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<document id="8E9C604CE4A7D559EF86FA66945C86E4" ID-DOI="10.5281/zenodo.170972" ID-GBIF-Dataset="f93b6cae-09b4-47ce-8eba-27d769ba4e26" ID-ISSN="1175­5326" ID-Zenodo-Dep="170972" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459717480872" checkinUser="plazi" docAuthor="Yu, A. &amp; Solodovnikov" docDate="2005" docId="03C087ECFFC2E209FED4F96803E7D502" docLanguage="en" docName="zt00902.pdf" docOrigin="Zootaxa 902" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Microsaurus Dejean 1833" docType="treatment" docVersion="9" lastPageNumber="3" masterDocId="FFF9FF94FFC3E20BFFDCFFA1010FD22B" masterDocTitle="New and little known species of Quedius from West Palaearctic (Coleoptera: Staphylinidae: Staphylininae)" masterLastPageNumber="13" masterPageNumber="1" pageNumber="2" updateTime="1698208227348" updateUser="plazi">
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<mods:title id="F1C9C62EEF874F9F6838E1FF40A0A32A">New and little known species of Quedius from West Palaearctic (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
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<mods:date id="658C176486FFAFDBE667CC7BF26AADE9">2005</mods:date>
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<paragraph id="8BD636FAFFC2E20AFED4F96802E4D4C8" blockId="1.[264,1003,1737,1763]" box="[264,1003,1737,1763]" pageId="1" pageNumber="2">
<heading id="D09E8196FFC2E20AFED4F96802E4D4C8" bold="true" box="[264,1003,1737,1763]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
<emphasis id="B91DEAE8FFC2E20AFED4F96802E4D4C8" bold="true" box="[264,1003,1737,1763]" pageId="1" pageNumber="2">
Subgenus
<taxonomicName id="4C694D79FFC2E20AFE54F96803DFD4C8" authority="Dejean, 1833" authorityName="Dejean" authorityYear="1833" box="[392,720,1737,1763]" class="Insecta" family="Staphylinidae" genus="Microsaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
Microsaurus
<bibRefCitation id="EFF84B0BFFC2E20AFDF2F96803DFD4C8" author="Dejean" box="[558,720,1737,1763]" pageId="1" pageNumber="12" refString="Dejean, P. F. M. A. (1833) Catalogue des Coleopteres de la collection de M. le Comte Dejean. Mequignon-Marvis, Paris, 176 pp." type="book" year="1833">Dejean, 1833</bibRefCitation>
</taxonomicName>
, sensu
<bibRefCitation id="EFF84B0BFFC2E20AFCF5F96802E4D4C8" author="Smetana" box="[809,1003,1737,1763]" pageId="1" pageNumber="13" refString="Smetana, A. (1971) Revision of the tribe Quediini of America north of Mexico (Coleoptera: Staphylinidae). Memoirs of the Entomological Society of Canada, 79, vi + 303 pp." type="book" year="1971">Smetana (1971)</bibRefCitation>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3736571FFC2E20AFED4F8B5006AD540" pageId="1" pageNumber="2" type="reference_group">
<paragraph id="8BD636FAFFC2E20AFED4F8B5006AD540" blockId="1.[264,1323,1812,1899]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="AB7911D4FFC2E20AFED4F8B5006AD540" pageId="1" pageNumber="2">
<taxonomicName id="4C694D79FFC2E20AFED4F8B500E1D560" ID-CoL="4R2KP" authority="Rambousek, 1915 Rambousek 1915: 27" authorityName="Rambousek, 1915 Rambousek" authorityPageNumber="27" authorityYear="1915" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="bernhaueri" subGenus="Microsaurus">
<emphasis id="B91DEAE8FFC2E20AFED4F8B5006ED500" box="[264,353,1812,1835]" italics="true" pageId="1" pageNumber="2">Quedius</emphasis>
(
<emphasis id="B91DEAE8FFC2E20AFEADF8B500F7D500" box="[369,504,1812,1835]" italics="true" pageId="1" pageNumber="2">Microsaurus</emphasis>
)
<emphasis id="B91DEAE8FFC2E20AFDDBF8B50371D500" box="[519,638,1812,1835]" italics="true" pageId="1" pageNumber="2">bernhaueri</emphasis>
<bibRefCitation id="EFF84B0BFFC2E20AFD58F8B5024AD500" author="Rambousek" box="[644,837,1812,1835]" pageId="1" pageNumber="13" refString="Rambousek, F. (1915) Tenebrobius (nov. subg. Quediorum) bernhaueri, novy druh ze str. Makedonie. (Col. Staphyl.). C asopis C eske Spole c nosti Entomologicke, 12, 54 - 56." type="journal article" year="1915">Rambousek, 1915</bibRefCitation>
<treatmentCitation id="0AC810EBFFC2E20AFED4F89500E1D560" author="Rambousek" box="[264,494,1844,1867]" page="27" pageId="1" pageNumber="2" year="1915">
<bibRefCitation id="EFF84B0BFFC2E20AFED4F89500C9D560" author="Rambousek" box="[264,454,1844,1867]" pageId="1" pageNumber="13" refString="Rambousek, F. (1915) Tenebrobius (nov. subg. Quediorum) bernhaueri, novy druh ze str. Makedonie. (Col. Staphyl.). C asopis C eske Spole c nosti Entomologicke, 12, 54 - 56." type="journal article" year="1915">Rambousek 1915</bibRefCitation>
: 27
</treatmentCitation>
</taxonomicName>
; 1916: 130;
<treatmentCitation id="0AC810EBFFC2E20AFDA9F895023FD560" author="Gridelli" box="[629,816,1844,1867]" page="33" pageId="1" pageNumber="2" year="1924">
<bibRefCitation id="EFF84B0BFFC2E20AFDA9F8950207D560" author="Gridelli" box="[629,776,1844,1867]" pageId="1" pageNumber="13" refString="Gridelli, E. (1924) Studi sul genere Quedius Steph. (Col. Staph.). Secondo contributo. Specie della regione paleartica. Memorie della Societa Entomologica Italiana, 3 (1), 5 - 112." type="journal article" year="1924">Gridelli 1924</bibRefCitation>
: 33
</treatmentCitation>
;
<treatmentCitation id="0AC810EBFFC2E20AFCE0F895050AD560" author="Coiffait" box="[828,1029,1844,1867]" page="181" pageId="1" pageNumber="2" year="1978">
<bibRefCitation id="EFF84B0BFFC2E20AFCE0F89502C0D560" author="Coiffait" box="[828,975,1844,1867]" pageId="1" pageNumber="12" refString="Coiffait, H. (1978) Coleopteres Staphylinidae de la region paleartique occidentale. III. Sous famille Staphylininae, tribu Quediini; sous famille Paederinae, tribu Pinophilini. Nouvelle Revue d'Entomologie, Supplement, 8 (4), 1 - 364." type="journal article" year="1978">Coiffait 1978</bibRefCitation>
: 181
</treatmentCitation>
;
<treatmentCitation id="0AC810EBFFC2E20AFBCDF895006ED540" author="Assing" page="114" pageId="1" pageNumber="2" year="2001">
<bibRefCitation id="EFF84B0BFFC2E20AFBCDF8950427D560" author="Assing" box="[1041,1320,1844,1867]" pageId="1" pageNumber="12" refString="Assing, V. &amp; Wunderle, P. (2001) On the Staphylinidae of Greece. II. New species and new records from central and northern Greece (Insecta: Coleoptera). Linzer biologische Beitrage, 33 (1), 103 - 136." type="journal article" year="2001">Assing &amp; Wunderle 2001</bibRefCitation>
: 114
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3736571FFC1E209FED4FEBA059FD14D" pageId="2" pageNumber="3" type="materials_examined">
<paragraph id="8BD636FAFFC1E209FED4FEBA05B6D385" blockId="2.[264,1324,283,1833]" pageId="2" pageNumber="3">
<emphasis id="B91DEAE8FFC1E209FED4FEBA00CDD31E" bold="true" box="[264,450,283,309]" pageId="2" pageNumber="3">
<typeStatus id="54D28858FFC1E209FED4FEBA004CD31E" box="[264,323,283,309]" pageId="2" pageNumber="3">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54D28858FFC1E209FE0FFEBD0347D31D" box="[467,584,284,310]" pageId="2" pageNumber="3" type="lectotype">Lectotype</typeStatus>
(here designated): ɗ,
<collectingCountry id="F37E766AFFC1E209FCBBFEBD0510D31D" box="[871,1055,284,310]" name="Macedonia" pageId="2" pageNumber="3">MACEDONIA</collectingCountry>
: “Maced.[
<collectingCountry id="F37E766AFFC1E209FB46FEBD042BD31D" box="[1178,1316,284,310]" name="Macedonia" pageId="2" pageNumber="3">Macedonia</collectingCountry>
] Perister alpin. 7. 14 Dr. Rambousek/
<taxonomicName id="4C694D79FFC1E209FD6AFEE50234D376" box="[694,827,324,349]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="13" phylum="Arthropoda" rank="species" species="bernhaueri">
<emphasis id="B91DEAE8FFC1E209FD6AFEE50234D376" box="[694,827,324,349]" italics="true" pageId="2" pageNumber="3">Bernhaueri</emphasis>
</taxonomicName>
m. Typ det. Rambousek” (
<collectionCode id="ED78AE3FFFC1E209FBAFFEE505B6D375" box="[1139,1209,324,350]" httpUri="http://grbio.org/cool/n1s9-2rrz" name="Slovak National Museum" pageId="2" pageNumber="3">SNM</collectionCode>
);
<typeStatus id="54D28858FFC1E209FB10FEE50059D3AD" pageId="2" pageNumber="3" type="paralectotype">paralectotype</typeStatus>
:
<collectingCountry id="F37E766AFFC1E209FEBAFECD0311D3AD" box="[358,542,364,390]" name="Macedonia" pageId="2" pageNumber="3">MACEDONIA</collectingCountry>
: ɗ, “Maced. [
<collectingCountry id="F37E766AFFC1E209FD10FECD025BD3AD" box="[716,852,364,390]" name="Macedonia" pageId="2" pageNumber="3">Macedonia</collectingCountry>
] Perister alpin. 7. 14 Dr. Rambousek/
<taxonomicName id="4C694D79FFC1E209FED4FE350082D386" box="[264,397,404,429]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="13" phylum="Arthropoda" rank="species" species="bernhaueri">
<emphasis id="B91DEAE8FFC1E209FED4FE350082D386" box="[264,397,404,429]" italics="true" pageId="2" pageNumber="3">Bernhaueri</emphasis>
</taxonomicName>
Ramb. Cotypus/ Chicago NHM M. Bernhauer Collection” (
<collectionCode id="ED78AE3FFFC1E209FB8DFE3505A2D385" LSID="urn:lsid:biocol.org:col:34795" box="[1105,1197,404,430]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="2" pageNumber="3">FMNH</collectionCode>
).
</paragraph>
<paragraph id="8BD636FAFFC1E209FEE4FE1A05FDD065" blockId="2.[264,1324,283,1833]" pageId="2" pageNumber="3">
<emphasis id="B91DEAE8FFC1E209FEE4FE1A0337D3FE" bold="true" box="[312,568,443,469]" pageId="2" pageNumber="3">Additional material.</emphasis>
<collectingCountry id="F37E766AFFC1E209FD9DFE1D03B7D3FD" box="[577,696,444,470]" name="Greece" pageId="2" pageNumber="3">GREECE</collectingCountry>
: 1 ɗ,
<taxonomicName id="4C694D79FFC1E209FCDDFE1D02F8D3FD" authority="Verno, Vitsi" authorityName="Verno, Vitsi" box="[769,1015,444,470]" class="Insecta" family="Sphingidae" genus="Florina" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Flórina, Vérno, Vitsi</taxonomicName>
SW Drossopigi, northern slope, valley of small creek,
<taxonomicName id="4C694D79FFC1E209FDB1FE4503B9D3D6" box="[621,694,484,509]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="genus">
<emphasis id="B91DEAE8FFC1E209FDB1FE4503B9D3D6" box="[621,694,484,509]" italics="true" pageId="2" pageNumber="3">Fagus</emphasis>
</taxonomicName>
­forest, snowpatch,
<date id="FFD7103AFFC1E209FC7AFE450524D3D5" box="[934,1067,484,510]" pageId="2" pageNumber="3" value="2000-04-13">13.IV.2000</date>
, 1410 m, 40°3850N, 21°2445E, leg. Zerche; 2 Ψ,
<taxonomicName id="4C694D79FFC1E209FD80FDAD02C6D00D" authority="Verno, Vitsi S Florina" authorityName="Verno, Vitsi S Florina" box="[604,969,524,550]" class="Insecta" family="Sphingidae" genus="Florina" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Flórina, Vérno, Vitsi S Flórina</taxonomicName>
, northern slope,
<taxonomicName id="4C694D79FFC1E209FB51FDAD05D9D00E" box="[1165,1238,524,549]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="genus">
<emphasis id="B91DEAE8FFC1E209FB51FDAD05D9D00E" box="[1165,1238,524,549]" italics="true" pageId="2" pageNumber="3">Fagus</emphasis>
</taxonomicName>
­forest, snowpatch,
<date id="FFD7103AFFC1E209FE48FD950316D065" box="[404,537,564,590]" pageId="2" pageNumber="3" value="2000-04-13">13.IV.2000</date>
, 1660 m, 40°3750N, 21°2403E, leg. Zerche and Behne (
<collectionCode id="ED78AE3FFFC1E209FB73FD9505E9D065" LSID="urn:lsid:biocol.org:col:33211" box="[1199,1254,564,590]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="2" pageNumber="3">DEI</collectionCode>
).
</paragraph>
<paragraph id="8BD636FAFFC1E209FEE4FDFA059FD14D" blockId="2.[264,1324,283,1833]" pageId="2" pageNumber="3">
<emphasis id="B91DEAE8FFC1E209FEE4FDFA0356D05E" bold="true" box="[312,601,603,629]" pageId="2" pageNumber="3">
<typeStatus id="54D28858FFC1E209FEE4FDFA00BBD05E" box="[312,436,603,629]" pageId="2" pageNumber="3" type="lectotype">Lectotype</typeStatus>
designation.
</emphasis>
In the original description, which was published in two almost identical papers, in Czech and
<collectingCountry id="F37E766AFFC1E209FD0DFD250220D0B5" box="[721,815,644,670]" name="Germany" pageId="2" pageNumber="3">German</collectingCountry>
(
<bibRefCitation id="EFF84B0BFFC1E209FC9DFD25051CD0B5" author="Rambousek" box="[833,1043,644,670]" pageId="2" pageNumber="13" refString="Rambousek, F. (1915) Tenebrobius (nov. subg. Quediorum) bernhaueri, novy druh ze str. Makedonie. (Col. Staphyl.). C asopis C eske Spole c nosti Entomologicke, 12, 54 - 56." type="journal article" year="1915">Rambousek 1915</bibRefCitation>
) and in
<collectingCountry id="F37E766AFFC1E209FBA4FD2505D9D0B5" box="[1144,1238,644,670]" name="Germany" pageId="2" pageNumber="3">German</collectingCountry>
(
<bibRefCitation id="EFF84B0BFFC1E209FB34FD2500ACD0ED" author="Rambousek" pageId="2" pageNumber="13" refString="Rambousek, F. (1916) Ein neues Subgenus der Gattung Quedius. Coleopterologische Rundschau, 11 / 12 (1915), 130 - 132." type="journal article" year="1916">Rambousek 1916</bibRefCitation>
), no information about the number of
<typeStatus id="54D28858FFC1E209FCABFD0D02A5D0ED" box="[887,938,684,710]" pageId="2" pageNumber="3">type</typeStatus>
specimens was provided. I was able to locate two specimens (both males) whose identity and labels exactly match the data provided in the original description. One of these
<typeStatus id="54D28858FFC1E209FC8CFD5D02B9D13D" box="[848,950,764,790]" pageId="2" pageNumber="3" type="syntype">syntypes</typeStatus>
(deposited in
<collectionCode id="ED78AE3FFFC1E209FB81FD5D0591D13D" box="[1117,1182,764,790]" httpUri="http://grbio.org/cool/n1s9-2rrz" name="Slovak National Museum" pageId="2" pageNumber="3">SNM</collectionCode>
) with Rambouseks
<typeStatus id="54D28858FFC1E209FEADFC8500ABD115" box="[369,420,804,830]" pageId="2" pageNumber="3">type</typeStatus>
label is designated here as the
<typeStatus id="54D28858FFC1E209FCF6FC850296D115" box="[810,921,804,830]" pageId="2" pageNumber="3" type="lectotype">lectotype</typeStatus>
. The other
<typeStatus id="54D28858FFC1E209FBF4FC85058CD115" box="[1064,1155,804,830]" pageId="2" pageNumber="3" type="syntype">syntype</typeStatus>
(deposited in
<collectionCode id="ED78AE3FFFC1E209FED4FCED0050D14D" LSID="urn:lsid:biocol.org:col:34795" box="[264,351,844,870]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="2" pageNumber="3">FMNH</collectionCode>
) with Bernhauers identification and “cotype” label is a
<typeStatus id="54D28858FFC1E209FC36FCED0583D14D" box="[1002,1164,844,870]" pageId="2" pageNumber="3" type="paralectotype">paralectotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C3736571FFC1E209FEE4FCD503E7D502" pageId="2" pageNumber="3" type="discussion">
<paragraph id="8BD636FAFFC1E209FEE4FCD503E7D502" blockId="2.[264,1324,283,1833]" pageId="2" pageNumber="3">
<emphasis id="B91DEAE8FFC1E209FEE4FCD500CCD1A5" bold="true" box="[312,451,884,910]" pageId="2" pageNumber="3">Discussion.</emphasis>
This remarkable species, described from the Perister range in
<collectingCountry id="F37E766AFFC1E209FB75FCD50424D1A5" box="[1193,1323,884,910]" name="Macedonia" pageId="2" pageNumber="3">Macedonia</collectingCountry>
early in the 20th century (
<bibRefCitation id="EFF84B0BFFC1E209FDE6FC3E0202D192" author="Rambousek" box="[570,781,927,953]" pageId="2" pageNumber="13" refString="Rambousek, F. (1915) Tenebrobius (nov. subg. Quediorum) bernhaueri, novy druh ze str. Makedonie. (Col. Staphyl.). C asopis C eske Spole c nosti Entomologicke, 12, 54 - 56." type="journal article" year="1915">Rambousek 1915</bibRefCitation>
,
<bibRefCitation id="EFF84B0BFFC1E209FCC0FC3E0255D192" author="Rambousek" box="[796,858,927,953]" pageId="2" pageNumber="13" refString="Rambousek, F. (1916) Ein neues Subgenus der Gattung Quedius. Coleopterologische Rundschau, 11 / 12 (1915), 130 - 132." type="journal article" year="1916">1916</bibRefCitation>
), was collected again only recently by
<bibRefCitation id="EFF84B0BFFC1E209FED4FC660366D1CA" author="Assing" box="[264,617,967,993]" pageId="2" pageNumber="12" refString="Assing, V. &amp; Wunderle, P. (2001) On the Staphylinidae of Greece. II. New species and new records from central and northern Greece (Insecta: Coleoptera). Linzer biologische Beitrage, 33 (1), 103 - 136." type="journal article" year="2001">Assing and Wunderle (2001)</bibRefCitation>
.
<bibRefCitation id="EFF84B0BFFC1E209FDABFC660253D1CA" author="Rambousek" box="[631,860,967,993]" pageId="2" pageNumber="13" refString="Rambousek, F. (1915) Tenebrobius (nov. subg. Quediorum) bernhaueri, novy druh ze str. Makedonie. (Col. Staphyl.). C asopis C eske Spole c nosti Entomologicke, 12, 54 - 56." type="journal article" year="1915">Rambousek (1915)</bibRefCitation>
found this species in soil from under large stones deeply immersed in the ground, at the elevation of
<quantity id="4C919B1FFFC1E209FC2FFC4E0596D622" box="[1011,1177,1007,1033]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="2" pageNumber="3" unit="m" value="1900.0" valueMax="2000.0" valueMin="1800.0">18002000 m</quantity>
.
<bibRefCitation id="EFF84B0BFFC1E209FB79FC4E00DFD61A" author="Assing" pageId="2" pageNumber="12" refString="Assing, V. &amp; Wunderle, P. (2001) On the Staphylinidae of Greece. II. New species and new records from central and northern Greece (Insecta: Coleoptera). Linzer biologische Beitrage, 33 (1), 103 - 136." type="journal article" year="2001">Assing and Wunderle (2001)</bibRefCitation>
sifted
<taxonomicName id="4C694D79FFC1E209FDFFFBB60201D61B" box="[547,782,1047,1072]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="bernhaueri">
<emphasis id="B91DEAE8FFC1E209FDFFFBB60201D61B" box="[547,782,1047,1072]" italics="true" pageId="2" pageNumber="3">Quedius bernhaueri</emphasis>
</taxonomicName>
from deep leaf litter in beech forest with calcareous soil, in northern and northwestern exposition at altitudes of
<date id="FFD7103AFFC1E209FBF8FB9E05DCD672" box="[1060,1235,1087,1113]" pageId="2" pageNumber="3" value="1800" valueMax="1900">
1800 and
<quantity id="4C919B1FFFC1E209FB44FB9E05DCD672" box="[1176,1235,1087,1113]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="2" pageNumber="3" unit="m" value="1900.0">1900</quantity>
</date>
m near the peak of Oros Vitsi in Florina (
<collectingCountry id="F37E766AFFC1E209FD4FFBC603E2D6AA" box="[659,749,1127,1153]" name="Greece" pageId="2" pageNumber="3">Greece</collectingCountry>
). New findings (from the same region in
<collectingCountry id="F37E766AFFC1E209FB13FBC6042BD6AA" box="[1231,1316,1127,1153]" name="Greece" pageId="2" pageNumber="3">Greece</collectingCountry>
) listed in this paper fit the known narrow distribution of
<taxonomicName id="4C694D79FFC1E209FC73FB2E0557D683" box="[943,1112,1167,1192]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="bernhaueri">
<emphasis id="B91DEAE8FFC1E209FC73FB2E0557D683" box="[943,1112,1167,1192]" italics="true" pageId="2" pageNumber="3">Q. bernhaueri</emphasis>
</taxonomicName>
, though record it from a slightly lower elevation (down to
<quantity id="4C919B1FFFC1E209FD2DFB160240D6FA" box="[753,847,1207,1233]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="2" pageNumber="3" unit="m" value="1400.0">1400 m</quantity>
).
<bibRefCitation id="EFF84B0BFFC1E209FCB8FB1605B5D6FA" author="Assing" box="[868,1210,1207,1233]" pageId="2" pageNumber="12" refString="Assing, V. &amp; Wunderle, P. (2001) On the Staphylinidae of Greece. II. New species and new records from central and northern Greece (Insecta: Coleoptera). Linzer biologische Beitrage, 33 (1), 103 - 136." type="journal article" year="2001">Assing and Wunderle (2001)</bibRefCitation>
provided illustrations of the habitus and male genitalia, based on their newly collected specimens. My examination of the
<typeStatus id="54D28858FFC1E209FDC0FAA60340D70A" box="[540,591,1287,1313]" pageId="2" pageNumber="3">type</typeStatus>
specimens (see above) confirms that the non­type based identification by
<bibRefCitation id="EFF84B0BFFC1E209FE7DFA8E03F7D762" author="Assing" box="[417,760,1327,1353]" pageId="2" pageNumber="12" refString="Assing, V. &amp; Wunderle, P. (2001) On the Staphylinidae of Greece. II. New species and new records from central and northern Greece (Insecta: Coleoptera). Linzer biologische Beitrage, 33 (1), 103 - 136." type="journal article" year="2001">Assing and Wunderle (2001)</bibRefCitation>
is correct. Because of the peculiar habitus features of
<taxonomicName id="4C694D79FFC1E209FEB4FAF60300D75B" box="[360,527,1367,1392]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="bernhaueri">
<emphasis id="B91DEAE8FFC1E209FEB4FAF60300D75B" box="[360,527,1367,1392]" italics="true" pageId="2" pageNumber="3">Q. bernhaueri</emphasis>
</taxonomicName>
(apparently associated with its endogean bionomics), its taxonomic position within the genus
<taxonomicName id="4C694D79FFC1E209FD9DFADE03ACD7B3" box="[577,675,1407,1432]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B91DEAE8FFC1E209FD9DFADE03ACD7B3" box="[577,675,1407,1432]" italics="true" pageId="2" pageNumber="3">Quedius</emphasis>
</taxonomicName>
is dubious.
<bibRefCitation id="EFF84B0BFFC1E209FCE4FADE0515D7B2" author="Rambousek" box="[824,1050,1407,1433]" pageId="2" pageNumber="13" refString="Rambousek, F. (1915) Tenebrobius (nov. subg. Quediorum) bernhaueri, novy druh ze str. Makedonie. (Col. Staphyl.). C asopis C eske Spole c nosti Entomologicke, 12, 54 - 56." type="journal article" year="1915">Rambousek (1915)</bibRefCitation>
assigned it to his new subgenus of
<emphasis id="B91DEAE8FFC1E209FE42FA060391D7EB" box="[414,670,1447,1472]" italics="true" pageId="2" pageNumber="3">
<taxonomicName id="4C694D79FFC1E209FE42FA06030DD7EB" box="[414,514,1447,1472]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
,
<taxonomicName id="4C694D79FFC1E209FDD3FA060391D7EB" box="[527,670,1447,1472]" class="Insecta" family="Staphylinidae" genus="Tenebrobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Tenebrobius</taxonomicName>
</emphasis>
. Later authors, starting with
<bibRefCitation id="EFF84B0BFFC1E209FC28FA0605A7D7EA" author="Gridelli" box="[1012,1192,1447,1473]" pageId="2" pageNumber="13" refString="Gridelli, E. (1924) Studi sul genere Quedius Steph. (Col. Staph.). Secondo contributo. Specie della regione paleartica. Memorie della Societa Entomologica Italiana, 3 (1), 5 - 112." type="journal article" year="1924">Gridelli (1924)</bibRefCitation>
, referred it to the subgenus
<taxonomicName id="4C694D79FFC1E209FE12FA6E036CD7C3" box="[462,611,1487,1512]" class="Insecta" family="Staphylinidae" genus="Microsaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B91DEAE8FFC1E209FE12FA6E036CD7C3" box="[462,611,1487,1512]" italics="true" pageId="2" pageNumber="3">Microsaurus</emphasis>
</taxonomicName>
. My examination of
<taxonomicName id="4C694D79FFC1E209FCB8FA6E0503D7C3" box="[868,1036,1487,1512]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="bernhaueri">
<emphasis id="B91DEAE8FFC1E209FCB8FA6E0503D7C3" box="[868,1036,1487,1512]" italics="true" pageId="2" pageNumber="3">Q. bernhaueri</emphasis>
</taxonomicName>
indicates that it is only such character as relatively small eyes, which makes it similar with
<taxonomicName id="4C694D79FFC1E209FBE7FA5605DFD43B" box="[1083,1232,1527,1552]" class="Insecta" family="Staphylinidae" genus="Microsaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B91DEAE8FFC1E209FBE7FA5605DFD43B" box="[1083,1232,1527,1552]" italics="true" pageId="2" pageNumber="3">Microsaurus</emphasis>
</taxonomicName>
. At the same time, unlike most species of
<taxonomicName id="4C694D79FFC1E209FD45F9BE0222D413" box="[665,813,1567,1592]" class="Insecta" family="Staphylinidae" genus="Microsaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B91DEAE8FFC1E209FD45F9BE0222D413" box="[665,813,1567,1592]" italics="true" pageId="2" pageNumber="3">Microsaurus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C694D79FFC1E209FCE0F9BE02EDD413" box="[828,994,1567,1592]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="bernhaueri">
<emphasis id="B91DEAE8FFC1E209FCE0F9BE02EDD413" box="[828,994,1567,1592]" italics="true" pageId="2" pageNumber="3">Q. bernhaueri</emphasis>
</taxonomicName>
has only one pair of setiferous punctures situated in front of posterior margin of head and posteromediad of posterior frontal puncture. Given the somewhat artificial and unclear limits between the currently accepted subgenera of
<taxonomicName id="4C694D79FFC1E209FDC8F9360379D49B" box="[532,630,1687,1712]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B91DEAE8FFC1E209FDC8F9360379D49B" box="[532,630,1687,1712]" italics="true" pageId="2" pageNumber="3">Quedius</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFF84B0BFFC1E209FD59F936023ED49A" author="Smetana" box="[645,817,1687,1713]" pageId="2" pageNumber="13" refString="Smetana, A. (1988) Revision of the tribes Quediini and Atanygnathini. Part II. The Himalayan region (Coleoptera: Staphylinidae). Quaestiones Entomologicae, 24, 163 - 464." type="journal article" year="1988">Smetana 1988</bibRefCitation>
;
<bibRefCitation id="EFF84B0BFFC1E209FCE1F9360527D49A" author="Solodovnikov" box="[829,1064,1687,1713]" pageId="2" pageNumber="13" refString="Solodovnikov, A. (2002) Taxonomy and faunistics of some species of Quedius Stephens, 1829 from the Caucasus and Asia Minor (Coleoptera: Staphylinidae). Koleopterologische Rundschau, 72, 137 - 158." type="journal article" year="2002">Solodovnikov 2002</bibRefCitation>
), phylogenetic affiliations of this spectacular European species must be revisited on the basis of a broad phylogenetic study of
<taxonomicName id="4C694D79FFC1E209FE3EF946034BD52B" box="[482,580,1767,1792]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B91DEAE8FFC1E209FE3EF946034BD52B" box="[482,580,1767,1792]" italics="true" pageId="2" pageNumber="3">Quedius</emphasis>
</taxonomicName>
and allies. Before this is done, I retain
<taxonomicName id="4C694D79FFC1E209FB8AF946040BD52B" box="[1110,1284,1767,1792]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="bernhaueri">
<emphasis id="B91DEAE8FFC1E209FB8AF946040BD52B" box="[1110,1284,1767,1792]" italics="true" pageId="2" pageNumber="3">Q. bernhaueri</emphasis>
</taxonomicName>
in
<taxonomicName id="4C694D79FFC1E209FED4F8AE0092D503" box="[264,413,1807,1832]" class="Insecta" family="Staphylinidae" genus="Microsaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B91DEAE8FFC1E209FED4F8AE0092D503" box="[264,413,1807,1832]" italics="true" pageId="2" pageNumber="3">Microsaurus</emphasis>
</taxonomicName>
following previous authors.
</paragraph>
</subSubSection>
</treatment>
</document>