424 lines
50 KiB
XML
424 lines
50 KiB
XML
<document id="86A44C525B1197EDEF5A97718CF0370E" ID-DOI="10.11646/zootaxa.5428.2.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="10847591" ID-ZooBank="47855779-AAC3-4380-A7D1-06BBE6BF3FF4" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1711009968178" checkinUser="plazi" docAuthor="Montalvo-Salazar, Jorge L. & López-García, Margarita M." docDate="2024" docId="03CF652B3814FFA8FF5BBB51FE60A61E" docLanguage="en" docName="zootaxa.5428.2.6.pdf" docOrigin="Zootaxa 5428 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5428.2.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ennya lamariela Montalvo-Salazar & Lopez-Garcia 2024, sp. nov." docType="treatment" docVersion="3" lastPageNumber="283" masterDocId="FFF61D53381FFFA6FFCCBE37FFCDA32F" masterDocTitle="Contributions on the treehopper genus Ennya Stål, 1866 (Hemiptera: Membracidae) with two new species from Ecuador" masterLastPageNumber="289" masterPageNumber="269" pageNumber="280" updateTime="1711132732293" updateUser="valdenar" zenodo-license-document="CLOSED">
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<mods:title id="BD1E9799670196C5B7DCA1C83A9DDF13">Contributions on the treehopper genus Ennya Stål, 1866 (Hemiptera: Membracidae) with two new species from Ecuador</mods:title>
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<mods:name id="951804AE90DA5C6026637C630D8565A4" type="personal">
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<mods:role id="C4A2BAC723600A1088C5C53E4D50C9E7">
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<mods:roleTerm id="693E957040319B0981902D8280E19027">Author</mods:roleTerm>
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<mods:namePart id="D7624D0B07C1495C6B03BCE58280EDBE">Montalvo-Salazar, Jorge L.</mods:namePart>
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<mods:nameIdentifier id="0CB0CC4C551E912D1439DCF655C02CF4" type="ORCID">0009-0003-1221-0304</mods:nameIdentifier>
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<mods:affiliation id="C24474A5EA895E5A51932527575FF0FA">Universidad San Francisco de Quito USFQ, Colegio de Ciencias Biológicas y Ambientales, Instituto de Biodiversidad Tropical IBIOTROP, Laboratorio de Zoología Terrestre, Museo de Zoología, Quito 170901, Ecuador</mods:affiliation>
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<mods:nameIdentifier id="E7841627B9B16535FE002AD71BF98A2D" type="email">jorgemontalvo2000@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name id="2A2CA69FC88660136AE613A74CB09824" type="personal">
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<mods:roleTerm id="B6EC1D57A8014F23E31A8DDBB0C77CBF">Author</mods:roleTerm>
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<mods:namePart id="8640F8ECCAFB4BA131505A3DEE0A7D6E">López-García, Margarita M.</mods:namePart>
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<mods:nameIdentifier id="14B33D389FEFAD40E0EE1D1B2BED60AF" type="ORCID">0000-0003-2796-2931</mods:nameIdentifier>
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<mods:affiliation id="0F2097D5AE7082771AA77DFF7FE5184B">Universidad San Francisco de Quito USFQ, Colegio de Ciencias Biológicas y Ambientales, Instituto de Biodiversidad Tropical IBIOTROP, Laboratorio de Zoología Terrestre, Museo de Zoología, Quito 170901, Ecuador</mods:affiliation>
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<mods:nameIdentifier id="7503DCAA544D4D1DF688C525702E6B36" type="email">mlopezg@usfq.edu.ec</mods:nameIdentifier>
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<mods:title id="A84DFBD12973C346A16AB2CBAB5404D9">Zootaxa</mods:title>
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<mods:part id="8C66820E68AB58882A5435FC094A2A90">
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<mods:date id="EEC93A792E62BB49F35A94D7A8958C3E">2024</mods:date>
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<mods:number id="AD5B0CE574BAB282F23A36BD62F657D3">2024-03-21</mods:number>
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<mods:number id="639A487C54ABA013CE1845C5E6B399AA">5428</mods:number>
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<mods:detail id="5A2AEE5FC63F779653CDBD639B8B21BB" type="issue">
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<mods:url id="765E962131AB374D6BCF3369BBA15412">http://dx.doi.org/10.11646/zootaxa.5428.2.6</mods:url>
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<mods:classification id="BF5F4C99FA5B3D75D3DD41266ECD7815">journal article</mods:classification>
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<mods:identifier id="91FF101AD94ACCEBAF831D63F95F674B" type="DOI">10.11646/zootaxa.5428.2.6</mods:identifier>
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<treatment id="03CF652B3814FFA8FF5BBB51FE60A61E" LSID="urn:lsid:plazi:treatment:03CF652B3814FFA8FF5BBB51FE60A61E" httpUri="http://treatment.plazi.org/id/03CF652B3814FFA8FF5BBB51FE60A61E" lastPageId="14" lastPageNumber="283" pageId="11" pageNumber="280">
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<subSubSection id="C37C87B63814FFADFF5BBB51FC2AA68B" pageId="11" pageNumber="280" type="nomenclature">
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<paragraph id="8BD9D43D3814FFADFF5BBB51FC49A650" blockId="11.[151,999,1380,1480]" box="[151,900,1380,1408]" pageId="11" pageNumber="280">
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<heading id="D09163513814FFADFF5BBB51FC49A650" bold="true" box="[151,900,1380,1408]" fontSize="11" level="1" pageId="11" pageNumber="280" reason="1">
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<emphasis id="B912082F3814FFADFF5BBB51FC49A650" bold="true" box="[151,900,1380,1408]" pageId="11" pageNumber="280">
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<taxonomicName id="4C66AFBE3814FFADFF5BBB51FCD3A650" authority="Montalvo-Salazar & Lopez-Garcia" authorityName="Montalvo-Salazar & Lopez-Garcia" authorityYear="2024" box="[151,798,1380,1408]" class="Insecta" family="Membracidae" genus="Ennya" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="280" phylum="Arthropoda" rank="species" species="lamariela" status="sp. nov.">
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<emphasis id="B912082F3814FFADFF5BBB51FEAFA6AF" bold="true" box="[151,354,1381,1408]" italics="true" pageId="11" pageNumber="280">Ennya lamariela</emphasis>
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Montalvo-Salazar & López-García
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</taxonomicName>
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<taxonomicNameLabel id="A221B5543814FFADFCE9BB52FC49A650" box="[805,900,1381,1407]" pageId="11" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BD9D43D3814FFADFF5BBBBEFC2AA68B" blockId="11.[151,999,1380,1480]" box="[151,999,1417,1444]" pageId="11" pageNumber="280">
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urn:lsid:zoobank.org:act:
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<uuid id="FFC0EEE83814FFADFE69BBBEFC2AA68B" box="[421,999,1417,1444]" pageId="11" pageNumber="280">A0BBDFFF-6A5A-42EF-BE3A-81AC53945EDB</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37C87B63814FFADFF5BBB9AFE5EA6E8" box="[151,403,1453,1480]" pageId="11" pageNumber="280" type="description">
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<paragraph id="8BD9D43D3814FFADFF5BBB9AFE5EA6E8" blockId="11.[151,999,1380,1480]" box="[151,403,1453,1480]" pageId="11" pageNumber="280">
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(
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<figureCitation id="135DC8B83814FFADFF53BB9AFF22A6E7" box="[159,239,1453,1480]" captionStart="FIGURE 7" captionStartId="12.[151,250,1765,1789]" captionTargetBox="[197,1391,287,1726]" captionTargetId="figure-55@12.[184,1404,274,1743]" captionTargetPageId="12" captionText="FIGURE 7. Ennya lamariela sp. nov. holotype male: A. frontal view, B. lateral view, C. dorsal view. Paratype melanic male: D. lateral view. Last-instar nymph: E. frontal view, F. lateral view. Scale bars: 1 mm (A, E–F) 2 mm (B–D)." figureDoi="http://doi.org/10.5281/zenodo.10847615" httpUri="https://zenodo.org/record/10847615/files/figure.png" pageId="11" pageNumber="280">Figs. 7</figureCitation>
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,
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<figureCitation id="135DC8B83814FFADFF37BB9AFEC7A6E8" box="[251,266,1453,1479]" captionStart="FIGURE 8" captionStartId="13.[151,250,1122,1146]" captionTargetBox="[151,1430,382,1093]" captionTargetId="figure-115@13.[151,1436,382,1100]" captionTargetPageId="13" captionText="FIGURE 8. Ennya lamariela sp. nov. Holotype male: A. aedeagus in lateral view, B. apex of aedeagus in anterior view, C. apex of aedeagus in posterior view, D. styles in dorsal view. Paratype female: E. gonoplac in lateral view, F. second valvulae in lateral view, G. first valvulae in lateral view. Scale bars: 0.1 mm (A, D–E) 0.05 mm (B–C) 0.2 mm (E–G)." figureDoi="http://doi.org/10.5281/zenodo.10847619" httpUri="https://zenodo.org/record/10847619/files/figure.png" pageId="11" pageNumber="280">8</figureCitation>
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,
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<figureCitation id="135DC8B83814FFADFEDBBB9AFE41A6E8" box="[279,396,1453,1479]" captionStart="FIGURE 10" captionStartId="17.[151,250,1231,1255]" captionTargetBox="[155,1436,322,1208]" captionTargetId="figure-52@17.[151,1436,322,1208]" captionTargetPageId="17" captionText="FIGURE 10. A. Locality records for the species of Ennya with known distribution in Ecuador (Red star: E. lamariela, Yellow star: E. allkukiru, Black point: E. chlorizans, Pink point: E. funkhouseri, Blue point: E. chrysura, Orange point: E. sobria, Gray point: E. scaromizzonoi), B. Forest patch in Pillaro, C–D. La Mariela forest in Moraspungo (Photo by David Brito)." figureDoi="http://doi.org/10.5281/zenodo.10847627" httpUri="https://zenodo.org/record/10847627/files/figure.png" pageId="11" pageNumber="280">10A, C–D</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C37C87B63814FFADFF5BBBC2FBCBA51B" pageId="11" pageNumber="280" type="diagnosis">
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<paragraph id="8BD9D43D3814FFADFF5BBBC2FBCBA51B" blockId="11.[151,1437,1525,2020]" pageId="11" pageNumber="280">
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<emphasis id="B912082F3814FFADFF5BBBC2FEDFA520" bold="true" box="[151,274,1525,1551]" pageId="11" pageNumber="280">Diagnosis.</emphasis>
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Metopidium in frontal view sub-quadrate; pronotum with three lateral carinae; dorsal process low and blunt, then sinuous but not produced and finally descending gradually to apex.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C37C87B63814FFA8FF0BB80AFC87A3D6" lastPageId="14" lastPageNumber="283" pageId="11" pageNumber="280" type="description">
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||
<paragraph id="8BD9D43D3814FFADFF0BB80AFCCAA424" blockId="11.[151,1437,1525,2020]" pageId="11" pageNumber="280">
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<emphasis id="B912082F3814FFADFF0BB80AFE94A578" bold="true" box="[199,345,1597,1623]" pageId="11" pageNumber="280">Description.</emphasis>
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<typeStatus id="54DD6A9F3814FFADFEADB80AFE04A578" box="[353,457,1597,1623]" pageId="11" pageNumber="280" type="holotype">Holotype</typeStatus>
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male.
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<emphasis id="B912082F3814FFADFDD4B809FD76A578" box="[536,699,1598,1623]" italics="true" pageId="11" pageNumber="280">Measurements</emphasis>
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(mm): Body length: 5.6; pronotum length 5.6; pronotum height: 1.8; distance between humeral angles: 2.8; head width: 2.2; head height: 9.4.
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<emphasis id="B912082F3814FFADFBC0B856FB80A554" box="[1036,1101,1633,1659]" italics="true" pageId="11" pageNumber="280">Color</emphasis>
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. Mostly fuscous green. Head brown, eyes gray mottled with black spots, ocelli pale yellow. Pronotum pale green with brown spots; metopidium fuscous; humeral angles, dorsal process and posterior apex darker. Forewing membrane smoky with a black apical patch; sclerotized area brown; forewing veins brown to black. Hind wings hyaline, apex smoky hyaline, veins black. Thoracic and abdominal sternites, and legs pale yellow.
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</paragraph>
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<paragraph id="8BD9D43D3814FFADFF0BB922FD21A4B4" blockId="11.[151,1437,1525,2020]" pageId="11" pageNumber="280">
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<emphasis id="B912082F3814FFADFF0BB922FEFFA400" box="[199,306,1813,1839]" italics="true" pageId="11" pageNumber="280">Sculpture</emphasis>
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. Head, pronotum, thoracic sternites and sclerotized areas of forewings densely and deeply punctate, and with sparse short setae, central carinae flanked by long erect setae. Pronotum reticulate with three longitudinal lateral carinae reaching the apex; metopidium with two lateral carinae; dorsal process with trasnversal carinae. Abdominal sternites and legs with dense pubescence.
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</paragraph>
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<paragraph id="8BD9D43D3814FFAAFF0BB992FE21A3D6" blockId="11.[151,1437,1525,2020]" lastBlockId="12.[151,1437,151,249]" lastPageId="12" lastPageNumber="281" pageId="11" pageNumber="280">
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<emphasis id="B912082F3814FFADFF0BB992FEC9A490" box="[199,260,1957,1983]" italics="true" pageId="11" pageNumber="280">Head</emphasis>
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. In frontal view: width (without eyes) approximately half of distance between tips of humeral angles; eye width approximately half vertex width; ocelli closer to coronal suture than to eyes; vertex slightly sinuated; clypeus diamond shaped with width approximately half vertex width; suprantennal ledges large and slightly concave. In ventral view clypeus ventrally convex, maxillary plates flattened with sinuate margins; rostrum reaching posterior margin of metathoracic coxae.
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</paragraph>
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<caption id="DF1984B53813FFAAFF5BB8D2FB2CA40E" ID-DOI="http://doi.org/10.5281/zenodo.10847615" ID-Zenodo-Dep="10847615" httpUri="https://zenodo.org/record/10847615/files/figure.png" pageId="12" pageNumber="281" startId="12.[151,250,1765,1789]" targetBox="[197,1391,287,1726]" targetPageId="12" targetType="figure">
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<paragraph id="8BD9D43D3813FFAAFF5BB8D2FB2CA40E" blockId="12.[151,1437,1765,1825]" pageId="12" pageNumber="281">
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<emphasis id="B912082F3813FFAAFF5BB8D2FEDEA5D2" bold="true" box="[151,275,1765,1789]" pageId="12" pageNumber="281">FIGURE 7.</emphasis>
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<taxonomicName id="4C66AFBE3813FFAAFED6B8D1FE08A5D2" authority="Montalvo-Salazar & López-García, 2024" authorityName="Montalvo-Salazar & López-García" authorityYear="2024" box="[282,453,1765,1789]" class="Insecta" family="Membracidae" genus="Ennya" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="281" phylum="Arthropoda" rank="species" species="lamariela" status="sp. nov.">
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<emphasis id="B912082F3813FFAAFED6B8D1FE08A5D2" box="[282,453,1765,1789]" italics="true" pageId="12" pageNumber="281">Ennya lamariela</emphasis>
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</taxonomicName>
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<emphasis id="B912082F3813FFAAFE07B8D2FDD3A5D2" bold="true" box="[459,542,1765,1789]" pageId="12" pageNumber="281">
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<taxonomicNameLabel id="A221B5543813FFAAFE07B8D2FDD3A5D2" box="[459,542,1765,1789]" pageId="12" pageNumber="281" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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holotype male: A. frontal view, B. lateral view, C. dorsal view. Paratype melanic male: D. lateral view. Last-instar nymph: E. frontal view, F. lateral view. Scale bars: 1 mm (A, E–F) 2 mm (B–D).
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</paragraph>
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</caption>
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<paragraph id="8BD9D43D3813FFABFF0BB944FD06A24A" blockId="12.[151,1437,1907,2042]" lastBlockId="13.[151,1437,151,357]" lastPageId="13" lastPageNumber="282" pageId="12" pageNumber="281">
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<emphasis id="B912082F3813FFAAFF0BB944FED5A4A2" box="[199,280,1907,1933]" italics="true" pageId="12" pageNumber="281">Thorax</emphasis>
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. Pronotum in frontal view with metopidium sub-quadrate, humeral angles blunt triangular produced; dorsal process triangular. In lateral view with metopodium convex; pronotum with low blunt dorsal process, base wider than height, emerging just behind humeral angles and directed upwards, then sinuate and finally descending gradually to acute apex, reaching tips of forewings; central carina sharp and percurrent throughout. In dorsal view diamond-shaped, posterior process elongate, lateral margins posterior to humeral angles weakly sinuous, converging to an acute apex. Forewings with five apical cells, third apical cell one fifth as long as first apical cell, one discoidal cell half length of first apical cell. Metathoracic femur longer and more slender than pro- and mesothoracic femora. Metathoracic tibia with cucullate setae in all three rows [row I irregular / II double / III double apically]. Metathoracic tibia longer than pro- and mesothoracic tibiae. First tarsomere of posterior legs longer than those of anterior and middle legs, with plantar cucullate setae ventrally.
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</paragraph>
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<caption id="DF1984B53812FFABFF5BBA55FB47A7ED" ID-DOI="http://doi.org/10.5281/zenodo.10847619" ID-Zenodo-Dep="10847619" httpUri="https://zenodo.org/record/10847619/files/figure.png" pageId="13" pageNumber="282" startId="13.[151,250,1122,1146]" targetBox="[151,1430,382,1093]" targetPageId="13" targetType="figure">
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<paragraph id="8BD9D43D3812FFABFF5BBA55FB47A7ED" blockId="13.[151,1436,1122,1218]" pageId="13" pageNumber="282">
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<emphasis id="B912082F3812FFABFF5BBA55FEDFA755" bold="true" box="[151,274,1122,1146]" pageId="13" pageNumber="282">FIGURE 8.</emphasis>
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<taxonomicName id="4C66AFBE3812FFABFEDBBA54FE0DA755" authority="Montalvo-Salazar & López-García, 2024" authorityName="Montalvo-Salazar & López-García" authorityYear="2024" box="[279,448,1122,1146]" class="Insecta" family="Membracidae" genus="Ennya" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="282" phylum="Arthropoda" rank="species" species="lamariela" status="sp. nov.">
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<emphasis id="B912082F3812FFABFEDBBA54FE0DA755" box="[279,448,1122,1146]" italics="true" pageId="13" pageNumber="282">Ennya lamariela</emphasis>
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</taxonomicName>
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<emphasis id="B912082F3812FFABFE09BA55FDDAA755" bold="true" box="[453,535,1122,1146]" pageId="13" pageNumber="282">
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<taxonomicNameLabel id="A221B5543812FFABFE09BA55FDDAA755" box="[453,535,1122,1146]" pageId="13" pageNumber="282" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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Holotype male: A. aedeagus in lateral view, B. apex of aedeagus in anterior view, C. apex of aedeagus in posterior view, D. styles in dorsal view. Paratype female: E. gonoplac in lateral view, F. second valvulae in lateral view, G. first valvulae in lateral view. Scale bars: 0.1 mm (A, D–E) 0.05 mm (B–C) 0.2 mm (E–G).
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</paragraph>
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</caption>
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<paragraph id="8BD9D43D3812FFABFF0BBB3BFD33A6BD" blockId="13.[151,1437,1292,2039]" pageId="13" pageNumber="282">
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<emphasis id="B912082F3812FFABFF0BBB3BFEFCA609" box="[199,305,1292,1318]" italics="true" pageId="13" pageNumber="282">Abdomen</emphasis>
|
||
. Aedeagus robust, cylindrical; in lateral view with basal margin truncate and dorsal margin deeply emarginate; posterior arm slightly narrowed toward apex in anterior and caudal view, 2 x as tall as anterior arm; corona with margin rounded, with numerous scale-like dentae on anterior and dorsal region of posterior arm. Styles curved ventrolaterally, apically recurved, tips rounded.
|
||
</paragraph>
|
||
<paragraph id="8BD9D43D3812FFABFF0BBBABFD75A5A1" blockId="13.[151,1437,1292,2039]" pageId="13" pageNumber="282">
|
||
<emphasis id="B912082F3812FFABFF0BBBABFEF7A699" bold="true" box="[199,314,1436,1462]" pageId="13" pageNumber="282">Variation</emphasis>
|
||
.
|
||
<emphasis id="B912082F3812FFABFE84BBAAFE26A699" box="[328,491,1437,1462]" italics="true" pageId="13" pageNumber="282">Measurements</emphasis>
|
||
. Male / Female (mm): Body length: 5.4–5.7/ 5.7–6,0; pronotum length 54–5.7 / 5.7– 6.0; pronotum height: 1.5–1.8 / 1.7–2.0; distance between humeral angles: 2.5–2.6 / 2.7–2.8; head width: 2.1–2.3 / 2.3–2.4; head height: 1.0 / 0.9–1.2. Females tend to be paler than males, mostly yellow-green, and some females pale are pink on the base of the metopidium and head. In some
|
||
<typeStatus id="54DD6A9F3812FFABFCA8B83EFC1FA50C" box="[868,978,1545,1571]" pageId="13" pageNumber="282" type="paratype">paratypes</typeStatus>
|
||
, the metopidium and anterior area of the dorsal process or just the dorsal process is darker than the rest of the pronotum.
|
||
<specimenCount id="9D601FB43812FFABFBEBB81BFB59A569" box="[1063,1172,1580,1606]" count="1" pageId="13" pageNumber="282" type="male">One male</specimenCount>
|
||
<typeStatus id="54DD6A9F3812FFABFB50B81AFB31A568" box="[1180,1276,1581,1607]" pageId="13" pageNumber="282" type="paratype">paratype</typeStatus>
|
||
is completely black mottled with brown spots. Transverse carinae may be absent or more than one. The pattern of the carinae of the dorsal process is different in each individual.
|
||
</paragraph>
|
||
<paragraph id="8BD9D43D3812FFABFF0BB8AFFC5AA5D4" blockId="13.[151,1437,1292,2039]" pageId="13" pageNumber="282">
|
||
<emphasis id="B912082F3812FFABFF0BB8AFFE48A59D" box="[199,389,1688,1714]" italics="true" pageId="13" pageNumber="282">Female genitalia</emphasis>
|
||
. Gonoplac with setae along ventral margin. First valvulae gradually narrowing toward a sharp apex. Second valvulae with parallel margins until the apical third where dorsal margin is abruptly widened; apex ending in a rounded point; dorsal margin minutely crenulate at apex.
|
||
</paragraph>
|
||
<paragraph id="8BD9D43D3812FFA8FF0BB933FC87A3D6" blockId="13.[151,1437,1292,2039]" lastBlockId="14.[151,1437,151,1329]" lastPageId="14" lastPageNumber="283" pageId="13" pageNumber="282">
|
||
<emphasis id="B912082F3812FFABFF0BB933FDF2A431" bold="true" box="[199,575,1796,1822]" pageId="13" pageNumber="282">Fifth-instar nymph description</emphasis>
|
||
.
|
||
<emphasis id="B912082F3812FFABFD9CB933FD5CA431" box="[592,657,1796,1822]" italics="true" pageId="13" pageNumber="282">Color</emphasis>
|
||
. Body mostly brown greenish to black, head with two longitudinal darker stripes, pronotum with central carinae black.
|
||
<emphasis id="B912082F3812FFABFD2AB91FFCB7A46D" box="[742,890,1832,1858]" italics="true" pageId="13" pageNumber="282">Overall body</emphasis>
|
||
. Dense chalazal setae simple, needlelike; dorsal contour of abdomen in lateral view straight to slightly convex. A pair of small protuberances close to the posterior margin of head.
|
||
<emphasis id="B912082F3812FFABFE9BB947FE07A4A5" box="[343,458,1904,1930]" italics="true" pageId="13" pageNumber="282">Prothorax</emphasis>
|
||
. Pre- and postmetopidum without scoli; pronotum with a short blunt dorsal process; posterior extension of pronotum surpassing posterior margin of metanotum, reaching to second segment of abdomen; in lateral view: metopidium concave, then straight and finally descending to posterior apex, pronotum trapezoidal; in dorsal view posterior process subtriangular, apex acute.
|
||
<emphasis id="B912082F3812FFABFCF9B9EBFC76A4D9" box="[821,955,2012,2038]" italics="true" pageId="13" pageNumber="282">Mesothorax</emphasis>
|
||
. Scoli absent; forewing pad costal margin slightly concave; forewing pad chalazae sparse. Metathorax. Scoli absent.
|
||
<emphasis id="B912082F3811FFA8FC24BEA0FBD0A39F" box="[1000,1053,151,176]" italics="true" pageId="14" pageNumber="283">Legs</emphasis>
|
||
. Chalazae on all surface of tibiae with enlarged chalazae on apex.
|
||
<emphasis id="B912082F3811FFA8FE37BE8CFDA8A3FA" box="[507,613,187,213]" italics="true" pageId="14" pageNumber="283">Abdomen</emphasis>
|
||
. Without scoli; chalazae subequal on all segments, lateral rows of enlarged chalazae absent; segment IX dorsal and ventral lengths equal.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37C87B63811FFA8FF0BBF34FD1EA186" pageId="14" pageNumber="283" type="discussion">
|
||
<paragraph id="8BD9D43D3811FFA8FF0BBF34FD1EA186" blockId="14.[151,1437,151,1329]" pageId="14" pageNumber="283">
|
||
<emphasis id="B912082F3811FFA8FF0BBF34FEF9A232" bold="true" box="[199,308,259,285]" pageId="14" pageNumber="283">Remarks</emphasis>
|
||
. This is the first known species of the genus
|
||
<taxonomicName id="4C66AFBE3811FFA8FCFFBF34FCB6A233" authorityName="Stal" authorityYear="1866" box="[819,891,259,284]" class="Insecta" family="Membracidae" genus="Ennya" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="283" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B912082F3811FFA8FCFFBF34FCB6A233" box="[819,891,259,284]" italics="true" pageId="14" pageNumber="283">Ennya</emphasis>
|
||
</taxonomicName>
|
||
in which the shape of the metopidium in frontal view is sub-quadrate; the other species exhibit a rounded shape.
|
||
<taxonomicName id="4C66AFBE3811FFA8FC4EBF10FB8DA26E" authority="Montalvo-Salazar & López-García, 2024" authorityName="Montalvo-Salazar & López-García" authorityYear="2024" box="[898,1088,295,321]" class="Insecta" family="Membracidae" genus="Ennya" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="283" phylum="Arthropoda" rank="species" species="lamariela" status="sp. nov.">
|
||
<emphasis id="B912082F3811FFA8FC4EBF10FB8DA26E" box="[898,1088,295,321]" italics="true" pageId="14" pageNumber="283">Ennya lamariela</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B912082F3811FFA8FB86BF10FB6AA26E" bold="true" box="[1098,1191,295,321]" pageId="14" pageNumber="283">
|
||
<taxonomicNameLabel id="A221B5543811FFA8FB86BF10FB6AA26E" box="[1098,1191,295,321]" pageId="14" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is most similar to
|
||
<taxonomicName id="4C66AFBE3811FFA8FA48BF10FF28A24A" baseAuthorityName="Sakakibara" baseAuthorityYear="1996" class="Insecta" family="Membracidae" genus="Ennya" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="283" phylum="Arthropoda" rank="species" species="andina">
|
||
<emphasis id="B912082F3811FFA8FA48BF10FF28A24A" italics="true" pageId="14" pageNumber="283">E. andina</emphasis>
|
||
</taxonomicName>
|
||
with the triangular dorsal process being short, blunt and directed upwards. However, the pronotum of
|
||
<taxonomicName id="4C66AFBE3811FFA8FA48BF7CFEC9A2A6" authority="Montalvo-Salazar & López-García, 2024" authorityName="Montalvo-Salazar & López-García" authorityYear="2024" class="Insecta" family="Membracidae" genus="Ennya" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="283" phylum="Arthropoda" rank="species" species="lamariela" status="sp. nov.">
|
||
<emphasis id="B912082F3811FFA8FA48BF7CFEC9A2A6" italics="true" pageId="14" pageNumber="283">E. lamariela</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B912082F3811FFA8FEC6BF58FEAEA2A6" bold="true" box="[266,355,367,393]" pageId="14" pageNumber="283">
|
||
<taxonomicNameLabel id="A221B5543811FFA8FEC6BF58FEAEA2A6" box="[266,355,367,393]" pageId="14" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has a second elevation not produced next to the dorsal process while, in
|
||
<emphasis id="B912082F3811FFA8FBB3BF58FB3CA2A6" box="[1151,1265,367,393]" italics="true" pageId="14" pageNumber="283">
|
||
<taxonomicName id="4C66AFBE3811FFA8FBB3BF58FB23A2A6" baseAuthorityName="Sakakibara" baseAuthorityYear="1996" box="[1151,1262,367,393]" class="Insecta" family="Membracidae" genus="Ennya" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="283" phylum="Arthropoda" rank="species" species="andina">E. andina</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
after the dorsal process the pronotum descends gradually to the apex.
|
||
<taxonomicName id="4C66AFBE3811FFA8FD21BFA4FC65A282" authority="Montalvo-Salazar & López-García, 2024" authorityName="Montalvo-Salazar & López-García" authorityYear="2024" box="[749,936,403,429]" class="Insecta" family="Membracidae" genus="Ennya" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="283" phylum="Arthropoda" rank="species" species="lamariela" status="sp. nov.">
|
||
<emphasis id="B912082F3811FFA8FD21BFA4FC65A282" box="[749,936,403,429]" italics="true" pageId="14" pageNumber="283">Ennya lamariela</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B912082F3811FFA8FC61BFA4FBCAA282" bold="true" box="[941,1031,403,429]" pageId="14" pageNumber="283">
|
||
<taxonomicNameLabel id="A221B5543811FFA8FC61BFA4FBCAA282" box="[941,1031,403,429]" pageId="14" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C66AFBE3811FFA8FB55BFA4FADCA282" box="[1177,1297,403,429]" class="Insecta" genus="Hille" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="283" phylum="Arthropoda" rank="species" species="pacifica">
|
||
<emphasis id="B912082F3811FFA8FB55BFA4FADCA282" box="[1177,1297,403,429]" italics="true" pageId="14" pageNumber="283">E. pacifica</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C66AFBE3811FFA8FAD2BFA4FA5BA282" authorityName="Sakakibara" authorityYear="1996" box="[1310,1430,403,429]" class="Insecta" family="Membracidae" genus="Ennya" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="283" phylum="Arthropoda" rank="species" species="bordoni">
|
||
<emphasis id="B912082F3811FFA8FAD2BFA4FA5BA282" box="[1310,1430,403,429]" italics="true" pageId="14" pageNumber="283">E. bordoni</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C66AFBE3811FFA8FF5BBF80FECCA2FF" box="[151,257,439,464]" class="Insecta" genus="Hille" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="283" phylum="Arthropoda" rank="species" species="notata">
|
||
<emphasis id="B912082F3811FFA8FF5BBF80FECCA2FF" box="[151,257,439,464]" italics="true" pageId="14" pageNumber="283">E. notata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B912082F3811FFA8FEDDBF80FE48A2FF" box="[273,389,439,464]" italics="true" pageId="14" pageNumber="283">
|
||
<taxonomicName id="4C66AFBE3811FFA8FEDDBF80FE4DA2FF" box="[273,384,439,464]" class="Insecta" genus="Hille" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="283" phylum="Arthropoda" rank="species" species="nutans">E. nutans</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C66AFBE3811FFA8FE0DBF80FDF4A2FE" box="[449,569,439,465]" class="Insecta" family="Membracidae" genus="Gelastogonia" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="283" phylum="Arthropoda" rank="species" species="gibbera">
|
||
<emphasis id="B912082F3811FFA8FE0DBF80FDF4A2FE" box="[449,569,439,465]" italics="true" pageId="14" pageNumber="283">E. gibbera</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B912082F3811FFA8FD8FBF81FD2CA2FE" bold="true" box="[579,737,438,465]" pageId="14" pageNumber="283">reinst. comb.</emphasis>
|
||
in having the anterior margin of the dorsal process not sub-perpendicular but oblique. The triangular dorsal process directed upwards in
|
||
<taxonomicName id="4C66AFBE3811FFA8FC3CBFECFBB6A2DA" authority="Montalvo-Salazar & López-García, 2024" authorityName="Montalvo-Salazar & López-García" authorityYear="2024" box="[1008,1147,475,501]" class="Insecta" family="Membracidae" genus="Ennya" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="283" phylum="Arthropoda" rank="species" species="lamariela" status="sp. nov.">
|
||
<emphasis id="B912082F3811FFA8FC3CBFECFBB6A2DA" box="[1008,1147,475,501]" italics="true" pageId="14" pageNumber="283">E. lamariela</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B912082F3811FFA8FB4EBFECFB11A2DA" bold="true" box="[1154,1244,475,501]" pageId="14" pageNumber="283">
|
||
<taxonomicNameLabel id="A221B5543811FFA8FB4EBFECFB11A2DA" box="[1154,1244,475,501]" pageId="14" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
separates it from
|
||
<taxonomicName id="4C66AFBE3811FFA8FF5BBFC8FE9EA136" box="[151,339,511,537]" class="Insecta" genus="Hille" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="283" phylum="Arthropoda" rank="species" species="ecuadoriensis">
|
||
<emphasis id="B912082F3811FFA8FF5BBFC8FE9EA136" box="[151,339,511,537]" italics="true" pageId="14" pageNumber="283">E. ecuadoriensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C66AFBE3811FFA8FEADBFC8FE3EA137" authorityName="Stal (NHRS" authorityYear="1869" box="[353,499,511,536]" class="Insecta" genus="Hille" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="283" phylum="Arthropoda" rank="species" species="conspersa">
|
||
<emphasis id="B912082F3811FFA8FEADBFC8FE3EA137" box="[353,499,511,536]" italics="true" pageId="14" pageNumber="283">E. conspersa</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<emphasis id="B912082F3811FFA8FDFEBFC8FD26A136" box="[562,747,511,537]" italics="true" pageId="14" pageNumber="283">
|
||
<taxonomicName id="4C66AFBE3811FFA8FDFEBFC8FD2AA136" box="[562,743,511,537]" class="Insecta" family="Membracidae" genus="Ennya" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="283" phylum="Arthropoda" rank="species" species="maculicornis">E. maculicornis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
which all have a dorsal process that is directed forward. The blunt dorsal process of
|
||
<taxonomicName id="4C66AFBE3811FFA8FE5ABC14FDEDA112" authority="Montalvo-Salazar & López-García, 2024" authorityName="Montalvo-Salazar & López-García" authorityYear="2024" box="[406,544,547,573]" class="Insecta" family="Membracidae" genus="Ennya" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="283" phylum="Arthropoda" rank="species" species="lamariela" status="sp. nov.">
|
||
<emphasis id="B912082F3811FFA8FE5ABC14FDEDA112" box="[406,544,547,573]" italics="true" pageId="14" pageNumber="283">E. lamariela</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B912082F3811FFA8FDE9BC14FDB2A112" bold="true" box="[549,639,547,573]" pageId="14" pageNumber="283">
|
||
<taxonomicNameLabel id="A221B5543811FFA8FDE9BC14FDB2A112" box="[549,639,547,573]" pageId="14" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
distinguishes it from
|
||
<taxonomicName id="4C66AFBE3811FFA8FCA0BC14FC1FA112" baseAuthorityName="Walker" baseAuthorityYear="1851" box="[876,978,547,573]" class="Insecta" family="Membracidae" genus="Ennya" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="283" phylum="Arthropoda" rank="species" species="sobria">
|
||
<emphasis id="B912082F3811FFA8FCA0BC14FC1FA112" box="[876,978,547,573]" italics="true" pageId="14" pageNumber="283">E. sobria</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C66AFBE3811FFA8FC12BC14FB8BA113" box="[990,1094,547,572]" class="Insecta" family="Membracidae" genus="Ennya" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="283" phylum="Arthropoda" rank="species" species="conica">
|
||
<emphasis id="B912082F3811FFA8FC12BC14FB8BA113" box="[990,1094,547,572]" italics="true" pageId="14" pageNumber="283">E. conica</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C66AFBE3811FFA8FB9EBC14FB04A112" box="[1106,1225,547,573]" class="Insecta" genus="Hille" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="283" phylum="Arthropoda" rank="species" species="dorsalis">
|
||
<emphasis id="B912082F3811FFA8FB9EBC14FB04A112" box="[1106,1225,547,573]" italics="true" pageId="14" pageNumber="283">E. dorsalis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C66AFBE3811FFA8FB1ABC14FA58A113" box="[1238,1429,547,572]" class="Insecta" family="Membracidae" genus="Ennya" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="283" phylum="Arthropoda" rank="species" species="scaramozzinoi">
|
||
<emphasis id="B912082F3811FFA8FB1ABC14FA58A113" box="[1238,1429,547,572]" italics="true" pageId="14" pageNumber="283">E. scaramozzinoi</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C66AFBE3811FFA8FF06BC70FE9CA14E" authority="Montalvo-Salazar & López-García, 2024" authorityName="Montalvo-Salazar & López-García" authorityYear="2024" box="[202,337,583,609]" class="Insecta" family="Membracidae" genus="Ennya" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="283" phylum="Arthropoda" rank="species" species="allkukiru" status="sp. nov.">
|
||
<emphasis id="B912082F3811FFA8FF06BC70FE9CA14E" box="[202,337,583,609]" italics="true" pageId="14" pageNumber="283">E. allkukiru</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B912082F3811FFA8FE97BC70FE75A14E" bold="true" box="[347,440,583,609]" pageId="14" pageNumber="283">
|
||
<taxonomicNameLabel id="A221B5543811FFA8FE97BC70FE75A14E" box="[347,440,583,609]" pageId="14" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, which all have an acute dorsal process. Lastly,
|
||
<taxonomicName id="4C66AFBE3811FFA8FC3CBC70FBB2A14E" authority="Montalvo-Salazar & López-García, 2024" authorityName="Montalvo-Salazar & López-García" authorityYear="2024" box="[1008,1151,583,609]" class="Insecta" family="Membracidae" genus="Ennya" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="283" phylum="Arthropoda" rank="species" species="lamariela" status="sp. nov.">
|
||
<emphasis id="B912082F3811FFA8FC3CBC70FBB2A14E" box="[1008,1151,583,609]" italics="true" pageId="14" pageNumber="283">E. lamariela</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B912082F3811FFA8FB45BC70FB2AA14E" bold="true" box="[1161,1255,583,609]" pageId="14" pageNumber="283">
|
||
<taxonomicNameLabel id="A221B5543811FFA8FB45BC70FB2AA14E" box="[1161,1255,583,609]" pageId="14" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C66AFBE3811FFA8FA48BC70FED6A1AA" authorityName="Sakakibara" authorityYear="1996" class="Insecta" family="Membracidae" genus="Ennya" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="283" phylum="Arthropoda" rank="species" species="colombiana">
|
||
<emphasis id="B912082F3811FFA8FA48BC70FED6A1AA" italics="true" pageId="14" pageNumber="283">E. colombiana</emphasis>
|
||
</taxonomicName>
|
||
by the presence of a dorsal process, and from
|
||
<taxonomicName id="4C66AFBE3811FFA8FCE9BC5CFC64A1AA" baseAuthorityName="Fairmaire" baseAuthorityYear="1846" box="[805,937,619,645]" class="Insecta" family="Membracidae" genus="Ennya" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="283" phylum="Arthropoda" rank="species" species="chrysura">
|
||
<emphasis id="B912082F3811FFA8FCE9BC5CFC64A1AA" box="[805,937,619,645]" italics="true" pageId="14" pageNumber="283">E. chrysura</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C66AFBE3811FFA8FC7BBC5CFB97A1AA" authorityName="Goding (USNM" authorityYear="1928" box="[951,1114,619,645]" class="Insecta" family="Membracidae" genus="Gelastogonia" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="283" phylum="Arthropoda" rank="species" species="funkhouseri">
|
||
<emphasis id="B912082F3811FFA8FC7BBC5CFB97A1AA" box="[951,1114,619,645]" italics="true" pageId="14" pageNumber="283">E. funkhouseri</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C66AFBE3811FFA8FB54BC5CFAE1A1AA" authorityName="Montalvo-Salazar & López-García" authorityYear="2024" box="[1176,1324,619,645]" class="Insecta" family="Membracidae" genus="Ennya" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="283" phylum="Arthropoda" rank="species" species="chlorizans">
|
||
<emphasis id="B912082F3811FFA8FB54BC5CFAE1A1AA" box="[1176,1324,619,645]" italics="true" pageId="14" pageNumber="283">E. chlorizans</emphasis>
|
||
</taxonomicName>
|
||
in having the dorsal process triangular instead of trapezoidal.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37C87B63811FFA8FF0BBC85FBE8A1E2" box="[199,1061,690,717]" pageId="14" pageNumber="283" type="distribution">
|
||
<paragraph id="8BD9D43D3811FFA8FF0BBC85FBE8A1E2" blockId="14.[151,1437,151,1329]" box="[199,1061,690,717]" pageId="14" pageNumber="283">
|
||
<emphasis id="B912082F3811FFA8FF0BBC85FE94A1E3" bold="true" box="[199,345,690,716]" pageId="14" pageNumber="283">Distribution</emphasis>
|
||
. Western foothills of the Ecuadorian Andes at
|
||
<quantity id="4C9E79D83811FFA8FCA4BC85FC62A1E2" box="[872,943,690,717]" metricMagnitude="2" metricUnit="m" metricValue="8.75" pageId="14" pageNumber="283" unit="m" value="875.0">875 m</quantity>
|
||
(
|
||
<figureCitation id="135DC8B83811FFA8FC72BC84FBD5A1E2" box="[958,1048,691,717]" captionStart="FIGURE 9" captionStartId="16.[151,250,1184,1208]" captionTargetBox="[164,1423,545,1161]" captionTargetId="figure-90@16.[164,1423,540,1161]" captionTargetPageId="16" captionText="FIGURE 9. Creonus fasciata (Oxygonia fasciata syntype): A. frontal view, B. lateral view, C. labels. Centronodus gounellei (Heliria gounellei syntype): D. frontal view, E. lateral view, F. labels. Scale bars: 1 mm (A–B) 2 mm (D–E). Photos by Laurent Fauvre © 2023 Muséum National d’Histoire Naturelle." figureDoi="http://doi.org/10.5281/zenodo.10847623" httpUri="https://zenodo.org/record/10847623/files/figure.png" pageId="14" pageNumber="283">Fig. 9A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37C87B63811FFA8FF0BBCE1FDFFA7C6" pageId="14" pageNumber="283" type="materials_examined">
|
||
<paragraph id="8BD9D43D3811FFA8FF0BBCE1FEF7A08A" blockId="14.[151,1437,151,1329]" pageId="14" pageNumber="283">
|
||
<emphasis id="B912082F3811FFA8FF0BBCE1FE63A1DF" bold="true" box="[199,430,726,752]" pageId="14" pageNumber="283">Material examined</emphasis>
|
||
.
|
||
<materialsCitation id="3B0EDE603811FFA8FE0CBCE0FE96A072" collectingDate="2023-02-26" collectorName="Ex. Manual" country="Ecuador" elevation="875" location="Moraspungo" pageId="14" pageNumber="283" specimenCount="2" stateProvince="Cotopaxi" typeStatus="holotype">
|
||
<emphasis id="B912082F3811FFA8FE0CBCE0FDE0A1DE" bold="true" box="[448,557,727,753]" pageId="14" pageNumber="283">
|
||
<typeStatus id="54DD6A9F3811FFA8FE0CBCE0FDE0A1DE" box="[448,557,727,753]" pageId="14" pageNumber="283" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
male from
|
||
<collectingCountry id="F37194AD3811FFA8FD77BCE0FC8AA1DE" box="[699,839,727,753]" name="Ecuador" pageId="14" pageNumber="283">ECUADOR</collectingCountry>
|
||
:
|
||
<collectingRegion id="49A21ADF3811FFA8FC9ABCE0FC0CA1DE" box="[854,961,727,753]" country="Ecuador" name="Cotopaxi" pageId="14" pageNumber="283">Cotopaxi</collectingRegion>
|
||
:
|
||
<location id="8EB982E63811FFA8FC1CBCE0FBADA1DE" LSID="urn:lsid:plazi:treatment:03CF652B3814FFA8FF5BBB51FE60A61E:8EB982E63811FFA8FC1CBCE0FBADA1DE" box="[976,1120,727,753]" country="Ecuador" name="Moraspungo" pageId="14" pageNumber="283" stateProvince="Cotopaxi">Moraspungo</location>
|
||
, La Mariela. “ECUADOR, Cotopaxi / Moraspungo
|
||
<quantity id="4C9E79D83811FFA8FE71BCCDFE33A03A" box="[445,510,762,789]" metricMagnitude="2" metricUnit="m" metricValue="8.75" pageId="14" pageNumber="283" unit="m" value="875.0">
|
||
<elevation id="004B330E3811FFA8FE71BCCDFE33A03A" box="[445,510,762,789]" metricMagnitude="2" metricUnit="m" metricValue="8.75" pageId="14" pageNumber="283" unit="m" value="875.0">875m</elevation>
|
||
</quantity>
|
||
/ Lat: -1.0866111; Lon: -79.1893888 / Montalvo, J. & López-García, M.M.
|
||
<date id="FFD8F2FD3811FFA8FF5BBD28FEDBA016" box="[151,278,799,825]" pageId="14" pageNumber="283" value="2023-02-26">
|
||
<collectingDate id="EF9C0B153811FFA8FF5BBD28FEDBA016" box="[151,278,799,825]" pageId="14" pageNumber="283" value="2023-02-26">26Feb2023</collectingDate>
|
||
</date>
|
||
/
|
||
<collectorName id="2693B1EB3811FFA8FEFCBD28FE7BA016" box="[304,438,799,825]" pageId="14" pageNumber="283">Ex. Manual</collectorName>
|
||
//
|
||
<taxonomicName id="4C66AFBE3811FFA8FE14BD28FD5CA016" authorityName="Montalvo-Salazar & López-García" authorityYear="2024" box="[472,657,799,825]" class="Insecta" family="Membracidae" genus="Ennya" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="283" phylum="Arthropoda" rank="species" species="lamariela">Ennya lamariela</taxonomicName>
|
||
/ Montalvo, J. & López-García, M.M. / HOLOTYPE (in red label) // ZSFQ-i11333”
|
||
</materialsCitation>
|
||
.
|
||
<emphasis id="B912082F3811FFA8FEA4BD74FE2DA072" bold="true" box="[360,480,835,861]" pageId="14" pageNumber="283">
|
||
<typeStatus id="54DD6A9F3811FFA8FEA4BD74FE2DA072" box="[360,480,835,861]" pageId="14" pageNumber="283" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
.
|
||
<specimenCount id="9D601FB43811FFA8FE3CBD74FD59A072" box="[496,660,835,861]" count="7" pageId="14" pageNumber="283" type="female">Seven females</specimenCount>
|
||
ZSFQ-i11333:11340 and
|
||
<materialsCitation id="3B0EDE603811FFA8FC0DBD74FE5FA0AE" collectingDate="2023-02-26" collectorName="Ex. Manual" country="Ecuador" elevation="875" location="Moraspungo" pageId="14" pageNumber="283" specimenCount="3" specimenCount-male="3" stateProvince="Cotopaxi" typeStatus="paratype">
|
||
<specimenCount id="9D601FB43811FFA8FC0DBD74FB8EA072" box="[961,1091,835,861]" count="3" pageId="14" pageNumber="283" type="male">three males</specimenCount>
|
||
ZSFQ-i11341:11343 with the same data as holotype
|
||
</materialsCitation>
|
||
.
|
||
<emphasis id="B912082F3811FFA8FE50BD51FD4AA0AF" bold="true" box="[412,647,870,896]" pageId="14" pageNumber="283">
|
||
<typeStatus id="54DD6A9F3811FFA8FE50BD51FD4AA0AF" box="[412,647,870,896]" isOtherTypeStatus="true" pageId="14" pageNumber="283">Additional material</typeStatus>
|
||
</emphasis>
|
||
. Six nymphs at different stages with the same data as holotype ZSFQi11344:11349.
|
||
</paragraph>
|
||
<paragraph id="8BD9D43D3811FFA8FF0BBD98FBA7A0C2" blockId="14.[151,1437,151,1329]" pageId="14" pageNumber="283">
|
||
<emphasis id="B912082F3811FFA8FF0BBD98FEDCA0E6" bold="true" box="[199,273,943,969]" pageId="14" pageNumber="283">Notes.</emphasis>
|
||
<typeStatus id="54DD6A9F3811FFA8FED5BD98FE4CA0E6" box="[281,385,943,969]" pageId="14" pageNumber="283" type="holotype">Holotype</typeStatus>
|
||
and
|
||
<typeStatus id="54DD6A9F3811FFA8FE75BD98FDE9A0E6" box="[441,548,943,969]" pageId="14" pageNumber="283" type="paratype">paratypes</typeStatus>
|
||
minuten-mounted, in excellent state of preservation.
|
||
<typeStatus id="54DD6A9F3811FFA8FB4CBD98FB25A0E6" box="[1152,1256,943,969]" pageId="14" pageNumber="283" type="holotype">Holotype</typeStatus>
|
||
and
|
||
<specimenCount id="9D601FB43811FFA8FAECBD98FA51A0E6" box="[1312,1436,943,969]" count="1" pageId="14" pageNumber="283" type="female">one female</specimenCount>
|
||
<typeStatus id="54DD6A9F3811FFA8FF5BBDE4FF3AA0C2" box="[151,247,979,1005]" pageId="14" pageNumber="283" type="paratype">paratype</typeStatus>
|
||
with dissected abdomens placed in vials with glycerol pinned with specimens.
|
||
</paragraph>
|
||
<paragraph id="8BD9D43D3811FFA8FF0BBDC0FA54A78E" blockId="14.[151,1437,151,1329]" pageId="14" pageNumber="283">
|
||
<emphasis id="B912082F3811FFA8FF0BBDC0FE4CA73F" bold="true" box="[199,385,1014,1041]" pageId="14" pageNumber="283">Natural history</emphasis>
|
||
.
|
||
<taxonomicName id="4C66AFBE3811FFA8FE5DBDC0FD83A73E" authority="Montalvo-Salazar & López-García, 2024" authorityName="Montalvo-Salazar & López-García" authorityYear="2024" box="[401,590,1015,1041]" class="Insecta" family="Membracidae" genus="Ennya" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="283" phylum="Arthropoda" rank="species" species="lamariela" status="sp. nov.">
|
||
<emphasis id="B912082F3811FFA8FE5DBDC0FD83A73E" box="[401,590,1015,1041]" italics="true" pageId="14" pageNumber="283">Ennya lamariela</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B912082F3811FFA8FD9BBDC0FD7EA73E" bold="true" box="[599,691,1015,1041]" pageId="14" pageNumber="283">
|
||
<taxonomicNameLabel id="A221B5543811FFA8FD9BBDC0FD7EA73E" box="[599,691,1015,1041]" pageId="14" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
inhabits the remnants of the mature forest of Occidental Foothill Forest of Northern Andes, in a zone extremely fragmented by banana (
|
||
<taxonomicName id="4C66AFBE3811FFA8FC0FBA2CFB66A71A" box="[963,1195,1051,1077]" class="Liliopsida" family="Musaceae" genus="Musa" isHybrid="true" kingdom="Plantae" order="Zingiberales" pageId="14" pageNumber="283" phylum="Tracheophyta" rank="species" species="paradisiaca">
|
||
<emphasis id="B912082F3811FFA8FC0FBA2CFB66A71A" box="[963,1195,1051,1077]" italics="true" pageId="14" pageNumber="283">Musa × paradisiaca</emphasis>
|
||
</taxonomicName>
|
||
L.) monoculture and livestock (
|
||
<figureCitation id="135DC8B83811FFA8FEC1BA08FE58A776" box="[269,405,1087,1113]" captionStart="FIGURE 9" captionStartId="16.[151,250,1184,1208]" captionTargetBox="[164,1423,545,1161]" captionTargetId="figure-90@16.[164,1423,540,1161]" captionTargetPageId="16" captionText="FIGURE 9. Creonus fasciata (Oxygonia fasciata syntype): A. frontal view, B. lateral view, C. labels. Centronodus gounellei (Heliria gounellei syntype): D. frontal view, E. lateral view, F. labels. Scale bars: 1 mm (A–B) 2 mm (D–E). Photos by Laurent Fauvre © 2023 Muséum National d’Histoire Naturelle." figureDoi="http://doi.org/10.5281/zenodo.10847623" httpUri="https://zenodo.org/record/10847623/files/figure.png" pageId="14" pageNumber="283">Figs. 9C–D</figureCitation>
|
||
). Adult specimens and nymphs at different stages were found in an aggregation on the low branches at
|
||
<quantity id="4C9E79D83811FFA8FED3BA55FEBFA752" box="[287,370,1122,1149]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="283" unit="cm" value="150.0">150 cm</quantity>
|
||
above the ground of an undetermined species of Caesalpinioideae D.C. (
|
||
<taxonomicName id="4C66AFBE3811FFA8FB61BA54FAD1A752" box="[1197,1308,1123,1149]" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" order="Fabales" pageId="14" pageNumber="283" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
|
||
). However, egg masses or scars were not found. The proportion of nymphs was higher than adults, as was females over males.
|
||
</paragraph>
|
||
<paragraph id="8BD9D43D3811FFA8FF0BBA9CFDFFA7C6" blockId="14.[151,1437,151,1329]" pageId="14" pageNumber="283">
|
||
Species of the genus
|
||
<taxonomicName id="4C66AFBE3811FFA8FE63BA9CFE3AA7EB" authorityName="Stal" authorityYear="1866" box="[431,503,1195,1220]" class="Insecta" family="Membracidae" genus="Ennya" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="283" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B912082F3811FFA8FE63BA9CFE3AA7EB" box="[431,503,1195,1220]" italics="true" pageId="14" pageNumber="283">Ennya</emphasis>
|
||
</taxonomicName>
|
||
in the Andes are usually found around
|
||
<quantity id="4C9E79D83811FFA8FC64BA9CFBCEA7EA" box="[936,1027,1195,1221]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="283" unit="m" value="2000.0">2,000 m</quantity>
|
||
(
|
||
<bibRefCitation id="EFF7A9CC3811FFA8FBDCBA9CFB38A7EA" author="Florez-V, C. & Wolff, M. I. & Cardona-Duque, J." box="[1040,1269,1194,1221]" pageId="14" pageNumber="283" pagination="1 - 261" refId="ref12066" refString="Florez-V, C., Wolff, M. I. & Cardona-Duque, J. (2015) Contribution to the taxonomy of the family Membracidae Rafinesque (Hemiptera: Auchenorrhyncha) in Colombia. Zootaxa, 3910 (1), 1 - 261. https: // doi. org / 10.11646 / zootaxa. 3910.1.1" type="journal article" year="2015">
|
||
Flórez-V
|
||
<emphasis id="B912082F3811FFA8FBB1BA9CFB79A7EA" box="[1149,1204,1195,1221]" italics="true" pageId="14" pageNumber="283">et al.</emphasis>
|
||
2015
|
||
</bibRefCitation>
|
||
), so this record is one of the lowest known altitudes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37C87B63811FFA8FF0BBAC4FE60A61E" pageId="14" pageNumber="283" type="etymology">
|
||
<paragraph id="8BD9D43D3811FFA8FF0BBAC4FE60A61E" blockId="14.[151,1437,151,1329]" pageId="14" pageNumber="283">
|
||
<emphasis id="B912082F3811FFA8FF0BBAC4FE85A622" bold="true" box="[199,328,1267,1293]" pageId="14" pageNumber="283">Etymology</emphasis>
|
||
. This species is named after its
|
||
<typeStatus id="54DD6A9F3811FFA8FD7ABAC4FD2BA622" box="[694,742,1267,1293]" pageId="14" pageNumber="283">type</typeStatus>
|
||
locality, La Mariela, a fragment of Western foothill forest in Moraspungo (
|
||
<collectingRegion id="49A21ADF3811FFA8FEF8BB20FE6CA61E" box="[308,417,1303,1329]" country="Ecuador" name="Cotopaxi" pageId="14" pageNumber="283">Cotopaxi</collectingRegion>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |