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<document id="F81E088A7A06002F5CDD6DAD2C98CAFD" ID-DOI="10.5281/zenodo.35388" ID-GBIF-Dataset="b184dfcc-9b5f-4abb-b401-0f45f76006f6" checkinTime="1450029366780" checkinUser="donat" docAuthor="Dijkstra, Klaas-Douwe B., Kipping, Jens &amp; Mézière, Nicolas" docDate="2015" docId="03A25264CADEFF20EE9AFD5F40CBFCDF" docLanguage="en" docName="Odonatologica_44-4-low_res.pdf.imf" docOrigin="Odonatologica 44 (4)" docTitle="Porpax mezierei Dijkstra &amp; Kipping, sp. nov." docType="treatment" docVersion="24" lastPageNumber="647" masterDocId="FF9B2A1CCA19FFEAEE22FFED4105FFB2" masterDocTitle="Sixty new dragonfly and damselfly species from Africa (Odonata)" masterLastPageNumber="678" masterPageNumber="447" pageNumber="644" updateTime="1695001031402" updateUser="admin">
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<mods:title id="4A6F7C67EB65D7AD25D4714916D8BB8F">Sixty new dragonfly and damselfly species from Africa (Odonata)</mods:title>
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<mods:namePart id="E8E5D206DB76367949D2E4239215B890">Dijkstra, Klaas-Douwe B.</mods:namePart>
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<mods:namePart id="61D1D2B2F01FC9A54DA798EB782D3BD4">Kipping, Jens</mods:namePart>
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<mods:namePart id="BD5CB543B8B0E795F105110F295472D6">Mézière, Nicolas</mods:namePart>
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<mods:title id="76C605676FFC8CA4ED1B39DACFB00059">Odonatologica</mods:title>
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<mods:part id="6EB02F32DD92F4FB6D9E059C5192148E">
<mods:date id="2ECC940A58C8FDC63FC7CC4C4005F204">2015</mods:date>
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<treatment id="03A25264CADEFF20EE9AFD5F40CBFCDF" ID-DOI="http://doi.org/10.5281/zenodo.5640300" ID-GBIF-Taxon="114923465" ID-Zenodo-Dep="5640300" LSID="urn:lsid:zoobank.org:act:03A25264-CADE-FF20-EE9A-FD5F40CBFCDF" httpUri="http://treatment.plazi.org/id/03A25264CADEFF20EE9AFD5F40CBFCDF" lastPageId="202" lastPageNumber="647" pageId="199" pageNumber="644">
<subSubSection id="C311B0F9CADEFF2DEE9AFD5F4260FD45" pageId="199" pageNumber="644" type="nomenclature">
<paragraph id="8BB4E372CADEFF2DEE9AFD5F4260FD45" pageId="199" pageNumber="644">
<heading id="D0FC541ECADEFF2DEE9AFD5F4260FD45" bold="true" centered="true" fontSize="11" level="2" pageId="199" pageNumber="644" reason="4">
<emphasis id="B97F3F60CADEFF2DEE9AFD5F42F7FD7F" bold="true" box="[184,1010,689,717]" pageId="199" pageNumber="644">
<taxonomicName id="4C0B98F1CADEFF2DEE9AFD5F4359FD7F" LSID-ZBK="urn:lsid:zoobank.org:act:03A25264-CADE-FF20-EE9A-FD5F40CBFCDF" authority="Dijkstra &amp; Kipping" authorityName="Dijkstra &amp; Kipping" box="[184,604,689,717]" class="Insecta" family="Libellulidae" genus="Porpax" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="199" pageNumber="644" phylum="Arthropoda" rank="species" species="mezierei" status="sp. nov.">
<emphasis id="B97F3F60CADEFF2DEE9AFD5F407EFD7E" bold="true" box="[184,379,690,716]" italics="true" pageId="199" pageNumber="644">Porpax mezierei</emphasis>
Dijkstra &amp; Kipping
</taxonomicName>
<taxonomicNameLabel id="A24C821BCADEFF2DEC41FD5F43C5FD7E" box="[611,704,690,716]" pageId="199" pageNumber="644" rank="species">sp. nov.</taxonomicNameLabel>
Blue-spotted Pricklyleg
</emphasis>
(
<typeStatus id="54B05DD0CADEFF2DEF69FD314083FD45" box="[331,390,732,759]" pageId="199" pageNumber="644">Type</typeStatus>
<figureCitation id="1330FFF7CADEFF2DEFAFFD3140F9FD45" box="[397,508,732,759]" captionStart="Photo 54" captionStartId="157.[159,225,1382,1406]" captionText="Photo 54. Small forest stream south of Bentor, Liberia. Type locality of Paragomphus lemperti. Photo: KDBD (26 - ii- - 2012)" pageId="199" pageNumber="644">Photo 54</figureCitation>
,
<figureCitation id="1330FFF7CADEFF2DEC24FD314384FD45" box="[518,641,732,759]" captionStart="Photo 52" captionStartId="150.[159,225,1147,1172]" captionText="Photo 52. Boubou Stream near Bongoville at the foot of the Batéké Plateau, Gabon. Habitat of Africocypha varicolor, Elattoneura aurifex, Ceriagrion obfuscans, Pseudagrion aureolum, Paragomphus darwalli, Malgassophlebia andzaba, Neodythemis infra, Porpax mezierei, and Trithemis hinnula. Photo: NM (20 - xi- 2011)" pageId="199" pageNumber="644">Photos 52</figureCitation>
,
<figureCitation id="1330FFF7CADEFF2DECA9FD3143A8FD45" box="[651,685,732,759]" captionStart="Photo 62" captionStartId="179.[781,848,1237,1261]" captionText="Photo 62. Andzaba Stream near Bongoville at the foot of the Batéké Plateau, Gabon. Habitat of Malgassophlebia andzaba, Elattoneura aurifex, Pseudagrion aureolum, and Lestinogomphus venustus. Photo: NM (28 - iii- 2010)" pageId="199" pageNumber="644">62</figureCitation>
,
<figureCitation id="1330FFF7CADEFF2DEC95FD314200FD45" box="[695,773,732,759]" captionStart-0="Photo 69" captionStart-1="Photo 70" captionStartId-0="201.[159,225,1527,1551]" captionStartId-1="202.[159,225,1381,1405]" captionTargetBox-0="[158,1035,766,1485]" captionTargetId-0="figure@201.[156,1038,763,1488]" captionTargetPageId-0="201" captionText-0="Photo 69. Porpax mezierei, male; Ekala, Gabon. Photo: NM (06 - xi- 2010)" captionText-1="Photo 70. Forest swamp at Ekala near Bongoville, Gabon. Type locality of Neodythemis infra, Porpax mezierei, and Trithemis hinnula with their discoverer Nicolas Mézière. Photo: JK (27 - i- 2012)" pageId="199" pageNumber="644">6970</figureCitation>
,
<figureCitation id="1330FFF7CADEFF2DED2DFD314258FD45" box="[783,861,732,759]" captionStart="Figure 31" captionStartId="189.[784,852,1410,1435]" captionTargetBox="[159,745,910,1550]" captionTargetId="figure@189.[156,749,904,1552]" captionTargetPageId="189" captionText="Figure 31. Secondary genitalia in lateral view of the saegeri - group of Orthetrum and three similar species." pageId="199" pageNumber="644">Fig. 31</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CADEFF2DEEBDFCE6401AFC96" box="[159,287,779,804]" pageId="199" pageNumber="644" type="description">
<paragraph id="8BB4E372CADEFF2DEEBDFCE6401AFC96" box="[159,287,779,804]" pageId="199" pageNumber="644">
<heading id="D0FC541ECADEFF2DEEBDFCE6401AFC96" bold="true" box="[159,287,779,804]" fontSize="11" level="3" pageId="199" pageNumber="644" reason="6">
<emphasis id="B97F3F60CADEFF2DEEBDFCE6401AFC96" bold="true" box="[159,287,779,804]" pageId="199" pageNumber="644">Taxonomy</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CADEFF2DEEBDFCC24209FBC6" pageId="199" pageNumber="644" type="discussion">
<paragraph id="8BB4E372CADEFF2DEEBDFCC24209FBC6" pageId="199" pageNumber="644">
<bibRefCitation id="7D106BD2E2BE0C31434AD5771C223832" author="Dijkstra K. - D. B." journalOrPublisher="Tijdschrift voor Entomologie" pagination="71 - 88" part="149" refId="ref66812" refString="Dijkstra K. - D. B. 2006. Taxonomy and biogeography of Porpax, a dragonfly genus centred in the Congo Basin (Odonata, Libellulidae). Tijdschrift voor Entomologie 149: 71 - 88" title="Taxonomy and biogeography of Porpax, a dragonfly genus centred in the Congo Basin (Odonata, Libellulidae)" type="journal article" year="2006">Dijkstra (2006)</bibRefCitation>
revised the genus
<taxonomicName id="4C0B98F1CADEFF2DEC16FCDD4221FCF8" authority="Karsch, 1896" authorityName="Karsch" authorityYear="1896" box="[564,804,815,842]" class="Insecta" family="Libellulidae" genus="Porpax" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="199" pageNumber="644" phylum="Arthropoda" rank="genus">
<emphasis id="B97F3F60CADEFF2DEC16FCDD4383FCF8" box="[564,646,816,842]" italics="true" pageId="199" pageNumber="644">Porpax</emphasis>
Karsch, 1896
</taxonomicName>
, separating five species principally on the basis of male markings and hind femur ornamentation.
<taxonomicName id="4C0B98F1CADEFF2DEEFEFC964364FC26" authority="Pinhey, 1966" authorityName="Pinhey" authorityYear="1966" box="[220,609,890,916]" class="Insecta" family="Libellulidae" genus="Porpax" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="199" pageNumber="644" phylum="Arthropoda" rank="species" species="garambensis">
<emphasis id="B97F3F60CADEFF2DEEFEFC9640C1FC26" box="[220,452,890,916]" italics="true" pageId="199" pageNumber="644">Porpax garambensis</emphasis>
Pinhey, 1966
</taxonomicName>
was the only species to differ also in the structure of male and female genitalia. The current taxon is morphologically and genetically close to that species but overlaps geographically, differs genetically, and has unique abdominal markings: while other
<taxonomicName id="4C0B98F1CADEFF2DED5DFC0642D4FBB6" box="[895,977,1003,1028]" class="Insecta" family="Libellulidae" genus="Porpax" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="199" pageNumber="644" phylum="Arthropoda" rank="genus">
<emphasis id="B97F3F60CADEFF2DED5DFC0642D4FBB6" box="[895,977,1003,1028]" italics="true" pageId="199" pageNumber="644">Porpax</emphasis>
</taxonomicName>
species are marked with greenish blue, it and the closely related
<taxonomicName id="4C0B98F1CADEFF2DED55FBE24070FBFD" authority="Ris, 1915" authorityName="Ris" authorityYear="1915" class="Insecta" family="Libellulidae" genus="Cyanothemis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="199" pageNumber="644" phylum="Arthropoda" rank="species" species="simpsoni">
<emphasis id="B97F3F60CADEFF2DED55FBE24006FBFD" italics="true" pageId="199" pageNumber="644">Cyanothemis simpsoni</emphasis>
Ris, 1915
</taxonomicName>
seem exceptional among nearly 1 500 libelluloid species for attaining deep blue coloration without pruinosity.
</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CADEFF22EEBDFB4842EEFDF0" lastPageId="200" lastPageNumber="645" pageId="199" pageNumber="644" type="materials_examined">
<paragraph id="8BB4E372CADEFF2DEEBDFB48406DFB0D" box="[159,360,1189,1215]" pageId="199" pageNumber="644">
<heading id="D0FC541ECADEFF2DEEBDFB48406DFB0D" bold="true" box="[159,360,1189,1215]" fontSize="11" level="3" pageId="199" pageNumber="644" reason="6">
<emphasis id="B97F3F60CADEFF2DEEBDFB48406DFB0D" bold="true" box="[159,360,1189,1215]" pageId="199" pageNumber="644">Material studied</emphasis>
</heading>
</paragraph>
<paragraph id="8BB4E372CADEFF2DEEBDFB264372FAE6" pageId="199" pageNumber="644">
<materialsCitation id="3B63E92FCADEFF2DEEBDFB264376FAE6" ID-GBIF-Occurrence="3396361307" collectingDate="2013-05-19" collectionCode="RMNH" collectorName="N. Mézière, J. Renoult &amp;, J. Lekogo" country="Gabon" elevation="393 m" httpUri="http://www.boldsystems.org/index.php/Public_RecordView?processid=NSPAF517-15" latitude="-1.6068" location="Bongoville-Léconi road after Ekala, old village of Boubou" longLatPrecision="7" longitude="13.9137" pageId="199" pageNumber="644" specimenCode="RMNH.INS.554425" specimenCount="1" specimenCount-male="1" stateProvince="Haut-Ogooué Province" typeStatus="holotype">
<typeStatus id="54B05DD0CADEFF2DEEBDFB264015FB57" box="[159,272,1227,1253]" pageId="199" pageNumber="644">
<emphasis id="B97F3F60CADEFF2DEEBDFB264015FB57" bold="true" box="[159,272,1227,1253]" pageId="199" pageNumber="644">Holotype</emphasis>
</typeStatus>
<emphasis id="B97F3F60CADEFF2DEF3BFB274028FB51" bold="true" box="[281,301,1226,1251]" pageId="199" pageNumber="644">
<specimenCount id="4CE97C2DDA0318252B7885BB2BFF0268" type="male"></specimenCount>
</emphasis>
<emphasis id="B97F3F60CADEFF2DEF0FFB264030FB56" bold="true" box="[301,309,1227,1252]" pageId="199" pageNumber="644">.</emphasis>
<specimenCode id="BB5FA8C9229AAE58E938A39339B86CA6">RMNH.INS.508827</specimenCode>
,
<collectingCountry id="F31CA3E2CADEFF2DEC16FB274383FB56" box="[564,646,1226,1252]" name="Gabon" pageId="199" pageNumber="644" value="Gabon">Gabon</collectingCountry>
,
<collectingRegion id="49CF2D90CADEFF2DECB1FB2442A2FB56" box="[659,935,1225,1252]" pageId="199" pageNumber="644">Haut-Ogooué Province</collectingRegion>
,
<location id="F6AD506173D0B46DE0C064E1B693AC4E" LSID="urn:lsid:plazi:treatment:03A25264CADEFF20EE9AFD5F40CBFCDF:F6AD506173D0B46DE0C064E1B693AC4E" country="Gabon" latitude="-1.6068" longLatPrecision="7" longitude="13.9137" name="Bongoville-Léconi road after Ekala, old village of Boubou" stateProvince="Haut-Ogooué Province">Bongoville-Léconi road after Ekala, old village of Boubou</location>
, ponds and humid forest near stream (
<figureCitation id="1330FFF7CADEFF2DEF14FAF940A6FA9D" box="[310,419,1300,1327]" captionStart="Photo 70" captionStartId="202.[159,225,1381,1405]" captionText="Photo 70. Forest swamp at Ekala near Bongoville, Gabon. Type locality of Neodythemis infra, Porpax mezierei, and Trithemis hinnula with their discoverer Nicolas Mézière. Photo: JK (27 - i- 2012)" pageId="199" pageNumber="644">Photo 70</figureCitation>
),
<quantity id="4CF34E97CADEFF2DEF94FAF840FBFA9D" box="[438,510,1301,1327]" metricMagnitude="2" metricUnit="m" metricValue="3.9299999999999997" pageId="199" pageNumber="644" unit="m" value="393.0">
<elevation id="7827ECED56F2D59F8E27023AB390B8AC" box="[438,510,1301,1327]" metricMagnitude="2" metricUnit="m" metricValue="3.9299999999999997" pageId="199" pageNumber="644" unit="m" value="393.0">393 m</elevation>
</quantity>
a.s.l. (
<geoCoordinate id="EE3F85B5CADEFF2DEC67FAF943A9FA9D" box="[581,684,1300,1327]" direction="south" orientation="latitude" pageId="199" pageNumber="644" precision="5" value="-1.6068">1.6068 ° S</geoCoordinate>
<geoCoordinate id="EE3F85B5CADEFF2DEC93FAF9422EFA9D" box="[689,811,1300,1327]" direction="east" orientation="longitude" pageId="199" pageNumber="644" precision="5" value="13.9137">13.9137 ° E</geoCoordinate>
),
<date id="FFB5C5B2CADEFF2DED1CFAF842B2FA9D" box="[830,951,1301,1327]" pageId="199" pageNumber="644" value="2013-05-19">
<collectingDate id="BB8A37A8D3E67A7592AE8C31C1A13579" box="[830,951,1301,1327]" pageId="199" pageNumber="644" value="2013-05-19">19 -v- 2013</collectingDate>
</date>
, leg.
<collectorName id="80CC7EABB90AC86F3DE408DF5C7E8F08">N. Mézière</collectorName>
,
<collectorName id="6DEEDDA3CE3F2205015042FFF32ECD55">J. Renoult &amp;</collectorName>
<collectorName id="F5AF4691282FE3F85389D7A063467ADC">J. Lekogo</collectorName>
,
<collectionCode id="ED1A7BB7CADEFF2DEC35FAD74376FAE6" box="[535,627,1338,1364]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" pageId="199" pageNumber="644">RMNH</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB4E372CADEFF2DEEBDFA8D4509F9BD" pageId="199" pageNumber="644">
<emphasis id="B97F3F60CADEFF2DEEBDFA8D4087FAC8" bold="true" box="[159,386,1376,1402]" pageId="199" pageNumber="644">Further material.</emphasis>
<materialsCitation id="3B63E92FCADEFF2DEFBAFAB2433CFA2D" ID-GBIF-Occurrence="1224548093" collectingDate="2013-05-19" collectionCode="RMNH" collectorName="N. Mézière, J. Renoult &amp;, J. Lekogo" country="Gabon" elevation="393 m" latitude="-1.6068" location="Bongoville-Léconi road after Ekala, old village of Boubou" longLatPrecision="7" longitude="13.9137" pageId="199" pageNumber="644" specimenCode="RMNH.INS.554425" specimenCount="1" specimenCount-male="1" stateProvince="Haut-Ogooué Province">
<collectingCountry id="F31CA3E2CADEFF2DEFBAFAB24305FAC8" box="[408,512,1375,1402]" name="Gabon" pageId="199" pageNumber="644" value="Gabon">GABON</collectingCountry>
(
<collectingRegion id="49CF2D90CADEFF2DEC3DFAB34243FAC8" box="[543,838,1374,1402]" pageId="199" pageNumber="644">Haut-Ogooué Province</collectingRegion>
):
<specimenCount id="9D0D28FBCADEFF2DED4BFAB2428EFACB" box="[873,907,1375,1402]" pageId="199" pageNumber="644" type="generic">1 ♂</specimenCount>
(
<specimenCode id="A84987AC10BF5CF9C7BB4F65FBD53C50">RMNH.INS.508823</specimenCode>
), as holotype,
<collectionCode id="ED1A7BB7CADEFF2DEFFFFA68433CFA2D" box="[477,569,1413,1439]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" pageId="199" pageNumber="644">RMNH</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B63E92FCADEFF2DEC64FA684060FA76" ID-GBIF-Occurrence="1224548086" collectingDate="2010-12-28" collectionCode="RMNH" collectorName="N. Mézière" country="Gabon" elevation="393 m" latitude="-1.6068" location="Bongoville-Léconi road after Ekala, old village of Boubou" longLatPrecision="7" longitude="13.9137" pageId="199" pageNumber="644" specimenCount="3" specimenCount-male="3" stateProvince="Haut-ogooue Province">
<specimenCount id="9D0D28FBCADEFF2DEC64FA68436DFA2C" box="[582,616,1412,1439]" pageId="199" pageNumber="644" type="generic">3 ♂</specimenCount>
, same locality,
<date id="FFB5C5B2CADEFF2DED06FA6942A8FA2D" box="[804,941,1412,1439]" pageId="199" pageNumber="644" value="2010-12-28">
<collectingDate id="5309BB4739BD3CA75EAC505D5D09F61A" box="[804,941,1412,1439]" pageId="199" pageNumber="644" value="2010-12-28">28 -xii- 2010</collectingDate>
</date>
, leg.
<collectorName id="93F6CED62EBBF6DA562E3573C98A9CB6">N. Mézière</collectorName>
,
<collectionCode id="ED1A7BB7CADEFF2DEF2BFA474060FA76" box="[265,357,1450,1476]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" pageId="199" pageNumber="644">RMNH</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B63E92FCADEFF2DEF4DFA474060F9BD" ID-GBIF-Occurrence="1224548101" collectingDate="2010-12-30" collectionCode="RMNH" collectorName="N. Mézière" country="Gabon" elevation="421 m" latitude="-1.6116" location="Bongoville-Léconi road, Boubou River, Ekala" longLatPrecision="7" longitude="13.9116" pageId="199" pageNumber="644" specimenCount="3" specimenCount-male="3" stateProvince="Haut-ogooue Province">
<specimenCount id="9D0D28FBCADEFF2DEF4DFA474094FA71" box="[367,401,1450,1476]" pageId="199" pageNumber="644" type="generic">3 ♂</specimenCount>
,
<location id="C6530E043B26D847FC05986ABE860CDC" LSID="urn:lsid:plazi:treatment:03A25264CADEFF20EE9AFD5F40CBFCDF:C6530E043B26D847FC05986ABE860CDC" country="Gabon" latitude="-1.6116" longLatPrecision="7" longitude="13.9116" name="Bongoville-Léconi road, Boubou River, Ekala" stateProvince="Haut-ogooue Province">Bongoville-Léconi road, Boubou River, Ekala</location>
, swamp close to the river,
<quantity id="4CF34E97CADEFF2DEF5DFA2240C8FA58" box="[383,461,1487,1514]" metricMagnitude="2" metricUnit="m" metricValue="4.21" pageId="199" pageNumber="644" unit="m" value="421.0">
<elevation id="6D28DEDE3988C97949439B72E76E8887" box="[383,461,1487,1514]" metricMagnitude="2" metricUnit="m" metricValue="4.21" pageId="199" pageNumber="644" unit="m" value="421.0">421 m</elevation>
</quantity>
a.s.l. (
<geoCoordinate id="EE3F85B5CADEFF2DEC3CFA224380FA58" box="[542,645,1487,1514]" direction="south" orientation="latitude" pageId="199" pageNumber="644" precision="5" value="-1.6116">1.6116 ° S</geoCoordinate>
<geoCoordinate id="EE3F85B5CADEFF2DECB2FA22420FFA58" box="[656,778,1487,1514]" direction="east" orientation="longitude" pageId="199" pageNumber="644" precision="5" value="13.9116">13.9116 ° E</geoCoordinate>
),
<date id="FFB5C5B2CADEFF2DED00FA2242A9FA58" box="[802,940,1487,1514]" pageId="199" pageNumber="644" value="2010-12-30">
<collectingDate id="80B11C0E0102FD174B18FF51C26CB5C2" box="[802,940,1487,1514]" pageId="199" pageNumber="644" value="2010-12-30">30 -xii- 2010</collectingDate>
</date>
, leg.
<collectorName id="43CEBA2D65FE068A9E15531B8A8D3C56">N. Mézière</collectorName>
,
<collectionCode id="ED1A7BB7CADEFF2DEF2BFA184060F9BD" box="[265,357,1525,1551]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" pageId="199" pageNumber="644">RMNH</collectionCode>
</materialsCitation>
.
<materialsCitation id="865BA52E031015E680E3CA863C9D5F47" ID-GBIF-Occurrence="3396361351" box="[366,1032,1524,1551]" collectingDate="2012-01-27" collectionCode="RMNH" collectorName="N. Mézière" country="Gabon" elevation="421 m" httpUri="http://www.boldsystems.org/index.php/Public_RecordView?processid=NSPAF453-15" latitude="-1.6116" location="Bongoville-Léconi road, Boubou River, Ekala" longitude="13.9116" pageId="199" pageNumber="644" specimenCode="RMNH.INS.554425" specimenCount="1" specimenCount-male="1" stateProvince="Haut-ogooue Province">
<specimenCount id="798402CF3F0A563C436C87CCCBB52F21" box="[366,401,1524,1551]" pageId="199" pageNumber="644" type="generic">1 ♂</specimenCount>
(
<specimenCode id="7C78C86AE5314AD587E691CC8E349F27">RMNH.INS.508160</specimenCode>
), same locality,
<date id="FFB5C5B2CADEFF2DED60FA1942B3F9BD" box="[834,950,1524,1551]" pageId="199" pageNumber="644" value="2012-01-27">
<collectingDate id="EB5ABE548DE3F1201ADE7AA850418890" box="[834,950,1524,1551]" pageId="199" pageNumber="644" value="2012-01-27">27 -i- 2012</collectingDate>
</date>
, leg.
<collectorName id="3F55DB3EDB1EA94CE521EE7993AB93C9">
N.
<paragraph id="8BB4E372CAD1FF22EEBDFF1142EEFDF0" pageId="200" pageNumber="645">
Mézière,
<collectionCode id="ED1A7BB7CAD1FF22EF2FFF13406CFEAA" box="[269,361,254,280]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" pageId="200" pageNumber="645">RMNH</collectionCode>
.
<materialsCitation id="3B63E92FCAD1FF22EF54FF1340C4FE8F" ID-GBIF-Occurrence="3396361343" collectingDate="2011-11-20" collectionCode="RMNH" collectorName="N. Mézière" country="Gabon" elevation="421 m" httpUri="http://www.boldsystems.org/index.php/Public_RecordView?processid=NSPAF393-15" latitude="-1.6116" location="Bongoville-Léconi road, Boubou River, Ekala" longitude="13.9116" pageId="200" pageNumber="645" specimenCode="RMNH.INS.554425" specimenCount="5" specimenCount-female="1" specimenCount-male="4">
<specimenCount id="9D0D28FBCAD1FF22EF54FF13409DFEA5" box="[374,408,253,280]" pageId="200" pageNumber="645" type="generic">4 ♂</specimenCount>
<specimenCount id="9341EAD6685F6ADD125567A5F3251ACA" type="female">1 ♀</specimenCount>
(
<specimenCode id="EE302419DF2B06DFA7E6DD8F3CD4D45F">RMNH.INS.506221</specimenCode>
), same locality,
<date id="FFB5C5B2CAD1FF22EDA4FF104502FEAA" box="[902,1031,253,280]" pageId="200" pageNumber="645" value="2011-11-20">
<collectingDate id="216E1B5A759422710764A38840CD6044" box="[902,1031,253,280]" pageId="200" pageNumber="645" value="2011-11-20">20 -xi- 2011</collectingDate>
</date>
, leg.
<collectorName id="606F815751EC3BB3F54F235677495415">N. Mézière</collectorName>
,
<collectionCode id="ED1A7BB7CAD1FF22EF47FECE40C4FE8F" box="[357,449,291,317]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" pageId="200" pageNumber="645">RMNH</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B63E92FCAD1FF22EFECFECE43D4FED0" ID-GBIF-Occurrence="1224548097" collectingDate="2012-01-27" collectionCode="RMNH" collectorName="N. Mézière, A. Günther, J. Kipping &amp;, H. Krahnstöver" country="Gabon" elevation="421 m" latitude="-1.6116" location="Bongoville-Léconi road, Boubou River, Ekala" longitude="13.9116" pageId="200" pageNumber="645" specimenCount="2" specimenCount-male="2" stateProvince="Haut-ogooue Province">
<specimenCount id="9D0D28FBCAD1FF22EFECFECE40F5FE8E" box="[462,496,290,317]" pageId="200" pageNumber="645" type="generic">2 ♂</specimenCount>
, same locality,
<date id="FFB5C5B2CAD1FF22EC8BFECF4218FE8F" box="[681,797,290,317]" pageId="200" pageNumber="645" value="2012-01-27">
<collectingDate id="F7B71C6D88B3F5FC29528048D384A614" box="[681,797,290,317]" pageId="200" pageNumber="645" value="2012-01-27">27 -i- 2012</collectingDate>
</date>
, leg.
<collectorName id="C7B2A8D6A9B52A6AACF96FE3585792C1">N. Mézière</collectorName>
,
<collectorName id="68828CFF115879C78DD8D52543865A49">A. Günther</collectorName>
,
<collectorName id="1BC08EF373295CBEA826EB53E4BE72C3">J. Kipping &amp;</collectorName>
<collectorName id="F131F7DA5291EF648ECD0D64754C0C47">H. Krahnstöver</collectorName>
,
<collectionCode id="ED1A7BB7CAD1FF22EC57FEA443D4FED0" box="[629,721,329,354]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" pageId="200" pageNumber="645">RMNH</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B63E92FCAD1FF22ECFFFEA5432CFE60" ID-GBIF-Occurrence="3396361330" collectingDate="2013-04-05" collectionCode="RMNH" collectorName="N. Mézière" country="Gabon" elevation="427" httpUri="http://www.boldsystems.org/index.php/Public_RecordView?processid=NSPAF510-15" latitude="-1.6133" location="Bongoville-Léconi road, Boubou River, Ekala" longLatPrecision="7" longitude="13.9124" pageId="200" pageNumber="645" specimenCode="RMNH.INS.508803" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D0D28FBCAD1FF22ECFFFEA543FAFED0" box="[733,767,328,354]" pageId="200" pageNumber="645" type="generic">1 ♂</specimenCount>
(
<specimenCode id="C370D2875EC3F51AD5EC66FE858203D8">RMNH.INS.508803</specimenCode>
),
<location id="F18A3221E6E5D1C044CF1C03D4D5A21E" LSID="urn:lsid:plazi:treatment:03A25264CADEFF20EE9AFD5F40CBFCDF:F18A3221E6E5D1C044CF1C03D4D5A21E" country="Gabon" latitude="-1.6133" longLatPrecision="7" longitude="13.9124" name="Bongoville-Léconi road, Boubou River, Ekala">Bongoville-Léconi road, Boubou River, Ekala</location>
, sandy forest stream (
<quantity id="4CF34E97CAD1FF22EDE1FE804509FE3A" box="[963,1036,365,392]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="200" pageNumber="645" unit="m" value="1.5" valueMax="2.0" valueMin="1.0">12 m</quantity>
wide,
<quantity id="4CF34E97CAD1FF22EEC4FE7E404FFE1F" box="[230,330,403,429]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="200" pageNumber="645" unit="cm" value="7.5" valueMax="10.0" valueMin="5.0">510 cm</quantity>
deep), muddy edges,
<quantity id="4CF34E97CAD1FF22EC72FE7E439FFE1F" box="[592,666,403,429]" metricMagnitude="2" metricUnit="m" metricValue="4.27" pageId="200" pageNumber="645" unit="m" value="427.0">
<elevation id="FA92ADDADB5E7538B45BB3B7A9B0C2FA" box="[592,666,403,429]" metricMagnitude="2" metricUnit="m" metricValue="4.27" pageId="200" pageNumber="645" unit="m" value="427.0">427 m</elevation>
</quantity>
a.s.l. (
<geoCoordinate id="EE3F85B5CAD1FF22ECC5FE7F4248FE1F" box="[743,845,402,429]" direction="south" orientation="latitude" pageId="200" pageNumber="645" precision="5" value="-1.6133">1.6133 ° S</geoCoordinate>
<geoCoordinate id="EE3F85B5CAD1FF22ED74FE7F42D5FE1F" box="[854,976,402,429]" direction="east" orientation="longitude" pageId="200" pageNumber="645" precision="5" value="13.9124">13.9124 ° E</geoCoordinate>
),
<collectingDate id="60227EC2116476459EECD478FDF342E8" value="2013-04-05">
<date id="5FC2DBECA15F7608FB8AE6C8B8193FD6" value="2013-04-05">05- iv- 2013</date>
</collectingDate>
, leg.
<collectorName id="74DAECA3BDBED9B6A6AB9D07152ECF38">N. Mézière</collectorName>
,
<collectionCode id="ED1A7BB7CAD1FF22EFEFFE54432CFE60" box="[461,553,441,466]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" pageId="200" pageNumber="645">RMNH</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B63E92FCAD1FF22EC16FE554060FDF0" ID-GBIF-Occurrence="1224548075" collectingDate="2009-02-05" collectionCode="RMNH" collectorName="N. Mézière" country="Gabon" elevation="520 m" latitude="-1.6001" location="Bakoumba road, village 3 km before Kounda, Mikouangna (Mouyeugue)" longLatPrecision="7" longitude="13.1584" pageId="200" pageNumber="645" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D0D28FBCAD1FF22EC16FE554353FE60" box="[564,598,440,466]" pageId="200" pageNumber="645" type="generic">1 ♂</specimenCount>
,
<location id="FBC7F6FA5BAD42ED97502869518E761D" LSID="urn:lsid:plazi:treatment:03A25264CADEFF20EE9AFD5F40CBFCDF:FBC7F6FA5BAD42ED97502869518E761D" country="Gabon" latitude="-1.6001" longLatPrecision="7" longitude="13.1584" name="Bakoumba road, village 3 km before Kounda, Mikouangna (Mouyeugue)">
Bakoumba road, village
<quantity id="4CF34E97CAD1FF22ED5DFE5542BEFE60" box="[895,955,440,466]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="200" pageNumber="645" unit="km" value="3.0">3 km</quantity>
before Kounda, Mikouangna (Mouyeugue)
</location>
, large standing water in forest clearing and cassava fields,
<quantity id="4CF34E97CAD1FF22EF8AFDEE40F7FDAF" box="[424,498,515,541]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="200" pageNumber="645" unit="m" value="520.0">
<elevation id="BDCA66FF751E8AB1A863095F5C7D938D" box="[424,498,515,541]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="200" pageNumber="645" unit="m" value="520.0">520 m</elevation>
</quantity>
a.s.l. (
<geoCoordinate id="EE3F85B5CAD1FF22EC1EFDEF43A6FDAF" box="[572,675,514,541]" direction="south" orientation="latitude" pageId="200" pageNumber="645" precision="5" value="-1.6001">1.6001 ° S</geoCoordinate>
<geoCoordinate id="EE3F85B5CAD1FF22EC88FDEF4221FDAF" box="[682,804,514,541]" direction="east" orientation="longitude" pageId="200" pageNumber="645" precision="5" value="13.1584">13.1584 ° E</geoCoordinate>
),
<date id="FFB5C5B2CAD1FF22ED1AFDEF42B6FDAF" box="[824,947,514,541]" pageId="200" pageNumber="645" value="2009-02-05">
<collectingDate id="79E9B7C500F9F96004E9CA68F7F6F7F4" box="[824,947,514,541]" pageId="200" pageNumber="645" value="2009-02-05">05-ii- 2009</collectingDate>
</date>
, leg.
<collectorName id="D59B816C4CB8A164DD86BF5D0D8C65BF">N. Mézière</collectorName>
,
<collectionCode id="ED1A7BB7CAD1FF22EF28FDC44060FDF0" box="[266,357,553,578]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" pageId="200" pageNumber="645">RMNH</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B63E92FCAD1FF22EF4DFDC542E2FDF0" ID-GBIF-Occurrence="1224548076" box="[367,999,551,578]" collectingDate="2009-04-05" collectionCode="RMNH" collectorName="N. Mézière" country="Gabon" elevation="520 m" latitude="-1.6001" location="Bakoumba road, village 3 km before Kounda, Mikouangna (Mouyeugue)" longLatPrecision="7" longitude="13.1584" pageId="200" pageNumber="645" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D0D28FBCAD1FF22EF4DFDC54097FDF0" box="[367,402,552,578]" pageId="200" pageNumber="645" type="generic">1 ♂</specimenCount>
, same locality,
<date id="FFB5C5B2CAD1FF22EC60FDC543C6FDF0" box="[578,707,552,578]" pageId="200" pageNumber="645" value="2009-04-05">
<collectingDate id="32045462A624970E8A16B5A7D029C2B7" box="[578,707,552,578]" pageId="200" pageNumber="645" value="2009-04-05">05-iv- 2009</collectingDate>
</date>
, leg.
<collectorName id="8B2831C33A4F02B97FBA4FFC057B9A35">N. Mézière</collectorName>
,
<collectionCode id="ED1A7BB7CAD1FF22EDA9FDC442E2FDF0" box="[907,999,553,578]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" pageId="200" pageNumber="645">RMNH</collectionCode>
</materialsCitation>
.
</paragraph>
</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CAD1FF20EEBDFD9E40C7FE1F" lastPageId="202" lastPageNumber="647" pageId="200" pageNumber="645" type="diagnosis">
<paragraph id="8BB4E372CAD1FF23EEBDFD9E4317FEAA" lastPageId="201" lastPageNumber="646" pageId="200" pageNumber="645">
<emphasis id="B97F3F60CAD1FF22EEBDFD9E400CFD3F" bold="true" box="[159,265,627,653]" pageId="200" pageNumber="645">Genetics</emphasis>
Three unique haplotypes (n= 5) distinct from but close to five of
<taxonomicName id="4C0B98F1CAD1FF22EDBFFD7441E1FD6A" class="Insecta" family="Libellulidae" genus="Porpax" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="200" pageNumber="645" phylum="Arthropoda" rank="species" species="garambensis">
<emphasis id="B97F3F60CAD1FF22EDBFFD7441E1FD6A" italics="true" pageId="200" pageNumber="645">P. garambensis</emphasis>
</taxonomicName>
(n= 6).
<heading id="D0FC541ECAD0FF23EEBDFF134317FEAA" bold="true" box="[159,530,254,280]" fontSize="11" level="3" pageId="201" pageNumber="646" reason="6">
<emphasis id="B97F3F60CAD0FF23EEBDFF134317FEAA" bold="true" box="[159,530,254,280]" pageId="201" pageNumber="646">Male morphological diagnosis</emphasis>
</heading>
</paragraph>
<caption id="DF74B3FACAD1FF22EEBDFA1A422AF9BD" box="[159,815,1527,1552]" pageId="200" pageNumber="645" targetBox="[158,1035,776,1486]" targetPageId="200">
<paragraph id="8BB4E372CAD1FF22EEBDFA1A422AF9BD" box="[159,815,1527,1552]" pageId="200" pageNumber="645">
<emphasis id="B97F3F60CAD1FF22EEBDFA1A4041F9BD" bold="true" box="[159,324,1527,1552]" pageId="200" pageNumber="645">
Type
<figureCitation id="1330FFF7CAD1FF22EEF8FA1A4045F9BD" box="[218,320,1527,1551]" captionStart="Photo 54" captionStartId="157.[159,225,1382,1406]" captionText="Photo 54. Small forest stream south of Bentor, Liberia. Type locality of Paragomphus lemperti. Photo: KDBD (26 - ii- - 2012)" pageId="200" pageNumber="645">Photo 54</figureCitation>
.
</emphasis>
Holotype of
<taxonomicName id="4C0B98F1CAD1FF22EFEDFA1A437EF9BD" box="[463,635,1527,1551]" class="Insecta" family="Libellulidae" genus="Porpax" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="200" pageNumber="645" phylum="Arthropoda" rank="species" species="mezierei" status="sp. nov.">
<emphasis id="B97F3F60CAD1FF22EFEDFA1A437EF9BD" box="[463,635,1527,1551]" italics="true" pageId="200" pageNumber="645">Porpax mezierei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C821BCAD1FF22ECA3FA1A43DDF9BD" box="[641,728,1527,1551]" pageId="200" pageNumber="645" rank="species">sp. nov.</taxonomicNameLabel>
, RMNH.
</paragraph>
</caption>
<paragraph id="8BB4E372CAD0FF20EEBDFECF40C7FE1F" lastPageId="202" lastPageNumber="647" pageId="201" pageNumber="646">
Structurally like its sympatric sister-species
<taxonomicName id="4C0B98F1CAD0FF23EC8BFECE4256FE8F" box="[681,851,290,317]" class="Insecta" family="Libellulidae" genus="Porpax" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="201" pageNumber="646" phylum="Arthropoda" rank="species" species="garambensis">
<emphasis id="B97F3F60CAD0FF23EC8BFECE4256FE8F" box="[681,851,290,317]" italics="true" pageId="201" pageNumber="646">P. garambensis</emphasis>
</taxonomicName>
by (a) the dorsally converging pale postdorsal stripes; (b) the densely hairy anterior side of the hind femur and trochanter; and (c) the pointed rather than rounded lobe of the hamule. However, many details are closer to the norm in this genus, such as (1) the larger size, Hw
<quantity id="4CF34E97CAD0FF23EC68FE5543EAFE60" box="[586,751,440,466]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" metricValueMax="2.55" metricValueMin="2.35" pageId="201" pageNumber="646" unit="mm" value="24.5" valueMax="25.5" valueMin="23.5">23.525.5 mm</quantity>
(n = 8) rather than 21.5 23.0 mm; (2) the large central pale spot on the spiracular dorsum; (3)the black area between the pale postdorsal stripes that is at least as wide as the stripes themselves; (4) the venter of the thorax with the largely pale poststernum and metepimera enclosed by black; (5) normally 2 rather than 1 Cux in Hw, although both species vary between 1 and 3; (6) 11 ½– 12 ½ rath- er than 8 ½– 10 ½ Ax in Fw; and (7) the absence of abdominal pruinosity. Unique in the genus by (8) the pale band on the frons being severed by black medially, as well as at the level of each antenna; (9) the black vertex with at most a weak dorsal spot, rather than a large and contrasting pale patch; (10)S 47 with lateral blue spots that extend onto the ventral part of the tergites, but without a ring-like pale marking on S 6; and (11) the blue dorsa of the cerci and epiproct.
</paragraph>
</subSubSection>
<caption id="DF74B3FACAD0FF23EEBDFA1A42A7F9A2" box="[159,930,1527,1552]" pageId="201" pageNumber="646" targetBox="[158,1035,766,1485]" targetPageId="201">
<paragraph id="8BB4E372CAD0FF23EEBDFA1A42A7F9A2" box="[159,930,1527,1552]" pageId="201" pageNumber="646">
<emphasis id="B97F3F60CAD0FF23EEBDFA1A400CF9BD" bold="true" box="[159,265,1527,1551]" pageId="201" pageNumber="646">Photo 69.</emphasis>
<taxonomicName id="4C0B98F1CAD0FF23EF2DFA1A40BEF9BD" box="[271,443,1527,1551]" class="Insecta" family="Libellulidae" genus="Porpax" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="201" pageNumber="646" phylum="Arthropoda" rank="species" species="mezierei">
<emphasis id="B97F3F60CAD0FF23EF2DFA1A40BEF9BD" box="[271,443,1527,1551]" italics="true" pageId="201" pageNumber="646">Porpax mezierei</emphasis>
</taxonomicName>
, male; Ekala, Gabon. Photo: NM (06-xi-2010)
</paragraph>
</caption>
<subSubSection id="C311B0F9CAD3FF20EEBDFE334509FDDA" pageId="202" pageNumber="647" type="etymology">
<paragraph id="8BB4E372CAD3FF20EEBDFE334021FE4A" box="[159,292,478,504]" pageId="202" pageNumber="647">
<heading id="D0FC541ECAD3FF20EEBDFE334021FE4A" bold="true" box="[159,292,478,504]" fontSize="11" level="3" pageId="202" pageNumber="647" reason="6">
<emphasis id="B97F3F60CAD3FF20EEBDFE334021FE4A" bold="true" box="[159,292,478,504]" pageId="202" pageNumber="647">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BB4E372CAD3FF20EEBDFDEF4509FDDA" pageId="202" pageNumber="647">Named in honour of our co-author Nicolas Mézière who, living in southeastern Gabon, has made many discoveries in this odonatologically unexplored part of Africa within a short time (masculine singular genitive noun).</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CAD3FF20EEBDFD7440CBFCDF" pageId="202" pageNumber="647" type="distribution">
<paragraph id="8BB4E372CAD3FF20EEBDFD744081FD01" box="[159,388,665,691]" pageId="202" pageNumber="647">
<heading id="D0FC541ECAD3FF20EEBDFD744081FD01" bold="true" box="[159,388,665,691]" fontSize="11" level="3" pageId="202" pageNumber="647" reason="6">
<emphasis id="B97F3F60CAD3FF20EEBDFD744081FD01" bold="true" box="[159,388,665,691]" pageId="202" pageNumber="647">Range and ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BB4E372CAD3FF20EEBDFD5040CBFCDF" pageId="202" pageNumber="647">
Recorded between 350 and
<quantity id="4CF34E97CAD3FF20EFC5FD534334FD6A" box="[487,561,702,728]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="202" pageNumber="647" unit="m" value="550.0">550 m</quantity>
a.s.l. in south-eastern Gabon. Males are found on sunny vegetation by swamps and pools that have much detritus, probably never dry out completely and lie on the forest edge. Not found with
<taxonomicName id="4C0B98F1CAD3FF20EEF8FCC34084FCFA" box="[218,385,813,840]" class="Insecta" family="Libellulidae" genus="Porpax" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="202" pageNumber="647" phylum="Arthropoda" rank="species" species="garambensis">
<emphasis id="B97F3F60CAD3FF20EEF8FCC34084FCFA" box="[218,385,813,840]" italics="true" pageId="202" pageNumber="647">P. garambensis</emphasis>
</taxonomicName>
as that species prefers similar but deeply shaded habitat with only small sun spots.
</paragraph>
</subSubSection>
</treatment>
</document>