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<document id="8D7D3A6C939641C24DA7860923700E23" ID-CLB-Dataset="5365" ID-DOI="10.11646/zootaxa.3812.1.1" ID-GBIF-Dataset="fd5ba21e-09ce-4ac6-b84f-56632ed93917" ID-ISSN="1175-5326" ID-Zenodo-Dep="4919079" ID-ZooBank="7847D731-9F66-4856-A79F-9435FED25B1D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623268636583" checkinUser="felipe" docAuthor="Murányi, Dávid, Gamboa, Maribet &amp; Orci, Kirill Márk" docDate="2014" docId="03F99336FF23FFC21BE45E4DD16FFA3A" docLanguage="en" docName="zt03812p082.pdf" docOrigin="Zootaxa 3812 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Zwicknia kovacsi Muranyi &amp; Gamboa 2014, sp. n." docType="treatment" docVersion="8" lastPageNumber="46" masterDocId="FFC0EB4EFF08FFEF1B735B43D539FFDD" masterDocTitle="Zwicknia gen. n., a new genus for the Capnia bifrons species group, with descriptions of three new species based on morphology, drumming signals and molecular genetics, and a synopsis of the West Palaearctic and Nearctic genera of Capniidae (Plecoptera)" masterLastPageNumber="82" masterPageNumber="1" pageNumber="44" updateTime="1699007786927" updateUser="ExternalLinkService">
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<mods:title id="D061CFA2E87F24F3A2AE00B7954F0987">Zwicknia gen. n., a new genus for the Capnia bifrons species group, with descriptions of three new species based on morphology, drumming signals and molecular genetics, and a synopsis of the West Palaearctic and Nearctic genera of Capniidae (Plecoptera)</mods:title>
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<mods:namePart id="584A78DD007F6EFB4DC2B2E704FE8E12">Murányi, Dávid</mods:namePart>
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<mods:namePart id="6001310FA4F1D4A0742C1CA00CD98037">Gamboa, Maribet</mods:namePart>
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<mods:namePart id="5A0A22322D50FE3C33FCAD1203DA3E0C">Orci, Kirill Márk</mods:namePart>
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<treatment id="03F99336FF23FFC21BE45E4DD16FFA3A" ID-DOI="http://doi.org/10.5281/zenodo.4919259" ID-GBIF-Taxon="183709173" ID-Zenodo-Dep="4919259" LSID="urn:lsid:plazi:treatment:03F99336FF23FFC21BE45E4DD16FFA3A" httpUri="http://treatment.plazi.org/id/03F99336FF23FFC21BE45E4DD16FFA3A" lastPageId="45" lastPageNumber="46" pageId="43" pageNumber="44">
<subSubSection id="C34A71ABFF23FFC41BE45E4DD7F8FAF5" box="[151,705,1294,1320]" pageId="43" pageNumber="44" type="nomenclature">
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<taxonomicName id="4C5059A3FF23FFC41BE45E4DD74BFAF5" ID-CoL="7GGL4" authority="Muranyi &amp; Gamboa" authorityName="Muranyi &amp; Gamboa" authorityYear="2014" box="[151,626,1294,1320]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="kovacsi" status="sp. nov.">
<emphasis id="B924FE32FF23FFC41BE45E4DD451FAF5" bold="true" box="[151,360,1294,1320]" italics="true" pageId="43" pageNumber="44">Zwicknia kovacsi</emphasis>
Murányi &amp; Gamboa
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,
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<paragraph id="8BEF2220FF23FFC41BE45E72D1D2FA97" blockId="43.[151,1259,1294,1354]" box="[151,1259,1329,1354]" pageId="43" pageNumber="44">
(
<figureCitation id="136B3EA5FF23FFC41BEC5E72D41EFA97" box="[159,295,1329,1354]" captionStart-0="FIGURE 49" captionStart-1="FIGURE 50" captionStart-2="FIGURE 51" captionStartId-0="24.[151,250,1960,1982]" captionStartId-1="25.[151,250,1969,1991]" captionStartId-2="26.[151,250,1010,1032]" captionTargetBox-0="[171,1394,199,1947]" captionTargetBox-1="[199,1377,195,1963]" captionTargetBox-2="[152,1425,208,974]" captionTargetId-0="figure-48@24.[151,1436,170,1981]" captionTargetId-1="figure-46@25.[199,1385,171,1983]" captionTargetId-2="figure-474@26.[151,1436,193,989]" captionTargetPageId-0="24" captionTargetPageId-1="25" captionTargetPageId-2="26" captionText-0="FIGURE 49. Dorsal habitus of male holotype Zwicknia kovacsi Murányi &amp; Gamboa, sp. n.; Romania, Maramureş County, Rodna Mts., Borşa-Staţiunea Borşa—scale 1 mm." captionText-1="FIGURE 50. Ventral habitus of male holotype Zwicknia kovacsi Murányi &amp; Gamboa, sp. n.; Romania, Maramureş County, Rodna Mts., Borşa-Staţiunea Borşa—scale 1 mm." captionText-2="FIGURE 51. Lateral habitus of male holotype Zwicknia kovacsi Murányi &amp; Gamboa, sp. n.; Romania, Maramureş County, Rodna Mts., Borşa-Staţiunea Borşa—scale 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4919095" figureDoi-1="http://doi.org/10.5281/zenodo.4919097" figureDoi-2="http://doi.org/10.5281/zenodo.4919099" httpUri-0="https://zenodo.org/record/4919095/files/figure.png" httpUri-1="https://zenodo.org/record/4919097/files/figure.png" httpUri-2="https://zenodo.org/record/4919099/files/figure.png" pageId="43" pageNumber="44">Figs. 49–51</figureCitation>
,
<figureCitation id="136B3EA5FF23FFC41A415E72D442FA97" box="[306,379,1329,1354]" captionStart="FIGURES 53–55" captionStartId="29.[151,264,1334,1357]" captionTargetBox="[171,1408,529,1298]" captionTargetId="figure-537@29.[151,1436,517,1313]" captionTargetPageId="29" captionText="FIGURES 53–55. Thoracal sclerites of male Zwicknia kovacsi Murányi &amp; Gamboa, sp. n.—53: ventral view; 54: lateral view; 55: dorsal view (P: prothoracic; Me: mesothoracic; M: metathoracic—Ab: anterior basalare; Aes: anepisternum; Al: alifer; Bs: basisternum; Cx: coxa; Epm: epimeron; Eps: episternum; Etn: eutrochantin; Fs: furcasternum; Fsa: furcasternal arm; Fsp: furcasternal pit; Kes: katepisternum; Lcs: lateral cervical sclerite; Pb: posterior basalare; Pfs: postfurcasternum; Pn: pronotum; Po: postcoxal bridge; Pr: precoxal bridge; Pra: prealar bridge; Prs: presternum; Psc: prescutum; Pscl: postscutellum; Sct: scutum; Scl: scutellum; Ss: spinasternum; St1: Sternite 1; Tg1: Tergite 1; Tgl: tegula; Tn: trochantin; Tr: trochanter—wings and subalar sclerites omitted)—not to scale." figureDoi="http://doi.org/10.5281/zenodo.4919104" httpUri="https://zenodo.org/record/4919104/files/figure.png" pageId="43" pageNumber="44">
<tableCitation id="C6D2179BFF23FFC41A415E72D442FA97" box="[306,379,1329,1354]" captionStart="TABLE 1" captionStartId="7.[147,247,602,628]" captionTargetBox="[167,1849,720,1573]" captionText="TABLE 1. Ventral sclerites of valid extant Capniidae genera. — Capniella not included because of lacking information; in case of two character states the first one refers to the type species; for abbreviations of sclerites see Figs. 53-55." httpUri="http://table.plazi.org/id/DF2F72A8FF0FFFE81BE05919D188FD46" pageId="43" pageNumber="44" tableUuid="DF2F72A8FF0FFFE81BE05919D188FD46">53–55</tableCitation>
</figureCitation>
,
<figureCitation id="136B3EA5FF23FFC41AF55E72D4F6FA97" box="[390,463,1329,1354]" captionStart="FIGURES 86–89" captionStartId="44.[151,264,1964,1987]" captionTargetBox="[237,1353,206,1945]" captionTargetId="figure-60@44.[219,1370,193,1959]" captionTargetPageId="44" captionText="FIGURES 86–89. Male terminalia of Zwicknia kovacsi Murányi &amp; Gamboa, sp. n.; paratype, Romania, Maramureş County, Rodna Mts., Borşa-Staţiunea Borşa—86: dorsal view; 87: ventral view; 88: lateral view; 89: caudal view—scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919122" httpUri="https://zenodo.org/record/4919122/files/figure.png" pageId="43" pageNumber="44">86–89</figureCitation>
,
<figureCitation id="136B3EA5FF23FFC41AA95E72D73FFA97" box="[474,518,1329,1354]" captionStart="FIGURES 110–113" captionStartId="55.[151,264,1953,1976]" captionTargetBox="[151,1436,1069,1940]" captionTargetId="figure-316@55.[151,1436,1069,1940]" captionTargetPageId="55" captionText="FIGURES 110–113. Lateral view of male epiproct of Zwicknia Murányi, gen. n.—110: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 111: Z. sevanica (Zhiltzova, 1964); 112: Z. tuberculata (Zhiltzova, 1964); 113: Z. turkestanica turkestanica (Kimmins, 1950)—scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919138" httpUri="https://zenodo.org/record/4919138/files/figure.png" pageId="43" pageNumber="44">110</figureCitation>
,
<figureCitation id="136B3EA5FF23FFC419625E72D704FA97" box="[529,573,1329,1354]" captionStart="FIGURES 114–119" captionStartId="56.[151,265,1540,1563]" captionTargetBox="[151,1436,193,1520]" captionTargetId="figure-203@56.[151,1436,193,1520]" captionTargetPageId="56" captionText="FIGURES 114–119. Dorsal view of male epiproct of Zwicknia Murányi, gen. n.—114: Z. bifrons (Newman, 1838); 115, 117: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 116: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 118: Z. sevanica (Zhiltzova, 1964); 119: Z. acuta Murányi &amp; Orci, sp. n.—scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919140" httpUri="https://zenodo.org/record/4919140/files/figure.png" pageId="43" pageNumber="44">116</figureCitation>
,
<figureCitation id="136B3EA5FF23FFC4193B5E72D74DFA97" box="[584,628,1329,1354]" captionStart="FIGURES 120–123" captionStartId="57.[151,264,1761,1784]" captionTargetBox="[165,1401,468,1695]" captionTargetId="figure-157@57.[151,1436,457,1720]" captionTargetPageId="57" captionText="FIGURES 120–123. Male terminalia of Zwicknia Murányi, gen. n., dorso-caudal view—120: Z. bifrons (Newman, 1838); 121: Z. acuta Murányi &amp; Orci, sp. n.; 122: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 123: Z. kovacsi Murányi &amp; Gamboa, sp. n.—scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919142" httpUri="https://zenodo.org/record/4919142/files/figure.png" pageId="43" pageNumber="44">123</figureCitation>
,
<figureCitation id="136B3EA5FF23FFC419F35E72D7DCFA97" box="[640,741,1329,1354]" captionStart="FIGURES 127–147" captionStartId="59.[151,264,1277,1300]" captionTargetBox="[154,1418,219,1236]" captionTargetId="figure-173@59.[151,1436,193,1256]" captionTargetPageId="59" captionText="FIGURES 127–147. Male epiproct of Zwicknia Murányi, gen. n.—127–130 Z. acuta Murányi &amp; Orci, sp. n.; 131–132: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 133–138: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 139–143: Z. bifrons (Newman, 1838); 144–145: Z. sevanica (Zhiltzova, 1964); 146: Z. tuberculata (Zhiltzova, 1964); 147: Z. turkestanica turkestanica (Kimmins, 1950); 127–131, 133, 135, 137, 139–147: lateral view; 132, 134, 136, 138: dorsal view (127: Slovakia, Krupinská Planina, Pôtor; 128: Hungary, Zemplén Mts., Kishuta; 129: Serbia, Fruska Gora, Vrdnik; 130: Serbia, Zlatibor Mts., Crni Rzav; 131–132: Romania, Rodna Mts., Borşa–Staţiunea Borşa; 133–134, 137–138: Croatia, Ivanščica Mts., Stari Golubovec; 135–136: Hungary, Mecsek Mts., Komló-Zobákpuszta; 139: Hungary, Börzsöny Mts., Szokolya-Királyrét; 140: Hungary, Mátra Mts., Gyöngyössolymos; 141: Hungary, Bükk Mts., Nagyvisnyó; 142: Hungary, Bakony Mts., Bakonybél; 143: Serbia, Maljen Mts., Brajkovići; 144–145: Turkey, Akyarma Pass; 146: Kazakhstan, Karatau Mts., Bajaldyrtal; 147: Kazakhstan, Tarbagatai Mts., Ayagos)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919146" httpUri="https://zenodo.org/record/4919146/files/figure.png" pageId="43" pageNumber="44">131–132</figureCitation>
,
<figureCitation id="136B3EA5FF23FFC419825E72D624FA97" box="[753,797,1329,1354]" captionStart="FIGURES 148–164" captionStartId="60.[151,264,1223,1246]" captionTargetBox="[166,1423,196,1203]" captionTargetId="figure-337@60.[151,1436,181,1215]" captionTargetPageId="60" captionText="FIGURES 148–164. Humps of male Tergite 9 of Zwicknia Murányi, gen. n., caudal view—148–151: Z. acuta Murányi &amp; Orci, sp. n.; 152–156: Z. bifrons (Newman, 1838); 157–159: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 160: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 161–162: Z. sevanica (Zhiltzova, 1964); 163: Z. tuberculata (Zhiltzova, 1964); 164: Z. turkestanica turkestanica (Kimmins, 1950) (148: Slovakia, Krupinská Planina, Pôtor; 149: Hungary, Zemplén Mts., Kishuta; 150: Serbia, Fruska Gora, Vrdnik; 151: Serbia, Zlatibor Mts., Crni Rzav; 152: Hungary, Börzsöny Mts., Szokolya-Királyrét; 153: Hungary, Mátra Mts., Gyöngyössolymos; 154: Hungary, Bükk Mts., Nagyvisnyó; 155: Hungary, Bakony Mts., Bakonybél; 156: Serbia, Maljen Mts., Brajkovići; 157, 159: Croatia, Ivanščica Mts., Stari Golubovec; 158: Hungary, Mecsek Mts., Komló-Zobákpuszta; 160: Romania, Rodna Mts., Borşa-Staţiunea Borşa; 161–162: Turkey, Akyarma Pass; 163: Kazakhstan, Karatau Mts., Bajaldyrtal; 164: Kazakhstan, Tarbagatai Mts., Ayagos)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919148" httpUri="https://zenodo.org/record/4919148/files/figure.png" pageId="43" pageNumber="44">160</figureCitation>
,
<figureCitation id="136B3EA5FF23FFC4185A5E72D66CFA97" box="[809,853,1329,1354]" captionStart="FIGURES 165–168" captionStartId="61.[151,264,1330,1353]" captionTargetBox="[162,1428,857,1282]" captionTargetId="figure-628@61.[151,1436,850,1309]" captionTargetPageId="61" captionText="FIGURES 165–168. Slow oscillograms of male calls of four Zwicknia Murányi, gen. n. species, showing call sequences versus sporadic call production—165: two call sequence of Z. bifrons (Newman, 1838), &quot;//&quot; indicates where 2 minutes 23 seconds of silence was cut out from the recording; 166: three call sequences of Z. acuta Murányi &amp; Orci, sp. n.; 167: sporadic calls of Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 168: sporadic call production in Z. kovacsi Murányi &amp; Gamboa, sp. n.—time scale is the same for the four oscillograms; for detailed data on the animals and recording circumstances, see Table 9." figureDoi="http://doi.org/10.5281/zenodo.4919154" httpUri="https://zenodo.org/record/4919154/files/figure.png" pageId="43" pageNumber="44">168</figureCitation>
,
<figureCitation id="136B3EA5FF23FFC418125E72D6FCFA97" box="[865,965,1329,1354]" captionStart="FIGURES 169–184" captionStartId="62.[151,264,1845,1868]" captionTargetBox="[201,1412,198,1820]" captionTargetId="figure-140@62.[168,1412,183,1822]" captionTargetPageId="62" captionText="FIGURES 169–184. Male drumming calls of Zwicknia Murányi, gen. n.—169–173: Z. bifrons (Newman, 1838); 174–178: Z. acuta Murányi &amp; Orci, sp. n.; 179–182: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 183–184: Z. kovacsi Murányi &amp; Gamboa, sp. n. (see detailed data on the animals and recording circumstances in Table 9)—Oscillograms in Figs. 169–179, 181, 183 have the same time axis resolution (time scale bar is presented in Fig. 169 for all of them), an other time scale is used in the faster oscillograms of Figs. 180, 182, 184 (time scale bar for them can be found in Fig. 184), those are zoomed sections of Figs. 179, 181, 183, respectively." figureDoi="http://doi.org/10.5281/zenodo.4919156" httpUri="https://zenodo.org/record/4919156/files/figure.png" pageId="43" pageNumber="44">183–184</figureCitation>
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<figureCitation id="136B3EA5FF23FFC418A25E72D10FFA97" box="[977,1078,1329,1354]" captionStart-0="FIGURE 190" captionStart-1="FIGURE 191" captionStart-2="FIGURE 192" captionStartId-0="64.[151,250,1125,1147]" captionStartId-1="65.[151,250,1357,1379]" captionStartId-2="66.[151,249,1180,1202]" captionTargetBox-0="[163,1430,185,1044]" captionTargetBox-1="[164,1416,208,1328]" captionTargetBox-2="[156,1424,193,1155]" captionTargetId-0="figure-392@64.[151,1436,177,1058]" captionTargetId-1="figure-231@65.[151,1436,193,1336]" captionTargetId-2="figure-240@66.[151,1433,185,1171]" captionTargetPageId-0="64" captionTargetPageId-1="65" captionTargetPageId-2="66" captionText-0="FIGURE 190. Inter-beat interval variation during male calls in the four examined Zwicknia Murányi, gen. n. species.—Box plot colors: white—Z. bifrons (Newman, 1838), red—Z. acuta Murányi &amp; Orci, sp. n., blue—Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n. In Z. kovacsi Murányi &amp; Gamboa, sp. n. we have drumming samples only from two specimens therefore in the case of that species two simple mark series are used to show inter-beat intervals in their calls (circles for specimen 1, and triangles for specimen 2). i1, i2, i3 and so on are labels containing the serial number of inter-beat intervals counting them from the beginning of each call. Box plots show the duration values measured in 7 specimens in Z. bifrons and Z. acuta, and 5 specimens in Z. rupprechti. Vertical axis has a logarithmic scale. Only calls recorded at an ambient air temperature of 17–19 oC were used. (see the measurement data in Appendix Tables 1–4)." captionText-1="FIGURE 191. A scatter plot showing the clear separation of male calls of Zwicknia Murányi, gen. n. along two drumming signal characters—Z. bifrons (Newman, 1838): triangles; Z. acuta Murányi &amp; Orci, sp. n.: circles; Z. kovacsi Murányi &amp; Gamboa, sp. n.: +; Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.: X—Each mark represents the mean values of the characters for a given specimen. Note that both axes have logarithmic scale. Air temperature varied between 15–21 oC in every species (for details see Tables 6–8)." captionText-2="FIGURE 192. Phylogenetic tree of cox1 including Zwicknia Murányi, gen. n. with our accession codes (300941-301003), and GenBank closest species (with alphanumeric accession codes) as outgroup. MP and ML had same topology. Numbers on braches are ML/MP bootstrap values&gt;50%. Species names right to the tree refer to morphological species. P-Distance table is showed." figureDoi-0="http://doi.org/10.5281/zenodo.4919160" figureDoi-1="http://doi.org/10.5281/zenodo.4919162" figureDoi-2="http://doi.org/10.5281/zenodo.4919164" httpUri-0="https://zenodo.org/record/4919160/files/figure.png" httpUri-1="https://zenodo.org/record/4919162/files/figure.png" httpUri-2="https://zenodo.org/record/4919164/files/figure.png" pageId="43" pageNumber="44">190–192</figureCitation>
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<figureCitation id="136B3EA5FF23FFC41F335E72D154FA97" box="[1088,1133,1329,1354]" captionStart="FIGURES 193–195" captionStartId="67.[151,264,1829,1852]" captionTargetBox="[199,1367,231,1794]" captionTargetId="figure-90@67.[199,1367,231,1810]" captionTargetPageId="67" captionText="FIGURES 193–195. Type localities of the new species described in Zwicknia Murányi, gen. n.—193: Zwicknia acuta Murányi &amp; Orci, sp. n.: Stará rieka, Slovakia, Banskobystrický Region, Krupinská Planina, Pôtor; 194: Zwicknia kovacsi Murányi &amp; Gamboa, sp. n.: Cimpoies Stream, Romania, Maramureş County, Rodna Mts., Borşa-Staţiunea Borşa; 195: Zwicknia rupprechti Murányi, Orci &amp; Gamboa, sp. n.: Völgységi Stream, Hungary, Baranya County, Mecsek Mts., Komló-Zobákpuszta." figureDoi="http://doi.org/10.5281/zenodo.4919166" httpUri="https://zenodo.org/record/4919166/files/figure.png" pageId="43" pageNumber="44">194</figureCitation>
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<figureCitation id="136B3EA5FF23FFC41F0B5E72D1E6FA97" box="[1144,1247,1329,1354]" captionStart="FIGURES 196–197" captionStartId="68.[151,264,1908,1931]" captionTargetBox="[188,1393,187,1898]" captionTargetId="figure-93@68.[188,1393,187,1898]" captionTargetPageId="68" captionText="FIGURES 196–197. Distribution of Zwicknia Murányi, gen. n.—196: global distribution; 197: studied populations of the Carpathian Basin and the northern Balkans—in Fig. 196, white dot on Z. bifrons symbols indicate populations studied by Rupprecht (1997), dotted lines indicate limits of s.l. Z. bifrons literature reports; in Fig. 197, white dot on symbols indicate populations where drumming signals were studied by us, white star indicate type localities." figureDoi="http://doi.org/10.5281/zenodo.4919168" httpUri="https://zenodo.org/record/4919168/files/figure.png" pageId="43" pageNumber="44">196–197</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34A71ABFF23FFC41BE45E37D61BF99E" pageId="43" pageNumber="44" type="diagnosis">
<paragraph id="8BEF2220FF23FFC41BE45E37D61BF99E" blockId="43.[151,1436,1396,1927]" pageId="43" pageNumber="44">
<emphasis id="B924FE32FF23FFC41BE45E37D428FA50" bold="true" box="[151,273,1396,1421]" pageId="43" pageNumber="44">Diagnosis.</emphasis>
Male epiproct: Ep-scl medially swollen, wide and blunt in dorsal view, tip straight in lateral view; ventral membranous section terminates far before the base in lateral view, apical spines thin and long, distributed also on the apex of Ep-scl. Process of male Tg 9: wide, perpendicularly elevated, 2 × wider than the swollen part of the Ep-scl, bearing two distinct apical horns. Males produce short (150–160 ms), monophasic, percussive calls. A call is composed of 6–7 beats that follow each other with nearly constant inter-beat intervals (27–34 ms at 17–18
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C ambient air temperature). Calls are produced sporadically.
</paragraph>
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<emphasis id="B924FE32FF23FFC41BB45D0DD447F9BA" bold="true" box="[199,382,1614,1639]" pageId="43" pageNumber="44">Type material.</emphasis>
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<figureCitation id="136B3EA5FF23FFC419A55DD5D65BF972" box="[726,866,1686,1711]" captionStart-0="FIGURE 49" captionStart-1="FIGURE 50" captionStart-2="FIGURE 51" captionStartId-0="24.[151,250,1960,1982]" captionStartId-1="25.[151,250,1969,1991]" captionStartId-2="26.[151,250,1010,1032]" captionTargetBox-0="[171,1394,199,1947]" captionTargetBox-1="[199,1377,195,1963]" captionTargetBox-2="[152,1425,208,974]" captionTargetId-0="figure-48@24.[151,1436,170,1981]" captionTargetId-1="figure-46@25.[199,1385,171,1983]" captionTargetId-2="figure-474@26.[151,1436,193,989]" captionTargetPageId-0="24" captionTargetPageId-1="25" captionTargetPageId-2="26" captionText-0="FIGURE 49. Dorsal habitus of male holotype Zwicknia kovacsi Murányi &amp; Gamboa, sp. n.; Romania, Maramureş County, Rodna Mts., Borşa-Staţiunea Borşa—scale 1 mm." captionText-1="FIGURE 50. Ventral habitus of male holotype Zwicknia kovacsi Murányi &amp; Gamboa, sp. n.; Romania, Maramureş County, Rodna Mts., Borşa-Staţiunea Borşa—scale 1 mm." captionText-2="FIGURE 51. Lateral habitus of male holotype Zwicknia kovacsi Murányi &amp; Gamboa, sp. n.; Romania, Maramureş County, Rodna Mts., Borşa-Staţiunea Borşa—scale 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4919095" figureDoi-1="http://doi.org/10.5281/zenodo.4919097" figureDoi-2="http://doi.org/10.5281/zenodo.4919099" httpUri-0="https://zenodo.org/record/4919095/files/figure.png" httpUri-1="https://zenodo.org/record/4919097/files/figure.png" httpUri-2="https://zenodo.org/record/4919099/files/figure.png" pageId="43" pageNumber="44">Figs. 49–51</figureCitation>
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<figureCitation id="136B3EA5FF23FFC418075DD5D687F972" box="[884,958,1686,1711]" captionStart="FIGURES 53–55" captionStartId="29.[151,264,1334,1357]" captionTargetBox="[171,1408,529,1298]" captionTargetId="figure-537@29.[151,1436,517,1313]" captionTargetPageId="29" captionText="FIGURES 53–55. Thoracal sclerites of male Zwicknia kovacsi Murányi &amp; Gamboa, sp. n.—53: ventral view; 54: lateral view; 55: dorsal view (P: prothoracic; Me: mesothoracic; M: metathoracic—Ab: anterior basalare; Aes: anepisternum; Al: alifer; Bs: basisternum; Cx: coxa; Epm: epimeron; Eps: episternum; Etn: eutrochantin; Fs: furcasternum; Fsa: furcasternal arm; Fsp: furcasternal pit; Kes: katepisternum; Lcs: lateral cervical sclerite; Pb: posterior basalare; Pfs: postfurcasternum; Pn: pronotum; Po: postcoxal bridge; Pr: precoxal bridge; Pra: prealar bridge; Prs: presternum; Psc: prescutum; Pscl: postscutellum; Sct: scutum; Scl: scutellum; Ss: spinasternum; St1: Sternite 1; Tg1: Tergite 1; Tgl: tegula; Tn: trochantin; Tr: trochanter—wings and subalar sclerites omitted)—not to scale." figureDoi="http://doi.org/10.5281/zenodo.4919104" httpUri="https://zenodo.org/record/4919104/files/figure.png" pageId="43" pageNumber="44">
<tableCitation id="C6D2179BFF23FFC418075DD5D687F972" box="[884,958,1686,1711]" captionStart="TABLE 1" captionStartId="7.[147,247,602,628]" captionTargetBox="[167,1849,720,1573]" captionText="TABLE 1. Ventral sclerites of valid extant Capniidae genera. — Capniella not included because of lacking information; in case of two character states the first one refers to the type species; for abbreviations of sclerites see Figs. 53-55." httpUri="http://table.plazi.org/id/DF2F72A8FF0FFFE81BE05919D188FD46" pageId="43" pageNumber="44" tableUuid="DF2F72A8FF0FFFE81BE05919D188FD46">53–55</tableCitation>
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<figureCitation id="136B3EA5FF23FFC41ABC5DF9D76EF90E" box="[463,599,1722,1747]" captionStart="FIGURES 86–89" captionStartId="44.[151,264,1964,1987]" captionTargetBox="[237,1353,206,1945]" captionTargetId="figure-60@44.[219,1370,193,1959]" captionTargetPageId="44" captionText="FIGURES 86–89. Male terminalia of Zwicknia kovacsi Murányi &amp; Gamboa, sp. n.; paratype, Romania, Maramureş County, Rodna Mts., Borşa-Staţiunea Borşa—86: dorsal view; 87: ventral view; 88: lateral view; 89: caudal view—scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919122" httpUri="https://zenodo.org/record/4919122/files/figure.png" pageId="43" pageNumber="44">Figs. 86–89</figureCitation>
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<figureCitation id="136B3EA5FF23FFC419105DF9D7A9F90E" box="[611,656,1722,1747]" captionStart="FIGURES 120–123" captionStartId="57.[151,264,1761,1784]" captionTargetBox="[165,1401,468,1695]" captionTargetId="figure-157@57.[151,1436,457,1720]" captionTargetPageId="57" captionText="FIGURES 120–123. Male terminalia of Zwicknia Murányi, gen. n., dorso-caudal view—120: Z. bifrons (Newman, 1838); 121: Z. acuta Murányi &amp; Orci, sp. n.; 122: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 123: Z. kovacsi Murányi &amp; Gamboa, sp. n.—scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919142" httpUri="https://zenodo.org/record/4919142/files/figure.png" pageId="43" pageNumber="44">123</figureCitation>
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<figureCitation id="136B3EA5FF23FFC419EF5DF9D638F90E" box="[668,769,1722,1747]" captionStart="FIGURES 127–147" captionStartId="59.[151,264,1277,1300]" captionTargetBox="[154,1418,219,1236]" captionTargetId="figure-173@59.[151,1436,193,1256]" captionTargetPageId="59" captionText="FIGURES 127–147. Male epiproct of Zwicknia Murányi, gen. n.—127–130 Z. acuta Murányi &amp; Orci, sp. n.; 131–132: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 133–138: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 139–143: Z. bifrons (Newman, 1838); 144–145: Z. sevanica (Zhiltzova, 1964); 146: Z. tuberculata (Zhiltzova, 1964); 147: Z. turkestanica turkestanica (Kimmins, 1950); 127–131, 133, 135, 137, 139–147: lateral view; 132, 134, 136, 138: dorsal view (127: Slovakia, Krupinská Planina, Pôtor; 128: Hungary, Zemplén Mts., Kishuta; 129: Serbia, Fruska Gora, Vrdnik; 130: Serbia, Zlatibor Mts., Crni Rzav; 131–132: Romania, Rodna Mts., Borşa–Staţiunea Borşa; 133–134, 137–138: Croatia, Ivanščica Mts., Stari Golubovec; 135–136: Hungary, Mecsek Mts., Komló-Zobákpuszta; 139: Hungary, Börzsöny Mts., Szokolya-Királyrét; 140: Hungary, Mátra Mts., Gyöngyössolymos; 141: Hungary, Bükk Mts., Nagyvisnyó; 142: Hungary, Bakony Mts., Bakonybél; 143: Serbia, Maljen Mts., Brajkovići; 144–145: Turkey, Akyarma Pass; 146: Kazakhstan, Karatau Mts., Bajaldyrtal; 147: Kazakhstan, Tarbagatai Mts., Ayagos)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919146" httpUri="https://zenodo.org/record/4919146/files/figure.png" pageId="43" pageNumber="44">131–132</figureCitation>
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<figureCitation id="136B3EA5FF23FFC4187E5DF9D603F90E" box="[781,826,1722,1747]" captionStart="FIGURES 148–164" captionStartId="60.[151,264,1223,1246]" captionTargetBox="[166,1423,196,1203]" captionTargetId="figure-337@60.[151,1436,181,1215]" captionTargetPageId="60" captionText="FIGURES 148–164. Humps of male Tergite 9 of Zwicknia Murányi, gen. n., caudal view—148–151: Z. acuta Murányi &amp; Orci, sp. n.; 152–156: Z. bifrons (Newman, 1838); 157–159: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 160: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 161–162: Z. sevanica (Zhiltzova, 1964); 163: Z. tuberculata (Zhiltzova, 1964); 164: Z. turkestanica turkestanica (Kimmins, 1950) (148: Slovakia, Krupinská Planina, Pôtor; 149: Hungary, Zemplén Mts., Kishuta; 150: Serbia, Fruska Gora, Vrdnik; 151: Serbia, Zlatibor Mts., Crni Rzav; 152: Hungary, Börzsöny Mts., Szokolya-Királyrét; 153: Hungary, Mátra Mts., Gyöngyössolymos; 154: Hungary, Bükk Mts., Nagyvisnyó; 155: Hungary, Bakony Mts., Bakonybél; 156: Serbia, Maljen Mts., Brajkovići; 157, 159: Croatia, Ivanščica Mts., Stari Golubovec; 158: Hungary, Mecsek Mts., Komló-Zobákpuszta; 160: Romania, Rodna Mts., Borşa-Staţiunea Borşa; 161–162: Turkey, Akyarma Pass; 163: Kazakhstan, Karatau Mts., Bajaldyrtal; 164: Kazakhstan, Tarbagatai Mts., Ayagos)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919148" httpUri="https://zenodo.org/record/4919148/files/figure.png" pageId="43" pageNumber="44">160</figureCitation>
, for molecular studies as 300988, drumming recorded as 2011/No.2),
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<collectionCode id="ED41BAE5FF23FFC41A0B5D9DD4EDF92A" box="[376,468,1758,1783]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="43" pageNumber="44">HNHM</collectionCode>
:
<specimenCode id="DBF68A5BFF23FFC41AAC5D9DD772F92A" box="[479,587,1758,1783]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="43" pageNumber="44">PLP3807</specimenCode>
; terminalia prepared for SEM to
<figureCitation id="136B3EA5FF23FFC418B05D9DD117F92A" box="[963,1070,1758,1783]" captionStart="FIGURES 110–113" captionStartId="55.[151,264,1953,1976]" captionTargetBox="[151,1436,1069,1940]" captionTargetId="figure-316@55.[151,1436,1069,1940]" captionTargetPageId="55" captionText="FIGURES 110–113. Lateral view of male epiproct of Zwicknia Murányi, gen. n.—110: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 111: Z. sevanica (Zhiltzova, 1964); 112: Z. tuberculata (Zhiltzova, 1964); 113: Z. turkestanica turkestanica (Kimmins, 1950)—scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919138" httpUri="https://zenodo.org/record/4919138/files/figure.png" pageId="43" pageNumber="44">Figs. 110</figureCitation>
,
<figureCitation id="136B3EA5FF23FFC41F495D9DD15FF92A" box="[1082,1126,1758,1783]" captionStart="FIGURES 114–119" captionStartId="56.[151,265,1540,1563]" captionTargetBox="[151,1436,193,1520]" captionTargetId="figure-203@56.[151,1436,193,1520]" captionTargetPageId="56" captionText="FIGURES 114–119. Dorsal view of male epiproct of Zwicknia Murányi, gen. n.—114: Z. bifrons (Newman, 1838); 115, 117: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 116: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 118: Z. sevanica (Zhiltzova, 1964); 119: Z. acuta Murányi &amp; Orci, sp. n.—scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919140" httpUri="https://zenodo.org/record/4919140/files/figure.png" pageId="43" pageNumber="44">116</figureCitation>
, used for molecular studies as 300987, drumming recorded as 2011/No.1,
<figureCitation id="136B3EA5FF23FFC419ED5C41D678F8C6" box="[670,833,1794,1819]" captionStart="FIGURES 169–184" captionStartId="62.[151,264,1845,1868]" captionTargetBox="[201,1412,198,1820]" captionTargetId="figure-140@62.[168,1412,183,1822]" captionTargetPageId="62" captionText="FIGURES 169–184. Male drumming calls of Zwicknia Murányi, gen. n.—169–173: Z. bifrons (Newman, 1838); 174–178: Z. acuta Murányi &amp; Orci, sp. n.; 179–182: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 183–184: Z. kovacsi Murányi &amp; Gamboa, sp. n. (see detailed data on the animals and recording circumstances in Table 9)—Oscillograms in Figs. 169–179, 181, 183 have the same time axis resolution (time scale bar is presented in Fig. 169 for all of them), an other time scale is used in the faster oscillograms of Figs. 180, 182, 184 (time scale bar for them can be found in Fig. 184), those are zoomed sections of Figs. 179, 181, 183, respectively." figureDoi="http://doi.org/10.5281/zenodo.4919156" httpUri="https://zenodo.org/record/4919156/files/figure.png" pageId="43" pageNumber="44">Figs. 183–184</figureCitation>
), 1f (
<collectionCode id="ED41BAE5FF23FFC4180F5C41D68DF8C6" box="[892,948,1794,1819]" country="France" name="University of Montpellier" pageId="43" pageNumber="44">MM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34A71ABFF23FFC21BB45C65D17BFB1A" lastPageId="45" lastPageNumber="46" pageId="43" pageNumber="44" type="description">
<paragraph id="8BEF2220FF23FFC41BB45C65D12DF85B" blockId="43.[151,1436,1396,1927]" pageId="43" pageNumber="44">
<emphasis id="B924FE32FF23FFC41BB45C65D461F8E2" bold="true" box="[199,344,1830,1855]" pageId="43" pageNumber="44">Description.</emphasis>
Head, thorax, appendages and basal segments of the abdomen generotypic. Males micropterous, females macropterous. Body length:
<typeStatus id="54EB9C82FF23FFC419375C09D79FF8BE" box="[580,678,1866,1891]" pageId="43" pageNumber="44" type="holotype">holotype</typeStatus>
5.5 mm, male
<typeStatus id="54EB9C82FF23FFC4182E5C08D6F1F8BE" box="[861,968,1867,1891]" pageId="43" pageNumber="44" type="paratype">paratypes</typeStatus>
5.5, female
<typeStatus id="54EB9C82FF23FFC41F135C08D1F9F8BE" box="[1120,1216,1867,1891]" pageId="43" pageNumber="44" type="paratype">paratype</typeStatus>
8.0 mm; forewing length:
<typeStatus id="54EB9C82FF23FFC41B9F5C2DD477F85A" box="[236,334,1902,1927]" pageId="43" pageNumber="44" type="holotype">holotype</typeStatus>
1.2 mm, male
<typeStatus id="54EB9C82FF23FFC41A855C2CD75BF85A" box="[502,610,1903,1927]" pageId="43" pageNumber="44" type="paratype">paratypes</typeStatus>
1.2–1.4 mm, female
<typeStatus id="54EB9C82FF23FFC418235C2CD689F85A" box="[848,944,1903,1927]" pageId="43" pageNumber="44" type="paratype">paratype</typeStatus>
7.0 mm.
</paragraph>
<caption id="DF2F72A8FF24FFC31BE45CEFD074F83C" ID-DOI="http://doi.org/10.5281/zenodo.4919122" ID-Zenodo-Dep="4919122" httpUri="https://zenodo.org/record/4919122/files/figure.png" pageId="44" pageNumber="45" startId="44.[151,264,1964,1987]" targetBox="[237,1353,206,1945]" targetPageId="44">
<paragraph id="8BEF2220FF24FFC31BE45CEFD074F83C" blockId="44.[151,1436,1964,2018]" pageId="44" pageNumber="45">
<emphasis id="B924FE32FF24FFC31BE45CEFD46CF81F" bold="true" box="[151,341,1964,1987]" pageId="44" pageNumber="45">FIGURES 86–89.</emphasis>
Male terminalia of
<taxonomicName id="4C5059A3FF24FFC319505CEED697F81E" authority="Muranyi &amp; Gamboa" authorityName="Muranyi &amp; Gamboa" authorityYear="2014" box="[547,942,1965,1987]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="kovacsi" status="sp. nov.">
<emphasis id="B924FE32FF24FFC319505CEED7EDF81E" box="[547,724,1965,1987]" italics="true" pageId="44" pageNumber="45">Zwicknia kovacsi</emphasis>
Murányi &amp; Gamboa
</taxonomicName>
,
<emphasis id="B924FE32FF24FFC318C95CEED6CDF81F" bold="true" box="[954,1012,1965,1986]" pageId="44" pageNumber="45">
<taxonomicNameLabel id="A2174349FF24FFC318C95CEED6CDF81F" box="[954,1012,1965,1986]" pageId="44" pageNumber="45" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; paratype, Romania, Maramureş County, Rodna Mts., Borşa-Staţiunea Borşa
<emphasis id="B924FE32FF24FFC31A8F5C8FD72BF83C" box="[508,530,1996,2017]" italics="true" pageId="44" pageNumber="45">—</emphasis>
86: dorsal view; 87: ventral view; 88: lateral view; 89: caudal view—scale 1 mm.
</paragraph>
</caption>
<paragraph id="8BEF2220FF25FFC21BB45BD4D741FE28" blockId="45.[151,1437,151,1511]" pageId="45" pageNumber="46">
Male terminalia (
<figureCitation id="136B3EA5FF25FFC21AF45BD4D737FF6D" box="[391,526,151,176]" captionStart="FIGURES 86–89" captionStartId="44.[151,264,1964,1987]" captionTargetBox="[237,1353,206,1945]" captionTargetId="figure-60@44.[219,1370,193,1959]" captionTargetPageId="44" captionText="FIGURES 86–89. Male terminalia of Zwicknia kovacsi Murányi &amp; Gamboa, sp. n.; paratype, Romania, Maramureş County, Rodna Mts., Borşa-Staţiunea Borşa—86: dorsal view; 87: ventral view; 88: lateral view; 89: caudal view—scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919122" httpUri="https://zenodo.org/record/4919122/files/figure.png" pageId="45" pageNumber="46">Figs. 86–89</figureCitation>
,
<figureCitation id="136B3EA5FF25FFC2196A5BD4D77EFF6D" box="[537,583,151,176]" captionStart="FIGURES 120–123" captionStartId="57.[151,264,1761,1784]" captionTargetBox="[165,1401,468,1695]" captionTargetId="figure-157@57.[151,1436,457,1720]" captionTargetPageId="57" captionText="FIGURES 120–123. Male terminalia of Zwicknia Murányi, gen. n., dorso-caudal view—120: Z. bifrons (Newman, 1838); 121: Z. acuta Murányi &amp; Orci, sp. n.; 122: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 123: Z. kovacsi Murányi &amp; Gamboa, sp. n.—scale 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919142" httpUri="https://zenodo.org/record/4919142/files/figure.png" pageId="45" pageNumber="46">123</figureCitation>
): Process of Tg 9 high and wide, its apex 1½× wider, membranous section in dorsal view 2 × wider than the medial swollen section of Ep-scl; apex perpendicularly elevated but the base slightly fold backwards; its sides are slightly sinuous, the apex broadly incised and bear two distinct horns; sides slightly indenting in caudal view, the membranous portion narrowest in the apical part (
<figureCitation id="136B3EA5FF25FFC21F6A5A47D145FEC0" box="[1049,1148,260,285]" captionStart="FIGURES 148–164" captionStartId="60.[151,264,1223,1246]" captionTargetBox="[166,1423,196,1203]" captionTargetId="figure-337@60.[151,1436,181,1215]" captionTargetPageId="60" captionText="FIGURES 148–164. Humps of male Tergite 9 of Zwicknia Murányi, gen. n., caudal view—148–151: Z. acuta Murányi &amp; Orci, sp. n.; 152–156: Z. bifrons (Newman, 1838); 157–159: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 160: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 161–162: Z. sevanica (Zhiltzova, 1964); 163: Z. tuberculata (Zhiltzova, 1964); 164: Z. turkestanica turkestanica (Kimmins, 1950) (148: Slovakia, Krupinská Planina, Pôtor; 149: Hungary, Zemplén Mts., Kishuta; 150: Serbia, Fruska Gora, Vrdnik; 151: Serbia, Zlatibor Mts., Crni Rzav; 152: Hungary, Börzsöny Mts., Szokolya-Királyrét; 153: Hungary, Mátra Mts., Gyöngyössolymos; 154: Hungary, Bükk Mts., Nagyvisnyó; 155: Hungary, Bakony Mts., Bakonybél; 156: Serbia, Maljen Mts., Brajkovići; 157, 159: Croatia, Ivanščica Mts., Stari Golubovec; 158: Hungary, Mecsek Mts., Komló-Zobákpuszta; 160: Romania, Rodna Mts., Borşa-Staţiunea Borşa; 161–162: Turkey, Akyarma Pass; 163: Kazakhstan, Karatau Mts., Bajaldyrtal; 164: Kazakhstan, Tarbagatai Mts., Ayagos)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919148" httpUri="https://zenodo.org/record/4919148/files/figure.png" pageId="45" pageNumber="46">Fig. 160</figureCitation>
). Tg 10, B-scl and Lb-scl generotypic. Ep-scl wide and blunt in dorsal view, medially swollen, its medial width is nearly the same like basal width; tip straight in lateral view, divided section short. Ventral membranous part between the division of Ep-scl ends far before the base in lateral view; apical spines thin and long, distributed not only on the membranous part but extend to the Ep-scl (
<figureCitation id="136B3EA5FF25FFC21AC85AD7D711FE70" box="[443,552,404,429]" captionStart="FIGURES 110–113" captionStartId="55.[151,264,1953,1976]" captionTargetBox="[151,1436,1069,1940]" captionTargetId="figure-316@55.[151,1436,1069,1940]" captionTargetPageId="55" captionText="FIGURES 110–113. Lateral view of male epiproct of Zwicknia Murányi, gen. n.—110: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 111: Z. sevanica (Zhiltzova, 1964); 112: Z. tuberculata (Zhiltzova, 1964); 113: Z. turkestanica turkestanica (Kimmins, 1950)—scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919138" httpUri="https://zenodo.org/record/4919138/files/figure.png" pageId="45" pageNumber="46">Figs. 110</figureCitation>
,
<figureCitation id="136B3EA5FF25FFC219465AD7D758FE70" box="[565,609,404,429]" captionStart="FIGURES 114–119" captionStartId="56.[151,265,1540,1563]" captionTargetBox="[151,1436,193,1520]" captionTargetId="figure-203@56.[151,1436,193,1520]" captionTargetPageId="56" captionText="FIGURES 114–119. Dorsal view of male epiproct of Zwicknia Murányi, gen. n.—114: Z. bifrons (Newman, 1838); 115, 117: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 116: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 118: Z. sevanica (Zhiltzova, 1964); 119: Z. acuta Murányi &amp; Orci, sp. n.—scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919140" httpUri="https://zenodo.org/record/4919140/files/figure.png" pageId="45" pageNumber="46">116</figureCitation>
,
<figureCitation id="136B3EA5FF25FFC2191D5AD7D7EDFE70" box="[622,724,404,429]" captionStart="FIGURES 127–147" captionStartId="59.[151,264,1277,1300]" captionTargetBox="[154,1418,219,1236]" captionTargetId="figure-173@59.[151,1436,193,1256]" captionTargetPageId="59" captionText="FIGURES 127–147. Male epiproct of Zwicknia Murányi, gen. n.—127–130 Z. acuta Murányi &amp; Orci, sp. n.; 131–132: Z. kovacsi Murányi &amp; Gamboa, sp. n.; 133–138: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 139–143: Z. bifrons (Newman, 1838); 144–145: Z. sevanica (Zhiltzova, 1964); 146: Z. tuberculata (Zhiltzova, 1964); 147: Z. turkestanica turkestanica (Kimmins, 1950); 127–131, 133, 135, 137, 139–147: lateral view; 132, 134, 136, 138: dorsal view (127: Slovakia, Krupinská Planina, Pôtor; 128: Hungary, Zemplén Mts., Kishuta; 129: Serbia, Fruska Gora, Vrdnik; 130: Serbia, Zlatibor Mts., Crni Rzav; 131–132: Romania, Rodna Mts., Borşa–Staţiunea Borşa; 133–134, 137–138: Croatia, Ivanščica Mts., Stari Golubovec; 135–136: Hungary, Mecsek Mts., Komló-Zobákpuszta; 139: Hungary, Börzsöny Mts., Szokolya-Királyrét; 140: Hungary, Mátra Mts., Gyöngyössolymos; 141: Hungary, Bükk Mts., Nagyvisnyó; 142: Hungary, Bakony Mts., Bakonybél; 143: Serbia, Maljen Mts., Brajkovići; 144–145: Turkey, Akyarma Pass; 146: Kazakhstan, Karatau Mts., Bajaldyrtal; 147: Kazakhstan, Tarbagatai Mts., Ayagos)—scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919146" httpUri="https://zenodo.org/record/4919146/files/figure.png" pageId="45" pageNumber="46">131–132</figureCitation>
). I-scl generotypic, Ec short and contorted in the three known specimens. St 9 not projecting medially, vesicle small. Sg rounded with not so pronounced triangular shape, tip rounded. Pp, Fp, Rp and cerci generotypic.
</paragraph>
<paragraph id="8BEF2220FF25FFC21BB45ABCD078FDE0" blockId="45.[151,1437,151,1511]" pageId="45" pageNumber="46">Female subgenital plate: Rectangular, posterior margin rounded, slightly incised and equal to the segment’s posterior margin. Antero-lateral recess distinct, the plate is entirely brown; lateral sclerites relatively large.</paragraph>
<paragraph id="8BEF2220FF25FFC21BB45904D4FCFCCA" blockId="45.[151,1437,151,1511]" pageId="45" pageNumber="46">
Drumming: Males produce single, short monophasic calls sporadically. Within each call, beat repetition frequency is nearly constant (see
<figureCitation id="136B3EA5FF25FFC2196A592FD74DFD58" box="[537,628,620,645]" captionStart="FIGURE 190" captionStartId="64.[151,250,1125,1147]" captionTargetBox="[163,1430,185,1044]" captionTargetId="figure-392@64.[151,1436,177,1058]" captionTargetPageId="64" captionText="FIGURE 190. Inter-beat interval variation during male calls in the four examined Zwicknia Murányi, gen. n. species.—Box plot colors: white—Z. bifrons (Newman, 1838), red—Z. acuta Murányi &amp; Orci, sp. n., blue—Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n. In Z. kovacsi Murányi &amp; Gamboa, sp. n. we have drumming samples only from two specimens therefore in the case of that species two simple mark series are used to show inter-beat intervals in their calls (circles for specimen 1, and triangles for specimen 2). i1, i2, i3 and so on are labels containing the serial number of inter-beat intervals counting them from the beginning of each call. Box plots show the duration values measured in 7 specimens in Z. bifrons and Z. acuta, and 5 specimens in Z. rupprechti. Vertical axis has a logarithmic scale. Only calls recorded at an ambient air temperature of 17–19 oC were used. (see the measurement data in Appendix Tables 1–4)." figureDoi="http://doi.org/10.5281/zenodo.4919160" httpUri="https://zenodo.org/record/4919160/files/figure.png" pageId="45" pageNumber="46">Fig 190</figureCitation>
, Appendix
<tableCitation id="C6D2179BFF25FFC21989592FD66AFD58" box="[762,851,620,645]" captionStart="TABLE 4" captionStartId="11.[147,247,534,560]" captionTargetBox="[387,1632,635,1702]" captionTargetId="graphics-128@11.[373,1650,628,1703]" captionText="TABLE 4. Male terminalia (besides epiproct) of valid extant Capniidae genera. — in case of two character states the first or the non-parenthesised one refers to the type species; for abbreviations of parts see Figs.7-10, 23, 35, 81." httpUri="http://table.plazi.org/id/DF2F72A8FF03FFE41BE05955D16DFD8F" pageId="45" pageNumber="46" tableUuid="DF2F72A8FF03FFE41BE05955D16DFD8F">Table 4</tableCitation>
). A call is composed of beats with nearly constant amplitude. See
<figureCitation id="136B3EA5FF25FFC21A3659CCD489FD75" box="[325,432,655,680]" captionStart="FIGURES 165–168" captionStartId="61.[151,264,1330,1353]" captionTargetBox="[162,1428,857,1282]" captionTargetId="figure-628@61.[151,1436,850,1309]" captionTargetPageId="61" captionText="FIGURES 165–168. Slow oscillograms of male calls of four Zwicknia Murányi, gen. n. species, showing call sequences versus sporadic call production—165: two call sequence of Z. bifrons (Newman, 1838), &quot;//&quot; indicates where 2 minutes 23 seconds of silence was cut out from the recording; 166: three call sequences of Z. acuta Murányi &amp; Orci, sp. n.; 167: sporadic calls of Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 168: sporadic call production in Z. kovacsi Murányi &amp; Gamboa, sp. n.—time scale is the same for the four oscillograms; for detailed data on the animals and recording circumstances, see Table 9." figureDoi="http://doi.org/10.5281/zenodo.4919154" httpUri="https://zenodo.org/record/4919154/files/figure.png" pageId="45" pageNumber="46">Figs. 168</figureCitation>
,
<figureCitation id="136B3EA5FF25FFC21ACF59CCD727FD75" box="[444,542,655,680]" captionStart="FIGURES 169–184" captionStartId="62.[151,264,1845,1868]" captionTargetBox="[201,1412,198,1820]" captionTargetId="figure-140@62.[168,1412,183,1822]" captionTargetPageId="62" captionText="FIGURES 169–184. Male drumming calls of Zwicknia Murányi, gen. n.—169–173: Z. bifrons (Newman, 1838); 174–178: Z. acuta Murányi &amp; Orci, sp. n.; 179–182: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 183–184: Z. kovacsi Murányi &amp; Gamboa, sp. n. (see detailed data on the animals and recording circumstances in Table 9)—Oscillograms in Figs. 169–179, 181, 183 have the same time axis resolution (time scale bar is presented in Fig. 169 for all of them), an other time scale is used in the faster oscillograms of Figs. 180, 182, 184 (time scale bar for them can be found in Fig. 184), those are zoomed sections of Figs. 179, 181, 183, respectively." figureDoi="http://doi.org/10.5281/zenodo.4919156" httpUri="https://zenodo.org/record/4919156/files/figure.png" pageId="45" pageNumber="46">183–184</figureCitation>
for the oscillographic pattern of the male drumming calls of this species. Mean values of the examined drumming call parameters for the two studied specimens (male 1, male 2): DC (ms): 157.4, 154; NBC: 6.3, 6; MBI (ms): 29.8, 30.8; air temperature (
<superScript id="7C258F68FF25FFC218425994D601FD38" attach="right" box="[817,824,727,741]" fontSize="6" pageId="45" pageNumber="46">o</superScript>
C): 17.7, 17.4. The pattern of male-female drumming duet is presently unknown.
</paragraph>
<paragraph id="8BEF2220FF25FFC21BB45861D63FFC82" blockId="45.[151,1437,151,1511]" pageId="45" pageNumber="46">
Genetics: The phylogeny (
<figureCitation id="136B3EA5FF25FFC21A845861D760FCE6" box="[503,601,802,827]" captionStart="FIGURE 192" captionStartId="66.[151,249,1180,1202]" captionTargetBox="[156,1424,193,1155]" captionTargetId="figure-240@66.[151,1433,185,1171]" captionTargetPageId="66" captionText="FIGURE 192. Phylogenetic tree of cox1 including Zwicknia Murányi, gen. n. with our accession codes (300941-301003), and GenBank closest species (with alphanumeric accession codes) as outgroup. MP and ML had same topology. Numbers on braches are ML/MP bootstrap values&gt;50%. Species names right to the tree refer to morphological species. P-Distance table is showed." figureDoi="http://doi.org/10.5281/zenodo.4919164" httpUri="https://zenodo.org/record/4919164/files/figure.png" pageId="45" pageNumber="46">Fig. 192</figureCitation>
) indicates a well-supported node for the Romanian population separated from other
<taxonomicName id="4C5059A3FF25FFC21A665805D445FC82" authorityName="Muranyi" authorityYear="2014" box="[277,380,838,863]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="B924FE32FF25FFC21A665805D445FC82" box="[277,380,838,863]" italics="true" pageId="45" pageNumber="46">Zwicknia</emphasis>
</taxonomicName>
populations ranging from 2%–3%.
</paragraph>
<paragraph id="8BEF2220FF25FFC21BB45829D17BFB1A" blockId="45.[151,1437,151,1511]" pageId="45" pageNumber="46">
<emphasis id="B924FE32FF25FFC21BB45829D405FC5F" bold="true" box="[199,316,874,899]" pageId="45" pageNumber="46">Affinities.</emphasis>
This species is morphologically distinct, and readily separated from other
<taxonomicName id="4C5059A3FF25FFC21FFA5829D1D6FC5E" authorityName="Muranyi" authorityYear="2014" box="[1161,1263,874,899]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="B924FE32FF25FFC21FFA5829D1D6FC5E" box="[1161,1263,874,899]" italics="true" pageId="45" pageNumber="46">Zwicknia</emphasis>
</taxonomicName>
known species on the basis of swollen Ep-scl and very wide, distinctly bicornuate process of Tg 9. Females of this species are difficult to reliably separate from other
<taxonomicName id="4C5059A3FF25FFC219F858F1D662FC16" box="[651,859,946,971]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B924FE32FF25FFC219F858F1D7CBFC16" box="[651,754,946,971]" italics="true" pageId="45" pageNumber="46">Zwicknia</emphasis>
species.
</taxonomicName>
The male drumming calls of this species are conspicuously different from those of
<taxonomicName id="4C5059A3FF25FFC2193B589BD78DFC32" authorityName="Murányi &amp; Gamboa &amp; Orci" authorityYear="2014" baseAuthorityName="Newman" baseAuthorityYear="1838" box="[584,692,982,1007]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="bifrons">
<emphasis id="B924FE32FF25FFC2193B589BD78DFC32" box="[584,692,982,1007]" italics="true" pageId="45" pageNumber="46">Z. bifrons</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5059A3FF25FFC2199E589BD673FC32" authorityName="Muranyi &amp; Orci" authorityYear="2014" box="[749,842,984,1007]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="acuta">
<emphasis id="B924FE32FF25FFC2199E589BD673FC32" box="[749,842,984,1007]" italics="true" pageId="45" pageNumber="46">Z. acuta</emphasis>
</taxonomicName>
which produce much longer calls with much slower beat repetition rate. Short call duration and fast beat repetition make the male calls of
<taxonomicName id="4C5059A3FF25FFC21F2558B8D1FEFBCE" authorityName="Muranyi &amp; Gamboa" authorityYear="2014" box="[1110,1223,1018,1043]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="kovacsi">
<emphasis id="B924FE32FF25FFC21F2558B8D1FEFBCE" box="[1110,1223,1018,1043]" italics="true" pageId="45" pageNumber="46">Z. kovacsi</emphasis>
</taxonomicName>
similar to the calls of
<taxonomicName id="4C5059A3FF25FFC21BC65F63D471FBEA" authorityName="Muranyi, Orci &amp; Gamboa" authorityYear="2014" box="[181,328,1054,1079]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="rupprechti">
<emphasis id="B924FE32FF25FFC21BC65F63D471FBEA" box="[181,328,1054,1079]" italics="true" pageId="45" pageNumber="46">Z. rupprechti</emphasis>
</taxonomicName>
. Despite the small sample of drumming calls available from
<taxonomicName id="4C5059A3FF25FFC218805F63D15DFBEA" authorityName="Muranyi &amp; Gamboa" authorityYear="2014" box="[1011,1124,1054,1079]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="kovacsi">
<emphasis id="B924FE32FF25FFC218805F63D15DFBEA" box="[1011,1124,1054,1079]" italics="true" pageId="45" pageNumber="46">Z. kovacsi</emphasis>
</taxonomicName>
our results suggest that this species produces longer calls composed of more beats, repeated more evenly and with longer inter-beat intervals, than in
<taxonomicName id="4C5059A3FF25FFC21B9F5F2BD446FBA2" authorityName="Muranyi, Orci &amp; Gamboa" authorityYear="2014" box="[236,383,1126,1151]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="rupprechti">
<emphasis id="B924FE32FF25FFC21B9F5F2BD446FBA2" box="[236,383,1126,1151]" italics="true" pageId="45" pageNumber="46">Z. rupprechti</emphasis>
</taxonomicName>
. Regarding its longer mean beat interval the male call of
<taxonomicName id="4C5059A3FF25FFC21F7A5F2BD142FBA2" authorityName="Muranyi &amp; Gamboa" authorityYear="2014" box="[1033,1147,1126,1151]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="kovacsi">
<emphasis id="B924FE32FF25FFC21F7A5F2BD142FBA2" box="[1033,1147,1126,1151]" italics="true" pageId="45" pageNumber="46">Z. kovacsi</emphasis>
</taxonomicName>
apparently is less closely similar to Rupprecht's &quot;Capbif&quot; drumming variant (
<bibRefCitation id="EFC15FD1FF25FFC219835FC9D692FB7E" author="Rupprecht, R." box="[752,939,1162,1187]" pageId="45" pageNumber="46" pagination="93 - 98" refId="ref45232" refString="Rupprecht, R. (1997) An attempt to explain different drumming signals within Capnia bifrons. In: Landolt, P. &amp; Sartori, M. (Eds.), Ephemeroptera &amp; Plecoptera, Biology-Ecology-Systematics. MTL-Mauron + Tinguely &amp; Lachat SA., Fribourg, pp. 93 - 98." type="book chapter" year="1997">Rupprecht 1997</bibRefCitation>
) than the male call of
<taxonomicName id="4C5059A3FF25FFC21FB25FC8D061FB7E" authorityName="Muranyi, Orci &amp; Gamboa" authorityYear="2014" box="[1217,1368,1162,1187]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="rupprechti">
<emphasis id="B924FE32FF25FFC21FB25FC8D061FB7E" box="[1217,1368,1162,1187]" italics="true" pageId="45" pageNumber="46">Z. rupprechti</emphasis>
</taxonomicName>
. This species is genetically distinct from the other species with 19 informative characters.
</paragraph>
</subSubSection>
<subSubSection id="C34A71ABFF25FFC21BB45F91D4D1FAA6" pageId="45" pageNumber="46" type="distribution">
<paragraph id="8BEF2220FF25FFC21BB45F91D4D1FAA6" blockId="45.[151,1437,151,1511]" pageId="45" pageNumber="46">
<emphasis id="B924FE32FF25FFC21BB45F91D4CDFB36" bold="true" box="[199,500,1234,1259]" pageId="45" pageNumber="46">Distribution and ecology.</emphasis>
<taxonomicName id="4C5059A3FF25FFC219705F91D7FCFB36" authorityName="Muranyi &amp; Gamboa" authorityYear="2014" box="[515,709,1234,1259]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="kovacsi">
<emphasis id="B924FE32FF25FFC219705F91D7FCFB36" box="[515,709,1234,1259]" italics="true" pageId="45" pageNumber="46">Zwicknia kovacsi</emphasis>
</taxonomicName>
occurs only in the Rodna Mts. of the Eastern Carpathians (
<figureCitation id="136B3EA5FF25FFC21E175F91D5C4FAD2" captionStart="FIGURES 196–197" captionStartId="68.[151,264,1908,1931]" captionTargetBox="[188,1393,187,1898]" captionTargetId="figure-93@68.[188,1393,187,1898]" captionTargetPageId="68" captionText="FIGURES 196–197. Distribution of Zwicknia Murányi, gen. n.—196: global distribution; 197: studied populations of the Carpathian Basin and the northern Balkans—in Fig. 196, white dot on Z. bifrons symbols indicate populations studied by Rupprecht (1997), dotted lines indicate limits of s.l. Z. bifrons literature reports; in Fig. 197, white dot on symbols indicate populations where drumming signals were studied by us, white star indicate type localities." figureDoi="http://doi.org/10.5281/zenodo.4919168" httpUri="https://zenodo.org/record/4919168/files/figure.png" pageId="45" pageNumber="46">Figs. 196–197</figureCitation>
). There are only four additional records of
<taxonomicName id="4C5059A3FF25FFC2187C5FB5D64FFAD2" authorityName="Muranyi" authorityYear="2014" box="[783,886,1270,1295]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="B924FE32FF25FFC2187C5FB5D64FFAD2" box="[783,886,1270,1295]" italics="true" pageId="45" pageNumber="46">Zwicknia</emphasis>
</taxonomicName>
from
<collectingCountry id="F34762B0FF25FFC218B55FB5D112FAD2" box="[966,1067,1270,1295]" name="Romania" pageId="45" pageNumber="46">Romania</collectingCountry>
(
<bibRefCitation id="EFC15FD1FF25FFC21F335FB5D188FAD2" author="Kis, B." box="[1088,1201,1270,1295]" pageId="45" pageNumber="46" pagination="1 - 271" refId="ref43129" refString="Kis, B. (1974) Plecoptera. Fauna Republicii Socialiste Romania, 8 (7), 1 - 271." type="journal article" year="1974">Kis 1974</bibRefCitation>
), the specimens we examined from
<collectingRegion id="4994ECC2FF25FFC21A395E59D4E0FAEE" box="[330,473,1306,1331]" country="Romania" name="Cluj" pageId="45" pageNumber="46">Cluj-Napoca</collectingRegion>
are
<emphasis id="B924FE32FF25FFC2197F5E58D7B9FAEE" box="[524,640,1306,1331]" italics="true" pageId="45" pageNumber="46">
<taxonomicName id="4C5059A3FF25FFC2197F5E58D745FAEE" authorityName="Murányi &amp; Gamboa &amp; Orci" authorityYear="2014" baseAuthorityName="Newman" baseAuthorityYear="1838" box="[524,636,1306,1331]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="bifrons">Z. bifrons</taxonomicName>
,
</emphasis>
and the figures of
<bibRefCitation id="EFC15FD1FF25FFC2182B5E59D6F4FAEE" author="Kis, B." box="[856,973,1306,1331]" pageId="45" pageNumber="46" pagination="1 - 271" refId="ref43129" refString="Kis, B. (1974) Plecoptera. Fauna Republicii Socialiste Romania, 8 (7), 1 - 271." type="journal article" year="1974">Kis (1974</bibRefCitation>
:
<figureCitation id="136B3EA5FF25FFC218AA5E59D15EFAEE" box="[985,1127,1306,1331]" captionStart="FIGURES 62–65" captionStartId="33.[151,264,1508,1531]" captionTargetBox="[151,1436,231,1487]" captionTargetId="figure-310@33.[151,1436,193,1487]" captionTargetPageId="33" captionText="FIGURES 62–65. Female genitalia of Zwicknia Murányi, gen. n.—62–63: Z. bifrons (Newman, 1838); 64: Z. acuta Murányi &amp; Orci, sp. n.; 65: Z. rupprechti Murányi, Orci &amp; Gamboa, sp. n.; 62: terminalia, ventral view; 63: inner genital organs, dorsal view; 64–65: subgenital plates, ventral view (gc: genital cavity; go: genital opening; Sg: subgenital plate)—scales 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4919112" httpUri="https://zenodo.org/record/4919112/files/figure.png" pageId="45" pageNumber="46">Fig. 64 A–C</figureCitation>
) probably also refer to this species. Adults were collected in early April at a single, fast flowing and medium-sized mountain stream at an elevation of
<quantity id="4CA88FC5FF25FFC21A505E21D453FAA7" box="[291,362,1378,1403]" metricMagnitude="2" metricUnit="m" metricValue="9.3" pageId="45" pageNumber="46" unit="m" value="930.0">930 m</quantity>
(
<figureCitation id="136B3EA5FF25FFC21A0A5E21D4E2FAA6" box="[377,475,1378,1403]" captionStart="FIGURES 193–195" captionStartId="67.[151,264,1829,1852]" captionTargetBox="[199,1367,231,1794]" captionTargetId="figure-90@67.[199,1367,231,1810]" captionTargetPageId="67" captionText="FIGURES 193–195. Type localities of the new species described in Zwicknia Murányi, gen. n.—193: Zwicknia acuta Murányi &amp; Orci, sp. n.: Stará rieka, Slovakia, Banskobystrický Region, Krupinská Planina, Pôtor; 194: Zwicknia kovacsi Murányi &amp; Gamboa, sp. n.: Cimpoies Stream, Romania, Maramureş County, Rodna Mts., Borşa-Staţiunea Borşa; 195: Zwicknia rupprechti Murányi, Orci &amp; Gamboa, sp. n.: Völgységi Stream, Hungary, Baranya County, Mecsek Mts., Komló-Zobákpuszta." figureDoi="http://doi.org/10.5281/zenodo.4919166" httpUri="https://zenodo.org/record/4919166/files/figure.png" pageId="45" pageNumber="46">Fig. 194</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34A71ABFF25FFC21BB45EC5D16FFA3A" pageId="45" pageNumber="46" type="etymology">
<paragraph id="8BEF2220FF25FFC21BB45EC5D16FFA3A" blockId="45.[151,1437,151,1511]" pageId="45" pageNumber="46">
<emphasis id="B924FE32FF25FFC21BB45EC5D477FA42" bold="true" box="[199,334,1414,1439]" pageId="45" pageNumber="46">Etymology.</emphasis>
The species is dedicated to Tibor Kovács, Gyöngyös,
<collectingCountry id="F34762B0FF25FFC218C45EC4D123FA42" box="[951,1050,1415,1439]" name="Hungary" pageId="45" pageNumber="46">Hungary</collectingCountry>
, in recognition of his contribution to our knowledge on the
<taxonomicName id="4C5059A3FF25FFC21AC75EE9D713FA1E" box="[436,554,1450,1475]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="order">Plecoptera</taxonomicName>
and other insects of the Carpathian Basin and the Balkans; furthermore, he is one of the collectors of the
<typeStatus id="54EB9C82FF25FFC21ABA5E8CD4C0FA3A" box="[457,505,1487,1511]" pageId="45" pageNumber="46">type</typeStatus>
series. Used as the genitive of a noun of male gender.
</paragraph>
</subSubSection>
</treatment>
</document>
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