182 lines
21 KiB
XML
182 lines
21 KiB
XML
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<mods:title id="62561639689CFFA1D339044FC75AE4F0">Contributions on the treehopper genus Ennya Stål, 1866 (Hemiptera: Membracidae) with two new species from Ecuador</mods:title>
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<mods:name id="8CC902CEF93AD4C1DD2078393367D03F" type="personal">
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<mods:namePart id="9C583EBB39F000EC3A3F265CAC7E53A1">Montalvo-Salazar, Jorge L.</mods:namePart>
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<mods:nameIdentifier id="017CF39580E3FAF1A1F0C58E439D8DBE" type="ORCID">0009-0003-1221-0304</mods:nameIdentifier>
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<mods:affiliation id="1BD4203783D21942B5185C7C88E90983">Universidad San Francisco de Quito USFQ, Colegio de Ciencias Biológicas y Ambientales, Instituto de Biodiversidad Tropical IBIOTROP, Laboratorio de Zoología Terrestre, Museo de Zoología, Quito 170901, Ecuador</mods:affiliation>
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<mods:nameIdentifier id="0F86B8C50CF00C7C83D779A45720F4B9" type="email">jorgemontalvo2000@gmail.com</mods:nameIdentifier>
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<mods:namePart id="9D5A562080DBEFFC9A460ADCB116B474">López-García, Margarita M.</mods:namePart>
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<mods:nameIdentifier id="39A8727D34D64EB0BAE95533C462C3F0" type="ORCID">0000-0003-2796-2931</mods:nameIdentifier>
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<mods:affiliation id="3BEF8A5D60A3EEF890B4F14A12BE2C5F">Universidad San Francisco de Quito USFQ, Colegio de Ciencias Biológicas y Ambientales, Instituto de Biodiversidad Tropical IBIOTROP, Laboratorio de Zoología Terrestre, Museo de Zoología, Quito 170901, Ecuador</mods:affiliation>
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<mods:nameIdentifier id="A1068DC870DABD10030457286738C2F5" type="email">mlopezg@usfq.edu.ec</mods:nameIdentifier>
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<mods:title id="439ABB5584491541DDD5C3914BE9AE60">Zootaxa</mods:title>
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<mods:part id="9270A5AA3063C548DF75848DCEB9D255">
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<mods:date id="55E3947B5BB0B158225F27687EAB2BE7">2024</mods:date>
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<mods:number id="6FB43A3BD3D33B9BDE7CF8B4A2150E55">2024-03-21</mods:number>
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<mods:number id="0737299BFEE4941241AD1DBD64576BDA">5428</mods:number>
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<mods:start id="9492C9ED0D789A9BD24319298701AAEA">269</mods:start>
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<treatment id="03CF652B3815FFACFF5BBCE7FD08A7A5" LSID="urn:lsid:plazi:treatment:03CF652B3815FFACFF5BBCE7FD08A7A5" httpUri="http://treatment.plazi.org/id/03CF652B3815FFACFF5BBCE7FD08A7A5" lastPageNumber="279" pageId="10" pageNumber="279">
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<subSubSection id="C37C87B63815FFACFF5BBCE7FC5BA1C6" box="[151,918,719,746]" pageId="10" pageNumber="279" type="nomenclature">
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<paragraph id="8BD9D43D3815FFACFF5BBCE7FC5BA1C6" blockId="10.[151,918,719,746]" box="[151,918,719,746]" pageId="10" pageNumber="279">
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<heading id="D09163513815FFACFF5BBCE7FC5BA1C6" bold="true" box="[151,918,719,746]" fontSize="11" level="1" pageId="10" pageNumber="279" reason="1">
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<emphasis id="B912082F3815FFACFF5BBCE7FC5BA1C6" bold="true" box="[151,918,719,746]" pageId="10" pageNumber="279">
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<taxonomicName id="4C66AFBE3815FFACFF5BBCE7FD89A1C6" authority="(Goding, 1928)" authorityName="Montalvo-Salazar & López-García" authorityYear="2024" baseAuthorityName="Goding" baseAuthorityYear="1928" box="[151,580,719,746]" class="Insecta" family="Membracidae" genus="Ennya" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="279" phylum="Arthropoda" rank="species" species="funkhouseri" status="stat. nov.">
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<emphasis id="B912082F3815FFACFF5BBCE7FE4FA1C5" bold="true" box="[151,386,719,746]" italics="true" pageId="10" pageNumber="279">Ennya funkhouseri</emphasis>
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(
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<bibRefCitation id="EFF7A9CC3815FFACFE5EBCF8FDF6A1C6" author="Goding, F. W." box="[402,571,719,745]" pageId="10" pageNumber="279" pagination="137 - 142" refId="ref12487" refString="Goding, F. W. (1928) New Membracidae VII. Bulletin of the Brooklyn Entomological Society, 23, 137 - 142." type="journal article" year="1928">Goding, 1928</bibRefCitation>
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)
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||
</taxonomicName>
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<taxonomicNameLabel id="A221B5543815FFACFD87BCE7FD7AA1C5" box="[587,695,720,746]" pageId="10" pageNumber="279">stat. rev.</taxonomicNameLabel>
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(
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<figureCitation id="135DC8B83815FFACFD0BBCF8FC81A1C6" box="[711,844,719,745]" captionStart="FIGURE 6" captionStartId="11.[151,250,1131,1155]" captionTargetBox="[151,1436,186,1108]" captionTargetId="figure-17@11.[151,1436,182,1108]" captionTargetPageId="11" captionText="FIGURE 6. Hille ecuadorensis Fowler lectotype: A. frontal view, B. lateral view, C. labels. Gelastogonia funkhouseri Goding syntype male: D. frontal view, E. lateral view, F. labels. Gelastogonia gibbera Goding syntype female: G. frontal view, H. lateral view, I. labels. Photographs by (A–C) © 2023 Natural History Museum Vienna, Hemiptera Image Collection / H. Bruckner, (D–F) © 2009 National Museum of Natural History, Smithsonian Institution / Thomas J. Henry, EZID, (G–I) © 2023 American Museum of Natural History / Ruth Salas." figureDoi="http://doi.org/10.5281/zenodo.10847611" httpUri="https://zenodo.org/record/10847611/files/figure.png" pageId="10" pageNumber="279">Figs. 6H–I</figureCitation>
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,
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<figureCitation id="135DC8B83815FFACFC96BCF8FC40A1C6" box="[858,909,719,745]" captionStart="FIGURE 10" captionStartId="17.[151,250,1231,1255]" captionTargetBox="[155,1436,322,1208]" captionTargetId="figure-52@17.[151,1436,322,1208]" captionTargetPageId="17" captionText="FIGURE 10. A. Locality records for the species of Ennya with known distribution in Ecuador (Red star: E. lamariela, Yellow star: E. allkukiru, Black point: E. chlorizans, Pink point: E. funkhouseri, Blue point: E. chrysura, Orange point: E. sobria, Gray point: E. scaromizzonoi), B. Forest patch in Pillaro, C–D. La Mariela forest in Moraspungo (Photo by David Brito)." figureDoi="http://doi.org/10.5281/zenodo.10847627" httpUri="https://zenodo.org/record/10847627/files/figure.png" pageId="10" pageNumber="279">10A</figureCitation>
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)
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C37C87B63815FFACFF5BBD23FBDBA046" pageId="10" pageNumber="279" type="reference_group">
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||
<paragraph id="8BD9D43D3815FFACFF5BBD23FDAEA003" blockId="10.[151,1046,788,873]" box="[151,611,788,812]" pageId="10" pageNumber="279">
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<treatmentCitationGroup id="AB76F3133815FFACFF5BBD23FDAEA003" box="[151,611,788,812]" pageId="10" pageNumber="279">
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<treatmentCitation id="0AC7F22C3815FFACFF5BBD23FDAEA003" author="Goding, F. W." box="[151,611,788,812]" page="138" pageId="10" pageNumber="279" year="1928">
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||
<taxonomicName id="4C66AFBE3815FFACFF5BBD23FDAEA003" authority="Goding, 1928: 138" authorityName="Goding (USNM" authorityPageNumber="138" authorityYear="1928" box="[151,611,788,812]" class="Insecta" family="Membracidae" genus="Gelastogonia" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="279" phylum="Arthropoda" rank="species" species="funkhouseri">
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<emphasis id="B912082F3815FFACFF5BBD23FE53A003" box="[151,414,788,812]" italics="true" pageId="10" pageNumber="279">Gelastogonia funkhouseri</emphasis>
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<bibRefCitation id="EFF7A9CC3815FFACFE68BD23FDAEA003" author="Goding, F. W." box="[420,611,788,812]" pageId="10" pageNumber="279" pagination="137 - 142" refId="ref12487" refString="Goding, F. W. (1928) New Membracidae VII. Bulletin of the Brooklyn Entomological Society, 23, 137 - 142." type="journal article" year="1928">Goding, 1928: 138</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="8BD9D43D3815FFACFF5BBD04FD3BA064" blockId="10.[151,1046,788,873]" box="[151,758,818,843]" pageId="10" pageNumber="279">
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||
<treatmentCitationGroup id="AB76F3133815FFACFF5BBD04FD3BA064" box="[151,758,818,843]" pageId="10" pageNumber="279">
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||
<taxonomicName id="4C66AFBE3815FFACFF5BBD04FE6AA064" authority="Goding" authorityName="Goding" authorityYear="2024" box="[151,423,819,843]" class="Insecta" family="Membracidae" genus="Ennya" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="279" phylum="Arthropoda" rank="species" species="funkhouseri">
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<emphasis id="B912082F3815FFACFF5BBD04FE9BA064" box="[151,342,819,843]" italics="true" pageId="10" pageNumber="279">Ennya funkhouseri</emphasis>
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Goding
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</taxonomicName>
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—
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<treatmentCitation id="0AC7F22C3815FFACFE0DBD04FD3BA064" author="Metcalf, Z. P. & Wade, V." box="[449,758,818,843]" page="1026" pageId="10" pageNumber="279" year="1965">
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<bibRefCitation id="EFF7A9CC3815FFACFE0DBD04FD3BA064" author="Metcalf, Z. P. & Wade, V." box="[449,758,818,843]" pageId="10" pageNumber="279" pagination="745 - 1552" refId="ref13006" refString="Metcalf, Z. P. & Wade, V. (1965) Membracoidea. In: General Catalogue of the Homoptera. 2. North Carolina State University, Raleigh, pp. 745 - 1552." type="book chapter" year="1965">Metcalf & Wade (1965: 1026)</bibRefCitation>
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</treatmentCitation>
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||
</treatmentCitationGroup>
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||
</paragraph>
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<paragraph id="8BD9D43D3815FFACFF5BBD65FBDBA046" blockId="10.[151,1046,788,873]" box="[151,1046,849,873]" pageId="10" pageNumber="279">
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<treatmentCitationGroup id="AB76F3133815FFACFF5BBD65FBDBA046" box="[151,1046,849,873]" pageId="10" pageNumber="279">
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<taxonomicName id="4C66AFBE3815FFACFF5BBD65FE34A046" ID-CoL="39W7Q" authority="Fallou Equals" authorityName="Fallou Equals" authorityYear="1890" box="[151,505,849,873]" class="Insecta" family="Membracidae" genus="Ennya" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="279" phylum="Arthropoda" rank="species" species="rufomaculata">
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<emphasis id="B912082F3815FFACFF5BBD65FEAAA046" box="[151,359,849,873]" italics="true" pageId="10" pageNumber="279">Ennya rufomaculata</emphasis>
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Fallou Equals
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</taxonomicName>
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<taxonomicName id="4C66AFBE3815FFACFE33BD65FCC2A046" authority="Goding" authorityName="Goding" authorityYear="2024" box="[511,783,849,873]" class="Insecta" family="Membracidae" genus="Ennya" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="279" phylum="Arthropoda" rank="species" species="funkhouseri">
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<emphasis id="B912082F3815FFACFE33BD65FD73A046" box="[511,702,849,873]" italics="true" pageId="10" pageNumber="279">Ennya funkhouseri</emphasis>
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Goding
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</taxonomicName>
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—
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<treatmentCitation id="0AC7F22C3815FFACFCE5BD66FBDBA046" author="Sakakibara, A. M." box="[809,1046,849,873]" page="95" pageId="10" pageNumber="279" year="1996">
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||
<bibRefCitation id="EFF7A9CC3815FFACFCE5BD66FBDBA046" author="Sakakibara, A. M." box="[809,1046,849,873]" pageId="10" pageNumber="279" pagination="93 - 108" refId="ref13235" refString="Sakakibara, A. M. (1996 a) On some Polyglyptini (Homoptera, Membracidae, Smiliinae): new genus, new species and taxonomic notes. Revista Brasileira de Zoologia, 13, 93 - 108. https: // doi. org / 10.1590 / S 0101 - 81751996000200015" type="journal article" year="1996">Sakakibara (1996a: 95)</bibRefCitation>
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</treatmentCitation>
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C37C87B63815FFACFF5BBDA0FA4EA09D" box="[151,1411,919,946]" pageId="10" pageNumber="279" type="materials_examined">
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||
<paragraph id="8BD9D43D3815FFACFF5BBDA0FA4EA09D" blockId="10.[151,1437,919,1162]" box="[151,1411,919,946]" pageId="10" pageNumber="279">
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||
<emphasis id="B912082F3815FFACFF5BBDA0FEB4A09E" bold="true" box="[151,377,919,945]" pageId="10" pageNumber="279">Material examined</emphasis>
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||
:
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||
<materialsCitation id="3B0EDE603815FFACFE44BDA0FAB2A09D" box="[392,1407,919,946]" collectionCode="USNM" country="Ecuador" location="Huigra" pageId="10" pageNumber="279" specimenCount="1" typeStatus="syntype">
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||
<emphasis id="B912082F3815FFACFE44BDA0FE25A09E" bold="true" box="[392,488,919,945]" pageId="10" pageNumber="279">
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||
<typeStatus id="54DD6A9F3815FFACFE44BDA0FE25A09E" box="[392,488,919,945]" pageId="10" pageNumber="279" type="syntype">Syntype</typeStatus>
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</emphasis>
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(photograph) male
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<taxonomicName id="4C66AFBE3815FFACFD0ABDA0FC25A09E" authorityName="Goding (USNM" authorityYear="1928" box="[710,1000,919,945]" class="Insecta" family="Membracidae" genus="Gelastogonia" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="279" phylum="Arthropoda" rank="species" species="funkhouseri">
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||
<emphasis id="B912082F3815FFACFD0ABDA0FC25A09E" box="[710,1000,919,945]" italics="true" pageId="10" pageNumber="279">Gelastogonia funkhouseri</emphasis>
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||
</taxonomicName>
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||
from
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||
<collectingCountry id="F37194AD3815FFACFBE1BDAFFB74A09D" box="[1069,1209,920,946]" name="Ecuador" pageId="10" pageNumber="279">ECUADOR</collectingCountry>
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||
:
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||
<location id="8EB982E63815FFACFB0FBDA0FADCA09E" LSID="urn:lsid:plazi:treatment:03CF652B3815FFACFF5BBCE7FD08A7A5:8EB982E63815FFACFB0FBDA0FADCA09E" box="[1219,1297,919,945]" country="Ecuador" name="Huigra" pageId="10" pageNumber="279">Huigra</location>
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||
(
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||
<collectionCode id="ED774CF83815FFACFAECBDAFFABAA09D" box="[1312,1399,920,946]" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="279" type="Museum">USNM</collectionCode>
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||
)
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||
</materialsCitation>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C37C87B63815FFACFF0BBD8CFD08A7A5" pageId="10" pageNumber="279" type="discussion">
|
||
<paragraph id="8BD9D43D3815FFACFF0BBD8CFAF6A732" blockId="10.[151,1437,919,1162]" pageId="10" pageNumber="279">
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<emphasis id="B912082F3815FFACFF0BBD8CFE9FA0FA" bold="true" box="[199,338,955,981]" pageId="10" pageNumber="279">Comments:</emphasis>
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||
<bibRefCitation id="EFF7A9CC3815FFACFE95BD8CFDF8A0FA" author="Sakakibara, A. M." box="[345,565,955,981]" pageId="10" pageNumber="279" pagination="93 - 108" refId="ref13235" refString="Sakakibara, A. M. (1996 a) On some Polyglyptini (Homoptera, Membracidae, Smiliinae): new genus, new species and taxonomic notes. Revista Brasileira de Zoologia, 13, 93 - 108. https: // doi. org / 10.1590 / S 0101 - 81751996000200015" type="journal article" year="1996">Sakakibara (1996a)</bibRefCitation>
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||
considered
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||
<taxonomicName id="4C66AFBE3815FFACFD71BD8BFC5CA0FA" authorityName="Goding" authorityYear="2024" box="[701,913,955,981]" class="Insecta" family="Membracidae" genus="Ennya" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="279" phylum="Arthropoda" rank="species" species="funkhouseri">
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<emphasis id="B912082F3815FFACFD71BD8BFC5CA0FA" box="[701,913,955,981]" italics="true" pageId="10" pageNumber="279">Ennya funkhouseri</emphasis>
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</taxonomicName>
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as the junior synonym of
|
||
<taxonomicName id="4C66AFBE3815FFACFB76BD8BFABCA0FA" authorityName="Fallou Equals" authorityYear="1890" box="[1210,1393,955,981]" class="Insecta" family="Membracidae" genus="Ennya" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="279" phylum="Arthropoda" rank="species" species="rufomaculata">
|
||
<emphasis id="B912082F3815FFACFB76BD8BFABCA0FA" box="[1210,1393,955,981]" italics="true" pageId="10" pageNumber="279">E. rufomaculata</emphasis>
|
||
</taxonomicName>
|
||
but the latter does not belong to the genus
|
||
<taxonomicName id="4C66AFBE3815FFACFD95BDD7FD6CA0D6" authorityName="Stal" authorityYear="1866" box="[601,673,992,1017]" class="Insecta" family="Membracidae" genus="Ennya" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="279" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B912082F3815FFACFD95BDD7FD6CA0D6" box="[601,673,992,1017]" italics="true" pageId="10" pageNumber="279">Ennya</emphasis>
|
||
</taxonomicName>
|
||
(see below).
|
||
<taxonomicName id="4C66AFBE3815FFACFCF2BDD7FBDEA0D6" authorityName="Goding" authorityYear="2024" box="[830,1043,991,1017]" class="Insecta" family="Membracidae" genus="Ennya" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="279" phylum="Arthropoda" rank="species" species="funkhouseri">
|
||
<emphasis id="B912082F3815FFACFCF2BDD7FBDEA0D6" box="[830,1043,991,1017]" italics="true" pageId="10" pageNumber="279">Ennya funkhouseri</emphasis>
|
||
</taxonomicName>
|
||
cannot be placed in
|
||
<taxonomicName id="4C66AFBE3815FFACFAC9BDE8FA51A0D6" authorityName="Kirkaldy" authorityYear="1904" box="[1285,1436,991,1017]" class="Insecta" family="Membracidae" genus="Gelastogonia" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="279" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B912082F3815FFACFAC9BDE8FA51A0D6" box="[1285,1436,991,1017]" italics="true" pageId="10" pageNumber="279">Gelastogonia</emphasis>
|
||
</taxonomicName>
|
||
because the pronotum lacks reticulate carinae and the dorsal process is not foliaceous (
|
||
<bibRefCitation id="EFF7A9CC3815FFACFBACBA34FAE2A732" author="Sakakibara, A. M." box="[1120,1327,1027,1053]" pageId="10" pageNumber="279" pagination="93 - 108" refId="ref13235" refString="Sakakibara, A. M. (1996 a) On some Polyglyptini (Homoptera, Membracidae, Smiliinae): new genus, new species and taxonomic notes. Revista Brasileira de Zoologia, 13, 93 - 108. https: // doi. org / 10.1590 / S 0101 - 81751996000200015" type="journal article" year="1996">Sakakibara 1996a</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD9D43D3815FFACFF0BBA1FFD08A7A5" blockId="10.[151,1437,919,1162]" pageId="10" pageNumber="279">
|
||
<taxonomicName id="4C66AFBE3815FFACFF0BBA1FFE54A76E" authorityName="Goding" authorityYear="2024" box="[199,409,1063,1089]" class="Insecta" family="Membracidae" genus="Ennya" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="279" phylum="Arthropoda" rank="species" species="funkhouseri">
|
||
<emphasis id="B912082F3815FFACFF0BBA1FFE54A76E" box="[199,409,1063,1089]" italics="true" pageId="10" pageNumber="279">Ennya funkhouseri</emphasis>
|
||
</taxonomicName>
|
||
differs from its congeners in having the dorsal process trapezoidal and longer than high with the anterior and posterior vertex rounded. This species is similar to
|
||
<taxonomicName id="4C66AFBE3815FFACFC49BA7BFBD4A74A" authorityName="Montalvo-Salazar & López-García" authorityYear="2024" box="[901,1049,1099,1125]" class="Insecta" family="Membracidae" genus="Ennya" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="279" phylum="Arthropoda" rank="species" species="chlorizans" status="stat. rev.">
|
||
<emphasis id="B912082F3815FFACFC49BA7BFBD4A74A" box="[901,1049,1099,1125]" italics="true" pageId="10" pageNumber="279">E. chlorizans</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B912082F3815FFACFBD3BA7BFB4BA749" bold="true" box="[1055,1158,1100,1126]" pageId="10" pageNumber="279">
|
||
<taxonomicNameLabel id="A221B5543815FFACFBD3BA7BFB4BA749" box="[1055,1158,1100,1126]" pageId="10" pageNumber="279">stat. rev.</taxonomicNameLabel>
|
||
</emphasis>
|
||
but the anterior vertex is not acute and not higher than the posterior vertex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |