNumber="4">Figs. 1A, 1C</figureCitation>
|
|
,
|
|
<figureCitation box="[143,175,1081,1100]" captionStart="Fig" captionStartId="2.[100,125,1940,1954]" captionTargetBox="[110,1479,1011,1907]" captionTargetId="figure-62@2.[106,1482,1007,1910]" captionTargetPageId="2" captionText="Fig. 2. Photomicrographs of Hybristodryinus viriosus Wang, Olmi and Gao, sp. nov., holotype ♀ CNU-HYM-MA2015109. A. Lateral view. B. Lateral view of head and mesosoma. C. Ventral view of head.D. Dorsal view of head and mesosoma. E. Anterior view of head. Abbreviations: mp, maxillary palpus; oc, occipital carina. Scale bars:1.0 mm for A; 0.4 mm for B, C, D and E." figureDoi="http://doi.org/10.5281/zenodo.4812999" httpUri="https://zenodo.org/record/4812999/files/figure.png" pageId="3" pageNumber="4">2A</figureCitation>
|
|
): 28 (coxa): 30 (trochanter): 56 (femur): 56 (tibia): 40 (tarsomere 1): 17 (tarsomere 2): 11 (tarsomere 3): 7 (tarsomere 4): 10 (tarsomere 5). Mesotrochanter short and broad, less than three times as long as broad: 15:6. Hind leg ratio (
|
|
<figureCitation box="[605,716,1164,1183]" captionStart="Fig" captionStartId="2.[100,125,907,921]" captionTargetBox="[110,1469,172,874]" captionTargetId="figure-0@2.[106,1482,166,878]" captionTargetPageId="2" captionText="Fig. 1. Photomicrographs and line drawing of Hybristodryinus viriosus Wang, Olmi and Gao, sp. nov., holotype ♀ CNU-HYM-MA2015109 preserved in Cenomanian-aged Myanmar (Burmese) amber. A. Ventral view. B. Dorsal view. C. Line drawing in ventral view. Antennae and legs are artificially extended from the body for easy recognition and comparison. Scale bars: 1.0 mm for A, B and C." figureDoi="http://doi.org/10.5281/zenodo.4812997" httpUri="https://zenodo.org/record/4812997/files/figure.png" pageId="3" pageNumber="4">Figs.1A, 1C</figureCitation>
|
|
,
|
|
<figureCitation box="[727,759,1164,1183]" captionStart="Fig" captionStartId="2.[100,125,1940,1954]" captionTargetBox="[110,1479,1011,1907]" captionTargetId="figure-62@2.[106,1482,1007,1910]" captionTargetPageId="2" captionText="Fig. 2. Photomicrographs of Hybristodryinus viriosus Wang, Olmi and Gao, sp. nov., holotype ♀ CNU-HYM-MA2015109. A. Lateral view. B. Lateral view of head and mesosoma. C. Ventral view of head.D. Dorsal view of head and mesosoma. E. Anterior view of head. Abbreviations: mp, maxillary palpus; oc, occipital carina. Scale bars:1.0 mm for A; 0.4 mm for B, C, D and E." figureDoi="http://doi.org/10.5281/zenodo.4812999" httpUri="https://zenodo.org/record/4812999/files/figure.png" pageId="3" pageNumber="4">2A</figureCitation>
|
|
): 44 (coxa): 13 (trochanter): 67 (femur): 50 (tibia): 46 (tarsomere 1): 24 (tarsomere 2): 16 (tarsomere 3): 9 (tarsomere 4): 16 (tarsomere 5).
|
|
</paragraph>
|
|
<caption ID-DOI="http://doi.org/10.5281/zenodo.4812997" ID-Zenodo-Dep="4812997" httpUri="https://zenodo.org/record/4812997/files/figure.png" pageId="2" pageNumber="3" startId="2.[100,125,907,921]" targetBox="[110,1469,172,874]" targetPageId="2">
|
|
<paragraph blockId="2.[100,1489,907,967]" pageId="2" pageNumber="3">
|
|
<emphasis bold="true" box="[100,146,907,921]" pageId="2" pageNumber="3">Fig. 1.</emphasis>
|
|
Photomicrographs and line drawing of
|
|
<taxonomicName authority="Wang, Olmi and Gao" authorityName="Wang, Olmi and Gao" authorityYear="2021" box="[462,813,907,922]" class="Insecta" family="Dryinidae" genus="Hybristodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="viriosus" status="sp. nov.">
|
|
<emphasis box="[462,644,907,922]" italics="true" pageId="2" pageNumber="3">Hybristodryinus viriosus</emphasis>
|
|
Wang, Olmi and Gao
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicNameLabel box="[821,883,908,922]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
|
|
, holotype ♀ CNU-HYM-MA2015109 preserved in Cenomanian-aged Myanmar (Burmese) amber. A. Ventral view. B. Dorsal view. C. Line drawing in ventral view. Antennae and legs are artificially extended from the body for easy recognition and comparison. Scale bars: 1.0 mm for A, Band C.
|
|
</paragraph>
|
|
</caption>
|
|
<caption ID-DOI="http://doi.org/10.5281/zenodo.4812999" ID-Zenodo-Dep="4812999" httpUri="https://zenodo.org/record/4812999/files/figure.png" pageId="2" pageNumber="3" startId="2.[100,125,1940,1954]" targetBox="[110,1479,1011,1907]" targetPageId="2">
|
|
<paragraph blockId="2.[100,1489,1939,2000]" pageId="2" pageNumber="3">
|
|
<emphasis bold="true" box="[100,149,1940,1954]" pageId="2" pageNumber="3">Fig. 2.</emphasis>
|
|
Photomicrographs of
|
|
<taxonomicName authority="Wang, Olmi and Gao" authorityName="Wang, Olmi and Gao" authorityYear="2021" box="[326,679,1939,1954]" class="Insecta" family="Dryinidae" genus="Hybristodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="viriosus" status="sp. nov.">
|
|
<emphasis box="[326,509,1939,1954]" italics="true" pageId="2" pageNumber="3">Hybristodryinus viriosus</emphasis>
|
|
Wang, Olmi and Gao
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicNameLabel box="[688,750,1940,1954]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
|
|
, holotype ♀ CNU-HYM-MA2015109. A. Lateral view. B. Lateral view of head and mesosoma. C. Ventral view of head. D. Dorsal view of head and mesosoma. E. Anterior view of head. Abbreviations: mp, maxillary palpus; oc, occipital carina. Scale bars: 1.0 mm for A; 0.4 mm for B, C, Dand E.
|
|
</paragraph>
|
|
</caption>
|
|
<caption ID-DOI="http://doi.org/10.5281/zenodo.4813001" ID-Zenodo-Dep="4813001" httpUri="https://zenodo.org/record/4813001/files/figure.png" pageId="3" pageNumber="4" startId="3.[100,125,919,933]" targetBox="[110,1478,181,887]" targetPageId="3">
|
|
<paragraph blockId="3.[100,1489,919,979]" pageId="3" pageNumber="4">
|
|
<emphasis bold="true" box="[100,148,919,933]" pageId="3" pageNumber="4">Fig. 3.</emphasis>
|
|
Photomicrographs and line drawings of
|
|
<taxonomicName authority="Wang, Olmi and Gao" authorityName="Wang, Olmi and Gao" authorityYear="2021" box="[466,813,919,934]" class="Insecta" family="Dryinidae" genus="Hybristodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="viriosus" status="sp. nov.">
|
|
<emphasis box="[466,648,919,934]" italics="true" pageId="3" pageNumber="4">Hybristodryinus viriosus</emphasis>
|
|
Wang, Olmi and Gao
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicNameLabel box="[820,881,920,934]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
|
, holotype ♀ CNU-HYM-MA2015109. A. Ventral view of mesosoma and fore leg. B. Right chela. C. Line drawing of right chela. D. Forewing venation. E. Line drawing of forewing venation. Abbreviations: a, arolium; ec, enlarged claw; rc, rudimentary claw; T4, protarsomere 4; T5, protarsomere 5; Pt, Pterostigma. Scale bars: 0.4 mm for A, Band C; 1.0 mm for Eand F.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph blockId="3.[100,771,1053,1602]" pageId="3" pageNumber="4">
|
|
Forewing hyaline, not darkened, with the usual venation of
|
|
<taxonomicName authorityName="Haliday" authorityYear="1833" box="[100,199,1304,1323]" class="Insecta" family="Dryinidae" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="subFamily" subFamily="Dryininae">Dryininae</taxonomicName>
|
|
(
|
|
<figureCitation box="[212,316,1304,1323]" captionStart="Fig" captionStartId="3.[100,125,919,933]" captionTargetBox="[110,1478,181,887]" captionTargetId="figure-512@3.[106,1482,178,890]" captionTargetPageId="3" captionText="Fig. 3. Photomicrographs and line drawings of Hybristodryinus viriosus Wang, Olmi and Gao, sp. nov., holotype ♀ CNU-HYM-MA2015109.A. Ventral view of mesosoma and fore leg. B. Right chela. C. Line drawing of right chela. D. Forewing venation. E. Line drawing of forewing venation. Abbreviations: a, arolium; ec, enlarged claw; rc, rudimentary claw; T4, protarsomere 4; T5, protarsomere 5; Pt, Pterostigma. Scale bars: 0.4 mm for A, B and C; 1.0 mm for E and F." figureDoi="http://doi.org/10.5281/zenodo.4813001" httpUri="https://zenodo.org/record/4813001/files/figure.png" pageId="3" pageNumber="4">Figs. 3D, E</figureCitation>
|
|
); pterostigma much longer than broad (40:6), metacarp (PostabR1) shorter than pterostigma (25:40); marginal cell 2R1 open; stigmal veins (2r-rs & Rs) with distal part (Rs) much longer than proximal part (2r-rs) (39:21). The spurious vein of secondary nature that runs anterior of the claval furrow between cu-a and end of the 3-Cu. Forewing with usual three basal cells (costal (C), radial (R) and first cubital (1Cu)) clearly enclosed by pigmented veins. Hind wing not distinct.
|
|
</paragraph>
|
|
<paragraph blockId="3.[100,771,1053,1602]" pageId="3" pageNumber="4">Metasomal petiole very short, much shorter than rest of metasoma (7:145). Metasoma slightly longer than mesosoma (145:140). Tibial spur formula 1/1/2.</paragraph>
|
|
<paragraph blockId="3.[100,770,1638,1823]" box="[103,267,1638,1658]" pageId="3" pageNumber="4">
|
|
<emphasis box="[103,157,1638,1658]" italics="true" pageId="3" pageNumber="4">Male.</emphasis>
|
|
Unknown.
|
|
</paragraph>
|
|
<paragraph blockId="3.[100,770,1638,1823]" box="[103,263,1665,1685]" pageId="3" pageNumber="4">
|
|
<emphasis box="[103,153,1665,1685]" italics="true" pageId="3" pageNumber="4">Host.</emphasis>
|
|
Unknown.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="3" pageNumber="4" type="discussion">
|
|
<paragraph blockId="3.[100,770,1638,1823]" pageId="3" pageNumber="4">
|
|
<emphasis box="[100,183,1691,1711]" italics="true" pageId="3" pageNumber="4">Remarks</emphasis>
|
|
. Because protrochanter is about 13 times as long as broad and the disc of pronotum is not excised medially, the new species is similar to
|
|
<emphasis box="[208,414,1746,1766]" italics="true" pageId="3" pageNumber="4">
|
|
<taxonomicName authorityName="Engel" authorityYear="2005" box="[208,363,1746,1766]" class="Insecta" family="Dryinidae" genus="Hybristodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Hybristodryinus</taxonomicName>
|
|
mon
|
|
</emphasis>
|
|
<bibRefCitation author="Perkovsky, E. E. & Olmi, M. & Muller, P. & Martynova, K. V." box="[423,659,1747,1767]" journalOrPublisher="Cretaceous Research" pageId="3" pageNumber="4" pagination="169 ‾ 189" part="99" refId="ref6295" refString="Perkovsky, E. E., Olmi, M., Muller, P., Martynova, K. V., 2019. A review of the genus Hybristodryinus Engel, 2005 (Hymenoptera, Dryinidae) from mid-Cretaceous Burmese amber, with a discussion on its phylogenetic significance. Cretaceous Research 99, 169 ‾ 189. https: // doi. org / 10.1016 / j. cretres. 2019.01.023." title="A review of the genus Hybristodryinus Engel, 2005 (Hymenoptera, Dryinidae) from mid-Cretaceous Burmese amber, with a discussion on its phylogenetic significance" type="journal article" year="2019">Perkovsky et al. (2019)</bibRefCitation>
|
|
. The main difference between the new species and
|
|
<emphasis box="[506,575,1774,1794]" italics="true" pageId="3" pageNumber="4">H. mon</emphasis>
|
|
is that POL is much longer than OPL in
|
|
<taxonomicName authorityName="Wang, Olmi and Gao" authorityYear="2021" box="[296,398,1802,1822]" class="Insecta" family="Dryinidae" genus="Hybristodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="viriosus">
|
|
<emphasis box="[296,398,1802,1822]" italics="true" pageId="3" pageNumber="4">H. viriosus</emphasis>
|
|
</taxonomicName>
|
|
and about as long as OPL in
|
|
<emphasis box="[690,761,1802,1822]" italics="true" pageId="3" pageNumber="4">H. mon</emphasis>
|
|
.
|
|
</paragraph>
|
|
<paragraph blockId="3.[100,770,1844,1892]" lastBlockId="3.[818,1487,1053,1100]" pageId="3" pageNumber="4">
|
|
Based on the description of
|
|
<taxonomicName authority="Wang & Olmi & Shih & Ren & Gao, 2021" authorityName="Wang & Olmi & Shih & Ren & Gao" authorityYear="2021" box="[410,646,1844,1864]" class="Insecta" family="Dryinidae" genus="Hybristodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="viriosus" status="sp. nov.">
|
|
<emphasis box="[410,646,1844,1864]" italics="true" pageId="3" pageNumber="4">Hybristodryinus viriosus</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel box="[651,728,1846,1865]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
|
, the keys to the females of
|
|
<taxonomicName authorityName="Engel" authorityYear="2005" box="[364,519,1872,1892]" class="Insecta" family="Dryinidae" genus="Hybristodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
|
<emphasis box="[364,519,1872,1892]" italics="true" pageId="3" pageNumber="4">Hybristodryinus</emphasis>
|
|
</taxonomicName>
|
|
from Myanmar amber published by
|
|
<bibRefCitation author="Perkovsky, E. E. & Olmi, M. & Muller, P. & Martynova, K. V." box="[954,1183,1053,1072]" journalOrPublisher="Cretaceous Research" pageId="3" pageNumber="4" pagination="169 ‾ 189" part="99" refId="ref6295" refString="Perkovsky, E. E., Olmi, M., Muller, P., Martynova, K. V., 2019. A review of the genus Hybristodryinus Engel, 2005 (Hymenoptera, Dryinidae) from mid-Cretaceous Burmese amber, with a discussion on its phylogenetic significance. Cretaceous Research 99, 169 ‾ 189. https: // doi. org / 10.1016 / j. cretres. 2019.01.023." title="A review of the genus Hybristodryinus Engel, 2005 (Hymenoptera, Dryinidae) from mid-Cretaceous Burmese amber, with a discussion on its phylogenetic significance" type="journal article" year="2019">Perkovsky et al. (2019)</bibRefCitation>
|
|
,
|
|
<bibRefCitation author="Tribull, C. & Barden, P. & Olmi, M." box="[1194,1388,1053,1072]" journalOrPublisher="Cretaceous Research" pageId="3" pageNumber="4" part="114" refId="ref6970" refString="Tribull, C., Barden, P., Olmi, M., 2020. Hybristodryinus moutesoe (Hymenoptera, Dryinidae), a new species from mid-Cretaceous Kachin (Burmese) amber. Cretaceous Research 114, 104528. https: // doi. org / 10.1016 / j. cretres. 2020.104528." title="Hybristodryinus moutesoe (Hymenoptera, Dryinidae), a new species from mid-Cretaceous Kachin (Burmese) amber" type="journal article" year="2020">Tribull et al. (2020)</bibRefCitation>
|
|
and
|
|
<bibRefCitation author="Olmi, M. & Chen, H. - Y. & Shih, C. K. & Muller, P. & Capradossi, L. & Ren, D. & Perkovsky, E. E. & Guglielmino, A." journalOrPublisher="Journal of Hymenoptera Research" pageId="3" pageNumber="4" pagination="43 ‾ 55" part="81" refId="ref5966" refString="Olmi, M., Chen, H. - Y., Shih, C. K., Muller, P., Capradossi, L., Ren, D., Perkovsky, E. E., Guglielmino, A., 2021. New species of Hybristodryinus Engel (Hymenoptera, Dryinidae) from mid-Cretaceous amber of northern Myanmar, with notes on their possible hosts. Journal of Hymenoptera Research 81, 43 ‾ 55. https: // doi. org / 10.3897 / jhr. 81.57792." title="New species of Hybristodryinus Engel (Hymenoptera, Dryinidae) from mid-Cretaceous amber of northern Myanmar, with notes on their possible hosts" type="journal article" year="2021">Olmi et al., 2021</bibRefCitation>
|
|
, must be modified by replacing couplet 7 as follows:
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="3" pageNumber="4" type="key">
|
|
<paragraph blockId="3.[822,1488,1136,1519]" pageId="3" pageNumber="4">
|
|
7. Protrochanter less than seven times as long as broad (greatest breadth) ¨¨¨¨¨¨¨¨¨¨¨¨¨¨¨¨¨¨¨¨¨¨¨¨
|
|
<taxonomicName authority="Perkovskyet al. (2019)" authorityName="Perkovskyet al." authorityYear="2019" box="[849,1208,1191,1212]" class="Insecta" family="Dryinidae" genus="Hybristodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="cretacicus">
|
|
<emphasis box="[849,972,1191,1211]" italics="true" pageId="3" pageNumber="4">H. cretacicus</emphasis>
|
|
<bibRefCitation author="Perkovsky, E. E. & Olmi, M. & Muller, P. & Martynova, K. V." box="[979,1208,1192,1212]" journalOrPublisher="Cretaceous Research" pageId="3" pageNumber="4" pagination="169 ‾ 189" part="99" refId="ref6295" refString="Perkovsky, E. E., Olmi, M., Muller, P., Martynova, K. V., 2019. A review of the genus Hybristodryinus Engel, 2005 (Hymenoptera, Dryinidae) from mid-Cretaceous Burmese amber, with a discussion on its phylogenetic significance. Cretaceous Research 99, 169 ‾ 189. https: // doi. org / 10.1016 / j. cretres. 2019.01.023." title="A review of the genus Hybristodryinus Engel, 2005 (Hymenoptera, Dryinidae) from mid-Cretaceous Burmese amber, with a discussion on its phylogenetic significance" type="journal article" year="2019">Perkovskyet al. (2019)</bibRefCitation>
|
|
</taxonomicName>
|
|
</paragraph>
|
|
<paragraph blockId="3.[822,1488,1136,1519]" pageId="3" pageNumber="4">- Protrochanter about eight-thirteen times as long as broad (greatest breadth) ¨¨¨¨¨¨¨¨¨¨¨¨¨¨¨¨¨¨¨ 8</paragraph>
|
|
<paragraph blockId="3.[822,1488,1136,1519]" pageId="3" pageNumber="4">
|
|
8. Disc of pronotum deeply longitudinally excised medially (
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<figureCitation box="[1421,1487,1276,1295]" captionStart="Fig" captionStartId="2.[100,125,1940,1954]" captionTargetBox="[110,1479,1011,1907]" captionTargetId="figure-62@2.[106,1482,1007,1910]" captionTargetPageId="2" captionText="Fig. 2. Photomicrographs of Hybristodryinus viriosus Wang, Olmi and Gao, sp. nov., holotype ♀ CNU-HYM-MA2015109. A. Lateral view. B. Lateral view of head and mesosoma. C. Ventral view of head.D. Dorsal view of head and mesosoma. E. Anterior view of head. Abbreviations: mp, maxillary palpus; oc, occipital carina. Scale bars:1.0 mm for A; 0.4 mm for B, C, D and E." figureDoi="http://doi.org/10.5281/zenodo.4812999" httpUri="https://zenodo.org/record/4812999/files/figure.png" pageId="3" pageNumber="4">Fig. 2A</figureCitation>
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in
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<bibRefCitation author="Tribull, C. & Barden, P. & Olmi, M." box="[892,1124,1304,1323]" journalOrPublisher="Cretaceous Research" pageId="3" pageNumber="4" part="114" refId="ref6970" refString="Tribull, C., Barden, P., Olmi, M., 2020. Hybristodryinus moutesoe (Hymenoptera, Dryinidae), a new species from mid-Cretaceous Kachin (Burmese) amber. Cretaceous Research 114, 104528. https: // doi. org / 10.1016 / j. cretres. 2020.104528." title="Hybristodryinus moutesoe (Hymenoptera, Dryinidae), a new species from mid-Cretaceous Kachin (Burmese) amber" type="journal article" year="2020">Tribull et al., 2020</bibRefCitation>
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) ¨¨¨¨¨¨¨¨¨¨¨¨¨¨¨ ¨¨¨¨¨¨¨¨¨¨¨¨¨
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<taxonomicName authority="Tribull et al. (2020)" authorityName="Tribull et al." authorityYear="2020" box="[1143,1464,1331,1351]" class="Insecta" family="Dryinidae" genus="Hybristodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="moutesoe">
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<emphasis box="[1143,1263,1331,1351]" italics="true" pageId="3" pageNumber="4">H. moutesoe</emphasis>
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<bibRefCitation author="Tribull, C. & Barden, P. & Olmi, M." box="[1269,1464,1332,1351]" journalOrPublisher="Cretaceous Research" pageId="3" pageNumber="4" part="114" refId="ref6970" refString="Tribull, C., Barden, P., Olmi, M., 2020. Hybristodryinus moutesoe (Hymenoptera, Dryinidae), a new species from mid-Cretaceous Kachin (Burmese) amber. Cretaceous Research 114, 104528. https: // doi. org / 10.1016 / j. cretres. 2020.104528." title="Hybristodryinus moutesoe (Hymenoptera, Dryinidae), a new species from mid-Cretaceous Kachin (Burmese) amber" type="journal article" year="2020">Tribull et al. (2020)</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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<paragraph blockId="3.[822,1488,1136,1519]" pageId="3" pageNumber="4">
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- Disc of pronotum not excised longitudinally medially (
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<figureCitation box="[1409,1483,1359,1379]" captionStart="Fig" captionStartId="2.[100,125,907,921]" captionTargetBox="[110,1469,172,874]" captionTargetId="figure-0@2.[106,1482,166,878]" captionTargetPageId="2" captionText="Fig. 1. Photomicrographs and line drawing of Hybristodryinus viriosus Wang, Olmi and Gao, sp. nov., holotype ♀ CNU-HYM-MA2015109 preserved in Cenomanian-aged Myanmar (Burmese) amber. A. Ventral view. B. Dorsal view. C. Line drawing in ventral view. Antennae and legs are artificially extended from the body for easy recognition and comparison. Scale bars: 1.0 mm for A, B and C." figureDoi="http://doi.org/10.5281/zenodo.4812997" httpUri="https://zenodo.org/record/4812997/files/figure.png" pageId="3" pageNumber="4">Figs 1B</figureCitation>
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,
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<figureCitation box="[849,879,1388,1407]" captionStart="Fig" captionStartId="2.[100,125,1940,1954]" captionTargetBox="[110,1479,1011,1907]" captionTargetId="figure-62@2.[106,1482,1007,1910]" captionTargetPageId="2" captionText="Fig. 2. Photomicrographs of Hybristodryinus viriosus Wang, Olmi and Gao, sp. nov., holotype ♀ CNU-HYM-MA2015109. A. Lateral view. B. Lateral view of head and mesosoma. C. Ventral view of head.D. Dorsal view of head and mesosoma. E. Anterior view of head. Abbreviations: mp, maxillary palpus; oc, occipital carina. Scale bars:1.0 mm for A; 0.4 mm for B, C, D and E." figureDoi="http://doi.org/10.5281/zenodo.4812999" httpUri="https://zenodo.org/record/4812999/files/figure.png" pageId="3" pageNumber="4">2D</figureCitation>
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) ¨¨¨¨¨¨¨¨¨¨¨¨¨¨¨¨¨¨¨¨¨¨¨¨¨¨ 9.
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</paragraph>
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<paragraph blockId="3.[822,1488,1136,1519]" pageId="3" pageNumber="4">
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9. Head with POL about as long as OPL ¨¨¨¨¨¨¨¨¨¨¨ ¨¨¨¨¨¨¨¨¨¨¨¨¨¨¨
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<emphasis box="[1185,1254,1442,1462]" italics="true" pageId="3" pageNumber="4">H. mon</emphasis>
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<bibRefCitation author="Perkovsky, E. E. & Olmi, M. & Muller, P. & Martynova, K. V." box="[1259,1482,1443,1463]" journalOrPublisher="Cretaceous Research" pageId="3" pageNumber="4" pagination="169 ‾ 189" part="99" refId="ref6295" refString="Perkovsky, E. E., Olmi, M., Muller, P., Martynova, K. V., 2019. A review of the genus Hybristodryinus Engel, 2005 (Hymenoptera, Dryinidae) from mid-Cretaceous Burmese amber, with a discussion on its phylogenetic significance. Cretaceous Research 99, 169 ‾ 189. https: // doi. org / 10.1016 / j. cretres. 2019.01.023." title="A review of the genus Hybristodryinus Engel, 2005 (Hymenoptera, Dryinidae) from mid-Cretaceous Burmese amber, with a discussion on its phylogenetic significance" type="journal article" year="2019">Perkovskyet al. (2019)</bibRefCitation>
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.
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</paragraph>
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<paragraph blockId="3.[822,1488,1136,1519]" pageId="3" pageNumber="4">
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- Head with POL much longer than OPL ¨¨¨¨¨¨¨¨¨¨¨ ¨¨¨¨¨¨¨¨¨¨¨
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<taxonomicName authority="Wang, Olmi and Gao" authorityName="Wang, Olmi and Gao" authorityYear="2021" box="[1092,1404,1498,1519]" class="Insecta" family="Dryinidae" genus="Hybristodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="viriosus" status="sp. nov.">
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<emphasis box="[1092,1193,1498,1518]" italics="true" pageId="3" pageNumber="4">H. viriosus</emphasis>
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Wang, Olmi and Gao
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</taxonomicName>
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,
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<taxonomicNameLabel box="[1413,1488,1500,1519]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |