783 lines
137 KiB
XML
783 lines
137 KiB
XML
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<mods:title id="39B7CEDD355E90D40637A0BBCCB75456">Redescription of Vieja hartwegi (Taylor & Miller 1980) (Teleostei: Cichlidae) from the Grijalva River basin, Mexico and Guatemala, with description of a rheophilic morph</mods:title>
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<mods:namePart id="7553995ECEB6E80D804CD5A2536F2846">Gómez-González, Adán E.</mods:namePart>
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<mods:namePart id="B2AD8C831AC38FDBA0AC626FA318A7D8">Álvarez, Fernando</mods:namePart>
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<treatment id="03C0878BFFE7FFA1E09D4388FC2DC096" ID-DOI="http://doi.org/10.5281/zenodo.5969912" ID-GBIF-Taxon="140417873" ID-Zenodo-Dep="5969912" LSID="urn:lsid:plazi:treatment:03C0878BFFE7FFA1E09D4388FC2DC096" httpUri="http://treatment.plazi.org/id/03C0878BFFE7FFA1E09D4388FC2DC096" lastPageId="14" lastPageNumber="385" pageId="5" pageNumber="376">
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<subSubSection id="C3736516FFE7FFAAE09D4388FC85C793" box="[151,831,220,247]" pageId="5" pageNumber="376" type="nomenclature">
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<paragraph id="8BD6369DFFE7FFAAE09D4388FC85C793" blockId="5.[151,831,220,280]" box="[151,831,220,247]" pageId="5" pageNumber="376">
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<heading id="D09E81F1FFE7FFAAE09D4388FC85C793" bold="true" box="[151,831,220,247]" fontSize="11" level="1" pageId="5" pageNumber="376" reason="1">
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<emphasis id="B91DEA8FFFE7FFAAE09D4388FC85C793" bold="true" box="[151,831,220,247]" pageId="5" pageNumber="376">
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Redescription of
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<taxonomicName id="4C694D1EFFE7FFAAE1664389FC85C793" ID-CoL="5BDC3" authority="(Taylor & Miller 1980)" baseAuthorityName="Taylor & Miller" baseAuthorityYear="1980" box="[364,831,220,247]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="hartwegi">
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<emphasis id="B91DEA8FFFE7FFAAE1664389FDA1C792" bold="true" box="[364,539,220,246]" italics="true" pageId="5" pageNumber="376">Vieja hartwegi</emphasis>
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(
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<bibRefCitation id="EFF84B6CFFE7FFAAE2204388FC8DC793" author="Taylor" box="[554,823,221,247]" pageId="5" pageNumber="376" refString="Taylor, J. N. & Miller, R. R. (1980) Two new cichlid fishes, genus Cichlasoma, from Chiapas, Mexico. Occasional Papers of the Museum of Zoology, University of Michigan, 693, 1 - 16." type="journal article" year="1980">Taylor & Miller 1980</bibRefCitation>
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)
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C3736516FFE7FFAAE09D43AAFE3DC67C" box="[151,391,255,280]" pageId="5" pageNumber="376" type="description">
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<paragraph id="8BD6369DFFE7FFAAE09D43AAFE3DC67C" blockId="5.[151,831,220,280]" box="[151,391,255,280]" pageId="5" pageNumber="376">
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<figureCitation id="13522A18FFE7FFAAE09D43AAFEBBC67C" box="[151,257,255,280]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE5" captionStart-3="FIGURE 6" captionStart-4="FIGURE 7" captionStart-5="FIGURE 8" captionStartId-0="6.[151,250,1858,1880]" captionStartId-1="7.[151,250,1125,1147]" captionStartId-2="8.[151,250,1823,1845]" captionStartId-3="9.[151,250,1685,1707]" captionStartId-4="10.[151,250,1908,1930]" captionStartId-5="11.[151,250,1153,1175]" captionTargetBox-0="[192,1411,225,1818]" captionTargetBox-1="[151,1437,193,1104]" captionTargetBox-2="[220,1316,219,1782]" captionTargetBox-3="[151,1436,235,1664]" captionTargetBox-4="[169,1418,1029,1870]" captionTargetBox-5="[162,1401,219,1104]" captionTargetId-0="figure@6.[151,1436,193,1837]" captionTargetId-1="figure@7.[151,1437,193,1104]" captionTargetId-2="figure@8.[203,1338,193,1801]" captionTargetId-3="figure@9.[151,1436,193,1664]" captionTargetId-4="figure@10.[150,1436,1021,1887]" captionTargetId-5="figure@11.[151,1436,193,1132]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionTargetPageId-4="10" captionTargetPageId-5="11" captionText-0="FIGURE 3. Live specimens of the typical morph of Vieja hartwegi, showing ontogenic variation in coloration pattern. A) Juvenile, 75 mm SL, MZUNICACH 6761, La Angostura Reservoir, La Concordia, Chiapas. B) Adult, 144 mm SL, aquarium specimen, Cacahuanó River, Ocozocoautla, Chiapas. C) Large adult, 250 mm SL, uncataloged specimen from a fishery, Grijalva River (main channel at Tres Picos), Copainalá, Chiapas (photo by J. M. López-Vila)." captionText-1="FIGURE 4. Live specimens of the rheophilic morph of Vieja hartwegi. A) Adult female in normal coloration, 97 mm SL, MZUNICACH 7161, Lagos de Colón, La Trinitaria, Chiapas (photo by M. A. Peralta-Meixueiro). B) Adult female in stress coloration, 65 mm SL, uncataloged specimen, Lagos de Colón, La Trinitaria, Chiapas. C) Adult female in stress coloration, 91 mm SL, MZUNICACH 7159, Tachinua River, Chicomuselo, Chiapas. D) Adult male in stress coloration, 80 mm SL, MZUNICACH 7157, Yayahuita River, Chicomuselo, Chiapas." captionText-2="FIGURE5. Fixed specimensof the typical Viejahartwegi morph,showing the variation inbody shapeand the ontogenetic and individual variation in coloration patterns. A) Holotype (adult in breeding coloration), UMMZ 207701. B) Chiquito River, Chiapa de Corzo (juv.), MZUNICACH 7053. C) El Zapote River, Chiapa de Corzo (juv.), MZUNICACH 7210. D) La Angostura Reservoir, La Concordia (juv.), MZUNICACH 7154. E) Totopac River, Tecpatán (subadult), MZUNICACH 1053. F) Lagos de Colón, La Trinitaria (subadult), MZUNICACH 7065.G) CopanóRiver, Ostuacán (adult), MZUNICACH 7208.H- I) Grijalva River (main cannel), Copainalá (adults), MZUNICACH 3145. J) Santo Domingo River, Chiapa de Corzo (adult), MZUNICACH 6245." captionText-3="FIGURE 6. Preserved specimens of the rheophilic Vieja hartwegi morph. A) Adult female, MZUNICACH 7161, Lagos de Colón, La Trinitaria, Chiapas. B) Adult female, MZUNICACH 7159 [post-mortem fixed specimen], Yayahuita River, Chicomuselo, Chiapas." captionText-4="FIGURE 7. Lower pharyngeal tooth plate and urohyal bone of the two morphs of V. hartwegi. A) Typical morph, MZUNICACH 6761.B)Rheophilic morph, MZUNICACH 7157(photos by K.Ramírez-Moreno [pharyngeal tooth plates] and G. Morales-Flores [urohyal bones])." captionText-5="FIGURE 8. Cleared and stained specimens of the two morphs of V. hartwegi. A) Typical morph, 89.3 mm SL, MZUNICACH 6761, La Angostura Reservoir, La Concordia, Chiapas. B) Rheophilic morph, 53.2 mm SL, MZUNICACH 7159, Tachinua River, Chicomuselo, Chiapas." httpUri-0="https://zenodo.org/record/1160372/files/figure.png" httpUri-1="https://zenodo.org/record/1160375/files/figure.png" httpUri-2="https://zenodo.org/record/1160377/files/figure.png" httpUri-3="https://zenodo.org/record/1160379/files/figure.png" httpUri-4="https://zenodo.org/record/1160381/files/figure.png" httpUri-5="https://zenodo.org/record/1160383/files/figure.png" pageId="5" pageNumber="376">Figs. 3–8</figureCitation>
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,
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<tableCitation id="C6EB0326FFE7FFAAE10743AAFED7C67C" box="[269,365,255,280]" captionStart="TABLE2" captionStartId="15.[138,213,471,491]" captionText="TABLE2. Morphometricdatafor the two morphsof Vieja hartwegi and related species. Measurements are presented as percent ofstandard length or percent ofhead length.SD=standard deviation,Min=Minimum,Max= Maximum." pageId="5" pageNumber="376">Tables 2</tableCitation>
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, 3
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</paragraph>
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</subSubSection>
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<subSubSection id="C3736516FFE7FFAAE09D421DFB3CC6FF" pageId="5" pageNumber="376" type="reference_group">
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<paragraph id="8BD6369DFFE7FFAAE09D421DFCF5C63A" blockId="5.[151,1178,328,411]" box="[151,847,328,350]" pageId="5" pageNumber="376">
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<taxonomicName id="4C694D1EFFE7FFAAE09D421DFDD0C63A" ID-CoL="5YT3G" authority="Taylor & Miller 1980: 8" authorityName="Taylor & Miller" authorityPageNumber="8" authorityYear="1980" box="[151,618,328,350]" class="Actinopterygii" family="Cichlidae" genus="Cichlasoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="hartwegi">
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<emphasis id="B91DEA8FFFE7FFAAE09D421DFECAC63A" box="[151,368,328,350]" italics="true" pageId="5" pageNumber="376">Cichlasoma hartwegi</emphasis>
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<bibRefCitation id="EFF84B6CFFE7FFAAE17C421DFDEEC63A" author="Taylor" box="[374,596,328,350]" pageId="5" pageNumber="376" refString="Taylor, J. N. & Miller, R. R. (1980) Two new cichlid fishes, genus Cichlasoma, from Chiapas, Mexico. Occasional Papers of the Museum of Zoology, University of Michigan, 693, 1 - 16." type="journal article" year="1980">Taylor & Miller 1980</bibRefCitation>
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: 8
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</taxonomicName>
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[original description].
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</paragraph>
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<paragraph id="8BD6369DFFE7FFAAE09D4233FB20C618" blockId="5.[151,1178,328,411]" box="[151,1178,358,380]" pageId="5" pageNumber="376">
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<taxonomicName id="4C694D1EFFE7FFAAE09D4233FDD9C618" ID-CoL="6TRQL" authority="(Taylor & Miller 1980)" baseAuthorityName="Taylor & Miller" baseAuthorityYear="1980" box="[151,611,358,380]" class="Actinopterygii" family="Cichlidae" genus="Paratheraps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="hartwegi">
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<emphasis id="B91DEA8FFFE7FFAAE09D4233FECCC618" box="[151,374,358,380]" italics="true" pageId="5" pageNumber="376">Paratheraps hartwegi</emphasis>
|
||
(
|
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<bibRefCitation id="EFF84B6CFFE7FFAAE1894233FDE6C618" author="Taylor" box="[387,604,358,380]" pageId="5" pageNumber="376" refString="Taylor, J. N. & Miller, R. R. (1980) Two new cichlid fishes, genus Cichlasoma, from Chiapas, Mexico. Occasional Papers of the Museum of Zoology, University of Michigan, 693, 1 - 16." type="journal article" year="1980">Taylor & Miller 1980</bibRefCitation>
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)
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</taxonomicName>
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||
—
|
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<bibRefCitation id="EFF84B6CFFE7FFAAE2774233FC13C618" author="Werner" box="[637,937,358,380]" pageId="5" pageNumber="376" refString="Werner, U. & Stawikowski, R. (1987) Ein neuer Buntbarsch aus Sudmexiko: Paratheraps breidohri, gen. nov., spec. nov. Die Aquarien-und Terrarien-Zeitschrift, 41, 20 - 23." type="journal article" year="1987">Werner & Stawikowski 1987</bibRefCitation>
|
||
: 20 [
|
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<taxonomicNameLabel id="A22E57F4FFE7FFAAE3D34232FB2BC618" box="[985,1169,358,380]" pageId="5" pageNumber="376">new combination</taxonomicNameLabel>
|
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].
|
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</paragraph>
|
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<paragraph id="8BD6369DFFE7FFAAE09D42D0FB3CC6FF" blockId="5.[151,1178,328,411]" box="[151,1158,389,411]" pageId="5" pageNumber="376">
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<taxonomicName id="4C694D1EFFE7FFAAE09D42D0FD38C6FF" ID-CoL="75PH5" authority="(Taylor & Miller 1980)" baseAuthorityName="Taylor & Miller" baseAuthorityYear="1980" box="[151,642,389,411]" class="Actinopterygii" family="Cichlidae" genus="Paraneetroplus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="hartwegi">
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||
<emphasis id="B91DEA8FFFE7FFAAE09D42D0FE2EC6FF" box="[151,404,389,411]" italics="true" pageId="5" pageNumber="376">Paraneetroplus hartwegi</emphasis>
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||
(
|
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<bibRefCitation id="EFF84B6CFFE7FFAAE1AB42D0FDC1C6FF" author="Taylor" box="[417,635,389,411]" pageId="5" pageNumber="376" refString="Taylor, J. N. & Miller, R. R. (1980) Two new cichlid fishes, genus Cichlasoma, from Chiapas, Mexico. Occasional Papers of the Museum of Zoology, University of Michigan, 693, 1 - 16." type="journal article" year="1980">Taylor & Miller 1980</bibRefCitation>
|
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)
|
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</taxonomicName>
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—
|
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<bibRefCitation id="EFF84B6CFFE7FFAAE29642D0FCC7C6FF" author="McMahan" box="[668,893,389,411]" pageId="5" pageNumber="376" refString="McMahan, C. D., Geheber, A. D. & Piller, K. R. (2010) Molecular Systematics of the Enigmatic Middle American Genus Vieja (Teleostei: Cichlidae). Molecular Phylogenetics and Evolution, 57, 1293 - 1300. https: // doi. org / 10.1016 / j. ympev. 2010.09.005" type="journal article" year="2010">
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McMahan
|
||
<emphasis id="B91DEA8FFFE7FFAAE30742D3FCFAC6FF" box="[781,832,389,411]" italics="true" pageId="5" pageNumber="376">et al.</emphasis>
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2010
|
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</bibRefCitation>
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: 1298 [
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<taxonomicNameLabel id="A22E57F4FFE7FFAAE3CF42D3FBC7C6FF" box="[965,1149,389,411]" pageId="5" pageNumber="376">new combination</taxonomicNameLabel>
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].
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</paragraph>
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|
||
<subSubSection id="C3736516FFE7FFAAE09D4292FB21C4D1" pageId="5" pageNumber="376" type="materials_examined">
|
||
<paragraph id="8BD6369DFFE7FFAAE09D4292FB21C4D1" blockId="5.[151,1437,455,2029]" pageId="5" pageNumber="376">
|
||
<emphasis id="B91DEA8FFFE7FFAAE09D4292FE38C684" bold="true" box="[151,386,455,480]" pageId="5" pageNumber="376">Material examined.</emphasis>
|
||
All from
|
||
<collectingRegion id="49ADF87FFFE7FFAAE1FF4292FDE8C684" box="[501,594,455,480]" country="Mexico" name="Chiapas" pageId="5" pageNumber="376">Chiapas</collectingRegion>
|
||
,
|
||
<collectingCountry id="F37E760DFFE7FFAAE2554292FD02C684" box="[607,696,455,480]" name="Mexico" pageId="5" pageNumber="376">
|
||
<collectingRegion id="49ADF87FFFE7FFAAE2554292FD02C684" box="[607,696,455,480]" country="Mexico" name="Mexico" pageId="5" pageNumber="376">Mexico</collectingRegion>
|
||
</collectingCountry>
|
||
, except
|
||
<collectionCode id="ED78AE58FFE7FFAAE31C429DFCDDC6BB" box="[790,871,456,479]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="5" pageNumber="376">FMNH</collectionCode>
|
||
131457; numbers in parentheses indicate number of specimens examined.
|
||
<emphasis id="B91DEA8FFFE7FFAAE1B942BEFC96C561" bold="true" box="[435,812,491,517]" pageId="5" pageNumber="376">
|
||
<taxonomicName id="4C694D1EFFE7FFAAE1B942BEFDE6C560" baseAuthorityName="Taylor & Miller" baseAuthorityYear="1980" box="[435,604,491,516]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="hartwegi">
|
||
<emphasis id="B91DEA8FFFE7FFAAE1B942BEFDE6C560" bold="true" box="[435,604,491,516]" italics="true" pageId="5" pageNumber="376">Vieja hartwegi</emphasis>
|
||
</taxonomicName>
|
||
(typical morph):
|
||
</emphasis>
|
||
<collectionCode id="ED78AE58FFE7FFAAE33C42B8FC37C560" box="[822,909,493,516]" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="5" pageNumber="376">UMMZ</collectionCode>
|
||
207701 (
|
||
<typeStatus id="54D2883FFFE7FFAAE3F142B9FBD9C561" box="[1019,1123,492,517]" pageId="5" pageNumber="376" type="holotype">holotype</typeStatus>
|
||
), Río Grande de Chiapa (= Grijalva River) about
|
||
<quantity id="4C919B78FFE7FFAAE1914145FE62C54C" box="[411,472,527,552]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="376" unit="km" value="1.0">1 km</quantity>
|
||
above bridge between Tuxtla Gutiérrez and Chiapa de Corzo;
|
||
<collectionCode id="ED78AE58FFE7FFAAE4C94145FAA0C543" box="[1219,1306,528,551]" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="5" pageNumber="376">UMMZ</collectionCode>
|
||
186400 (
|
||
<specimenCount id="9D6FFD14FFE7FFAAE5874145FEBEC528" pageId="5" pageNumber="376" type="generic" typeStatus="paratypes">2 paratypes</specimenCount>
|
||
) Frío River, above mouth of Chiapa River, Acala;
|
||
<collectionCode id="ED78AE58FFE7FFAAE3424160FC23C528" box="[840,921,565,588]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="5" pageNumber="376">FMNH</collectionCode>
|
||
93578 (
|
||
<specimenCount id="9D6FFD14FFE7FFAAE3F14161FBC8C528" box="[1019,1138,564,588]" pageId="5" pageNumber="376" type="generic" typeStatus="paratype">1 paratype</specimenCount>
|
||
) Salado River,
|
||
<quantity id="4C919B78FFE7FFAAE5294161FAE4C529" box="[1315,1374,564,589]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="376" unit="km" value="1.0">1 km</quantity>
|
||
from Chiapilla; MZUNICACH 1053 (1), 1065 (2), 1125 (1), 1156 (1) Totopac River, Tecpatán; MZUNICACH 1560 (1), 1639 (1), 7158 (2) La Venta River, Ocozocoautla; MZUNICACH 1789 (2) Malpaso Reservoir, Ocozocoautla; MZUNICACH 3045 (3) Río Grijalva, Chicoasén; MZUNICACH 3145 (5) Grijalva River, Copainalá; MZUNICACH 5947 (1), 6245 (1) Santo Domingo River, Chiapa de Corzo; MZUNICACH 7053 (1) Chiquito River, Chiapa de Corzo; MZUNICACH 7153 (3) Yayahuita River, Chicomuselo; MZUNICACH 6761 (2 c&s), 7154 (8) La Angostura Reservoir, La Concordia; MZUNICACH 7208 (1) Copanó River, Ostuacán;
|
||
<collectionCode id="ED78AE58FFE7FFAAE5084059FAE6C440" box="[1282,1372,780,804]" pageId="5" pageNumber="376">ECOSC</collectionCode>
|
||
7542 (2), 7544 (1) Blanco River, Venustiano Carranza;
|
||
<collectionCode id="ED78AE58FFE7FFAAE2CA4065FCA0C42C" box="[704,794,816,840]" pageId="5" pageNumber="376">ECOSC</collectionCode>
|
||
7543 (3), 7548 (3) Lagos de
|
||
<collectingRegion id="49ADF87FFFE7FFAAE455407AFB1CC42C" box="[1119,1190,815,840]" country="Panama" name="Colon" pageId="5" pageNumber="376">Colón</collectingRegion>
|
||
, La Trinitaria;
|
||
<collectionCode id="ED78AE58FFE7FFAAE5414065FA26C423" box="[1355,1436,816,839]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="5" pageNumber="376">FMNH</collectionCode>
|
||
131457 (2) Lagartero River, Nentón,
|
||
<collectingRegion id="49ADF87FFFE7FFAAE2704001FC97C409" box="[634,813,852,877]" country="Guatemala" name="Huehuetenango" pageId="5" pageNumber="376">Huehuetenango</collectingRegion>
|
||
,
|
||
<collectingCountry id="F37E760DFFE7FFAAE34F4001FC78C409" box="[837,962,852,877]" name="Guatemala" pageId="5" pageNumber="376">
|
||
<collectingRegion id="49ADF87FFFE7FFAAE34F4001FC78C409" box="[837,962,852,877]" country="Guatemala" name="Guatemala" pageId="5" pageNumber="376">Guatemala</collectingRegion>
|
||
</collectingCountry>
|
||
.
|
||
<emphasis id="B91DEA8FFFE7FFAAE3D34006FA26C409" bold="true" box="[985,1436,851,877]" pageId="5" pageNumber="376">
|
||
<taxonomicName id="4C694D1EFFE7FFAAE3D34006FB37C408" baseAuthorityName="Taylor & Miller" baseAuthorityYear="1980" box="[985,1165,851,876]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="hartwegi">
|
||
<emphasis id="B91DEA8FFFE7FFAAE3D34006FB37C408" bold="true" box="[985,1165,851,876]" italics="true" pageId="5" pageNumber="376">Vieja hartwegi</emphasis>
|
||
</taxonomicName>
|
||
(rheophilic morph):
|
||
</emphasis>
|
||
MZUNICACH 7157 (1 + 1 c&s), 7160 (1), 7166 (1) Yayahuita River, Chicomuselo; MZUNICACH 7159 (1 + 1 c&s) Tachinua River, Chicomuselo; MZUNICACH 7161 (4) Lagos de
|
||
<collectingRegion id="49ADF87FFFE7FFAAE3B940C9FC40C4D1" box="[947,1018,924,949]" country="Panama" name="Colon" pageId="5" pageNumber="376">Colón</collectingRegion>
|
||
, La Trinitaria.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3736516FFE7FFAAE0CD40EAFC38C2B4" pageId="5" pageNumber="376" type="diagnosis">
|
||
<paragraph id="8BD6369DFFE7FFAAE0CD40EAFD31C3D4" blockId="5.[151,1437,455,2029]" pageId="5" pageNumber="376">
|
||
<emphasis id="B91DEA8FFFE7FFAAE0CD40EAFEFBC4BC" bold="true" box="[199,321,959,984]" pageId="5" pageNumber="376">Diagnosis.</emphasis>
|
||
The typical morph of
|
||
<taxonomicName id="4C694D1EFFE7FFAAE2454094FD69C4BC" baseAuthorityName="Taylor & Miller" baseAuthorityYear="1980" box="[591,723,960,984]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="hartwegi">
|
||
<emphasis id="B91DEA8FFFE7FFAAE2454094FD69C4BC" box="[591,723,960,984]" italics="true" pageId="5" pageNumber="376">V. hartwegi</emphasis>
|
||
</taxonomicName>
|
||
usually possesses a dorsolateral or second stripe thinner and horizontal (parallel) to the midlateral stripe (
|
||
<figureCitation id="13522A18FFE7FFAAE29940B1FD6CC498" box="[659,726,996,1021]" captionStart="FIGURE 3" captionStartId="6.[151,250,1858,1880]" captionTargetBox="[192,1411,225,1818]" captionTargetId="figure@6.[151,1436,193,1837]" captionTargetPageId="6" captionText="FIGURE 3. Live specimens of the typical morph of Vieja hartwegi, showing ontogenic variation in coloration pattern. A) Juvenile, 75 mm SL, MZUNICACH 6761, La Angostura Reservoir, La Concordia, Chiapas. B) Adult, 144 mm SL, aquarium specimen, Cacahuanó River, Ocozocoautla, Chiapas. C) Large adult, 250 mm SL, uncataloged specimen from a fishery, Grijalva River (main channel at Tres Picos), Copainalá, Chiapas (photo by J. M. López-Vila)." httpUri="https://zenodo.org/record/1160372/files/figure.png" pageId="5" pageNumber="376">Fig. 3</figureCitation>
|
||
B-C); in contrast,
|
||
<taxonomicName id="4C694D1EFFE7FFAAE3AC40B0FB88C498" baseAuthorityName="Steindachner" baseAuthorityYear="1864" box="[934,1074,996,1020]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="bifasciata">
|
||
<emphasis id="B91DEA8FFFE7FFAAE3AC40B0FB88C498" box="[934,1074,996,1020]" italics="true" pageId="5" pageNumber="376">V. bifasciata</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13522A18FFE7FFAAE44940B1FB26C498" box="[1091,1180,996,1021]" captionStart="FIGURE 9" captionStartId="12.[151,250,1919,1941]" captionTargetBox="[164,1432,476,1890]" captionTargetId="figure@12.[151,1436,445,1898]" captionTargetPageId="12" captionText="FIGURE 9. Comparison to species closely related to V. hartwegi: A) V. bifasciata, 140 mm SL, ECOSC 4677, Tzendales River, Ocosingo, Chiapas.B) V.breidohri, 105 mm SL, MZUNICACH 7065, Lagos de Colón, La Trinitaria, Chiapas." httpUri="https://zenodo.org/record/1160385/files/figure.png" pageId="5" pageNumber="376">Fig. 9A</figureCitation>
|
||
) has a broader second stripe that is slightly angled dorsally. The two stripes are separated by a narrow space (two thirds or half of a scale) or are often fused in
|
||
<taxonomicName id="4C694D1EFFE7FFAAE1864778FDB4C320" baseAuthorityName="Taylor & Miller" baseAuthorityYear="1980" box="[396,526,1068,1092]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="hartwegi">
|
||
<emphasis id="B91DEA8FFFE7FFAAE1864778FDB4C320" box="[396,526,1068,1092]" italics="true" pageId="5" pageNumber="376">V. hartwegi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13522A18FFE7FFAAE2154779FDD9C321" box="[543,611,1068,1093]" captionStart="FIGURE5" captionStartId="8.[151,250,1823,1845]" captionTargetBox="[220,1316,219,1782]" captionTargetId="figure@8.[203,1338,193,1801]" captionTargetPageId="8" captionText="FIGURE5. Fixed specimensof the typical Viejahartwegi morph,showing the variation inbody shapeand the ontogenetic and individual variation in coloration patterns. A) Holotype (adult in breeding coloration), UMMZ 207701. B) Chiquito River, Chiapa de Corzo (juv.), MZUNICACH 7053. C) El Zapote River, Chiapa de Corzo (juv.), MZUNICACH 7210. D) La Angostura Reservoir, La Concordia (juv.), MZUNICACH 7154. E) Totopac River, Tecpatán (subadult), MZUNICACH 1053. F) Lagos de Colón, La Trinitaria (subadult), MZUNICACH 7065.G) CopanóRiver, Ostuacán (adult), MZUNICACH 7208.H- I) Grijalva River (main cannel), Copainalá (adults), MZUNICACH 3145. J) Santo Domingo River, Chiapa de Corzo (adult), MZUNICACH 6245." httpUri="https://zenodo.org/record/1160377/files/figure.png" pageId="5" pageNumber="376">Fig. 5</figureCitation>
|
||
D-J), whereas in
|
||
<taxonomicName id="4C694D1EFFE7FFAAE3214778FC0DC320" baseAuthorityName="Steindachner" baseAuthorityYear="1864" box="[811,951,1068,1092]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="bifasciata">
|
||
<emphasis id="B91DEA8FFFE7FFAAE3214778FC0DC320" box="[811,951,1068,1092]" italics="true" pageId="5" pageNumber="376">V. bifasciata</emphasis>
|
||
</taxonomicName>
|
||
the space is broader (one and half or two scales).
|
||
<taxonomicName id="4C694D1EFFE7FFAAE0FE4705FE1FC30C" baseAuthorityName="Steindachner" baseAuthorityYear="1864" box="[244,421,1104,1128]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="bifasciata">
|
||
<emphasis id="B91DEA8FFFE7FFAAE0FE4705FE1FC30C" box="[244,421,1104,1128]" italics="true" pageId="5" pageNumber="376">Vieja bifasciata</emphasis>
|
||
</taxonomicName>
|
||
usually has a large dark opercular blotch, extending to the border of the eye (
|
||
<figureCitation id="13522A18FFE7FFAAE538471AFA35C30C" box="[1330,1423,1103,1128]" captionStart="FIGURE 9" captionStartId="12.[151,250,1919,1941]" captionTargetBox="[164,1432,476,1890]" captionTargetId="figure@12.[151,1436,445,1898]" captionTargetPageId="12" captionText="FIGURE 9. Comparison to species closely related to V. hartwegi: A) V. bifasciata, 140 mm SL, ECOSC 4677, Tzendales River, Ocosingo, Chiapas.B) V.breidohri, 105 mm SL, MZUNICACH 7065, Lagos de Colón, La Trinitaria, Chiapas." httpUri="https://zenodo.org/record/1160385/files/figure.png" pageId="5" pageNumber="376">Fig. 9A</figureCitation>
|
||
), whereas
|
||
<taxonomicName id="4C694D1EFFE7FFAAE0F14720FEC6C3E8" baseAuthorityName="Taylor & Miller" baseAuthorityYear="1980" box="[251,380,1140,1164]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="hartwegi">
|
||
<emphasis id="B91DEA8FFFE7FFAAE0F14720FEC6C3E8" box="[251,380,1140,1164]" italics="true" pageId="5" pageNumber="376">V. hartwegi</emphasis>
|
||
</taxonomicName>
|
||
has a simple crescent shape (
|
||
<figureCitation id="13522A18FFE7FFAAE2C44721FCAAC3E8" box="[718,784,1140,1165]" captionStart="FIGURE 3" captionStartId="6.[151,250,1858,1880]" captionTargetBox="[192,1411,225,1818]" captionTargetId="figure@6.[151,1436,193,1837]" captionTargetPageId="6" captionText="FIGURE 3. Live specimens of the typical morph of Vieja hartwegi, showing ontogenic variation in coloration pattern. A) Juvenile, 75 mm SL, MZUNICACH 6761, La Angostura Reservoir, La Concordia, Chiapas. B) Adult, 144 mm SL, aquarium specimen, Cacahuanó River, Ocozocoautla, Chiapas. C) Large adult, 250 mm SL, uncataloged specimen from a fishery, Grijalva River (main channel at Tres Picos), Copainalá, Chiapas (photo by J. M. López-Vila)." httpUri="https://zenodo.org/record/1160372/files/figure.png" pageId="5" pageNumber="376">Fig. 3</figureCitation>
|
||
B-C). Additionally,
|
||
<taxonomicName id="4C694D1EFFE7FFAAE3FE4720FBC5C3E8" baseAuthorityName="Steindachner" baseAuthorityYear="1864" box="[1012,1151,1140,1164]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="bifasciata">
|
||
<emphasis id="B91DEA8FFFE7FFAAE3FE4720FBC5C3E8" box="[1012,1151,1140,1164]" italics="true" pageId="5" pageNumber="376">V. bifasciata</emphasis>
|
||
</taxonomicName>
|
||
has a much deeper body with a more rounded shape than
|
||
<emphasis id="B91DEA8FFFE7FFAAE20E47CCFD31C3D4" box="[516,651,1176,1200]" italics="true" pageId="5" pageNumber="376">
|
||
<taxonomicName id="4C694D1EFFE7FFAAE20E47CCFD3DC3D4" baseAuthorityName="Taylor & Miller" baseAuthorityYear="1980" box="[516,647,1176,1200]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="hartwegi">V. hartwegi</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BD6369DFFE7FFAAE0CD47E8FC38C2B4" blockId="5.[151,1437,455,2029]" pageId="5" pageNumber="376">
|
||
<taxonomicName id="4C694D1EFFE7FFAAE0CD47E8FECAC3B0" baseAuthorityName="Werner & Stawikowski" baseAuthorityYear="1987" box="[199,368,1212,1236]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="breidohri">
|
||
<emphasis id="B91DEA8FFFE7FFAAE0CD47E8FECAC3B0" box="[199,368,1212,1236]" italics="true" pageId="5" pageNumber="376">Vieja breidohri</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from
|
||
<taxonomicName id="4C694D1EFFE7FFAAE2AB47E8FCA6C3B0" baseAuthorityName="Taylor & Miller" baseAuthorityYear="1980" box="[673,796,1212,1236]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="hartwegi">
|
||
<emphasis id="B91DEA8FFFE7FFAAE2AB47E8FCA6C3B0" box="[673,796,1212,1236]" italics="true" pageId="5" pageNumber="376">V. hartwegi</emphasis>
|
||
</taxonomicName>
|
||
by the presence of molariform or subconical pharyngeal teeth, versus conical. Additionally, lateral blotches or a second stripe above the midlateral stripe are absent in
|
||
<taxonomicName id="4C694D1EFFE7FFAAE58D47B4FEBAC278" baseAuthorityName="Werner & Stawikowski" baseAuthorityYear="1987" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="breidohri">
|
||
<emphasis id="B91DEA8FFFE7FFAAE58D47B4FA2EC39C" box="[1415,1428,1249,1272]" italics="true" pageId="5" pageNumber="376">V</emphasis>
|
||
.
|
||
<emphasis id="B91DEA8FFFE7FFAAE09D4651FEBAC278" box="[151,256,1284,1308]" italics="true" pageId="5" pageNumber="376">breidohri</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13522A18FFE7FFAAE1054651FED2C278" box="[271,360,1284,1309]" captionStart="FIGURE 9" captionStartId="12.[151,250,1919,1941]" captionTargetBox="[164,1432,476,1890]" captionTargetId="figure@12.[151,1436,445,1898]" captionTargetPageId="12" captionText="FIGURE 9. Comparison to species closely related to V. hartwegi: A) V. bifasciata, 140 mm SL, ECOSC 4677, Tzendales River, Ocosingo, Chiapas.B) V.breidohri, 105 mm SL, MZUNICACH 7065, Lagos de Colón, La Trinitaria, Chiapas." httpUri="https://zenodo.org/record/1160385/files/figure.png" pageId="5" pageNumber="376">Fig. 9B</figureCitation>
|
||
). The midlateral stripe in
|
||
<taxonomicName id="4C694D1EFFE7FFAAE2874650FCA8C278" baseAuthorityName="Werner & Stawikowski" baseAuthorityYear="1987" box="[653,786,1284,1308]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="breidohri">
|
||
<emphasis id="B91DEA8FFFE7FFAAE2874650FCA8C278" box="[653,786,1284,1308]" italics="true" pageId="5" pageNumber="376">V. breidohri</emphasis>
|
||
</taxonomicName>
|
||
is markedly disrupted, formed by a series of separated or connected blotches that fade anteriorly, not reaching the opercle (
|
||
<figureCitation id="13522A18FFE7FFAAE3784672FC71C224" box="[882,971,1319,1344]" captionStart="FIGURE 9" captionStartId="12.[151,250,1919,1941]" captionTargetBox="[164,1432,476,1890]" captionTargetId="figure@12.[151,1436,445,1898]" captionTargetPageId="12" captionText="FIGURE 9. Comparison to species closely related to V. hartwegi: A) V. bifasciata, 140 mm SL, ECOSC 4677, Tzendales River, Ocosingo, Chiapas.B) V.breidohri, 105 mm SL, MZUNICACH 7065, Lagos de Colón, La Trinitaria, Chiapas." httpUri="https://zenodo.org/record/1160385/files/figure.png" pageId="5" pageNumber="376">Fig. 9B</figureCitation>
|
||
). Juvenile specimens of
|
||
<taxonomicName id="4C694D1EFFE7FFAAE4D6467CFAE1C224" baseAuthorityName="Taylor & Miller" baseAuthorityYear="1980" box="[1244,1371,1320,1344]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="hartwegi">
|
||
<emphasis id="B91DEA8FFFE7FFAAE4D6467CFAE1C224" box="[1244,1371,1320,1344]" italics="true" pageId="5" pageNumber="376">V. hartwegi</emphasis>
|
||
</taxonomicName>
|
||
often possess a disrupted midlateral stripe below or at the middle of the body; in this case the two species are more difficult to distinguish.
|
||
<taxonomicName id="4C694D1EFFE7FFAAE1A84625FDF2C2EC" baseAuthorityName="Taylor & Miller" baseAuthorityYear="1980" box="[418,584,1392,1416]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="hartwegi">
|
||
<emphasis id="B91DEA8FFFE7FFAAE1A84625FDF2C2EC" box="[418,584,1392,1416]" italics="true" pageId="5" pageNumber="376">Vieja hartwegi</emphasis>
|
||
</taxonomicName>
|
||
possesses red spots or blotches on head (
|
||
<figureCitation id="13522A18FFE7FFAAE42E463AFBDCC2EC" box="[1060,1126,1391,1416]" captionStart="FIGURE 3" captionStartId="6.[151,250,1858,1880]" captionTargetBox="[192,1411,225,1818]" captionTargetId="figure@6.[151,1436,193,1837]" captionTargetPageId="6" captionText="FIGURE 3. Live specimens of the typical morph of Vieja hartwegi, showing ontogenic variation in coloration pattern. A) Juvenile, 75 mm SL, MZUNICACH 6761, La Angostura Reservoir, La Concordia, Chiapas. B) Adult, 144 mm SL, aquarium specimen, Cacahuanó River, Ocozocoautla, Chiapas. C) Large adult, 250 mm SL, uncataloged specimen from a fishery, Grijalva River (main channel at Tres Picos), Copainalá, Chiapas (photo by J. M. López-Vila)." httpUri="https://zenodo.org/record/1160372/files/figure.png" pageId="5" pageNumber="376">Fig. 3</figureCitation>
|
||
A-C), whereas
|
||
<taxonomicName id="4C694D1EFFE7FFAAE51C4624FA26C2EC" baseAuthorityName="Werner & Stawikowski" baseAuthorityYear="1987" box="[1302,1436,1392,1416]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="breidohri">
|
||
<emphasis id="B91DEA8FFFE7FFAAE51C4624FA26C2EC" box="[1302,1436,1392,1416]" italics="true" pageId="5" pageNumber="376">V. breidohri</emphasis>
|
||
</taxonomicName>
|
||
has small dark dots on sides of the head (
|
||
<figureCitation id="13522A18FFE7FFAAE26246C1FD78C2C8" box="[616,706,1428,1453]" captionStart="FIGURE 9" captionStartId="12.[151,250,1919,1941]" captionTargetBox="[164,1432,476,1890]" captionTargetId="figure@12.[151,1436,445,1898]" captionTargetPageId="12" captionText="FIGURE 9. Comparison to species closely related to V. hartwegi: A) V. bifasciata, 140 mm SL, ECOSC 4677, Tzendales River, Ocosingo, Chiapas.B) V.breidohri, 105 mm SL, MZUNICACH 7065, Lagos de Colón, La Trinitaria, Chiapas." httpUri="https://zenodo.org/record/1160385/files/figure.png" pageId="5" pageNumber="376">Fig. 9B</figureCitation>
|
||
).
|
||
<taxonomicName id="4C694D1EFFE7FFAAE2DF46C0FCC0C2C8" baseAuthorityName="Taylor & Miller" baseAuthorityYear="1980" box="[725,890,1428,1452]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="hartwegi">
|
||
<emphasis id="B91DEA8FFFE7FFAAE2DF46C0FCC0C2C8" box="[725,890,1428,1452]" italics="true" pageId="5" pageNumber="376">Vieja hartwegi</emphasis>
|
||
</taxonomicName>
|
||
possesses 29–31 total vertebrae (15 precaudal + 14–16 caudal), and
|
||
<taxonomicName id="4C694D1EFFE7FFAAE17E46ECFE42C2B4" baseAuthorityName="Werner & Stawikowski" baseAuthorityYear="1987" box="[372,504,1464,1488]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="breidohri">
|
||
<emphasis id="B91DEA8FFFE7FFAAE17E46ECFE42C2B4" box="[372,504,1464,1488]" italics="true" pageId="5" pageNumber="376">V. breidohri</emphasis>
|
||
</taxonomicName>
|
||
has 28 (14 precaudal + 14 caudal).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3736516FFE7FFA8E0CD4689FE53C153" lastPageId="7" lastPageNumber="378" pageId="5" pageNumber="376" type="discussion">
|
||
<paragraph id="8BD6369DFFE7FFAAE0CD4689FB63C17C" blockId="5.[151,1437,455,2029]" pageId="5" pageNumber="376">
|
||
All other species of
|
||
<taxonomicName id="4C694D1EFFE7FFAAE1A14688FE5EC290" authorityName="Fernandez-Yepez" authorityYear="1969" box="[427,484,1501,1524]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91DEA8FFFE7FFAAE1A14688FE5EC290" box="[427,484,1501,1524]" italics="true" pageId="5" pageNumber="376">Vieja</emphasis>
|
||
</taxonomicName>
|
||
with a midlateral stripe along the side of the body (
|
||
<emphasis id="B91DEA8FFFE7FFAAE4274688FAF3C290" box="[1069,1353,1500,1524]" italics="true" pageId="5" pageNumber="376">
|
||
<taxonomicName id="4C694D1EFFE7FFAAE4274688FB06C290" baseAuthorityName="Günther" baseAuthorityYear="1860" box="[1069,1212,1500,1524]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="fenestrata">V. fenestrata</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C694D1EFFE7FFAAE4C24688FAF3C290" baseAuthorityName="Günther" baseAuthorityYear="1864" box="[1224,1353,1500,1524]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="guttulata">V. guttulata</taxonomicName>
|
||
</emphasis>
|
||
, and
|
||
<taxonomicName id="4C694D1EFFE7FFAAE58D4688FF58C17C" baseAuthorityName="Meek" baseAuthorityYear="1905" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="zonata">
|
||
<emphasis id="B91DEA8FFFE7FFAAE58D4688FF58C17C" italics="true" pageId="5" pageNumber="376">V. zonata</emphasis>
|
||
</taxonomicName>
|
||
) can be differentiated from
|
||
<taxonomicName id="4C694D1EFFE7FFAAE2164554FD21C17C" baseAuthorityName="Taylor & Miller" baseAuthorityYear="1980" box="[540,667,1536,1560]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="hartwegi">
|
||
<emphasis id="B91DEA8FFFE7FFAAE2164554FD21C17C" box="[540,667,1536,1560]" italics="true" pageId="5" pageNumber="376">V. hartwegi</emphasis>
|
||
</taxonomicName>
|
||
by the absence of the second or dorsolateral stripe.
|
||
</paragraph>
|
||
<paragraph id="8BD6369DFFE7FFA8E0CD4571FE53C153" blockId="5.[151,1437,455,2029]" lastBlockId="7.[151,1437,1314,2029]" lastPageId="7" lastPageNumber="378" pageId="5" pageNumber="376">
|
||
The rheophilic
|
||
<taxonomicName id="4C694D1EFFE7FFAAE1734570FE46C158" baseAuthorityName="Taylor & Miller" baseAuthorityYear="1980" box="[377,508,1572,1596]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="hartwegi">
|
||
<emphasis id="B91DEA8FFFE7FFAAE1734570FE46C158" box="[377,508,1572,1596]" italics="true" pageId="5" pageNumber="376">V. hartwegi</emphasis>
|
||
</taxonomicName>
|
||
morph differs from the typical morph and all other species of
|
||
<taxonomicName id="4C694D1EFFE7FFAAE4EB4570FAA0C158" authorityName="Fernandez-Yepez" authorityYear="1969" box="[1249,1306,1573,1596]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91DEA8FFFE7FFAAE4EB4570FAA0C158" box="[1249,1306,1573,1596]" italics="true" pageId="5" pageNumber="376">Vieja</emphasis>
|
||
</taxonomicName>
|
||
by a more elongated body shape (
|
||
<figureCitation id="13522A18FFE7FFAAE1944512FE57C104" box="[414,493,1607,1632]" captionStart="FIGURE 4" captionStartId="7.[151,250,1125,1147]" captionTargetBox="[151,1437,193,1104]" captionTargetId="figure@7.[151,1437,193,1104]" captionTargetPageId="7" captionText="FIGURE 4. Live specimens of the rheophilic morph of Vieja hartwegi. A) Adult female in normal coloration, 97 mm SL, MZUNICACH 7161, Lagos de Colón, La Trinitaria, Chiapas (photo by M. A. Peralta-Meixueiro). B) Adult female in stress coloration, 65 mm SL, uncataloged specimen, Lagos de Colón, La Trinitaria, Chiapas. C) Adult female in stress coloration, 91 mm SL, MZUNICACH 7159, Tachinua River, Chicomuselo, Chiapas. D) Adult male in stress coloration, 80 mm SL, MZUNICACH 7157, Yayahuita River, Chicomuselo, Chiapas." httpUri="https://zenodo.org/record/1160375/files/figure.png" pageId="5" pageNumber="376">Figs. 4</figureCitation>
|
||
&
|
||
<figureCitation id="13522A18FFE7FFAAE21F4512FD9FC104" box="[533,549,1607,1632]" captionStart="FIGURE 6" captionStartId="9.[151,250,1685,1707]" captionTargetBox="[151,1436,235,1664]" captionTargetId="figure@9.[151,1436,193,1664]" captionTargetPageId="9" captionText="FIGURE 6. Preserved specimens of the rheophilic Vieja hartwegi morph. A) Adult female, MZUNICACH 7161, Lagos de Colón, La Trinitaria, Chiapas. B) Adult female, MZUNICACH 7159 [post-mortem fixed specimen], Yayahuita River, Chicomuselo, Chiapas." httpUri="https://zenodo.org/record/1160379/files/figure.png" pageId="5" pageNumber="376">6</figureCitation>
|
||
), which contrasts with a deeper body in the typical morph of
|
||
<taxonomicName id="4C694D1EFFE7FFAAE4E0451CFAD1C104" baseAuthorityName="Taylor & Miller" baseAuthorityYear="1980" box="[1258,1387,1608,1632]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="hartwegi">
|
||
<emphasis id="B91DEA8FFFE7FFAAE4E0451CFB42C104" box="[1258,1272,1609,1632]" italics="true" pageId="5" pageNumber="376">V</emphasis>
|
||
.
|
||
<emphasis id="B91DEA8FFFE7FFAAE50D451DFAD1C104" box="[1287,1387,1608,1632]" italics="true" pageId="5" pageNumber="376">hartwegi</emphasis>
|
||
</taxonomicName>
|
||
and other species of
|
||
<taxonomicName id="4C694D1EFFE7FFAAE1534538FE2AC1E0" authorityName="Fernandez-Yepez" authorityYear="1969" box="[345,400,1645,1668]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91DEA8FFFE7FFAAE1534538FE2AC1E0" box="[345,400,1645,1668]" italics="true" pageId="5" pageNumber="376">Vieja</emphasis>
|
||
</taxonomicName>
|
||
. The body shape of specimens is similar to that of other rheophilic herichthyine cichlid species, such as
|
||
<taxonomicName id="4C694D1EFFE7FFAAE14645C5FE43C1CC" authorityName="Regan" authorityYear="1905" box="[332,505,1680,1704]" class="Actinopterygii" family="Cichlidae" genus="Paraneetroplus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91DEA8FFFE7FFAAE14645C5FE43C1CC" box="[332,505,1680,1704]" italics="true" pageId="5" pageNumber="376">Paraneetroplus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B91DEA8FFFE7FFAAE22445C5FD17C1CC" box="[558,685,1680,1704]" italics="true" pageId="5" pageNumber="376">
|
||
<taxonomicName id="4C694D1EFFE7FFAAE22445C5FD12C1CC" authorityName="" authorityYear="2015" box="[558,680,1680,1704]" class="Actinopterygii" family="Cichlidae" genus="Rheoheros" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="genus">Rheoheros</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The rheophilic morph also differs from all members of
|
||
<taxonomicName id="4C694D1EFFE7FFAAE51245C5FAEBC1C3" authorityName="Fernandez-Yepez" authorityYear="1969" box="[1304,1361,1680,1703]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91DEA8FFFE7FFAAE51245C5FAEBC1C3" box="[1304,1361,1680,1703]" italics="true" pageId="5" pageNumber="376">Vieja</emphasis>
|
||
</taxonomicName>
|
||
by the presence of a simple great spot (
|
||
<figureCitation id="13522A18FFE7FFAAE20845E1FDFFC1A8" box="[514,581,1716,1741]" captionStart="FIGURE 4" captionStartId="7.[151,250,1125,1147]" captionTargetBox="[151,1437,193,1104]" captionTargetId="figure@7.[151,1437,193,1104]" captionTargetPageId="7" captionText="FIGURE 4. Live specimens of the rheophilic morph of Vieja hartwegi. A) Adult female in normal coloration, 97 mm SL, MZUNICACH 7161, Lagos de Colón, La Trinitaria, Chiapas (photo by M. A. Peralta-Meixueiro). B) Adult female in stress coloration, 65 mm SL, uncataloged specimen, Lagos de Colón, La Trinitaria, Chiapas. C) Adult female in stress coloration, 91 mm SL, MZUNICACH 7159, Tachinua River, Chicomuselo, Chiapas. D) Adult male in stress coloration, 80 mm SL, MZUNICACH 7157, Yayahuita River, Chicomuselo, Chiapas." httpUri="https://zenodo.org/record/1160375/files/figure.png" pageId="5" pageNumber="376">Fig. 4</figureCitation>
|
||
A-C) or an ocellus (
|
||
<figureCitation id="13522A18FFE7FFAAE32F45E1FCC7C1A8" box="[805,893,1716,1741]" captionStart="FIGURE 4" captionStartId="7.[151,250,1125,1147]" captionTargetBox="[151,1437,193,1104]" captionTargetId="figure@7.[151,1437,193,1104]" captionTargetPageId="7" captionText="FIGURE 4. Live specimens of the rheophilic morph of Vieja hartwegi. A) Adult female in normal coloration, 97 mm SL, MZUNICACH 7161, Lagos de Colón, La Trinitaria, Chiapas (photo by M. A. Peralta-Meixueiro). B) Adult female in stress coloration, 65 mm SL, uncataloged specimen, Lagos de Colón, La Trinitaria, Chiapas. C) Adult female in stress coloration, 91 mm SL, MZUNICACH 7159, Tachinua River, Chicomuselo, Chiapas. D) Adult male in stress coloration, 80 mm SL, MZUNICACH 7157, Yayahuita River, Chicomuselo, Chiapas." httpUri="https://zenodo.org/record/1160375/files/figure.png" pageId="5" pageNumber="376">Fig. 4D</figureCitation>
|
||
) at the center of the base of the caudal peduncle and on the origin of the caudal fin; other species of
|
||
<taxonomicName id="4C694D1EFFE7FFAAE2D6458DFCA9C18B" authorityName="Fernandez-Yepez" authorityYear="1969" box="[732,787,1752,1775]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91DEA8FFFE7FFAAE2D6458DFCA9C18B" box="[732,787,1752,1775]" italics="true" pageId="5" pageNumber="376">Vieja</emphasis>
|
||
</taxonomicName>
|
||
have a simple caudal spot (
|
||
<taxonomicName id="4C694D1EFFE7FFAAE447458CFB40C194" baseAuthorityName="Regan" baseAuthorityYear="1905" box="[1101,1274,1752,1776]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="maculicauda">
|
||
<emphasis id="B91DEA8FFFE7FFAAE447458CFB40C194" box="[1101,1274,1752,1776]" italics="true" pageId="5" pageNumber="376">V. maculicauda</emphasis>
|
||
</taxonomicName>
|
||
) or the caudal spot is fused with the midlateral stripe (the rest of the species; e.g.
|
||
<taxonomicName id="4C694D1EFFE7FFAAE38945A8FBB0C070" baseAuthorityName="Günther" baseAuthorityYear="1862" box="[899,1034,1788,1812]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="melanura">
|
||
<emphasis id="B91DEA8FFFE7FFAAE38945A8FC2AC070" box="[899,912,1789,1812]" italics="true" pageId="5" pageNumber="376">V</emphasis>
|
||
.
|
||
<emphasis id="B91DEA8FFFE7FFAAE39445A9FBB0C070" box="[926,1034,1788,1812]" italics="true" pageId="5" pageNumber="376">melanura</emphasis>
|
||
</taxonomicName>
|
||
). Also, in the rheophilic morph, the midlateral stripe is absent; only a midlateral blotch on the fifth bar is noticeable. This coloration pattern is similar to the syntopic
|
||
<taxonomicName id="4C694D1EFFE7FFAAE1584411FDCBC038" baseAuthorityName="Taylor & Miller" baseAuthorityYear="1980" box="[338,625,1860,1884]" class="Actinopterygii" family="Cichlidae" genus="Chiapaheros" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="grammodes">
|
||
<emphasis id="B91DEA8FFFE7FFAAE1584411FDCBC038" box="[338,625,1860,1884]" italics="true" pageId="5" pageNumber="376">Chiapaheros grammodes</emphasis>
|
||
</taxonomicName>
|
||
but can be differentiated by the great spot or ocellus on the caudal peduncle of the rheophilic
|
||
<taxonomicName id="4C694D1EFFE7FFAAE1DB443CFDEEC0E4" baseAuthorityName="Taylor & Miller" baseAuthorityYear="1980" box="[465,596,1896,1920]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="hartwegi">
|
||
<emphasis id="B91DEA8FFFE7FFAAE1DB443CFDEEC0E4" box="[465,596,1896,1920]" italics="true" pageId="5" pageNumber="376">V. hartwegi</emphasis>
|
||
</taxonomicName>
|
||
morph (
|
||
<figureCitation id="13522A18FFE7FFAAE2B14432FCBBC0E4" box="[699,769,1895,1920]" captionStart="FIGURE 6" captionStartId="9.[151,250,1685,1707]" captionTargetBox="[151,1436,235,1664]" captionTargetId="figure@9.[151,1436,193,1664]" captionTargetPageId="9" captionText="FIGURE 6. Preserved specimens of the rheophilic Vieja hartwegi morph. A) Adult female, MZUNICACH 7161, Lagos de Colón, La Trinitaria, Chiapas. B) Adult female, MZUNICACH 7159 [post-mortem fixed specimen], Yayahuita River, Chicomuselo, Chiapas." httpUri="https://zenodo.org/record/1160379/files/figure.png" pageId="5" pageNumber="376">Fig. 6</figureCitation>
|
||
) versus the simple oblong blotch in
|
||
<taxonomicName id="4C694D1EFFE7FFAAE4B8443DFAE3C0E4" baseAuthorityName="Taylor & Miller" baseAuthorityYear="1980" box="[1202,1369,1896,1920]" class="Actinopterygii" family="Cichlidae" genus="Chiapaheros" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="grammodes">
|
||
<emphasis id="B91DEA8FFFE7FFAAE4B8443DFAE3C0E4" box="[1202,1369,1896,1920]" italics="true" pageId="5" pageNumber="376">C. grammodes</emphasis>
|
||
</taxonomicName>
|
||
. This difference is always noticeable between juvenile and adult specimens. The midlateral blotch in
|
||
<taxonomicName id="4C694D1EFFE7FFAAE4B744D9FADAC0C0" baseAuthorityName="Taylor & Miller" baseAuthorityYear="1980" box="[1213,1376,1932,1956]" class="Actinopterygii" family="Cichlidae" genus="Chiapaheros" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="grammodes">
|
||
<emphasis id="B91DEA8FFFE7FFAAE4B744D9FB6AC0C7" box="[1213,1232,1932,1955]" italics="true" pageId="5" pageNumber="376">C</emphasis>
|
||
.
|
||
<emphasis id="B91DEA8FFFE7FFAAE4D744D9FADAC0C0" box="[1245,1376,1932,1956]" italics="true" pageId="5" pageNumber="376">grammodes</emphasis>
|
||
</taxonomicName>
|
||
is on the sixth bar versus the fifth bar in the rheophilic
|
||
<taxonomicName id="4C694D1EFFE7FFAAE2C244E4FCF2C0AC" baseAuthorityName="Taylor & Miller" baseAuthorityYear="1980" box="[712,840,1968,1992]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="hartwegi">
|
||
<emphasis id="B91DEA8FFFE7FFAAE2C244E4FCF2C0AC" box="[712,840,1968,1992]" italics="true" pageId="5" pageNumber="376">V. hartwegi</emphasis>
|
||
</taxonomicName>
|
||
morph.
|
||
<taxonomicName id="4C694D1EFFE7FFAAE3AD44E5FB7BC0AC" baseAuthorityName="Taylor & Miller" baseAuthorityYear="1980" box="[935,1217,1968,1992]" class="Actinopterygii" family="Cichlidae" genus="Chiapaheros" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="grammodes">
|
||
<emphasis id="B91DEA8FFFE7FFAAE3AD44E5FB7BC0AC" box="[935,1217,1968,1992]" italics="true" pageId="5" pageNumber="376">Chiapaheros grammodes</emphasis>
|
||
</taxonomicName>
|
||
has a series of thin brownish lines across interorbital region, snout and cheeks (the main character to diagnose this monotypic genus); in contrast, the rheophilic morph possesses two dark interorbital bars. Additionally,
|
||
<taxonomicName id="4C694D1EFFE5FFA8E4464676FB4AC25F" baseAuthorityName="Taylor & Miller" baseAuthorityYear="1980" box="[1100,1264,1315,1339]" class="Actinopterygii" family="Cichlidae" genus="Chiapaheros" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="378" phylum="Chordata" rank="species" species="grammodes">
|
||
<emphasis id="B91DEA8FFFE5FFA8E4464676FB4AC25F" box="[1100,1264,1315,1339]" italics="true" pageId="7" pageNumber="378">C. grammodes</emphasis>
|
||
</taxonomicName>
|
||
differs notably from the genus
|
||
<taxonomicName id="4C694D1EFFE5FFA8E14F4612FEC6C23A" authorityName="Fernandez-Yepez" authorityYear="1969" box="[325,380,1351,1374]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="378" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91DEA8FFFE5FFA8E14F4612FEC6C23A" box="[325,380,1351,1374]" italics="true" pageId="7" pageNumber="378">Vieja</emphasis>
|
||
</taxonomicName>
|
||
by the large head and enlarged snout, a large mouth with prognathous lower jaw. In preserved specimens, the rheophilic morph possesses a horizontal stripe above the middle of the body (
|
||
<figureCitation id="13522A18FFE5FFA8E4CE463FFAB0C2E7" box="[1220,1290,1386,1411]" captionStart="FIGURE 6" captionStartId="9.[151,250,1685,1707]" captionTargetBox="[151,1436,235,1664]" captionTargetId="figure@9.[151,1436,193,1664]" captionTargetPageId="9" captionText="FIGURE 6. Preserved specimens of the rheophilic Vieja hartwegi morph. A) Adult female, MZUNICACH 7161, Lagos de Colón, La Trinitaria, Chiapas. B) Adult female, MZUNICACH 7159 [post-mortem fixed specimen], Yayahuita River, Chicomuselo, Chiapas." httpUri="https://zenodo.org/record/1160379/files/figure.png" pageId="7" pageNumber="378">Fig. 6</figureCitation>
|
||
), and in live specimens the band is noticeable under stress conditions (
|
||
<figureCitation id="13522A18FFE5FFA8E32646DBFCCBC2C3" box="[812,881,1422,1447]" captionStart="FIGURE 4" captionStartId="7.[151,250,1125,1147]" captionTargetBox="[151,1437,193,1104]" captionTargetId="figure@7.[151,1437,193,1104]" captionTargetPageId="7" captionText="FIGURE 4. Live specimens of the rheophilic morph of Vieja hartwegi. A) Adult female in normal coloration, 97 mm SL, MZUNICACH 7161, Lagos de Colón, La Trinitaria, Chiapas (photo by M. A. Peralta-Meixueiro). B) Adult female in stress coloration, 65 mm SL, uncataloged specimen, Lagos de Colón, La Trinitaria, Chiapas. C) Adult female in stress coloration, 91 mm SL, MZUNICACH 7159, Tachinua River, Chicomuselo, Chiapas. D) Adult male in stress coloration, 80 mm SL, MZUNICACH 7157, Yayahuita River, Chicomuselo, Chiapas." httpUri="https://zenodo.org/record/1160375/files/figure.png" pageId="7" pageNumber="378">Fig. 4</figureCitation>
|
||
B-D). This stripe is similar to the dorsolateral or second stripe of
|
||
<taxonomicName id="4C694D1EFFE5FFA8E15846E6FE69C2AF" baseAuthorityName="Taylor & Miller" baseAuthorityYear="1980" box="[338,467,1459,1483]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="378" phylum="Chordata" rank="species" species="hartwegi">
|
||
<emphasis id="B91DEA8FFFE5FFA8E15846E6FE69C2AF" box="[338,467,1459,1483]" italics="true" pageId="7" pageNumber="378">V. hartwegi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C694D1EFFE5FFA8E20146E6FD2CC2AF" baseAuthorityName="Steindachner" baseAuthorityYear="1864" box="[523,662,1459,1483]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="378" phylum="Chordata" rank="species" species="bifasciata">
|
||
<emphasis id="B91DEA8FFFE5FFA8E20146E6FD2CC2AF" box="[523,662,1459,1483]" italics="true" pageId="7" pageNumber="378">V. bifasciata</emphasis>
|
||
</taxonomicName>
|
||
, formed by the fusion of lateral blotches and always separated from the caudal blotch (
|
||
<figureCitation id="13522A18FFE5FFA8E17B4683FE00C28B" box="[369,442,1494,1519]" captionStart="FIGURE 6" captionStartId="9.[151,250,1685,1707]" captionTargetBox="[151,1436,235,1664]" captionTargetId="figure@9.[151,1436,193,1664]" captionTargetPageId="9" captionText="FIGURE 6. Preserved specimens of the rheophilic Vieja hartwegi morph. A) Adult female, MZUNICACH 7161, Lagos de Colón, La Trinitaria, Chiapas. B) Adult female, MZUNICACH 7159 [post-mortem fixed specimen], Yayahuita River, Chicomuselo, Chiapas." httpUri="https://zenodo.org/record/1160379/files/figure.png" pageId="7" pageNumber="378">Fig. 6</figureCitation>
|
||
). Other members of
|
||
<taxonomicName id="4C694D1EFFE5FFA8E2B8468DFD51C28B" authorityName="Fernandez-Yepez" authorityYear="1969" box="[690,747,1496,1519]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="378" phylum="Chordata" rank="genus">
|
||
<emphasis id="B91DEA8FFFE5FFA8E2B8468DFD51C28B" box="[690,747,1496,1519]" italics="true" pageId="7" pageNumber="378">Vieja</emphasis>
|
||
</taxonomicName>
|
||
possess a strong midlateral stripe at level of mid-body (
|
||
<taxonomicName id="4C694D1EFFE5FFA8E58D468DFEBAC177" baseAuthorityName="Werner & Stawikowski" baseAuthorityYear="1987" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="378" phylum="Chordata" rank="species" species="breidohri">
|
||
<emphasis id="B91DEA8FFFE5FFA8E58D468DFEBAC177" italics="true" pageId="7" pageNumber="378">V. breidohri</emphasis>
|
||
</taxonomicName>
|
||
), below mid-body (
|
||
<emphasis id="B91DEA8FFFE5FFA8E1ED46AEFBA5C176" box="[487,1055,1531,1555]" italics="true" pageId="7" pageNumber="378">
|
||
<taxonomicName id="4C694D1EFFE5FFA8E1ED46AEFDCCC177" baseAuthorityName="Steindachner" baseAuthorityYear="1864" box="[487,630,1531,1555]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="378" phylum="Chordata" rank="species" species="bifasciata">V. bifasciata</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C694D1EFFE5FFA8E28F46AEFCACC177" baseAuthorityName="Günther" baseAuthorityYear="1860" box="[645,790,1531,1555]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="378" phylum="Chordata" rank="species" species="fenestrata">V. fenestrata</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C694D1EFFE5FFA8E32946AEFC13C177" baseAuthorityName="Günther" baseAuthorityYear="1864" box="[803,937,1531,1555]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="378" phylum="Chordata" rank="species" species="guttulata">V. guttulata</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C694D1EFFE5FFA8E3BD46AEFBA5C176" baseAuthorityName="Meek" baseAuthorityYear="1905" box="[951,1055,1531,1554]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="378" phylum="Chordata" rank="species" species="zonata">V. zonata</taxonomicName>
|
||
</emphasis>
|
||
), or variable, as in the typical
|
||
<taxonomicName id="4C694D1EFFE5FFA8E58D46AEFF41C153" baseAuthorityName="Taylor & Miller" baseAuthorityYear="1980" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="378" phylum="Chordata" rank="species" species="hartwegi">
|
||
<emphasis id="B91DEA8FFFE5FFA8E58D46AEFF41C153" italics="true" pageId="7" pageNumber="378">V. hartwegi</emphasis>
|
||
</taxonomicName>
|
||
morph (
|
||
<figureCitation id="13522A18FFE5FFA8E150454BFE1DC153" box="[346,423,1566,1591]" captionStart="FIGURE 3" captionStartId="6.[151,250,1858,1880]" captionTargetBox="[192,1411,225,1818]" captionTargetId="figure@6.[151,1436,193,1837]" captionTargetPageId="6" captionText="FIGURE 3. Live specimens of the typical morph of Vieja hartwegi, showing ontogenic variation in coloration pattern. A) Juvenile, 75 mm SL, MZUNICACH 6761, La Angostura Reservoir, La Concordia, Chiapas. B) Adult, 144 mm SL, aquarium specimen, Cacahuanó River, Ocozocoautla, Chiapas. C) Large adult, 250 mm SL, uncataloged specimen from a fishery, Grijalva River (main channel at Tres Picos), Copainalá, Chiapas (photo by J. M. López-Vila)." httpUri="https://zenodo.org/record/1160372/files/figure.png" pageId="7" pageNumber="378">Figs. 3</figureCitation>
|
||
&
|
||
<figureCitation id="13522A18FFE5FFA8E1C1454BFE61C153" box="[459,475,1566,1591]" captionStart="FIGURE5" captionStartId="8.[151,250,1823,1845]" captionTargetBox="[220,1316,219,1782]" captionTargetId="figure@8.[203,1338,193,1801]" captionTargetPageId="8" captionText="FIGURE5. Fixed specimensof the typical Viejahartwegi morph,showing the variation inbody shapeand the ontogenetic and individual variation in coloration patterns. A) Holotype (adult in breeding coloration), UMMZ 207701. B) Chiquito River, Chiapa de Corzo (juv.), MZUNICACH 7053. C) El Zapote River, Chiapa de Corzo (juv.), MZUNICACH 7210. D) La Angostura Reservoir, La Concordia (juv.), MZUNICACH 7154. E) Totopac River, Tecpatán (subadult), MZUNICACH 1053. F) Lagos de Colón, La Trinitaria (subadult), MZUNICACH 7065.G) CopanóRiver, Ostuacán (adult), MZUNICACH 7208.H- I) Grijalva River (main cannel), Copainalá (adults), MZUNICACH 3145. J) Santo Domingo River, Chiapa de Corzo (adult), MZUNICACH 6245." httpUri="https://zenodo.org/record/1160377/files/figure.png" pageId="7" pageNumber="378">5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF166615FFE4FFA9E09D4417FBF0C0D0" httpUri="https://zenodo.org/record/1160372/files/figure.png" pageId="6" pageNumber="377" targetBox="[192,1411,225,1818]" targetPageId="6">
|
||
<paragraph id="8BD6369DFFE4FFA9E09D4417FBF0C0D0" blockId="6.[151,1436,1858,1972]" pageId="6" pageNumber="377">
|
||
<emphasis id="B91DEA8FFFE4FFA9E09D4417FEACC033" bold="true" box="[151,278,1858,1880]" pageId="6" pageNumber="377">FIGURE 3.</emphasis>
|
||
Live specimens of the typical morph of
|
||
<taxonomicName id="4C694D1EFFE4FFA9E2C24416FCE5C03C" baseAuthorityName="Taylor & Miller" baseAuthorityYear="1980" box="[712,863,1858,1880]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="377" phylum="Chordata" rank="species" species="hartwegi">
|
||
<emphasis id="B91DEA8FFFE4FFA9E2C24416FCE5C03C" box="[712,863,1858,1880]" italics="true" pageId="6" pageNumber="377">Vieja hartwegi</emphasis>
|
||
</taxonomicName>
|
||
, showing ontogenic variation in coloration pattern. A) Juvenile, 75 mm SL, MZUNICACH 6761, La Angostura Reservoir, La Concordia, Chiapas. B) Adult, 144 mm SL, aquarium specimen, Cacahuanó River, Ocozocoautla, Chiapas. C) Large adult, 250 mm SL, uncataloged specimen from a fishery, Grijalva River (main channel at Tres Picos), Copainalá, Chiapas (photo by J. M. López-Vila).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF166615FFE5FFA8E09D4730FCABC392" httpUri="https://zenodo.org/record/1160375/files/figure.png" pageId="7" pageNumber="378" targetBox="[151,1437,193,1104]" targetPageId="7">
|
||
<paragraph id="8BD6369DFFE5FFA8E09D4730FCABC392" blockId="7.[151,1436,1125,1270]" pageId="7" pageNumber="378">
|
||
<emphasis id="B91DEA8FFFE5FFA8E09D4730FEAFC31E" bold="true" box="[151,277,1125,1147]" pageId="7" pageNumber="378">FIGURE 4.</emphasis>
|
||
Live specimens of the rheophilic morph of
|
||
<emphasis id="B91DEA8FFFE5FFA8E2ED4733FC3FC31F" box="[743,901,1125,1147]" italics="true" pageId="7" pageNumber="378">
|
||
<taxonomicName id="4C694D1EFFE5FFA8E2ED4733FC3BC31F" baseAuthorityName="Taylor & Miller" baseAuthorityYear="1980" box="[743,897,1125,1147]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="378" phylum="Chordata" rank="species" species="hartwegi">Vieja hartwegi</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
A) Adult female in normal coloration, 97 mm SL, MZUNICACH 7161, Lagos de Colón, La Trinitaria, Chiapas (photo by M. A. Peralta-Meixueiro). B) Adult female in stress coloration, 65 mm SL, uncataloged specimen, Lagos de Colón, La Trinitaria, Chiapas. C) Adult female in stress coloration, 91 mm SL, MZUNICACH 7159, Tachinua River, Chicomuselo, Chiapas. D) Adult male in stress coloration, 80 mm SL, MZUNICACH 7157, Yayahuita River, Chicomuselo, Chiapas.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3736516FFE5FFA2E0CD4517FC24C1D9" lastPageId="13" lastPageNumber="384" pageId="7" pageNumber="378" type="description">
|
||
<paragraph id="8BD6369DFFE5FFA6E0CD4517FE81C0F1" blockId="7.[151,1437,1314,2029]" lastBlockId="9.[151,1437,1808,2013]" lastPageId="9" lastPageNumber="380" pageId="7" pageNumber="378">
|
||
<emphasis id="B91DEA8FFFE5FFA8E0CD4517FEE3C13F" bold="true" box="[199,345,1602,1627]" pageId="7" pageNumber="378">Description.</emphasis>
|
||
Morphometric and meristic data summarized in
|
||
<tableCitation id="C6EB0326FFE5FFA8E3984517FBA8C13F" box="[914,1042,1602,1627]" captionStart-0="TABLE 1" captionStart-1="TABLE2" captionStartId-0="4.[120,200,499,520]" captionStartId-1="15.[138,213,471,491]" captionText-0="TABLE 1.GenBank accession numbersforCyt-b sequences used in thisstudy and locationsof the newly sequences.Codesin brackets correspond toindividualsof V.hartwegi in Fig.. °Sample from near typelocality.* Indicates rheophilic morph." captionText-1="TABLE2. Morphometricdatafor the two morphsof Vieja hartwegi and related species. Measurements are presented as percent ofstandard length or percent ofhead length.SD=standard deviation,Min=Minimum,Max= Maximum." pageId="7" pageNumber="378">Tables 1–2</tableCitation>
|
||
. Maximum size
|
||
<quantity id="4C919B78FFE5FFA8E4DF4516FA8CC13E" box="[1237,1334,1603,1627]" metricMagnitude="-1" metricUnit="m" metricValue="3.4" pageId="7" pageNumber="378" unit="mm" value="340.0">340 mm</quantity>
|
||
SL with weight of
|
||
<quantity id="4C919B78FFE5FFA8E1024532FEEFC1E4" box="[264,341,1639,1664]" metricMagnitude="0" metricUnit="kg" metricValue="1.028" pageId="7" pageNumber="378" unit="g" value="1028.0">1028 g</quantity>
|
||
in Grijalva River main channel, municipality of Chicoasén,
|
||
<collectingRegion id="49ADF87FFFE5FFA8E3F34533FBE9C11B" box="[1017,1107,1638,1663]" country="Mexico" name="Chiapas" pageId="7" pageNumber="378">Chiapas</collectingRegion>
|
||
(observation from fishery La Cuevita). Meristics for the typical
|
||
<taxonomicName id="4C694D1EFFE5FFA8E21745DEFD27C1C7" baseAuthorityName="Taylor & Miller" baseAuthorityYear="1980" box="[541,669,1675,1699]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="378" phylum="Chordata" rank="species" species="hartwegi">
|
||
<emphasis id="B91DEA8FFFE5FFA8E21745DEFD27C1C7" box="[541,669,1675,1699]" italics="true" pageId="7" pageNumber="378">V. hartwegi</emphasis>
|
||
</taxonomicName>
|
||
morph (
|
||
<figureCitation id="13522A18FFE5FFA8E2F445DFFCF6C1C7" box="[766,844,1674,1699]" captionStart="FIGURE 3" captionStartId="6.[151,250,1858,1880]" captionTargetBox="[192,1411,225,1818]" captionTargetId="figure@6.[151,1436,193,1837]" captionTargetPageId="6" captionText="FIGURE 3. Live specimens of the typical morph of Vieja hartwegi, showing ontogenic variation in coloration pattern. A) Juvenile, 75 mm SL, MZUNICACH 6761, La Angostura Reservoir, La Concordia, Chiapas. B) Adult, 144 mm SL, aquarium specimen, Cacahuanó River, Ocozocoautla, Chiapas. C) Large adult, 250 mm SL, uncataloged specimen from a fishery, Grijalva River (main channel at Tres Picos), Copainalá, Chiapas (photo by J. M. López-Vila)." httpUri="https://zenodo.org/record/1160372/files/figure.png" pageId="7" pageNumber="378">Figs. 3</figureCitation>
|
||
&
|
||
<figureCitation id="13522A18FFE5FFA8E37845DFFC38C1C7" box="[882,898,1674,1699]" captionStart="FIGURE5" captionStartId="8.[151,250,1823,1845]" captionTargetBox="[220,1316,219,1782]" captionTargetId="figure@8.[203,1338,193,1801]" captionTargetPageId="8" captionText="FIGURE5. Fixed specimensof the typical Viejahartwegi morph,showing the variation inbody shapeand the ontogenetic and individual variation in coloration patterns. A) Holotype (adult in breeding coloration), UMMZ 207701. B) Chiquito River, Chiapa de Corzo (juv.), MZUNICACH 7053. C) El Zapote River, Chiapa de Corzo (juv.), MZUNICACH 7210. D) La Angostura Reservoir, La Concordia (juv.), MZUNICACH 7154. E) Totopac River, Tecpatán (subadult), MZUNICACH 1053. F) Lagos de Colón, La Trinitaria (subadult), MZUNICACH 7065.G) CopanóRiver, Ostuacán (adult), MZUNICACH 7208.H- I) Grijalva River (main cannel), Copainalá (adults), MZUNICACH 3145. J) Santo Domingo River, Chiapa de Corzo (adult), MZUNICACH 6245." httpUri="https://zenodo.org/record/1160377/files/figure.png" pageId="7" pageNumber="378">5</figureCitation>
|
||
):
|
||
<emphasis id="B91DEA8FFFE5FFA8E39345DEFC6FC1C6" box="[921,981,1675,1698]" italics="true" pageId="7" pageNumber="378">Fins:</emphasis>
|
||
D XVI–XVIII (mode 17), 11–14 (mode 12); A VI–VII (mode 6), 8–10 (mode 9); pectoral rays 15–17 (mode 16).
|
||
<emphasis id="B91DEA8FFFE5FFA8E3D345FAFB93C1A3" box="[985,1065,1711,1735]" italics="true" pageId="7" pageNumber="378">Scales:</emphasis>
|
||
Lateral series 30–34 (mode 32); upper lateral line 16–23 (mode 20); lower lateral line 8–15 (mode 12); in one specimen, pored scales of upper lateral line disrupted by non-pored scales (MZUNICACH 7158, specimen 2); subsidiary pored scales on caudal fin, usually 0–3 (mode 2) continuous with lower lateral line or interrupted by one non-pored scale; isolated pored scales (1–3) on the caudal-fin base, usually 1–2 scales above or below the subsidiary pored scales; predorsal squamation pattern irregular; scale row overlap 2–4 (mode 2); pectoral-pelvic scale rows 6–8 (mode 7); scale rows from lateral line to base of first dorsal-fin ray (not including the scaly sheet along the base), 3–4 1/2 (mode 3 1/2); scale rows from lateral line to base of first dorsal-fin spine 5–7 (mode 6); scale rows from origin of anal fin to lower lateral line 8–10 (mode 9); circumpeduncular scales 16–22 (mode 20); scale rows on cheek 5–7 (mode 5), one specimen with irregular pattern, not forming rows (ECOSC 7543, specimen 1). Cheek and opercle fully scaled, scales cycloid; dorsal portion of head and predorsal scales cycloid; base of dorsal and anal fins and scaly sheath along fin bases cycloid; lateral body scales ctenoid; lateral chest scales ctenoid, smaller than lateral body scales; ventral chest scales cycloid.
|
||
</paragraph>
|
||
<caption id="DF166615FFEAFFA7E09D444AFECAC089" httpUri="https://zenodo.org/record/1160377/files/figure.png" pageId="8" pageNumber="379" targetBox="[220,1316,219,1782]" targetPageId="8">
|
||
<paragraph id="8BD6369DFFEAFFA7E09D444AFECAC089" blockId="8.[151,1436,1823,2029]" pageId="8" pageNumber="379">
|
||
<emphasis id="B91DEA8FFFEAFFA7E09D444AFEA8C050" bold="true" box="[151,274,1823,1845]" pageId="8" pageNumber="379">FIGURE 5.</emphasis>
|
||
Fixed specimens of the typical
|
||
<taxonomicName id="4C694D1EFFEAFFA7E2584475FD5FC051" baseAuthorityName="Taylor & Miller" baseAuthorityYear="1980" box="[594,741,1823,1845]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="379" phylum="Chordata" rank="species" species="hartwegi">
|
||
<emphasis id="B91DEA8FFFEAFFA7E2584475FD5FC051" box="[594,741,1823,1845]" italics="true" pageId="8" pageNumber="379">Vieja hartwegi</emphasis>
|
||
</taxonomicName>
|
||
morph, showing the variation in body shape and the ontogenetic and individual variation in coloration patterns. A) Holotype (adult in breeding coloration), UMMZ 207701. B) Chiquito River, Chiapa de Corzo (juv.), MZUNICACH 7053. C) El Zapote River, Chiapa de Corzo (juv.), MZUNICACH 7210. D) La Angostura Reservoir, La Concordia (juv.), MZUNICACH 7154. E) Totopac River, Tecpatán (subadult), MZUNICACH 1053. F) Lagos de Colón, La Trinitaria (subadult), MZUNICACH 7065. G) Copanó River, Ostuacán (adult), MZUNICACH 7208. H- I) Grijalva River (main cannel), Copainalá (adults), MZUNICACH 3145. J) Santo Domingo River, Chiapa de Corzo (adult), MZUNICACH 6245.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF166615FFEBFFA6E09D45C0FE38C18C" httpUri="https://zenodo.org/record/1160379/files/figure.png" pageId="9" pageNumber="380" targetBox="[151,1436,235,1664]" targetPageId="9">
|
||
<paragraph id="8BD6369DFFEBFFA6E09D45C0FE38C18C" blockId="9.[151,1436,1685,1768]" pageId="9" pageNumber="380">
|
||
<emphasis id="B91DEA8FFFEBFFA6E09D45C0FEAFC1CE" bold="true" box="[151,277,1685,1707]" pageId="9" pageNumber="380">FIGURE 6.</emphasis>
|
||
Preserved specimens of the rheophilic
|
||
<taxonomicName id="4C694D1EFFEBFFA6E2BB45C3FCFDC1CF" baseAuthorityName="Taylor & Miller" baseAuthorityYear="1980" box="[689,839,1685,1707]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="380" phylum="Chordata" rank="species" species="hartwegi">
|
||
<emphasis id="B91DEA8FFFEBFFA6E2BB45C3FCFDC1CF" box="[689,839,1685,1707]" italics="true" pageId="9" pageNumber="380">Vieja hartwegi</emphasis>
|
||
</taxonomicName>
|
||
morph. A) Adult female, MZUNICACH 7161, Lagos de Colón, La Trinitaria, Chiapas. B) Adult female, MZUNICACH 7159 [post-mortem fixed specimen], Yayahuita River, Chicomuselo, Chiapas.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD6369DFFEBFFA5E0CD44F5FD5CC6EC" blockId="9.[151,1437,1808,2013]" lastBlockId="10.[151,1437,151,968]" lastPageId="10" lastPageNumber="381" pageId="9" pageNumber="380">
|
||
Body shape moderately deep or slightly slender (
|
||
<figureCitation id="13522A18FFEBFFA6E2E444F5FC81C0DC" box="[750,827,1952,1977]" captionStart="FIGURE 3" captionStartId="6.[151,250,1858,1880]" captionTargetBox="[192,1411,225,1818]" captionTargetId="figure@6.[151,1436,193,1837]" captionTargetPageId="6" captionText="FIGURE 3. Live specimens of the typical morph of Vieja hartwegi, showing ontogenic variation in coloration pattern. A) Juvenile, 75 mm SL, MZUNICACH 6761, La Angostura Reservoir, La Concordia, Chiapas. B) Adult, 144 mm SL, aquarium specimen, Cacahuanó River, Ocozocoautla, Chiapas. C) Large adult, 250 mm SL, uncataloged specimen from a fishery, Grijalva River (main channel at Tres Picos), Copainalá, Chiapas (photo by J. M. López-Vila)." httpUri="https://zenodo.org/record/1160372/files/figure.png" pageId="9" pageNumber="380">Figs. 3</figureCitation>
|
||
&
|
||
<figureCitation id="13522A18FFEBFFA6E35544F5FCD4C0DD" box="[863,878,1952,1977]" captionStart="FIGURE5" captionStartId="8.[151,250,1823,1845]" captionTargetBox="[220,1316,219,1782]" captionTargetId="figure@8.[203,1338,193,1801]" captionTargetPageId="8" captionText="FIGURE5. Fixed specimensof the typical Viejahartwegi morph,showing the variation inbody shapeand the ontogenetic and individual variation in coloration patterns. A) Holotype (adult in breeding coloration), UMMZ 207701. B) Chiquito River, Chiapa de Corzo (juv.), MZUNICACH 7053. C) El Zapote River, Chiapa de Corzo (juv.), MZUNICACH 7210. D) La Angostura Reservoir, La Concordia (juv.), MZUNICACH 7154. E) Totopac River, Tecpatán (subadult), MZUNICACH 1053. F) Lagos de Colón, La Trinitaria (subadult), MZUNICACH 7065.G) CopanóRiver, Ostuacán (adult), MZUNICACH 7208.H- I) Grijalva River (main cannel), Copainalá (adults), MZUNICACH 3145. J) Santo Domingo River, Chiapa de Corzo (adult), MZUNICACH 6245." httpUri="https://zenodo.org/record/1160377/files/figure.png" pageId="9" pageNumber="380">5</figureCitation>
|
||
), in very large adults (>
|
||
<quantity id="4C919B78FFEBFFA6E47D44F5FB60C0DC" box="[1143,1242,1952,1976]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="380" unit="mm" value="200.0">200 mm</quantity>
|
||
SL) body deeper (
|
||
<figureCitation id="13522A18FFEBFFA6E0954491FF40C0B8" box="[159,250,1988,2013]" captionStart="FIGURE 3" captionStartId="6.[151,250,1858,1880]" captionTargetBox="[192,1411,225,1818]" captionTargetId="figure@6.[151,1436,193,1837]" captionTargetPageId="6" captionText="FIGURE 3. Live specimens of the typical morph of Vieja hartwegi, showing ontogenic variation in coloration pattern. A) Juvenile, 75 mm SL, MZUNICACH 6761, La Angostura Reservoir, La Concordia, Chiapas. B) Adult, 144 mm SL, aquarium specimen, Cacahuanó River, Ocozocoautla, Chiapas. C) Large adult, 250 mm SL, uncataloged specimen from a fishery, Grijalva River (main channel at Tres Picos), Copainalá, Chiapas (photo by J. M. López-Vila)." httpUri="https://zenodo.org/record/1160372/files/figure.png" pageId="9" pageNumber="380">Fig. 3C</figureCitation>
|
||
); laterally compresssed; predorsal contour steep, dorsal head profile in front of orbit usually convex in juveniles, straight or slightly rounded in adults; the individuals with elongated snout, concavity present over the eyes; prepelvic contour gently convex; caudal peduncle short, deeper than long. Mouth small, usually low in position in juveniles and preadults, terminal and slightly oblique in large adults; individuals of upper reaches with more slender bodies, mouth usually terminal (
|
||
<figureCitation id="13522A18FFE8FFA5E2954251FD56C679" box="[671,748,260,285]" captionStart="FIGURE5" captionStartId="8.[151,250,1823,1845]" captionTargetBox="[220,1316,219,1782]" captionTargetId="figure@8.[203,1338,193,1801]" captionTargetPageId="8" captionText="FIGURE5. Fixed specimensof the typical Viejahartwegi morph,showing the variation inbody shapeand the ontogenetic and individual variation in coloration patterns. A) Holotype (adult in breeding coloration), UMMZ 207701. B) Chiquito River, Chiapa de Corzo (juv.), MZUNICACH 7053. C) El Zapote River, Chiapa de Corzo (juv.), MZUNICACH 7210. D) La Angostura Reservoir, La Concordia (juv.), MZUNICACH 7154. E) Totopac River, Tecpatán (subadult), MZUNICACH 1053. F) Lagos de Colón, La Trinitaria (subadult), MZUNICACH 7065.G) CopanóRiver, Ostuacán (adult), MZUNICACH 7208.H- I) Grijalva River (main cannel), Copainalá (adults), MZUNICACH 3145. J) Santo Domingo River, Chiapa de Corzo (adult), MZUNICACH 6245." httpUri="https://zenodo.org/record/1160377/files/figure.png" pageId="10" pageNumber="381">Figs. 5</figureCitation>
|
||
F-G). Upper jaw in juveniles and preadults usually projected beyond lower jaw (
|
||
<figureCitation id="13522A18FFE8FFA5E1794272FE0CC624" box="[371,438,295,320]" captionStart="FIGURE5" captionStartId="8.[151,250,1823,1845]" captionTargetBox="[220,1316,219,1782]" captionTargetId="figure@8.[203,1338,193,1801]" captionTargetPageId="8" captionText="FIGURE5. Fixed specimensof the typical Viejahartwegi morph,showing the variation inbody shapeand the ontogenetic and individual variation in coloration patterns. A) Holotype (adult in breeding coloration), UMMZ 207701. B) Chiquito River, Chiapa de Corzo (juv.), MZUNICACH 7053. C) El Zapote River, Chiapa de Corzo (juv.), MZUNICACH 7210. D) La Angostura Reservoir, La Concordia (juv.), MZUNICACH 7154. E) Totopac River, Tecpatán (subadult), MZUNICACH 1053. F) Lagos de Colón, La Trinitaria (subadult), MZUNICACH 7065.G) CopanóRiver, Ostuacán (adult), MZUNICACH 7208.H- I) Grijalva River (main cannel), Copainalá (adults), MZUNICACH 3145. J) Santo Domingo River, Chiapa de Corzo (adult), MZUNICACH 6245." httpUri="https://zenodo.org/record/1160377/files/figure.png" pageId="10" pageNumber="381">Fig. 5</figureCitation>
|
||
A-D), large adults isognathous (
|
||
<figureCitation id="13522A18FFE8FFA5E3294272FCDCC624" box="[803,870,295,320]" captionStart="FIGURE5" captionStartId="8.[151,250,1823,1845]" captionTargetBox="[220,1316,219,1782]" captionTargetId="figure@8.[203,1338,193,1801]" captionTargetPageId="8" captionText="FIGURE5. Fixed specimensof the typical Viejahartwegi morph,showing the variation inbody shapeand the ontogenetic and individual variation in coloration patterns. A) Holotype (adult in breeding coloration), UMMZ 207701. B) Chiquito River, Chiapa de Corzo (juv.), MZUNICACH 7053. C) El Zapote River, Chiapa de Corzo (juv.), MZUNICACH 7210. D) La Angostura Reservoir, La Concordia (juv.), MZUNICACH 7154. E) Totopac River, Tecpatán (subadult), MZUNICACH 1053. F) Lagos de Colón, La Trinitaria (subadult), MZUNICACH 7065.G) CopanóRiver, Ostuacán (adult), MZUNICACH 7208.H- I) Grijalva River (main cannel), Copainalá (adults), MZUNICACH 3145. J) Santo Domingo River, Chiapa de Corzo (adult), MZUNICACH 6245." httpUri="https://zenodo.org/record/1160377/files/figure.png" pageId="10" pageNumber="381">Fig. 5</figureCitation>
|
||
G-J). Snout moderately pronounced or sometimes flattish. Lips of both jaws moderately thickened. Frenum well developed. Maxillary cleft position, variable but usually at level of dorsal margin of pectoral fin base.
|
||
</paragraph>
|
||
<paragraph id="8BD6369DFFE8FFA5E0CD42C1FCA3C5E1" blockId="10.[151,1437,151,968]" pageId="10" pageNumber="381">
|
||
<emphasis id="B91DEA8FFFE8FFA5E0CD42C1FE62C6C8" box="[199,472,404,428]" italics="true" pageId="10" pageNumber="381">Gillrakers on first arch:</emphasis>
|
||
lower limb 7–11 (mode 9), one specimen found in sulphidic environment possessed only two gillrakers (MZUNICACH 1065, specimen 1); upper limb 1–3 (mode 2); total 9–13 (mode 11), one specimen with only 5 (see line above). Gillrakers moderately elongate, shorter and thickened in adults and more elongated in juveniles; tips pointed or rounded, commonly anterior gillrakers on lower limb shorter and rounded and posterior ones larger and pointed. One specimen, with posterior gillrakers bifurcated on lower limb (MZUNICACH 3145, specimen 1). Outer teeth in both jaws recurved, conical; anterior teeth enlarged with small posterior cusp; 3–4 irregular series of inner smaller teeth.
|
||
</paragraph>
|
||
<paragraph id="8BD6369DFFE8FFA5E0CD41C5FAAFC4AC" blockId="10.[151,1437,151,968]" pageId="10" pageNumber="381">
|
||
Lower pharyngeal tooth-plate (
|
||
<figureCitation id="13522A18FFE8FFA5E22141DAFD32C5CC" box="[555,648,655,680]" captionStart="FIGURE 7" captionStartId="10.[151,250,1908,1930]" captionTargetBox="[169,1418,1029,1870]" captionTargetId="figure@10.[150,1436,1021,1887]" captionTargetPageId="10" captionText="FIGURE 7. Lower pharyngeal tooth plate and urohyal bone of the two morphs of V. hartwegi. A) Typical morph, MZUNICACH 6761.B)Rheophilic morph, MZUNICACH 7157(photos by K.Ramírez-Moreno [pharyngeal tooth plates] and G. Morales-Flores [urohyal bones])." httpUri="https://zenodo.org/record/1160381/files/figure.png" pageId="10" pageNumber="381">Fig. 7A</figureCitation>
|
||
), dissected from one specimen (MZUNICACH 6761,
|
||
<quantity id="4C919B78FFE8FFA5E4F141C5FAD8C5C3" box="[1275,1378,656,680]" metricMagnitude="-2" metricUnit="m" metricValue="8.93" pageId="10" pageNumber="381" unit="mm" value="89.3">89.3 mm</quantity>
|
||
SL), dorsoventrally compressed, with short anterior process and posterior processes slightly enlarged; wider than long; posterior margin concave at center and convex at margins; teeth conical, enlarged posteriorly and shorter anteriorly, with antrorse cusp; second cusp small present at the base; three medial rows on each side, mainly the first, notably bigger and more robust posteriorly than the rest; 26 total (12/
|
||
<quantity id="4C919B78FFE8FFA5E3354075FCCDC45C" box="[831,887,799,824]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" pageId="10" pageNumber="381" unit="in" value="14.0">14 in</quantity>
|
||
each side) teeth in posterior row; 7 and 6 teeth in each near median row. Urohyal bone with posterior edge concave, dorsally slight convex with a short, curved spine (
|
||
<figureCitation id="13522A18FFE8FFA5E0954032FF42C4E4" box="[159,248,871,896]" captionStart="FIGURE 7" captionStartId="10.[151,250,1908,1930]" captionTargetBox="[169,1418,1029,1870]" captionTargetId="figure@10.[150,1436,1021,1887]" captionTargetPageId="10" captionText="FIGURE 7. Lower pharyngeal tooth plate and urohyal bone of the two morphs of V. hartwegi. A) Typical morph, MZUNICACH 6761.B)Rheophilic morph, MZUNICACH 7157(photos by K.Ramírez-Moreno [pharyngeal tooth plates] and G. Morales-Flores [urohyal bones])." httpUri="https://zenodo.org/record/1160381/files/figure.png" pageId="10" pageNumber="381">Fig. 7A</figureCitation>
|
||
). Axial skeleton, based on 2 c&s specimens: vertebral counts, 15 precaudal and 14–15 caudal, total 29–30; ribs, 9 epipleurals and 13 pleurals; 2 supraneurals, between the supraoccipital crest and first dorsal pterygiophore; pterygiophores, 27 dorsal and 12–13 anal; the first anal pterygiophore contains 2 or 3 spines (
|
||
<figureCitation id="13522A18FFE8FFA5E4A440FAFAB2C4AC" box="[1198,1288,943,968]" captionStart="FIGURE 8" captionStartId="11.[151,250,1153,1175]" captionTargetBox="[162,1401,219,1104]" captionTargetId="figure@11.[151,1436,193,1132]" captionTargetPageId="11" captionText="FIGURE 8. Cleared and stained specimens of the two morphs of V. hartwegi. A) Typical morph, 89.3 mm SL, MZUNICACH 6761, La Angostura Reservoir, La Concordia, Chiapas. B) Rheophilic morph, 53.2 mm SL, MZUNICACH 7159, Tachinua River, Chicomuselo, Chiapas." httpUri="https://zenodo.org/record/1160383/files/figure.png" pageId="10" pageNumber="381">Fig. 8A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF166615FFE8FFA5E09D4421FDB9C0A3" httpUri="https://zenodo.org/record/1160381/files/figure.png" pageId="10" pageNumber="381" targetBox="[169,1418,1029,1870]" targetPageId="10">
|
||
<paragraph id="8BD6369DFFE8FFA5E09D4421FDB9C0A3" blockId="10.[151,1436,1908,1991]" pageId="10" pageNumber="381">
|
||
<emphasis id="B91DEA8FFFE8FFA5E09D4421FEA0C0EE" bold="true" box="[151,282,1908,1930]" pageId="10" pageNumber="381">FIGURE 7.</emphasis>
|
||
Lower pharyngeal tooth plate and urohyal bone of the two morphs of
|
||
<emphasis id="B91DEA8FFFE8FFA5E4494420FB7FC0EE" box="[1091,1221,1908,1930]" italics="true" pageId="10" pageNumber="381">
|
||
<taxonomicName id="4C694D1EFFE8FFA5E4494420FB7BC0EE" baseAuthorityName="Taylor & Miller" baseAuthorityYear="1980" box="[1091,1217,1908,1930]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="381" phylum="Chordata" rank="species" species="hartwegi">V. hartwegi</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
A) Typical morph, MZUNICACH 6761. B) Rheophilic morph, MZUNICACH 7157 (photos by K. Ramírez-Moreno [pharyngeal tooth plates] and G. Morales-Flores [urohyal bones]).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF166615FFE9FFA4E09D47D4FE7CC3B0" httpUri="https://zenodo.org/record/1160383/files/figure.png" pageId="11" pageNumber="382" targetBox="[162,1401,219,1104]" targetPageId="11">
|
||
<paragraph id="8BD6369DFFE9FFA4E09D47D4FE7CC3B0" blockId="11.[151,1435,1153,1236]" pageId="11" pageNumber="382">
|
||
<emphasis id="B91DEA8FFFE9FFA4E09D47D4FEA9C3F2" bold="true" box="[151,275,1153,1175]" pageId="11" pageNumber="382">FIGURE 8.</emphasis>
|
||
Cleared and stained specimens of the two morphs of
|
||
<taxonomicName id="4C694D1EFFE9FFA4E33E47D7FC1FC3F3" baseAuthorityName="Taylor & Miller" baseAuthorityYear="1980" box="[820,933,1153,1175]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="382" phylum="Chordata" rank="species" species="hartwegi">
|
||
<emphasis id="B91DEA8FFFE9FFA4E33E47D7FC1FC3F3" box="[820,933,1153,1175]" italics="true" pageId="11" pageNumber="382">V. hartwegi</emphasis>
|
||
</taxonomicName>
|
||
. A) Typical morph, 89.3 mm SL, MZUNICACH 6761, La Angostura Reservoir, La Concordia, Chiapas. B) Rheophilic morph, 53.2 mm SL, MZUNICACH 7159, Tachinua River, Chicomuselo, Chiapas.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD6369DFFE9FFA4E0CD4654FE81C124" blockId="11.[151,1437,1281,2032]" pageId="11" pageNumber="382">
|
||
Rheophilic morph smaller, largest known
|
||
<quantity id="4C919B78FFE9FFA4E2A34657FCAAC27D" box="[681,784,1282,1306]" metricMagnitude="-2" metricUnit="m" metricValue="9.709999999999999" pageId="11" pageNumber="382" unit="mm" value="97.1">97.1 mm</quantity>
|
||
SL, with following meristic data:
|
||
<emphasis id="B91DEA8FFFE9FFA4E4954657FB60C27D" box="[1183,1242,1282,1305]" italics="true" pageId="11" pageNumber="382">Fins:</emphasis>
|
||
D XVIII, 11–13 (mode 11), A V–VII (mode 6), 8–10 (mode 9); pectoral rays 15–16 (mode 16).
|
||
<emphasis id="B91DEA8FFFE9FFA4E4184670FBD9C259" box="[1042,1123,1317,1341]" italics="true" pageId="11" pageNumber="382">Scales:</emphasis>
|
||
Lateral series 31–34 (mode 31); upper lateral line 20–21 (mode 21); lower lateral line 9–13 (mode 12), in two cases, pored scales disrupted by non-pored scales and usually 1–3 (mode 2) subsidiary pored scales on caudal fin; isolated pored scales on caudalfin base, absent; predorsal squamation pattern irregular; scale row overlap 1–3 (mode 2); pectoral-pelvic scale rows 7–8 (mode 7); scale rows from lateral line to base of first dorsal-fin ray 3–3 1/2 (mode 3 1/2); scale rows from lateral line to base of first dorsal-fin spine 6–7 (mode 6); scale rows from origin of anal fin to lower lateral line 8–9 (mode 9); circumpeduncular scales 18–21 (mode 20); scale rows on cheek 6–7 (mode 7). Squamation pattern similar to typical morph.
|
||
</paragraph>
|
||
<paragraph id="8BD6369DFFE9FFA4E0CD451EFDF9C190" blockId="11.[151,1437,1281,2032]" pageId="11" pageNumber="382">
|
||
Body shape slender or moderately deep (
|
||
<figureCitation id="13522A18FFE9FFA4E293451EFD5DC107" box="[665,743,1611,1636]" captionStart="FIGURE 4" captionStartId="7.[151,250,1125,1147]" captionTargetBox="[151,1437,193,1104]" captionTargetId="figure@7.[151,1437,193,1104]" captionTargetPageId="7" captionText="FIGURE 4. Live specimens of the rheophilic morph of Vieja hartwegi. A) Adult female in normal coloration, 97 mm SL, MZUNICACH 7161, Lagos de Colón, La Trinitaria, Chiapas (photo by M. A. Peralta-Meixueiro). B) Adult female in stress coloration, 65 mm SL, uncataloged specimen, Lagos de Colón, La Trinitaria, Chiapas. C) Adult female in stress coloration, 91 mm SL, MZUNICACH 7159, Tachinua River, Chicomuselo, Chiapas. D) Adult male in stress coloration, 80 mm SL, MZUNICACH 7157, Yayahuita River, Chicomuselo, Chiapas." httpUri="https://zenodo.org/record/1160375/files/figure.png" pageId="11" pageNumber="382">Figs. 4</figureCitation>
|
||
&
|
||
<figureCitation id="13522A18FFE9FFA4E304451EFCA4C100" box="[782,798,1611,1636]" captionStart="FIGURE 6" captionStartId="9.[151,250,1685,1707]" captionTargetBox="[151,1436,235,1664]" captionTargetId="figure@9.[151,1436,193,1664]" captionTargetPageId="9" captionText="FIGURE 6. Preserved specimens of the rheophilic Vieja hartwegi morph. A) Adult female, MZUNICACH 7161, Lagos de Colón, La Trinitaria, Chiapas. B) Adult female, MZUNICACH 7159 [post-mortem fixed specimen], Yayahuita River, Chicomuselo, Chiapas." httpUri="https://zenodo.org/record/1160379/files/figure.png" pageId="11" pageNumber="382">6</figureCitation>
|
||
); predorsal contour slightly steep, dorsal head profile in front of orbit usually straight or slightly convex; prepelvic contour gently convex; caudal peduncle short, deeper than long. Mouth small, terminal and isognathous in all sizes. Snout moderately pronunced. Lips of both jaws moderately thickened. Frenum well developed. Maxillary cleft position, usually ventral or sometimes at level of dorsal margin of the pectoral-fin base.
|
||
</paragraph>
|
||
<paragraph id="8BD6369DFFE9FFA4E0CD45AAFB2DC0E0" blockId="11.[151,1437,1281,2032]" pageId="11" pageNumber="382">
|
||
<emphasis id="B91DEA8FFFE9FFA4E0CD45AAFE5FC073" box="[199,485,1791,1815]" italics="true" pageId="11" pageNumber="382">Gillrakers on first arch:</emphasis>
|
||
lower limb 8–10 (mode 9); upper limb 3–4 (mode 3); total 11–13 (mode 13). Gillrakers moderately elongated; tips pointed or rounded, commonly anterior gillrakers on lower limb shorter and rounded, and posterior larger and pointed. Outer teeth in both jaws recurved and longer than typical morph, with posterior little cusp, anteriorly increasing in size; 3–4 irregular series of inner smaller teeth.
|
||
</paragraph>
|
||
<paragraph id="8BD6369DFFE9FFA3E0CD44C5FC49C6EC" blockId="11.[151,1437,1281,2032]" lastBlockId="12.[151,1436,151,392]" lastPageId="12" lastPageNumber="383" pageId="11" pageNumber="382">
|
||
Lower pharyngeal tooth-plate (
|
||
<figureCitation id="13522A18FFE9FFA4E22744DAFD3FC0C3" box="[557,645,1935,1960]" captionStart="FIGURE 7" captionStartId="10.[151,250,1908,1930]" captionTargetBox="[169,1418,1029,1870]" captionTargetId="figure@10.[150,1436,1021,1887]" captionTargetPageId="10" captionText="FIGURE 7. Lower pharyngeal tooth plate and urohyal bone of the two morphs of V. hartwegi. A) Typical morph, MZUNICACH 6761.B)Rheophilic morph, MZUNICACH 7157(photos by K.Ramírez-Moreno [pharyngeal tooth plates] and G. Morales-Flores [urohyal bones])." httpUri="https://zenodo.org/record/1160381/files/figure.png" pageId="11" pageNumber="382">Fig. 7B</figureCitation>
|
||
) dissected from two specimens (MZUNICACH 7157,
|
||
<quantity id="4C919B78FFE9FFA4E50944DAFAD0C0C3" box="[1283,1386,1935,1960]" metricMagnitude="-2" metricUnit="m" metricValue="6.12" pageId="11" pageNumber="382" unit="mm" value="61.2">61.2 mm</quantity>
|
||
SL; 7159,
|
||
<quantity id="4C919B78FFE9FFA4E0E944E6FEF7C0AF" box="[227,333,1971,1996]" metricMagnitude="-2" metricUnit="m" metricValue="5.33" pageId="11" pageNumber="382" unit="mm" value="53.3">53.3 mm</quantity>
|
||
SL), dorsoventrally compressed, with short anterior process and posterior processes slightly enlarged; wider than large; posterior margin concave at center and very convex at margins; teeth conical, enlarged posteriorly and shorter anteriorly, with antrorse cusp; second cusp small but strong, present at the base; four or three medial rows, mainly the first, notably larger and more robust posteriorly than the rest; 22–24 (
|
||
<date id="FFD7105DFFEEFFA3E4E343E9FA26C7B0" box="[1257,1436,188,213]" pageId="12" pageNumber="383" value="2012-11-11" valueMax="2012-12-11" valueMin="2012-11-11">
|
||
11/11 and 12/
|
||
<quantity id="4C919B78FFEEFFA3E58B43E9FA26C7B0" box="[1409,1436,188,212]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="12" pageNumber="383" unit="in" value="12.0">12</quantity>
|
||
</date>
|
||
in each side) total teeth in posterior row; 8/7 and 6/7 teeth in near median rows. Urohyal bone with posterior edge concave, dorsally convex with a strong curved spine (
|
||
<figureCitation id="13522A18FFEEFFA3E2F44251FCE2C678" box="[766,856,260,285]" captionStart="FIGURE 7" captionStartId="10.[151,250,1908,1930]" captionTargetBox="[169,1418,1029,1870]" captionTargetId="figure@10.[150,1436,1021,1887]" captionTargetPageId="10" captionText="FIGURE 7. Lower pharyngeal tooth plate and urohyal bone of the two morphs of V. hartwegi. A) Typical morph, MZUNICACH 6761.B)Rheophilic morph, MZUNICACH 7157(photos by K.Ramírez-Moreno [pharyngeal tooth plates] and G. Morales-Flores [urohyal bones])." httpUri="https://zenodo.org/record/1160381/files/figure.png" pageId="12" pageNumber="383">Fig. 7B</figureCitation>
|
||
). Axial skeleton, based on two c&s specimens and one radiograph: vertebral counts, 15 precaudal and 15(1)–16(2) caudal, total 30(1)–31(2); ribs, 12 epipleurals and 13 pleurals; 2 supraneurals, between the supraoccipital crest and first dorsal pterygiophore; pterygiophores, 30 dorsals and 15 anals; the first anal pterygiophore contains 2 spines (
|
||
<figureCitation id="13522A18FFEEFFA3E384423AFC5FC6EC" box="[910,997,367,392]" captionStart="FIGURE 8" captionStartId="11.[151,250,1153,1175]" captionTargetBox="[162,1401,219,1104]" captionTargetId="figure@11.[151,1436,193,1132]" captionTargetPageId="11" captionText="FIGURE 8. Cleared and stained specimens of the two morphs of V. hartwegi. A) Typical morph, 89.3 mm SL, MZUNICACH 6761, La Angostura Reservoir, La Concordia, Chiapas. B) Rheophilic morph, 53.2 mm SL, MZUNICACH 7159, Tachinua River, Chicomuselo, Chiapas." httpUri="https://zenodo.org/record/1160383/files/figure.png" pageId="12" pageNumber="383">Fig. 8B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF166615FFEEFFA3E09D442AFA85C0D0" httpUri="https://zenodo.org/record/1160385/files/figure.png" pageId="12" pageNumber="383" targetBox="[164,1432,476,1890]" targetPageId="12">
|
||
<paragraph id="8BD6369DFFEEFFA3E09D442AFA85C0D0" blockId="12.[151,1436,1919,1972]" pageId="12" pageNumber="383">
|
||
<emphasis id="B91DEA8FFFEEFFA3E09D442AFEAFC0F0" bold="true" box="[151,277,1919,1941]" pageId="12" pageNumber="383">FIGURE 9.</emphasis>
|
||
Comparison to species closely related to
|
||
<taxonomicName id="4C694D1EFFEEFFA3E2DA44D4FCFCC0F1" baseAuthorityName="Taylor & Miller" baseAuthorityYear="1980" box="[720,838,1919,1942]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="383" phylum="Chordata" rank="species" species="hartwegi">
|
||
<emphasis id="B91DEA8FFFEEFFA3E2DA44D4FCFCC0F1" box="[720,838,1919,1942]" italics="true" pageId="12" pageNumber="383">V. hartwegi</emphasis>
|
||
</taxonomicName>
|
||
: A)
|
||
<emphasis id="B91DEA8FFFEEFFA3E37144D4FBBBC0F1" box="[891,1025,1919,1942]" italics="true" pageId="12" pageNumber="383">
|
||
<taxonomicName id="4C694D1EFFEEFFA3E37144D4FC44C0F1" baseAuthorityName="Steindachner" baseAuthorityYear="1864" box="[891,1022,1919,1942]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="383" phylum="Chordata" rank="species" species="bifasciata">V. bifasciata</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
140 mm SL, ECOSC 4677, Tzendales River, Ocosingo, Chiapas. B)
|
||
<emphasis id="B91DEA8FFFEEFFA3E1CC44CAFDFEC0D0" box="[454,580,1950,1972]" italics="true" pageId="12" pageNumber="383">
|
||
<taxonomicName id="4C694D1EFFEEFFA3E1CC44CAFDFAC0D0" baseAuthorityName="Werner & Stawikowski" baseAuthorityYear="1987" box="[454,576,1950,1972]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="383" phylum="Chordata" rank="species" species="breidohri">V. breidohri</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
105 mm SL, MZUNICACH 7065, Lagos de Colón, La Trinitaria, Chiapas.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD6369DFFEFFFA2E0CD43C2FB7CC559" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="384">
|
||
<emphasis id="B91DEA8FFFEFFFA2E0CD43C2FDC7C7D4" bold="true" box="[199,637,151,176]" pageId="13" pageNumber="384">Coloration in preserved specimens.</emphasis>
|
||
<taxonomicName id="4C694D1EFFEFFFA2E28143CDFC8EC7D4" baseAuthorityName="Taylor & Miller" baseAuthorityYear="1980" box="[651,820,152,176]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="384" phylum="Chordata" rank="species" species="hartwegi">
|
||
<emphasis id="B91DEA8FFFEFFFA2E28143CDFC8EC7D4" box="[651,820,152,176]" italics="true" pageId="13" pageNumber="384">Vieja hartwegi</emphasis>
|
||
</taxonomicName>
|
||
has grayish background color, turning to brownish depending on time since preservation. Head usually somewhat darker than body. Ventral portion uniformly lighter. Typical morph with distinctive midlateral stripe extending from caudal-fin base to opercle, running at level (
|
||
<figureCitation id="13522A18FFEFFFA2E565438AFF01C679" captionStart="FIGURE5" captionStartId="8.[151,250,1823,1845]" captionTargetBox="[220,1316,219,1782]" captionTargetId="figure@8.[203,1338,193,1801]" captionTargetPageId="8" captionText="FIGURE5. Fixed specimensof the typical Viejahartwegi morph,showing the variation inbody shapeand the ontogenetic and individual variation in coloration patterns. A) Holotype (adult in breeding coloration), UMMZ 207701. B) Chiquito River, Chiapa de Corzo (juv.), MZUNICACH 7053. C) El Zapote River, Chiapa de Corzo (juv.), MZUNICACH 7210. D) La Angostura Reservoir, La Concordia (juv.), MZUNICACH 7154. E) Totopac River, Tecpatán (subadult), MZUNICACH 1053. F) Lagos de Colón, La Trinitaria (subadult), MZUNICACH 7065.G) CopanóRiver, Ostuacán (adult), MZUNICACH 7208.H- I) Grijalva River (main cannel), Copainalá (adults), MZUNICACH 3145. J) Santo Domingo River, Chiapa de Corzo (adult), MZUNICACH 6245." httpUri="https://zenodo.org/record/1160377/files/figure.png" pageId="13" pageNumber="384">Fig. 5A</figureCitation>
|
||
, C-D) or usually below mid-body (
|
||
<figureCitation id="13522A18FFEFFFA2E2624251FD7FC679" box="[616,709,260,285]" captionStart="FIGURE5" captionStartId="8.[151,250,1823,1845]" captionTargetBox="[220,1316,219,1782]" captionTargetId="figure@8.[203,1338,193,1801]" captionTargetPageId="8" captionText="FIGURE5. Fixed specimensof the typical Viejahartwegi morph,showing the variation inbody shapeand the ontogenetic and individual variation in coloration patterns. A) Holotype (adult in breeding coloration), UMMZ 207701. B) Chiquito River, Chiapa de Corzo (juv.), MZUNICACH 7053. C) El Zapote River, Chiapa de Corzo (juv.), MZUNICACH 7210. D) La Angostura Reservoir, La Concordia (juv.), MZUNICACH 7154. E) Totopac River, Tecpatán (subadult), MZUNICACH 1053. F) Lagos de Colón, La Trinitaria (subadult), MZUNICACH 7065.G) CopanóRiver, Ostuacán (adult), MZUNICACH 7208.H- I) Grijalva River (main cannel), Copainalá (adults), MZUNICACH 3145. J) Santo Domingo River, Chiapa de Corzo (adult), MZUNICACH 6245." httpUri="https://zenodo.org/record/1160377/files/figure.png" pageId="13" pageNumber="384">Fig. 5B</figureCitation>
|
||
, E-J); a well-developed dorsolateral or second stripe slightly separated (
|
||
<figureCitation id="13522A18FFEFFFA2E11C4272FE0FC624" box="[278,437,295,320]" captionStart="FIGURE5" captionStartId="8.[151,250,1823,1845]" captionTargetBox="[220,1316,219,1782]" captionTargetId="figure@8.[203,1338,193,1801]" captionTargetPageId="8" captionText="FIGURE5. Fixed specimensof the typical Viejahartwegi morph,showing the variation inbody shapeand the ontogenetic and individual variation in coloration patterns. A) Holotype (adult in breeding coloration), UMMZ 207701. B) Chiquito River, Chiapa de Corzo (juv.), MZUNICACH 7053. C) El Zapote River, Chiapa de Corzo (juv.), MZUNICACH 7210. D) La Angostura Reservoir, La Concordia (juv.), MZUNICACH 7154. E) Totopac River, Tecpatán (subadult), MZUNICACH 1053. F) Lagos de Colón, La Trinitaria (subadult), MZUNICACH 7065.G) CopanóRiver, Ostuacán (adult), MZUNICACH 7208.H- I) Grijalva River (main cannel), Copainalá (adults), MZUNICACH 3145. J) Santo Domingo River, Chiapa de Corzo (adult), MZUNICACH 6245." httpUri="https://zenodo.org/record/1160377/files/figure.png" pageId="13" pageNumber="384">Fig. 5E, G, J</figureCitation>
|
||
), partially fusioned anteriorly (
|
||
<figureCitation id="13522A18FFEFFFA2E3204272FC32C624" box="[810,904,295,320]" captionStart="FIGURE5" captionStartId="8.[151,250,1823,1845]" captionTargetBox="[220,1316,219,1782]" captionTargetId="figure@8.[203,1338,193,1801]" captionTargetPageId="8" captionText="FIGURE5. Fixed specimensof the typical Viejahartwegi morph,showing the variation inbody shapeand the ontogenetic and individual variation in coloration patterns. A) Holotype (adult in breeding coloration), UMMZ 207701. B) Chiquito River, Chiapa de Corzo (juv.), MZUNICACH 7053. C) El Zapote River, Chiapa de Corzo (juv.), MZUNICACH 7210. D) La Angostura Reservoir, La Concordia (juv.), MZUNICACH 7154. E) Totopac River, Tecpatán (subadult), MZUNICACH 1053. F) Lagos de Colón, La Trinitaria (subadult), MZUNICACH 7065.G) CopanóRiver, Ostuacán (adult), MZUNICACH 7208.H- I) Grijalva River (main cannel), Copainalá (adults), MZUNICACH 3145. J) Santo Domingo River, Chiapa de Corzo (adult), MZUNICACH 6245." httpUri="https://zenodo.org/record/1160377/files/figure.png" pageId="13" pageNumber="384">Fig. 5H</figureCitation>
|
||
) or sometimes fully fused (
|
||
<figureCitation id="13522A18FFEFFFA2E4D04272FAEFC624" box="[1242,1365,295,320]" captionStart="FIGURE5" captionStartId="8.[151,250,1823,1845]" captionTargetBox="[220,1316,219,1782]" captionTargetId="figure@8.[203,1338,193,1801]" captionTargetPageId="8" captionText="FIGURE5. Fixed specimensof the typical Viejahartwegi morph,showing the variation inbody shapeand the ontogenetic and individual variation in coloration patterns. A) Holotype (adult in breeding coloration), UMMZ 207701. B) Chiquito River, Chiapa de Corzo (juv.), MZUNICACH 7053. C) El Zapote River, Chiapa de Corzo (juv.), MZUNICACH 7210. D) La Angostura Reservoir, La Concordia (juv.), MZUNICACH 7154. E) Totopac River, Tecpatán (subadult), MZUNICACH 1053. F) Lagos de Colón, La Trinitaria (subadult), MZUNICACH 7065.G) CopanóRiver, Ostuacán (adult), MZUNICACH 7208.H- I) Grijalva River (main cannel), Copainalá (adults), MZUNICACH 3145. J) Santo Domingo River, Chiapa de Corzo (adult), MZUNICACH 6245." httpUri="https://zenodo.org/record/1160377/files/figure.png" pageId="13" pageNumber="384">Fig. 5D, I</figureCitation>
|
||
) with midlateral stripe in adults. In adults, second stripe formed by the fusion of blotches. In juveniles and subadults, midlateral stripe usually disrupted; commonly three to five irregular blotches above midlateral stripe (
|
||
<figureCitation id="13522A18FFEFFFA2E50F423AFAFCC6EC" box="[1285,1350,367,392]" captionStart="FIGURE5" captionStartId="8.[151,250,1823,1845]" captionTargetBox="[220,1316,219,1782]" captionTargetId="figure@8.[203,1338,193,1801]" captionTargetPageId="8" captionText="FIGURE5. Fixed specimensof the typical Viejahartwegi morph,showing the variation inbody shapeand the ontogenetic and individual variation in coloration patterns. A) Holotype (adult in breeding coloration), UMMZ 207701. B) Chiquito River, Chiapa de Corzo (juv.), MZUNICACH 7053. C) El Zapote River, Chiapa de Corzo (juv.), MZUNICACH 7210. D) La Angostura Reservoir, La Concordia (juv.), MZUNICACH 7154. E) Totopac River, Tecpatán (subadult), MZUNICACH 1053. F) Lagos de Colón, La Trinitaria (subadult), MZUNICACH 7065.G) CopanóRiver, Ostuacán (adult), MZUNICACH 7208.H- I) Grijalva River (main cannel), Copainalá (adults), MZUNICACH 3145. J) Santo Domingo River, Chiapa de Corzo (adult), MZUNICACH 6245." httpUri="https://zenodo.org/record/1160377/files/figure.png" pageId="13" pageNumber="384">Fig. 5</figureCitation>
|
||
B-C, F) or rarely absent in some specimens; in small juveniles, cross-bars arranged along midlateral stripe. Adults in breeding coloration presents five cross-bars along midlateral stripe, lower jaw and belly darker (as the
|
||
<typeStatus id="54D2883FFFEFFFA2E53942E2FA22C6B4" box="[1331,1432,439,464]" pageId="13" pageNumber="384" type="holotype">holotype</typeStatus>
|
||
,
|
||
<figureCitation id="13522A18FFEFFFA2E09D4289FF48C691" box="[151,242,476,501]" captionStart="FIGURE5" captionStartId="8.[151,250,1823,1845]" captionTargetBox="[220,1316,219,1782]" captionTargetId="figure@8.[203,1338,193,1801]" captionTargetPageId="8" captionText="FIGURE5. Fixed specimensof the typical Viejahartwegi morph,showing the variation inbody shapeand the ontogenetic and individual variation in coloration patterns. A) Holotype (adult in breeding coloration), UMMZ 207701. B) Chiquito River, Chiapa de Corzo (juv.), MZUNICACH 7053. C) El Zapote River, Chiapa de Corzo (juv.), MZUNICACH 7210. D) La Angostura Reservoir, La Concordia (juv.), MZUNICACH 7154. E) Totopac River, Tecpatán (subadult), MZUNICACH 1053. F) Lagos de Colón, La Trinitaria (subadult), MZUNICACH 7065.G) CopanóRiver, Ostuacán (adult), MZUNICACH 7208.H- I) Grijalva River (main cannel), Copainalá (adults), MZUNICACH 3145. J) Santo Domingo River, Chiapa de Corzo (adult), MZUNICACH 6245." httpUri="https://zenodo.org/record/1160377/files/figure.png" pageId="13" pageNumber="384">Fig. 5A</figureCitation>
|
||
). Interorbital bars commonly not evident in fixed specimens. Black crescent-shaped operculum marking present in adults, in juveniles and preadults usually weak or absent. In some specimens, brown spots present on cheeks. Soft portions of dorsal and anal fins and caudal fin, dark brown spotted in both morphs.
|
||
</paragraph>
|
||
<paragraph id="8BD6369DFFEFFFA2E0CD4112FDE9C471" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="384">
|
||
Rheophilic morph with distinctive dorsolateral stripe present above of the middle of the body, extending from margin of the opercle to the level of dorsal fin posterior end (
|
||
<figureCitation id="13522A18FFEFFFA2E3364139FC2FC5E1" box="[828,917,620,645]" captionStart="FIGURE 6" captionStartId="9.[151,250,1685,1707]" captionTargetBox="[151,1436,235,1664]" captionTargetId="figure@9.[151,1436,193,1664]" captionTargetPageId="9" captionText="FIGURE 6. Preserved specimens of the rheophilic Vieja hartwegi morph. A) Adult female, MZUNICACH 7161, Lagos de Colón, La Trinitaria, Chiapas. B) Adult female, MZUNICACH 7159 [post-mortem fixed specimen], Yayahuita River, Chicomuselo, Chiapas." httpUri="https://zenodo.org/record/1160379/files/figure.png" pageId="13" pageNumber="384">Fig. 6A</figureCitation>
|
||
); this stripe is similar to dorsolateral or second stripe in typical morph, formed by the fusion of lateral blotches. Great rounded spot at center of caudal peduncle and base of caudal fin, often disrupted with dorsolateral stripe or sometimes confluent, but always remarkable (
|
||
<figureCitation id="13522A18FFEFFFA2E56541E1FF1DC594" captionStart="FIGURE 6" captionStartId="9.[151,250,1685,1707]" captionTargetBox="[151,1436,235,1664]" captionTargetId="figure@9.[151,1436,193,1664]" captionTargetPageId="9" captionText="FIGURE 6. Preserved specimens of the rheophilic Vieja hartwegi morph. A) Adult female, MZUNICACH 7161, Lagos de Colón, La Trinitaria, Chiapas. B) Adult female, MZUNICACH 7159 [post-mortem fixed specimen], Yayahuita River, Chicomuselo, Chiapas." httpUri="https://zenodo.org/record/1160379/files/figure.png" pageId="13" pageNumber="384">Fig. 6</figureCitation>
|
||
). Eight diffuse bars along the body, usually six bars notable, especially in juveniles (
|
||
<figureCitation id="13522A18FFEFFFA2E4604182FB7EC594" box="[1130,1220,727,752]" captionStart="FIGURE 6" captionStartId="9.[151,250,1685,1707]" captionTargetBox="[151,1436,235,1664]" captionTargetId="figure@9.[151,1436,193,1664]" captionTargetPageId="9" captionText="FIGURE 6. Preserved specimens of the rheophilic Vieja hartwegi morph. A) Adult female, MZUNICACH 7161, Lagos de Colón, La Trinitaria, Chiapas. B) Adult female, MZUNICACH 7159 [post-mortem fixed specimen], Yayahuita River, Chicomuselo, Chiapas." httpUri="https://zenodo.org/record/1160379/files/figure.png" pageId="13" pageNumber="384">Fig. 6B</figureCitation>
|
||
). Usually, series of 4–6 dark dots below dorsolateral stripe.
|
||
</paragraph>
|
||
<paragraph id="8BD6369DFFEFFFA2E0CD404AFBAEC3A0" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="384">
|
||
<emphasis id="B91DEA8FFFEFFFA2E0CD404AFE1BC45C" bold="true" box="[199,417,799,824]" pageId="13" pageNumber="384">Coloration in life.</emphasis>
|
||
<taxonomicName id="4C694D1EFFEFFFA2E1A64075FDE9C45C" baseAuthorityName="Taylor & Miller" baseAuthorityYear="1980" box="[428,595,800,824]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="384" phylum="Chordata" rank="species" species="hartwegi">
|
||
<emphasis id="B91DEA8FFFEFFFA2E1A64075FDE9C45C" box="[428,595,800,824]" italics="true" pageId="13" pageNumber="384">Vieja hartwegi</emphasis>
|
||
</taxonomicName>
|
||
exhibits a highly variable coloration throughout its distributional range. Populations in upper Grijalva show background color that varies from silver-gray to olive-gray; in middle and lower Grijalva it varies from olive-gray to yellowish (see Conkel 1993: 58). Head dorsally olive-gray to yellowish; in juveniles and sub-adults red spots present on preoperculum, operculum, cheek, and lachrymal (
|
||
<figureCitation id="13522A18FFEFFFA2E51A40D9FAD4C4C0" box="[1296,1390,908,933]" captionStart="FIGURE 3" captionStartId="6.[151,250,1858,1880]" captionTargetBox="[192,1411,225,1818]" captionTargetId="figure@6.[151,1436,193,1837]" captionTargetPageId="6" captionText="FIGURE 3. Live specimens of the typical morph of Vieja hartwegi, showing ontogenic variation in coloration pattern. A) Juvenile, 75 mm SL, MZUNICACH 6761, La Angostura Reservoir, La Concordia, Chiapas. B) Adult, 144 mm SL, aquarium specimen, Cacahuanó River, Ocozocoautla, Chiapas. C) Large adult, 250 mm SL, uncataloged specimen from a fishery, Grijalva River (main channel at Tres Picos), Copainalá, Chiapas (photo by J. M. López-Vila)." httpUri="https://zenodo.org/record/1160372/files/figure.png" pageId="13" pageNumber="384">Fig. 3A</figureCitation>
|
||
); in adults red blotches cover entire head (
|
||
<figureCitation id="13522A18FFEFFFA2E24740FAFD2AC4AC" box="[589,656,943,968]" captionStart="FIGURE 3" captionStartId="6.[151,250,1858,1880]" captionTargetBox="[192,1411,225,1818]" captionTargetId="figure@6.[151,1436,193,1837]" captionTargetPageId="6" captionText="FIGURE 3. Live specimens of the typical morph of Vieja hartwegi, showing ontogenic variation in coloration pattern. A) Juvenile, 75 mm SL, MZUNICACH 6761, La Angostura Reservoir, La Concordia, Chiapas. B) Adult, 144 mm SL, aquarium specimen, Cacahuanó River, Ocozocoautla, Chiapas. C) Large adult, 250 mm SL, uncataloged specimen from a fishery, Grijalva River (main channel at Tres Picos), Copainalá, Chiapas (photo by J. M. López-Vila)." httpUri="https://zenodo.org/record/1160372/files/figure.png" pageId="13" pageNumber="384">Fig. 3</figureCitation>
|
||
B-C). Side of head bluish and iridescent base color, more vibrant in sub-adults and adults. Two dark interorbital bars present. In large adults red dots at the center of the scales cover all of the body continuing onto dorsal, anal, and caudal fins (
|
||
<figureCitation id="13522A18FFEFFFA2E33E40A2FC35C374" box="[820,911,1015,1040]" captionStart="FIGURE 3" captionStartId="6.[151,250,1858,1880]" captionTargetBox="[192,1411,225,1818]" captionTargetId="figure@6.[151,1436,193,1837]" captionTargetPageId="6" captionText="FIGURE 3. Live specimens of the typical morph of Vieja hartwegi, showing ontogenic variation in coloration pattern. A) Juvenile, 75 mm SL, MZUNICACH 6761, La Angostura Reservoir, La Concordia, Chiapas. B) Adult, 144 mm SL, aquarium specimen, Cacahuanó River, Ocozocoautla, Chiapas. C) Large adult, 250 mm SL, uncataloged specimen from a fishery, Grijalva River (main channel at Tres Picos), Copainalá, Chiapas (photo by J. M. López-Vila)." httpUri="https://zenodo.org/record/1160372/files/figure.png" pageId="13" pageNumber="384">Fig. 3C</figureCitation>
|
||
). Belly usually red in adults. In juveniles and subadults, dots along the body brownish red. Midlateral and dorsolateral (or blotches) stripes black; sometimes the two bars fully fused. Pectoral fins transparent or slightly yellowish; pelvic fins grayish. Dorsal and anal fins gray translucent in juveniles and sub-adults and yellowish in adults. In very large adults, spiny portion of the dorsal fin, golden. Border of dorsal fin red and anal fin dark. Caudal fin gray and translucent, with red border in large adults (
|
||
<figureCitation id="13522A18FFEFFFA2E09547F9FF4DC3A0" box="[159,247,1196,1221]" captionStart="FIGURE 3" captionStartId="6.[151,250,1858,1880]" captionTargetBox="[192,1411,225,1818]" captionTargetId="figure@6.[151,1436,193,1837]" captionTargetPageId="6" captionText="FIGURE 3. Live specimens of the typical morph of Vieja hartwegi, showing ontogenic variation in coloration pattern. A) Juvenile, 75 mm SL, MZUNICACH 6761, La Angostura Reservoir, La Concordia, Chiapas. B) Adult, 144 mm SL, aquarium specimen, Cacahuanó River, Ocozocoautla, Chiapas. C) Large adult, 250 mm SL, uncataloged specimen from a fishery, Grijalva River (main channel at Tres Picos), Copainalá, Chiapas (photo by J. M. López-Vila)." httpUri="https://zenodo.org/record/1160372/files/figure.png" pageId="13" pageNumber="384">Fig. 3C</figureCitation>
|
||
). Breeding coloration is shown in
|
||
<bibRefCitation id="EFF84B6CFFEFFFA2E27247F8FBAAC3A0" author="Werner" box="[632,1040,1196,1221]" pageId="13" pageNumber="384" refString="Werner, U. & Stawikowski, R. (1987) Ein neuer Buntbarsch aus Sudmexiko: Paratheraps breidohri, gen. nov., spec. nov. Die Aquarien-und Terrarien-Zeitschrift, 41, 20 - 23." type="journal article" year="1987">Werner and Stawikowski (1987: 23)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD6369DFFEFFFA2E0CD479AFC24C1D9" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="384">
|
||
Rheophilic morph has olive gray base color. Head olive gray dorsally with red or brownish orange spots or blotches. Two dark interorbital bars evident. Side of head bluish and iridescent, covered by variable brownish orange or reddish spots or blotches (usually forming a reticular pattern) all over opercle, preopercle, cheek, and lachrymal (
|
||
<figureCitation id="13522A18FFEFFFA2E11F4669FEE0C230" box="[277,346,1340,1365]" captionStart="FIGURE 4" captionStartId="7.[151,250,1125,1147]" captionTargetBox="[151,1437,193,1104]" captionTargetId="figure@7.[151,1437,193,1104]" captionTargetPageId="7" captionText="FIGURE 4. Live specimens of the rheophilic morph of Vieja hartwegi. A) Adult female in normal coloration, 97 mm SL, MZUNICACH 7161, Lagos de Colón, La Trinitaria, Chiapas (photo by M. A. Peralta-Meixueiro). B) Adult female in stress coloration, 65 mm SL, uncataloged specimen, Lagos de Colón, La Trinitaria, Chiapas. C) Adult female in stress coloration, 91 mm SL, MZUNICACH 7159, Tachinua River, Chicomuselo, Chiapas. D) Adult male in stress coloration, 80 mm SL, MZUNICACH 7157, Yayahuita River, Chicomuselo, Chiapas." httpUri="https://zenodo.org/record/1160375/files/figure.png" pageId="13" pageNumber="384">Fig. 4</figureCitation>
|
||
). In some specimens, bluish iridescent horizontal stripe prominent below eye (
|
||
<figureCitation id="13522A18FFEFFFA2E4CE4669FAA6C230" box="[1220,1308,1340,1365]" captionStart="FIGURE 4" captionStartId="7.[151,250,1125,1147]" captionTargetBox="[151,1437,193,1104]" captionTargetId="figure@7.[151,1437,193,1104]" captionTargetPageId="7" captionText="FIGURE 4. Live specimens of the rheophilic morph of Vieja hartwegi. A) Adult female in normal coloration, 97 mm SL, MZUNICACH 7161, Lagos de Colón, La Trinitaria, Chiapas (photo by M. A. Peralta-Meixueiro). B) Adult female in stress coloration, 65 mm SL, uncataloged specimen, Lagos de Colón, La Trinitaria, Chiapas. C) Adult female in stress coloration, 91 mm SL, MZUNICACH 7159, Tachinua River, Chicomuselo, Chiapas. D) Adult male in stress coloration, 80 mm SL, MZUNICACH 7157, Yayahuita River, Chicomuselo, Chiapas." httpUri="https://zenodo.org/record/1160375/files/figure.png" pageId="13" pageNumber="384">Fig. 4C</figureCitation>
|
||
). In normal coloration, two dark marks notable: a great rounded spot at center of the caudal peduncle and base of caudal fin (
|
||
<figureCitation id="13522A18FFEFFFA2E09546D1FF5CC2F8" box="[159,230,1412,1437]" captionStart="FIGURE 4" captionStartId="7.[151,250,1125,1147]" captionTargetBox="[151,1437,193,1104]" captionTargetId="figure@7.[151,1437,193,1104]" captionTargetPageId="7" captionText="FIGURE 4. Live specimens of the rheophilic morph of Vieja hartwegi. A) Adult female in normal coloration, 97 mm SL, MZUNICACH 7161, Lagos de Colón, La Trinitaria, Chiapas (photo by M. A. Peralta-Meixueiro). B) Adult female in stress coloration, 65 mm SL, uncataloged specimen, Lagos de Colón, La Trinitaria, Chiapas. C) Adult female in stress coloration, 91 mm SL, MZUNICACH 7159, Tachinua River, Chicomuselo, Chiapas. D) Adult male in stress coloration, 80 mm SL, MZUNICACH 7157, Yayahuita River, Chicomuselo, Chiapas." httpUri="https://zenodo.org/record/1160375/files/figure.png" pageId="13" pageNumber="384">Fig. 4</figureCitation>
|
||
A-C; some specimens present an ocellus, with iridescent light blue around the spot, probably a sexual dimorfism in males
|
||
<figureCitation id="13522A18FFEFFFA2E17446F2FE6DC2A4" box="[382,471,1447,1472]" captionStart="FIGURE 4" captionStartId="7.[151,250,1125,1147]" captionTargetBox="[151,1437,193,1104]" captionTargetId="figure@7.[151,1437,193,1104]" captionTargetPageId="7" captionText="FIGURE 4. Live specimens of the rheophilic morph of Vieja hartwegi. A) Adult female in normal coloration, 97 mm SL, MZUNICACH 7161, Lagos de Colón, La Trinitaria, Chiapas (photo by M. A. Peralta-Meixueiro). B) Adult female in stress coloration, 65 mm SL, uncataloged specimen, Lagos de Colón, La Trinitaria, Chiapas. C) Adult female in stress coloration, 91 mm SL, MZUNICACH 7159, Tachinua River, Chicomuselo, Chiapas. D) Adult male in stress coloration, 80 mm SL, MZUNICACH 7157, Yayahuita River, Chicomuselo, Chiapas." httpUri="https://zenodo.org/record/1160375/files/figure.png" pageId="13" pageNumber="384">Fig. 4D</figureCitation>
|
||
) and irregular blotch on the middle of the body. Under stress, coloration consists of dark vertical bars along the body and series of seven blotches along the bars, forming a general dorsolateral stripe above the middle of the body with no fusion of caudal spot or ocellus (
|
||
<figureCitation id="13522A18FFEFFFA2E3BF46BAFC43C16C" box="[949,1017,1519,1544]" captionStart="FIGURE 4" captionStartId="7.[151,250,1125,1147]" captionTargetBox="[151,1437,193,1104]" captionTargetId="figure@7.[151,1437,193,1104]" captionTargetPageId="7" captionText="FIGURE 4. Live specimens of the rheophilic morph of Vieja hartwegi. A) Adult female in normal coloration, 97 mm SL, MZUNICACH 7161, Lagos de Colón, La Trinitaria, Chiapas (photo by M. A. Peralta-Meixueiro). B) Adult female in stress coloration, 65 mm SL, uncataloged specimen, Lagos de Colón, La Trinitaria, Chiapas. C) Adult female in stress coloration, 91 mm SL, MZUNICACH 7159, Tachinua River, Chicomuselo, Chiapas. D) Adult male in stress coloration, 80 mm SL, MZUNICACH 7157, Yayahuita River, Chicomuselo, Chiapas." httpUri="https://zenodo.org/record/1160375/files/figure.png" pageId="13" pageNumber="384">Fig. 4</figureCitation>
|
||
B-D). Distal portions of lateral scales reddish or brownish orange, whereas proximal portions iridescent light blue, forming regular series of longitudinal stripes along each scale row. Pectoral fins transparent or slightly yellowish; pelvic fins pale or dark grayish. Dorsal and anal fins brownish or yellowish and translucent with bluish iridescent dots. Border of dorsal fin red with bluish line below the red and across the spiny portion of the fin. Border of anal fin dark; caudal fin gray and translucent with regular series of brownish and sometimes bluish iridescent dots.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3736516FFEFFFA2E0CD4592FC51C0B8" pageId="13" pageNumber="384" type="distribution">
|
||
<paragraph id="8BD6369DFFEFFFA2E0CD4592FC51C0B8" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="384">
|
||
<emphasis id="B91DEA8FFFEFFFA2E0CD4592FE4CC184" bold="true" box="[199,502,1735,1760]" pageId="13" pageNumber="384">Distribution and habitat.</emphasis>
|
||
Found on the Atlantic slope of
|
||
<collectingRegion id="49ADF87FFFEFFFA2E3664592FC73C184" box="[876,969,1735,1760]" country="Mexico" name="Chiapas" pageId="13" pageNumber="384">Chiapas</collectingRegion>
|
||
,
|
||
<collectingCountry id="F37E760DFFEFFFA2E3DD4592FB96C184" box="[983,1068,1735,1760]" name="Mexico" pageId="13" pageNumber="384">
|
||
<collectingRegion id="49ADF87FFFEFFFA2E3DD4592FB96C184" box="[983,1068,1735,1760]" country="Mexico" name="Mexico" pageId="13" pageNumber="384">Mexico</collectingRegion>
|
||
</collectingCountry>
|
||
and
|
||
<collectingCountry id="F37E760DFFEFFFA2E4624592FB5EC184" box="[1128,1252,1735,1760]" name="Guatemala" pageId="13" pageNumber="384">
|
||
<collectingRegion id="49ADF87FFFEFFFA2E4624592FB5EC184" box="[1128,1252,1735,1760]" country="Guatemala" name="Guatemala" pageId="13" pageNumber="384">Guatemala</collectingRegion>
|
||
</collectingCountry>
|
||
, endemic to the Grijalva River basin.
|
||
<bibRefCitation id="EFF84B6CFFEFFFA2E18745B9FDD3C061" box="[397,617,1772,1797]" pageId="13" pageNumber="384" refString="Miller, R. R., Minckley, W. L. & Norris, S. M. (2005) Freshwater Fishes of Mexico. University of Chicago Press, Chicago, Illinois, 652 pp." type="book">
|
||
Miller
|
||
<emphasis id="B91DEA8FFFEFFFA2E1D645B8FDACC060" box="[476,534,1772,1796]" italics="true" pageId="13" pageNumber="384">et al.</emphasis>
|
||
(2005)
|
||
</bibRefCitation>
|
||
considered its distribution only in the middle and upper portions of the Grijalva.
|
||
<bibRefCitation id="EFF84B6CFFEFFFA2E108445AFDE1C04C" box="[258,603,1807,1832]" pageId="13" pageNumber="384" refString="Gomez-Gonzalez, A. E., Velazquez-Velazquez, E., Anzueto-Calvo, M. J. & Maza-Cruz, M. F. (2015) Fishes of the Grijalva River basin of Mexico and Guatemala. Check List, 11, 1726. https: // doi. org / 10.15560 / 11.1726" type="journal article">
|
||
Gómez-González
|
||
<emphasis id="B91DEA8FFFEFFFA2E1C44444FDB2C04C" box="[462,520,1808,1832]" italics="true" pageId="13" pageNumber="384">et al.</emphasis>
|
||
(2015)
|
||
</bibRefCitation>
|
||
documented its presence in the lower Grijalva in Ángel Albino Corzo or Peñitas reservoir and tributaries. It also inhabits the upper reaches of the lower Grijalva outside of reservoirs (
|
||
<figureCitation id="13522A18FFEFFFA2E5654461FF0CC014" captionStart="FIGURE 10" captionStartId="14.[151,250,1455,1477]" captionTargetBox="[155,1433,200,1427]" captionTargetId="figure@14.[151,1436,193,1433]" captionTargetPageId="14" captionText="FIGURE 10. Map showing distribution of the two morphs of V. hartwegi in the Grijalva River basin, Mexico and Guatemala. The white square represents the type locality, black circles are localities of the typical morph and white circles are localities of the rheophilic morph." httpUri="https://zenodo.org/record/1160387/files/figure.png" pageId="13" pageNumber="384">Fig. 10</figureCitation>
|
||
). Elevations typically range from
|
||
<quantity id="4C919B78FFEFFFA2E24F440DFD52C014" box="[581,744,1880,1904]" metricMagnitude="2" metricUnit="m" metricValue="4.0" metricValueMax="7.0" metricValueMin="1.0" pageId="13" pageNumber="384" unit="m" value="400.0" valueMax="700.0" valueMin="100.0">100 to 700 m</quantity>
|
||
; however, an exceptional record exists from
|
||
<quantity id="4C919B78FFEFFFA2E4FC4402FAF5C014" box="[1270,1359,1879,1904]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="13" pageNumber="384" unit="m" value="1600.0">1600 m</quantity>
|
||
in the isolated Laguna Verde, municipality of Coapilla,
|
||
<collectingRegion id="49ADF87FFFEFFFA2E2C74429FC9DC0F1" box="[717,807,1916,1941]" country="Mexico" name="Chiapas" pageId="13" pageNumber="384">Chiapas</collectingRegion>
|
||
(MZUNICACH 1239, 1242). The rheophilic morph is restricted to tributaries of the far upper Grijalva River basin, only known in Lagartero River, near the Guatemalan border, and the Tachinua and Yayahuita rivers in
|
||
<collectingRegion id="49ADF87FFFEFFFA2E2B74491FCA0C0B9" box="[701,794,1988,2013]" country="Mexico" name="Chiapas" pageId="13" pageNumber="384">Chiapas</collectingRegion>
|
||
,
|
||
<collectingCountry id="F37E760DFFEFFFA2E32F4491FCC0C0B9" box="[805,890,1988,2013]" name="Mexico" pageId="13" pageNumber="384">
|
||
<collectingRegion id="49ADF87FFFEFFFA2E32F4491FCC0C0B9" box="[805,890,1988,2013]" country="Mexico" name="Mexico" pageId="13" pageNumber="384">Mexico</collectingRegion>
|
||
</collectingCountry>
|
||
(
|
||
<figureCitation id="13522A18FFEFFFA2E3804491FC67C0B8" box="[906,989,1988,2013]" captionStart="FIGURE 10" captionStartId="14.[151,250,1455,1477]" captionTargetBox="[155,1433,200,1427]" captionTargetId="figure@14.[151,1436,193,1433]" captionTargetPageId="14" captionText="FIGURE 10. Map showing distribution of the two morphs of V. hartwegi in the Grijalva River basin, Mexico and Guatemala. The white square represents the type locality, black circles are localities of the typical morph and white circles are localities of the rheophilic morph." httpUri="https://zenodo.org/record/1160387/files/figure.png" pageId="13" pageNumber="384">Fig. 10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF166615FFECFFA1E09D46FAFEC9C166" httpUri="https://zenodo.org/record/1160387/files/figure.png" pageId="14" pageNumber="385" targetBox="[155,1433,200,1427]" targetPageId="14">
|
||
<paragraph id="8BD6369DFFECFFA1E09D46FAFEC9C166" blockId="14.[151,1437,1455,1538]" pageId="14" pageNumber="385">
|
||
<emphasis id="B91DEA8FFFECFFA1E09D46FAFE9AC2A0" bold="true" box="[151,288,1455,1477]" pageId="14" pageNumber="385">FIGURE 10.</emphasis>
|
||
Map showing distribution of the two morphs of
|
||
<taxonomicName id="4C694D1EFFECFFA1E31846E5FC3FC2A1" baseAuthorityName="Taylor & Miller" baseAuthorityYear="1980" box="[786,901,1455,1477]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="385" phylum="Chordata" rank="species" species="hartwegi">
|
||
<emphasis id="B91DEA8FFFECFFA1E31846E5FC3FC2A1" box="[786,901,1455,1477]" italics="true" pageId="14" pageNumber="385">V. hartwegi</emphasis>
|
||
</taxonomicName>
|
||
in the Grijalva River basin, Mexico and Guatemala. The white square represents the type locality, black circles are localities of the typical morph and white circles are localities of the rheophilic morph.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3736516FFECFFA1E0CD457CFC2DC096" pageId="14" pageNumber="385" type="biology_ecology">
|
||
<paragraph id="8BD6369DFFECFFA1E0CD457CFE08C07E" blockId="14.[151,1437,1577,2034]" pageId="14" pageNumber="385">
|
||
The typical
|
||
<taxonomicName id="4C694D1EFFECFFA1E146457FFE71C126" baseAuthorityName="Taylor & Miller" baseAuthorityYear="1980" box="[332,459,1578,1602]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="385" phylum="Chordata" rank="species" species="hartwegi">
|
||
<emphasis id="B91DEA8FFFECFFA1E146457FFE71C126" box="[332,459,1578,1602]" italics="true" pageId="14" pageNumber="385">V. hartwegi</emphasis>
|
||
</taxonomicName>
|
||
morph inhabits a great variety of habitats. Large adults prefer deeper zones of the main river channel (>
|
||
<quantity id="4C919B78FFECFFA1E15A451BFE38C101" box="[336,386,1614,1638]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="385" unit="m" value="2.0">2 m</quantity>
|
||
depth) with variable but mainly moderate current and rocky or sandy substrates, usually without aquatic vegetation. Juveniles and subadults prefer shallow waters with rocky bottoms, sand, silt and mud in rivers and streams with swift to slow currents, usually without aquatic vegetation but covered by riverine vegetation. Also inhabits lakes and reservoirs. The species is also tolerant to sulphidic environments; it has been recorded in El Azufre and Baños del Carmen, municipalities of Tecpatán and Venustiano Carranza, respectively, both in
|
||
<collectingRegion id="49ADF87FFFECFFA1E0E64454FEF3C07E" box="[236,329,1793,1818]" country="Mexico" name="Chiapas" pageId="14" pageNumber="385">Chiapas</collectingRegion>
|
||
,
|
||
<collectingCountry id="F37E760DFFECFFA1E15F4454FE14C07E" box="[341,430,1793,1818]" name="Mexico" pageId="14" pageNumber="385">
|
||
<collectingRegion id="49ADF87FFFECFFA1E15F4454FE14C07E" box="[341,430,1793,1818]" country="Mexico" name="Mexico" pageId="14" pageNumber="385">Mexico</collectingRegion>
|
||
</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD6369DFFECFFA1E0CD4470FC2DC096" blockId="14.[151,1437,1577,2034]" pageId="14" pageNumber="385">
|
||
The rheophilic morph primarily inhabits rocky bottoms covered with boulders and rocks of various sizes and wooden debris, aquatic vegetation absent, in depths of
|
||
<quantity id="4C919B78FFECFFA1E30A441FFC38C005" box="[768,898,1866,1890]" metricMagnitude="-1" metricUnit="m" metricValue="6.5" metricValueMax="10.0" metricValueMin="3.0" pageId="14" pageNumber="385" unit="cm" value="65.0" valueMax="100.0" valueMin="30.0">30–100 cm</quantity>
|
||
; moderate to fast currents, with clear or slightly murky waters. This morph of
|
||
<taxonomicName id="4C694D1EFFECFFA1E1E7443BFDD4C0E1" baseAuthorityName="Taylor & Miller" baseAuthorityYear="1980" box="[493,622,1901,1925]" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="385" phylum="Chordata" rank="species" species="hartwegi">
|
||
<emphasis id="B91DEA8FFFECFFA1E1E7443BFDD4C0E1" box="[493,622,1901,1925]" italics="true" pageId="14" pageNumber="385">V. hartwegi</emphasis>
|
||
</taxonomicName>
|
||
is sympatric with the typical morph; however, the two forms are likely segregated because of habitat preferences and food habits. There are too few specimens to robustly analyze the food preferences, but slight differences in pharyngeal jaw dentition may be related to diet. Additionally,
|
||
<taxonomicName id="4C694D1EFFECFFA1E58D44E3FEBAC096" baseAuthorityName="Werner & Stawikowski" baseAuthorityYear="1987" class="Actinopterygii" family="Cichlidae" genus="Vieja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="385" phylum="Chordata" rank="species" species="breidohri">
|
||
<emphasis id="B91DEA8FFFECFFA1E58D44E3FEBAC096" italics="true" pageId="14" pageNumber="385">V. breidohri</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C694D1EFFECFFA1E13C448FFE63C096" baseAuthorityName="Taylor & Miller" baseAuthorityYear="1980" box="[310,473,2010,2034]" class="Actinopterygii" family="Cichlidae" genus="Chiapaheros" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="385" phylum="Chordata" rank="species" species="grammodes">
|
||
<emphasis id="B91DEA8FFFECFFA1E13C448FFE63C096" box="[310,473,2010,2034]" italics="true" pageId="14" pageNumber="385">C. grammodes</emphasis>
|
||
</taxonomicName>
|
||
are found in sympatry with this morph.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |