249 lines
35 KiB
XML
249 lines
35 KiB
XML
<document id="C2B67F785CE17023A9AED72D6D5AA303" ID-ISSN="11755334" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625278689308" checkinUser="felipe" docAuthor="Paulsen, M. J. & Mondaca, José" docDate="2006" docId="03D03943FFF5FFB1FEF4BCC6FB80FDF9" docLanguage="en" docName="zt01191p019.pdf" docOrigin="Zootaxa 1191 (1191)" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Hilophyllus martinezi Paulsen & Mondaca 2006, new species" docType="treatment" docUuid="ED0447EE-E91B-4D31-A999-105272C49723" docUuidSource="ZooBank" docVersion="4" lastPageNumber="18" masterDocId="FFE9413BFFFAFFA0FFFCB923FF85FFDF" masterDocTitle="Revision of the South American Ceratognathini (Coleoptera: Lucanidae: Aesalinae) with the description of a new genus and a new species" masterLastPageNumber="19" masterPageNumber="1" pageNumber="16" updateTime="1698985734564" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="F46A43ED6D7D4EE40551EDA7A4ECAB22">
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<mods:title id="74CD1D2707574A1CBDE945C65F6AA2C4">Revision of the South American Ceratognathini (Coleoptera: Lucanidae: Aesalinae) with the description of a new genus and a new species</mods:title>
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<mods:name id="B78F8DE3C8066D674750D33626F67954" type="personal">
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<mods:role id="3115069B49D9C7FC600D45DD19A8DD6C">
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<mods:roleTerm id="1E5BAD462CE9FCA3D70F8EF711B49844">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C7CC78EE812D8DAE40A43EB42AAA7DBF">Paulsen, M. J.</mods:namePart>
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<mods:affiliation id="F83A8739570F7BDA838A3C8991F5553D">Division of Entomology, University of Nebraska State Museum, W 436 Nebraska Hall, Lincoln, NE, USA,</mods:affiliation>
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<mods:name id="12D09FE8275B60F14BA4CC5247235F31" type="personal">
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<mods:roleTerm id="B0DE9107F1844F471C50C4E34EE66A57">Author</mods:roleTerm>
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<mods:namePart id="DEF9CF5F08200CE210866C3BB167286C">Mondaca, José</mods:namePart>
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<mods:typeOfResource id="E74658495924E64CE5C08C33F588B5D9">text</mods:typeOfResource>
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<mods:titleInfo id="94D0101F3BAC65C8CBDE808E128DF95F">
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<mods:title id="10EEA14C5154624862EB5D7C24EF7A9C">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="EAD9157ECD68E48DB1D2A207A638DB86">
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<mods:date id="B8E8D05DC585E1F85A21F96E089599A1">2006</mods:date>
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<mods:number id="CFD5B38362407B24379A470D0C1E6847">2006-05-01</mods:number>
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<mods:detail id="9CC38EBCA2D0F6DD3AEA61D8B2E919BC" type="volume">
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<mods:number id="510946ABDEC3402DF56F7E5DB0A1DFD7">1191</mods:number>
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<mods:detail id="35007BCE9E42B19D32E19018C738998E" type="issue">
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<mods:number id="A7CFB31A4FC89BAF141D8460D5EC9591">1191</mods:number>
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<mods:classification id="CA2867B935002614440EB7BE5534823E">journal article</mods:classification>
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<mods:identifier id="D1D80A10C1A21F174984A25C8221CED3" type="ISSN">11755334</mods:identifier>
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<mods:identifier id="F9B93020700E6DC76FB7C6172204612F" type="ZooBank">829B214A-C06F-42CB-B1A8-D729C631DF55</mods:identifier>
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<treatment id="03D03943FFF5FFB1FEF4BCC6FB80FDF9" ID-DOI="http://doi.org/10.5281/zenodo.5064404" ID-Zenodo-Dep="5064404" LSID="urn:lsid:zoobank.org:act:ED0447EE-E91B-4D31-A999-105272C49723" httpUri="http://treatment.plazi.org/id/03D03943FFF5FFB1FEF4BCC6FB80FDF9" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16">
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<subSubSection id="C363DBDEFFF5FFAFFEF4BCC6FC36FA21" box="[264,947,1508,1535]" pageId="15" pageNumber="16" type="nomenclature">
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<paragraph id="8BC68855FFF5FFAFFEF4BCC6FC36FA21" blockId="15.[264,947,1508,1569]" box="[264,947,1508,1535]" pageId="15" pageNumber="16">
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<heading id="D08E3F39FFF5FFAFFEF4BCC6FC36FA21" bold="true" box="[264,947,1508,1535]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
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<emphasis id="B90D5447FFF5FFAFFEF4BCC6FC36FA21" bold="true" box="[264,947,1508,1535]" pageId="15" pageNumber="16">
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<taxonomicName id="4C79F3D6FFF5FFAFFEF4BCC6FC93FA21" ID-CoL="3LXHZ" authority="Paulsen & Mondaca" authorityName="Paulsen & Mondaca" authorityYear="2006" box="[264,790,1508,1535]" class="Insecta" family="Lucanidae" genus="Hilophyllus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="martinezi" status="sp. nov.">
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<emphasis id="B90D5447FFF5FFAFFEF4BCC6FD8AFA20" bold="true" box="[264,527,1509,1535]" italics="true" pageId="15" pageNumber="16">Hilophyllus martinezi</emphasis>
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Paulsen & Mondaca
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</taxonomicName>
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,
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<taxonomicNameLabel id="A23EE93CFFF5FFAFFCDFBCC7FC36FA21" box="[803,947,1508,1534]" pageId="15" pageNumber="16" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C363DBDEFFF5FFB0FEF3BF24FD83F9D1" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="description">
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<paragraph id="8BC68855FFF5FFAFFEF3BF24FDCBF9FE" blockId="15.[264,947,1508,1569]" box="[271,590,1543,1569]" pageId="15" pageNumber="16">
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(
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<figureCitation id="134294D0FFF5FFAFFEE4BF24FEEFF9FE" box="[280,362,1543,1569]" captionStart="FIGURES 1–6" captionStartId="8.[264,382,1611,1635]" captionTargetBox="[272,1285,617,1590]" captionTargetId="figure-176@8.[264,1323,609,1604]" captionTargetPageId="8" captionText="FIGURES 1–6. 1, dorsal habitus of male Hilophyllus penai. 2, dorsal habitus of male H. argentinensis. 3, dorsal habitus of male holotype H. martinezi, n.sp. 4, dorsal view of head of female H. penai showing smaller antennal club, eyes, and mandibles. 5, dorsal view of head of male H. penai showing eyes and mandibles. 6, dorsal view of head of male H. argentinensis showing large eyes and bent, interlocking mandibles. Scale bar = 2 mm (Figs. 1–3) or 1 mm (Figs. 4–6)." pageId="15" pageNumber="16">Figs. 3</figureCitation>
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,
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<figureCitation id="134294D0FFF5FFAFFE8BBF24FE47F9FE" box="[375,450,1543,1569]" captionStart="FIGURES 13–18" captionStartId="13.[264,383,1475,1499]" captionTargetBox="[264,1323,567,1451]" captionTargetId="figure-216@13.[264,1323,555,1451]" captionTargetPageId="13" captionText="FIGURES 13–18. 13, detail of elytral surface of H. argentinensis showing scale, puncture, and smooth surface structure. Scale bar = 100 m. 14, detail of elytral surface of H. martinezi showing scale, puncture, and minutelywrinkled surface structure. Scale bar = 100 m. 15, right antenna of male H. argentinensis. Scale bar = 200 m. 16, right antenna of male H. martinezi. Scale bar = 200 m. 17, scape of right antenna of male H. martinezi. Scale bar = 50 m. 18, external surface of apex of right paramere of male H. martinezi showing two minute teeth. Scale bar = 50 m." pageId="15" pageNumber="16">14, 18</figureCitation>
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,
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<figureCitation id="134294D0FFF5FFAFFE33BF24FDC3F9FE" box="[463,582,1543,1569]" captionStart-0="FIGURES 19–24" captionStart-1="FIGURE 25" captionStartId-0="14.[264,383,1638,1662]" captionStartId-1="15.[264,368,1372,1396]" captionTargetBox-0="[300,1293,973,1617]" captionTargetBox-1="[296,1310,326,1355]" captionTargetId-0="figure-317@14.[300,1293,967,1617]" captionTargetId-1="figure-88@15.[264,1323,326,1367]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionText-0="FIGURES 19–24. 19, ventral view of male genitalia, H. penai. 20, ventral view of male genitalia, H. argentinensis. 21, ventral view of male genitalia, H. martinezi. 22, female genitalia of H. penai. Terminology as in Holloway (1998): BC = bursa copulatrix, H = hemisternite, H9 = ninth hemisternite, SD = spermathecal duct, SG? = spermathecal gland (not seen), SP = spermatheca, ST = stylus. 23, outline of anterior portion of pronotum of H. argentinensis, showing lack of produced anterior angles. 24, outline of anterior portion of pronotum of H. martinezi, showing produced anterior angles." captionText-1="FIGURE 25. Localities from which Hilophyllus specimens were examined: H. penai (asterisks), H. argentinensis (squares), and H. martinezi (triangle)." pageId="15" pageNumber="16">21, 24–25</figureCitation>
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)
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</paragraph>
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<paragraph id="8BC68855FFF5FFAFFEF4BF7BFE2CF9AE" blockId="15.[264,1322,1624,1769]" box="[264,425,1624,1649]" pageId="15" pageNumber="16">
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<emphasis id="B90D5447FFF5FFAFFEF4BF7BFE2CF9AE" box="[264,425,1624,1649]" italics="true" pageId="15" pageNumber="16">Type material</emphasis>
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</paragraph>
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<paragraph id="8BC68855FFF5FFAFFEC4BF5CFC76F936" blockId="15.[264,1322,1624,1769]" pageId="15" pageNumber="16">
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<materialsCitation id="3B118208FFF5FFAFFEC4BF5CFC76F936" collectingDate="1988-11-29" collectorName="S. Roitman" country="Chile" location="Los Lleuques" pageId="15" pageNumber="16" specimenCount="2" typeStatus="holotype">
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<typeStatus id="54C236F7FFF5FFAFFEC4BF5CFE20F946" box="[312,421,1663,1689]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
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male (MNNC) labeled: a) “
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<collectingCountry id="F36EC8C5FFF5FFAFFD09BF5CFCCBF946" box="[757,846,1663,1689]" name="Chile" pageId="15" pageNumber="16">CHILE</collectingCountry>
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ÑUBLE /
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<location id="8EA6DE8EFFF5FFAFFC28BF5CFBF1F946" LSID="urn:lsid:plazi:treatment:03D03943FFF5FFB1FEF4BCC6FB80FDF9:8EA6DE8EFFF5FFAFFC28BF5CFBF1F946" box="[980,1140,1663,1689]" country="Chile" name="Los Lleuques" pageId="15" pageNumber="16">Los Lleuques</location>
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/
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<date id="FFC7AE95FFF5FFAFFB73BF5CFA9DF946" box="[1167,1304,1663,1689]" pageId="15" pageNumber="16" value="1988-11-29">
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<collectingDate id="EF83577DFFF5FFAFFB73BF5CFA9DF946" box="[1167,1304,1663,1689]" pageId="15" pageNumber="16" value="1988-11-29">29XI1988</collectingDate>
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</date>
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/ Colección MNHN” [=MNNC,
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<collectorName id="268CED83FFF5FFAFFD74BF84FC97F91E" box="[648,786,1703,1729]" pageId="15" pageNumber="16">S. Roitman</collectorName>
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collector]; b) on red paper, “
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<emphasis id="B90D5447FFF5FFAFFB70BF8BFEF2F936" italics="true" pageId="15" pageNumber="16">
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<taxonomicName id="4C79F3D6FFF5FFAFFB70BF8BFA90F91E" authorityName="Paulsen and Mondaca" authorityYear="2006" box="[1164,1301,1704,1729]" class="Insecta" family="Lucanidae" genus="Hilophyllus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">Hilophyllus</taxonomicName>
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/
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<taxonomicName id="4C79F3D6FFF5FFAFFEF4BFF3FEF2F936" authorityName="Paulsen & Mondaca" authorityYear="2006" box="[264,375,1744,1769]" class="Insecta" family="Lucanidae" genus="Hilophyllus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="martinezi">martinezi</taxonomicName>
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</emphasis>
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(male symbol)
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<emphasis id="B90D5447FFF5FFAFFDCCBFF3FDBDF936" box="[560,568,1744,1769]" italics="true" pageId="15" pageNumber="16">/</emphasis>
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Paulsen & Mondaca /
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<typeStatus id="54C236F7FFF5FFAFFCBFBFECFC60F936" box="[835,997,1743,1769]" pageId="15" pageNumber="16" type="holotype">HOLOTYPE</typeStatus>
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”.
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</materialsCitation>
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</paragraph>
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<paragraph id="8BC68855FFEAFFB0FEF4B83FFE17FEEA" blockId="16.[264,1324,284,950]" box="[264,402,284,309]" pageId="16" pageNumber="17">
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<emphasis id="B90D5447FFEAFFB0FEF4B83FFE17FEEA" box="[264,402,284,309]" italics="true" pageId="16" pageNumber="17">Description</emphasis>
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</paragraph>
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<paragraph id="8BC68855FFEAFFB0FEC4B867FC6CFC51" blockId="16.[264,1324,284,950]" pageId="16" pageNumber="17">
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<typeStatus id="54C236F7FFEAFFB0FEC4B867FE20FE81" box="[312,421,324,350]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
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male.
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<emphasis id="B90D5447FFEAFFB0FDFAB867FDE3FE81" bold="true" box="[518,614,324,350]" italics="true" pageId="16" pageNumber="17">Length:</emphasis>
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8.0 mm.
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<emphasis id="B90D5447FFEAFFB0FD11B867FCBAFE82" box="[749,831,324,350]" italics="true" pageId="16" pageNumber="17">
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<emphasis id="B90D5447FFEAFFB0FD11B867FCB0FE81" bold="true" box="[749,821,324,350]" italics="true" pageId="16" pageNumber="17">Width</emphasis>
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:
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</emphasis>
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<quantity id="4C8125B0FFEAFFB0FCB6B867FC2EFE81" box="[842,939,324,350]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="16" pageNumber="17" unit="mm" value="2.9">2.9 mm</quantity>
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.
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<emphasis id="B90D5447FFEAFFB0FC39B867FBFCFE81" bold="true" box="[965,1145,324,350]" italics="true" pageId="16" pageNumber="17">Color/Surface:</emphasis>
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Dark reddishbrown, weakly shining. Dorsum (
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<figureCitation id="134294D0FFEAFFB0FD63B84FFD63FE59" box="[671,742,364,390]" captionStart="FIGURES 1–6" captionStartId="8.[264,382,1611,1635]" captionTargetBox="[272,1285,617,1590]" captionTargetId="figure-176@8.[264,1323,609,1604]" captionTargetPageId="8" captionText="FIGURES 1–6. 1, dorsal habitus of male Hilophyllus penai. 2, dorsal habitus of male H. argentinensis. 3, dorsal habitus of male holotype H. martinezi, n.sp. 4, dorsal view of head of female H. penai showing smaller antennal club, eyes, and mandibles. 5, dorsal view of head of male H. penai showing eyes and mandibles. 6, dorsal view of head of male H. argentinensis showing large eyes and bent, interlocking mandibles. Scale bar = 2 mm (Figs. 1–3) or 1 mm (Figs. 4–6)." pageId="16" pageNumber="17">Fig. 3</figureCitation>
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) almost uniformly covered with large punctures, most with a single elongate, apically acute scale (
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<figureCitation id="134294D0FFEAFFB0FC9FB8B7FC3BFE71" box="[867,958,404,430]" captionStart="FIGURES 13–18" captionStartId="13.[264,383,1475,1499]" captionTargetBox="[264,1323,567,1451]" captionTargetId="figure-216@13.[264,1323,555,1451]" captionTargetPageId="13" captionText="FIGURES 13–18. 13, detail of elytral surface of H. argentinensis showing scale, puncture, and smooth surface structure. Scale bar = 100 m. 14, detail of elytral surface of H. martinezi showing scale, puncture, and minutelywrinkled surface structure. Scale bar = 100 m. 15, right antenna of male H. argentinensis. Scale bar = 200 m. 16, right antenna of male H. martinezi. Scale bar = 200 m. 17, scape of right antenna of male H. martinezi. Scale bar = 50 m. 18, external surface of apex of right paramere of male H. martinezi showing two minute teeth. Scale bar = 50 m." pageId="16" pageNumber="17">Fig. 14</figureCitation>
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).
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<emphasis id="B90D5447FFEAFFB0FC1DB8B7FBAEFE71" bold="true" box="[993,1067,404,430]" italics="true" pageId="16" pageNumber="17">Head:</emphasis>
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Eyes large, width of eyes together in dorsal view greater than narrowest interocular width. Mandibles small, externally rounded, not abruptly curved inward at middle; dorsal tooth acute; ventral tooth of right mandible minute (apex of right mandible broken). Antennal club segments elongate, longer than 1.5 times length of head (
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<figureCitation id="134294D0FFEAFFB0FC98BB17FC47FD91" box="[868,962,564,590]" captionStart="FIGURES 13–18" captionStartId="13.[264,383,1475,1499]" captionTargetBox="[264,1323,567,1451]" captionTargetId="figure-216@13.[264,1323,555,1451]" captionTargetPageId="13" captionText="FIGURES 13–18. 13, detail of elytral surface of H. argentinensis showing scale, puncture, and smooth surface structure. Scale bar = 100 m. 14, detail of elytral surface of H. martinezi showing scale, puncture, and minutelywrinkled surface structure. Scale bar = 100 m. 15, right antenna of male H. argentinensis. Scale bar = 200 m. 16, right antenna of male H. martinezi. Scale bar = 200 m. 17, scape of right antenna of male H. martinezi. Scale bar = 50 m. 18, external surface of apex of right paramere of male H. martinezi showing two minute teeth. Scale bar = 50 m." pageId="16" pageNumber="17">Fig. 16</figureCitation>
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).
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<emphasis id="B90D5447FFEAFFB0FC10BB17FBEAFD91" bold="true" box="[1004,1135,564,590]" italics="true" pageId="16" pageNumber="17">Pronotum:</emphasis>
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Sides strongly explanate, explanate margin subequal to eye in width. Anterior angle produced beyond anterior margin (
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<figureCitation id="134294D0FFEAFFB0FE31BBA7FDA0FD41" box="[461,549,644,670]" captionStart="FIGURES 19–24" captionStartId="14.[264,383,1638,1662]" captionTargetBox="[300,1293,973,1617]" captionTargetId="figure-317@14.[300,1293,967,1617]" captionTargetPageId="14" captionText="FIGURES 19–24. 19, ventral view of male genitalia, H. penai. 20, ventral view of male genitalia, H. argentinensis. 21, ventral view of male genitalia, H. martinezi. 22, female genitalia of H. penai. Terminology as in Holloway (1998): BC = bursa copulatrix, H = hemisternite, H9 = ninth hemisternite, SD = spermathecal duct, SG? = spermathecal gland (not seen), SP = spermatheca, ST = stylus. 23, outline of anterior portion of pronotum of H. argentinensis, showing lack of produced anterior angles. 24, outline of anterior portion of pronotum of H. martinezi, showing produced anterior angles." pageId="16" pageNumber="17">Fig. 24</figureCitation>
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). Lateral margin broadly rounded, widest before middle.
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<emphasis id="B90D5447FFEAFFB0FB2ABBA7FAAFFD41" bold="true" box="[1238,1322,644,670]" italics="true" pageId="16" pageNumber="17">Elytra:</emphasis>
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Shape elongate. Almost uniformly punctate, most punctures with scales, punctures without scales forming longitudinal rows (not patches), scales brown and testaceous. Sutural interval and lateral margin elevated apically. Surface between punctures roughened (
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<figureCitation id="134294D0FFEAFFB0FE68BA07FE6EFCE1" box="[404,491,804,830]" captionStart="FIGURES 13–18" captionStartId="13.[264,383,1475,1499]" captionTargetBox="[264,1323,567,1451]" captionTargetId="figure-216@13.[264,1323,555,1451]" captionTargetPageId="13" captionText="FIGURES 13–18. 13, detail of elytral surface of H. argentinensis showing scale, puncture, and smooth surface structure. Scale bar = 100 m. 14, detail of elytral surface of H. martinezi showing scale, puncture, and minutelywrinkled surface structure. Scale bar = 100 m. 15, right antenna of male H. argentinensis. Scale bar = 200 m. 16, right antenna of male H. martinezi. Scale bar = 200 m. 17, scape of right antenna of male H. martinezi. Scale bar = 50 m. 18, external surface of apex of right paramere of male H. martinezi showing two minute teeth. Scale bar = 50 m." pageId="16" pageNumber="17">Fig. 14</figureCitation>
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), appearing wrinkled at 40x.
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<emphasis id="B90D5447FFEAFFB0FCB2BA07FC14FCE1" bold="true" box="[846,913,804,830]" italics="true" pageId="16" pageNumber="17">Legs:</emphasis>
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Mesotibiae lacking external tooth.
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<emphasis id="B90D5447FFEAFFB0FEF4BA6FFE41FCB9" bold="true" box="[264,452,844,870]" italics="true" pageId="16" pageNumber="17">Male genitalia:</emphasis>
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See
|
||
<figureCitation id="134294D0FFEAFFB0FDF9BA6FFDDAFCB9" box="[517,607,844,870]" captionStart="FIGURES 19–24" captionStartId="14.[264,383,1638,1662]" captionTargetBox="[300,1293,973,1617]" captionTargetId="figure-317@14.[300,1293,967,1617]" captionTargetPageId="14" captionText="FIGURES 19–24. 19, ventral view of male genitalia, H. penai. 20, ventral view of male genitalia, H. argentinensis. 21, ventral view of male genitalia, H. martinezi. 22, female genitalia of H. penai. Terminology as in Holloway (1998): BC = bursa copulatrix, H = hemisternite, H9 = ninth hemisternite, SD = spermathecal duct, SG? = spermathecal gland (not seen), SP = spermatheca, ST = stylus. 23, outline of anterior portion of pronotum of H. argentinensis, showing lack of produced anterior angles. 24, outline of anterior portion of pronotum of H. martinezi, showing produced anterior angles." pageId="16" pageNumber="17">Fig. 21</figureCitation>
|
||
. Parameres with 2 small, external teeth at apex (
|
||
<figureCitation id="134294D0FFEAFFB0FB3EBA6FFA99FCB9" box="[1218,1308,844,870]" captionStart="FIGURES 13–18" captionStartId="13.[264,383,1475,1499]" captionTargetBox="[264,1323,567,1451]" captionTargetId="figure-216@13.[264,1323,555,1451]" captionTargetPageId="13" captionText="FIGURES 13–18. 13, detail of elytral surface of H. argentinensis showing scale, puncture, and smooth surface structure. Scale bar = 100 m. 14, detail of elytral surface of H. martinezi showing scale, puncture, and minutelywrinkled surface structure. Scale bar = 100 m. 15, right antenna of male H. argentinensis. Scale bar = 200 m. 16, right antenna of male H. martinezi. Scale bar = 200 m. 17, scape of right antenna of male H. martinezi. Scale bar = 50 m. 18, external surface of apex of right paramere of male H. martinezi showing two minute teeth. Scale bar = 50 m." pageId="16" pageNumber="17">Fig. 18</figureCitation>
|
||
), median lobe cylindrical toward apex, not laterally compressed.
|
||
</paragraph>
|
||
<paragraph id="8BC68855FFEAFFB0FEC4BABFFD8FFC69" blockId="16.[264,1324,284,950]" box="[312,522,924,950]" pageId="16" pageNumber="17">Female unknown.</paragraph>
|
||
<paragraph id="8BC68855FFEAFFB0FEF4BACFFEFAFBDA" blockId="16.[264,1323,1004,1470]" box="[264,383,1004,1029]" pageId="16" pageNumber="17">
|
||
<emphasis id="B90D5447FFEAFFB0FEF4BACFFEFAFBDA" box="[264,383,1004,1029]" italics="true" pageId="16" pageNumber="17">Diagnosis</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC68855FFEAFFB0FEC4BD37FC67FA61" blockId="16.[264,1323,1004,1470]" pageId="16" pageNumber="17">
|
||
This species is distinguished from
|
||
<taxonomicName id="4C79F3D6FFEAFFB0FD31BD37FCB7FBF2" authorityName="Paulsen & Mondaca" authorityYear="2006" baseAuthorityName="Paulsen & Mondaca" baseAuthorityYear="2006" box="[717,818,1044,1069]" class="Insecta" family="Lucanidae" genus="Hilophyllus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="penai">
|
||
<emphasis id="B90D5447FFEAFFB0FD31BD37FCB7FBF2" box="[717,818,1044,1069]" italics="true" pageId="16" pageNumber="17">H. penai</emphasis>
|
||
</taxonomicName>
|
||
by its unarmed mesotibiae. It is, however, externally quite similar to
|
||
<emphasis id="B90D5447FFEAFFB0FDC1BD1FFC80FB8A" box="[573,773,1084,1109]" italics="true" pageId="16" pageNumber="17">
|
||
<taxonomicName id="4C79F3D6FFEAFFB0FDC1BD1FFC84FB8A" box="[573,769,1084,1109]" class="Insecta" family="Lucanidae" genus="Ceratognathus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="argentinensis">H. argentinensis</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The most notable external difference between these two species is in the anterior angles of the pronotum, which are produced beyond the anterior pronotal margin in
|
||
<taxonomicName id="4C79F3D6FFEAFFB0FDA1BDAFFD76FB7A" authorityName="Paulsen & Mondaca" authorityYear="2006" box="[605,755,1164,1189]" class="Insecta" family="Lucanidae" genus="Hilophyllus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="martinezi">
|
||
<emphasis id="B90D5447FFEAFFB0FDA1BDAFFD76FB7A" box="[605,755,1164,1189]" italics="true" pageId="16" pageNumber="17">H. martinezi</emphasis>
|
||
</taxonomicName>
|
||
but more or less contiguous with the anterior margin of the pronotum in
|
||
<taxonomicName id="4C79F3D6FFEAFFB0FDBFBD97FC86FB12" box="[579,771,1204,1229]" class="Insecta" family="Lucanidae" genus="Ceratognathus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="argentinensis">
|
||
<emphasis id="B90D5447FFEAFFB0FDBFBD97FC86FB12" box="[579,771,1204,1229]" italics="true" pageId="16" pageNumber="17">H. argentinensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="134294D0FFEAFFB0FCE8BD97FCEDFB11" box="[788,872,1204,1230]" captionStart="FIGURES 19–24" captionStartId="14.[264,383,1638,1662]" captionTargetBox="[300,1293,973,1617]" captionTargetId="figure-317@14.[300,1293,967,1617]" captionTargetPageId="14" captionText="FIGURES 19–24. 19, ventral view of male genitalia, H. penai. 20, ventral view of male genitalia, H. argentinensis. 21, ventral view of male genitalia, H. martinezi. 22, female genitalia of H. penai. Terminology as in Holloway (1998): BC = bursa copulatrix, H = hemisternite, H9 = ninth hemisternite, SD = spermathecal duct, SG? = spermathecal gland (not seen), SP = spermatheca, ST = stylus. 23, outline of anterior portion of pronotum of H. argentinensis, showing lack of produced anterior angles. 24, outline of anterior portion of pronotum of H. martinezi, showing produced anterior angles." pageId="16" pageNumber="17">Fig. 23</figureCitation>
|
||
). This gives the overall impression of a smaller pronotum in
|
||
<emphasis id="B90D5447FFEAFFB0FDEDBDFFFD2EFB2A" box="[529,683,1244,1269]" italics="true" pageId="16" pageNumber="17">
|
||
<taxonomicName id="4C79F3D6FFEAFFB0FDEDBDFFFD22FB2A" authorityName="Paulsen & Mondaca" authorityYear="2006" box="[529,679,1244,1269]" class="Insecta" family="Lucanidae" genus="Hilophyllus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="martinezi">H. martinezi</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Ideally we would prefer more material upon which to base a new species. However, the form of the male genitalia (
|
||
<figureCitation id="134294D0FFEAFFB0FBC6BC27FB4AFAC1" box="[1082,1231,1284,1310]" captionStart="FIGURES 19–24" captionStartId="14.[264,383,1638,1662]" captionTargetBox="[300,1293,973,1617]" captionTargetId="figure-317@14.[300,1293,967,1617]" captionTargetPageId="14" captionText="FIGURES 19–24. 19, ventral view of male genitalia, H. penai. 20, ventral view of male genitalia, H. argentinensis. 21, ventral view of male genitalia, H. martinezi. 22, female genitalia of H. penai. Terminology as in Holloway (1998): BC = bursa copulatrix, H = hemisternite, H9 = ninth hemisternite, SD = spermathecal duct, SG? = spermathecal gland (not seen), SP = spermatheca, ST = stylus. 23, outline of anterior portion of pronotum of H. argentinensis, showing lack of produced anterior angles. 24, outline of anterior portion of pronotum of H. martinezi, showing produced anterior angles." pageId="16" pageNumber="17">Figs. 20–21</figureCitation>
|
||
) in
|
||
<taxonomicName id="4C79F3D6FFEAFFB0FAF2BC27FE20FA9A" class="Insecta" family="Lucanidae" genus="Ceratognathus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="argentinensis">
|
||
<emphasis id="B90D5447FFEAFFB0FAF2BC27FE20FA9A" italics="true" pageId="16" pageNumber="17">H. argentinensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C79F3D6FFEAFFB0FE16BC0FFD07FA9A" authorityName="Paulsen & Mondaca" authorityYear="2006" box="[490,642,1324,1349]" class="Insecta" family="Lucanidae" genus="Hilophyllus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="martinezi">
|
||
<emphasis id="B90D5447FFEAFFB0FE16BC0FFD07FA9A" box="[490,642,1324,1349]" italics="true" pageId="16" pageNumber="17">H. martinezi</emphasis>
|
||
</taxonomicName>
|
||
are vastly different. Based on the constancy of male genitalic form (
|
||
<emphasis id="B90D5447FFEAFFB0FE35BC77FE71FAB2" box="[457,500,1364,1389]" italics="true" pageId="16" pageNumber="17">i.e.,</emphasis>
|
||
lack of variation) observed in
|
||
<taxonomicName id="4C79F3D6FFEAFFB0FC80BC77FC63FAB2" authorityName="Paulsen & Mondaca" authorityYear="2006" baseAuthorityName="Paulsen & Mondaca" baseAuthorityYear="2006" box="[892,998,1364,1389]" class="Insecta" family="Lucanidae" genus="Hilophyllus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="penai">
|
||
<emphasis id="B90D5447FFEAFFB0FC80BC77FC63FAB2" box="[892,998,1364,1389]" italics="true" pageId="16" pageNumber="17">H. penai</emphasis>
|
||
</taxonomicName>
|
||
(n = 4), as well as in
|
||
<taxonomicName id="4C79F3D6FFEAFFB0FAF2BC77FE20FA4A" class="Insecta" family="Lucanidae" genus="Ceratognathus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="argentinensis">
|
||
<emphasis id="B90D5447FFEAFFB0FAF2BC77FE20FA4A" italics="true" pageId="16" pageNumber="17">H. argentinensis</emphasis>
|
||
</taxonomicName>
|
||
(n = 9) across its range, we believe that the unique male genitalic form of
|
||
<taxonomicName id="4C79F3D6FFEAFFB0FAF2BC5FFEF2FA62" authorityName="Paulsen & Mondaca" authorityYear="2006" class="Insecta" family="Lucanidae" genus="Hilophyllus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="martinezi">
|
||
<emphasis id="B90D5447FFEAFFB0FAF2BC5FFEF2FA62" italics="true" pageId="16" pageNumber="17">H. martinezi</emphasis>
|
||
</taxonomicName>
|
||
is compelling evidence that this is a distinct species.
|
||
</paragraph>
|
||
<paragraph id="8BC68855FFEAFFB0FEF4BCD7FD83F9D1" blockId="16.[264,901,1524,1710]" box="[264,518,1524,1550]" pageId="16" pageNumber="17">
|
||
<emphasis id="B90D5447FFEAFFB0FEF4BCD7FE12F9D2" box="[264,407,1524,1549]" italics="true" pageId="16" pageNumber="17">Distribution</emphasis>
|
||
(
|
||
<figureCitation id="134294D0FFEAFFB0FE5BBCD7FE79F9D1" box="[423,508,1524,1550]" captionStart="FIGURE 25" captionStartId="15.[264,368,1372,1396]" captionTargetBox="[296,1310,326,1355]" captionTargetId="figure-88@15.[264,1323,326,1367]" captionTargetPageId="15" captionText="FIGURE 25. Localities from which Hilophyllus specimens were examined: H. penai (asterisks), H. argentinensis (squares), and H. martinezi (triangle)." pageId="16" pageNumber="17">Fig. 25</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C363DBDEFFEAFFB1FEC4BF3FFB80FDF9" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="materials_examined">
|
||
<paragraph id="8BC68855FFEAFFB0FEC4BF3FFC00F9E9" blockId="16.[264,901,1524,1710]" box="[312,901,1564,1590]" pageId="16" pageNumber="17">
|
||
<collectingCountry id="F36EC8C5FFEAFFB0FEC4BF3FFE17F9E9" box="[312,402,1564,1590]" name="Chile" pageId="16" pageNumber="17">CHILE</collectingCountry>
|
||
. The species is known from central Chile.
|
||
</paragraph>
|
||
<paragraph id="8BC68855FFEAFFB0FEC4BF67FD62F981" blockId="16.[264,901,1524,1710]" box="[312,743,1604,1630]" pageId="16" pageNumber="17">
|
||
Locality data:
|
||
<specimenCount id="9D7F43DCFFEAFFB0FE1DBF67FDE3F981" box="[481,614,1604,1630]" pageId="16" pageNumber="17" type="generic">1 specimen</specimenCount>
|
||
examined.
|
||
</paragraph>
|
||
<paragraph id="8BC68855FFEAFFB0FEC4BF4FFCBEF959" blockId="16.[264,901,1524,1710]" box="[312,827,1644,1670]" pageId="16" pageNumber="17">
|
||
<collectingCountry id="F36EC8C5FFEAFFB0FEC4BF4FFE0BF959" box="[312,398,1644,1670]" name="Chile" pageId="16" pageNumber="17">CHILE</collectingCountry>
|
||
(1). VIII Región (1): Los Lleuques.
|
||
</paragraph>
|
||
<paragraph id="8BC68855FFEAFFB0FEC4BFB7FD7DF971" blockId="16.[264,901,1524,1710]" box="[312,760,1684,1710]" pageId="16" pageNumber="17">Temporal distribution. November (1).</paragraph>
|
||
<paragraph id="8BC68855FFEAFFB0FEF4BFC7FE00F922" blockId="16.[264,1323,1764,1870]" box="[264,389,1764,1789]" pageId="16" pageNumber="17">
|
||
<emphasis id="B90D5447FFEAFFB0FEF4BFC7FE00F922" box="[264,389,1764,1789]" italics="true" pageId="16" pageNumber="17">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC68855FFEAFFB0FEC4BE2FFCAFF891" blockId="16.[264,1323,1764,1870]" pageId="16" pageNumber="17">
|
||
The species is named
|
||
<taxonomicName id="4C79F3D6FFEAFFB0FDC7BE2FFD2FF8FA" authorityName="Paulsen & Mondaca" authorityYear="2006" box="[571,682,1804,1829]" class="Insecta" family="Lucanidae" genus="Hilophyllus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="martinezi">
|
||
<emphasis id="B90D5447FFEAFFB0FDC7BE2FFD2FF8FA" box="[571,682,1804,1829]" italics="true" pageId="16" pageNumber="17">martinezi</emphasis>
|
||
</taxonomicName>
|
||
in honor of Antonio Martínez and his contributions to the knowledge of South American
|
||
<taxonomicName id="4C79F3D6FFEAFFB0FD5EBE17FCA3F891" authorityName="Linnaeus" authorityYear="1758" box="[674,806,1844,1870]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC68855FFEBFFB1FEF4B83FFEEBFEEA" blockId="17.[264,1323,284,550]" box="[264,366,284,309]" pageId="17" pageNumber="18">
|
||
<emphasis id="B90D5447FFEBFFB1FEF4B83FFEEBFEEA" box="[264,366,284,309]" italics="true" pageId="17" pageNumber="18">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC68855FFEBFFB1FEC4B867FB80FDF9" blockId="17.[264,1323,284,550]" pageId="17" pageNumber="18">
|
||
This single specimen from Los Lleuques, Ñuble, is similar externally to
|
||
<taxonomicName id="4C79F3D6FFEBFFB1FAF2B867FE20FE5A" class="Insecta" family="Lucanidae" genus="Ceratognathus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="argentinensis">
|
||
<emphasis id="B90D5447FFEBFFB1FAF2B867FE20FE5A" italics="true" pageId="17" pageNumber="18">H. argentinensis</emphasis>
|
||
</taxonomicName>
|
||
. However, the male genitalia (
|
||
<figureCitation id="134294D0FFEBFFB1FCCBB84FFC17FE59" box="[823,914,364,390]" captionStart="FIGURES 19–24" captionStartId="14.[264,383,1638,1662]" captionTargetBox="[300,1293,973,1617]" captionTargetId="figure-317@14.[300,1293,967,1617]" captionTargetPageId="14" captionText="FIGURES 19–24. 19, ventral view of male genitalia, H. penai. 20, ventral view of male genitalia, H. argentinensis. 21, ventral view of male genitalia, H. martinezi. 22, female genitalia of H. penai. Terminology as in Holloway (1998): BC = bursa copulatrix, H = hemisternite, H9 = ninth hemisternite, SD = spermathecal duct, SG? = spermathecal gland (not seen), SP = spermatheca, ST = stylus. 23, outline of anterior portion of pronotum of H. argentinensis, showing lack of produced anterior angles. 24, outline of anterior portion of pronotum of H. martinezi, showing produced anterior angles." pageId="17" pageNumber="18">Fig. 21</figureCitation>
|
||
) are distinct. The apex of each paramere bears two minute, external teeth (
|
||
<figureCitation id="134294D0FFEBFFB1FCE5B8B7FCF4FE71" box="[793,881,404,430]" captionStart="FIGURES 13–18" captionStartId="13.[264,383,1475,1499]" captionTargetBox="[264,1323,567,1451]" captionTargetId="figure-216@13.[264,1323,555,1451]" captionTargetPageId="13" captionText="FIGURES 13–18. 13, detail of elytral surface of H. argentinensis showing scale, puncture, and smooth surface structure. Scale bar = 100 m. 14, detail of elytral surface of H. martinezi showing scale, puncture, and minutelywrinkled surface structure. Scale bar = 100 m. 15, right antenna of male H. argentinensis. Scale bar = 200 m. 16, right antenna of male H. martinezi. Scale bar = 200 m. 17, scape of right antenna of male H. martinezi. Scale bar = 50 m. 18, external surface of apex of right paramere of male H. martinezi showing two minute teeth. Scale bar = 50 m." pageId="17" pageNumber="18">Fig. 18</figureCitation>
|
||
) rather than the large external tooth present on
|
||
<taxonomicName id="4C79F3D6FFEBFFB1FE75B89FFDCFFE0A" box="[393,586,444,469]" class="Insecta" family="Lucanidae" genus="Ceratognathus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="argentinensis">
|
||
<emphasis id="B90D5447FFEBFFB1FE75B89FFDCFFE0A" box="[393,586,444,469]" italics="true" pageId="17" pageNumber="18">H. argentinensis</emphasis>
|
||
</taxonomicName>
|
||
. Furthermore, the parameres are strongly narrowed before the apex and the median lobe is broad and subcylindrical to the apex, not strongly laterally compressed. Nothing is known about the biology of this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |