541 lines
72 KiB
XML
541 lines
72 KiB
XML
<document id="DCFF56BBC5DD06D2DCAD56DA864B882B" ID-CLB-Dataset="22100" ID-DOI="10.5852/ejt.2017.321" ID-GBIF-Dataset="caeead37-b6ce-4f03-96d2-ae8c8beb8573" ID-ISSN="2118-9773" ID-Zenodo-Dep="3829282" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589562339121" checkinUser="carolina" docAuthor="Maxwell, Erin E. & Stumpf, Sebastian" docDate="2017" docId="03B22572FF92FF86FD81F9D4BEDACAED" docLanguage="en" docName="ejt-2017-321.pdf.imf" docOrigin="European Journal of Taxonomy 321 (321)" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Saurorhynchus acutus" docType="treatment" docVersion="8" lastPageNumber="8" masterDocId="FF8B5D0AFF91FF81FFB1FFF9BF47CD01" masterDocTitle="Revision of Saurorhynchus (Actinopterygii: Saurichthyidae) from the Early Jurassic of England and Germany" masterLastPageNumber="29" masterPageNumber="1" pageNumber="4" updateTime="1698839856834" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="DC7F80F7E1BADBFF801DB9AB26135F22">Revision of Saurorhynchus (Actinopterygii: Saurichthyidae) from the Early Jurassic of England and Germany</mods:title>
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<mods:namePart id="7BFAE134DC9B89A416D0D86D51B11D17">Maxwell, Erin E.</mods:namePart>
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<mods:affiliation id="BE95E6E583DA3F190FD135D3E1F973F9">Staatliches Museum für Naturkunde Stuttgart, Rosenstein 1, 70191 Stuttgart, Germany. Institut für Geographie und Geologie, Ernst-Moritz-Arndt-Universität Greifswald, Friedrich-Ludwig-Jahn Strasse 17 a, 17489 Greifswald, Germany.</mods:affiliation>
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<mods:nameIdentifier id="11219CDEE26CCDB48E1AC7E749EBA9D9" type="email">erin.maxwell@smns-bw.de</mods:nameIdentifier>
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<mods:namePart id="D4DDFFC5C4031F4AA68D4BEE3DE27FD3">Stumpf, Sebastian</mods:namePart>
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<mods:affiliation id="696B7260F052B60FB78F6F1970E6A5E5">Email: sebastian. stumpf @ stud. uni-greifswald. de & urn: lsid: zoobank. org: author: 8 DADF 851 - CFA 5 - 469 E- 9628 - AD 020803 B 424</mods:affiliation>
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<mods:nameIdentifier id="F019D36F7D1C2B492FCE2A81EC0C3C11" type="email">sebastian.stumpf@stud.uni-greifswald.de</mods:nameIdentifier>
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<mods:title id="A7AEF396D5134FB54E90FA3F35B6AFA7">European Journal of Taxonomy</mods:title>
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<treatment id="03B22572FF92FF86FD81F9D4BEDACAED" ID-DOI="http://doi.org/10.5281/zenodo.3848068" ID-GBIF-Taxon="164159997" ID-Zenodo-Dep="3848068" LSID="urn:lsid:plazi:treatment:03B22572FF92FF86FD81F9D4BEDACAED" httpUri="http://treatment.plazi.org/id/03B22572FF92FF86FD81F9D4BEDACAED" lastPageId="7" lastPageNumber="8" pageId="3" pageNumber="4">
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<subSubSection id="C301C7EFFF92FF82FD81F9D4BB44CB49" box="[560,1027,1581,1608]" pageId="3" pageNumber="4" type="nomenclature">
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<paragraph id="8BA49464FF92FF82FD81F9D4BB44CB49" blockId="3.[560,1027,1581,1644]" box="[560,1027,1581,1608]" pageId="3" pageNumber="4">
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<heading id="D0EC2308FF92FF82FD81F9D4BB44CB49" box="[560,1027,1581,1608]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="4" reason="2">
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<taxonomicName id="4C1BEFE7FF92FF82FD81F9D4BB44CB49" authority="(Agassiz, 1844)" baseAuthorityName="Agassiz" baseAuthorityYear="1844" box="[560,1027,1581,1608]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="acutus">
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<emphasis id="B96F4876FF92FF82FD81F9D4BC04CB49" bold="true" box="[560,835,1581,1608]" italics="true" pageId="3" pageNumber="4">Saurorhynchus acutus</emphasis>
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(
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<bibRefCitation id="EF8AE995FF92FF82FCE3F9D7BCBCCB49" author="Agassiz L." box="[850,1019,1582,1608]" pageId="3" pageNumber="4" refId="ref12560" refString="Agassiz L. 1844. Recherches sur les Poissons Fossiles. II. Contenant l'Histoire de l'Ordre des Ganoides. Vol. 2, Part 2. Petitpierre, Neuchatel." type="book" year="1844">Agassiz, 1844</bibRefCitation>
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)
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C301C7EFFF92FF82FD3FF9A8BCE2CB6D" box="[654,933,1617,1644]" pageId="3" pageNumber="4" type="description">
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<paragraph id="8BA49464FF92FF82FD3FF9A8BCE2CB6D" blockId="3.[560,1027,1581,1644]" box="[654,933,1617,1644]" pageId="3" pageNumber="4">
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<figureCitation id="132088E1FF92FF82FD3FF9A8BDA9CB6A" box="[654,750,1617,1643]" captionStart="Fig" captionStartId="5.[189,232,1858,1884]" captionTargetBox="[274,1313,297,1817]" captionTargetId="figure@5.[274,1313,297,1817]" captionTargetPageId="5" captionText="Fig. 1. Holotype specimens of Early Jurassic saurichthyids. A. Saurorhynchus acutus (Agassiz, 1844) (NHMUK PV P 4268). B. Saurorhynchus brevirostris (Woodward, 1895) (NHMUK PV OR 40726). C. Saurorhynchus anningae sp. nov. (NHMUK PV P 3791). D–E. Saurorhynchus hauffi sp. nov. (SMNS 55057). Scale bars: A, C = 20 mm; E = 50 mm. Photos A–C © The Trustees of the Natural History Museum, London." figureDoi="http://doi.org/10.5281/zenodo.3829284" httpUri="https://zenodo.org/record/3829284/files/figure.png" pageId="3" pageNumber="4">Figs 1A</figureCitation>
|
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,
|
||
<figureCitation id="132088E1FF92FF82FD4BF9A8BC66CB6A" box="[762,801,1617,1643]" captionStart="Fig" captionStartId="6.[189,232,1647,1673]" captionTargetBox="[205,1385,332,1597]" captionTargetId="figure@6.[204,1385,329,1597]" captionTargetPageId="6" captionText="Fig. 2. Reconstruction of the dermal skull in Early Jurassic saurichthyids. A. Saurorhynchus brevirostris (Woodward, 1895). B. Saurorhynchus anningae sp. nov. C. Saurorhynchus hauffi sp. nov. D. Saurorhynchus acutus (Agassiz, 1844). Not to scale. Dashed lines indicate sensory canals, grey lines indicate bones that were present but where the exact location of the sutural contact is unclear, and question marks indicate areas of uncertainty. Abbreviations: ang = angular; bs = basisphenoid; cl = cleithrum; d = dentary; dh = dermohyal; dpt = dermopterotic; dsp = dermophenotic; f = frontal; io = infraorbital; io.c = infraorbital sensory canal; l = lacrimal; l.ex = lateral extrascapular; m = maxilla; md.c = mandibular sensory canal; mio.c = medial branch of the infraorbital sensory canal; n = neomorph; na-ao = nasaloantorbital; op = opercle; p = parietal; pm = rostropremaxilla; pop = preopercle; ps = parasphenoid; sa = supraangular; scl = supracleithrum; so = supraorbital; so.c = supraorbital sensory canal." figureDoi="http://doi.org/10.5281/zenodo.3829286" httpUri="https://zenodo.org/record/3829286/files/figure.png" pageId="3" pageNumber="4">2D</figureCitation>
|
||
,
|
||
<figureCitation id="132088E1FF92FF82FC9CF9A8BC13CB6A" box="[813,852,1617,1643]" captionStart="Fig" captionStartId="10.[189,232,1683,1709]" captionTargetBox="[189,1399,933,1642]" captionTargetId="figure@10.[189,1399,929,1642]" captionTargetPageId="10" captionText="Fig. 4. Orbitotemporal region, illustrating variation in the braincase of Early Jurassic saurichthyids. A. Saurorhynchus anningae sp. nov. (based on NHMUK PV P 36227). B, E. Saurorhynchus brevirostris (Woodward, 1895) (photo pertains to NHMUK PV P 4878, mirrored). C, F. Saurorhynchus hauffi sp. nov. (photo is of SMNS 51888, mirrored). D. Saurorhynchus acutus (Agassiz, 1844) (based on SMNS 87737). Abbreviations: asc = ascending process of the parasphenoid; bs = basisphenoid; dpt = dermopterotic; dsp = dermosphenotic; f.ic+ep = foramen for the internal carotid artery and efferent pseudobranchial artery; f.oma = f.om = foramen for the great ophthalmic artery; my = posterior myodome; n = neomorph; nc = neurocranium; orb = orbit; psr = parasphenoid rostrum; rot = foramen for the lateral otic ramus; spi = ventral opening of the spiracular canal; tf.c = trigeminofacial chamber. Scale bars: E–F = 2 mm. E. Photo © The Trustees of the Natural History Museum, London." figureDoi="http://doi.org/10.5281/zenodo.3829291" httpUri="https://zenodo.org/record/3829291/files/figure.png" pageId="3" pageNumber="4">4D</figureCitation>
|
||
,
|
||
<figureCitation id="132088E1FF92FF82FCEEF9A8BC2BCB6D" box="[863,876,1617,1644]" captionStart="Fig" captionStartId="13.[189,232,1790,1816]" captionTargetBox="[189,1399,869,1754]" captionTargetId="figure@13.[189,1399,869,1754]" captionTargetPageId="13" captionText="Fig. 5. Characters distinguishing Saurorhynchus acutus (Agassiz, 1844) and S. hauffi sp. nov. A. S. acutus, narial region (SMNS 56923). B. S. acutus, lateral extrascapular-dermopterotic contact (SMNS 57039). C. S. hauffi sp. nov., narial region (SMNS 51888). D. S. hauffi sp. nov., lateral extrascapular-dermopterotic contact (SMNS 53980, mirrored). Abbreviations: dpt = dermopterotic; dh = dermohyal; f = frontal; l.ex = lateral extrascapular; m = maxilla; na-ao = nasaloantorbital; orb = orbit; pm = rostropremaxilla; pop = preopercle; pop* = damaged preopercle; ps = parasphenoid; so = supraorbital. Scale bars: A, C–D = 2 mm; B = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3829293" httpUri="https://zenodo.org/record/3829293/files/figure.png" pageId="3" pageNumber="4">5</figureCitation>
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||
A–B
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C301C7EFFF92FF82FF0CF96BBBAACA10" pageId="3" pageNumber="4" type="reference_group">
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||
<paragraph id="8BA49464FF92FF82FF0CF96BBC0ACBAC" blockId="3.[189,845,1682,1744]" box="[189,845,1682,1709]" pageId="3" pageNumber="4">
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||
<treatmentCitationGroup id="AB0BB34AFF92FF82FF0CF96BBC0ACBAC" box="[189,845,1682,1709]" pageId="3" pageNumber="4">
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<treatmentCitation id="0ABAB275FF92FF82FF0CF96BBDE7CBAC" author="Agassiz L." box="[189,672,1682,1709]" page="142" pageId="3" pageNumber="4" year="1844">
|
||
<taxonomicName id="4C1BEFE7FF92FF82FF0CF96BBDE7CBAC" authority="Agassiz, 1844: 142" authorityName="Agassiz" authorityPageNumber="142" authorityYear="1844" box="[189,672,1682,1709]" class="Actinopterygii" family="Aspidorhynchidae" genus="Belonostomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aspidorhychiformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="acutus">
|
||
<emphasis id="B96F4876FF92FF82FF0CF96BBEF1CBAD" box="[189,438,1682,1708]" italics="true" pageId="3" pageNumber="4">Belonostomus acutus</emphasis>
|
||
<bibRefCitation id="EF8AE995FF92FF82FE0AF96BBDE7CBAC" author="Agassiz L." box="[443,672,1682,1709]" pageId="3" pageNumber="4" refId="ref12560" refString="Agassiz L. 1844. Recherches sur les Poissons Fossiles. II. Contenant l'Histoire de l'Ordre des Ganoides. Vol. 2, Part 2. Petitpierre, Neuchatel." type="book" year="1844">Agassiz, 1844: 142</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
, pl. 47a, fig. 4.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF92FF82FF0CF94FBC07CBCE" blockId="3.[189,845,1682,1744]" box="[189,832,1717,1744]" pageId="3" pageNumber="4">
|
||
<treatmentCitationGroup id="AB0BB34AFF92FF82FF0CF94FBC07CBCE" box="[189,832,1717,1744]" pageId="3" pageNumber="4">
|
||
<treatmentCitation id="0ABAB275FF92FF82FF0CF94FBDE3CBD1" box="[189,676,1717,1744]" pageId="3" pageNumber="4">
|
||
<taxonomicName id="4C1BEFE7FF92FF82FF0CF94FBDE3CBD1" authority="Stensio, 1925" authorityName="Stensio" authorityYear="1925" box="[189,676,1717,1744]" class="Actinopterygii" family="Saurichthyidae" genus="Acidorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="brevirostris">
|
||
<emphasis id="B96F4876FF92FF82FF0CF94FBEBCCBD1" box="[189,507,1718,1744]" italics="true" pageId="3" pageNumber="4">Acidorhynchus brevirostris</emphasis>
|
||
<bibRefCitation id="EF8AE995FF92FF82FDB3F94FBDE3CBD1" author="Stensio E." box="[514,676,1717,1744]" pageId="3" pageNumber="4" pagination="1 - 126" refId="ref14959" refString="Stensio E. 1925. Triassic fishes from Spitzbergen, Part II. Kungliga Svenska Vetenskapsakademiens Handlingar Tredje Serien 2 (1): 1 - 126." type="journal article" year="1925">Stensiö, 1925</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
, partim: 176.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF92FF82FF0CF90EBBAACA10" blockId="3.[189,1261,1783,1810]" box="[189,1261,1783,1810]" pageId="3" pageNumber="4">
|
||
<treatmentCitationGroup id="AB0BB34AFF92FF82FF0CF90EBBAACA10" box="[189,1261,1783,1810]" pageId="3" pageNumber="4">
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||
<taxonomicName id="4C1BEFE7FF92FF82FF0CF90EBEBCCA10" authorityName="Stensio" authorityYear="1925" box="[189,507,1783,1809]" class="Actinopterygii" family="Saurichthyidae" genus="Acidorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="brevirostris">
|
||
<emphasis id="B96F4876FF92FF82FF0CF90EBEBCCA10" box="[189,507,1783,1809]" italics="true" pageId="3" pageNumber="4">Acidorhynchus brevirostris</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0ABAB275FF92FF82FDA9F90EBDD9CA10" box="[536,670,1783,1809]" pageId="3" pageNumber="4">
|
||
<bibRefCitation id="EF8AE995FF92FF82FDA9F90EBDD9CA10" author="Hauff B." box="[536,670,1783,1809]" pageId="3" pageNumber="4" pagination="214 - 248" refId="ref13522" refString="Hauff B. 1938. Uber Acidorhynchus aus den Posidonienschiefern von Holzmaden. Palaontologische Zeitschrift 20: 214 - 248. https: // doi. org / 10.1007 / BF 03041918" type="journal article" year="1938">Hauff 1938</bibRefCitation>
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||
</treatmentCitation>
|
||
, partim: pl. 22, fig. d. —
|
||
<treatmentCitation id="0ABAB275FF92FF82FC78F901BB0ACA10" box="[969,1101,1783,1810]" pageId="3" pageNumber="4">
|
||
<bibRefCitation id="EF8AE995FF92FF82FC78F901BB0ACA10" author="Wenz S." box="[969,1101,1783,1810]" pageId="3" pageNumber="4" pagination="1 - 276" refId="ref15053" refString="Wenz S. 1967. Complements a l'etude des poissons actinopterygiens du Jurassique Francais. Cahiers de Paleontologie 1967: 1 - 276." type="journal article" year="1967">Wenz 1967</bibRefCitation>
|
||
</treatmentCitation>
|
||
: pl. II, fig. A.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C301C7EFFF92FF85FF0CF8C1BA29CC91" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="diagnosis">
|
||
<paragraph id="8BA49464FF92FF82FF0CF8C1BEDDCA53" blockId="3.[189,410,1848,1874]" box="[189,410,1848,1874]" pageId="3" pageNumber="4">
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<heading id="D0EC2308FF92FF82FF0CF8C1BEDDCA53" bold="true" box="[189,410,1848,1874]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
|
||
<emphasis id="B96F4876FF92FF82FF0CF8C1BEDDCA53" bold="true" box="[189,410,1848,1874]" pageId="3" pageNumber="4">Revised diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF92FF85FF0CF89EBA29CC91" blockId="3.[189,1398,1895,2027]" lastBlockId="4.[189,1398,267,401]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
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||
Anterior narial opening narrow and elongate; maxilla strongly dorsoventrally compressed and ventrally deflected suborbitally such that the contact with the premaxilla occurs at an angle; posterior dorsal skull parallel to the long axis of the skull; postorbital segment approximately equal in length to the maximum depth of the lower jaw; parasphenoid edentulous ventral to the orbit; foramen for the internal carotid and efferent pseudobranchial arteries centered within the basisphenoid (sensu
|
||
<bibRefCitation id="EF8AE995FF95FF85FB35FEF5BA55CC27" author="Wenz S." box="[1156,1298,267,294]" pageId="4" pageNumber="5" pagination="1 - 276" refId="ref15053" refString="Wenz S. 1967. Complements a l'etude des poissons actinopterygiens du Jurassique Francais. Cahiers de Paleontologie 1967: 1 - 276." type="journal article" year="1967">Wenz 1967</bibRefCitation>
|
||
); lateral extrascapular fused to lateral dermopterotic and not extending dorsal to the hyomandibula; posterior edge of the mandible strongly sinusoidal; angle between the posterior and ventral edges of the mandible less than 80 degrees; subnarial laniary dentition absent; acrodin caps of the posterior laniaries straight.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C301C7EFFF95FF85FF0CFE41BD1DCE3D" pageId="4" pageNumber="5" type="materials_examined">
|
||
<paragraph id="8BA49464FF95FF85FF0CFE41BECBCCD3" blockId="4.[189,396,440,466]" box="[189,396,440,466]" pageId="4" pageNumber="5">
|
||
<heading id="D0EC2308FF95FF85FF0CFE41BECBCCD3" bold="true" box="[189,396,440,466]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
|
||
<emphasis id="B96F4876FF95FF85FF0CFE41BECBCCD3" bold="true" box="[189,396,440,466]" pageId="4" pageNumber="5">Material studied</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<materialsCitation id="3B739E39FF95FF85FF5BFE1EBB7FCF25" ID-GBIF-Occurrence="2620280301" collectionCode="(NHMUK" country="United Kingdom" location="Toarcian of Whitby" pageId="4" pageNumber="5" specimenCode="NHMUK PV P 4268" specimenCount="1" typeStatus="holotype">
|
||
<paragraph id="8BA49464FF95FF85FF5BFE1EBE1BCF00" blockId="4.[189,1080,487,549]" box="[234,348,487,513]" pageId="4" pageNumber="5">
|
||
<emphasis id="B96F4876FF95FF85FF5BFE1EBE1BCF00" bold="true" box="[234,348,487,513]" pageId="4" pageNumber="5">
|
||
<typeStatus id="54A02AC6FF95FF85FF5BFE1EBE1BCF00" box="[234,348,487,513]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF95FF85FF0CFDF2BB7FCF25" blockId="4.[189,1080,487,549]" box="[189,1080,522,549]" pageId="4" pageNumber="5">
|
||
<collectingCountry id="F30CD4F4FF95FF85FF0CFDF2BE84CF25" box="[189,451,522,549]" name="United Kingdom" pageId="4" pageNumber="5">UNITED KINGDOM</collectingCountry>
|
||
:
|
||
<location id="8EC4C2BFFF95FF85FE7FFDF3BDF4CF25" LSID="urn:lsid:plazi:treatment:03B22572FF92FF86FD81F9D4BEDACAED:8EC4C2BFFF95FF85FE7FFDF3BDF4CF25" box="[462,691,522,548]" country="United Kingdom" name="Toarcian of Whitby" pageId="4" pageNumber="5">Toarcian of Whitby</location>
|
||
<collectionCode id="ED0A0CA1FF95FF85FD0BFDF3BC74CF25" box="[698,819,522,548]" pageId="4" pageNumber="5">(NHMUK</collectionCode>
|
||
PV P 4268,
|
||
<figureCitation id="132088E1FF95FF85FC7FFDF3BB6DCF25" box="[974,1066,522,548]" captionStart="Fig" captionStartId="5.[189,232,1858,1884]" captionTargetBox="[274,1313,297,1817]" captionTargetId="figure@5.[274,1313,297,1817]" captionTargetPageId="5" captionText="Fig. 1. Holotype specimens of Early Jurassic saurichthyids. A. Saurorhynchus acutus (Agassiz, 1844) (NHMUK PV P 4268). B. Saurorhynchus brevirostris (Woodward, 1895) (NHMUK PV OR 40726). C. Saurorhynchus anningae sp. nov. (NHMUK PV P 3791). D–E. Saurorhynchus hauffi sp. nov. (SMNS 55057). Scale bars: A, C = 20 mm; E = 50 mm. Photos A–C © The Trustees of the Natural History Museum, London." figureDoi="http://doi.org/10.5281/zenodo.3829284" httpUri="https://zenodo.org/record/3829284/files/figure.png" pageId="4" pageNumber="5">Fig. 1A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</materialsCitation>
|
||
<paragraph id="8BA49464FF95FF85FF5BFDB5BE8CCF67" blockId="4.[189,1399,588,828]" box="[234,459,588,614]" pageId="4" pageNumber="5">
|
||
<emphasis id="B96F4876FF95FF85FF5BFDB5BE8CCF67" bold="true" box="[234,459,588,614]" pageId="4" pageNumber="5">Referred material</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF95FF85FF0CFD89BD1DCE3D" blockId="4.[189,1399,588,828]" pageId="4" pageNumber="5">
|
||
<materialsCitation id="3B739E39FF95FF85FF0CFD89BA31CF8B" ID-GBIF-Occurrence="3763234309" box="[189,1398,623,650]" collectionCode="A" country="Germany" location="Bisingen" municipality="Bisingen" pageId="4" pageNumber="5" specimenCode="SMNS 56923" specimenCount="1">
|
||
<collectingCountry id="F30CD4F4FF95FF85FF0CFD89BE14CF8B" box="[189,339,624,650]" name="Germany" pageId="4" pageNumber="5">GERMANY</collectingCountry>
|
||
: all referred material consists of isolated skulls.
|
||
<collectingMunicipality id="6BC00E1EFF95FF85FC75FD89BB6BCF8B" box="[964,1068,624,650]" pageId="4" pageNumber="5">Bisingen</collectingMunicipality>
|
||
(
|
||
<specimenCode id="DBBD3C1FFF95FF85FBF2FD89BBB7CF8B" box="[1091,1264,623,650]" pageId="4" pageNumber="5">SMNS 56923</specimenCode>
|
||
,
|
||
<figureCitation id="132088E1FF95FF85FAB2FD89BA2FCF8B" box="[1283,1384,623,650]" captionStart="Fig" captionStartId="13.[189,232,1790,1816]" captionTargetBox="[189,1399,869,1754]" captionTargetId="figure@13.[189,1399,869,1754]" captionTargetPageId="13" captionText="Fig. 5. Characters distinguishing Saurorhynchus acutus (Agassiz, 1844) and S. hauffi sp. nov. A. S. acutus, narial region (SMNS 56923). B. S. acutus, lateral extrascapular-dermopterotic contact (SMNS 57039). C. S. hauffi sp. nov., narial region (SMNS 51888). D. S. hauffi sp. nov., lateral extrascapular-dermopterotic contact (SMNS 53980, mirrored). Abbreviations: dpt = dermopterotic; dh = dermohyal; f = frontal; l.ex = lateral extrascapular; m = maxilla; na-ao = nasaloantorbital; orb = orbit; pm = rostropremaxilla; pop = preopercle; pop* = damaged preopercle; ps = parasphenoid; so = supraorbital. Scale bars: A, C–D = 2 mm; B = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3829293" httpUri="https://zenodo.org/record/3829293/files/figure.png" pageId="4" pageNumber="5">
|
||
Fig. 5
|
||
<collectionCode id="ED0A0CA1FF95FF85FAFEFD96BA2FCF8B" box="[1359,1384,623,650]" pageId="4" pageNumber="5">A</collectionCode>
|
||
</figureCitation>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B739E39FF95FF85FF0CFD6ABC74CFAC" ID-GBIF-Occurrence="3763234310" box="[189,819,659,686]" collectionCode="B" location="Holzmaden" pageId="4" pageNumber="5" specimenCode="SMNS 57039, SMNS 88007" specimenCount="1">
|
||
<location id="8EC4C2BFFF95FF85FF0CFD6ABE03CFAC" LSID="urn:lsid:plazi:treatment:03B22572FF92FF86FD81F9D4BEDACAED:8EC4C2BFFF95FF85FF0CFD6ABE03CFAC" box="[189,324,659,685]" name="Holzmaden" pageId="4" pageNumber="5">Holzmaden</location>
|
||
(
|
||
<specimenCode id="DBBD3C1FFF95FF85FEE4FD6ABEBFCFAF" box="[341,504,659,686]" pageId="4" pageNumber="5">SMNS 57039</specimenCode>
|
||
(
|
||
<figureCitation id="132088E1FF95FF85FDBBFD6ABD20CFAF" box="[522,615,659,686]" captionStart="Fig" captionStartId="13.[189,232,1790,1816]" captionTargetBox="[189,1399,869,1754]" captionTargetId="figure@13.[189,1399,869,1754]" captionTargetPageId="13" captionText="Fig. 5. Characters distinguishing Saurorhynchus acutus (Agassiz, 1844) and S. hauffi sp. nov. A. S. acutus, narial region (SMNS 56923). B. S. acutus, lateral extrascapular-dermopterotic contact (SMNS 57039). C. S. hauffi sp. nov., narial region (SMNS 51888). D. S. hauffi sp. nov., lateral extrascapular-dermopterotic contact (SMNS 53980, mirrored). Abbreviations: dpt = dermopterotic; dh = dermohyal; f = frontal; l.ex = lateral extrascapular; m = maxilla; na-ao = nasaloantorbital; orb = orbit; pm = rostropremaxilla; pop = preopercle; pop* = damaged preopercle; ps = parasphenoid; so = supraorbital. Scale bars: A, C–D = 2 mm; B = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3829293" httpUri="https://zenodo.org/record/3829293/files/figure.png" pageId="4" pageNumber="5">
|
||
Fig. 5
|
||
<collectionCode id="ED0A0CA1FF95FF85FDE3FD6ABD20CFAF" box="[594,615,659,686]" pageId="4" pageNumber="5">B</collectionCode>
|
||
</figureCitation>
|
||
),
|
||
<specimenCode id="DBBD3C1FFF95FF85FDCEFD6ABC62CFAC" box="[639,805,659,685]" pageId="4" pageNumber="5">SMNS 88007</specimenCode>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B739E39FF95FF85FC8DFD6ABBE9CFAC" ID-GBIF-Occurrence="3763234322" box="[828,1198,659,691]" location="Holzmaden" municipality="Holzmaden" pageId="4" pageNumber="5" specimenCount="1">
|
||
<collectingMunicipality id="6BC00E1EFF95FF85FC8DFD6ABC81CFAC" box="[828,966,659,685]" pageId="4" pageNumber="5">Holzmaden</collectingMunicipality>
|
||
, εII
|
||
<subScript id="179F9621FF95FF85FC42FD5CBCBBCFB2" attach="left" box="[1011,1020,677,691]" fontSize="6" pageId="4" pageNumber="5">3</subScript>
|
||
(PMU 30009);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B739E39FF95FF85FB06FD6ABEACCFD0" ID-GBIF-Occurrence="3763234320" collectionCode="SMNS, MHH" location="Holzmaden" pageId="4" pageNumber="5" specimenCode="SMNS 50924, MHH 17" specimenCount="1">
|
||
<location id="8EC4C2BFFF95FF85FB06FD6ABA06CFAC" LSID="urn:lsid:plazi:treatment:03B22572FF92FF86FD81F9D4BEDACAED:8EC4C2BFFF95FF85FB06FD6ABA06CFAC" box="[1207,1345,659,685]" name="Holzmaden" pageId="4" pageNumber="5">Holzmaden</location>
|
||
, εII
|
||
<subScript id="179F9621FF95FF85FADFFD5CBA30CFB2" attach="left" box="[1390,1399,677,691]" fontSize="6" pageId="4" pageNumber="5">4</subScript>
|
||
(
|
||
<specimenCode id="DBBD3C1FFF95FF85FF74FD4EBE2FCFD3" box="[197,360,695,722]" pageId="4" pageNumber="5">SMNS 50924</specimenCode>
|
||
,
|
||
<specimenCode id="DBBD3C1FFF95FF85FEC3FD4EBE9BCFD0" box="[370,476,695,721]" pageId="4" pageNumber="5">MHH 17</specimenCode>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B739E39FF95FF85FE40FD4EBB41CFD0" ID-GBIF-Occurrence="3763234318" box="[497,1030,695,727]" collectionCode="SMNS" location="Holzmaden" pageId="4" pageNumber="5" specimenCode="SMNS 55324" specimenCount="1">
|
||
<location id="8EC4C2BFFF95FF85FE40FD4EBD3CCFD0" LSID="urn:lsid:plazi:treatment:03B22572FF92FF86FD81F9D4BEDACAED:8EC4C2BFFF95FF85FE40FD4EBD3CCFD0" box="[497,635,695,721]" name="Holzmaden" pageId="4" pageNumber="5">Holzmaden</location>
|
||
, εII
|
||
<subScript id="179F9621FF95FF85FD14FD30BDE9CFD6" attach="left" box="[677,686,713,727]" fontSize="6" pageId="4" pageNumber="5">6</subScript>
|
||
(
|
||
<specimenCode id="DBBD3C1FFF95FF85FD08FD4EBC1BCFD3" box="[697,860,695,722]" pageId="4" pageNumber="5">SMNS 55324</specimenCode>
|
||
, PMU 30010);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B739E39FF95FF85FBBDFD4EBB90CFF5" ID-GBIF-Occurrence="3763234311" collectionCode="NHMUK, PV, P, GPIT, OS" location="Ohmden" pageId="4" pageNumber="5" specimenCode="SMNS 55319, SMNS 87738" specimenCount="1">
|
||
<location id="8EC4C2BFFF95FF85FBBDFD4EBB36CFD0" LSID="urn:lsid:plazi:treatment:03B22572FF92FF86FD81F9D4BEDACAED:8EC4C2BFFF95FF85FBBDFD4EBB36CFD0" box="[1036,1137,695,721]" name="Ohmden" pageId="4" pageNumber="5">Ohmden</location>
|
||
(
|
||
<collectionCode id="ED0A0CA1FF95FF85FBCEFD4EBBB7CFD0" box="[1151,1264,695,721]" collectionName="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="4" pageNumber="5" type="Museum">NHMUK</collectionCode>
|
||
<collectionCode id="ED0A0CA1FF95FF85FB44FD4EBA68CFD0" box="[1269,1327,695,721]" pageId="4" pageNumber="5">PV P</collectionCode>
|
||
3792,
|
||
<collectionCode id="ED0A0CA1FF95FF85FF0CFD22BE6BCFF4" box="[189,300,731,757]" collectionName="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="4" pageNumber="5" type="Museum">NHMUK</collectionCode>
|
||
<collectionCode id="ED0A0CA1FF95FF85FE80FD22BE11CFF4" box="[305,342,731,757]" pageId="4" pageNumber="5">PV</collectionCode>
|
||
P 36222,
|
||
<collectionCode id="ED0A0CA1FF95FF85FE74FD22BD73CFF4" box="[453,564,731,757]" collectionName="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="4" pageNumber="5" type="Museum">NHMUK</collectionCode>
|
||
<collectionCode id="ED0A0CA1FF95FF85FD88FD22BD19CFF4" box="[569,606,731,757]" pageId="4" pageNumber="5">PV</collectionCode>
|
||
P 36223,
|
||
<specimenCode id="DBBD3C1FFF95FF85FD7CFD23BC29CFF4" box="[717,878,730,757]" pageId="4" pageNumber="5">SMNS 55319</specimenCode>
|
||
,
|
||
<specimenCode id="DBBD3C1FFF95FF85FCC9FD23BB5ECFF5" box="[888,1049,730,756]" pageId="4" pageNumber="5">SMNS 87738</specimenCode>
|
||
,
|
||
<collectionCode id="ED0A0CA1FF95FF85FB92FD23BB2ECFF5" box="[1059,1129,730,756]" country="0" httpUri="http://grbio.org/cool/b26r-wc8g" name="Institut und Museum fur Geologie und Palaeontologie, Universitat Tuebingen" pageId="4" pageNumber="5">GPIT</collectionCode>
|
||
/
|
||
<collectionCode id="ED0A0CA1FF95FF85FBC3FD23BBD0CFF5" box="[1138,1175,730,756]" pageId="4" pageNumber="5">OS</collectionCode>
|
||
/133);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B739E39FF95FF85FB6CFD23BE3FCE18" ID-GBIF-Occurrence="3763234301" collectionCode="SMNS" location="Ohmden" municipality="Ohmden" pageId="4" pageNumber="5" specimenCode="SMNS 51009" specimenCount="1">
|
||
<collectingMunicipality id="6BC00E1EFF95FF85FB6CFD23BA02CFF5" box="[1245,1349,730,756]" pageId="4" pageNumber="5">Ohmden</collectingMunicipality>
|
||
, εII
|
||
<subScript id="179F9621FF95FF85FADFFD15BA30CFFB" attach="left" box="[1390,1399,748,762]" fontSize="6" pageId="4" pageNumber="5">3</subScript>
|
||
(
|
||
<specimenCode id="DBBD3C1FFF95FF85FF74FD07BE2CCE18" box="[197,363,766,793]" pageId="4" pageNumber="5">SMNS 51009</specimenCode>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B739E39FF95FF85FECFFD07BD90CE18" ID-GBIF-Occurrence="3763234307" box="[382,727,766,798]" collectionCode="SMNS" location="Ohmden" pageId="4" pageNumber="5" specimenCode="SMNS 50268" specimenCount="1">
|
||
<location id="8EC4C2BFFF95FF85FECFFD07BEA1CE19" LSID="urn:lsid:plazi:treatment:03B22572FF92FF86FD81F9D4BEDACAED:8EC4C2BFFF95FF85FECFFD07BEA1CE19" box="[382,486,766,792]" name="Ohmden" pageId="4" pageNumber="5">Ohmden</location>
|
||
, εII
|
||
<subScript id="179F9621FF95FF85FDA1FCE9BD5ECE1F" attach="left" box="[528,537,784,798]" fontSize="6" pageId="4" pageNumber="5">4</subScript>
|
||
(
|
||
<specimenCode id="DBBD3C1FFF95FF85FD95FD07BD8ECE18" box="[548,713,766,793]" pageId="4" pageNumber="5">SMNS 50268</specimenCode>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B739E39FF95FF85FD6CFD07BBA5CE18" ID-GBIF-Occurrence="3763234312" box="[733,1250,766,798]" collectionCode="SMNS" location="Ohmden" pageId="4" pageNumber="5" specimenCode="SMNS 96927" specimenCount="1">
|
||
<location id="8EC4C2BFFF95FF85FD6CFD07BC02CE19" LSID="urn:lsid:plazi:treatment:03B22572FF92FF86FD81F9D4BEDACAED:8EC4C2BFFF95FF85FD6CFD07BC02CE19" box="[733,837,766,792]" name="Ohmden" pageId="4" pageNumber="5">Ohmden</location>
|
||
, εII
|
||
<subScript id="179F9621FF95FF85FCDEFCE9BC3FCE1F" attach="left" box="[879,888,784,798]" fontSize="6" pageId="4" pageNumber="5">5</subScript>
|
||
(
|
||
<specimenCode id="DBBD3C1FFF95FF85FC32FD07BB61CE19" box="[899,1062,766,792]" pageId="4" pageNumber="5">SMNS 96927</specimenCode>
|
||
,
|
||
<specimenCode id="DBBD3C1FFF95FF85FB80FD07BB92CE18" box="[1073,1237,766,793]" pageId="4" pageNumber="5">SMNS 96927</specimenCode>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B739E39FF95FF85FB59FD07BD1DCE3D" ID-GBIF-Occurrence="3763234315" collectionCode="GZG" location="Zone" municipality="Schandelah" pageId="4" pageNumber="5" specimenCount="1">
|
||
<collectingMunicipality id="6BC00E1EFF95FF85FB59FD07BA35CE19" box="[1256,1394,766,792]" pageId="4" pageNumber="5">Schandelah</collectingMunicipality>
|
||
,
|
||
<emphasis id="B96F4876FF95FF85FF0CFCDBBE09CE3A" box="[189,334,802,827]" italics="true" pageId="4" pageNumber="5">serpentinum</emphasis>
|
||
<location id="8EC4C2BFFF95FF85FEE4FCDBBED6CE3D" LSID="urn:lsid:plazi:treatment:03B22572FF92FF86FD81F9D4BEDACAED:8EC4C2BFFF95FF85FEE4FCDBBED6CE3D" box="[341,401,802,828]" municipality="Schandelah" name="Zone" pageId="4" pageNumber="5">Zone</location>
|
||
(
|
||
<collectionCode id="ED0A0CA1FF95FF85FE10FCDBBEA5CE3D" box="[417,482,802,828]" pageId="4" pageNumber="5">GZG</collectionCode>
|
||
.V.27932).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C301C7EFFF95FF86FF0CFC9ABC50C944" lastPageId="7" lastPageNumber="8" pageId="4" pageNumber="5" type="description">
|
||
<paragraph id="8BA49464FF95FF85FF0CFC9ABE09CE7C" blockId="4.[189,334,867,893]" box="[189,334,867,893]" pageId="4" pageNumber="5">
|
||
<heading id="D0EC2308FF95FF85FF0CFC9ABE09CE7C" bold="true" box="[189,334,867,893]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
|
||
<emphasis id="B96F4876FF95FF85FF0CFC9ABE09CE7C" bold="true" box="[189,334,867,893]" pageId="4" pageNumber="5">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF95FF85FF0CFC6BBD04C916" blockId="4.[189,1398,914,1047]" pageId="4" pageNumber="5">
|
||
<taxonomicName id="4C1BEFE7FF95FF85FF0CFC6BBE86CEAD" baseAuthorityName="Agassiz" baseAuthorityYear="1844" box="[189,449,914,940]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="acutus">
|
||
<emphasis id="B96F4876FF95FF85FF0CFC6BBE86CEAD" box="[189,449,914,940]" italics="true" pageId="4" pageNumber="5">Saurorhynchus acutus</emphasis>
|
||
</taxonomicName>
|
||
is anatomically very similar to the Toarcian
|
||
<taxonomicName id="4C1BEFE7FF95FF85FC7FFC6ABB68CEAD" authority="Maxwell & Stumpf, 2017" authorityName="Maxwell & Stumpf" authorityYear="2017" box="[974,1071,914,941]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="hauffi" status="sp. nov.">
|
||
<emphasis id="B96F4876FF95FF85FC7FFC6ABB68CEAD" box="[974,1071,914,941]" italics="true" pageId="4" pageNumber="5">S. hauffi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25CF50DFF95FF85FB87FC6ABBD4CEAC" box="[1078,1171,915,941]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, described in detail in a subsequent section. The following brief description is designed only to differentiate the two species, to clarify general osteological nomenclature within
|
||
<taxonomicName id="4C1BEFE7FF95FF85FCA4FC20BC82CEF2" authorityName="Reis" authorityYear="1892" box="[789,965,985,1011]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||
<emphasis id="B96F4876FF95FF85FCA4FC20BC82CEF2" box="[789,965,985,1011]" italics="true" pageId="4" pageNumber="5">Saurorhynchus</emphasis>
|
||
</taxonomicName>
|
||
, and to describe parts of the anatomy not preserved in
|
||
<taxonomicName id="4C1BEFE7FF95FF85FE30FC04BEA5C916" authority="Maxwell & Stumpf, 2017" authorityName="Maxwell & Stumpf" authorityYear="2017" box="[385,482,1021,1047]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="hauffi" status="sp. nov.">
|
||
<emphasis id="B96F4876FF95FF85FE30FC04BEA5C916" box="[385,482,1021,1047]" italics="true" pageId="4" pageNumber="5">S. hauffi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25CF50DFF95FF85FE58FC04BD04C916" box="[489,579,1021,1047]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF95FF85FF0CFBC6BE13C80B" blockId="4.[189,1399,1087,1290]" pageId="4" pageNumber="5">
|
||
No well-preserved postcranial material is available for
|
||
<taxonomicName id="4C1BEFE7FF95FF85FCDBFBC6BB33C958" baseAuthorityName="Agassiz" baseAuthorityYear="1844" box="[874,1140,1087,1113]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="acutus">
|
||
<emphasis id="B96F4876FF95FF85FCDBFBC6BB33C958" box="[874,1140,1087,1113]" italics="true" pageId="4" pageNumber="5">Saurorhynchus acutus</emphasis>
|
||
</taxonomicName>
|
||
. Skull length (tip of rostrum to jaw joint, and corresponding to the length of the dermatocranium when measured along the dorsal midline) of the largest referred specimen is
|
||
<quantity id="4CE33981FF95FF85FCB9FB7FBC2EC9A1" box="[776,873,1158,1185]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" pageId="4" pageNumber="5" unit="mm" value="125.0">125 mm</quantity>
|
||
(
|
||
<figureCitation id="132088E1FF95FF85FCC6FB7FBB45C9A1" box="[887,1026,1158,1184]" captionStart="Appendix 1" captionStartId="27.[189,311,267,293]" captionTargetPageId="27" captionText="Appendix 1. Select cranial measurements for Early Jurassic species of Saurorhynchus Reis, 1892." figureDoi="http://doi.org/10.5281/zenodo.3829299" httpUri="https://zenodo.org/record/3829299/files/figure.png" pageId="4" pageNumber="5">Appendix 1</figureCitation>
|
||
). Assuming similar proportions to
|
||
<taxonomicName id="4C1BEFE7FF95FF85FF6DFB53BE79C9C2" authority="Maxwell & Stumpf, 2017" authorityName="Maxwell & Stumpf" authorityYear="2017" box="[220,318,1193,1220]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="hauffi" status="sp. nov.">
|
||
<emphasis id="B96F4876FF95FF85FF6DFB53BE79C9C2" box="[220,318,1193,1220]" italics="true" pageId="4" pageNumber="5">S. hauffi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25CF50DFF95FF85FEF6FB53BEE2C9C5" box="[327,421,1194,1220]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, fork length is estimated at
|
||
<quantity id="4CE33981FF95FF85FD40FB50BC7CC9C5" box="[753,827,1193,1220]" metricMagnitude="-1" metricUnit="m" metricValue="4.4" pageId="4" pageNumber="5" unit="cm" value="44.0">44 cm</quantity>
|
||
for large individuals of
|
||
<taxonomicName id="4C1BEFE7FF95FF85FBEFFB53BB8EC9C2" baseAuthorityName="Agassiz" baseAuthorityYear="1844" box="[1118,1225,1194,1220]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="acutus">
|
||
<emphasis id="B96F4876FF95FF85FBEFFB53BB8EC9C2" box="[1118,1225,1194,1220]" italics="true" pageId="4" pageNumber="5">S. acutus</emphasis>
|
||
</taxonomicName>
|
||
. All the skulls examined in the course of this study were severely compressed, making the cranial sutures difficult to differentiate.
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF95FF85FF0CFACBBB0DCB8C" blockId="4.[189,1399,1330,1677]" pageId="4" pageNumber="5">
|
||
There is no clear evidence for interrostral bones. The anterior narial opening is an elongate anteroventrally inclined slit. When the skull is viewed dorsolaterally, it is evident based on the orientation and lipping along the anterior external narial opening that it is functionally situated more dorsally than the posterior narial opening. The suborbital bar of the maxilla is strongly dorsoventrally compressed ventral to the orbit and is also ventrally deflected such that it meets the rostropremaxilla at an angle ventral to the external narial opening. This creates the impression of proportionately large, round orbits. The suture between the maxilla and premaxilla is interdigitating, differing from some of the Toarcian material of
|
||
<taxonomicName id="4C1BEFE7FF95FF85FF0CF9D2BE2ACB44" authorityName="Reis" authorityYear="1892" box="[189,365,1579,1605]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||
<emphasis id="B96F4876FF95FF85FF0CF9D2BE2ACB44" box="[189,365,1579,1605]" italics="true" pageId="4" pageNumber="5">Saurorhynchus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F30CD4F4FF95FF85FE07F9D5BD41CB47" box="[438,518,1580,1606]" name="France" pageId="4" pageNumber="5">France</collectingCountry>
|
||
and Northern
|
||
<collectingCountry id="F30CD4F4FF95FF85FD03F9D2BC58CB44" box="[690,799,1579,1605]" name="Germany" pageId="4" pageNumber="5">Germany</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EF8AE995FF95FF85FC9EF9D2BCFECB44" author="Wenz S." box="[815,953,1579,1605]" pageId="4" pageNumber="5" pagination="1 - 276" refId="ref15053" refString="Wenz S. 1967. Complements a l'etude des poissons actinopterygiens du Jurassique Francais. Cahiers de Paleontologie 1967: 1 - 276." type="journal article" year="1967">Wenz 1967</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8AE995FF95FF85FC77F9D2BB0CCB47" author="Thies D." box="[966,1099,1579,1606]" pageId="4" pageNumber="5" pagination="183 - 203" refId="ref14987" refString="Thies D. 1985. Funde von Acidorhynchus brevirostris (Woodward 1895) aus dem Posidonienschiefer (Unter-Toarcium) NW-Deutschlands. Palaeontographica Abt. A 187 (4 - 6): 183 - 203." type="journal article" year="1985">Thies 1985</bibRefCitation>
|
||
). In some specimens, the ornamentation on the skull roof is drastically reduced and simplified, consisting mainly of pitting and being slightly more reticular only in the region of the parietals (e.g.,
|
||
<figureCitation id="132088E1FF95FF85FC50F98BBB7BCB8C" box="[993,1084,1650,1677]" captionStart="Fig" captionStartId="13.[189,232,1790,1816]" captionTargetBox="[189,1399,869,1754]" captionTargetId="figure@13.[189,1399,869,1754]" captionTargetPageId="13" captionText="Fig. 5. Characters distinguishing Saurorhynchus acutus (Agassiz, 1844) and S. hauffi sp. nov. A. S. acutus, narial region (SMNS 56923). B. S. acutus, lateral extrascapular-dermopterotic contact (SMNS 57039). C. S. hauffi sp. nov., narial region (SMNS 51888). D. S. hauffi sp. nov., lateral extrascapular-dermopterotic contact (SMNS 53980, mirrored). Abbreviations: dpt = dermopterotic; dh = dermohyal; f = frontal; l.ex = lateral extrascapular; m = maxilla; na-ao = nasaloantorbital; orb = orbit; pm = rostropremaxilla; pop = preopercle; pop* = damaged preopercle; ps = parasphenoid; so = supraorbital. Scale bars: A, C–D = 2 mm; B = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3829293" httpUri="https://zenodo.org/record/3829293/files/figure.png" pageId="4" pageNumber="5">Fig. 5B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF95FF86FF0CF94DBC86CCBB" blockId="4.[189,1399,1716,2027]" lastBlockId="7.[189,1399,267,442]" lastPageId="7" lastPageNumber="8" pageId="4" pageNumber="5">
|
||
Unlike in
|
||
<emphasis id="B96F4876FF95FF85FE8AF94CBEACCBCF" box="[315,491,1716,1743]" italics="true" pageId="4" pageNumber="5">
|
||
<taxonomicName id="4C1BEFE7FF95FF85FE8AF94CBEA0CBCF" authorityName="Stensio" authorityYear="1925" box="[315,487,1716,1743]" class="Actinopterygii" family="Saurichthyidae" genus="Acidorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="brevirostris">S. brevirostris</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
the dorsal skull roof extends posteriorly without strong dorsal deflection. The element sutured to the posterolateral dermopterotic has been documented in
|
||
<taxonomicName id="4C1BEFE7FF95FF85FB2FF921BA09CBF3" authorityName="Reis" authorityYear="1892" box="[1182,1358,1752,1778]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||
<emphasis id="B96F4876FF95FF85FB2FF921BA09CBF3" box="[1182,1358,1752,1778]" italics="true" pageId="4" pageNumber="5">Saurorhynchus</emphasis>
|
||
</taxonomicName>
|
||
by several authors, and has been variously interpreted as the squamosal (
|
||
<bibRefCitation id="EF8AE995FF95FF85FC42F902BB2ACA14" author="Reis O. M." box="[1011,1133,1787,1813]" pageId="4" pageNumber="5" pagination="143 - 171" refId="ref14400" refString="Reis O. M. 1892. Zur Osteologie und Systematik der Belonorhynchiden und Tetragololepiden. Geognostische Jahreshefte 4: 143 - 171." type="journal article" year="1892">Reis 1892</bibRefCitation>
|
||
), fused suprascapularsupracleithrum (
|
||
<bibRefCitation id="EF8AE995FF95FF85FECEF8E6BD77CA3B" author="Gardiner B. G." box="[383,560,1823,1850]" pageId="4" pageNumber="5" pagination="241 - 384" refId="ref13301" refString="Gardiner B. G. 1960. A revision of certain actinopterygian and coelacanth fishes, chiefly from the Lower Lias. Bulletin of the British Museum (Natural History): Geology 4 (7): 241 - 384." type="journal article" year="1960">Gardiner 1960</bibRefCitation>
|
||
), or extrascapular (
|
||
<bibRefCitation id="EF8AE995FF95FF85FCADF8E6BCEECA38" author="Hauff B." box="[796,937,1823,1849]" pageId="4" pageNumber="5" pagination="214 - 248" refId="ref13522" refString="Hauff B. 1938. Uber Acidorhynchus aus den Posidonienschiefern von Holzmaden. Palaontologische Zeitschrift 20: 214 - 248. https: // doi. org / 10.1007 / BF 03041918" type="journal article" year="1938">Hauff 1938</bibRefCitation>
|
||
). It does not participate in the cranial midline, and extends slightly further posteriorly than the dermopterotic. We interpret this element as a lateral extrascapular, following the discussion about the homology in
|
||
<bibRefCitation id="EF8AE995FF95FF85FBBAF89EBA7DCA80" author="Kogan I. & Romano C." box="[1035,1338,1894,1921]" pageId="4" pageNumber="5" pagination="1151886" refId="ref13737" refString="Kogan I. & Romano C. 2016. Redescription of Saurichthys madagascariensis Piveteau, 1944 - 45 (Actinopterygii, Early Triassic), with implications for the early saurichthyid morphotype. Journal of Vertebrate Paleontology 36 (4): e 1151886. https: // doi. org / 10.1080 / 02724634.2016.1151886" type="journal article" year="2016">Kogan & Romano (2016)</bibRefCitation>
|
||
. The posterior elongation of the skull roof in
|
||
<taxonomicName id="4C1BEFE7FF95FF85FD10F873BC4ACAA2" baseAuthorityName="Agassiz" baseAuthorityYear="1844" box="[673,781,1930,1956]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="acutus">
|
||
<emphasis id="B96F4876FF95FF85FD10F873BC4ACAA2" box="[673,781,1930,1956]" italics="true" pageId="4" pageNumber="5">S. acutus</emphasis>
|
||
</taxonomicName>
|
||
relative to
|
||
<taxonomicName id="4C1BEFE7FF95FF85FC29F873BCBCCAA5" authority="Maxwell & Stumpf, 2017" authorityName="Maxwell & Stumpf" authorityYear="2017" box="[920,1019,1930,1956]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="hauffi" status="sp. nov.">
|
||
<emphasis id="B96F4876FF95FF85FC29F873BCBCCAA5" box="[920,1019,1930,1956]" italics="true" pageId="4" pageNumber="5">S. hauffi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25CF50DFF95FF85FBB5F873BB26CAA5" box="[1028,1121,1930,1956]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is partly accomplished through the posterior displacement of the lateral extrascapular relative to the dermopterotic (
|
||
<figureCitation id="132088E1FF95FF85FAA5F854BF91CAEA" captionStart="Fig" captionStartId="13.[189,232,1790,1816]" captionTargetBox="[189,1399,869,1754]" captionTargetId="figure@13.[189,1399,869,1754]" captionTargetPageId="13" captionText="Fig. 5. Characters distinguishing Saurorhynchus acutus (Agassiz, 1844) and S. hauffi sp. nov. A. S. acutus, narial region (SMNS 56923). B. S. acutus, lateral extrascapular-dermopterotic contact (SMNS 57039). C. S. hauffi sp. nov., narial region (SMNS 51888). D. S. hauffi sp. nov., lateral extrascapular-dermopterotic contact (SMNS 53980, mirrored). Abbreviations: dpt = dermopterotic; dh = dermohyal; f = frontal; l.ex = lateral extrascapular; m = maxilla; na-ao = nasaloantorbital; orb = orbit; pm = rostropremaxilla; pop = preopercle; pop* = damaged preopercle; ps = parasphenoid; so = supraorbital. Scale bars: A, C–D = 2 mm; B = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3829293" httpUri="https://zenodo.org/record/3829293/files/figure.png" pageId="4" pageNumber="5">Fig. 5B, D</figureCitation>
|
||
). The lateral extrascapular and the dermopterotic are fused, such that it is difficult to see the suture along parts of its length. On the ventral surface of the dermal skull roof, two ridges run longitudinally between the orbits, dividing the interorbital surface into thirds. On the lateral side of the ridges, numerous foramina are located between the ridge and the ventral surface of the frontals. Such ridges have been interpreted as part of the neurocranium in the orbitotemporal region (
|
||
<bibRefCitation id="EF8AE995FF96FF86FC45FE82BBACCC97" author="Maxwell E. E. & Diependaal H. & Winkelhorst H. & Goris G. & Klein N." box="[1012,1259,379,406]" pageId="7" pageNumber="8" pagination="119 - 134" refId="ref14125" refString="Maxwell E. E., Diependaal H., Winkelhorst H., Goris G. & Klein N. 2016. A new species of Saurichthys (Actinopterygii: Saurichthyidae) from the Middle Triassic of Winterswijk, The Netherlands. Neues Jahrbuch fur Geologie und Palaontologie Abhandlung 280 (2): 119 - 134. https: // doi. org / 10.1127 / njgpa / 2016 / 0569" type="journal article" year="2016">
|
||
Maxwell
|
||
<emphasis id="B96F4876FF96FF86FBD7FE85BBE4CC94" box="[1126,1187,379,405]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
|
||
2016
|
||
</bibRefCitation>
|
||
). The space between these ridges is filled with broken, cancellous bone tissue.
|
||
</paragraph>
|
||
<caption id="DF64C4ECFF94FF84FF0CF8BBBED2CAE8" ID-DOI="http://doi.org/10.5281/zenodo.3829284" ID-Zenodo-Dep="3829284" httpUri="https://zenodo.org/record/3829284/files/figure.png" pageId="5" pageNumber="6" startId="5.[189,232,1858,1884]" targetBox="[274,1313,297,1817]" targetPageId="5">
|
||
<paragraph id="8BA49464FF94FF84FF0CF8BBBED2CAE8" blockId="5.[189,1399,1858,2025]" pageId="5" pageNumber="6">
|
||
<emphasis id="B96F4876FF94FF84FF0CF8BBBE4BCA5D" bold="true" box="[189,268,1858,1884]" pageId="5" pageNumber="6">Fig. 1.</emphasis>
|
||
Holotype specimens of Early Jurassic saurichthyids.
|
||
<emphasis id="B96F4876FF94FF84FC39F8BBBCDACA5D" bold="true" box="[904,925,1858,1884]" pageId="5" pageNumber="6">A</emphasis>
|
||
.
|
||
<taxonomicName id="4C1BEFE7FF94FF84FC1CF8BBBA31CA5D" authority="(Agassiz, 1844)" baseAuthorityName="Agassiz" baseAuthorityYear="1844" box="[941,1398,1858,1884]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="acutus">
|
||
<emphasis id="B96F4876FF94FF84FC1CF8BBBBF4CA5D" box="[941,1203,1858,1884]" italics="true" pageId="5" pageNumber="6">Saurorhynchus acutus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8AE995FF94FF84FB75F8BBBA29CA5D" author="Agassiz L." box="[1220,1390,1858,1884]" pageId="5" pageNumber="6" refId="ref12560" refString="Agassiz L. 1844. Recherches sur les Poissons Fossiles. II. Contenant l'Histoire de l'Ordre des Ganoides. Vol. 2, Part 2. Petitpierre, Neuchatel." type="book" year="1844">Agassiz, 1844</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(NHMUK PV P 4268).
|
||
<emphasis id="B96F4876FF94FF84FE6FF89CBEB5CA7E" bold="true" box="[478,498,1893,1919]" pageId="5" pageNumber="6">B</emphasis>
|
||
.
|
||
<taxonomicName id="4C1BEFE7FF94FF84FDB2F89CBB6ACA81" authority="(Woodward, 1895)" baseAuthorityName="Woodward" baseAuthorityYear="1895" box="[515,1069,1893,1920]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="brevirostris">
|
||
<emphasis id="B96F4876FF94FF84FDB2F89CBC03CA7E" box="[515,836,1893,1919]" italics="true" pageId="5" pageNumber="6">Saurorhynchus brevirostris</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8AE995FF94FF84FCE7F89CBB62CA81" author="Woodward A. S." box="[854,1061,1893,1920]" pageId="5" pageNumber="6" refId="ref15257" refString="Woodward A. S. 1895. Catalogue of the Fossil Fishes in the British Museum (Natural History). Part III. British Museum (Natural History), London." type="book" year="1895">Woodward, 1895</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(NHMUK PV OR 40726).
|
||
<emphasis id="B96F4876FF94FF84FF0CF871BF95CAA3" bold="true" box="[189,210,1928,1954]" pageId="5" pageNumber="6">C</emphasis>
|
||
.
|
||
<taxonomicName id="4C1BEFE7FF94FF84FF51F871BD44CAA3" authority="Maxwell & Stumpf, 2017" authorityName="Maxwell & Stumpf" authorityYear="2017" box="[224,515,1928,1954]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="anningae" status="sp. nov.">
|
||
<emphasis id="B96F4876FF94FF84FF51F871BD44CAA3" box="[224,515,1928,1954]" italics="true" pageId="5" pageNumber="6">Saurorhynchus anningae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25CF50DFF94FF84FDB8F870BD24CAA2" box="[521,611,1929,1955]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(NHMUK PV P 3791).
|
||
<emphasis id="B96F4876FF94FF84FCCBF870BCF6CAA2" bold="true" box="[890,945,1929,1955]" pageId="5" pageNumber="6">D–E</emphasis>
|
||
.
|
||
<taxonomicName id="4C1BEFE7FF94FF84FC0FF871BBFFCAA3" authority="Maxwell & Stumpf, 2017" authorityName="Maxwell & Stumpf" authorityYear="2017" box="[958,1208,1928,1954]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="hauffi" status="sp. nov.">
|
||
<emphasis id="B96F4876FF94FF84FC0FF871BBFFCAA3" box="[958,1208,1928,1954]" italics="true" pageId="5" pageNumber="6">Saurorhynchus hauffi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25CF50DFF94FF84FB0FF870BA50CAA2" box="[1214,1303,1929,1955]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(SMNS 55057). Scale bars: A, C = 20 mm; E = 50 mm. Photos A–C © The Trustees of the Natural History Museum, London.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF64C4ECFF97FF87FF0CF996BE44CAE8" ID-DOI="http://doi.org/10.5281/zenodo.3829286" ID-Zenodo-Dep="3829286" httpUri="https://zenodo.org/record/3829286/files/figure.png" pageId="6" pageNumber="7" startId="6.[189,232,1647,1673]" targetBox="[205,1385,332,1597]" targetPageId="6">
|
||
<paragraph id="8BA49464FF97FF87FF0CF996BE44CAE8" blockId="6.[189,1399,1647,2025]" pageId="6" pageNumber="7">
|
||
<emphasis id="B96F4876FF97FF87FF0CF996BE52CB88" bold="true" box="[189,277,1647,1673]" pageId="6" pageNumber="7">Fig. 2.</emphasis>
|
||
Reconstruction of the dermal skull in Early Jurassic saurichthyids.
|
||
<emphasis id="B96F4876FF97FF87FB29F996BBEACB88" bold="true" box="[1176,1197,1647,1673]" pageId="6" pageNumber="7">A</emphasis>
|
||
.
|
||
<taxonomicName id="4C1BEFE7FF97FF87FB77F996BD69CBAC" authority="(Woodward, 1895)" baseAuthorityName="Woodward" baseAuthorityYear="1895" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="brevirostris">
|
||
<emphasis id="B96F4876FF97FF87FB77F996BE03CBAD" italics="true" pageId="6" pageNumber="7">Saurorhynchus brevirostris</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8AE995FF97FF87FEE5F96BBD62CBAC" author="Woodward A. S." box="[340,549,1682,1709]" pageId="6" pageNumber="7" refId="ref15257" refString="Woodward A. S. 1895. Catalogue of the Fossil Fishes in the British Museum (Natural History). Part III. British Museum (Natural History), London." type="book" year="1895">Woodward, 1895</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B96F4876FF97FF87FD8AF96BBD08CBAD" bold="true" box="[571,591,1682,1708]" pageId="6" pageNumber="7">B</emphasis>
|
||
.
|
||
<taxonomicName id="4C1BEFE7FF97FF87FDEFF96BBCC3CBAD" authority="Maxwell & Stumpf, 2017" authorityName="Maxwell & Stumpf" authorityYear="2017" box="[606,900,1682,1708]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="anningae" status="sp. nov.">
|
||
<emphasis id="B96F4876FF97FF87FDEFF96BBCC3CBAD" box="[606,900,1682,1708]" italics="true" pageId="6" pageNumber="7">Saurorhynchus anningae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25CF50DFF97FF87FC3CF96ABCAECBAC" box="[909,1001,1683,1709]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="B96F4876FF97FF87FC40F96BBB41CBAD" bold="true" box="[1009,1030,1682,1708]" pageId="6" pageNumber="7">C</emphasis>
|
||
.
|
||
<taxonomicName id="4C1BEFE7FF97FF87FBA6F96BBA55CBAD" authority="Maxwell & Stumpf, 2017" authorityName="Maxwell & Stumpf" authorityYear="2017" box="[1047,1298,1682,1708]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="hauffi" status="sp. nov.">
|
||
<emphasis id="B96F4876FF97FF87FBA6F96BBA55CBAD" box="[1047,1298,1682,1708]" italics="true" pageId="6" pageNumber="7">Saurorhynchus hauffi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25CF50DFF97FF87FAAAF96ABA30CBAC" box="[1307,1399,1683,1709]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="B96F4876FF97FF87FF0CF94CBF95CBCE" bold="true" box="[189,210,1717,1743]" pageId="6" pageNumber="7">D</emphasis>
|
||
.
|
||
<taxonomicName id="4C1BEFE7FF97FF87FF54F94CBDFCCBCE" authority="(Agassiz, 1844)" baseAuthorityName="Agassiz" baseAuthorityYear="1844" box="[229,699,1717,1743]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="acutus">
|
||
<emphasis id="B96F4876FF97FF87FF54F94CBEA9CBCE" box="[229,494,1717,1743]" italics="true" pageId="6" pageNumber="7">Saurorhynchus acutus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8AE995FF97FF87FDB0F94CBDF5CBCE" author="Agassiz L." box="[513,690,1717,1743]" pageId="6" pageNumber="7" refId="ref12560" refString="Agassiz L. 1844. Recherches sur les Poissons Fossiles. II. Contenant l'Histoire de l'Ordre des Ganoides. Vol. 2, Part 2. Petitpierre, Neuchatel." type="book" year="1844">Agassiz, 1844</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Not to scale. Dashed lines indicate sensory canals, grey lines indicate bones that were present but where the exact location of the sutural contact is unclear, and question marks indicate areas of uncertainty. Abbreviations: ang = angular; bs = basisphenoid; cl = cleithrum; d = dentary; dh = dermohyal; dpt = dermopterotic; dsp = dermophenotic; f = frontal; io = infraorbital; io.c = infraorbital sensory canal; l = lacrimal; l.ex = lateral extrascapular; m = maxilla; md.c = mandibular sensory canal; mio.c = medial branch of the infraorbital sensory canal; n = neomorph; na-ao = nasaloantorbital; op = opercle; p = parietal; pm = rostropremaxilla; pop = preopercle; ps = parasphenoid; sa = supraangular; scl = supracleithrum; so = supraorbital; so.c = supraorbital sensory canal.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA49464FF96FF86FF0CFE1DBB47CE02" blockId="7.[189,1399,484,772]" pageId="7" pageNumber="8">The palate is exposed in a single specimen (SMNS 50924). The dermopalatine extends as far anteriorly as the posterior external narial opening. It is convex in ventral view, broadening in a medial direction. It bears denticles; these are smaller and blunter than those of the other tooth-bearing elements. Anteriorly, it articulates with the maxilla laterally; posteriorly it articulates with the parasphenoid medially and, posterior to this, the entopterygoid. There is no lateral palatal foramen, but because the parasphenoid is not anteriorly expanded, a medial foramen exists between the anterior dermopalatine and parasphenoid. The ectopterygoid is very small and triangular. The portion of the parasphenoid posterior to the anterior edge of the orbit is edentulous; more anteriorly it bears small denticles.</paragraph>
|
||
<paragraph id="8BA49464FF96FF86FF0CFCD4BDF0CEB7" blockId="7.[189,1399,813,950]" pageId="7" pageNumber="8">The anteriormost point of the angular is located on the ventral half of the lower jaw. The ventral part of the angular-dentary suture is posteroventrally angled and straight, forming a simple ‘v’. The teeth are largest at the midpoint of the rostrum and become much smaller anteriorly. In no specimens was corrugation of the collar ganoine observed.</paragraph>
|
||
<paragraph id="8BA49464FF96FF86FF0CFC19BC50C944" blockId="7.[189,1399,992,1093]" pageId="7" pageNumber="8">
|
||
The opercle is preserved in a single specimen (PMU 30010). This element is
|
||
<quantity id="4CE33981FF96FF86FB86FC19BBD8CEFA" box="[1079,1183,992,1019]" metricMagnitude="-2" metricUnit="m" metricValue="1.3599999999999999" pageId="7" pageNumber="8" unit="mm" value="13.6">13.6 mm</quantity>
|
||
high and
|
||
<quantity id="4CE33981FF96FF86FABEFC19BA30CEFA" box="[1295,1399,992,1019]" metricMagnitude="-2" metricUnit="m" metricValue="1.39" pageId="7" pageNumber="8" unit="mm" value="13.9">13.9 mm</quantity>
|
||
long, and ornamented with elongate pits radiating from the palatoquadrate articulation. In overall shape it is more ovate than in
|
||
<taxonomicName id="4C1BEFE7FF96FF86FE63FBD2BC57C944" baseAuthorityName="Woodward" baseAuthorityYear="1895" box="[466,784,1067,1093]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="brevirostris">
|
||
<emphasis id="B96F4876FF96FF86FE63FBD2BC57C944" box="[466,784,1067,1093]" italics="true" pageId="7" pageNumber="8">Saurorhynchus brevirostris</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C301C7EFFF96FF86FF0CFB97BBF9CA2D" pageId="7" pageNumber="8" type="discussion">
|
||
<paragraph id="8BA49464FF96FF86FF0CFB97BE68C989" blockId="7.[189,303,1134,1160]" box="[189,303,1134,1160]" pageId="7" pageNumber="8">
|
||
<heading id="D0EC2308FF96FF86FF0CFB97BE68C989" bold="true" box="[189,303,1134,1160]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
|
||
<emphasis id="B96F4876FF96FF86FF0CFB97BE68C989" bold="true" box="[189,303,1134,1160]" pageId="7" pageNumber="8">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF96FF86FF0CFB66BD6AC875" blockId="7.[189,1399,1183,1396]" pageId="7" pageNumber="8">
|
||
<bibRefCitation id="EF8AE995FF96FF86FF0CFB66BED2C9BB" author="Woodward A. S." box="[189,405,1183,1210]" pageId="7" pageNumber="8" refId="ref15257" refString="Woodward A. S. 1895. Catalogue of the Fossil Fishes in the British Museum (Natural History). Part III. British Museum (Natural History), London." type="book" year="1895">Woodward (1895)</bibRefCitation>
|
||
outlined a series of characters to diagnose
|
||
<taxonomicName id="4C1BEFE7FF96FF86FC2FFB66BB4CC9B8" baseAuthorityName="Agassiz" baseAuthorityYear="1844" box="[926,1035,1183,1209]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="acutus">
|
||
<emphasis id="B96F4876FF96FF86FC2FFB66BB4CC9B8" box="[926,1035,1183,1209]" italics="true" pageId="7" pageNumber="8">S. acutus</emphasis>
|
||
</taxonomicName>
|
||
, the majority of which cannot be observed in the
|
||
<typeStatus id="54A02AC6FF96FF86FE11FB3CBE94C9DE" box="[416,467,1221,1247]" pageId="7" pageNumber="8">type</typeStatus>
|
||
material and were drawn from observation of specimens from the much older Charmouth Mudstone Formation. Upon re-examination of the
|
||
<typeStatus id="54A02AC6FF96FF86FC24FB13BC8FC805" box="[917,968,1258,1284]" pageId="7" pageNumber="8">type</typeStatus>
|
||
material,
|
||
<bibRefCitation id="EF8AE995FF96FF86FB8AFB10BA48C805" author="Woodward A. S." box="[1083,1295,1257,1284]" pageId="7" pageNumber="8" pagination="455 - 472" refId="ref15290" refString="Woodward A. S. 1899. On the fossil fishes of the Upper Lias of Whitby. Part IV. Proceedings of the Yorkshire Geological and Polytechnic Society 13: 455 - 472." type="journal article" year="1899">Woodward (1899)</bibRefCitation>
|
||
clarified that the posterior (backwards) extension of the skull roof was the main feature uniting the Toarcian
|
||
<typeStatus id="54A02AC6FF96FF86FAF5FAF6BA30C828" box="[1348,1399,1295,1321]" pageId="7" pageNumber="8">type</typeStatus>
|
||
and Sinemurian referred specimens, even though he had reservations that a species was likely to have such a long stratigraphic range.
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF96FF86FF0CFA64BBF9CA2D" blockId="7.[189,1399,1437,1836]" pageId="7" pageNumber="8">
|
||
Our observations suggest that the type specimen of
|
||
<taxonomicName id="4C1BEFE7FF96FF86FCA8FA64BCC4C8B6" baseAuthorityName="Agassiz" baseAuthorityYear="1844" box="[793,899,1437,1463]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="acutus">
|
||
<emphasis id="B96F4876FF96FF86FCA8FA64BCC4C8B6" box="[793,899,1437,1463]" italics="true" pageId="7" pageNumber="8">S. acutus</emphasis>
|
||
</taxonomicName>
|
||
can be unambiguously differentiated from the older Sinemurian material by the short distance between the posterior edge of the orbit and the articular facet for the hyomandibula relative to the length of the orbit. These measurements fall along a single anteroposterior line, and thus are not expected to be significantly influenced by distortion. In addition, the narial opening in the
|
||
<typeStatus id="54A02AC6FF96FF86FDE0F9CBBDFFCB4D" box="[593,696,1586,1612]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
|
||
specimen of
|
||
<taxonomicName id="4C1BEFE7FF96FF86FCE4F9CABCF8CB4D" baseAuthorityName="Agassiz" baseAuthorityYear="1844" box="[853,959,1587,1613]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="acutus">
|
||
<emphasis id="B96F4876FF96FF86FCE4F9CABCF8CB4D" box="[853,959,1587,1613]" italics="true" pageId="7" pageNumber="8">S. acutus</emphasis>
|
||
</taxonomicName>
|
||
is narrower than in the material from the Charmouth Mudstone. The
|
||
<typeStatus id="54A02AC6FF96FF86FD81F9A1BDCCCB73" box="[560,651,1624,1650]" pageId="7" pageNumber="8" type="syntype">syntype</typeStatus>
|
||
specimen figured by
|
||
<bibRefCitation id="EF8AE995FF96FF86FC3AF9AEBB06CB73" author="Agassiz L." box="[907,1089,1623,1650]" pageId="7" pageNumber="8" refId="ref12560" refString="Agassiz L. 1844. Recherches sur les Poissons Fossiles. II. Contenant l'Histoire de l'Ordre des Ganoides. Vol. 2, Part 2. Petitpierre, Neuchatel." type="book" year="1844">Agassiz (1844)</bibRefCitation>
|
||
, NHMUK PV P 961a and its (unfigured) counterslab NHMUK PV P 36219 are non-diagnostic below family level. The
|
||
<typeStatus id="54A02AC6FF96FF86FAA1F984BA30CB96" box="[1296,1399,1661,1687]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
|
||
is consistent with a large number of specimens of
|
||
<taxonomicName id="4C1BEFE7FF96FF86FCA9F95BBC8FCBBD" authorityName="Reis" authorityYear="1892" box="[792,968,1698,1724]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">
|
||
<emphasis id="B96F4876FF96FF86FCA9F95BBC8FCBBD" box="[792,968,1698,1724]" italics="true" pageId="7" pageNumber="8">Saurorhynchus</emphasis>
|
||
</taxonomicName>
|
||
from the Toarcian of
|
||
<collectingCountry id="F30CD4F4FF96FF86FB61F95BBA79CBBD" box="[1232,1342,1698,1724]" name="Germany" pageId="7" pageNumber="8">Germany</collectingCountry>
|
||
, and when the additional anatomical information present in this referred material is considered it becomes clear that large differences exist between the Sinemurian and Toarcian specimens. The revised diagnosis is constructed based on the referred material from
|
||
<collectingCountry id="F30CD4F4FF96FF86FCBDF8EBBC3DCA2D" box="[780,890,1810,1836]" name="Germany" pageId="7" pageNumber="8">Germany</collectingCountry>
|
||
, as well as on the
|
||
<typeStatus id="54A02AC6FF96FF86FBFEF8EBBBFDCA2D" box="[1103,1210,1810,1836]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C301C7EFFF96FF86FF0CF8ACBEDACAED" pageId="7" pageNumber="8" type="distribution">
|
||
<paragraph id="8BA49464FF96FF86FF0CF8ACBE08CA6E" blockId="7.[189,335,1877,1903]" box="[189,335,1877,1903]" pageId="7" pageNumber="8">
|
||
<heading id="D0EC2308FF96FF86FF0CF8ACBE08CA6E" bold="true" box="[189,335,1877,1903]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
|
||
<emphasis id="B96F4876FF96FF86FF0CF8ACBE08CA6E" bold="true" box="[189,335,1877,1903]" pageId="7" pageNumber="8">Occurrence</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF96FF86FF0CF87FBEDACAED" blockId="7.[189,1398,1926,2028]" pageId="7" pageNumber="8">
|
||
Early Jurassic, Toarcian, Whitby (
|
||
<typeStatus id="54A02AC6FF96FF86FDE2F87FBDC0CAA1" box="[595,647,1926,1952]" pageId="7" pageNumber="8">type</typeStatus>
|
||
locality); Toarcian,
|
||
<taxonomicName id="4C1BEFE7FF96FF86FCCBF87EBB20CAA0" authority="Zone" authorityName="Zone" box="[890,1127,1927,1953]" class="Cephalopoda" family="Dactylioceratidae" genus="Dactylioceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="tenuicostatum">
|
||
<emphasis id="B96F4876FF96FF86FCCBF87EBB58CAA1" box="[890,1055,1927,1952]" italics="true" pageId="7" pageNumber="8">tenuicostatum</emphasis>
|
||
Zone
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1BEFE7FF96FF86FBC4F87FBA31CAA1" authority="Subzone" authorityName="Subzone" box="[1141,1398,1926,1952]" class="Cephalopoda" family="Dactylioceratidae" genus="Dactylioceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="semicelatum">
|
||
<emphasis id="B96F4876FF96FF86FBC4F87FBA4FCAA1" box="[1141,1288,1926,1952]" italics="true" pageId="7" pageNumber="8">semicelatum</emphasis>
|
||
Subzone
|
||
</taxonomicName>
|
||
to
|
||
<emphasis id="B96F4876FF96FF86FF50F855BE35CAC4" box="[225,370,1964,1989]" italics="true" pageId="7" pageNumber="8">serpentinum</emphasis>
|
||
Zone,
|
||
<collectingRegion id="49DF5A86FF96FF86FE7EF852BD84CAC4" box="[463,707,1963,1989]" country="Germany" name="Baden-Wuerttemberg" pageId="7" pageNumber="8">Baden-Württemberg</collectingRegion>
|
||
,
|
||
<collectingCountry id="F30CD4F4FF96FF86FD65F855BC02CAC7" box="[724,837,1964,1990]" name="Germany" pageId="7" pageNumber="8">Germany</collectingCountry>
|
||
;
|
||
<emphasis id="B96F4876FF96FF86FCE6F855BCAFCAC4" box="[855,1000,1964,1989]" italics="true" pageId="7" pageNumber="8">serpentinum</emphasis>
|
||
Zone,
|
||
<collectingRegion id="49DF5A86FF96FF86FBF4F855BBBFCAC7" box="[1093,1272,1964,1990]" country="Germany" name="Niedersachsen" pageId="7" pageNumber="8">Lower Saxony</collectingRegion>
|
||
,
|
||
<collectingCountry id="F30CD4F4FF96FF86FAB8F855BA31CAC7" box="[1289,1398,1964,1990]" name="Germany" pageId="7" pageNumber="8">Germany</collectingCountry>
|
||
(referred material).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |