333 lines
43 KiB
XML
333 lines
43 KiB
XML
<document id="88E475CBECE7907A5DC655EB3CD21420" ID-DOI="10.5281/zenodo.158749" ID-GBIF-Dataset="6b8ee1ef-2734-4c25-b99a-892baab60742" ID-ISSN="11755326" ID-Zenodo-Dep="158749" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461024960850" checkinUser="plazi" docAuthor="Shear, William A. & Leonard, William P." docDate="2004" docId="03E43043FFE3FFCBFEB6EF4AFAAFFBEE" docLanguage="en" docName="zt00609.pdf" docOrigin="Zootaxa 609" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Leschius mcallisteri Shear & Leonard, 2004, new species" docType="treatment" docVersion="9" lastPageNumber="6" masterDocId="FFDD483BFFE1FFCEFFBEEE56FFB1FFE8" masterDocTitle="The milliped family Anthroleucosomatidae new to North America: Leschius mcallisteri, n. gen., n. sp. (Diplopoda: Chordeumatida: Anthroleucosomatoidea)" masterLastPageNumber="7" masterPageNumber="1" pageNumber="3" updateTime="1698571675019" updateUser="plazi">
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<mods:titleInfo id="DF58A5178AF78FB5B888EB5620A7C823">
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<mods:title id="AFA57C75F43944297EFBDDCF76C7DE4E">The milliped family Anthroleucosomatidae new to North America: Leschius mcallisteri, n. gen., n. sp. (Diplopoda: Chordeumatida: Anthroleucosomatoidea)</mods:title>
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<mods:namePart id="6DA92E0DEDB6B3029D2C1C2ADBEB70CE">Shear, William A.</mods:namePart>
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<mods:namePart id="B55D9694B4852FEA9015789E196C4B3D">Leonard, William P.</mods:namePart>
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<mods:typeOfResource id="DECBA9F90DE2D0231E7D11D81CC771C1">text</mods:typeOfResource>
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<mods:title id="1F6F69EEDC315E7D8DCE757E7FCE5D21">Zootaxa</mods:title>
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<mods:part id="333F92B030F624F2659EDD2A9F0AA0BD">
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<mods:date id="30B4DA49298D87050D6E10A812F9B626">2004</mods:date>
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<mods:number id="53245B155C52FF11D0C80A2607945FEC">609</mods:number>
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<mods:identifier id="3793A260D0509E761FB8DC597E697266" type="DOI">10.5281/zenodo.158749</mods:identifier>
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<treatment id="03E43043FFE3FFCBFEB6EF4AFAAFFBEE" ID-DOI="http://doi.org/10.5281/zenodo.6272929" ID-GBIF-Taxon="119606123" ID-Zenodo-Dep="6272929" LSID="urn:lsid:plazi:treatment:03E43043FFE3FFCBFEB6EF4AFAAFFBEE" httpUri="http://treatment.plazi.org/id/03E43043FFE3FFCBFEB6EF4AFAAFFBEE" lastPageId="5" lastPageNumber="6" pageId="2" pageNumber="3">
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<subSubSection id="C357D2DEFFE3FFCCFEB6EF4AFEC4FEB1" pageId="2" pageNumber="3" type="nomenclature">
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<paragraph id="8BF28155FFE3FFCCFEB6EF4AFD29FEDD" blockId="2.[264,664,283,345]" box="[264,664,283,310]" pageId="2" pageNumber="3">
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<heading id="D0BA3639FFE3FFCCFEB6EF4AFD29FEDD" bold="true" box="[264,664,283,310]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
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<emphasis id="B9395D47FFE3FFCCFEB6EF4AFD29FEDD" bold="true" box="[264,664,283,310]" pageId="2" pageNumber="3">
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<emphasis id="B9395D47FFE3FFCCFEB6EF4AFDB0FEDE" bold="true" box="[264,513,284,310]" italics="true" pageId="2" pageNumber="3">
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<taxonomicName id="4C4DFAD6FFE3FFCCFEB6EF4AFE4AFEDE" ID-CoL="6Q3DD" box="[264,507,284,310]" class="Diplopoda" family="Anthroleucosomatidae" genus="Leschius" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="mcallisteri" status="sp. nov.">Leschius mcallisteri</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="A20AE03CFFE3FFCCFDB7EF4DFD29FEDD" box="[521,664,283,309]" pageId="2" pageNumber="3" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF28155FFE3FFCCFEB6EF69FEC4FEB1" blockId="2.[264,664,283,345]" box="[264,373,319,345]" pageId="2" pageNumber="3">
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<figureCitation id="13769DD0FFE3FFCCFEB6EF69FEC4FEB1" box="[264,373,319,345]" captionStart="FIGURES 1 – 6" captionStartId="4.[264,383,1731,1755]" captionTargetBox="[274,1307,319,1720]" captionTargetId="figure@4.[247,1339,302,1729]" captionTargetPageId="4" captionText="FIGURES 1 – 6. Leschius mcallisteri, n. sp. 1. Anterior gonopods, anterior view; 2. The same, posterior view; 3. Posterior gonopod (legpair 9), anterior view.; 4. The same, posterior view; 5. Legpair 10 of male, anterior view; 6. Cyphopods, posterior view. Scale line for figures 1 – 5 = 0.1 mm, for figure 6, 0.5 mm." httpUri="https://zenodo.org/record/158750/files/figure.png" pageId="2" pageNumber="3">Figs. 1–5</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C357D2DEFFE3FFCBFEB6EFD9FAAFFBEE" lastPageId="5" lastPageNumber="6" pageId="2" pageNumber="3" type="description">
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<paragraph id="8BF28155FFE3FFCCFEB6EFD9FC8FFDA1" blockId="2.[264,1323,399,1905]" pageId="2" pageNumber="3">
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<typeStatus id="54F63FF7FFE3FFCCFEB6EFD9FEE3FE41" box="[264,338,399,425]" pageId="2" pageNumber="3">Types</typeStatus>
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: Male
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<typeStatus id="54F63FF7FFE3FFCCFE18EFD9FDBCFE41" box="[422,525,399,425]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
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(FMNH),
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<specimenCount id="9D4B4ADCFFE3FFCCFD2EEFD9FD53FE41" box="[656,738,399,425]" pageId="2" pageNumber="3" type="male">4 male</specimenCount>
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and
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<specimenCount id="9D4B4ADCFFE3FFCCFC9EEFD9FC38FE41" box="[800,905,399,425]" pageId="2" pageNumber="3" type="female">4 female</specimenCount>
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<typeStatus id="54F63FF7FFE3FFCCFC2CEFD9FBB3FE41" box="[914,1026,399,425]" pageId="2" pageNumber="3" type="paratype">paratypes</typeStatus>
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(FMNH, AMNH, CAS) from leaf litter at McAllister Springs, near Olympia, Washington,
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<collectingCountry id="F35AC1C5FFE3FFCCFB8DEFE1FBC0FE39" box="[1075,1137,439,465]" name="United States of America" pageId="2" pageNumber="3">USA</collectingCountry>
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, (
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<geoCoordinate id="EE79E792FFE3FFCCFB37EFE1FA97FE39" box="[1161,1318,439,465]" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="1" value="47.0491">N47°02.946’</geoCoordinate>
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,
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<geoCoordinate id="EE79E792FFE3FFCCFEB6EF89FE0BFE11" box="[264,442,479,505]" direction="west" orientation="longitude" pageId="2" pageNumber="3" precision="1" value="-122.72797">W122°43.678’</geoCoordinate>
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), collected
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<date id="FFF3A795FFE3FFCCFDF4EF89FC96FE11" box="[586,807,479,505]" pageId="2" pageNumber="3" value="2004-02-22">22 February 2004</date>
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by W. P. Leonard;
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<specimenCount id="9D4B4ADCFFE3FFCCFB9AEF89FBC8FE11" box="[1060,1145,479,505]" pageId="2" pageNumber="3" type="male">2 male</specimenCount>
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and
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<specimenCount id="9D4B4ADCFFE3FFCCFB03EF89FA98FE11" box="[1213,1321,479,505]" pageId="2" pageNumber="3" type="female">3 female</specimenCount>
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<typeStatus id="54F63FF7FFE3FFCCFEB6EC51FEC9FDC9" box="[264,376,519,545]" pageId="2" pageNumber="3" type="paratype">paratypes</typeStatus>
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(WAS) from near the same locality (
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<geoCoordinate id="EE79E792FFE3FFCCFC8BEC51FC63FDC9" box="[821,978,519,545]" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="1" value="47.047283">N47°02.837’</geoCoordinate>
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,
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<geoCoordinate id="EE79E792FFE3FFCCFC61EC51FB23FDC9" box="[991,1170,519,545]" direction="west" orientation="longitude" pageId="2" pageNumber="3" precision="1" value="-122.7287">W122°43.722’</geoCoordinate>
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), collected
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<date id="FFF3A795FFE3FFCCFAA4EC51FE05FDA1" pageId="2" pageNumber="3" value="2004-02-07">7 February 2004</date>
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by W. P. Leonard and C. Richart.
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</paragraph>
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<paragraph id="8BF28155FFE3FFCCFE86EC01FDC3FD01" blockId="2.[264,1323,399,1905]" pageId="2" pageNumber="3">
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Etymology: The species is named for Olympia naturalist Kelly R. McAllister (1956–), descendant of Chief Scadawah of the Cowlitz Indian Tribe of Washington,
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<collectingCountry id="F35AC1C5FFE3FFCCFB34EC29FB79FD71" box="[1162,1224,639,665]" name="United States of America" pageId="2" pageNumber="3">USA</collectingCountry>
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, and the McAllister Clan, one the first European families to homestead the Puget Sound Region in 1852, near McAllister Springs.
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</paragraph>
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<paragraph id="8BF28155FFE3FFCCFE86ECA1FDDDF941" blockId="2.[264,1323,399,1905]" pageId="2" pageNumber="3">
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Description: Male (
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<figureCitation id="13769DD0FFE3FFCCFDA3ECA1FD3EFCF9" box="[541,655,759,785]" captionStart="FIGURES 1 – 6" captionStartId="4.[264,383,1731,1755]" captionTargetBox="[274,1307,319,1720]" captionTargetId="figure@4.[247,1339,302,1729]" captionTargetPageId="4" captionText="FIGURES 1 – 6. Leschius mcallisteri, n. sp. 1. Anterior gonopods, anterior view; 2. The same, posterior view; 3. Posterior gonopod (legpair 9), anterior view.; 4. The same, posterior view; 5. Legpair 10 of male, anterior view; 6. Cyphopods, posterior view. Scale line for figures 1 – 5 = 0.1 mm, for figure 6, 0.5 mm." httpUri="https://zenodo.org/record/158750/files/figure.png" pageId="2" pageNumber="3">Figs. 1–5</figureCitation>
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): Twentysix segments, at least two segments anterior to epiproct legless. Length, 3.6 mm, width 0.32 mm. Third antennomere about 4 times longer than wide. Ocelli 5 or 6, in single row plus single ocellus below anterior ocelli of single row, moderately wellformed, lightly pigmented. Trunk segments with smooth, shining metaterga, segmental setae smooth, acute, about 1/2 width of terga, two outer setae on each side on common tubercle, all setae strongly curved, directed medially, arching over metaterga. Legpairs 1 and 2 with tarsal combs, coxae 2 with vas deferentia opening through coxae on short tubes. Legpairs 3–7 somewhat incrassate compared to more posterior legpairs, podomeres unmodified. Anterior gonopods in anterior view (
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<figureCitation id="13769DD0FFE3FFCCFB27EA61FB51FBB9" box="[1177,1248,1079,1105]" captionStart="FIGURES 1 – 6" captionStartId="4.[264,383,1731,1755]" captionTargetBox="[274,1307,319,1720]" captionTargetId="figure@4.[247,1339,302,1729]" captionTargetPageId="4" captionText="FIGURES 1 – 6. Leschius mcallisteri, n. sp. 1. Anterior gonopods, anterior view; 2. The same, posterior view; 3. Posterior gonopod (legpair 9), anterior view.; 4. The same, posterior view; 5. Legpair 10 of male, anterior view; 6. Cyphopods, posterior view. Scale line for figures 1 – 5 = 0.1 mm, for figure 6, 0.5 mm." httpUri="https://zenodo.org/record/158750/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
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) with large, blocky coxae (
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<emphasis id="B9395D47FFE3FFCCFDBBEA08FDA3FB90" bold="true" box="[517,530,1118,1144]" pageId="2" pageNumber="3">c</emphasis>
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) set on anteriorly projecting sternal shelf, coxites (
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<emphasis id="B9395D47FFE3FFCCFBC0EA08FB2BFB90" bold="true" box="[1150,1178,1118,1144]" pageId="2" pageNumber="3">cx</emphasis>
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) bladelike, about 4 times longer than wide. In posterior view (
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<figureCitation id="13769DD0FFE3FFCCFCCEEAD1FC0BFB49" box="[880,954,1159,1185]" captionStart="FIGURES 1 – 6" captionStartId="4.[264,383,1731,1755]" captionTargetBox="[274,1307,319,1720]" captionTargetId="figure@4.[247,1339,302,1729]" captionTargetPageId="4" captionText="FIGURES 1 – 6. Leschius mcallisteri, n. sp. 1. Anterior gonopods, anterior view; 2. The same, posterior view; 3. Posterior gonopod (legpair 9), anterior view.; 4. The same, posterior view; 5. Legpair 10 of male, anterior view; 6. Cyphopods, posterior view. Scale line for figures 1 – 5 = 0.1 mm, for figure 6, 0.5 mm." httpUri="https://zenodo.org/record/158750/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
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), sternum greatly enlarged and produced as massive posterior sternal process (
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<emphasis id="B9395D47FFE3FFCCFC92EAF8FC86FB20" bold="true" box="[812,823,1198,1224]" pageId="2" pageNumber="3">s</emphasis>
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), bearing anteriolateral, short, forcepslike projections (these projections grasp posteriorly projecting knobs from coxites), posterior face of process with paired gland channels emerging through pores clearly visible subapically; gland channels continue to large central glandular mass between and dorsal to tracheal apodemes. Posterior gonopods (
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<figureCitation id="13769DD0FFE3FFCCFD6EEB19FCF0FA81" box="[720,833,1359,1385]" captionStart="FIGURES 1 – 6" captionStartId="4.[264,383,1731,1755]" captionTargetBox="[274,1307,319,1720]" captionTargetId="figure@4.[247,1339,302,1729]" captionTargetPageId="4" captionText="FIGURES 1 – 6. Leschius mcallisteri, n. sp. 1. Anterior gonopods, anterior view; 2. The same, posterior view; 3. Posterior gonopod (legpair 9), anterior view.; 4. The same, posterior view; 5. Legpair 10 of male, anterior view; 6. Cyphopods, posterior view. Scale line for figures 1 – 5 = 0.1 mm, for figure 6, 0.5 mm." httpUri="https://zenodo.org/record/158750/files/figure.png" pageId="2" pageNumber="3">Figs. 3, 4</figureCitation>
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) with broad sternum, medially excavated to receive sternal process of anterior gonopods. Coxae large, projecting posteriorly; 4 coxites on each side, coxite 1 short, acute, basally fused to coxite 2, latter broader, longer, more bladelike, coxite 3 slightly sinuous acute rod, coxite 4 divided into 3 or 4 fimbrialike parts.
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<emphasis id="B9395D47FFE3FFCCFE2BEBB9FE5BF9E0" box="[405,490,1519,1544]" italics="true" pageId="2" pageNumber="3">In situ,</emphasis>
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anterior gonopods inclined posteriorly, sternal process fits tightly in median sternal excavation of posterior gonopods, posterior gonopod coxites extend laterally and anteriorly around anterior gonopod coxites. Legpair 10 (
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<figureCitation id="13769DD0FFE3FFCCFBFDE869FB3EF9B1" box="[1091,1167,1599,1625]" captionStart="FIGURES 1 – 6" captionStartId="4.[264,383,1731,1755]" captionTargetBox="[274,1307,319,1720]" captionTargetId="figure@4.[247,1339,302,1729]" captionTargetPageId="4" captionText="FIGURES 1 – 6. Leschius mcallisteri, n. sp. 1. Anterior gonopods, anterior view; 2. The same, posterior view; 3. Posterior gonopod (legpair 9), anterior view.; 4. The same, posterior view; 5. Legpair 10 of male, anterior view; 6. Cyphopods, posterior view. Scale line for figures 1 – 5 = 0.1 mm, for figure 6, 0.5 mm." httpUri="https://zenodo.org/record/158750/files/figure.png" pageId="2" pageNumber="3">Fig. 5</figureCitation>
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) with coxae slightly enlarged, bearing large glands; legpair 11 similar but coxae and glands smaller. Coloration white, ocelli black.
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</paragraph>
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<paragraph id="8BF28155FFE3FFCCFE86E8E1FDAFF911" blockId="2.[264,1323,399,1905]" pageId="2" pageNumber="3">
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Female: length, 4.0 mm, width 0.37 mm. Nonsexual characters as described for male. Cyphopods as in
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<figureCitation id="13769DD0FFE3FFCCFE6CE889FDABF911" box="[466,538,1759,1785]" captionStart="FIGURES 1 – 6" captionStartId="4.[264,383,1731,1755]" captionTargetBox="[274,1307,319,1720]" captionTargetId="figure@4.[247,1339,302,1729]" captionTargetPageId="4" captionText="FIGURES 1 – 6. Leschius mcallisteri, n. sp. 1. Anterior gonopods, anterior view; 2. The same, posterior view; 3. Posterior gonopod (legpair 9), anterior view.; 4. The same, posterior view; 5. Legpair 10 of male, anterior view; 6. Cyphopods, posterior view. Scale line for figures 1 – 5 = 0.1 mm, for figure 6, 0.5 mm." httpUri="https://zenodo.org/record/158750/files/figure.png" pageId="2" pageNumber="3">Fig. 6</figureCitation>
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.
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</paragraph>
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<paragraph id="8BF28155FFE3FFCDFE86E951FC62FE16" blockId="2.[264,1323,399,1905]" lastBlockId="3.[264,1325,284,1790]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
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Natural History Observations: Specimens were handcollected while using an OptiVisor
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<date id="FFF3A795FFE3FFCCFE8BE979FEE8F8A1" box="[309,345,1839,1865]" pageId="2" pageNumber="3">3X</date>
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magnifying visor to search leaf litter along a steep, eastfacing slope in the lower Nisqually River Valley.
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<taxonomicName id="4C4DFAD6FFE3FFCCFD8EE901FCACF898" box="[560,797,1879,1904]" class="Diplopoda" family="Anthroleucosomatidae" genus="Leschius" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="mcallisteri">
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<emphasis id="B9395D47FFE3FFCCFD8EE901FCACF898" box="[560,797,1879,1904]" italics="true" pageId="2" pageNumber="3">Leschius mcallisteri</emphasis>
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</taxonomicName>
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appeared to be limited to the forested “bottoms” along the base of the slope and near perennial springs which form the headwaters of tributaries to McAllister Creek. The site is vegetated by mature secondgrowth forest dominated by bigleaf maple (
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<taxonomicName id="4C4DFAD6FFE2FFCDFDD8EF3AFCF7FE6D" box="[614,838,364,389]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" kingdom="Plantae" order="Sapindales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="macrophylum">
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<emphasis id="B9395D47FFE2FFCDFDD8EF3AFCF7FE6D" box="[614,838,364,389]" italics="true" pageId="3" pageNumber="4">Acer macrophylum</emphasis>
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</taxonomicName>
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), red alder (
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<taxonomicName id="4C4DFAD6FFE2FFCDFC65EF3AFBCFFE6D" box="[987,1150,364,389]" class="Magnoliopsida" family="Betulaceae" genus="Alnus" kingdom="Plantae" order="Fagales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="rubrum">
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<emphasis id="B9395D47FFE2FFCDFC65EF3AFBCFFE6D" box="[987,1150,364,389]" italics="true" pageId="3" pageNumber="4">Alnus rubrum</emphasis>
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</taxonomicName>
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), western redcedar (
|
||
<taxonomicName id="4C4DFAD6FFE2FFCDFEE5EFC2FE4BFE45" box="[347,506,404,429]" class="Pinopsida" family="Cupressaceae" genus="Thuja" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="plicata">
|
||
<emphasis id="B9395D47FFE2FFCDFEE5EFC2FE4BFE45" box="[347,506,404,429]" italics="true" pageId="3" pageNumber="4">Thuja plicata</emphasis>
|
||
</taxonomicName>
|
||
), and sword fern (
|
||
<taxonomicName id="4C4DFAD6FFE2FFCDFD65EFC2FC6AFE45" box="[731,987,404,429]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="munitum">
|
||
<emphasis id="B9395D47FFE2FFCDFD65EFC2FC6AFE45" box="[731,987,404,429]" italics="true" pageId="3" pageNumber="4">Polystichum munitum</emphasis>
|
||
</taxonomicName>
|
||
). We think the presence of perennial springfed streams and mature forest at the
|
||
<typeStatus id="54F63FF7FFE2FFCDFC3AEFEAFC06FE3E" box="[900,951,444,470]" pageId="3" pageNumber="4">type</typeStatus>
|
||
locality are important in maintaining habitat conditions necessary for the species’ survival.
|
||
</paragraph>
|
||
<paragraph id="8BF28155FFE2FFCDFE86EC5AFDA1FD06" blockId="3.[264,1325,284,1790]" pageId="3" pageNumber="4">
|
||
Discussion: Among sympatric or nearly sympatric millipeds,
|
||
<taxonomicName id="4C4DFAD6FFE2FFCDFB9BEC5AFB74FDCD" box="[1061,1221,524,549]" class="Diplopoda" family="Anthroleucosomatidae" genus="Leschius" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="mcallisteri">
|
||
<emphasis id="B9395D47FFE2FFCDFB9BEC5AFB74FDCD" box="[1061,1221,524,549]" italics="true" pageId="3" pageNumber="4">L. mcallisteri</emphasis>
|
||
</taxonomicName>
|
||
may be confused only with the recently described
|
||
<taxonomicName id="4C4DFAD6FFE2FFCDFD41EC62FB66FDA6" authority="Shear and Leonard" authorityName="Shear and Leonard" box="[767,1239,564,590]" class="Diplopoda" family="Microlympiidae" genus="Microlympia" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="echina">
|
||
<emphasis id="B9395D47FFE2FFCDFD41EC62FC5DFDA5" box="[767,1004,564,589]" italics="true" pageId="3" pageNumber="4">Microlympia echina</emphasis>
|
||
Shear and Leonard
|
||
</taxonomicName>
|
||
, which has 28 segments, up to 8 ocelli arranged in 2 rows, and entirely different gonopods (
|
||
<bibRefCitation id="EFDCFCA4FFE2FFCDFB5AEC0AFE54FD76" author="Shear" pageId="3" pageNumber="7" refString="Shear, W. A., & Leonard, W. P. (2003) Microlympiidae, a new milliped family from North America, and Microlympia echina, new genus and species (Diplopoda: Chordeumatida: Brannerioidea). Zootaxa, 243, 1015011." type="journal article" year="2003">Shear and Leonard 2003</bibRefCitation>
|
||
). Undescribed new species of minute, white
|
||
<taxonomicName id="4C4DFAD6FFE2FFCDFC41ECD2FBC6FD76" box="[1023,1143,644,670]" class="Diplopoda" family="Caseyidae" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="family">Caseyidae</taxonomicName>
|
||
are also found in the region, but have 28 or 30 segments, much shorter segmental setae not set on tubercles, and only 2 ocelli.
|
||
</paragraph>
|
||
<paragraph id="8BF28155FFE2FFCDFE86ECAAFDC5FAF6" blockId="3.[264,1325,284,1790]" pageId="3" pageNumber="4">
|
||
The Anthroleucosomini (
|
||
<emphasis id="B9395D47FFE2FFCDFDDCECAAFD14FCFD" box="[610,677,764,789]" italics="true" pageId="3" pageNumber="4">recte:</emphasis>
|
||
Anthroleucosomatini) was set up in 1899 by Verhoeff as a tribe; he raised it to a family in 1910. It has been incorrectly spelled as “Antroleucosomatidae” by some authors (i.e.,
|
||
<bibRefCitation id="EFDCFCA4FFE2FFCDFD2DED1AFC82FC8E" author="Strasser" box="[659,819,844,870]" pageId="3" pageNumber="7" refString="Strasser, K. (1970) Uber enige Diplopoden aus dem westlichen Kaukasus. Revue Suisse de Zoologie, 77, 199 - - 205." type="journal volume" year="1970">Strasser 1970</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFDCFCA4FFE2FFCDFC81ED1AFC4DFC8E" author="Golovatch" box="[831,1020,844,870]" pageId="3" pageNumber="6" refString="Golovatch, S. I. (1981) Some new forms of millipedes (Diplopoda) from the Caucasus. Annales zoologici (Warszawa), 36, 105 - 116." type="journal article" year="1981">Golovatch 1981</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFDCFCA4FFE2FFCDFBB6ED1AFBF6FC8E" author="Golovatch" box="[1032,1095,844,870]" pageId="3" pageNumber="6" refString="Golovatch, S. I. (1984) Two new genera of cave-dwelling millipedes (Diplopoda), with remarks on the millipede fauna of West Caucasian caves. International Journal of Speleology, 14, 39 - 50." type="journal article" year="1984">1984</bibRefCitation>
|
||
);
|
||
<bibRefCitation id="EFDCFCA4FFE2FFCDFBE5ED1AFB48FC8E" author="Jeekel" box="[1115,1273,844,870]" pageId="3" pageNumber="6" refString="Jeekel, C. A. W. (1970) Nomenclator generum et familiarum Diplopodorum: A list of the genus and family-group names in the Class Diplopoda from the 10 th edition of Linnaeus, 1758, to the end of 1957. Monographieen van de Nederlandse Entomologische Vereniging, Amsterdam, 412 pp." type="book" year="1970">Jeekel (1970)</bibRefCitation>
|
||
and Hoffman (1979) establish the correct spelling. Hoffman (1979) proposed an expanded version of the family, including
|
||
<taxonomicName id="4C4DFAD6FFE2FFCDFD02EDCAFC3DFC5E" box="[700,908,924,950]" class="Diplopoda" family="Origmatogonidae" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="family">Origmatogonidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4DFAD6FFE2FFCDFC1AEDCAFBFCFC5E" box="[932,1101,924,950]" class="Diplopoda" family="Anthogonidae" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="family">Anthogonidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4DFAD6FFE2FFCDFBDBEDCAFB6AFC5E" box="[1125,1243,924,950]" class="Diplopoda" family="Haasiidae" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="family">Haasiidae</taxonomicName>
|
||
, Scutogonidae, Brachychaetumatidae,
|
||
<taxonomicName id="4C4DFAD6FFE2FFCDFD1DED92FC14FC36" box="[675,933,964,990]" class="Diplopoda" family="Macrochaeteumatidae" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="family">Macrochaeteumatidae</taxonomicName>
|
||
and Chaemosomatidae as subfamilies. Shear (2000) moved some of these families to the superfamily Brannerioidea and restored others to full family statuses in the Superfamily Anthroleucosomatoidea. The superfamily remains poorly studied, and we anticipate a wholesale rearrangement in the future. A core group of genera originally included in the family occurs from
|
||
<collectingCountry id="F35AC1C5FFE2FFCDFB6CEA32FAB9FB96" box="[1234,1288,1124,1150]" name="Italy" pageId="3" pageNumber="4">Italy</collectingCountry>
|
||
to
|
||
<collectingCountry id="F35AC1C5FFE2FFCDFEB6EADAFEEFFB4E" box="[264,350,1164,1190]" name="Greece" pageId="3" pageNumber="4">Greece</collectingCountry>
|
||
,
|
||
<collectingCountry id="F35AC1C5FFE2FFCDFED7EADAFE7FFB4E" box="[361,462,1164,1190]" name="Bulgaria" pageId="3" pageNumber="4">Bulgaria</collectingCountry>
|
||
and western
|
||
<collectingCountry id="F35AC1C5FFE2FFCDFDD6EADAFD0DFB4E" box="[616,700,1164,1190]" name="Turkey" pageId="3" pageNumber="4">Turkey</collectingCountry>
|
||
(
|
||
<emphasis id="B9395D47FFE2FFCDFD73EADAFD49FB4D" box="[717,760,1164,1189]" italics="true" pageId="3" pageNumber="4">i.e.,</emphasis>
|
||
<bibRefCitation id="EFDCFCA4FFE2FFCDFD40EADAFC10FB4E" author="Strasser" box="[766,929,1164,1190]" pageId="3" pageNumber="7" refString="Strasser, K. (1960) Diplopoden aus Alpen-, Appeninen- und Balkanlandern. Fragmenta entomologica, 3, 95 - 140." type="journal article" year="1960">Strasser 1960</bibRefCitation>
|
||
). More recently, genera and species have been described that extend the family’s distribution to
|
||
<collectingCountry id="F35AC1C5FFE2FFCDFC46EAE2FB96FB26" box="[1016,1063,1204,1230]" name="Iran" pageId="3" pageNumber="4">Iran</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFDCFCA4FFE2FFCDFB88EAE2FB6DFB26" author="Mauries" box="[1078,1244,1204,1230]" pageId="3" pageNumber="6" refString="Mauries, J. - P. (1982) Diplopod de l'Iran: Persedicus n. gen. (Craspedosomida: Anthroleucosomatidae). Senckenbergiana biologia, 62, 385 - 390." type="journal article" year="1982">Mauriès 1982</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFDCFCA4FFE2FFCDFB56EAE2FEF7FB1E" author="Shear" pageId="3" pageNumber="7" refString="Shear, W. A. (1988) Systematic position of the milliped species Alloiopus solitarius Attems and the genus Ghilarovia Gulicka (Chordeumatida, Anthroleucosomatidae). Myriapodologica, 2, 51 - 58." type="journal article" year="1988">Shear 1988</bibRefCitation>
|
||
), the Georgian Caucasus (
|
||
<bibRefCitation id="EFDCFCA4FFE2FFCDFDC8EA8AFCA8FB1E" author="Strasser" box="[630,793,1244,1270]" pageId="3" pageNumber="7" refString="Strasser, K. (1970) Uber enige Diplopoden aus dem westlichen Kaukasus. Revue Suisse de Zoologie, 77, 199 - - 205." type="journal volume" year="1970">Strasser 1970</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFDCFCA4FFE2FFCDFC9AEA8AFC52FB1E" author="Golovatch" box="[804,995,1244,1270]" pageId="3" pageNumber="6" refString="Golovatch, S. I. (1981) Some new forms of millipedes (Diplopoda) from the Caucasus. Annales zoologici (Warszawa), 36, 105 - 116." type="journal article" year="1981">Golovatch 1981</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFDCFCA4FFE2FFCDFC50EA8AFB9DFB1E" author="Golovatch" box="[1006,1068,1244,1270]" pageId="3" pageNumber="6" refString="Golovatch, S. I. (1984) Two new genera of cave-dwelling millipedes (Diplopoda), with remarks on the millipede fauna of West Caucasian caves. International Journal of Speleology, 14, 39 - 50." type="journal article" year="1984">1984</bibRefCitation>
|
||
), and the Novosibirsk region of Siberia (
|
||
<bibRefCitation id="EFDCFCA4FFE2FFCDFE62EB52FDD9FAF6" author="Shear" box="[476,616,1284,1310]" pageId="3" pageNumber="7" refString="Shear, W. A. (1988) Systematic position of the milliped species Alloiopus solitarius Attems and the genus Ghilarovia Gulicka (Chordeumatida, Anthroleucosomatidae). Myriapodologica, 2, 51 - 58." type="journal article" year="1988">Shear 1988</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF28155FFE2FFCDFE86EB7AFCFBF916" blockId="3.[264,1325,284,1790]" pageId="3" pageNumber="4">
|
||
Of the described genera,
|
||
<taxonomicName id="4C4DFAD6FFE2FFCDFDE1EB7AFD77FAAD" box="[607,710,1324,1349]" class="Diplopoda" family="Anthroleucosomatidae" genus="Leschius" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9395D47FFE2FFCDFDE1EB7AFD77FAAD" box="[607,710,1324,1349]" italics="true" pageId="3" pageNumber="4">Leschius</emphasis>
|
||
</taxonomicName>
|
||
seems closer in gonopod design to three monotypic genera known from the Caucasus:
|
||
<taxonomicName id="4C4DFAD6FFE2FFCDFD06EB02FBB0FA86" authority="Strasser 1971" authorityName="Strasser" authorityYear="1971" box="[696,1025,1364,1390]" class="Diplopoda" family="Anthroleucosomatidae" genus="Caucaseuma" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9395D47FFE2FFCDFD06EB02FCE1FA85" box="[696,848,1364,1389]" italics="true" pageId="3" pageNumber="4">Caucaseuma</emphasis>
|
||
Strasser 1971
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4DFAD6FFE2FFCDFBADEB02FEF4FA7E" authority="Golovatch 1981" authorityName="Golovatch" authorityYear="1981" class="Diplopoda" family="Anthroleucosomatidae" genus="Adshardicus" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9395D47FFE2FFCDFBADEB02FB14FA85" box="[1043,1189,1364,1389]" italics="true" pageId="3" pageNumber="4">Adshardicus</emphasis>
|
||
<bibRefCitation id="EFDCFCA4FFE2FFCDFB0EEB02FEF4FA7E" author="Golovatch" pageId="3" pageNumber="6" refString="Golovatch, S. I. (1981) Some new forms of millipedes (Diplopoda) from the Caucasus. Annales zoologici (Warszawa), 36, 105 - 116." type="journal article" year="1981">Golovatch 1981</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C4DFAD6FFE2FFCDFE37EB2AFD68FA7E" authority="Golovatch 1985" authorityName="Golovatch" authorityYear="1985" box="[393,729,1404,1430]" class="Diplopoda" family="Anthroleucosomatidae" genus="Ratcheuma" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9395D47FFE2FFCDFE37EB2AFDBCFA7D" box="[393,525,1404,1429]" italics="true" pageId="3" pageNumber="4">Ratcheuma</emphasis>
|
||
Golovatch 1985
|
||
</taxonomicName>
|
||
. All feature a large, backward projecting sternal lobe, and in both
|
||
<taxonomicName id="4C4DFAD6FFE2FFCDFE62EBF2FDDFFA55" box="[476,622,1444,1469]" class="Diplopoda" family="Anthroleucosomatidae" genus="Adshardicus" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9395D47FFE2FFCDFE62EBF2FDDFFA55" box="[476,622,1444,1469]" italics="true" pageId="3" pageNumber="4">Adshardicus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B9395D47FFE2FFCDFD17EBF2FC84FA55" box="[681,821,1444,1469]" italics="true" pageId="3" pageNumber="4">
|
||
<taxonomicName id="4C4DFAD6FFE2FFCDFD17EBF2FC81FA55" box="[681,816,1444,1469]" class="Diplopoda" family="Anthroleucosomatidae" genus="Ratcheuma" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Ratcheuma</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
there is evidence of a glandular complex, with paired ducts entering the sternal lobe. However, in each of these three genera, the posterior gonopods are closely appressed to one another, not widely separated as in
|
||
<emphasis id="B9395D47FFE2FFCDFAADEBA2FE3CF9DD" italics="true" pageId="3" pageNumber="4">
|
||
<taxonomicName id="4C4DFAD6FFE2FFCDFAADEBA2FE38F9DD" class="Diplopoda" family="Anthroleucosomatidae" genus="Leschius" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="mcallisteri">L. mcallisteri</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The posterior gonopods, or ninth legpair, of
|
||
<taxonomicName id="4C4DFAD6FFE2FFCDFC7CE84AFBE2F9DD" box="[962,1107,1564,1589]" class="Diplopoda" family="Anthroleucosomatidae" genus="Adshardicus" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9395D47FFE2FFCDFC7CE84AFBE2F9DD" box="[962,1107,1564,1589]" italics="true" pageId="3" pageNumber="4">Adshardicus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4DFAD6FFE2FFCDFB2DE84AFA9BF9DD" box="[1171,1322,1564,1589]" class="Diplopoda" family="Anthroleucosomatidae" genus="Caucaseuma" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9395D47FFE2FFCDFB2DE84AFA9BF9DD" box="[1171,1322,1564,1589]" italics="true" pageId="3" pageNumber="4">Caucaseuma</emphasis>
|
||
</taxonomicName>
|
||
have very small telopodite remnants; these are entirely absent in
|
||
<taxonomicName id="4C4DFAD6FFE2FFCDFBBEE812FB35F9B5" box="[1024,1156,1604,1629]" class="Diplopoda" family="Anthroleucosomatidae" genus="Ratcheuma" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9395D47FFE2FFCDFBBEE812FB35F9B5" box="[1024,1156,1604,1629]" italics="true" pageId="3" pageNumber="4">Ratcheuma</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B9395D47FFE2FFCDFB02E812FA9BF9B5" box="[1212,1322,1604,1629]" italics="true" pageId="3" pageNumber="4">
|
||
<taxonomicName id="4C4DFAD6FFE2FFCDFB02E812FA97F9B5" box="[1212,1318,1604,1629]" class="Diplopoda" family="Anthroleucosomatidae" genus="Leschius" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Leschius</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
with
|
||
<taxonomicName id="4C4DFAD6FFE2FFCDFEF4E83AFE00F96D" box="[330,433,1644,1669]" class="Diplopoda" family="Anthroleucosomatidae" genus="Leschius" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9395D47FFE2FFCDFEF4E83AFE00F96D" box="[330,433,1644,1669]" italics="true" pageId="3" pageNumber="4">Leschius</emphasis>
|
||
</taxonomicName>
|
||
having much more complex posterior gonopod coxites than any other anthroleucosomatids. Because of the presence of these coxites, it seems appropriate to postulate that the ninth legs have a role to play in spermatophore transfer, and thus accurately can be referred to as “posterior gonopods.”
|
||
</paragraph>
|
||
<caption id="DF32D1DDFFE5FFCAFEB6E895FDAFF8AA" httpUri="https://zenodo.org/record/158750/files/figure.png" pageId="4" pageNumber="5" targetBox="[274,1307,319,1720]" targetPageId="4">
|
||
<paragraph id="8BF28155FFE5FFCAFEB6E895FDAFF8AA" blockId="4.[264,1323,1731,1858]" pageId="4" pageNumber="5">
|
||
<emphasis id="B9395D47FFE5FFCAFEB6E895FE04F933" bold="true" box="[264,437,1731,1755]" pageId="4" pageNumber="5">FIGURES 1–6.</emphasis>
|
||
<emphasis id="B9395D47FFE5FFCAFE7BE895FD12F932" box="[453,675,1731,1754]" italics="true" pageId="4" pageNumber="5">
|
||
<taxonomicName id="4C4DFAD6FFE5FFCAFE7BE895FD2EF932" box="[453,671,1731,1754]" class="Diplopoda" family="Anthroleucosomatidae" genus="Leschius" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="mcallisteri" status="sp. nov.">Leschius mcallisteri</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<taxonomicNameLabel id="A20AE03CFFE5FFCAFD15E895FD57F932" box="[683,742,1731,1754]" pageId="4" pageNumber="5" rank="species">n. sp.</taxonomicNameLabel>
|
||
1. Anterior gonopods, anterior view; 2. The same, posterior view; 3. Posterior gonopod (legpair 9), anterior view.; 4. The same, posterior view; 5. Legpair 10 of male, anterior view; 6. Cyphopods, posterior view. Scale line for figures 1–5 = 0.1 mm, for figure 6, 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF28155FFE4FFCBFE86EF4AFC35FE3D" blockId="5.[264,1323,284,1030]" pageId="5" pageNumber="6">
|
||
The geographically nearest occurring anthroleucosomatid species to
|
||
<taxonomicName id="4C4DFAD6FFE4FFCBFBDCEF4AFB4FFEDD" box="[1122,1278,284,309]" class="Diplopoda" family="Anthroleucosomatidae" genus="Leschius" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="mcallisteri">
|
||
<emphasis id="B9395D47FFE4FFCBFBDCEF4AFB4FFEDD" box="[1122,1278,284,309]" italics="true" pageId="5" pageNumber="6">L. mcallisteri</emphasis>
|
||
</taxonomicName>
|
||
are
|
||
<taxonomicName id="4C4DFAD6FFE4FFCBFEB6EF12FDF4FEB6" authority="Gulicka" authorityName="Gulicka" box="[264,581,324,350]" class="Diplopoda" family="Anthroleucosomatidae" genus="Ghilarovia" higherTaxonomySource="CoL" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="kygae">
|
||
<emphasis id="B9395D47FFE4FFCBFEB6EF12FE68FEB5" box="[264,473,324,349]" italics="true" pageId="5" pageNumber="6">Ghilarovia kygae</emphasis>
|
||
Gulicka
|
||
</taxonomicName>
|
||
(Altai Mountains) and
|
||
<taxonomicName id="4C4DFAD6FFE4FFCBFCCCEF12FBCDFEB6" authority="Shear" authorityName="Shear" box="[882,1148,324,350]" class="Diplopoda" family="Anthroleucosomatidae" genus="Ghilarovia" higherTaxonomySource="CoL" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="novosibirica">
|
||
<emphasis id="B9395D47FFE4FFCBFCCCEF12FB9BFEB5" box="[882,1066,324,349]" italics="true" pageId="5" pageNumber="6">G. novosibirica</emphasis>
|
||
Shear
|
||
</taxonomicName>
|
||
(Novosibirsk;
|
||
<bibRefCitation id="EFDCFCA4FFE4FFCBFEB6EF3AFE27FE6E" author="Shear" box="[264,406,364,390]" pageId="5" pageNumber="7" refString="Shear, W. A. (1988) Systematic position of the milliped species Alloiopus solitarius Attems and the genus Ghilarovia Gulicka (Chordeumatida, Anthroleucosomatidae). Myriapodologica, 2, 51 - 58." type="journal article" year="1988">Shear 1988</bibRefCitation>
|
||
). Both species have complex coxites in the posterior gonopods, but with prominent, 2articled telopodites as well. A variety of pregonopodal leg modifications occur in all the mentioned genera, but not in
|
||
<emphasis id="B9395D47FFE4FFCBFCA8EFEAFC35FE3D" box="[790,900,444,469]" italics="true" pageId="5" pageNumber="6">
|
||
<taxonomicName id="4C4DFAD6FFE4FFCBFCA8EFEAFC30FE3D" box="[790,897,444,469]" class="Diplopoda" family="Anthroleucosomatidae" genus="Leschius" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Leschius</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF28155FFE4FFCBFE86EFB2FB38FD06" blockId="5.[264,1323,284,1030]" pageId="5" pageNumber="6">
|
||
An intriguing possible connection exists between the aforementioned genera with glands opening on the sternal process,
|
||
<emphasis id="B9395D47FFE4FFCBFD71EC5AFC8DFDCD" box="[719,828,524,549]" italics="true" pageId="5" pageNumber="6">
|
||
<taxonomicName id="4C4DFAD6FFE4FFCBFD71EC5AFC89FDCD" box="[719,824,524,549]" class="Diplopoda" family="Anthroleucosomatidae" genus="Leschius" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Leschius</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and the enigmatic
|
||
<taxonomicName id="4C4DFAD6FFE4FFCBFBA3EC5AFABAFDCE" box="[1053,1291,524,550]" class="Diplopoda" family="Speophilosomatidae" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="family">Speophilosomatidae</taxonomicName>
|
||
of
|
||
<collectingCountry id="F35AC1C5FFE4FFCBFEB6EC62FEFEFDA6" box="[264,335,564,590]" name="Japan" pageId="5" pageNumber="6">Japan</collectingCountry>
|
||
. Speophilosomatids have simple anterior gonopods, with separate free coxites on either side and a large, median sternal process bearing apical gland pores (
|
||
<bibRefCitation id="EFDCFCA4FFE4FFCBFBD6EC0AFE69FD76" author="Shear" pageId="5" pageNumber="7" refString="Shear, W. A., Tsurusaki, N. & Tanabe, T. (1994) Japanese chordeumatid millipeds. I. On the genus Speophilosoma Takukawa (Diplopoda, Chordeumatida, Speophilosomatidae). Myriapodologica, 3, 25 - 36." type="journal article" year="1994">Shear, Tsurusaki and Tanabe 1994</bibRefCitation>
|
||
). If these features are homologous, it may be that
|
||
<taxonomicName id="4C4DFAD6FFE4FFCBFB83ECD2FA9AFD76" box="[1085,1323,644,670]" class="Diplopoda" family="Speophilosomatidae" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="family">Speophilosomatidae</taxonomicName>
|
||
belongs in Anthroleucosomatoidea. However, speophilosomatids have uniquely modified seventh legpairs that suggest an alliance with
|
||
<taxonomicName id="4C4DFAD6FFE4FFCBFC9EEC82FB34FD06" authority="Shear 2000" authorityName="Shear" authorityYear="2000" box="[800,1157,724,750]" class="Diplopoda" family="Chordeumatidae" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="family">Chordeumatidae (Shear 2000)</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF28155FFE4FFCBFE86ECAAFAAFFBEE" blockId="5.[264,1323,284,1030]" pageId="5" pageNumber="6">
|
||
Thus while we are confident that
|
||
<taxonomicName id="4C4DFAD6FFE4FFCBFD01ECAAFC97FCFD" box="[703,806,764,789]" class="Diplopoda" family="Anthroleucosomatidae" genus="Leschius" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9395D47FFE4FFCBFD01ECAAFC97FCFD" box="[703,806,764,789]" italics="true" pageId="5" pageNumber="6">Leschius</emphasis>
|
||
</taxonomicName>
|
||
is best placed in the Family
|
||
<taxonomicName id="4C4DFAD6FFE4FFCBFBCAECAAFED9FCD6" class="Diplopoda" family="Anthroleucosomatidae" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="family">Anthroleucosomatidae</taxonomicName>
|
||
, it is disjunct both geographically and morphologically from the rest of the family. A complete restudy and reclassification of the present
|
||
<taxonomicName id="4C4DFAD6FFE4FFCBFBA4ED1AFA96FC8E" box="[1050,1319,844,870]" class="Diplopoda" family="Anthroleucosomatidae" kingdom="Animalia" order="Chordeumatida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="family">Anthroleucosomatidae</taxonomicName>
|
||
, restricted as it now is (Shear 2000) from Hoffman’s much expanded version (
|
||
<bibRefCitation id="EFDCFCA4FFE4FFCBFB06ED22FEF5FC5E" author="Hoffman" pageId="5" pageNumber="6" refString="Hoffman, R. L. (1980) Classification of the Diplopoda. Museum d'Histoire Naturelle, Geneve, 237 pp." type="book" year="1980">Hoffman, 1980</bibRefCitation>
|
||
) will, we think, result in at least four new familylevel taxa. But with this whole group of millipeds so imperfectly known, we are presently content just to include our new genus and species, and leave the settlement of its exact taxonomic position for the future.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |