179 lines
18 KiB
XML
179 lines
18 KiB
XML
<document id="E0C9D4CEC6AEA770882677D63C63E7FF" ID-DOI="10.11646/zootaxa.2618.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5304087" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629699783393" checkinUser="felipe" docAuthor="Oláh, János & Johanson, Kjell Arne" docDate="2010" docId="03E487BBFFDE2B01FF3CFD3938931B73" docLanguage="en" docName="zt02618p035.pdf" docOrigin="Zootaxa 2618 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2618.1.1" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Nyctiophylax (Paranyctiophylax) anryutah Oláh & Johanson 2010, new species" docType="treatment" docVersion="4" lastPageNumber="15" masterDocId="FFDDFFC3FFD32B0FFFABFF9B39571F41" masterDocTitle="Fifteen new Trichoptera (Insecta) species from Sumatra, Indonesia" masterLastPageNumber="35" masterPageNumber="1" pageNumber="14" updateTime="1699097725960" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:mods id="0BAFF96FB097E67856E366A66C008BB7" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="E698D9EA77E844951142269185E7C50D">
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<mods:title id="9EACBFD5970B1BD9DE34B098B2EA7EAF">Fifteen new Trichoptera (Insecta) species from Sumatra, Indonesia</mods:title>
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</mods:titleInfo>
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<mods:name id="31AA5523E35D6D6FD4E3AA65F9CA61BD" type="personal">
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<mods:roleTerm id="786FFE49FFACB4190B50D3164CC97AD5">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="DEA7285EBA94C05BB53670A600C12811">Oláh, János</mods:namePart>
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<mods:affiliation id="5E6F1F6986278A3F485FE0FA82129C30">Szent István University, Gödöll, Centre of Environmental Health. Residence postal address: Tarján u. 28, H- 4032 Debrecen, Hungary. E-mail: profolah @ gmail. com</mods:affiliation>
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<mods:name id="9328A3B98C55DF0AF3F9C70B4BE4CD24" type="personal">
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<mods:roleTerm id="37C42FAE67DA43FEED8FF94886581410">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="EF75BA6D45E5833402D4F6945CA8D443">Johanson, Kjell Arne</mods:namePart>
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<mods:affiliation id="B1EF7F10EDA48D9037C160BA387D8070">Swedish Museum of Natural History, Entomology Department, Box 50007, S- 10405 Stockholm, Sweden.</mods:affiliation>
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<mods:typeOfResource id="A8257AEB26EC4E2E017DFBEDD48E0A0E">text</mods:typeOfResource>
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<mods:titleInfo id="3D4896B096AB3A1EA0E00D3A8FB52098">
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<mods:title id="3091027587DEC285CD4987C6EC0D66E1">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="68B0AE90FCA6B2F11F172C342AC60C21">
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<mods:date id="1485DB85A225C18FBCAB408F5CFD5447">2010</mods:date>
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<mods:number id="03AA47A6B3C40C42394F7F192F0BBBF6">2010-09-20</mods:number>
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<mods:detail id="4DCCAEE98617EF027E242C04536C5411" type="volume">
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<mods:number id="8601ABB820508919F933FFBC4286AA2A">2618</mods:number>
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<mods:url id="40C1883718E55175AB4727FEA7FF727C">https://biotaxa.org/Zootaxa/article/view/zootaxa.2618.1.1</mods:url>
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</mods:location>
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<mods:classification id="1206C3434B5C9665775807A88E7DB81E">journal article</mods:classification>
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<mods:identifier id="42B5A23CB4422863DA8FB7FCCAACF24F" type="DOI">10.11646/zootaxa.2618.1.1</mods:identifier>
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<mods:identifier id="F16B0B37E8D8EF5C9C3EB050256FD7D9" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="C1A54551E257E953714C55F2F7BF6141" type="Zenodo-Dep">5304087</mods:identifier>
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<treatment id="03E487BBFFDE2B01FF3CFD3938931B73" ID-DOI="http://doi.org/10.5281/zenodo.5311080" ID-Zenodo-Dep="5311080" LSID="urn:lsid:plazi:treatment:03E487BBFFDE2B01FF3CFD3938931B73" httpUri="http://treatment.plazi.org/id/03E487BBFFDE2B01FF3CFD3938931B73" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
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<subSubSection id="C3576526FFDE2B02FF3CFD393A6C1DFD" box="[151,827,674,700]" pageId="13" pageNumber="14" type="nomenclature">
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<paragraph id="8BF236ADFFDE2B02FF3CFD393A6C1DFD" blockId="13.[151,827,674,735]" box="[151,827,674,700]" pageId="13" pageNumber="14">
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<heading id="D0BA81C1FFDE2B02FF3CFD393A6C1DFD" bold="true" box="[151,827,674,700]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
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||
<emphasis id="B939EABFFFDE2B02FF3CFD393A6C1DFD" bold="true" box="[151,827,674,700]" pageId="13" pageNumber="14">
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<taxonomicName id="4C4D4D2EFFDE2B02FF3CFD393BCB1DFD" authority="Oláh & Johanson, 2010" authorityName="Oláh & Johanson" authorityYear="2010" box="[151,668,674,700]" class="Insecta" family="Polycentropodidae" genus="Nyctiophylax" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="anryutah" status="sp. nov." subGenus="Paranyctiophylax">
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<emphasis id="B939EABFFFDE2B02FF3CFD3938601DFD" bold="true" box="[151,311,674,700]" italics="true" pageId="13" pageNumber="14">Nyctiophylax</emphasis>
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(
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<emphasis id="B939EABFFFDE2B02FEECFD393B4B1DFD" bold="true" box="[327,540,674,700]" italics="true" pageId="13" pageNumber="14">Paranyctiophylax</emphasis>
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)
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<emphasis id="B939EABFFFDE2B02FD85FD393BCB1DFD" bold="true" box="[558,668,674,700]" italics="true" pageId="13" pageNumber="14">anryutah</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A20A57C4FFDE2B02FD00FD393A6C1DFD" box="[683,827,674,700]" pageId="13" pageNumber="14" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3576526FFDE2B01FF3CFD5E3D5C1C36" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="description">
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<paragraph id="8BF236ADFFDE2B02FF3CFD5E384D1D9E" blockId="13.[151,827,674,735]" box="[151,282,709,735]" pageId="13" pageNumber="14">
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<figureCitation id="13762A28FFDE2B02FF3CFD5E384D1D9E" box="[151,282,709,735]" captionStart="FIGURES 19–22" captionStartId="13.[151,270,1936,1960]" captionTargetBox="[190,1396,994,1872]" captionTargetId="figure-288@13.[151,1436,961,1914]" captionTargetPageId="13" captionText="FIGURES 19–22. Nyctiophylax (Paranyctiophylax) anryutah, new species, holotype. 19—genitalia, left lateral; 20— genitalia, dorsal; 21—right gonocoxite, ventral; 22—phallic apparatus, left lateral." figureDoi="http://doi.org/10.5281/zenodo.5304109" httpUri="https://zenodo.org/record/5304109/files/figure.png" pageId="13" pageNumber="14">Figs 19–22</figureCitation>
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</paragraph>
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<paragraph id="8BF236ADFFDE2B02FF3CFC8B3C421CDB" blockId="13.[151,1437,784,922]" pageId="13" pageNumber="14">
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This species belongs in the
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<taxonomicName id="4C4D4D2EFFDE2B02FE48FC8B3A971C68" authorityName="Schmid" authorityYear="1958" box="[483,960,784,810]" class="Insecta" family="Polycentropodidae" genus="Nyctiophylax" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="abaya" subGenus="Paranyctiophylax">
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<emphasis id="B939EABFFFDE2B02FE48FC8B3BD71C68" box="[483,640,784,809]" italics="true" pageId="13" pageNumber="14">Nyctiophylax</emphasis>
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(
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<emphasis id="B939EABFFFDE2B02FD39FC8B3A321C68" box="[658,869,784,809]" italics="true" pageId="13" pageNumber="14">Paranyctiophylax</emphasis>
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)
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<emphasis id="B939EABFFFDE2B02FCD3FC8B3A971C68" box="[888,960,784,809]" italics="true" pageId="13" pageNumber="14">abaya</emphasis>
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</taxonomicName>
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species group and is most similar to
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<taxonomicName id="4C4D4D2EFFDE2B02FA2AFC8B38D91C0E" authority="Malicky" authorityName="Malicky" authorityYear="1999" class="Insecta" family="Polycentropodidae" genus="Nyctiophylax" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="archemoros">
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<emphasis id="B939EABFFFDE2B02FA2AFC8B38751C0F" italics="true" pageId="13" pageNumber="14">N. archemoros</emphasis>
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Malicky
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</taxonomicName>
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, from which it differs by having long, crossing, spine-like dorsal paraproctal processes present. In
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<taxonomicName id="4C4D4D2EFFDE2B02FE8BFCC138851C32" authorityName="Malicky" authorityYear="1999" box="[288,466,858,883]" class="Insecta" family="Polycentropodidae" genus="Nyctiophylax" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="archemoros">
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<emphasis id="B939EABFFFDE2B02FE8BFCC138851C32" box="[288,466,858,883]" italics="true" pageId="13" pageNumber="14">N. archemoros</emphasis>
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</taxonomicName>
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the dorsal paraproctal processes are absent. Similar, but not crossing structures were originally described for other species of this species group as parameres of the phallic organ.
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</paragraph>
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<caption id="DF326625FFDE2B02FF3CF80B3D511886" ID-DOI="http://doi.org/10.5281/zenodo.5304109" ID-Zenodo-Dep="5304109" httpUri="https://zenodo.org/record/5304109/files/figure.png" pageId="13" pageNumber="14" startId="13.[151,270,1936,1960]" targetBox="[190,1396,994,1872]" targetPageId="13">
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<paragraph id="8BF236ADFFDE2B02FF3CF80B3D511886" blockId="13.[151,1435,1936,1991]" pageId="13" pageNumber="14">
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<emphasis id="B939EABFFFDE2B02FF3CF80B383718E9" bold="true" box="[151,352,1936,1960]" pageId="13" pageNumber="14">FIGURES 19–22.</emphasis>
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<taxonomicName id="4C4D4D2EFFDE2B02FECCF80A3A6F18E9" authority="Oláh & Johanson, 2010" authorityName="Oláh & Johanson" authorityYear="2010" box="[359,824,1937,1960]" class="Insecta" family="Polycentropodidae" genus="Nyctiophylax" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="anryutah" status="sp. nov." subGenus="Paranyctiophylax">
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||
<emphasis id="B939EABFFFDE2B02FECCF80A3A6F18E9" box="[359,824,1937,1960]" italics="true" pageId="13" pageNumber="14">Nyctiophylax (Paranyctiophylax) anryutah</emphasis>
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||
</taxonomicName>
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,
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<emphasis id="B939EABFFFDE2B02FCEDF80B3A9B18E9" bold="true" box="[838,972,1936,1960]" pageId="13" pageNumber="14">
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||
<taxonomicNameLabel id="A20A57C4FFDE2B02FCEDF80B3A9B18E9" box="[838,972,1936,1960]" pageId="13" pageNumber="14" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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, holotype. 19—genitalia, left lateral; 20— genitalia, dorsal; 21—right gonocoxite, ventral; 22—phallic apparatus, left lateral.
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</paragraph>
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</caption>
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<paragraph id="8BF236ADFFDD2B01FF6EFF0338B51E2D" blockId="14.[151,1437,152,1074]" pageId="14" pageNumber="15">
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Male. Body uniformly pale brown. Maxillary palp formula II-I-IV-III-V, segment III originates subapically from mesal face of segment II. Forewing length 4.2 mm; brown, with hypertrophied longitudinal
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<emphasis id="B939EABFFFDD2B01FF3CFF7939FE1FBA" box="[151,169,226,251]" italics="true" pageId="14" pageNumber="15">R</emphasis>
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vein and 2 hyaline spots: small, rounded spot below discoidal cell around crossvein
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<emphasis id="B939EABFFFDD2B01FB09FF793D9B1FBA" box="[1186,1228,226,251]" italics="true" pageId="14" pageNumber="15">r-m</emphasis>
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in subradial cell; and large triangular spot along crossvein
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<emphasis id="B939EABFFFDD2B01FD39FE933B991E60" box="[658,718,264,289]" italics="true" pageId="14" pageNumber="15">m-cu</emphasis>
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and base of fork
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<emphasis id="B939EABFFFDD2B01FC0DFE933AE91E60" box="[934,958,264,289]" italics="true" pageId="14" pageNumber="15">M</emphasis>
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; discoidal cell and median cell closed; apical forks 2, 3, 4, and 5 present;
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<emphasis id="B939EABFFFDD2B01FD87FEB63B1B1E07" box="[556,588,301,326]" italics="true" pageId="14" pageNumber="15">A1</emphasis>
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,
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<emphasis id="B939EABFFFDD2B01FDF0FEB63B2C1E07" box="[603,635,301,326]" italics="true" pageId="14" pageNumber="15">A2</emphasis>
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, and
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<emphasis id="B939EABFFFDD2B01FD17FEB63B8A1E07" box="[700,733,301,326]" italics="true" pageId="14" pageNumber="15">A3</emphasis>
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looping. Hind wing discoidal cell closed, median cell open, apical forks 2 and 5 present.
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</paragraph>
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<paragraph id="8BF236ADFFDD2B01FF6EFEE338FE1ED3" blockId="14.[151,1437,152,1074]" box="[197,425,376,402]" pageId="14" pageNumber="15">Spur formula 3,4,4.</paragraph>
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<paragraph id="8BF236ADFFDD2B01FF6EFE063D5C1C36" blockId="14.[151,1437,152,1074]" pageId="14" pageNumber="15">Male genitalia. Sternite IX robust, subtriangular in lateral view, with well-developed concavity on each apicodorsal corner at fulcrum. Tergite IX indiscernible, possibly fused to paraproctal body. Paraproctal body with strongly developed dorsal and ventral paraproctal processes. Segment X membranous, weakly visible, partially covered with minute microtrichae. Cerci setose, subtriangular in lateral view; long digitate in dorsal view; shorter than gonopods and ventral paraproctal processes. Paraproctal complex well developed, highly sclerotized. Dorsal paraproctal processes forming pair of large, spine-like structures crossing dorsally, arching dorsad in lateral view; pair of large ventral paraproctal processes directed posterad, each with rounded apex and with slightly ventrad-curving subapical setose hump dorsally. Gonopods with large, quadrangular “elbow” ventrobasally, continuing into almost filiform apical half in lateral view; in ventral view directed mesad, geniculate at meeting point of large “elbow” and filiform apical half. Phallic apparatus located dorsally in genitalia, fixed, guided by paraproctal processes and segment X; tube forming sclerotized phallotheca continuing into matrix of membranous apical half. Weakly discernible phallotremal sclerites present at apical ending of phallotheca and embedded in phallic membrane.</paragraph>
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</subSubSection>
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<subSubSection id="C3576526FFDD2B01FF6EFC193AA51C83" pageId="14" pageNumber="15" type="materials_examined">
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||
<paragraph id="8BF236ADFFDD2B01FF6EFC193AA51C83" blockId="14.[151,1437,152,1074]" pageId="14" pageNumber="15">
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||
<materialsCitation id="3B253CF0FFDD2B01FF6EFC193AA51C83" collectingDate="2008-06-17" collectingDateMax="2008-06-19" collectingDateMin="2008-06-17" collectorName="J. Olah" country="Indonesia" county="Bukit Barisan Selatan National Park" elevation="800" location="Way Pitias" municipality="Barisan Mts." pageId="14" pageNumber="15" specimenCount="1" specimenCount-male="1" stateProvince="Sumatera Utara" typeStatus="holotype">
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<emphasis id="B939EABFFFDD2B01FF6EFC19382D1CDD" bold="true" box="[197,378,898,924]" pageId="14" pageNumber="15">
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<typeStatus id="54F6880FFFDD2B01FF6EFC1938601CDD" box="[197,311,898,924]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
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male
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||
</emphasis>
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:
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<emphasis id="B939EABFFFDD2B01FE21FC193B671CDD" bold="true" box="[394,560,898,924]" pageId="14" pageNumber="15">
|
||
<collectingCountry id="F35A763DFFDD2B01FE21FC193B671CDD" box="[394,560,898,924]" name="Indonesia" pageId="14" pageNumber="15">INDONESIA</collectingCountry>
|
||
</emphasis>
|
||
(
|
||
<collectingRegion id="4989F84FFFDD2B01FD94FC193BFF1CDD" box="[575,680,898,924]" country="Indonesia" name="Sumatera Utara" pageId="14" pageNumber="15">Sumatra</collectingRegion>
|
||
):
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<collectingCounty id="62934E21FFDD2B01FD10FC193D301CDD" box="[699,1127,898,924]" pageId="14" pageNumber="15">Bukit Barisan Selatan National Park</collectingCounty>
|
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,
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<collectingMunicipality id="6B96ACD7FFDD2B01FBD8FC193C5C1CDD" box="[1139,1291,898,924]" pageId="14" pageNumber="15">Barisan Mts.</collectingMunicipality>
|
||
,
|
||
<location id="8E926076FFDD2B01FABCFC193CCF1CDD" LSID="urn:lsid:plazi:treatment:03E487BBFFDE2B01FF3CFD3938931B73:8E926076FFDD2B01FABCFC193CCF1CDD" box="[1303,1432,898,924]" country="Indonesia" county="Bukit Barisan Selatan National Park" municipality="Barisan Mts." name="Way Pitias" pageId="14" pageNumber="15" stateProvince="Sumatera Utara">Way Pitias</location>
|
||
,
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<quantity id="4CB59B48FFDD2B01FF3CFC3339B31C83" box="[151,228,936,962]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="14" pageNumber="15" unit="m" value="800.0">
|
||
<elevation id="0060D19EFFDD2B01FF3CFC3339B31C83" box="[151,228,936,962]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="14" pageNumber="15" unit="m" value="800.0">800 m</elevation>
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</quantity>
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,
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||
<date id="FFF3106DFFDD2B01FF44FC3338CC1C83" box="[239,411,936,962]" pageId="14" pageNumber="15" value="2008-06-17" valueMax="2008-06-19" valueMin="2008-06-17">
|
||
<collectingDate id="EFB7E985FFDD2B01FF44FC3338CC1C83" box="[239,411,936,962]" pageId="14" pageNumber="15" value="2008-06-17" valueMax="2008-06-19" valueMin="2008-06-17">17–19.vi.2008</collectingDate>
|
||
</date>
|
||
,
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<collectingMethod id="520C4EBAFFDD2B01FE0CFC333B461C83" box="[423,529,936,962]" pageId="14" pageNumber="15">light trap</collectingMethod>
|
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[
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<collectorName id="26B8537BFFDD2B01FD89FC333B2E1C83" box="[546,633,936,962]" pageId="14" pageNumber="15">J. Oláh</collectorName>
|
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, Jr.]—
|
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<specimenCount id="9D4BFD24FFDD2B01FD6FFC333A441C83" box="[708,787,936,962]" pageId="14" pageNumber="15" type="male">1 male</specimenCount>
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(OPC, in alcohol).
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||
</materialsCitation>
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||
</paragraph>
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</subSubSection>
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<subSubSection id="C3576526FFDD2B01FF6EFC563B321CA6" box="[197,613,973,999]" pageId="14" pageNumber="15" type="distribution">
|
||
<paragraph id="8BF236ADFFDD2B01FF6EFC563B321CA6" blockId="14.[151,1437,152,1074]" box="[197,613,973,999]" pageId="14" pageNumber="15">
|
||
<emphasis id="B939EABFFFDD2B01FF6EFC5638301CA6" bold="true" box="[197,359,973,999]" pageId="14" pageNumber="15">Distribution:</emphasis>
|
||
<collectingCountry id="F35A763DFFDD2B01FEC4FC5638B61CA6" box="[367,481,973,999]" name="Indonesia" pageId="14" pageNumber="15">Indonesia</collectingCountry>
|
||
(
|
||
<collectingRegion id="4989F84FFFDD2B01FE5AFC563B0E1CA6" box="[497,601,973,999]" country="Indonesia" name="Sumatera Utara" pageId="14" pageNumber="15">Sumatra</collectingRegion>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3576526FFDD2B01FF6EFC6938931B73" pageId="14" pageNumber="15" type="etymology">
|
||
<paragraph id="8BF236ADFFDD2B01FF6EFC6938931B73" blockId="14.[151,1437,152,1074]" pageId="14" pageNumber="15">
|
||
<emphasis id="B939EABFFFDD2B01FF6EFC6938001B4D" bold="true" box="[197,343,1010,1036]" pageId="14" pageNumber="15">Etymology:</emphasis>
|
||
<emphasis id="B939EABFFFDD2B01FECBFC6938981B4A" box="[352,463,1010,1035]" italics="true" pageId="14" pageNumber="15">Anryutah</emphasis>
|
||
, from Sanskrit, meaning “crossness,” referring to the crossing dorsal paraproctal processes in the genitalia.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |