treatments-xml/data/03/C0/87/03C087FBFFC3C23EFF54A90D3ACFCEAB.xml

<document id="B43C1C374561FD0E777188246C327A6D" ID-DOI="10.35929/RSZ.0108" ID-ISSN="0035-418" ID-Zenodo-Dep="10264611" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1701773453759" checkinUser="felipe" docAuthor="Puechmaille, Sébastien J., Dool, Serena, Beuneux, Gregory &amp; Ruedi, Manuel" docDate="2023" docId="03C087FBFFC3C23EFF54A90D3ACFCEAB" docLanguage="en" docName="RevsuisseZool.130.2.335-351.pdf" docOrigin="Revue suisse de Zoologie 130 (2)" docStyle="DocumentStyle:984CC6410E0DE5B23EAFEE8246BF4C6B.4:RevSuisseZool.2020-.journal_article.0cover" docStyleId="984CC6410E0DE5B23EAFEE8246BF4C6B" docStyleName="RevSuisseZool.2020-.journal_article.0cover" docStyleVersion="3" docTitle="Myotis nustrale Puechmaille &amp; Dool &amp; Beuneux &amp; Ruedi 2023, sp. nov." docType="treatment" docUuid="ACD0B3CB-EE94-423D-9CE7-707B8A675AE5" docUuidSource="ZooBank" docVersion="3" lastPageNumber="346" masterDocId="FFF9FF83FFC4C235FF82AE4C3E7FCB4A" masterDocTitle="Newly described and already endangered: a new mammal species endemic to Corsica" masterLastPageNumber="351" masterPageNumber="335" pageNumber="342" updateTime="1701798505739" updateUser="ExternalLinkService">
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<mods:title id="920406DD3B036F129BCC1AD766270678">Newly described and already endangered: a new mammal species endemic to Corsica</mods:title>
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<mods:namePart id="0214CEE078A3FCEF972A069D8E33369D">Puechmaille, Sébastien J.</mods:namePart>
<mods:affiliation id="D881BB487A10D4349D61250556BBA051">ISEM, University of Montpellier, CNRS, EPHE, IRD, F- 34090 Montpellier, France</mods:affiliation>
<mods:nameIdentifier id="21AE068ACAB796B7AF51DDB83CA5C05C" type="email">sebastien.puechmaille@umontpellier.fr</mods:nameIdentifier>
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<mods:namePart id="A2FDA3C9395237C36E392512E8BCB677">Dool, Serena</mods:namePart>
<mods:affiliation id="FCCA014A84BF483AA846903E15CF302F">Groupe Chiroptères de Midi-Pyrénées (CEN Occitanie-GCMP), F- 31076 Toulouse, France</mods:affiliation>
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<mods:namePart id="7B835766987A84700A7A52E2B1B2D8A5">Beuneux, Gregory</mods:namePart>
<mods:affiliation id="013E5D7B3C7D2E0591B4207108AAAFC8">Groupe Chiroptères Corse, BP 37, F- 20250 Corte, France</mods:affiliation>
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<mods:namePart id="42AF9CEF9C7440DFA0BD7C41E727170B">Ruedi, Manuel</mods:namePart>
<mods:affiliation id="15E176995EDC5008C5C29B190C422B82">Muséum d’histoire naturelle de Genève, C. P. 6434, CH- 1211 Genève 6, Suisse</mods:affiliation>
<mods:nameIdentifier id="454424D69BABD3185D2925EB30DE7BB6" type="email">manuel.ruedi@ville-ge.ch</mods:nameIdentifier>
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<paragraph id="8BD636EDFFC3C232FF54A90D3C74CC30" blockId="7.[214,693,1857,1948]" pageId="7" pageNumber="342">
<heading id="D09E8181FFC3C232FF54A90D3C74CC30" bold="true" centered="true" fontSize="10" level="2" pageId="7" pageNumber="342" reason="2">
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<taxonomicName id="4C694D6EFFC3C232FF54A90D3FFECC13" authority="Puechmaille &amp; Dool &amp; Beuneux &amp; Ruedi, 2023" authorityName="Puechmaille &amp; Dool &amp; Beuneux &amp; Ruedi" authorityYear="2023" box="[214,385,1857,1881]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="7" pageNumber="342" phylum="Chordata" rank="species" species="nustrale" status="sp. nov.">
<emphasis id="B91DEAFFFFC3C232FF54A90D3FFECC13" bold="true" box="[214,385,1857,1881]" italics="true" pageId="7" pageNumber="342">Myotis nustrale</emphasis>
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<taxonomicNameLabel id="A22E5784FFC3C232FE0AA90E3FA2CC10" box="[392,477,1858,1882]" pageId="7" pageNumber="342" rank="species">sp. nov.</taxonomicNameLabel>
Ruedi, Beuneux &amp; Puechmaille
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<paragraph id="8BD636EDFFC3C232FE1BA9C83F8DCCD6" blockId="7.[214,693,1857,1948]" box="[409,498,1924,1948]" pageId="7" pageNumber="342">
<figureCitation id="13522A68FFC3C232FE1BA9C83F8DCCD6" box="[409,498,1924,1948]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="8.[151,186,1011,1033]" captionStartId-1="10.[151,186,1263,1285]" captionStartId-2="11.[151,186,1212,1234]" captionTargetBox-0="[151,1436,261,969]" captionTargetBox-1="[151,1436,257,1218]" captionTargetBox-2="[151,1436,257,1160]" captionTargetId-0="figure-497@8.[151,1436,261,969]" captionTargetId-1="figure-309@10.[151,1436,257,1218]" captionTargetId-2="figure-317@11.[151,1436,257,1160]" captionTargetPageId-0="8" captionTargetPageId-1="10" captionTargetPageId-2="11" captionText-0="Fig. 4. Lateral and occlusal views of the skull and mandible of the holotype female Myotis nustrale sp. nov. (MNHN-ZM-2023-12). The scale is 5 mm." captionText-1="Fig. 5. External characters of Myotis nustrale sp. nov. Panels are close-ups of the female holotype (MNHN-ZM-2023-12), except the lower right one which is a picture from a released adult individual (Photo © Yann Le Bris). The upper left panel illustrates the stiff hairs running along the uropatagium, viewed from below. The upper right panel is the right foot with wing insertion to the base of the outer toe, in ventral view. The lower left panel illustrates the long and straight tragus and unnotched ear of the female holotype. On the portrait of the live individual, notice the black chin spot on the lower lip." captionText-2="Fig. 6. Portraits and hind feet (dorsal view) of an adult female M. nustrale sp. nov. caught near Bavella, in southern Corsica (left) and an adult female M. crypticus from the Jura mountains in Switzerland (right). Notice that the latter individual has a faint line bordering the lower lip, while the former has a much more conspicuous chin spot. Wing insertion to the outer toe is, however, identical in both species. For the left individual, the wing was not stretched while it was stretched for the right individual." figureDoi-0="http://doi.org/10.5281/zenodo.10264621" figureDoi-1="http://doi.org/10.5281/zenodo.10264625" figureDoi-2="http://doi.org/10.5281/zenodo.10264627" httpUri-0="https://zenodo.org/record/10264621/files/figure.png" httpUri-1="https://zenodo.org/record/10264625/files/figure.png" httpUri-2="https://zenodo.org/record/10264627/files/figure.png" pageId="7" pageNumber="342">Figs 4-6</figureCitation>
</paragraph>
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<emphasis id="B91DEAFFFFC3C232FF15A9F83C49CC81" box="[151,566,1972,1996]" italics="true" pageId="7" pageNumber="342">
ZooBank Life Science Identifier (
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):
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urn:lsid:zoobank. org:pub: 
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<subSubSection id="C3736566FFC3C232FF15A6493A89CC37" pageId="7" pageNumber="342" type="materials_examined">
<paragraph id="8BD636EDFFC3C232FF15A6493B71CC07" blockId="7.[151,756,2053,2077]" lastBlockId="7.[832,1437,1080,1869]" pageId="7" pageNumber="342">
<materialsCitation id="3B013CB0FFC3C232FF15A6493B71CC07" collectingDate="2006-07-25" collectionCode="MNHN" collectorName="Julien Barataud" country="France" elevation="1100" latitude="42.27833" location="Albertacce" longitude="8.911944" pageId="7" pageNumber="342" specimenCode="MNHN-ZM-2023-12, MNA008_CO_ALB" specimenCount="1" stateProvince="Haute-Corse" typeStatus="holotype">
<emphasis id="B91DEAFFFFC3C232FF15A6493F75C357" bold="true" box="[151,266,2053,2077]" pageId="7" pageNumber="342">
<typeStatus id="54D2884FFFC3C232FF15A6493F7BC357" box="[151,260,2053,2077]" pageId="7" pageNumber="342" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="9D6FFD64FFC3C232FE99A6493F93C357" box="[283,492,2053,2077]" pageId="7" pageNumber="342">One adult female</specimenCount>
(
<specimenCode id="DBCF9E96FFC3C232FD86A6493C8FC357" box="[516,752,2053,2077]" pageId="7" pageNumber="342">MNHN-ZM-2023-12</specimenCode>
, with field number SP.C.37 and sample number 
<specimenCode id="DBCF9E96FFC3C232FCC2AA143A61CF3A" box="[832,1054,1112,1136]" pageId="7" pageNumber="342">MNA008_CO_ALB</specimenCode>
) found dead by 
<collectorName id="269C533BFFC3C232FB6EAA143BE3CF3A" box="[1260,1436,1112,1136]" pageId="7" pageNumber="342">Julien Barataud</collectorName>
on the 
<collectingDate id="EF93E9C5FFC3C232FC0DAA353A26CFDB" box="[911,1113,1145,1169]" pageId="7" pageNumber="342">
25th of 
<date id="FFD7102DFFC3C232FC6BAA353A26CFDB" box="[1001,1113,1145,1169]" pageId="7" pageNumber="342" value="2006-07">July 2006</date>
</collectingDate>
. The whole carcass has been preserved in alcohol but is currently in a poor state, with large portions of the dorsal and ventral fur missing and the patagium of the left wing torn apart. Epiphyses are fully fused. The skull has been removed and preserved separately; it is intact, except for the right zygomatic arch that is broken (
<figureCitation id="13522A68FFC3C232FBB7AB703A00CE1F" box="[1077,1151,1340,1365]" captionStart="Fig" captionStartId="8.[151,186,1011,1033]" captionTargetBox="[151,1436,261,969]" captionTargetId="figure-497@8.[151,1436,261,969]" captionTargetPageId="8" captionText="Fig. 4. Lateral and occlusal views of the skull and mandible of the holotype female Myotis nustrale sp. nov. (MNHN-ZM-2023-12). The scale is 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10264621" httpUri="https://zenodo.org/record/10264621/files/figure.png" pageId="7" pageNumber="342">Fig. 4</figureCitation>
); teeth are only slightly worn. External measurements (in mm; 
<tableCitation id="C6EB0356FFC3C232FB7CAB113B2BCE3F" box="[1278,1364,1373,1397]" captionStart="Table 2" captionStartId="7.[153,206,257,279]" captionTargetPageId="7" captionText="Table 2. External measurements (mean ± SD, minimum and maximum values) recorded in the four compared species within the M. nattereri species complex. See text for definitions of the variables and acronyms. Besides ears which appear to be slightly longer in Myotis nustrale sp. nov., none of the external characters differ from those measured in the other species." httpUri="http://table.plazi.org/id/DF166665FFC3C232FF1BAF4D3B43CA1B" pageId="7" pageNumber="342" tableUuid="DF166665FFC3C232FF1BAF4D3B43CA1B">Table 2</tableCitation>
) were taken on the fluid-preserved carcass: FA 39.5, HB 42, TL 40; HF 7.8, TIB 17.0, EAR 18.5, TRA 10.9. Skull measurements (in mm; 
<tableCitation id="C6EB0356FFC3C232FBC4ABF33AE7CE9D" box="[1094,1176,1471,1495]" captionStart="Table 3" captionStartId="9.[151,204,257,279]" captionTargetPageId="9" captionText="Table 3. Descriptive statistics (mean ± SD, minimum and maximum values) of 12 cranial and four mandibular variables measured in four species within the M. nattereri species complex. See text for definitions of variables and acronyms. Measurements of the single known skull of Myotis nustrale sp. nov. are very much like those measured in other species from the continent." httpUri="http://table.plazi.org/id/DF166665FFCDC23CFF15AF4D3B20CA1B" pageId="7" pageNumber="342" tableUuid="DF166665FFCDC23CFF15AF4D3B20CA1B">Table 3</tableCitation>
) are: GTL 15.62; CBL 14.49; CCL 13.47; CM3 5.77; M3M3W 6.26; CCW 3.92; ZB 9.64; BCW 7.86; MAB 7.94; POC 3.93; ROL 4.79; M1M3 3.50; ml 11.14; cm3L 6.04; m 
<quantity id="4C919B08FFC3C232FAA2A86D3B3FCD73" box="[1312,1344,1569,1593]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="342" unit="m" value="1.0">1m</quantity>
3L 3.81; coh 3.29. Tissue samples from this specimen are stored in alcohol at -20°C. DNA sequences from this specimen are deposited in the European Nucleotide Archive and include regions of the nuclear introns SLC38A7 (
<accessionNumber id="943AAB0EFFC3C232FCC5A8883DBACD96" box="[839,965,1732,1756]" pageId="7" pageNumber="342">OX958065</accessionNumber>
), ABHD11 (
<accessionNumber id="943AAB0EFFC3C232FB0DA8883B6ECD96" box="[1167,1297,1732,1756]" pageId="7" pageNumber="342">OX958096</accessionNumber>
), ACOX2 (
<accessionNumber id="943AAB0EFFC3C232FCC5A8A83DBECDB6" box="[839,961,1764,1788]" pageId="7" pageNumber="342">OX958129</accessionNumber>
) and ROGDI (
<accessionNumber id="943AAB0EFFC3C232FBDCA8A83AA4CDB6" box="[1118,1243,1764,1788]" pageId="7" pageNumber="342">OX958185</accessionNumber>
). 
<emphasis id="B91DEAFFFFC3C232FCC2A9583D8DCC66" bold="true" box="[832,1010,1812,1836]" pageId="7" pageNumber="342">Type locality:</emphasis>
<location id="8EB66036FFC3C232FB8FA9583AF8CC66" LSID="urn:lsid:plazi:treatment:03C087FBFFC3C23EFF54A90D3ACFCEAB:8EB66036FFC3C232FB8FA9583AF8CC66" box="[1037,1159,1812,1836]" country="France" name="Albertacce" pageId="7" pageNumber="342" stateProvince="Haute-Corse">Albertacce</location>
, 
<collectingRegion id="49ADF80FFFC3C232FB27A9583B4CCC66" box="[1189,1331,1812,1836]" pageId="7" pageNumber="342">Haute-Corse</collectingRegion>
, 
<collectingCountry id="F37E767DFFC3C232FAD3A9593BE4CC66" box="[1361,1435,1813,1836]" name="France" pageId="7" pageNumber="342">France</collectingCountry>
(
<geoCoordinate id="EE5D502AFFC3C232FCCAA9793DB2CC07" box="[840,973,1845,1869]" degrees="42" direction="north" minutes="16" orientation="latitude" pageId="7" pageNumber="342" precision="15" seconds="42" value="42.27833">42°16’42”N</geoCoordinate>
, 
<geoCoordinate id="EE5D502AFFC3C232FC5AA9793A32CC07" box="[984,1101,1845,1869]" degrees="8" direction="east" minutes="54" orientation="longitude" pageId="7" pageNumber="342" precision="15" seconds="43" value="8.911944">8°54’43”E</geoCoordinate>
), ca. 
<quantity id="4C919B08FFC3C232FB04A9793AA9CC06" box="[1158,1238,1845,1869]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="7" pageNumber="342" unit="m" value="1100.0">
<elevation id="0044D1DEFFC3C232FB04A9793AA9CC06" box="[1158,1238,1845,1869]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="7" pageNumber="342" unit="m" value="1100.0">1100 m</elevation>
</quantity>
a.s.l.
</materialsCitation>
</paragraph>
<paragraph id="8BD636EDFFC3C232FCC2A9293A89CC37" blockId="7.[832,1270,1893,1917]" box="[832,1270,1893,1917]" pageId="7" pageNumber="342">
<emphasis id="B91DEAFFFFC3C232FCC2A9293DC4CC37" bold="true" box="[832,955,1893,1917]" pageId="7" pageNumber="342">
<typeStatus id="54D2884FFFC3C232FCC2A9293DC9CC37" box="[832,950,1893,1917]" pageId="7" pageNumber="342" type="paratype">Paratypes</typeStatus>
:
</emphasis>
No 
<typeStatus id="54D2884FFFC3C232FC6AA9293A31CC37" box="[1000,1102,1893,1917]" pageId="7" pageNumber="342" type="paratype">paratypes</typeStatus>
are designated.
</paragraph>
</subSubSection>
<subSubSection id="C3736566FFC3C232FCC2A9D83B17CC87" pageId="7" pageNumber="342" type="distribution">
<paragraph id="8BD636EDFFC3C232FCC2A9D83B17CC87" blockId="7.[832,1436,1940,1997]" pageId="7" pageNumber="342">
<emphasis id="B91DEAFFFFC3C232FCC2A9D83DAACCE6" bold="true" box="[832,981,1940,1964]" pageId="7" pageNumber="342">Distribution:</emphasis>
This species is endemic to the island of 
<collectingCountry id="F37E767DFFC3C232FCC2A9F93DEECC87" box="[832,913,1973,1997]" name="France" pageId="7" pageNumber="342">Corsica</collectingCountry>
(
<collectingCountry id="F37E767DFFC3C232FC1DA9F93D92CC87" box="[927,1005,1973,1997]" name="France" pageId="7" pageNumber="342">France</collectingCountry>
), in the Mediterranean Sea (
<figureCitation id="13522A68FFC3C232FA9BA9F93B27CC87" box="[1305,1368,1973,1997]" captionStart="Fig" captionStartId="3.[831,866,1404,1426]" captionTargetBox="[901,1422,260,1364]" captionTargetId="figure-564@3.[831,1436,254,1373]" captionTargetPageId="3" captionText="Fig. 1. Topographic map of Corsica (France) showing the current records of M. nustrale sp. nov. on this Mediterranean island (Courtois et al., 2011; Groupe Chiroptères Corse, 2019). The black dots represent capture sites, while the red stars indicate the presence of lactating females, including those living in the only known nursery roost of Ghisoni. A double circle indicates the location where the holotype of M.nustrale sp. nov. was found. Shades of grey represent areas above 100 m a.s.l." figureDoi="http://doi.org/10.5281/zenodo.10264613" httpUri="https://zenodo.org/record/10264613/files/figure.png" pageId="7" pageNumber="342">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3736566FFC3C23DFCC2A9A83A33CFA7" lastPageId="8" lastPageNumber="343" pageId="7" pageNumber="342" type="diagnosis">
<paragraph id="8BD636EDFFC3C23DFCC2A9A83F19CE25" blockId="7.[832,1438,2020,2077]" lastBlockId="8.[151,758,1138,2077]" lastPageId="8" lastPageNumber="343" pageId="7" pageNumber="342">
<emphasis id="B91DEAFFFFC3C232FCC2A9A83DC7CCB6" bold="true" box="[832,952,2020,2044]" pageId="7" pageNumber="342">Diagnosis:</emphasis>
Medium sized 
<taxonomicName id="4C694D6EFFC3C232FBF4A9A93AC0CCB6" box="[1142,1215,2021,2044]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="7" pageNumber="342" phylum="Chordata" rank="genus">
<emphasis id="B91DEAFFFFC3C232FBF4A9A93AC0CCB6" box="[1142,1215,2021,2044]" italics="true" pageId="7" pageNumber="342">Myotis</emphasis>
</taxonomicName>
(
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W 
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; FA 
<quantity id="4C919B08FFC3C232FCC2A6493D9FC357" box="[832,992,2053,2077]" metricMagnitude="-2" metricUnit="m" metricValue="3.9849999999999994" metricValueMax="4.13" metricValueMin="3.84" pageId="7" pageNumber="342" unit="mm" value="39.849999999999994" valueMax="41.3" valueMin="38.4">38.4-41.3 mm</quantity>
) with relatively long, unnotched ears; tragus straight, reaching beyond half the length of the conch (
<figureCitation id="13522A68FFCCC23DFF6AAADF3F53CFE0" box="[232,300,1170,1195]" captionStart="Fig" captionStartId="10.[151,186,1263,1285]" captionTargetBox="[151,1436,257,1218]" captionTargetId="figure-309@10.[151,1436,257,1218]" captionTargetPageId="10" captionText="Fig. 5. External characters of Myotis nustrale sp. nov. Panels are close-ups of the female holotype (MNHN-ZM-2023-12), except the lower right one which is a picture from a released adult individual (Photo © Yann Le Bris). The upper left panel illustrates the stiff hairs running along the uropatagium, viewed from below. The upper right panel is the right foot with wing insertion to the base of the outer toe, in ventral view. The lower left panel illustrates the long and straight tragus and unnotched ear of the female holotype. On the portrait of the live individual, notice the black chin spot on the lower lip." figureDoi="http://doi.org/10.5281/zenodo.10264625" httpUri="https://zenodo.org/record/10264625/files/figure.png" pageId="8" pageNumber="343">Fig. 5</figureCitation>
). The long, unkeeled and S-shaped calcar running along the proximal half of the uropatagium, as well as the stiff hairs visible on the trailing edge of the uropatagium are typical features of all taxa in the 
<taxonomicName id="4C694D6EFFCCC23DFD29AAB93E8CCE67" baseAuthorityName="Kuhl" baseAuthorityYear="1817" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="343" phylum="Chordata" rank="species" species="nattereri">
<emphasis id="B91DEAFFFFCCC23DFD29AAB93E8CCE67" italics="true" pageId="8" pageNumber="343">Myotis nattereri</emphasis>
</taxonomicName>
species complex. Stiff hairs of the uropatagium occur in two rows, with only few hairs being reflected backwards (
<figureCitation id="13522A68FFCCC23DFE9AAB1B3F26CE25" box="[280,345,1367,1391]" captionStart="Fig" captionStartId="10.[151,186,1263,1285]" captionTargetBox="[151,1436,257,1218]" captionTargetId="figure-309@10.[151,1436,257,1218]" captionTargetPageId="10" captionText="Fig. 5. External characters of Myotis nustrale sp. nov. Panels are close-ups of the female holotype (MNHN-ZM-2023-12), except the lower right one which is a picture from a released adult individual (Photo © Yann Le Bris). The upper left panel illustrates the stiff hairs running along the uropatagium, viewed from below. The upper right panel is the right foot with wing insertion to the base of the outer toe, in ventral view. The lower left panel illustrates the long and straight tragus and unnotched ear of the female holotype. On the portrait of the live individual, notice the black chin spot on the lower lip." figureDoi="http://doi.org/10.5281/zenodo.10264625" httpUri="https://zenodo.org/record/10264625/files/figure.png" pageId="8" pageNumber="343">Fig. 5</figureCitation>
).
</paragraph>
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<paragraph id="8BD636EDFFCCC23DFF15ADBF3FE6CF6C" blockId="8.[151,1437,1011,1062]" pageId="8" pageNumber="343">
Fig. 4. Lateral and occlusal views of the skull and mandible of the holotype female 
<taxonomicName id="4C694D6EFFCCC23DFC48ADB83A1ECF42" authority="Puechmaille &amp; Dool &amp; Beuneux &amp; Ruedi, 2023" authorityName="Puechmaille &amp; Dool &amp; Beuneux &amp; Ruedi" authorityYear="2023" box="[970,1121,1011,1033]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="343" phylum="Chordata" rank="species" species="nustrale" status="sp. nov.">
<emphasis id="B91DEAFFFFCCC23DFC48ADB83A1ECF42" box="[970,1121,1011,1033]" italics="true" pageId="8" pageNumber="343">Myotis nustrale</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E5784FFCCC23DFBEAADB83ACCCF43" box="[1128,1203,1012,1033]" pageId="8" pageNumber="343" rank="species">sp. nov.</taxonomicNameLabel>
(MNHN-ZM-2023-12). The scale is 5 mm.
</paragraph>
</caption>
<paragraph id="8BD636EDFFCCC23DFF15AB3B3C3FCC52" blockId="8.[151,758,1138,2077]" pageId="8" pageNumber="343">
Wing membrane (plagiopatagium) attaching to the base of the toe, as in 
<emphasis id="B91DEAFFFFCCC23DFE00ABD53C39CEFA" box="[386,582,1433,1456]" italics="true" pageId="8" pageNumber="343">
<taxonomicName id="4C694D6EFFCCC23DFE00ABD53C7ACEFA" baseAuthorityName="Kuhl" baseAuthorityYear="1817" box="[386,517,1433,1456]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="343" phylum="Chordata" rank="species" species="nattereri">M. nattereri</taxonomicName>
<taxonomicNameLabel id="A22E5784FFCCC23DFD8EABD53C39CEFA" box="[524,582,1433,1456]" pageId="8" pageNumber="343" sensu="stricto">s. str.</taxonomicNameLabel>
</emphasis>
or 
<taxonomicName id="4C694D6EFFCCC23DFDEFABD53C89CEFA" box="[621,758,1433,1456]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="343" phylum="Chordata" rank="species" species="crypticus">
<emphasis id="B91DEAFFFFCCC23DFDEFABD53C89CEFA" box="[621,758,1433,1456]" italics="true" pageId="8" pageNumber="343">M. crypticus</emphasis>
</taxonomicName>
(
<figureCitation id="13522A68FFCCC23DFF1DABF53E9ECE9B" box="[159,225,1465,1489]" captionStart="Fig" captionStartId="11.[151,186,1212,1234]" captionTargetBox="[151,1436,257,1160]" captionTargetId="figure-317@11.[151,1436,257,1160]" captionTargetPageId="11" captionText="Fig. 6. Portraits and hind feet (dorsal view) of an adult female M. nustrale sp. nov. caught near Bavella, in southern Corsica (left) and an adult female M. crypticus from the Jura mountains in Switzerland (right). Notice that the latter individual has a faint line bordering the lower lip, while the former has a much more conspicuous chin spot. Wing insertion to the outer toe is, however, identical in both species. For the left individual, the wing was not stretched while it was stretched for the right individual." figureDoi="http://doi.org/10.5281/zenodo.10264627" httpUri="https://zenodo.org/record/10264627/files/figure.png" pageId="8" pageNumber="343">Fig. 6</figureCitation>
), but unlike in 
<taxonomicName id="4C694D6EFFCCC23DFE07ABF63C73CE9B" authorityName="Cabrera" authorityYear="1904" box="[389,524,1465,1489]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="343" phylum="Chordata" rank="species" species="escalerai">
<emphasis id="B91DEAFFFFCCC23DFE07ABF63C73CE9B" box="[389,524,1465,1489]" italics="true" pageId="8" pageNumber="343">M. escalerai</emphasis>
</taxonomicName>
or 
<taxonomicName id="4C694D6EFFCCC23DFDB2ABF63CD4CE9B" box="[560,683,1466,1489]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="343" phylum="Chordata" rank="species" species="zenatius">
<emphasis id="B91DEAFFFFCCC23DFDB2ABF63CD4CE9B" box="[560,683,1466,1489]" italics="true" pageId="8" pageNumber="343">M. zenatius</emphasis>
</taxonomicName>
which attaches to the base of metatarsus (
<bibRefCitation id="EFF84B1CFFCCC23DFDA1AB963EAECD58" author="Puechmaille S. J. &amp; Allegrini B. &amp; Boston E. &amp; Dubourg-Savage M. - J. &amp; Evin A. &amp; Knochel A. &amp; Le Bris Y. &amp; Lecoq V. &amp; Lemaire M. &amp; Rist D. &amp; Teeling E. C." pageId="8" pageNumber="343" pagination="224 - 228" refId="ref13158" refString="Puechmaille S. J., Allegrini B., Boston E., Dubourg-Savage M. - J., Evin A., Knochel A., Le Bris Y., Lecoq V., Lemaire M., Rist D., Teeling E. C. 2012. Genetic analyses reveal further cryptic lineages within the Myotis nattereri species complex. Mammalian Biology 77: 224 - 228." type="journal article" year="2012">
Puechmaille 
<emphasis id="B91DEAFFFFCCC23DFD36AB963C91CEB8" box="[692,750,1498,1522]" italics="true" pageId="8" pageNumber="343">et al.</emphasis>
, 2012
</bibRefCitation>
). Pelage greyish-brown above and whitish below, with a sharp demarcation line running along the sides of the body. Feet relatively small, half or less the length of tibia. Wing and tail membranes essentially naked and ochraceous-brown. Face hairy, except around eyes. Muzzle relatively flat and pointed. A prominent darkpigmented chin spot is visible on the lower lip in adult individuals (
<figureCitation id="13522A68FFCCC23DFE9CA8933F1CCDBD" box="[286,355,1759,1783]" captionStart="Fig" captionStartId="10.[151,186,1263,1285]" captionTargetBox="[151,1436,257,1218]" captionTargetId="figure-309@10.[151,1436,257,1218]" captionTargetPageId="10" captionText="Fig. 5. External characters of Myotis nustrale sp. nov. Panels are close-ups of the female holotype (MNHN-ZM-2023-12), except the lower right one which is a picture from a released adult individual (Photo © Yann Le Bris). The upper left panel illustrates the stiff hairs running along the uropatagium, viewed from below. The upper right panel is the right foot with wing insertion to the base of the outer toe, in ventral view. The lower left panel illustrates the long and straight tragus and unnotched ear of the female holotype. On the portrait of the live individual, notice the black chin spot on the lower lip." figureDoi="http://doi.org/10.5281/zenodo.10264625" httpUri="https://zenodo.org/record/10264625/files/figure.png" pageId="8" pageNumber="343">Figs 5</figureCitation>
and 
<figureCitation id="13522A68FFCCC23DFE19A8933FD6CDBD" box="[411,425,1759,1783]" captionStart="Fig" captionStartId="11.[151,186,1212,1234]" captionTargetBox="[151,1436,257,1160]" captionTargetId="figure-317@11.[151,1436,257,1160]" captionTargetPageId="11" captionText="Fig. 6. Portraits and hind feet (dorsal view) of an adult female M. nustrale sp. nov. caught near Bavella, in southern Corsica (left) and an adult female M. crypticus from the Jura mountains in Switzerland (right). Notice that the latter individual has a faint line bordering the lower lip, while the former has a much more conspicuous chin spot. Wing insertion to the outer toe is, however, identical in both species. For the left individual, the wing was not stretched while it was stretched for the right individual." figureDoi="http://doi.org/10.5281/zenodo.10264627" httpUri="https://zenodo.org/record/10264627/files/figure.png" pageId="8" pageNumber="343">6</figureCitation>
) which distinguish them from all other European 
<taxonomicName id="4C694D6EFFCCC23DFEE1A94D3FD3CC52" box="[355,428,1793,1816]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="343" phylum="Chordata" rank="genus">
<emphasis id="B91DEAFFFFCCC23DFEE1A94D3FD3CC52" box="[355,428,1793,1816]" italics="true" pageId="8" pageNumber="343">Myotis</emphasis>
</taxonomicName>
in this group.
</paragraph>
<paragraph id="8BD636EDFFCCC23DFF15A96C3DC4CFE1" blockId="8.[151,758,1138,2077]" lastBlockId="8.[832,1437,1138,1261]" pageId="8" pageNumber="343">
Skull and dentition (
<figureCitation id="13522A68FFCCC23DFEF2A96C3FCFCC73" box="[368,432,1824,1849]" captionStart="Fig" captionStartId="8.[151,186,1011,1033]" captionTargetBox="[151,1436,261,969]" captionTargetId="figure-497@8.[151,1436,261,969]" captionTargetPageId="8" captionText="Fig. 4. Lateral and occlusal views of the skull and mandible of the holotype female Myotis nustrale sp. nov. (MNHN-ZM-2023-12). The scale is 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10264621" httpUri="https://zenodo.org/record/10264621/files/figure.png" pageId="8" pageNumber="343">Fig. 4</figureCitation>
) like that of other members of the 
<taxonomicName id="4C694D6EFFCCC23DFF3DA90E3F3FCC13" baseAuthorityName="Kuhl" baseAuthorityYear="1817" box="[191,320,1858,1881]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="343" phylum="Chordata" rank="species" species="nattereri">
<emphasis id="B91DEAFFFFCCC23DFF3DA90E3F3FCC13" box="[191,320,1858,1881]" italics="true" pageId="8" pageNumber="343">M. nattereri</emphasis>
</taxonomicName>
species complex (
<collectionCode id="ED78AE28FFCCC23DFD84A90D3C45CC13" box="[518,570,1857,1881]" pageId="8" pageNumber="343">GLS</collectionCode>
<quantity id="4C919B08FFCCC23DFDC3A90D3CCCCC13" box="[577,691,1857,1881]" metricMagnitude="-2" metricUnit="m" metricValue="1.5619999999999998" pageId="8" pageNumber="343" unit="mm" value="15.62">15.62 mm</quantity>
; 
<collectionCode id="ED78AE28FFCCC23DFD3CA90D3C95CC13" box="[702,746,1857,1881]" country="China" name="Chongqing Museum" pageId="8" pageNumber="343">CM</collectionCode>
3 
<quantity id="4C919B08FFCCC23DFF15A92E3E86CC33" box="[151,249,1890,1914]" metricMagnitude="-3" metricUnit="m" metricValue="5.77" pageId="8" pageNumber="343" unit="mm" value="5.77">5.77 mm</quantity>
; ml 
<quantity id="4C919B08FFCCC23DFEA5A92E3FE9CC30" box="[295,406,1890,1914]" metricMagnitude="-2" metricUnit="m" metricValue="1.114" pageId="8" pageNumber="343" unit="mm" value="11.14">11.14 mm</quantity>
). Neurocranium globose without visible crests. The frontal part raises sharply above the orbits and becomes relatively flat near apex. Rostrum nearly flat. Teeth strong, uncrowded, with long canines, third premolars and molars. All premolars in-line with the tooth row; the second premolar (both upper and lower jaw) much smaller than the first (
<figureCitation id="13522A68FFCCC23DFB2EAA3E3A8FCFC0" box="[1196,1264,1138,1162]" captionStart="Fig" captionStartId="8.[151,186,1011,1033]" captionTargetBox="[151,1436,261,969]" captionTargetId="figure-497@8.[151,1436,261,969]" captionTargetPageId="8" captionText="Fig. 4. Lateral and occlusal views of the skull and mandible of the holotype female Myotis nustrale sp. nov. (MNHN-ZM-2023-12). The scale is 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10264621" httpUri="https://zenodo.org/record/10264621/files/figure.png" pageId="8" pageNumber="343">Fig. 4</figureCitation>
). Lower molars myotodont.
</paragraph>
<paragraph id="8BD636EDFFCCC23DFCC2AAF83A33CFA7" blockId="8.[832,1437,1138,1261]" pageId="8" pageNumber="343">As the only examined specimen is a female, the shape of the baculum is unknown.</paragraph>
</subSubSection>
<subSubSection id="C3736566FFCCC23DFCC2AB493DD8CE14" pageId="8" pageNumber="343" type="etymology">
<paragraph id="8BD636EDFFCCC23DFCC2AB493DD8CE14" blockId="8.[832,1436,1284,1374]" pageId="8" pageNumber="343">
<emphasis id="B91DEAFFFFCCC23DFCC2AB493DBBCE57" bold="true" box="[832,964,1285,1309]" pageId="8" pageNumber="343">Etymology:</emphasis>
The specific epithet 
<taxonomicName id="4C694D6EFFCCC23DFB32AB483B77CE56" authorityName="Puechmaille &amp; Dool &amp; Beuneux &amp; Ruedi" authorityYear="2023" box="[1200,1288,1284,1308]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="343" phylum="Chordata" rank="species" species="nustrale">
<emphasis id="B91DEAFFFFCCC23DFB32AB483B77CE56" box="[1200,1288,1284,1308]" italics="true" pageId="8" pageNumber="343">nustrale</emphasis>
</taxonomicName>
is a pronoun in apposition meaning “ours - le notre” in the Corsican language.
</paragraph>
</subSubSection>
<subSubSection id="C3736566FFCCC23FFCC2AB3A3C33CC54" lastPageId="10" lastPageNumber="345" pageId="8" pageNumber="343" type="description">
<paragraph id="8BD636EDFFCCC23FFCC2AB3A3C33CC54" blockId="8.[832,1437,1398,2077]" lastBlockId="10.[151,756,1476,1822]" lastPageId="10" lastPageNumber="345" pageId="8" pageNumber="343">
<emphasis id="B91DEAFFFFCCC23DFCC2AB3A3DB2CEC4" bold="true" box="[832,973,1398,1422]" pageId="8" pageNumber="343">Description:</emphasis>
The new species of 
<taxonomicName id="4C694D6EFFCCC23DFB3BAB3B3B7DCEC4" box="[1209,1282,1399,1422]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="343" phylum="Chordata" rank="genus">
<emphasis id="B91DEAFFFFCCC23DFB3BAB3B3B7DCEC4" box="[1209,1282,1399,1422]" italics="true" pageId="8" pageNumber="343">Myotis</emphasis>
</taxonomicName>
from 
<collectingCountry id="F37E767DFFCCC23DFAC8AB3A3BE3CEC4" box="[1354,1436,1398,1422]" name="France" pageId="8" pageNumber="343">Corsica</collectingCountry>
shares external morphological characters with all other members of the 
<taxonomicName id="4C694D6EFFCCC23DFB8CABF43AB8CE85" baseAuthorityName="Kuhl" baseAuthorityYear="1817" box="[1038,1223,1464,1487]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="343" phylum="Chordata" rank="species" species="nattereri">
<emphasis id="B91DEAFFFFCCC23DFB8CABF43AB8CE85" box="[1038,1223,1464,1487]" italics="true" pageId="8" pageNumber="343">Myotis nattereri</emphasis>
</taxonomicName>
species complex, including a long straight tragus, unnotched ears (
<figureCitation id="13522A68FFCCC23DFACFAB943BF0CEBA" box="[1357,1423,1496,1520]" captionStart="Fig" captionStartId="10.[151,186,1263,1285]" captionTargetBox="[151,1436,257,1218]" captionTargetId="figure-309@10.[151,1436,257,1218]" captionTargetPageId="10" captionText="Fig. 5. External characters of Myotis nustrale sp. nov. Panels are close-ups of the female holotype (MNHN-ZM-2023-12), except the lower right one which is a picture from a released adult individual (Photo © Yann Le Bris). The upper left panel illustrates the stiff hairs running along the uropatagium, viewed from below. The upper right panel is the right foot with wing insertion to the base of the outer toe, in ventral view. The lower left panel illustrates the long and straight tragus and unnotched ear of the female holotype. On the portrait of the live individual, notice the black chin spot on the lower lip." figureDoi="http://doi.org/10.5281/zenodo.10264625" httpUri="https://zenodo.org/record/10264625/files/figure.png" pageId="8" pageNumber="343">Fig. 5</figureCitation>
), a long, S-curved, unkeeled calcar and the presence of stiff hairs along the trailing edge of the uropatagium. However, its wing insertion at the base of the toe (
<figureCitation id="13522A68FFCCC23DFCCAA8173DF0CD39" box="[840,911,1627,1651]" captionStart="Fig" captionStartId="11.[151,186,1212,1234]" captionTargetBox="[151,1436,257,1160]" captionTargetId="figure-317@11.[151,1436,257,1160]" captionTargetPageId="11" captionText="Fig. 6. Portraits and hind feet (dorsal view) of an adult female M. nustrale sp. nov. caught near Bavella, in southern Corsica (left) and an adult female M. crypticus from the Jura mountains in Switzerland (right). Notice that the latter individual has a faint line bordering the lower lip, while the former has a much more conspicuous chin spot. Wing insertion to the outer toe is, however, identical in both species. For the left individual, the wing was not stretched while it was stretched for the right individual." figureDoi="http://doi.org/10.5281/zenodo.10264627" httpUri="https://zenodo.org/record/10264627/files/figure.png" pageId="8" pageNumber="343">Fig. 6</figureCitation>
) and double row of stiff hairs bordering the uropatagium on the ventral side classify the Corsican species into the morpho-group comprising 
<emphasis id="B91DEAFFFFCCC23DFA9BA8D23D07CD9F" italics="true" pageId="8" pageNumber="343">
<taxonomicName id="4C694D6EFFCCC23DFA9BA8D23BE3CDFF" baseAuthorityName="Kuhl" baseAuthorityYear="1817" box="[1305,1436,1694,1717]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="343" phylum="Chordata" rank="species" species="nattereri">M. nattereri</taxonomicName>
<taxonomicNameLabel id="A22E5784FFCCC23DFCC2A8F23D07CD9F" box="[832,888,1726,1749]" pageId="8" pageNumber="343" sensu="stricto">s. str.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="4C694D6EFFCCC23DFC2EA8F23A4DCD9F" box="[940,1074,1726,1749]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="343" phylum="Chordata" rank="species" species="crypticus">
<emphasis id="B91DEAFFFFCCC23DFC2EA8F23A4DCD9F" box="[940,1074,1726,1749]" italics="true" pageId="8" pageNumber="343">M. crypticus</emphasis>
</taxonomicName>
. Morphological comparisons will therefore concentrate on the distinction of 
<taxonomicName id="4C694D6EFFCCC23DFA9FA8933BE3CDBC" authorityName="Puechmaille &amp; Dool &amp; Beuneux &amp; Ruedi" authorityYear="2023" box="[1309,1436,1758,1782]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="343" phylum="Chordata" rank="species" species="nustrale">
<emphasis id="B91DEAFFFFCCC23DFA9FA8933BE3CDBC" box="[1309,1436,1758,1782]" italics="true" pageId="8" pageNumber="343">M. nustrale</emphasis>
</taxonomicName>
from 
<emphasis id="B91DEAFFFFCCC23DFC00A94C3A27CC5D" box="[898,1112,1792,1815]" italics="true" pageId="8" pageNumber="343">
<taxonomicName id="4C694D6EFFCCC23DFC00A94C3A75CC5D" baseAuthorityName="Kuhl" baseAuthorityYear="1817" box="[898,1034,1792,1815]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="343" phylum="Chordata" rank="species" species="nattereri">M. nattereri</taxonomicName>
<taxonomicNameLabel id="A22E5784FFCCC23DFB9AA94C3A27CC5D" box="[1048,1112,1792,1815]" pageId="8" pageNumber="343" sensu="stricto">s. str.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="4C694D6EFFCCC23DFB19A94C3B56CC5D" box="[1179,1321,1792,1815]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="343" phylum="Chordata" rank="species" species="crypticus">
<emphasis id="B91DEAFFFFCCC23DFB19A94C3B56CC5D" box="[1179,1321,1792,1815]" italics="true" pageId="8" pageNumber="343">M. crypticus</emphasis>
</taxonomicName>
, all three being distributed exclusively in Europe and western 
<collectingCountry id="F37E767DFFCCC23DFCC2A90D3DF3CC13" box="[832,908,1857,1881]" name="Turkey" pageId="8" pageNumber="343">Turkey</collectingCountry>
(
<bibRefCitation id="EFF84B1CFFCCC23DFC1FA90D3AF0CC13" author="Coraman E. &amp; Dietz C. &amp; Hempel E. &amp; Ghazaryan A. &amp; Levin E. &amp; Presetnik P. &amp; Zagmajster M. &amp; Mayer F." box="[925,1167,1857,1881]" pageId="8" pageNumber="343" pagination="343 - 354" refId="ref11453" refString="Coraman E., Dietz C., Hempel E., Ghazaryan A., Levin E., Presetnik P., Zagmajster M., Mayer F. 2019. Reticulate evolutionary history of a Western Palaearctic Bat Complex explained by multiple mtDNA introgressions in secondary contacts. Journal of Biogeography 46: 343 - 354." type="journal article" year="2019">
Çoraman 
<emphasis id="B91DEAFFFFCCC23DFB8EA90E3A3ACC13" box="[1036,1093,1857,1881]" italics="true" pageId="8" pageNumber="343">et al.</emphasis>
, 2019
</bibRefCitation>
; 
<bibRefCitation id="EFF84B1CFFCCC23DFB1FA90D3BEECC13" author="Uvizl M. &amp; Benda P." box="[1181,1425,1857,1881]" pageId="8" pageNumber="343" pagination="963 - 977" refId="ref14678" refString="Uvizl M., Benda P. 2021. Diversity and distribution of the Myotis nattereri complex (Chiroptera: Vespertilionidae) in the Middle East: filling the gaps. Mammalian Biology 101: 963 - 977." type="journal article" year="2021">Uvizl &amp; Benda, 2021</bibRefCitation>
). Except for the genetically very distinct but similarsized 
<taxonomicName id="4C694D6EFFCCC23DFC06A9CF3D8FCCD0" authorityName="Harrison" authorityYear="1964" box="[900,1008,1922,1946]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="343" phylum="Chordata" rank="species" species="hoveli">
<emphasis id="B91DEAFFFFCCC23DFC06A9CF3D8FCCD0" box="[900,1008,1922,1946]" italics="true" pageId="8" pageNumber="343">M. hoveli</emphasis>
</taxonomicName>
from the Levant, Asian members of this species complex are indeed larger in most respects (
<bibRefCitation id="EFF84B1CFFCCC23DFCC5A9883A2ECC96" author="Uvizl M. &amp; Benda P." box="[839,1105,1988,2012]" pageId="8" pageNumber="343" pagination="963 - 977" refId="ref14678" refString="Uvizl M., Benda P. 2021. Diversity and distribution of the Myotis nattereri complex (Chiroptera: Vespertilionidae) in the Middle East: filling the gaps. Mammalian Biology 101: 963 - 977." type="journal article" year="2021">Uvizl &amp; Benda, 2021</bibRefCitation>
). Externally, 
<taxonomicName id="4C694D6EFFCCC23DFB71A9893B05CC96" authorityName="Puechmaille &amp; Dool &amp; Beuneux &amp; Ruedi" authorityYear="2023" box="[1267,1402,1988,2012]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="343" phylum="Chordata" rank="species" species="nustrale">
<emphasis id="B91DEAFFFFCCC23DFB71A9893B05CC96" box="[1267,1402,1988,2012]" italics="true" pageId="8" pageNumber="343">M. nustrale</emphasis>
</taxonomicName>
is very similar to 
<emphasis id="B91DEAFFFFCCC23DFC7AA9A93AB2CCB6" box="[1016,1229,2021,2044]" italics="true" pageId="8" pageNumber="343">
<taxonomicName id="4C694D6EFFCCC23DFC7AA9A93AFFCCB6" baseAuthorityName="Kuhl" baseAuthorityYear="1817" box="[1016,1152,2021,2044]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="343" phylum="Chordata" rank="species" species="nattereri">M. nattereri</taxonomicName>
<taxonomicNameLabel id="A22E5784FFCCC23DFB0CA9A93AB2CCB6" box="[1166,1229,2021,2044]" pageId="8" pageNumber="343" sensu="stricto">s. str.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="4C694D6EFFCCC23DFA8DA9A93BE2CCB6" box="[1295,1437,2021,2044]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="343" phylum="Chordata" rank="species" species="crypticus">
<emphasis id="B91DEAFFFFCCC23DFA8DA9A93BE2CCB6" box="[1295,1437,2021,2044]" italics="true" pageId="8" pageNumber="343">M. crypticus</emphasis>
</taxonomicName>
and probably cannot be reliably distinguished from them, except by the presence of a conspicuous blackish spot present on the lower lip of all adult individuals examined in 
<collectingCountry id="F37E767DFFCDC23CFEA5A8D13F07CDFF" box="[295,376,1693,1717]" name="France" pageId="9" pageNumber="344">Corsica</collectingCountry>
(
<figureCitation id="13522A68FFCDC23CFE0BA8D13FB0CDFF" box="[393,463,1693,1717]" captionStart="Fig" captionStartId="10.[151,186,1263,1285]" captionTargetBox="[151,1436,257,1218]" captionTargetId="figure-309@10.[151,1436,257,1218]" captionTargetPageId="10" captionText="Fig. 5. External characters of Myotis nustrale sp. nov. Panels are close-ups of the female holotype (MNHN-ZM-2023-12), except the lower right one which is a picture from a released adult individual (Photo © Yann Le Bris). The upper left panel illustrates the stiff hairs running along the uropatagium, viewed from below. The upper right panel is the right foot with wing insertion to the base of the outer toe, in ventral view. The lower left panel illustrates the long and straight tragus and unnotched ear of the female holotype. On the portrait of the live individual, notice the black chin spot on the lower lip." figureDoi="http://doi.org/10.5281/zenodo.10264625" httpUri="https://zenodo.org/record/10264625/files/figure.png" pageId="9" pageNumber="344">Figs 5</figureCitation>
and 
<figureCitation id="13522A68FFCDC23CFD8BA8D13C66CDFF" box="[521,537,1693,1717]" captionStart="Fig" captionStartId="11.[151,186,1212,1234]" captionTargetBox="[151,1436,257,1160]" captionTargetId="figure-317@11.[151,1436,257,1160]" captionTargetPageId="11" captionText="Fig. 6. Portraits and hind feet (dorsal view) of an adult female M. nustrale sp. nov. caught near Bavella, in southern Corsica (left) and an adult female M. crypticus from the Jura mountains in Switzerland (right). Notice that the latter individual has a faint line bordering the lower lip, while the former has a much more conspicuous chin spot. Wing insertion to the outer toe is, however, identical in both species. For the left individual, the wing was not stretched while it was stretched for the right individual." figureDoi="http://doi.org/10.5281/zenodo.10264627" httpUri="https://zenodo.org/record/10264627/files/figure.png" pageId="9" pageNumber="344">6</figureCitation>
). Such a pigmented spot is present in the lower lip of young individuals of a few other 
<taxonomicName id="4C694D6EFFCDC23CFEC0A8923F9DCDBF" box="[322,482,1757,1781]" pageId="9" pageNumber="344">
<emphasis id="B91DEAFFFFCDC23CFEC0A8923FF4CDBF" box="[322,395,1758,1781]" italics="true" pageId="9" pageNumber="344">Myotis</emphasis>
species
</taxonomicName>
(e.g. in 
<taxonomicName id="4C694D6EFFCDC23CFDC4A8923C92CDBF" baseAuthorityName="Kuhl" baseAuthorityYear="1817" box="[582,749,1757,1781]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="9" pageNumber="344" phylum="Chordata" rank="species" species="daubentonii">
<emphasis id="B91DEAFFFFCDC23CFDC4A8923C92CDBF" box="[582,749,1757,1781]" italics="true" pageId="9" pageNumber="344">M. daubentonii</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C694D6EFFCDC23CFF15A8B23F4ECC5F" baseAuthorityName="Boie" baseAuthorityYear="1825" box="[151,305,1789,1813]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="9" pageNumber="344" phylum="Chordata" rank="species" species="dasycneme">
<emphasis id="B91DEAFFFFCDC23CFF15A8B23F4ECC5F" box="[151,305,1789,1813]" italics="true" pageId="9" pageNumber="344">M. dasycneme</emphasis>
</taxonomicName>
) but vanishes when they attain adulthood (
<bibRefCitation id="EFF84B1CFFCDC23CFF1CA9513F15CC7F" author="Richardson P. W." box="[158,362,1821,1845]" pageId="9" pageNumber="344" pagination="307 - 309" refId="ref13633" refString="Richardson P. W. 1994. A new method of distinguishing Daubenton's bats (Myotis daubentonii) up to one year old from adults. Journal of Zoology 233: 307 - 309." type="journal article" year="1994">Richardson, 1994</bibRefCitation>
; 
<bibRefCitation id="EFF84B1CFFCDC23CFEFCA9523C96CC7F" author="Haarsma A. - J. &amp; Van Alphen J." box="[382,745,1821,1845]" pageId="9" pageNumber="344" pagination="97 - 107" refId="ref12055" refString="Haarsma A. - J., Van Alphen J. 2009. Chin-spot as an indicator of age in pond bats. Lutra 52: 97 - 107." type="journal article" year="2009">Haarsma &amp; Van Alphen, 2009</bibRefCitation>
). 
<taxonomicName id="4C694D6EFFCDC23CFF15A9723F5FCC1F" authorityName="Puechmaille &amp; Dool &amp; Beuneux &amp; Ruedi" authorityYear="2023" box="[151,288,1853,1877]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="9" pageNumber="344" phylum="Chordata" rank="species" species="nustrale">
<emphasis id="B91DEAFFFFCDC23CFF15A9723F5FCC1F" box="[151,288,1853,1877]" italics="true" pageId="9" pageNumber="344">M. nustrale</emphasis>
</taxonomicName>
has relatively longer ears and tragus than 
<taxonomicName id="4C694D6EFFCDC23CFF57A9123F20CC3F" baseAuthorityName="Kuhl" baseAuthorityYear="1817" box="[213,351,1886,1909]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="9" pageNumber="344" phylum="Chordata" rank="species" species="nattereri">
<emphasis id="B91DEAFFFFCDC23CFF57A9123F20CC3F" box="[213,351,1886,1909]" italics="true" pageId="9" pageNumber="344">M. nattereri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E5784FFCDC23CFEECA9123FCFCC3F" box="[366,432,1886,1909]" pageId="9" pageNumber="344" sensu="stricto">s. str.</taxonomicNameLabel>
or 
<taxonomicName id="4C694D6EFFCDC23CFE66A9123C0CCC3F" box="[484,627,1886,1909]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="9" pageNumber="344" phylum="Chordata" rank="species" species="crypticus">
<emphasis id="B91DEAFFFFCDC23CFE66A9123C0CCC3F" box="[484,627,1886,1909]" italics="true" pageId="9" pageNumber="344">M. crypticus</emphasis>
</taxonomicName>
(
<tableCitation id="C6EB0356FFCDC23CFD08A9113C99CC3F" box="[650,742,1885,1909]" captionStart="Table 2" captionStartId="7.[153,206,257,279]" captionTargetPageId="7" captionText="Table 2. External measurements (mean ± SD, minimum and maximum values) recorded in the four compared species within the M. nattereri species complex. See text for definitions of the variables and acronyms. Besides ears which appear to be slightly longer in Myotis nustrale sp. nov., none of the external characters differ from those measured in the other species." httpUri="http://table.plazi.org/id/DF166665FFC3C232FF1BAF4D3B43CA1B" pageId="9" pageNumber="344" tableUuid="DF166665FFC3C232FF1BAF4D3B43CA1B">Table 2</tableCitation>
), but because a single individual has been measured for these characters so far, individual variation may blur these apparent differences. The same applies to the shape of the tragus, the margins of which appear slightly more linear in the Corsican species than in the other continental taxa; tragus shape in these taxa, however, are subject to substantial individual variation. Fur colouration, greyish-brown above and whitish below, is also similar in all three species. Although craniodental dimensions of all three species are again largely overlapping (
<tableCitation id="C6EB0356FFCDC23CFB24A8B13B7FCC5F" box="[1190,1280,1789,1813]" captionStart="Table 3" captionStartId="9.[151,204,257,279]" captionTargetPageId="9" captionText="Table 3. Descriptive statistics (mean ± SD, minimum and maximum values) of 12 cranial and four mandibular variables measured in four species within the M. nattereri species complex. See text for definitions of variables and acronyms. Measurements of the single known skull of Myotis nustrale sp. nov. are very much like those measured in other species from the continent." httpUri="http://table.plazi.org/id/DF166665FFCDC23CFF15AF4D3B20CA1B" pageId="9" pageNumber="344" tableUuid="DF166665FFCDC23CFF15AF4D3B20CA1B">Table 3</tableCitation>
), the general shape of the skull of 
<taxonomicName id="4C694D6EFFCDC23CFBBCA9523ABFCC7F" authorityName="Puechmaille &amp; Dool &amp; Beuneux &amp; Ruedi" authorityYear="2023" box="[1086,1216,1821,1845]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="9" pageNumber="344" phylum="Chordata" rank="species" species="nustrale">
<emphasis id="B91DEAFFFFCDC23CFBBCA9523ABFCC7F" box="[1086,1216,1821,1845]" italics="true" pageId="9" pageNumber="344">M. nustrale</emphasis>
</taxonomicName>
is more similar to that of 
<taxonomicName id="4C694D6EFFCDC23CFC1EA9723A55CC1F" box="[924,1066,1854,1877]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="9" pageNumber="344" phylum="Chordata" rank="species" species="crypticus">
<emphasis id="B91DEAFFFFCDC23CFC1EA9723A55CC1F" box="[924,1066,1854,1877]" italics="true" pageId="9" pageNumber="344">M. crypticus</emphasis>
</taxonomicName>
than to the slightly larger and more robust skull of 
<taxonomicName id="4C694D6EFFCDC23CFBC0A9123AB3CC3F" baseAuthorityName="Kuhl" baseAuthorityYear="1817" box="[1090,1228,1886,1909]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="9" pageNumber="344" phylum="Chordata" rank="species" species="nattereri">
<emphasis id="B91DEAFFFFCDC23CFBC0A9123AB3CC3F" box="[1090,1228,1886,1909]" italics="true" pageId="9" pageNumber="344">M. nattereri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E5784FFCDC23CFB59A9123B63CC3F" box="[1243,1308,1886,1909]" pageId="9" pageNumber="344" sensu="stricto">s. str.</taxonomicNameLabel>
Sequences of the mitochondrial 
<emphasis id="B91DEAFFFFCDC23CFBB9A9313AACCCDF" box="[1083,1235,1917,1941]" italics="true" pageId="9" pageNumber="344">cytochrome b</emphasis>
gene, forming a strongly supported monophyletic group, are unique in the Corsican species and diverge by an average 7.5 % Kimura-2-parameter distance from its closest relative 
<taxonomicName id="4C694D6EFFCDC23CFCBDA9B23A8BC35F" authority="(Puechmaille et al., 2012)" baseAuthorityName="Puechmaille" baseAuthorityYear="2012" box="[831,1268,2045,2069]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="9" pageNumber="344" phylum="Chordata" rank="species" species="crypticus">
<emphasis id="B91DEAFFFFCDC23CFCBDA9B23DB7C35F" box="[831,968,2046,2069]" italics="true" pageId="9" pageNumber="344">M. crypticus</emphasis>
(
<bibRefCitation id="EFF84B1CFFCDC23CFC5BA9B13A93C35F" author="Puechmaille S. J. &amp; Allegrini B. &amp; Boston E. &amp; Dubourg-Savage M. - J. &amp; Evin A. &amp; Knochel A. &amp; Le Bris Y. &amp; Lecoq V. &amp; Lemaire M. &amp; Rist D. &amp; Teeling E. C." box="[985,1260,2045,2069]" pageId="9" pageNumber="344" pagination="224 - 228" refId="ref13158" refString="Puechmaille S. J., Allegrini B., Boston E., Dubourg-Savage M. - J., Evin A., Knochel A., Le Bris Y., Lecoq V., Lemaire M., Rist D., Teeling E. C. 2012. Genetic analyses reveal further cryptic lineages within the Myotis nattereri species complex. Mammalian Biology 77: 224 - 228." type="journal article" year="2012">
Puechmaille 
<emphasis id="B91DEAFFFFCDC23CFBEBA9B23ADDC35F" box="[1129,1186,2045,2069]" italics="true" pageId="9" pageNumber="344">et al.</emphasis>
, 2012
</bibRefCitation>
)
</taxonomicName>
. Nuclear genes and microsatellite data for 
<taxonomicName id="4C694D6EFFCEC23FFE61AB893C15CE96" authorityName="Puechmaille &amp; Dool &amp; Beuneux &amp; Ruedi" authorityYear="2023" box="[483,618,1476,1500]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="345" phylum="Chordata" rank="species" species="nustrale">
<emphasis id="B91DEAFFFFCEC23FFE61AB893C15CE96" box="[483,618,1476,1500]" italics="true" pageId="10" pageNumber="345">M. nustrale</emphasis>
</taxonomicName>
also differ considerably from other taxa in the 
<taxonomicName id="4C694D6EFFCEC23FFD9FABA93CE1CEB6" baseAuthorityName="Kuhl" baseAuthorityYear="1817" box="[541,670,1509,1532]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="345" phylum="Chordata" rank="species" species="nattereri">
<emphasis id="B91DEAFFFFCEC23FFD9FABA93CE1CEB6" box="[541,670,1509,1532]" italics="true" pageId="10" pageNumber="345">M. nattereri</emphasis>
</taxonomicName>
species complex (
<figureCitation id="13522A68FFCEC23FFE85A8493F30CD57" box="[263,335,1541,1565]" captionStart="Fig" captionStartId="5.[151,186,1644,1666]" captionTargetBox="[157,1436,261,1609]" captionTargetId="figure-147@5.[151,1436,261,1609]" captionTargetPageId="5" captionText="Fig. 2. Bayesian topology based on five partitioned nuclear introns (SLC38A7, ABHD11, ACOX2, COPS7A and ROGDI). The outgroup (Pipistrellus pipistrellus) is not shown. Bayesian posterior probabilities (BPP) &lt;60 are not shown, neither are within species BPP values. The holotype of Myotis nustrale sp. nov. (sample number MNA008_CO_ALB) is indicated by a white arrow." figureDoi="http://doi.org/10.5281/zenodo.10264615" httpUri="https://zenodo.org/record/10264615/files/figure.png" pageId="10" pageNumber="345">Figs 2</figureCitation>
and 
<figureCitation id="13522A68FFCEC23FFE0CA8493FE4CD57" box="[398,411,1541,1565]" captionStart="Fig" captionStartId="6.[151,186,1632,1654]" captionTargetBox="[151,1436,947,1588]" captionTargetId="figure-348@6.[151,1436,947,1588]" captionTargetPageId="6" captionText="Fig. 3. PCA showing (A) the clear separation between M. crypticus (Mcry; green circles) and M. nustrale sp. nov. individuals (Mnus; blue squares), and (B), the differentiation between populations within M. nustrale sp. nov. Horizontal and vertical axis explain 23.9 and 8.5% of the variance, respectively, in (A) and 17.1 and 13.9% of the variance in (B)." figureDoi="http://doi.org/10.5281/zenodo.10264619" httpUri="https://zenodo.org/record/10264619/files/figure.png" pageId="10" pageNumber="345">3</figureCitation>
) and suggest that 
<taxonomicName id="4C694D6EFFCEC23FFDF1A8493C8BCD57" authorityName="Puechmaille &amp; Dool &amp; Beuneux &amp; Ruedi" authorityYear="2023" box="[627,756,1541,1565]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="345" phylum="Chordata" rank="species" species="nustrale">
<emphasis id="B91DEAFFFFCEC23FFDF1A8493C8BCD57" box="[627,756,1541,1565]" italics="true" pageId="10" pageNumber="345">M. nustrale</emphasis>
</taxonomicName>
may be basal to other members of this clade (
<figureCitation id="13522A68FFCEC23FFD23A8693C98CD77" box="[673,743,1573,1597]" captionStart="Fig" captionStartId="5.[151,186,1644,1666]" captionTargetBox="[157,1436,261,1609]" captionTargetId="figure-147@5.[151,1436,261,1609]" captionTargetPageId="5" captionText="Fig. 2. Bayesian topology based on five partitioned nuclear introns (SLC38A7, ABHD11, ACOX2, COPS7A and ROGDI). The outgroup (Pipistrellus pipistrellus) is not shown. Bayesian posterior probabilities (BPP) &lt;60 are not shown, neither are within species BPP values. The holotype of Myotis nustrale sp. nov. (sample number MNA008_CO_ALB) is indicated by a white arrow." figureDoi="http://doi.org/10.5281/zenodo.10264615" httpUri="https://zenodo.org/record/10264615/files/figure.png" pageId="10" pageNumber="345">Fig. 2</figureCitation>
). Recordings of echolocation calls of 
<taxonomicName id="4C694D6EFFCEC23FFDC8A80A3CAFCD17" authorityName="Puechmaille &amp; Dool &amp; Beuneux &amp; Ruedi" authorityYear="2023" box="[586,720,1605,1629]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="345" phylum="Chordata" rank="species" species="nustrale">
<emphasis id="B91DEAFFFFCEC23FFDC8A80A3CAFCD17" box="[586,720,1605,1629]" italics="true" pageId="10" pageNumber="345">M. nustrale</emphasis>
</taxonomicName>
in 
<collectingCountry id="F37E767DFFCEC23FFF15A8293E97CD37" box="[151,232,1637,1661]" name="France" pageId="10" pageNumber="345">Corsica</collectingCountry>
show the typical acoustic signature of members of the 
<taxonomicName id="4C694D6EFFCEC23FFF64A8CA3F13CDD7" baseAuthorityName="Kuhl" baseAuthorityYear="1817" box="[230,364,1670,1693]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="345" phylum="Chordata" rank="species" species="nattereri">
<emphasis id="B91DEAFFFFCEC23FFF64A8CA3F13CDD7" box="[230,364,1670,1693]" italics="true" pageId="10" pageNumber="345">M. nattereri</emphasis>
</taxonomicName>
species complex (
<bibRefCitation id="EFF84B1CFFCEC23FFDC0A8C93EAFCDF7" author="Russo D. &amp; Jones G." pageId="10" pageNumber="345" pagination="91 - 103" refId="ref13941" refString="Russo D., Jones G. 2002. Identification of twenty-two bat species (Mammalia: Chiroptera) from Italy by analysis of time-expanded recordings of echolocation calls. Journal of Zoology, London 258: 91 - 103." type="journal article" year="2002">Russo &amp; Jones, 2002</bibRefCitation>
), i.e. short and strongly frequency modulated calls dominated by sweeps ranging from&gt;105 kHz down to ca. 25 kHz frequencies, but again these calls are too plastic to serve for species identification.
</paragraph>
</subSubSection>
<caption id="DF166665FFCDC23CFF15AF4D3B20CA1B" ID-Table-UUID="DF166665FFCDC23CFF15AF4D3B20CA1B" httpUri="http://table.plazi.org/id/DF166665FFCDC23CFF15AF4D3B20CA1B" pageId="9" pageNumber="344" startId="9.[151,204,257,279]" targetBox="[161,1373,382,1522]" targetIsTable="true" targetPageId="9" targetType="table">
<paragraph id="8BD636EDFFCDC23CFF15AF4D3B20CA1B" blockId="9.[151,1437,257,337]" pageId="9" pageNumber="344">
Table 3. Descriptive statistics (mean ± SD, minimum and maximum values) of 12 cranial and four mandibular variables measured in four species within the 
<taxonomicName id="4C694D6EFFCDC23CFE50AF533C3ACA7E" baseAuthorityName="Kuhl" baseAuthorityYear="1817" box="[466,581,287,308]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="9" pageNumber="344" phylum="Chordata" rank="species" species="nattereri">
<emphasis id="B91DEAFFFFCDC23CFE50AF533C3ACA7E" box="[466,581,287,308]" italics="true" pageId="9" pageNumber="344">M. nattereri</emphasis>
</taxonomicName>
species complex. See text for definitions of variables and acronyms. Measurements of the single known skull of 
<taxonomicName id="4C694D6EFFCDC23CFE49AF703C1ECA1A" authority="Puechmaille &amp; Dool &amp; Beuneux &amp; Ruedi, 2023" authorityName="Puechmaille &amp; Dool &amp; Beuneux &amp; Ruedi" authorityYear="2023" box="[459,609,315,337]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="9" pageNumber="344" phylum="Chordata" rank="species" species="nustrale" status="sp. nov.">
<emphasis id="B91DEAFFFFCDC23CFE49AF703C1ECA1A" box="[459,609,315,337]" italics="true" pageId="9" pageNumber="344">Myotis nustrale</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E5784FFCDC23CFDE5AF773CCDCA1A" box="[615,690,315,336]" pageId="9" pageNumber="344" rank="species">sp. nov.</taxonomicNameLabel>
are very much like those measured in other species from the continent.
</paragraph>
</caption>
<paragraph id="8BD636EDFFCDC23CFF20AF323B2ECEB8" pageId="9" pageNumber="344">
<table id="F969C44DFFCD3DCAFF23AF323B22CEB8" box="[161,1373,382,1522]" gridcols="5" gridrows="17" pageId="9" pageNumber="344">
<tr id="355934AFFFCD3DCAFF23AF323B22CAFB" box="[161,1373,382,433]" gridrow="0" pageId="9" pageNumber="344">
<th id="76885DD3FFCD3DCAFF23AF323F61CAFB" box="[161,286,382,433]" gridcol="0" gridrow="0" pageId="9" pageNumber="344">Craniodental variable</th>
<th id="76885DD3FFCD3DCAFE1EAF323C40CAFB" box="[412,575,382,433]" gridcol="1" gridrow="0" pageId="9" pageNumber="344">
<taxonomicName id="4C694D6EFFCDC23CFE20AF333C46CAD9" authority="Puechmaille &amp; Dool &amp; Beuneux &amp; Ruedi, 2023" authorityName="Puechmaille &amp; Dool &amp; Beuneux &amp; Ruedi" authorityYear="2023" box="[418,569,382,404]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="9" pageNumber="344" phylum="Chordata" rank="species" species="nustrale" status="sp. nov.">
<emphasis id="B91DEAFFFFCDC23CFE20AF333C46CAD9" box="[418,569,382,404]" italics="true" pageId="9" pageNumber="344">Myotis nustrale</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E5784FFCDC23CFE1EAFD03F99CAFB" box="[412,486,412,433]" pageId="9" pageNumber="344" rank="species">sp. nov.</taxonomicNameLabel>
holotype
</th>
<th id="76885DD3FFCD3DCAFD32AF323D3BCAFB" box="[688,836,382,433]" gridcol="2" gridrow="0" pageId="9" pageNumber="344">
<taxonomicName id="4C694D6EFFCDC23CFD3CAF333D49CADE" box="[702,822,383,404]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="9" pageNumber="344" phylum="Chordata" rank="species" species="crypticus">
<emphasis id="B91DEAFFFFCDC23CFD3CAF333D49CADE" box="[702,822,383,404]" italics="true" pageId="9" pageNumber="344">M. crypticus</emphasis>
</taxonomicName>
(n = 21)
</th>
<th id="76885DD3FFCD3DCAFC33AF323A22CAFB" box="[945,1117,382,433]" gridcol="3" gridrow="0" pageId="9" pageNumber="344">
<emphasis id="B91DEAFFFFCDC23CFC33AF333A22CADE" box="[945,1117,383,404]" italics="true" pageId="9" pageNumber="344">
<taxonomicName id="4C694D6EFFCDC23CFC33AF333A5BCADE" baseAuthorityName="Kuhl" baseAuthorityYear="1817" box="[945,1060,383,404]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="9" pageNumber="344" phylum="Chordata" rank="species" species="nattereri">M. nattereri</taxonomicName>
<taxonomicNameLabel id="A22E5784FFCDC23CFBA8AF333A22CADE" box="[1066,1117,383,404]" pageId="9" pageNumber="344" sensu="stricto">s. str.</taxonomicNameLabel>
</emphasis>
(n = 14)
</th>
<th id="76885DD3FFCD3DCAFB4BAF323B22CAFB" box="[1225,1373,382,433]" gridcol="4" gridrow="0" pageId="9" pageNumber="344">
<taxonomicName id="4C694D6EFFCDC23CFB55AF333B30CAD9" authorityName="Cabrera" authorityYear="1904" box="[1239,1359,382,404]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="9" pageNumber="344" phylum="Chordata" rank="species" species="escalerai">
<emphasis id="B91DEAFFFFCDC23CFB55AF333B30CAD9" box="[1239,1359,382,404]" italics="true" pageId="9" pageNumber="344">M. escalerai</emphasis>
</taxonomicName>
(n = 12)
</th>
</tr>
<tr id="355934AFFFCD3DCAFF23AF8E3B22CABF" box="[161,1373,450,501]" gridrow="1" pageId="9" pageNumber="344">
<th id="76885DD3FFCD3DCAFF23AF8E3F61CABF" box="[161,286,450,501]" gridcol="0" gridrow="1" pageId="9" pageNumber="344">GTL</th>
<td id="76885DD3FFCD3DCAFE1EAF8E3C40CABF" box="[412,575,450,501]" gridcol="1" gridrow="1" pageId="9" pageNumber="344">15.62</td>
<td id="76885DD3FFCD3DCAFD32AF8E3D3BCABF" box="[688,836,450,501]" gridcol="2" gridrow="1" pageId="9" pageNumber="344">15.53 ± 0.34 (14.99 – 16.44)</td>
<td id="76885DD3FFCD3DCAFC33AF8E3A22CABF" box="[945,1117,450,501]" gridcol="3" gridrow="1" pageId="9" pageNumber="344">15.65 ± 0.27 (15.09 – 16.03)</td>
<td id="76885DD3FFCD3DCAFB4BAF8E3B22CABF" box="[1225,1373,450,501]" gridcol="4" gridrow="1" pageId="9" pageNumber="344">15.59 ± 0.66 (15.21 – 15.95)</td>
</tr>
<tr id="355934AFFFCD3DCAFF23AC4A3B22C973" box="[161,1373,518,569]" gridrow="2" pageId="9" pageNumber="344">
<th id="76885DD3FFCD3DCAFF23AC4A3F61C973" box="[161,286,518,569]" gridcol="0" gridrow="2" pageId="9" pageNumber="344">CBL</th>
<td id="76885DD3FFCD3DCAFE1EAC4A3C40C973" box="[412,575,518,569]" gridcol="1" gridrow="2" pageId="9" pageNumber="344">14.49</td>
<td id="76885DD3FFCD3DCAFD32AC4A3D3BC973" box="[688,836,518,569]" gridcol="2" gridrow="2" pageId="9" pageNumber="344">14.46 ± 0.31 (13.94 – 15.20)</td>
<td id="76885DD3FFCD3DCAFC33AC4A3A22C973" box="[945,1117,518,569]" gridcol="3" gridrow="2" pageId="9" pageNumber="344">14.70 ± 0.32 (14.00 – 15.12)</td>
<td id="76885DD3FFCD3DCAFB4BAC4A3B22C973" box="[1225,1373,518,569]" gridcol="4" gridrow="2" pageId="9" pageNumber="344">14.52 ± 0.28 (14.17 – 14.91)</td>
</tr>
<tr id="355934AFFFCD3DCAFF23AC063B22C937" box="[161,1373,586,637]" gridrow="3" pageId="9" pageNumber="344">
<th id="76885DD3FFCD3DCAFF23AC063F61C937" box="[161,286,586,637]" gridcol="0" gridrow="3" pageId="9" pageNumber="344">CCL</th>
<td id="76885DD3FFCD3DCAFE1EAC063C40C937" box="[412,575,586,637]" gridcol="1" gridrow="3" pageId="9" pageNumber="344">13.47</td>
<td id="76885DD3FFCD3DCAFD32AC063D3BC937" box="[688,836,586,637]" gridcol="2" gridrow="3" pageId="9" pageNumber="344">13.52 ± 0.30 (12.90 – 14.12)</td>
<td id="76885DD3FFCD3DCAFC33AC063A22C937" box="[945,1117,586,637]" gridcol="3" gridrow="3" pageId="9" pageNumber="344">13.72 ± 0.30 (13.05 – 14.14)</td>
<td id="76885DD3FFCD3DCAFB4BAC063B22C937" box="[1225,1373,586,637]" gridcol="4" gridrow="3" pageId="9" pageNumber="344">13.58 ± 0.20 (13.27 – 13.96)</td>
</tr>
<tr id="355934AFFFCD3DCAFF23ACC23B22C98B" box="[161,1373,654,705]" gridrow="4" pageId="9" pageNumber="344">
<th id="76885DD3FFCD3DCAFF23ACC23F61C98B" box="[161,286,654,705]" gridcol="0" gridrow="4" pageId="9" pageNumber="344">CM3</th>
<td id="76885DD3FFCD3DCAFE1EACC23C40C98B" box="[412,575,654,705]" gridcol="1" gridrow="4" pageId="9" pageNumber="344">5.77</td>
<td id="76885DD3FFCD3DCAFD32ACC23D3BC98B" box="[688,836,654,705]" gridcol="2" gridrow="4" pageId="9" pageNumber="344">6.00 ± 0.03 (5.64 – 6.33)</td>
<td id="76885DD3FFCD3DCAFC33ACC23A22C98B" box="[945,1117,654,705]" gridcol="3" gridrow="4" pageId="9" pageNumber="344">6.13 ± 0.14 (5.86 – 6.38)</td>
<td id="76885DD3FFCD3DCAFB4BACC23B22C98B" box="[1225,1373,654,705]" gridcol="4" gridrow="4" pageId="9" pageNumber="344">5.99 ± 0.04 (5.77 – 6.27)</td>
</tr>
<tr id="355934AFFFCD3DCAFF23AC9E3B22C84F" box="[161,1373,722,773]" gridrow="5" pageId="9" pageNumber="344">
<th id="76885DD3FFCD3DCAFF23AC9E3F61C84F" box="[161,286,722,773]" gridcol="0" gridrow="5" pageId="9" pageNumber="344">M3M3W</th>
<td id="76885DD3FFCD3DCAFE1EAC9E3C40C84F" box="[412,575,722,773]" gridcol="1" gridrow="5" pageId="9" pageNumber="344">6.26</td>
<td id="76885DD3FFCD3DCAFD32AC9E3D3BC84F" box="[688,836,722,773]" gridcol="2" gridrow="5" pageId="9" pageNumber="344">6.22 ± 0.03 (5.92 – 6.44)</td>
<td id="76885DD3FFCD3DCAFC33AC9E3A22C84F" box="[945,1117,722,773]" gridcol="3" gridrow="5" pageId="9" pageNumber="344">6.47 ± 0.15 (6.22 – 6.73)</td>
<td id="76885DD3FFCD3DCAFB4BAC9E3B22C84F" box="[1225,1373,722,773]" gridcol="4" gridrow="5" pageId="9" pageNumber="344">6.23 ± 0.05 (5.93 – 6.53)</td>
</tr>
<tr id="355934AFFFCD3DCAFF23AD5B3B22C800" box="[161,1373,791,842]" gridrow="6" pageId="9" pageNumber="344">
<th id="76885DD3FFCD3DCAFF23AD5B3F61C800" box="[161,286,791,842]" gridcol="0" gridrow="6" pageId="9" pageNumber="344">CCW</th>
<td id="76885DD3FFCD3DCAFE1EAD5B3C40C800" box="[412,575,791,842]" gridcol="1" gridrow="6" pageId="9" pageNumber="344">3.92</td>
<td id="76885DD3FFCD3DCAFD32AD5B3D3BC800" box="[688,836,791,842]" gridcol="2" gridrow="6" pageId="9" pageNumber="344">3.90 ± 0.03 (3.57 – 4.19)</td>
<td id="76885DD3FFCD3DCAFC33AD5B3A22C800" box="[945,1117,791,842]" gridcol="3" gridrow="6" pageId="9" pageNumber="344">4.03 ± 0.08 (3.88 – 4.14)</td>
<td id="76885DD3FFCD3DCAFB4BAD5B3B22C800" box="[1225,1373,791,842]" gridcol="4" gridrow="6" pageId="9" pageNumber="344">4.09 ± 0.03 (3.95 – 4.33)</td>
</tr>
<tr id="355934AFFFCD3DCAFF23AD163B22C8C4" box="[161,1373,858,910]" gridrow="7" pageId="9" pageNumber="344">
<th id="76885DD3FFCD3DCAFF23AD163F61C8C4" box="[161,286,858,910]" gridcol="0" gridrow="7" pageId="9" pageNumber="344">ZB</th>
<td id="76885DD3FFCD3DCAFE1EAD163C40C8C4" box="[412,575,858,910]" gridcol="1" gridrow="7" pageId="9" pageNumber="344">9.64</td>
<td id="76885DD3FFCD3DCAFD32AD163D3BC8C4" box="[688,836,858,910]" gridcol="2" gridrow="7" pageId="9" pageNumber="344">9.60 ± 0.05 (9.26 – 10.06)</td>
<td id="76885DD3FFCD3DCAFC33AD163A22C8C4" box="[945,1117,858,910]" gridcol="3" gridrow="7" pageId="9" pageNumber="344">9.90 ± 0.23 (9.54 – 10.37)</td>
<td id="76885DD3FFCD3DCAFB4BAD163B22C8C4" box="[1225,1373,858,910]" gridcol="4" gridrow="7" pageId="9" pageNumber="344">9.70 ± 0.05 (9.40 – 10.13)</td>
</tr>
<tr id="355934AFFFCD3DCAFF23ADD33B22C898" box="[161,1373,927,978]" gridrow="8" pageId="9" pageNumber="344">
<th id="76885DD3FFCD3DCAFF23ADD33F61C898" box="[161,286,927,978]" gridcol="0" gridrow="8" pageId="9" pageNumber="344">BCW</th>
<td id="76885DD3FFCD3DCAFE1EADD33C40C898" box="[412,575,927,978]" gridcol="1" gridrow="8" pageId="9" pageNumber="344">7.86</td>
<td id="76885DD3FFCD3DCAFD32ADD33D3BC898" box="[688,836,927,978]" gridcol="2" gridrow="8" pageId="9" pageNumber="344">7.77 ± 0.16 (7.47 – 8.10)</td>
<td id="76885DD3FFCD3DCAFC33ADD33A22C898" box="[945,1117,927,978]" gridcol="3" gridrow="8" pageId="9" pageNumber="344">7.89 ± 0.15 (7.64 – 8.23)</td>
<td id="76885DD3FFCD3DCAFB4BADD33B22C898" box="[1225,1373,927,978]" gridcol="4" gridrow="8" pageId="9" pageNumber="344">7.60 ± 0.09 (7.41 – 7.78)</td>
</tr>
<tr id="355934AFFFCD3DCAFF23ADAF3B22CF5C" box="[161,1373,995,1046]" gridrow="9" pageId="9" pageNumber="344">
<th id="76885DD3FFCD3DCAFF23ADAF3F61CF5C" box="[161,286,995,1046]" gridcol="0" gridrow="9" pageId="9" pageNumber="344">MAB</th>
<td id="76885DD3FFCD3DCAFE1EADAF3C40CF5C" box="[412,575,995,1046]" gridcol="1" gridrow="9" pageId="9" pageNumber="344">7.94</td>
<td id="76885DD3FFCD3DCAFD32ADAF3D3BCF5C" box="[688,836,995,1046]" gridcol="2" gridrow="9" pageId="9" pageNumber="344">7.73 ± 0.13 (7.54 – 7.98)</td>
<td id="76885DD3FFCD3DCAFC33ADAF3A22CF5C" box="[945,1117,995,1046]" gridcol="3" gridrow="9" pageId="9" pageNumber="344">7.94 ± 0.16 (7.66 – 8.31)</td>
<td id="76885DD3FFCD3DCAFB4BADAF3B22CF5C" box="[1225,1373,995,1046]" gridcol="4" gridrow="9" pageId="9" pageNumber="344">7.77 ± 0.12 (7.49 – 7.90)</td>
</tr>
<tr id="355934AFFFCD3DCAFF23AA6B3B22CF10" box="[161,1373,1063,1114]" gridrow="10" pageId="9" pageNumber="344">
<th id="76885DD3FFCD3DCAFF23AA6B3F61CF10" box="[161,286,1063,1114]" gridcol="0" gridrow="10" pageId="9" pageNumber="344">POC</th>
<td id="76885DD3FFCD3DCAFE1EAA6B3C40CF10" box="[412,575,1063,1114]" gridcol="1" gridrow="10" pageId="9" pageNumber="344">3.93</td>
<td id="76885DD3FFCD3DCAFD32AA6B3D3BCF10" box="[688,836,1063,1114]" gridcol="2" gridrow="10" pageId="9" pageNumber="344">3.77 ± 0.07 (3.62 – 3.89)</td>
<td id="76885DD3FFCD3DCAFC33AA6B3A22CF10" box="[945,1117,1063,1114]" gridcol="3" gridrow="10" pageId="9" pageNumber="344">3.95 ± 0.13 (3.69 – 4.16)</td>
<td id="76885DD3FFCD3DCAFB4BAA6B3B22CF10" box="[1225,1373,1063,1114]" gridcol="4" gridrow="10" pageId="9" pageNumber="344">3.60 ± 0.08 (3.47 – 3.71)</td>
</tr>
<tr id="355934AFFFCD3DCAFF23AA273B22CFD4" box="[161,1373,1131,1182]" gridrow="11" pageId="9" pageNumber="344">
<th id="76885DD3FFCD3DCAFF23AA273F61CFD4" box="[161,286,1131,1182]" gridcol="0" gridrow="11" pageId="9" pageNumber="344">ROL</th>
<td id="76885DD3FFCD3DCAFE1EAA273C40CFD4" box="[412,575,1131,1182]" gridcol="1" gridrow="11" pageId="9" pageNumber="344">4.79</td>
<td id="76885DD3FFCD3DCAFD32AA273D3BCFD4" box="[688,836,1131,1182]" gridcol="2" gridrow="11" pageId="9" pageNumber="344">4.76 ± 0.20 (4.46 – 5.35)</td>
<td id="76885DD3FFCD3DCAFC33AA273A22CFD4" box="[945,1117,1131,1182]" gridcol="3" gridrow="11" pageId="9" pageNumber="344">4.69 ± 0.25 (4.08 – 4.99)</td>
<td id="76885DD3FFCD3DCAFB4BAA273B22CFD4" box="[1225,1373,1131,1182]" gridcol="4" gridrow="11" pageId="9" pageNumber="344">4.63 ± 0.09 (4.50 – 4.76)</td>
</tr>
<tr id="355934AFFFCD3DCAFF23AAE33B22CFA8" box="[161,1373,1199,1250]" gridrow="12" pageId="9" pageNumber="344">
<th id="76885DD3FFCD3DCAFF23AAE33F61CFA8" box="[161,286,1199,1250]" gridcol="0" gridrow="12" pageId="9" pageNumber="344">M1M3</th>
<td id="76885DD3FFCD3DCAFE1EAAE33C40CFA8" box="[412,575,1199,1250]" gridcol="1" gridrow="12" pageId="9" pageNumber="344">3.50</td>
<td id="76885DD3FFCD3DCAFD32AAE33D3BCFA8" box="[688,836,1199,1250]" gridcol="2" gridrow="12" pageId="9" pageNumber="344">3.48 ± 0.12 (3.30 – 3.71)</td>
<td id="76885DD3FFCD3DCAFC33AAE33A22CFA8" box="[945,1117,1199,1250]" gridcol="3" gridrow="12" pageId="9" pageNumber="344">3.54 ± 0.086 (3.35 – 3.67)</td>
<td id="76885DD3FFCD3DCAFB4BAAE33B22CFA8" box="[1225,1373,1199,1250]" gridcol="4" gridrow="12" pageId="9" pageNumber="344">3.53 ± 0.08 (3.40 – 3.68)</td>
</tr>
<tr id="355934AFFFCD3DCAFF23AABF3B22CE6C" box="[161,1373,1267,1318]" gridrow="13" pageId="9" pageNumber="344">
<th id="76885DD3FFCD3DCAFF23AABF3F61CE6C" box="[161,286,1267,1318]" gridcol="0" gridrow="13" pageId="9" pageNumber="344">ml</th>
<td id="76885DD3FFCD3DCAFE1EAABF3C40CE6C" box="[412,575,1267,1318]" gridcol="1" gridrow="13" pageId="9" pageNumber="344">11.14</td>
<td id="76885DD3FFCD3DCAFD32AABF3D3BCE6C" box="[688,836,1267,1318]" gridcol="2" gridrow="13" pageId="9" pageNumber="344">11.26 ± 0.30 (10.76 – 11.92)</td>
<td id="76885DD3FFCD3DCAFC33AABF3A22CE6C" box="[945,1117,1267,1318]" gridcol="3" gridrow="13" pageId="9" pageNumber="344">11.38 ± 0.31 (10.55 – 11.73)</td>
<td id="76885DD3FFCD3DCAFB4BAABF3B22CE6C" box="[1225,1373,1267,1318]" gridcol="4" gridrow="13" pageId="9" pageNumber="344">11.36 ± 0.20 (11.04 – 11.84)</td>
</tr>
<tr id="355934AFFFCD3DCAFF23AB7B3B22CE20" box="[161,1373,1335,1386]" gridrow="14" pageId="9" pageNumber="344">
<th id="76885DD3FFCD3DCAFF23AB7B3F61CE20" box="[161,286,1335,1386]" gridcol="0" gridrow="14" pageId="9" pageNumber="344">cm3L</th>
<td id="76885DD3FFCD3DCAFE1EAB7B3C40CE20" box="[412,575,1335,1386]" gridcol="1" gridrow="14" pageId="9" pageNumber="344">6.04</td>
<td id="76885DD3FFCD3DCAFD32AB7B3D3BCE20" box="[688,836,1335,1386]" gridcol="2" gridrow="14" pageId="9" pageNumber="344">6.40 ± 0.20 (6.00 – 6.87)</td>
<td id="76885DD3FFCD3DCAFC33AB7B3A22CE20" box="[945,1117,1335,1386]" gridcol="3" gridrow="14" pageId="9" pageNumber="344">6.43 ± 0.12 (6.13 – 6.61)</td>
<td id="76885DD3FFCD3DCAFB4BAB7B3B22CE20" box="[1225,1373,1335,1386]" gridcol="4" gridrow="14" pageId="9" pageNumber="344">6.38 ± 0.16 (6.17 – 6.64)</td>
</tr>
<tr id="355934AFFFCD3DCAFF23AB373B22CEE4" box="[161,1373,1403,1454]" gridrow="15" pageId="9" pageNumber="344">
<th id="76885DD3FFCD3DCAFF23AB373F61CEE4" box="[161,286,1403,1454]" gridcol="0" gridrow="15" pageId="9" pageNumber="344">m1m3L</th>
<td id="76885DD3FFCD3DCAFE1EAB373C40CEE4" box="[412,575,1403,1454]" gridcol="1" gridrow="15" pageId="9" pageNumber="344">3.81</td>
<td id="76885DD3FFCD3DCAFD32AB373D3BCEE4" box="[688,836,1403,1454]" gridcol="2" gridrow="15" pageId="9" pageNumber="344">3.88 ± 0.10 (3.64 – 4.05)</td>
<td id="76885DD3FFCD3DCAFC33AB373A22CEE4" box="[945,1117,1403,1454]" gridcol="3" gridrow="15" pageId="9" pageNumber="344">3.89 ± 0.12 (3.66 – 4.08)</td>
<td id="76885DD3FFCD3DCAFB4BAB373B22CEE4" box="[1225,1373,1403,1454]" gridcol="4" gridrow="15" pageId="9" pageNumber="344">3.90 ± 0.08 (3.81 – 4.06)</td>
</tr>
<tr id="355934AFFFCD3DCAFF23ABF33B22CEB8" box="[161,1373,1471,1522]" gridrow="16" pageId="9" pageNumber="344">
<th id="76885DD3FFCD3DCAFF23ABF33F61CEB8" box="[161,286,1471,1522]" gridcol="0" gridrow="16" pageId="9" pageNumber="344">coh</th>
<td id="76885DD3FFCD3DCAFE1EABF33C40CEB8" box="[412,575,1471,1522]" gridcol="1" gridrow="16" pageId="9" pageNumber="344">3.29</td>
<td id="76885DD3FFCD3DCAFD32ABF33D3BCEB8" box="[688,836,1471,1522]" gridcol="2" gridrow="16" pageId="9" pageNumber="344">3.21 ± 0.10 (2.98 – 3.44)</td>
<td id="76885DD3FFCD3DCAFC33ABF33A22CEB8" box="[945,1117,1471,1522]" gridcol="3" gridrow="16" pageId="9" pageNumber="344">3.31 ± 0.10 (3.12 – 3.47)</td>
<td id="76885DD3FFCD3DCAFB4BABF33B22CEB8" box="[1225,1373,1471,1522]" gridcol="4" gridrow="16" pageId="9" pageNumber="344">3.45 ± 0.10 (3.29 – 3.57)</td>
</tr>
</table>
</paragraph>
<caption id="DF166665FFCEC23FFF15AAA33AF3CE32" ID-DOI="http://doi.org/10.5281/zenodo.10264625" ID-Zenodo-Dep="10264625" httpUri="https://zenodo.org/record/10264625/files/figure.png" pageId="10" pageNumber="345" startId="10.[151,186,1263,1285]" targetBox="[151,1436,257,1218]" targetPageId="10" targetType="figure">
<paragraph id="8BD636EDFFCEC23FFF15AAA33AF3CE32" blockId="10.[151,1435,1263,1400]" pageId="10" pageNumber="345">
Fig. 5. External characters of 
<taxonomicName id="4C694D6EFFCEC23FFE39AAA33C2DCE4E" authority="Puechmaille &amp; Dool &amp; Beuneux &amp; Ruedi, 2023" authorityName="Puechmaille &amp; Dool &amp; Beuneux &amp; Ruedi" authorityYear="2023" box="[443,594,1263,1284]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="345" phylum="Chordata" rank="species" species="nustrale" status="sp. nov.">
<emphasis id="B91DEAFFFFCEC23FFE39AAA33C2DCE4E" box="[443,594,1263,1284]" italics="true" pageId="10" pageNumber="345">Myotis nustrale</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E5784FFCEC23FFDDAAAA33CDDCE4E" box="[600,674,1263,1284]" pageId="10" pageNumber="345" rank="species">sp. nov.</taxonomicNameLabel>
Panels are close-ups of the female holotype (MNHN-ZM-2023-12), except the lower right one which is a picture from a released adult individual (Photo © Yann Le Bris). The upper left panel illustrates the stiff hairs running along the uropatagium, viewed from below. The upper right panel is the right foot with wing insertion to the base of the outer toe, in ventral view. The lower left panel illustrates the long and straight tragus and unnotched ear of the female holotype. On the portrait of the live individual, notice the black chin spot on the lower lip.
</paragraph>
</caption>
<subSubSection id="C3736566FFCEC23FFF15A9793F9BCCE4" pageId="10" pageNumber="345" type="vernacular_names">
<paragraph id="8BD636EDFFCEC23FFF15A9793F9BCCE4" blockId="10.[151,756,1845,1966]" pageId="10" pageNumber="345">
<emphasis id="B91DEAFFFFCEC23FFF15A9793C7CCC07" bold="true" box="[151,515,1845,1869]" pageId="10" pageNumber="345">Proposed vernacular names:</emphasis>
Korsican Mausohr (German), Corsican 
<taxonomicName id="4C694D6EFFCEC23FFEFEA9193C48CC27" authority="(English)" baseAuthorityName="English" box="[380,567,1877,1901]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="345" phylum="Chordata" rank="species" species="myotis">myotis (English)</taxonomicName>
, murin de 
<collectingCountry id="F37E767DFFCEC23FFD37A9193C8CCC27" box="[693,755,1877,1901]" name="France" pageId="10" pageNumber="345">Corse</collectingCountry>
(French), murin 
<taxonomicName id="4C694D6EFFCEC23FFEF0A9393C24CCC7" authority="(Corsican)" baseAuthorityName="Corsican" box="[370,603,1909,1933]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="345" phylum="Chordata" rank="species" species="nustrale">nustrale (Corsican)</taxonomicName>
, murciélago ratonero de 
<collectingCountry id="F37E767DFFCEC23FFE95A9DA3F0FCCE4" box="[279,368,1942,1966]" name="France" pageId="10" pageNumber="345">Córcega</collectingCountry>
(Spanish).
</paragraph>
</subSubSection>
<subSubSection id="C3736566FFCEC23EFF15A9893C74CDEF" lastPageId="11" lastPageNumber="346" pageId="10" pageNumber="345" type="biology_ecology">
<paragraph id="8BD636EDFFCEC23EFF15A9893F8FCEAA" blockId="10.[151,756,1989,2077]" lastBlockId="11.[151,756,1414,1701]" lastPageId="11" lastPageNumber="346" pageId="10" pageNumber="345">
<emphasis id="B91DEAFFFFCEC23FFF15A9893F1CCC97" bold="true" box="[151,355,1989,2013]" pageId="10" pageNumber="345">Natural history:</emphasis>
Several females equipped with transmitters were radio tracked in spring and summer in two mountainous areas of central 
<collectingCountry id="F37E767DFFCEC23FFDB8A6493CEFC357" box="[570,656,2053,2077]" name="France" pageId="10" pageNumber="345">Corsica</collectingCountry>
: Ghisoni and Bavella (
<bibRefCitation id="EFF84B1CFFCEC23FFC5AAB8B3B32CE95" author="Groupe Chiropteres Corse" box="[984,1357,1479,1503]" pageId="10" pageNumber="345" refId="ref12005" refString="Groupe Chiropteres Corse. 2019. Programme d'etude concernant l'amelioration des connaissances sur l'ecologie d'une nouvelle espece de chauve-souris endemique a la Corse: Myotis sp. C (ex- Myotis nattereri) - Annee 2018. (Groupe Chiropteres Corse, Corte, France), 9 pp." type="book" year="2019">
Groupe Chiroptères 
<collectingCountry id="F37E767DFFCEC23FFB43AB8B3B7DCE95" box="[1217,1282,1479,1503]" name="France" pageId="10" pageNumber="345">Corse</collectingCountry>
, 2019
</bibRefCitation>
). Most records were gathered from forested areas above 
<quantity id="4C919B08FFCEC23FFAD5ABA43BE3CD4A" box="[1367,1436,1512,1536]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="345" unit="m" value="500.0">500 m</quantity>
a.s.l., although a few observations have been recorded near sea level (
<figureCitation id="13522A68FFCEC23FFC77A8603A43CD0E" box="[1013,1084,1580,1604]" captionStart="Fig" captionStartId="3.[831,866,1404,1426]" captionTargetBox="[901,1422,260,1364]" captionTargetId="figure-564@3.[831,1436,254,1373]" captionTargetPageId="3" captionText="Fig. 1. Topographic map of Corsica (France) showing the current records of M. nustrale sp. nov. on this Mediterranean island (Courtois et al., 2011; Groupe Chiroptères Corse, 2019). The black dots represent capture sites, while the red stars indicate the presence of lactating females, including those living in the only known nursery roost of Ghisoni. A double circle indicates the location where the holotype of M.nustrale sp. nov. was found. Shades of grey represent areas above 100 m a.s.l." figureDoi="http://doi.org/10.5281/zenodo.10264613" httpUri="https://zenodo.org/record/10264613/files/figure.png" pageId="10" pageNumber="345">Fig. 1</figureCitation>
). These animals were hunting in various habitats located within 
<quantity id="4C919B08FFCEC23FFB49A8023B78CD2C" box="[1227,1287,1614,1638]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="345" unit="km" value="8.0">8 km</quantity>
of their day roosts. Hunting territories were used regularly by single individuals and were either located in dense maquis populated with evergreen oak (
<taxonomicName id="4C694D6EFFCEC23FFB7DA8F83BF3CD81" box="[1279,1420,1715,1739]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="10" pageNumber="345" phylum="Tracheophyta" rank="species" species="ilex">
<emphasis id="B91DEAFFFFCEC23FFB7DA8F83BF3CD81" box="[1279,1420,1715,1739]" italics="true" pageId="10" pageNumber="345">Quercus ilex</emphasis>
</taxonomicName>
), ash (
<taxonomicName id="4C694D6EFFCEC23FFCFBA89A3A6DCDA7" box="[889,1042,1750,1773]" class="Magnoliopsida" family="Oleaceae" genus="Fraxinus" kingdom="Plantae" order="Lamiales" pageId="10" pageNumber="345" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B91DEAFFFFCEC23FFCFBA89A3DA5CDA7" box="[889,986,1750,1773]" italics="true" pageId="10" pageNumber="345">Fraxinus</emphasis>
spp.
</taxonomicName>
) and abundant undergrowth, or in more open forests with Corsican pines (
<taxonomicName id="4C694D6EFFCEC23FFA8DA8B43BECCC45" box="[1295,1427,1784,1807]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="10" pageNumber="345" phylum="Tracheophyta" rank="species" species="nigra">
<emphasis id="B91DEAFFFFCEC23FFA8DA8B43BECCC45" box="[1295,1427,1784,1807]" italics="true" pageId="10" pageNumber="345">Pinus nigra</emphasis>
</taxonomicName>
) surrounded by ferns. The bats were hunting both high in the canopy and closer to the ground. All known day roosts (
<figureCitation id="13522A68FFCEC23FFC52A9103A67CC3E" box="[976,1048,1884,1908]" captionStart="Fig" captionStartId="3.[831,866,1404,1426]" captionTargetBox="[901,1422,260,1364]" captionTargetId="figure-564@3.[831,1436,254,1373]" captionTargetPageId="3" captionText="Fig. 1. Topographic map of Corsica (France) showing the current records of M. nustrale sp. nov. on this Mediterranean island (Courtois et al., 2011; Groupe Chiroptères Corse, 2019). The black dots represent capture sites, while the red stars indicate the presence of lactating females, including those living in the only known nursery roost of Ghisoni. A double circle indicates the location where the holotype of M.nustrale sp. nov. was found. Shades of grey represent areas above 100 m a.s.l." figureDoi="http://doi.org/10.5281/zenodo.10264613" httpUri="https://zenodo.org/record/10264613/files/figure.png" pageId="10" pageNumber="345">Fig. 1</figureCitation>
) were in cliffs, rocky outcrops, stone bridges or in an artificial tunnel (where the only recorded nursery colony of ca. 60 individuals was established), denoting a strong rupicolous roosting behaviour for this Corsican myotis. This clear affinity for rocky habitats is similar to the roosting behaviour of 
<taxonomicName id="4C694D6EFFCFC23EFF38ABCB3F39CED4" authorityName="Cabrera" authorityYear="1904" box="[186,326,1414,1438]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="346" phylum="Chordata" rank="species" species="escalerai">
<emphasis id="B91DEAFFFFCFC23EFF38ABCB3F39CED4" box="[186,326,1414,1438]" italics="true" pageId="11" pageNumber="346">M. escalerai</emphasis>
</taxonomicName>
, but is distinct from the typical treeroosting behaviour of 
<taxonomicName id="4C694D6EFFCFC23EFE21ABEB3C51CEF4" baseAuthorityName="Kuhl" baseAuthorityYear="1817" box="[419,558,1447,1470]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="346" phylum="Chordata" rank="species" species="nattereri">
<emphasis id="B91DEAFFFFCFC23EFE21ABEB3C51CEF4" box="[419,558,1447,1470]" italics="true" pageId="11" pageNumber="346">M. nattereri</emphasis>
</taxonomicName>
or 
<taxonomicName id="4C694D6EFFCFC23EFDE6ABEB3F92CEAA" authority="(Agirre-Mendi &amp; Ibanez, 2012)" baseAuthorityName="Agirre-Mendi &amp; Ibanez" baseAuthorityYear="2012" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="346" phylum="Chordata" rank="species" species="crypticus">
<emphasis id="B91DEAFFFFCFC23EFDE6ABEB3C8BCEF4" box="[612,756,1447,1470]" italics="true" pageId="11" pageNumber="346">M. crypticus</emphasis>
(
<bibRefCitation id="EFF84B1CFFCFC23EFF1CAB843F9ACEAA" author="Agirre-Mendi P. T. &amp; Ibanez C." box="[158,485,1480,1504]" journalOrPublisher="Barbastella" pageId="11" pageNumber="346" pagination="8 - 11" part="5" refId="ref10944" refString="Agirre-Mendi P. T., Ibanez C. 2012. Primeros datos sobre la distribucion de Myotis cf. nattereri y Myotis escalerai Cabrera, 1904 (Chiroptera: Vespertilionidae) en la Comunidad Autonoma de La Rioja. Barbastella 5: 8 - 11." title="Primeros datos sobre la distribucion de Myotis cf. nattereri y Myotis escalerai Cabrera, 1904 (Chiroptera: Vespertilionidae) en la Comunidad Autonoma de La Rioja" type="journal article" year="2012">Agirre-Mendi &amp; Ibáñez, 2012</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<caption id="DF166665FFCFC23EFF15AAF03B0DCE62" ID-DOI="http://doi.org/10.5281/zenodo.10264627" ID-Zenodo-Dep="10264627" httpUri="https://zenodo.org/record/10264627/files/figure.png" pageId="11" pageNumber="346" startId="11.[151,186,1212,1234]" targetBox="[151,1436,257,1160]" targetPageId="11" targetType="figure">
<paragraph id="8BD636EDFFCFC23EFF15AAF03B0DCE62" blockId="11.[151,1435,1212,1320]" pageId="11" pageNumber="346">
Fig. 6. Portraits and hind feet (dorsal view) of an adult female 
<taxonomicName id="4C694D6EFFCFC23EFD76AAF03D1CCF9B" authority="Puechmaille &amp; Dool &amp; Beuneux &amp; Ruedi, 2023" authorityName="Puechmaille &amp; Dool &amp; Beuneux &amp; Ruedi" authorityYear="2023" box="[756,867,1212,1233]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="346" phylum="Chordata" rank="species" species="nustrale" status="sp. nov.">
<emphasis id="B91DEAFFFFCFC23EFD76AAF03D1CCF9B" box="[756,867,1212,1233]" italics="true" pageId="11" pageNumber="346">M. nustrale</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E5784FFCFC23EFCEAAAF03DCDCF9B" box="[872,946,1212,1233]" pageId="11" pageNumber="346" rank="species">sp. nov.</taxonomicNameLabel>
caught near Bavella, in southern Corsica (left) and an adult female 
<taxonomicName id="4C694D6EFFCFC23EFE02AA953F85CFA4" baseAuthorityName="Agirre-Mendi &amp; Ibanez" baseAuthorityYear="2012" box="[384,506,1241,1262]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="346" phylum="Chordata" rank="species" species="crypticus">
<emphasis id="B91DEAFFFFCFC23EFE02AA953F85CFA4" box="[384,506,1241,1262]" italics="true" pageId="11" pageNumber="346">M. crypticus</emphasis>
</taxonomicName>
from the Jura mountains in Switzerland (right). Notice that the latter individual has a faint line bordering the lower lip, while the former has a much more conspicuous chin spot. Wing insertion to the outer toe is, however, identical in both species. For the left individual, the wing was not stretched while it was stretched for the right individual.
</paragraph>
</caption>
<paragraph id="8BD636EDFFCFC23EFF15ABA43C74CDEF" blockId="11.[151,756,1414,1701]" pageId="11" pageNumber="346">
Other bat species recorded in the same hunting habitats as those frequented by the Corsican myotis included 
<emphasis id="B91DEAFFFFCFC23EFF15A8673C8BCD29" italics="true" pageId="11" pageNumber="346">
<taxonomicName id="4C694D6EFFCFC23EFF15A8673F0ECD08" baseAuthorityName="Schreber" baseAuthorityYear="1774" box="[151,369,1579,1602]" class="Mammalia" family="Vespertilionidae" genus="Eptesicus" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="346" phylum="Chordata" rank="species" species="serotinus">Eptesicus serotinus</taxonomicName>
, 
<taxonomicName id="4C694D6EFFCFC23EFE00A8673C5ACD08" baseAuthorityName="Bonaparte" baseAuthorityYear="1837" box="[386,549,1579,1602]" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="346" phylum="Chordata" rank="species" species="savii">Hypsugo savii</taxonomicName>
, 
<taxonomicName id="4C694D6EFFCFC23EFDB4A8663C90CD08" baseAuthorityName="Kuhl" baseAuthorityYear="1817" box="[566,751,1578,1602]" class="Mammalia" family="Vespertilionidae" genus="Nyctalus" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="346" phylum="Chordata" rank="species" species="leisleri">Nyctalus leisleri</taxonomicName>
, 
<taxonomicName id="4C694D6EFFCFC23EFF15A8073C42CD29" authority=", Rhinolophus" authorityName="Rhinolophus" box="[151,573,1611,1635]" class="Mammalia" family="Vespertilionidae" genus="Pipistrellus" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="346" phylum="Chordata" rank="species" species="pipistrellus">Pipistrellus pipistrellus, Rhinolophus</taxonomicName>
ferrumequinum
</emphasis>
and 
<taxonomicName id="4C694D6EFFCFC23EFF4BA8203FDECDCE" baseAuthorityName="J. Fischer" baseAuthorityYear="1829" box="[201,417,1644,1668]" class="Mammalia" family="Vespertilionidae" genus="Plecotus" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="346" phylum="Chordata" rank="species" species="austriacus">
<emphasis id="B91DEAFFFFCFC23EFF4BA8203FDECDCE" box="[201,417,1644,1668]" italics="true" pageId="11" pageNumber="346">Plecotus austriacus</emphasis>
</taxonomicName>
, but not other 
<taxonomicName id="4C694D6EFFCFC23EFDD0A8213C8CCDCE" box="[594,755,1644,1668]" pageId="11" pageNumber="346">
<emphasis id="B91DEAFFFFCFC23EFDD0A8213CE4CDCE" box="[594,667,1645,1668]" italics="true" pageId="11" pageNumber="346">Myotis</emphasis>
species
</taxonomicName>
(
<bibRefCitation id="EFF84B1CFFCFC23EFF1CA8C13F80CDEF" author="Groupe Chiropteres Corse" box="[158,511,1677,1701]" pageId="11" pageNumber="346" refId="ref12005" refString="Groupe Chiropteres Corse. 2019. Programme d'etude concernant l'amelioration des connaissances sur l'ecologie d'une nouvelle espece de chauve-souris endemique a la Corse: Myotis sp. C (ex- Myotis nattereri) - Annee 2018. (Groupe Chiropteres Corse, Corte, France), 9 pp." type="book" year="2019">
Groupe Chiroptères 
<collectingCountry id="F37E767DFFCFC23EFEF8A8C13FC4CDEF" box="[378,443,1677,1701]" name="France" pageId="11" pageNumber="346">Corse</collectingCountry>
, 2019
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3736566FFCFC23EFF15A8F03ACFCEAB" pageId="11" pageNumber="346" type="conservation">
<paragraph id="8BD636EDFFCFC23EFF15A8F03ACFCEAB" blockId="11.[151,756,1724,2077]" lastBlockId="11.[831,1436,1415,1505]" pageId="11" pageNumber="346">
<emphasis id="B91DEAFFFFCFC23EFF15A8F03F46CD9E" bold="true" box="[151,313,1724,1748]" pageId="11" pageNumber="346">Conservation:</emphasis>
This endemic species is apparently very rare (
<bibRefCitation id="EFF84B1CFFCFC23EFF57A8923FB8CDBC" author="Courtois J. - Y. &amp; Rist D. &amp; Beuneux G." box="[213,455,1758,1782]" pageId="11" pageNumber="346" refId="ref11538" refString="Courtois J. - Y., Rist D., Beuneux G. 2011. Les chauves-souris de Corse. Albiana, Ajaccio, France, 167 pp." type="book" year="2011">
Courtois 
<emphasis id="B91DEAFFFFCFC23EFEC3A8923F02CDBC" box="[321,381,1758,1782]" italics="true" pageId="11" pageNumber="346">et al.</emphasis>
, 2011
</bibRefCitation>
) and owing to its limited distribution range in the mountains of 
<collectingCountry id="F37E767DFFCFC23EFDC2A8B23CEECC5C" box="[576,657,1790,1814]" name="France" pageId="11" pageNumber="346">Corsica</collectingCountry>
(
<figureCitation id="13522A68FFCFC23EFD21A8B23C9CCC5D" box="[675,739,1790,1815]" captionStart="Fig" captionStartId="3.[831,866,1404,1426]" captionTargetBox="[901,1422,260,1364]" captionTargetId="figure-564@3.[831,1436,254,1373]" captionTargetPageId="3" captionText="Fig. 1. Topographic map of Corsica (France) showing the current records of M. nustrale sp. nov. on this Mediterranean island (Courtois et al., 2011; Groupe Chiroptères Corse, 2019). The black dots represent capture sites, while the red stars indicate the presence of lactating females, including those living in the only known nursery roost of Ghisoni. A double circle indicates the location where the holotype of M.nustrale sp. nov. was found. Shades of grey represent areas above 100 m a.s.l." figureDoi="http://doi.org/10.5281/zenodo.10264613" httpUri="https://zenodo.org/record/10264613/files/figure.png" pageId="11" pageNumber="346">Fig. 1</figureCitation>
), should be considered as globally endangered. Based on our current knowledge, the species has an area of occupancy of 
<quantity id="4C919B08FFCFC23EFEB2A92D3FE8CC33" box="[304,407,1889,1913]" metricMagnitude="6" metricUnit="m" metricValue="1.3" pageId="11" pageNumber="346" unit="km" value="1300.0">1,300 km</quantity>
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and an extent of occurrence of 
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. For the present and given its broad tolerance for diverse forested habitats as hunting grounds, humaninduced habitat change is not foreseen as a threat for the species. Forest fires might, however, temporarily limit hunting habitats. Rocky outcrops used as day roosts do not appear to be limiting factors in 
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, but they could be subject to disturbance from rock climbing or abseiling and associated activities.
</paragraph>
</subSubSection>
</treatment>
</document>