337 lines
42 KiB
XML
337 lines
42 KiB
XML
<document id="4E0E8A9642D39A4494B8630BDC32F6FE" ID-CLB-Dataset="183564" ID-DOI="10.11646/zootaxa.5205.4.2" ID-GBIF-Dataset="b9f30bf3-81d2-4052-8062-0a3f0b8d3288" ID-ISSN="1175-5326" ID-Zenodo-Dep="7306960" ID-ZooBank="AEB5F4C6-254C-4A22-86F2-B7C5697A72A7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1667981219887" checkinUser="plazi" docAuthor="Pham, Phu Van, Keith, Denis, Pham, Nhi Thi & Cao, Nga Quynh Thi" docDate="2022" docId="03C087917C11FF9DF8EEFE74FD11FE1F" docLanguage="en" docName="zootaxa.5205.4.2.pdf" docOrigin="Zootaxa 5205 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Miridiba (Miridiba) quangnamensis P. V. Pham & Keith 2022, new species" docType="treatment" docUuid="8E9A0286-D3C8-4C19-83B2-56A4E09585A9" docUuidSource="ZooBank" docVersion="6" lastPageNumber="337" masterDocId="FFF9FFE97C14FF9BF879FFC2FFCBFF96" masterDocTitle="Threenew species ofthe genus MiridibaReitter, 1902 (Scarabaeidae: Melolonthinae) from Vietnam along with a new country record" masterLastPageNumber="342" masterPageNumber="331" pageNumber="336" updateTime="1699411074940" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C5BE1A2D8CC2D1E465D275DF412CE99A">Threenew species ofthe genus MiridibaReitter, 1902 (Scarabaeidae: Melolonthinae) from Vietnam along with a new country record</mods:title>
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<mods:name id="010684F0B846CF0174E6D2F1041EE4D2" type="personal">
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<mods:namePart id="709E6CECEF89996C82A5DE3F672C8478">Pham, Phu Van</mods:namePart>
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<mods:affiliation id="5465D593CCF6B242A55B9F995F02E94B">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Techlonogy, 18 Hoang Quoc Viet, Cau Giay, Hanoi, Vietnam.</mods:affiliation>
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<mods:namePart id="B3312EC20B0155C6009D319C5FE7BFDB">Keith, Denis</mods:namePart>
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<mods:affiliation id="A51AD52B59248CD602E95393E612D034">120, rue Gabriel Péri, F- 28000 Chartres, France.</mods:affiliation>
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<mods:namePart id="64EAB2F276D564999549CEEA88437BB0">Pham, Nhi Thi</mods:namePart>
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<mods:affiliation id="C3AED8355C1146C6E134596C4F10022F">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Techlonogy, 18 Hoang Quoc Viet, Cau Giay, Hanoi, Vietnam. & Graduate University of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Hanoi, Vietnam</mods:affiliation>
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<mods:namePart id="343EBAD1A38B269D41EE00A208C59634">Cao, Nga Quynh Thi</mods:namePart>
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<mods:affiliation id="CC2485AC278BB60CA61C2D6BC59AA637">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Techlonogy, 18 Hoang Quoc Viet, Cau Giay, Hanoi, Vietnam.</mods:affiliation>
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<mods:title id="72BB02BA143B77DA648861D88AEF95D0">Zootaxa</mods:title>
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<mods:date id="0F370DD31F73D9568C8639B0CD816280">2022</mods:date>
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<mods:number id="15FDF5F4DEA7343B848DAE59E265DC78">2022-11-09</mods:number>
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<treatment id="03C087917C11FF9DF8EEFE74FD11FE1F" ID-DOI="http://doi.org/10.5281/zenodo.7306956" ID-GBIF-Taxon="204721519" ID-Zenodo-Dep="7306956" LSID="urn:lsid:zoobank.org:act:8E9A0286-D3C8-4C19-83B2-56A4E09585A9" httpUri="http://treatment.plazi.org/id/03C087917C11FF9DF8EEFE74FD11FE1F" lastPageId="6" lastPageNumber="337" pageId="5" pageNumber="336">
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<taxonomicName id="4C694D047C11FF9EF8EEFE74FC89FE46" authority="P.V. Pham & Keith" authorityName="P. V. Pham & Keith" authorityYear="2022" box="[151,834,438,465]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="species" species="quangnamensis" status="sp. nov." subGenus="Miridiba">
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<emphasis id="B91DEA957C11FF9EF8EEFE74FECAFE47" bold="true" box="[151,257,438,465]" italics="true" pageId="5" pageNumber="336">Miridiba</emphasis>
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(
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<emphasis id="B91DEA957C11FF9EF96BFE74FEB7FE47" bold="true" box="[274,380,438,465]" italics="true" pageId="5" pageNumber="336">Miridiba</emphasis>
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)
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<emphasis id="B91DEA957C11FF9EF9F4FE75FD87FE47" bold="true" box="[397,588,439,465]" italics="true" pageId="5" pageNumber="336">quangnamensis</emphasis>
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P.V. Pham & Keith
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</taxonomicName>
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,
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<taxonomicNameLabel id="A22E57EE7C11FF9EFB36FE75FC14FE46" box="[847,991,438,465]" pageId="5" pageNumber="336" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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urn:lsid:zoobank.org:act:
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<subSubSection id="C373650C7C11FF9EF8EEFE3BFEE1FD85" box="[151,298,505,531]" pageId="5" pageNumber="336" type="description">
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<paragraph id="8BD636877C11FF9EF8EEFE3BFEE1FD85" blockId="5.[151,991,438,531]" box="[151,298,505,531]" pageId="5" pageNumber="336">
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(
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<figureCitation id="13522A027C11FF9EF8E6FE3BFF2CFD85" box="[159,231,505,531]" captionStart="FIGURE 3" captionStartId="6.[151,250,1848,1872]" captionTargetBox="[152,1437,418,1807]" captionTargetId="figure-114@6.[151,1437,417,1825]" captionTargetPageId="6" captionText="FIGURE 3. Miridiba quangnamensis P.V. Pham & Keith, new species. a, habitus; b, pronotum and head; c, meso- and metasternum; d, scutellum; e, foretibia; f, hindtibia; g, pygidium." figureDoi="http://doi.org/10.5281/zenodo.7306968" httpUri="https://zenodo.org/record/7306968/files/figure.png" pageId="5" pageNumber="336">Figs 3</figureCitation>
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,
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<figureCitation id="13522A027C11FF9EF88DFE3BFEC9FD85" box="[244,258,505,531]" captionStart="FIGURE 4" captionStartId="7.[151,250,1184,1208]" captionTargetBox="[162,1426,227,1113]" captionTargetId="figure-18@7.[162,1426,181,1160]" captionTargetPageId="7" captionText="FIGURE 4. Male genitalia of Miridiba apicespinosa P.V. Pham & Keith, new species (a–c), Miridiba ngoclinhensis P.V. Pham & Keith, new species (d–f) and Miridiba quangnamensis P.V. Pham & Keith, new species (g–i). a, d, g, parameres in lateral view; b, e, h, parameres in dorsal view; c, f, i, parameres in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7306970" httpUri="https://zenodo.org/record/7306970/files/figure.png" pageId="5" pageNumber="336">4</figureCitation>
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g-i)
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</paragraph>
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</subSubSection>
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<subSubSection id="C373650C7C11FF9EF8EEFD83FD9EFDE9" pageId="5" pageNumber="336" type="materials_examined">
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<paragraph id="8BD636877C11FF9EF8EEFD83FD9EFDE9" blockId="5.[151,1437,577,2008]" pageId="5" pageNumber="336">
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<emphasis id="B91DEA957C11FF9EF8EEFD83FE8CFDCD" bold="true" box="[151,327,577,603]" pageId="5" pageNumber="336">Type material.</emphasis>
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<materialsCitation id="3B013CDA7C11FF9EF936FD83FD9EFDE9" ID-GBIF-Occurrence="3962977311" collectingDate="2005-04-28" collectionCode="IEBR" county="Nam Giang District" location="Song Thanh" municipality="Cha Vai" pageId="5" pageNumber="336" specimenCount="1" stateProvince="Quang Nam" typeStatus="holotype">
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<emphasis id="B91DEA957C11FF9EF936FD83FE0DFDCD" bold="true" box="[335,454,577,603]" pageId="5" pageNumber="336">
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<typeStatus id="54D288257C11FF9EF936FD83FE0BFDCD" box="[335,448,577,603]" pageId="5" pageNumber="336" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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male (
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<collectionCode id="ED78AE427C11FF9EFA6CFD83FD91FDCD" box="[533,602,577,603]" pageId="5" pageNumber="336">IEBR</collectionCode>
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),
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<collectingRegion id="49ADF8657C11FF9EFA09FD83FCA3FDCD" box="[624,872,577,604]" country="Vietnam" name="Quang Nam" pageId="5" pageNumber="336">Quang Nam Province</collectingRegion>
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,
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<collectingCounty id="62B74E0B7C11FF9EFB0CFD80FB92FDCD" box="[885,1113,577,604]" pageId="5" pageNumber="336">Nam Giang District</collectingCounty>
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,
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<collectingMunicipality id="6BB2ACFD7C11FF9EFC1CFD83FB08FDCD" box="[1125,1219,577,603]" pageId="5" pageNumber="336">Cha Vai</collectingMunicipality>
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,
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<location id="8EB6605C7C11FF9EFCA9FD83FA97FDCD" LSID="urn:lsid:plazi:treatment:03C087917C11FF9DF8EEFE74FD11FE1F:8EB6605C7C11FF9EFCA9FD83FA97FDCD" box="[1232,1372,577,603]" county="Nam Giang District" municipality="Cha Vai" name="Song Thanh" pageId="5" pageNumber="336" stateProvince="Quang Nam">Song Thanh</location>
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,
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<collectingMethod id="52284E907C11FF9EFD10FD83FF0EFD16" pageId="5" pageNumber="336">light trap</collectingMethod>
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,
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<date id="FFD710477C11FF9EF8A8FDA7FE82FD16" box="[209,329,613,640]" pageId="5" pageNumber="336" value="2005-04-28">
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<collectingDate id="EF93E9AF7C11FF9EF8A8FDA7FE82FD16" box="[209,329,613,640]" pageId="5" pageNumber="336" value="2005-04-28">
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28.iv.
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<quantity id="4C919B627C11FF9EF975FDA7FE82FD16" box="[268,329,613,640]" metricMagnitude="6" metricUnit="m" metricValue="2.0055199999999997" pageId="5" pageNumber="336" unit="m" value="2005520.0">2005</quantity>
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</collectingDate>
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</date>
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, 520 m above sea level.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C373650C7C11FF9EF8BEFD4BFE69F8B5" pageId="5" pageNumber="336" type="description">
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<paragraph id="8BD636877C11FF9EF8BEFD4BFF34FC99" blockId="5.[151,1437,577,2008]" pageId="5" pageNumber="336">
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<emphasis id="B91DEA957C11FF9EF8BEFD4BFD91FD35" bold="true" box="[199,602,649,675]" pageId="5" pageNumber="336">
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Description of the
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<typeStatus id="54D288257C11FF9EF9D9FD4BFDCCFD35" box="[416,519,649,675]" pageId="5" pageNumber="336" type="holotype">holotype</typeStatus>
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(male)
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</emphasis>
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: Body length:
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<quantity id="4C919B627C11FF9EFA87FD4BFCAAFD32" box="[766,865,649,676]" metricMagnitude="-2" metricUnit="m" metricValue="1.63" pageId="5" pageNumber="336" unit="mm" value="16.3">16.3 mm</quantity>
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(from anterior margin of clypeus to apex of elytra); body width:
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<quantity id="4C919B627C11FF9EF95DFD6CFEB2FD5E" box="[292,377,686,712]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="5" pageNumber="336" unit="mm" value="7.7">7.7 mm</quantity>
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||
(width across humeri). Color dark reddish brown on head, pronotum, and scutellum; yellowish brown to dark reddish brown on elytra, ventral surface, and legs; moderately shiny; dorsal surface almost glabrous (
|
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<figureCitation id="13522A027C11FF9EF8E6FD37FF3AFC99" box="[159,241,757,783]" captionStart="FIGURE 3" captionStartId="6.[151,250,1848,1872]" captionTargetBox="[152,1437,418,1807]" captionTargetId="figure-114@6.[151,1437,417,1825]" captionTargetPageId="6" captionText="FIGURE 3. Miridiba quangnamensis P.V. Pham & Keith, new species. a, habitus; b, pronotum and head; c, meso- and metasternum; d, scutellum; e, foretibia; f, hindtibia; g, pygidium." figureDoi="http://doi.org/10.5281/zenodo.7306968" httpUri="https://zenodo.org/record/7306968/files/figure.png" pageId="5" pageNumber="336">Fig. 3a</figureCitation>
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).
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</paragraph>
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<paragraph id="8BD636877C11FF9EF8BEFCDBFC51FC71" blockId="5.[151,1437,577,2008]" pageId="5" pageNumber="336">
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<emphasis id="B91DEA957C11FF9EF8BEFCDBFEC5FCA5" bold="true" box="[199,270,793,819]" pageId="5" pageNumber="336">Head.</emphasis>
|
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Surface densely punctate; punctures large; clypeus transverse, with apex moderately bilobed and reflexed, widened at base, length five times greater wide (
|
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<figureCitation id="13522A027C11FF9EFAC8FCFFFCCFFCC1" box="[689,772,829,855]" captionStart="FIGURE 3" captionStartId="6.[151,250,1848,1872]" captionTargetBox="[152,1437,418,1807]" captionTargetId="figure-114@6.[151,1437,417,1825]" captionTargetPageId="6" captionText="FIGURE 3. Miridiba quangnamensis P.V. Pham & Keith, new species. a, habitus; b, pronotum and head; c, meso- and metasternum; d, scutellum; e, foretibia; f, hindtibia; g, pygidium." figureDoi="http://doi.org/10.5281/zenodo.7306968" httpUri="https://zenodo.org/record/7306968/files/figure.png" pageId="5" pageNumber="336">Fig. 3b</figureCitation>
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||
). Clypeofrontal suture visible, moderately sinuated. Frontal carina moderately raised, weakly curved backwards, reaching periocular margins (
|
||
<figureCitation id="13522A027C11FF9EFC56FCA3FB48FCED" box="[1071,1155,865,891]" captionStart="FIGURE 3" captionStartId="6.[151,250,1848,1872]" captionTargetBox="[152,1437,418,1807]" captionTargetId="figure-114@6.[151,1437,417,1825]" captionTargetPageId="6" captionText="FIGURE 3. Miridiba quangnamensis P.V. Pham & Keith, new species. a, habitus; b, pronotum and head; c, meso- and metasternum; d, scutellum; e, foretibia; f, hindtibia; g, pygidium." figureDoi="http://doi.org/10.5281/zenodo.7306968" httpUri="https://zenodo.org/record/7306968/files/figure.png" pageId="5" pageNumber="336">Fig. 3b</figureCitation>
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||
). Basal area of head with punctures smaller than those on forehead, punctures with short erect setae (
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<figureCitation id="13522A027C11FF9EFB99FC47FBFFFC09" box="[992,1076,901,927]" captionStart="FIGURE 3" captionStartId="6.[151,250,1848,1872]" captionTargetBox="[152,1437,418,1807]" captionTargetId="figure-114@6.[151,1437,417,1825]" captionTargetPageId="6" captionText="FIGURE 3. Miridiba quangnamensis P.V. Pham & Keith, new species. a, habitus; b, pronotum and head; c, meso- and metasternum; d, scutellum; e, foretibia; f, hindtibia; g, pygidium." figureDoi="http://doi.org/10.5281/zenodo.7306968" httpUri="https://zenodo.org/record/7306968/files/figure.png" pageId="5" pageNumber="336">Fig. 3b</figureCitation>
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||
). Antenna with 9 antennomeres, antennal club composed of 3 antennomeres, club longer than length of antennomeres 2–6 combined (2.5/
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<quantity id="4C919B627C11FF9EFD4EFC6BFA5BFC55" box="[1335,1424,937,964]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="5" pageNumber="336" unit="mm" value="1.7">1.7 mm</quantity>
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). Last segment of maxillary palp elongated, dorsal surface with callus.
|
||
</paragraph>
|
||
<paragraph id="8BD636877C11FF9EF8BEFC33FC01FBC5" blockId="5.[151,1437,577,2008]" pageId="5" pageNumber="336">
|
||
<emphasis id="B91DEA957C11FF9EF8BEFC33FE8CFB9D" bold="true" box="[199,327,1009,1035]" pageId="5" pageNumber="336">Pronotum.</emphasis>
|
||
Transversal, widest at middle; anterolateral angles moderately protuberant; basolateral angles round; lateral margin smooth; anterior margin completely bordered; basal margin finely flanged and smooth, without pubescence; setiferous punctures smaller than those on clypeus (
|
||
<figureCitation id="13522A027C11FF9EFB10FBFBFC76FBC5" box="[873,957,1081,1107]" captionStart="FIGURE 3" captionStartId="6.[151,250,1848,1872]" captionTargetBox="[152,1437,418,1807]" captionTargetId="figure-114@6.[151,1437,417,1825]" captionTargetPageId="6" captionText="FIGURE 3. Miridiba quangnamensis P.V. Pham & Keith, new species. a, habitus; b, pronotum and head; c, meso- and metasternum; d, scutellum; e, foretibia; f, hindtibia; g, pygidium." figureDoi="http://doi.org/10.5281/zenodo.7306968" httpUri="https://zenodo.org/record/7306968/files/figure.png" pageId="5" pageNumber="336">Fig. 3b</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD636877C11FF9EF8BEFB9FFEF2FB0D" blockId="5.[151,1437,577,2008]" pageId="5" pageNumber="336">
|
||
<emphasis id="B91DEA957C11FF9EF8BEFB9FFEF4FBE1" bold="true" box="[199,319,1117,1143]" pageId="5" pageNumber="336">Scutellum</emphasis>
|
||
. Triangular, flat; strongly and densely puncate except on side margins; nearly two times wider than long (
|
||
<figureCitation id="13522A027C11FF9EF8A1FB43FEE7FB0D" box="[216,300,1153,1179]" captionStart="FIGURE 3" captionStartId="6.[151,250,1848,1872]" captionTargetBox="[152,1437,418,1807]" captionTargetId="figure-114@6.[151,1437,417,1825]" captionTargetPageId="6" captionText="FIGURE 3. Miridiba quangnamensis P.V. Pham & Keith, new species. a, habitus; b, pronotum and head; c, meso- and metasternum; d, scutellum; e, foretibia; f, hindtibia; g, pygidium." figureDoi="http://doi.org/10.5281/zenodo.7306968" httpUri="https://zenodo.org/record/7306968/files/figure.png" pageId="5" pageNumber="336">Fig. 3d</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD636877C11FF9EF8BEFB67FAC2FB72" blockId="5.[151,1437,577,2008]" pageId="5" pageNumber="336">
|
||
<emphasis id="B91DEA957C11FF9EF8BEFB67FED1FB29" bold="true" box="[199,282,1189,1215]" pageId="5" pageNumber="336">Elytra.</emphasis>
|
||
Elytra nearly parallel, slightly widened posteriorly. Punctures dense near the base, less posteriad; base of elytra bearing several long setae. Epipleura end at one fifth posteriorly bearing a row of long setae.
|
||
</paragraph>
|
||
<paragraph id="8BD636877C11FF9EF8BEFB2FFD28FA49" blockId="5.[151,1437,577,2008]" pageId="5" pageNumber="336">
|
||
<emphasis id="B91DEA957C11FF9EF8BEFB2FFECDFA91" bold="true" box="[199,262,1261,1287]" pageId="5" pageNumber="336">Legs.</emphasis>
|
||
Protibiae tridentate, teeth equidistant, tip of teeth moderately round; basal tooth small, middle tooth two third as apical teeth; internal apical spur inserted in front of middle tooth, reaching basal one-third of protarsomere 1; dorsal carina of protibia reaching the apex of apical tooth, and with a branch reaching the apex of second tooth (
|
||
<figureCitation id="13522A027C11FF9EF8E6FA9BFF3EFAE5" box="[159,245,1369,1395]" captionStart="FIGURE 3" captionStartId="6.[151,250,1848,1872]" captionTargetBox="[152,1437,418,1807]" captionTargetId="figure-114@6.[151,1437,417,1825]" captionTargetPageId="6" captionText="FIGURE 3. Miridiba quangnamensis P.V. Pham & Keith, new species. a, habitus; b, pronotum and head; c, meso- and metasternum; d, scutellum; e, foretibia; f, hindtibia; g, pygidium." figureDoi="http://doi.org/10.5281/zenodo.7306968" httpUri="https://zenodo.org/record/7306968/files/figure.png" pageId="5" pageNumber="336">Fig. 3e</figureCitation>
|
||
). Profemora, mesofemora, and metafemora moderately covered with hair-like, long setae; metafemora strongly broadened anteriorly, slightly convex. Metatibia with two apical spurs, upper spur moderately longer than lower spur, and reaching one fourth of mesotarsomeres 2 (
|
||
<figureCitation id="13522A027C11FF9EFB3DFA63FC5CFA2D" box="[836,919,1441,1467]" captionStart="FIGURE 3" captionStartId="6.[151,250,1848,1872]" captionTargetBox="[152,1437,418,1807]" captionTargetId="figure-114@6.[151,1437,417,1825]" captionTargetPageId="6" captionText="FIGURE 3. Miridiba quangnamensis P.V. Pham & Keith, new species. a, habitus; b, pronotum and head; c, meso- and metasternum; d, scutellum; e, foretibia; f, hindtibia; g, pygidium." figureDoi="http://doi.org/10.5281/zenodo.7306968" httpUri="https://zenodo.org/record/7306968/files/figure.png" pageId="5" pageNumber="336">Fig. 3f</figureCitation>
|
||
). Meso- and metatibiae with strong teeth on external margins and complete carina at apical third.
|
||
</paragraph>
|
||
<paragraph id="8BD636877C11FF9EF8BEFA2BFF1CF9E6" blockId="5.[151,1437,577,2008]" pageId="5" pageNumber="336">
|
||
<emphasis id="B91DEA957C11FF9EF8BEFA2BFE43F995" bold="true" box="[199,392,1513,1539]" pageId="5" pageNumber="336">Ventral surface.</emphasis>
|
||
Meso- and metasternum with dense fine punctures and yellowish hair (
|
||
<figureCitation id="13522A027C11FF9EFCC4FA2BFADBF995" box="[1213,1296,1513,1539]" captionStart="FIGURE 3" captionStartId="6.[151,250,1848,1872]" captionTargetBox="[152,1437,418,1807]" captionTargetId="figure-114@6.[151,1437,417,1825]" captionTargetPageId="6" captionText="FIGURE 3. Miridiba quangnamensis P.V. Pham & Keith, new species. a, habitus; b, pronotum and head; c, meso- and metasternum; d, scutellum; e, foretibia; f, hindtibia; g, pygidium." figureDoi="http://doi.org/10.5281/zenodo.7306968" httpUri="https://zenodo.org/record/7306968/files/figure.png" pageId="5" pageNumber="336">Fig. 3c</figureCitation>
|
||
). Abdomen: second and third visible abdomenal sternites densely covered with punctures, and long setae; those on fourth and fifth sternites sparserly punctate and setiferous, particularly on median parts; sixth sternite covered with longer setae.
|
||
</paragraph>
|
||
<paragraph id="8BD636877C11FF9EF8BEF9BBFC76F921" blockId="5.[151,1437,577,2008]" pageId="5" pageNumber="336">
|
||
<emphasis id="B91DEA957C11FF9EF8BEF9BBFE89F905" bold="true" box="[199,322,1657,1683]" pageId="5" pageNumber="336">Pygidium.</emphasis>
|
||
Slightly convex, triangular, margin bordered, apical margin moderately reflexed; dorsal surface with punctures identical to those on elytra with long and erect setae (
|
||
<figureCitation id="13522A027C11FF9EFB18F95FFC7BF921" box="[865,944,1693,1719]" captionStart="FIGURE 3" captionStartId="6.[151,250,1848,1872]" captionTargetBox="[152,1437,418,1807]" captionTargetId="figure-114@6.[151,1437,417,1825]" captionTargetPageId="6" captionText="FIGURE 3. Miridiba quangnamensis P.V. Pham & Keith, new species. a, habitus; b, pronotum and head; c, meso- and metasternum; d, scutellum; e, foretibia; f, hindtibia; g, pygidium." figureDoi="http://doi.org/10.5281/zenodo.7306968" httpUri="https://zenodo.org/record/7306968/files/figure.png" pageId="5" pageNumber="336">Fig. 3f</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD636877C11FF9EF8BEF903FED1F969" blockId="5.[151,1437,577,2008]" pageId="5" pageNumber="336">
|
||
<emphasis id="B91DEA957C11FF9EF8BEF903FEF1F94D" bold="true" box="[199,314,1729,1755]" pageId="5" pageNumber="336">Aedeagus</emphasis>
|
||
. Curved downwards, symmetrical in lateral view, bird's beak-like, without acicular expansion at base (
|
||
<figureCitation id="13522A027C11FF9EF8E6F927FEC5F969" box="[159,270,1765,1791]" captionStart="FIGURE 4" captionStartId="7.[151,250,1184,1208]" captionTargetBox="[162,1426,227,1113]" captionTargetId="figure-18@7.[162,1426,181,1160]" captionTargetPageId="7" captionText="FIGURE 4. Male genitalia of Miridiba apicespinosa P.V. Pham & Keith, new species (a–c), Miridiba ngoclinhensis P.V. Pham & Keith, new species (d–f) and Miridiba quangnamensis P.V. Pham & Keith, new species (g–i). a, d, g, parameres in lateral view; b, e, h, parameres in dorsal view; c, f, i, parameres in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7306970" httpUri="https://zenodo.org/record/7306970/files/figure.png" pageId="5" pageNumber="336">
|
||
Figs
|
||
<quantity id="4C919B627C11FF9EF8A1F927FF3DF969" box="[216,246,1765,1791]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="5" pageNumber="336" unit="g" value="4.0">4g</quantity>
|
||
–i
|
||
</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD636877C11FF9EF8BEF8CBFE69F8B5" blockId="5.[151,1437,577,2008]" box="[199,418,1801,1827]" pageId="5" pageNumber="336">
|
||
<emphasis id="B91DEA957C11FF9EF8BEF8CBFEEEF8B5" bold="true" box="[199,293,1801,1827]" pageId="5" pageNumber="336">Female.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C373650C7C11FF9DF8BEF8EFFE81FED7" lastPageId="6" lastPageNumber="337" pageId="5" pageNumber="336" type="discussion">
|
||
<paragraph id="8BD636877C11FF9DF8BEF8EFFE81FED7" blockId="5.[151,1437,577,2008]" lastBlockId="6.[151,1436,150,393]" lastPageId="6" lastPageNumber="337" pageId="5" pageNumber="336">
|
||
<emphasis id="B91DEA957C11FF9EF8BEF8EFFEF0F8D1" bold="true" box="[199,315,1837,1863]" pageId="5" pageNumber="336">Remarks.</emphasis>
|
||
<taxonomicName id="4C694D047C11FF9EF93DF8EFFD95F8D1" authority="Pham & Keith & Pham & Cao, 2022" authorityName="Pham & Keith & Pham & Cao" authorityYear="2022" box="[324,606,1837,1863]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="species" species="quangnamensis" status="sp. nov.">
|
||
<emphasis id="B91DEA957C11FF9EF93DF8EFFD95F8D1" box="[324,606,1837,1863]" italics="true" pageId="5" pageNumber="336">Miridiba quangnamensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B91DEA957C11FF9EFA14F8EFFD33F8D1" bold="true" box="[621,760,1837,1863]" pageId="5" pageNumber="336">
|
||
<taxonomicNameLabel id="A22E57EE7C11FF9EFA14F8EFFD33F8D1" box="[621,760,1837,1863]" pageId="5" pageNumber="336" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
is closely similar to
|
||
<taxonomicName id="4C694D047C11FF9EFB91F8ECFADEF8D1" authority="Keith, 2006" authorityName="Keith" authorityYear="2006" box="[1000,1301,1837,1863]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="species" species="hanoiensis">
|
||
<emphasis id="B91DEA957C11FF9EFB91F8ECFB4CF8D1" box="[1000,1159,1837,1863]" italics="true" pageId="5" pageNumber="336">M. hanoiensis</emphasis>
|
||
<bibRefCitation id="EFF84B767C11FF9EFCF6F8EFFADEF8D1" author="Keith, D." box="[1167,1301,1837,1863]" pageId="5" pageNumber="336" pagination="43 - 58" refId="ref7422" refString="Keith, D. (2006) Especes nouvelles ou meconnues de Melolonthidae asiatiques. Symbioses, Nouvelle Serie, 17, 43 - 58." type="journal article" year="2006">Keith, 2006</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13522A027C11FF9EFD5CF8EFFA5FF8DE" box="[1317,1428,1837,1864]" captionStart="FIGURE 5" captionStartId="9.[151,250,1852,1876]" captionTargetBox="[185,1404,205,1800]" captionTargetId="figure-18@9.[151,1436,181,1828]" captionTargetPageId="9" captionText="FIGURE 5. Miridiba pseudosinensis Keith, 2010 (a–c) and Miridiba hanoiensis Keith, 2006 (d–f). a, d, habitus; b, e, parameres in lateral view; c, f, parameres in dorsal view. Figures 5d–f photographed by Olivier Montreuil. Not in scale." figureDoi="http://doi.org/10.5281/zenodo.7306972" httpUri="https://zenodo.org/record/7306972/files/figure.png" pageId="5" pageNumber="336">Figs 5d–f</figureCitation>
|
||
) but they can be separated by the following characters combined: posterior pronotal margin without pubesence in
|
||
<taxonomicName id="4C694D047C11FF9EF8EEF8B4FEA7F819" authority="Pham & Keith & Pham & Cao, 2022" authorityName="Pham & Keith & Pham & Cao" authorityYear="2022" box="[151,364,1910,1935]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="species" species="quangnamensis" status="sp. nov.">
|
||
<emphasis id="B91DEA957C11FF9EF8EEF8B4FEA7F819" box="[151,364,1910,1935]" italics="true" pageId="5" pageNumber="336">M. quangnamensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B91DEA957C11FF9EF902F8B7FDCDF819" bold="true" box="[379,518,1909,1935]" pageId="5" pageNumber="336">
|
||
<taxonomicNameLabel id="A22E57EE7C11FF9EF902F8B7FDCDF819" box="[379,518,1909,1935]" pageId="5" pageNumber="336" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
(pubescent in
|
||
<taxonomicName id="4C694D047C11FF9EFAD7F8B4FC89F819" baseAuthorityName="Keith" baseAuthorityYear="2006" box="[686,834,1909,1936]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="species" species="hanoienis">
|
||
<emphasis id="B91DEA957C11FF9EFAD7F8B4FD0EF819" box="[686,709,1910,1935]" italics="true" pageId="5" pageNumber="336">M</emphasis>
|
||
.
|
||
<emphasis id="B91DEA957C11FF9EFAACF8B7FC89F819" box="[725,834,1909,1935]" italics="true" pageId="5" pageNumber="336">hanoienis</emphasis>
|
||
</taxonomicName>
|
||
); clypeo-frontal suture visible in
|
||
<taxonomicName id="4C694D047C11FF9EFCB9F8B4FA5EF819" authority="Pham & Keith & Pham & Cao, 2022" authorityName="Pham & Keith & Pham & Cao" authorityYear="2022" box="[1216,1429,1910,1935]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="species" species="quangnamensis" status="sp. nov.">
|
||
<emphasis id="B91DEA957C11FF9EFCB9F8B4FA5EF819" box="[1216,1429,1910,1935]" italics="true" pageId="5" pageNumber="336">M. quangnamensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B91DEA957C11FF9EF8EEF85BFEE8F825" bold="true" box="[151,291,1945,1971]" pageId="5" pageNumber="336">
|
||
<taxonomicNameLabel id="A22E57EE7C11FF9EF8EEF85BFEE8F825" box="[151,291,1945,1971]" pageId="5" pageNumber="336" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
(invisible in
|
||
<taxonomicName id="4C694D047C11FF9EF9C6F858FD94F825" authorityName="Keith" authorityYear="2006" box="[447,607,1945,1971]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="species" species="hanoiensis">
|
||
<emphasis id="B91DEA957C11FF9EF9C6F858FD94F825" box="[447,607,1945,1971]" italics="true" pageId="5" pageNumber="336">M. hanoiensis</emphasis>
|
||
</taxonomicName>
|
||
); antenna club longer than length of antennomeres 2–6 combined in
|
||
<taxonomicName id="4C694D047C11FF9EFD07F858FE8EF841" authority="Pham & Keith & Pham & Cao, 2022" authorityName="Pham & Keith & Pham & Cao" authorityYear="2022" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="species" species="quangnamensis" status="sp. nov.">
|
||
<emphasis id="B91DEA957C11FF9EFD07F858FE8EF841" italics="true" pageId="5" pageNumber="336">M. quangnamensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B91DEA957C11FF9EF92AF87FFE10F841" bold="true" box="[339,475,1981,2007]" pageId="5" pageNumber="336">
|
||
<taxonomicNameLabel id="A22E57EE7C11FF9EF92AF87FFE10F841" box="[339,475,1981,2007]" pageId="5" pageNumber="336" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="13522A027C11FF9EF993F87FFDF0F841" box="[490,571,1981,2007]" captionStart="FIGURE 3" captionStartId="6.[151,250,1848,1872]" captionTargetBox="[152,1437,418,1807]" captionTargetId="figure-114@6.[151,1437,417,1825]" captionTargetPageId="6" captionText="FIGURE 3. Miridiba quangnamensis P.V. Pham & Keith, new species. a, habitus; b, pronotum and head; c, meso- and metasternum; d, scutellum; e, foretibia; f, hindtibia; g, pygidium." figureDoi="http://doi.org/10.5281/zenodo.7306968" httpUri="https://zenodo.org/record/7306968/files/figure.png" pageId="5" pageNumber="336">Fig. 3d</figureCitation>
|
||
) (almost equal in
|
||
<taxonomicName id="4C694D047C11FF9EFB7AF87CFC6BF841" authorityName="Keith" authorityYear="2006" box="[771,928,1981,2007]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="species" species="hanoiensis">
|
||
<emphasis id="B91DEA957C11FF9EFB7AF87CFC6BF841" box="[771,928,1981,2007]" italics="true" pageId="5" pageNumber="336">M. hanoiensis</emphasis>
|
||
</taxonomicName>
|
||
); anterolateral angles moderately protuberant in
|
||
<taxonomicName id="4C694D047C12FF9DF8CEFF55FE46FF26" authorityName="Pham & Keith & Pham & Cao" authorityYear="2022" box="[183,397,151,176]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="337" phylum="Arthropoda" rank="species" species="quangnamensis">
|
||
<emphasis id="B91DEA957C12FF9DF8CEFF55FE46FF26" box="[183,397,151,176]" italics="true" pageId="6" pageNumber="337">M. quangnamensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13522A027C12FF9DF9E6FF55FE38FF27" box="[415,499,151,177]" captionStart="FIGURE 3" captionStartId="6.[151,250,1848,1872]" captionTargetBox="[152,1437,418,1807]" captionTargetId="figure-114@6.[151,1437,417,1825]" captionTargetPageId="6" captionText="FIGURE 3. Miridiba quangnamensis P.V. Pham & Keith, new species. a, habitus; b, pronotum and head; c, meso- and metasternum; d, scutellum; e, foretibia; f, hindtibia; g, pygidium." figureDoi="http://doi.org/10.5281/zenodo.7306968" httpUri="https://zenodo.org/record/7306968/files/figure.png" pageId="6" pageNumber="337">Fig. 3b</figureCitation>
|
||
) (nearly rectangular in
|
||
<emphasis id="B91DEA957C12FF9DFB7CFF55FC77FF26" box="[773,956,151,177]" italics="true" pageId="6" pageNumber="337">
|
||
<taxonomicName id="4C694D047C12FF9DFB7CFF55FC6EFF27" authorityName="Keith" authorityYear="2006" box="[773,933,151,177]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="337" phylum="Arthropoda" rank="species" species="hanoiensis">M. hanoiensis</taxonomicName>
|
||
–
|
||
</emphasis>
|
||
<figureCitation id="13522A027C12FF9DFBBFFF55FBD0FF27" box="[966,1051,150,177]" captionStart="FIGURE 5" captionStartId="9.[151,250,1852,1876]" captionTargetBox="[185,1404,205,1800]" captionTargetId="figure-18@9.[151,1436,181,1828]" captionTargetPageId="9" captionText="FIGURE 5. Miridiba pseudosinensis Keith, 2010 (a–c) and Miridiba hanoiensis Keith, 2006 (d–f). a, d, habitus; b, e, parameres in lateral view; c, f, parameres in dorsal view. Figures 5d–f photographed by Olivier Montreuil. Not in scale." figureDoi="http://doi.org/10.5281/zenodo.7306972" httpUri="https://zenodo.org/record/7306972/files/figure.png" pageId="6" pageNumber="337">Fig. 5d</figureCitation>
|
||
); median limits between sternites slightly opaque in in
|
||
<taxonomicName id="4C694D047C12FF9DF9F8FF79FD98FF42" authority="Pham & Keith & Pham & Cao, 2022" authorityName="Pham & Keith & Pham & Cao" authorityYear="2022" box="[385,595,187,212]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="337" phylum="Arthropoda" rank="species" species="quangnamensis" status="sp. nov.">
|
||
<emphasis id="B91DEA957C12FF9DF9F8FF79FD98FF42" box="[385,595,187,212]" italics="true" pageId="6" pageNumber="337">M. quangnamensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B91DEA957C12FF9DFA18FF79FD22FF42" bold="true" box="[609,745,186,213]" pageId="6" pageNumber="337">
|
||
<taxonomicNameLabel id="A22E57EE7C12FF9DFA18FF79FD22FF42" box="[609,745,186,213]" pageId="6" pageNumber="337" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
(invisible in
|
||
<taxonomicName id="4C694D047C12FF9DFB05FF79FBD3FF43" authorityName="Keith" authorityYear="2006" box="[892,1048,187,213]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="337" phylum="Arthropoda" rank="species" species="hanoiensis">
|
||
<emphasis id="B91DEA957C12FF9DFB05FF79FBD3FF43" box="[892,1048,187,213]" italics="true" pageId="6" pageNumber="337">M. hanoiensis</emphasis>
|
||
</taxonomicName>
|
||
), and the shape of parameres (
|
||
<figureCitation id="13522A027C12FF9DFD13FF79FF0EFF6F" captionStart="FIGURE 5" captionStartId="9.[151,250,1852,1876]" captionTargetBox="[185,1404,205,1800]" captionTargetId="figure-18@9.[151,1436,181,1828]" captionTargetPageId="9" captionText="FIGURE 5. Miridiba pseudosinensis Keith, 2010 (a–c) and Miridiba hanoiensis Keith, 2006 (d–f). a, d, habitus; b, e, parameres in lateral view; c, f, parameres in dorsal view. Figures 5d–f photographed by Olivier Montreuil. Not in scale." figureDoi="http://doi.org/10.5281/zenodo.7306972" httpUri="https://zenodo.org/record/7306972/files/figure.png" pageId="6" pageNumber="337">Figs 5e,f</figureCitation>
|
||
;
|
||
<figureCitation id="13522A027C12FF9DF8ABFF1DFEC3FF6F" box="[210,264,223,249]" captionStart="FIGURE 4" captionStartId="7.[151,250,1184,1208]" captionTargetBox="[162,1426,227,1113]" captionTargetId="figure-18@7.[162,1426,181,1160]" captionTargetPageId="7" captionText="FIGURE 4. Male genitalia of Miridiba apicespinosa P.V. Pham & Keith, new species (a–c), Miridiba ngoclinhensis P.V. Pham & Keith, new species (d–f) and Miridiba quangnamensis P.V. Pham & Keith, new species (g–i). a, d, g, parameres in lateral view; b, e, h, parameres in dorsal view; c, f, i, parameres in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7306970" httpUri="https://zenodo.org/record/7306970/files/figure.png" pageId="6" pageNumber="337">
|
||
<quantity id="4C919B627C12FF9DF8ABFF1DFF3BFF6F" box="[210,240,223,249]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="6" pageNumber="337" unit="g" value="4.0">4g</quantity>
|
||
–i
|
||
</figureCitation>
|
||
). The morphological characters of
|
||
<taxonomicName id="4C694D047C12FF9DFAEDFF1DFCA2FF6E" authority="Pham & Keith & Pham & Cao, 2022" authorityName="Pham & Keith & Pham & Cao" authorityYear="2022" box="[660,873,223,248]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="337" phylum="Arthropoda" rank="species" species="quangnamensis" status="sp. nov.">
|
||
<emphasis id="B91DEA957C12FF9DFAEDFF1DFCA2FF6E" box="[660,873,223,248]" italics="true" pageId="6" pageNumber="337">M. quangnamensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91DEA957C12FF9DFB08FF1DFC37FF6E" bold="true" box="[881,1020,222,249]" pageId="6" pageNumber="337">
|
||
<taxonomicNameLabel id="A22E57EE7C12FF9DFB08FF1DFC37FF6E" box="[881,1020,222,249]" pageId="6" pageNumber="337" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
do not characterize any morphotype proposed by
|
||
<bibRefCitation id="EFF84B767C12FF9DF952FEC1FD86FE8B" author="Gao, C. B. & Coca-Abia, M. M." box="[299,589,259,285]" pageId="6" pageNumber="337" pagination="1 - 94" refId="ref7217" refString="Gao, C. B. & Coca-Abia, M. M. (2021) Revision of the genus Miridiba Reitter, 1902 (Coleoptera, Scarabaeidae, Melolonthinae). Genital morphotypes and new taxonomic data. European Journal of Taxonomy, 749, 1 - 94. https: // doi. org / 10.5852 / ejt. 2021.749.1355" type="journal article" year="2021">Gao & Coca-Abia (2021)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFF84B767C12FF9DFAFFFEC1FC86FE8B" author="Gao, C. B. & Idrees, A. & Li, J." box="[646,845,259,285]" pageId="6" pageNumber="337" pagination="101 - 109" refId="ref7279" refString="Gao, C. B., Idrees, A. & Li, J. (2022) The Miridiba lamellata group (Coleoptera: Scarabaeidae) with a new species from Yunnan, China. The Coleopterists Bulletin, 76 (1), 101 - 109. https: // doi. org / 10.1649 / 0010 - 065 X- 76.1.101" type="journal article" year="2022">
|
||
Gao
|
||
<emphasis id="B91DEA957C12FF9DFAC4FEC1FD3CFE8B" box="[701,759,259,285]" italics="true" pageId="6" pageNumber="337">et al.</emphasis>
|
||
(2022)
|
||
</bibRefCitation>
|
||
, so the new species remains as
|
||
<emphasis id="B91DEA957C12FF9DFCCEFEC1FA98FE8B" box="[1207,1363,259,285]" italics="true" pageId="6" pageNumber="337">incertae sedis</emphasis>
|
||
in the genus
|
||
<taxonomicName id="4C694D047C12FF9DF899FEE5FE8FFED7" authorityName="Reitter" authorityYear="1902" box="[224,324,295,321]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="337" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91DEA957C12FF9DF899FEE5FE8FFED7" box="[224,324,295,321]" italics="true" pageId="6" pageNumber="337">Miridiba</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C373650C7C12FF9DF8BEFE89FB07FEF3" box="[199,1228,331,357]" pageId="6" pageNumber="337" type="etymology">
|
||
<paragraph id="8BD636877C12FF9DF8BEFE89FB07FEF3" blockId="6.[151,1436,150,393]" box="[199,1228,331,357]" pageId="6" pageNumber="337">
|
||
<emphasis id="B91DEA957C12FF9DF8BEFE89FE85FEF3" bold="true" box="[199,334,331,357]" pageId="6" pageNumber="337">Etymology.</emphasis>
|
||
This species is named after the locality of the
|
||
<typeStatus id="54D288257C12FF9DFB2FFE89FC73FEF3" box="[854,952,331,357]" pageId="6" pageNumber="337" type="holotype">holotype</typeStatus>
|
||
(
|
||
<collectingRegion id="49ADF8657C12FF9DFBBEFE89FB0AFEF3" box="[967,1217,331,357]" country="Vietnam" name="Quang Nam" pageId="6" pageNumber="337">Quang Nam Province</collectingRegion>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C373650C7C12FF9DF8BEFEACFD11FE1F" box="[199,730,366,393]" pageId="6" pageNumber="337" type="distribution">
|
||
<paragraph id="8BD636877C12FF9DF8BEFEACFD11FE1F" blockId="6.[151,1436,150,393]" box="[199,730,366,393]" pageId="6" pageNumber="337">
|
||
<emphasis id="B91DEA957C12FF9DF8BEFEACFEAAFE1E" bold="true" box="[199,353,366,392]" pageId="6" pageNumber="337">Distribution.</emphasis>
|
||
<collectingCountry id="F37E76177C12FF9DF91EFEADFE0DFE1F" box="[359,454,367,393]" name="Vietnam" pageId="6" pageNumber="337">Vietnam</collectingCountry>
|
||
(
|
||
<collectingRegion id="49ADF8657C12FF9DF9ACFEADFD04FE1F" box="[469,719,367,393]" country="Vietnam" name="Quang Nam" pageId="6" pageNumber="337">Quang Nam Province</collectingRegion>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |