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<document id="7C1022123A81935820DBE6E2618347A4" ID-CLB-Dataset="39188" ID-DOI="10.11646/zootaxa.4105.6.1" ID-GBIF-Dataset="9f3191f8-cd86-4a17-ad5b-0c454e0acb8d" ID-ISSN="1175-5326" ID-Zenodo-Dep="271319" ID-ZooBank="DCB6A5BB-46C9-4D05-8B4A-C6E4CBABB6F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461796872515" checkinUser="plazi" docAuthor="Taylor, Robert W. &amp; Alpert, Gary D." docDate="2016" docId="03C087E7456FFFA2FF3CF8A2FA9A4B08" docLanguage="en" docName="zootaxa.4105.6.1.pdf.imf" docOrigin="Zootaxa 4105 (6)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Metapone krombeini M.R. Smith 1947" docType="treatment" docVersion="17" lastPageNumber="526" masterDocId="FFF9FF9F4577FFBBFFABFFA4FF9D4811" masterDocTitle="The myrmicine ant genus Metapone Forel (Hymenoptera: Formicidae): a global taxonomic review with descriptions of twelve new species" masterLastPageNumber="545" masterPageNumber="501" pageNumber="525" updateTime="1698662027110" updateUser="ExternalLinkService">
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<mods:title id="EDEEAAD410632D6B4134448AC27ED7A2">The myrmicine ant genus Metapone Forel (Hymenoptera: Formicidae): a global taxonomic review with descriptions of twelve new species</mods:title>
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<treatment id="03C087E7456FFFA2FF3CF8A2FA9A4B08" ID-DOI="http://doi.org/10.5281/zenodo.5641464" ID-GBIF-Taxon="119651718" ID-Zenodo-Dep="5641464" LSID="urn:lsid:plazi:treatment:03C087E7456FFFA2FF3CF8A2FA9A4B08" httpUri="http://treatment.plazi.org/id/03C087E7456FFFA2FF3CF8A2FA9A4B08" lastPageId="25" lastPageNumber="526" pageId="24" pageNumber="525">
<subSubSection id="C373657A456FFFA3FF3CF8A2FD3D4F31" box="[151,672,1798,1824]" pageId="24" pageNumber="525" type="nomenclature">
<paragraph id="8BD636F1456FFFA3FF3CF8A2FD3D4F31" blockId="24.[151,672,1798,1858]" box="[151,672,1798,1824]" pageId="24" pageNumber="525">
<heading id="D09E819D456FFFA3FF3CF8A2FD3D4F31" bold="true" box="[151,672,1798,1824]" fontSize="11" level="1" pageId="24" pageNumber="525" reason="1">
<emphasis id="B91DEAE3456FFFA3FF3CF8A2FD3D4F31" bold="true" box="[151,672,1798,1824]" pageId="24" pageNumber="525">
15.
<taxonomicName id="4C694D72456FFFA3FF68F8A2FD3D4F31" ID-CoL="42CPL" authority="M.R. Smith, 1947" authorityName="M.R. Smith" authorityYear="1947" box="[195,672,1798,1824]" class="Insecta" family="Formicidae" genus="Metapone" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="525" phylum="Arthropoda" rank="species" species="krombeini">
<emphasis id="B91DEAE3456FFFA3FF68F8A2FE234F31" bold="true" box="[195,446,1798,1824]" italics="true" pageId="24" pageNumber="525">Metapone krombeini</emphasis>
<bibRefCitation id="EFF84B00456FFFA3FE6EF8A2FD3D4F31" author="Smith" box="[453,672,1798,1824]" pageId="24" pageNumber="525" refString="Smith, M. R. (1947) A new species of Metapone Forel from New Guinea (Hymenoptera, Formicidae) Proceedings of the Entomological Society of Washington, 49, 75 - 77." type="journal article" year="1947">M.R. Smith, 1947</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C373657A456FFFA3FF3CF88DFEBA4F53" box="[151,295,1833,1858]" pageId="24" pageNumber="525" type="description">
<paragraph id="8BD636F1456FFFA3FF3CF88DFEBA4F53" blockId="24.[151,672,1798,1858]" box="[151,295,1833,1858]" pageId="24" pageNumber="525">
(
<figureCitation id="13522A74456FFFA3FF34F88DFF6F4F53" box="[159,242,1833,1858]" captionStart="FIGURES 71 75" captionStartId="26.[151,264,1910,1933]" captionTargetBox="[151,1436,193,1888]" captionTargetId="figure@26.[151,1436,193,1890]" captionTargetPageId="26" captionText="FIGURES 71 75. Metapone krombeini, holotype gyne; FIGURES 76 80. Worker, see text for dimensions." httpUri="https://zenodo.org/record/271335/files/figure.png" pageId="24" pageNumber="525">Figs 71</figureCitation>
80)
</paragraph>
</subSubSection>
<subSubSection id="C373657A456FFFA3FF3CF8D1FB344F9B" box="[151,1193,1905,1930]" pageId="24" pageNumber="525" type="reference_group">
<paragraph id="8BD636F1456FFFA3FF3CF8D1FB344F9B" blockId="24.[151,1193,1905,1930]" box="[151,1193,1905,1930]" pageId="24" pageNumber="525">
<treatmentCitationGroup id="AB7911DF456FFFA3FF3CF8D1FB344F9B" box="[151,1193,1905,1930]" pageId="24" pageNumber="525">
<taxonomicName id="4C694D72456FFFA3FF3CF8D1FDDA4F9B" ID-CoL="42CPL" authority="M.R. Smith, 1947:76" authorityName="M.R. Smith" authorityPageNumber="76" authorityYear="1947" box="[151,583,1908,1930]" class="Insecta" family="Formicidae" genus="Metapone" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="525" phylum="Arthropoda" rank="species" species="krombeini">
<emphasis id="B91DEAE3456FFFA3FF3CF8D1FEF54F9B" box="[151,360,1908,1930]" italics="true" pageId="24" pageNumber="525">Metapone krombeini</emphasis>
<treatmentCitation id="0AC810E0456FFFA3FEC5F8D1FDDA4F9B" author="Smith" box="[366,583,1908,1930]" page="76" pageId="24" pageNumber="525" year="1947">
<bibRefCitation id="EFF84B00456FFFA3FEC5F8D1FDB54F9B" author="Smith" box="[366,552,1908,1930]" pageId="24" pageNumber="525" refString="Smith, M. R. (1947) A new species of Metapone Forel from New Guinea (Hymenoptera, Formicidae) Proceedings of the Entomological Society of Washington, 49, 75 - 77." type="journal article" year="1947">M.R. Smith, 1947</bibRefCitation>
:76
</treatmentCitation>
</taxonomicName>
, gyne;
<typeStatus id="54D28853456FFFA3FD3BF8D1FD5F4F9B" box="[656,706,1909,1930]" pageId="24" pageNumber="525">Type</typeStatus>
locality: New
<collectingCountry id="F37E7661456FFFA3FCF3F8D0FC3E4F9B" box="[856,931,1908,1930]" name="Guinea-Bissau" pageId="24" pageNumber="525">Guinea</collectingCountry>
,
<emphasis id="B91DEAE3456FFFA3FC05F8D0FB8B4F9B" box="[942,1046,1908,1930]" italics="true" pageId="24" pageNumber="525">Milne Bay</emphasis>
[10°S, 150°E].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C373657A456FFFA2FF3CF812FF6249C7" lastPageId="25" lastPageNumber="526" pageId="24" pageNumber="525" type="materials_examined">
<paragraph id="8BD636F1456FFFA2FF3CF812FF6249C7" blockId="24.[151,1436,1974,2038]" lastBlockId="25.[151,1437,151,794]" lastPageId="25" lastPageNumber="526" pageId="24" pageNumber="525">
<emphasis id="B91DEAE3456FFFA3FF3CF812FE184FDE" bold="true" box="[151,389,1974,1999]" pageId="24" pageNumber="525">Material examined.</emphasis>
<collectingCountry id="F37E7661456FFFA3FE3AF813FDB34FDE" box="[401,558,1975,1999]" name="Indonesia" pageId="24" pageNumber="525">INDONESIA</collectingCountry>
: West
<collectingCountry id="F37E7661456FFFA3FD2BF813FD594FDF" box="[640,708,1975,1998]" name="Papua New Guinea" pageId="24" pageNumber="525">Papua</collectingCountry>
(as “New
<collectingCountry id="F37E7661456FFFA3FCE1F812FC024FDE" box="[842,927,1974,1999]" name="Guinea-Bissau" pageId="24" pageNumber="525">Guinea</collectingCountry>
: Neth”):
<emphasis id="B91DEAE3456FFFA3FBA5F813FBE04FDF" box="[1038,1149,1975,1998]" italics="true" pageId="24" pageNumber="525">River Tor</emphasis>
(
<emphasis id="B91DEAE3456FFFA3FB39F812FB454FDE" box="[1170,1240,1974,1999]" italics="true" pageId="24" pageNumber="525">mouth</emphasis>
),
<quantity id="4C919B14456FFFA3FB5FF812FAB84FDE" box="[1268,1317,1974,1999]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="24" pageNumber="525" unit="km" value="4.0">4km</quantity>
E of Hol Maffen [
<geoCoordinate id="EE5D5036456FFFA3FF57F87AFED84FE7" box="[252,325,2010,2038]" direction="south" orientation="latitude" pageId="24" pageNumber="525" precision="925" value="-1.9666667">1°58S</geoCoordinate>
x
<geoCoordinate id="EE5D5036456FFFA3FECEF87AFE504FE7" box="[357,461,2010,2038]" direction="east" orientation="longitude" pageId="24" pageNumber="525" precision="925" value="138.91667">138°55E</geoCoordinate>
] (T.C. Maa,
<date id="FFD71031456FFFA3FDC8F879FD794FE7" box="[611,740,2013,2038]" pageId="24" pageNumber="525" value="1959-07-04">VII-4-1959</date>
) alate gyne (LACMNH).
<collectingCountry id="F37E7661456FFFA3FBA4F87AFAB64FE7" box="[1039,1323,2014,2038]" name="Papua New Guinea" pageId="24" pageNumber="525">
<collectionCode id="ED78AE34456FFFA3FBA4F87AFBF54FE7" box="[1039,1128,2014,2038]" pageId="24" pageNumber="525">PAPUA</collectionCode>
<collectionCode id="ED78AE34456FFFA3FBDAF87AFB2C4FE4" box="[1137,1201,2014,2037]" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="24" pageNumber="525">NEW</collectionCode>
<collectionCode id="ED78AE34456FFFA3FB10F87AFAB64FE7" box="[1211,1323,2014,2038]" pageId="24" pageNumber="525">GUINEA</collectionCode>
</collectingCountry>
: Northern Province:
<emphasis id="B91DEAE3456EFFA2FEA0FF33FE1D48A1" box="[267,384,151,176]" italics="true" pageId="25" pageNumber="526">Milne Bay</emphasis>
, KB Mission (K.V. Krombein,
<date id="FFD71031456EFFA2FD40FF33FC3648A1" box="[747,939,151,176]" pageId="25" pageNumber="526" value="1944-03-26" valueMax="1944-03-28" valueMin="1944-03-26">2628 Mar 1944</date>
)
<typeStatus id="54D28853456EFFA2FC16FF33FB8248A1" box="[957,1055,151,176]" pageId="25" pageNumber="526" type="holotype">holotype</typeStatus>
alate gyne (USNMNH,
<typeStatus id="54D28853456EFFA2FA92FF3CFAED48A1" box="[1337,1392,152,176]" pageId="25" pageNumber="526">Type</typeStatus>
No 58003): Madang Province:
<emphasis id="B91DEAE3456EFFA2FE65FF1AFDCE48C6" box="[462,595,190,215]" italics="true" pageId="25" pageNumber="526">Ohu village</emphasis>
,
<geoCoordinate id="EE5D5036456EFFA2FDC9FF1AFD3148C6" box="[610,684,187,215]" direction="south" orientation="latitude" pageId="25" pageNumber="526" precision="925" value="-5.4333334">5°26S</geoCoordinate>
<geoCoordinate id="EE5D5036456EFFA2FD1FFF1AFC8748C6" box="[692,794,187,215]" direction="east" orientation="longitude" pageId="25" pageNumber="526" precision="925" value="146.13333">145°68E</geoCoordinate>
(M. Janda, hand collection ex rotten log,
<quantity id="4C919B14456EFFA2FB52FF1BFAEE48C6" box="[1273,1395,191,215]" metricMagnitude="2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="25" pageNumber="526" unit="m" value="150.0" valueMax="200.0" valueMin="100.0">100200m</quantity>
,
<date id="FFD71031456EFFA2FA2BFF1BFF2148EA" pageId="25" pageNumber="526" value="1922-08">22 viii</date>
200'''2)
<collectionCode id="ED78AE34456EFFA2FEB7FF47FEF848EA" LSID="urn:lsid:biocol.org:col:32981" box="[284,357,227,251]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="25" pageNumber="526">ANIC</collectionCode>
: West New
<collectingCountry id="F37E7661456EFFA2FE41FF46FDA548EA" box="[490,568,226,251]" name="United Kingdom" pageId="25" pageNumber="526">Britain</collectingCountry>
Province:
<emphasis id="B91DEAE3456EFFA2FD19FF46FCD148EA" box="[690,844,226,251]" italics="true" pageId="25" pageNumber="526">
Umboi
<collectingCountry id="F37E7661456EFFA2FCADFF46FCD148EA" box="[774,844,226,251]" name="Iceland" pageId="25" pageNumber="526">Island</collectingCountry>
</emphasis>
[
<geoCoordinate id="EE5D5036456EFFA2FCF0FF46FC3748EA" box="[859,938,226,251]" direction="south" orientation="latitude" pageId="25" pageNumber="526" precision="925" value="-5.5333333">5°32S</geoCoordinate>
,
<geoCoordinate id="EE5D5036456EFFA2FC1DFF46FBB948EA" box="[950,1060,226,251]" direction="east" orientation="longitude" pageId="25" pageNumber="526" precision="925" value="147.86667">147°52E</geoCoordinate>
]:
<emphasis id="B91DEAE3456EFFA2FB9CFF46FA9748EA" box="[1079,1290,226,251]" italics="true" pageId="25" pageNumber="526">
<quantity id="4C919B14456EFFA2FB9CFF46FBF848EA" box="[1079,1125,226,251]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="25" pageNumber="526" unit="km" value="1.0">1km</quantity>
N of Awelkom
</emphasis>
,
<quantity id="4C919B14456EFFA2FAB3FF46FAC548EA" box="[1304,1368,226,251]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="25" pageNumber="526" unit="m" value="600.0">600m</quantity>
(G.A. Samuelson,
<date id="FFD71031456EFFA2FE88FEA2FE5F490E" box="[291,450,262,287]" pageId="25" pageNumber="526" value="1967-02-21" valueMax="1967-02-28" valueMin="1967-02-21">2128.II.1967</date>
) alate gyne (LACMNH). The River Tor gyne is badly damaged by pinning, with the detached right forewing, postpetiole and gaster glued to the data label. It closely matches the
<taxonomicName id="4C694D72456EFFA2FB11FE8EFAB64952" box="[1210,1323,298,323]" class="Insecta" family="Formicidae" genus="Metapone" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="526" phylum="Arthropoda" rank="species" species="krombeini">
<emphasis id="B91DEAE3456EFFA2FB11FE8EFAB64952" box="[1210,1323,298,323]" italics="true" pageId="25" pageNumber="526">krombeini</emphasis>
</taxonomicName>
<typeStatus id="54D28853456EFFA2FA98FE8EFA054952" box="[1331,1432,298,323]" pageId="25" pageNumber="526" type="holotype">holotype</typeStatus>
, and has a label by K. Yasumatsu identifying it as that species. An
<collectionCode id="ED78AE34456EFFA2FC0AFEEBFC6D4976" box="[929,1008,335,359]" pageId="25" pageNumber="526">MCZC</collectionCode>
alate gyne from
<collectingCountry id="F37E7661456EFFA2FB11FEEBFACA4976" box="[1210,1367,335,359]" name="Indonesia" pageId="25" pageNumber="526">INDONESIA</collectingCountry>
: West
<collectingCountry id="F37E7661456EFFA2FF3CFED2FF46499C" box="[151,219,374,397]" name="Papua New Guinea" pageId="25" pageNumber="526">Papua</collectingCountry>
(as “New
<collectingCountry id="F37E7661456EFFA2FEFFFED1FE34499F" box="[340,425,373,398]" name="Guinea-Bissau" pageId="25" pageNumber="526">Guinea</collectingCountry>
: Neth”) Vogelkop: Manokwari (“Menoekwari” [5°20', 134°05'],
<quantity id="4C919B14456EFFA2FBD3FED1FB36499F" box="[1144,1195,373,398]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="25" pageNumber="526" unit="m" value="75.0">75m</quantity>
(D. Elmo Hardy,
<date id="FFD71031456EFFA2FADEFED1FF6849A3" pageId="25" pageNumber="526" value="1957-07-18">vii- 18-1957</date>
) is provisionally identified as
<taxonomicName id="4C694D72456EFFA2FDE6FE3EFD7E49A3" box="[589,739,409,434]" class="Insecta" family="Formicidae" genus="Metapone" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="526" phylum="Arthropoda" rank="species" species="krombeini">
<emphasis id="B91DEAE3456EFFA2FDE6FE3EFD7E49A3" box="[589,739,409,434]" italics="true" pageId="25" pageNumber="526">M. krombeini</emphasis>
</taxonomicName>
. It generally agrees with the
<typeStatus id="54D28853456EFFA2FB86FE3EFBC049A3" box="[1069,1117,410,434]" pageId="25" pageNumber="526">type</typeStatus>
but the proboscis is slightly different.
</paragraph>
</subSubSection>
<subSubSection id="C373657A456EFFA2FF6CFE45FDAC4A53" pageId="25" pageNumber="526" type="description">
<paragraph id="8BD636F1456EFFA2FF6CFE45FDAC4A53" blockId="25.[151,1437,151,794]" pageId="25" pageNumber="526">
<emphasis id="B91DEAE3456EFFA2FF6CFE45FE3F49EB" bold="true" box="[199,418,481,506]" pageId="25" pageNumber="526">Worker diagnosis.</emphasis>
General and diagnostic features as in original description (
<bibRefCitation id="EFF84B00456EFFA2FBECFE45FB4949EB" author="Smith" box="[1095,1236,481,506]" pageId="25" pageNumber="526" refString="Smith, M. R. (1947) A new species of Metapone Forel from New Guinea (Hymenoptera, Formicidae) Proceedings of the Entomological Society of Washington, 49, 75 - 77." type="journal article" year="1947">Smith, 1947</bibRefCitation>
) and Figs 7680. Similar to gynes from New
<collectingCountry id="F37E7661456EFFA2FE43FDA1FDA44A0F" box="[488,569,517,542]" name="Guinea-Bissau" pageId="25" pageNumber="526">Guinea</collectingCountry>
and
<collectingCountry id="F37E7661456EFFA2FDD1FDA1FD654A0F" box="[634,760,517,542]" name="Micronesia" pageId="25" pageNumber="526">Micronesia</collectingCountry>
assigned below to the smaller
<taxonomicName id="4C694D72456EFFA2FBDBFDA2FB524A0F" box="[1136,1231,517,542]" class="Insecta" family="Formicidae" genus="Metapone" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="526" phylum="Arthropoda" rank="species" species="truki">
<emphasis id="B91DEAE3456EFFA2FBDBFDA2FB524A0F" box="[1136,1231,517,542]" italics="true" pageId="25" pageNumber="526">M. truki</emphasis>
</taxonomicName>
, with differences itemised in the key to species above.
</paragraph>
</subSubSection>
<subSubSection id="C373657A456EFFA2FF6CFDE9FA9A4B08" pageId="25" pageNumber="526" type="diagnosis">
<paragraph id="8BD636F1456EFFA2FF6CFDE9FDE54AE7" blockId="25.[151,1437,151,794]" pageId="25" pageNumber="526">
<emphasis id="B91DEAE3456EFFA2FF6CFDE9FE1B4A77" bold="true" box="[199,390,589,614]" pageId="25" pageNumber="526">Gyne diagnosis.</emphasis>
General features as in original description (
<bibRefCitation id="EFF84B00456EFFA2FC2BFDE9FB904A77" author="Smith" box="[896,1037,589,614]" pageId="25" pageNumber="526" refString="Smith, M. R. (1947) A new species of Metapone Forel from New Guinea (Hymenoptera, Formicidae) Proceedings of the Entomological Society of Washington, 49, 75 - 77." type="journal article" year="1947">Smith, 1947</bibRefCitation>
) and
<figureCitation id="13522A74456EFFA2FBFBFDE9FB4E4A77" box="[1104,1235,589,614]" captionStart="FIGURES 71 75" captionStartId="26.[151,264,1910,1933]" captionTargetBox="[151,1436,193,1888]" captionTargetId="figure@26.[151,1436,193,1890]" captionTargetPageId="26" captionText="FIGURES 71 75. Metapone krombeini, holotype gyne; FIGURES 76 80. Worker, see text for dimensions." httpUri="https://zenodo.org/record/271335/files/figure.png" pageId="25" pageNumber="526">Figs 7175</figureCitation>
. Similar to gynes from New
<collectingCountry id="F37E7661456EFFA2FEBFFDD5FEF84A9B" box="[276,357,625,650]" name="Guinea-Bissau" pageId="25" pageNumber="526">Guinea</collectingCountry>
and
<collectingCountry id="F37E7661456EFFA2FE0BFDD5FD834A9B" box="[416,542,625,650]" name="Micronesia" pageId="25" pageNumber="526">Micronesia</collectingCountry>
assigned below to the smaller
<taxonomicName id="4C694D72456EFFA2FC28FDD6FC424A9B" box="[899,991,625,650]" class="Insecta" family="Formicidae" genus="Metapone" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="526" phylum="Arthropoda" rank="species" species="truki">
<emphasis id="B91DEAE3456EFFA2FC28FDD6FC424A9B" box="[899,991,625,650]" italics="true" pageId="25" pageNumber="526">M. truki</emphasis>
</taxonomicName>
. The two species differ as specified in couplet 5 of the key to Melanesian species above. Colour dark brown, the gaster reddish-brown, TL: 3.36, HL: 1.72, HW (across eyes): 1.22, CI: 71, CpL: 0.65, CpI: 38, WL: 2.84, PetL: 0.78, PetW: 0.62, PetH: 0.98, PpetL: 0.64, PpetW: 0.79, PpetH: 0.74, GW: 1.34.
</paragraph>
<paragraph id="8BD636F1456EFFA2FF6CFCA5FA9A4B08" blockId="25.[151,1437,151,794]" box="[199,1287,769,794]" pageId="25" pageNumber="526">
Five New Guinean workers and a male (LACMNH) are provisionally identified as
<taxonomicName id="4C694D72456EFFA2FBC1FCA6FA9D4B0B" box="[1130,1280,769,794]" class="Insecta" family="Formicidae" genus="Metapone" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="526" phylum="Arthropoda" rank="species" species="krombeini">
<emphasis id="B91DEAE3456EFFA2FBC1FCA6FA9D4B0B" box="[1130,1280,769,794]" italics="true" pageId="25" pageNumber="526">M. krombeini</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>