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<document id="3EC1D56A7B2B0A5C4DA292B01EFA1C51" ID-DOI="10.5281/zenodo.35388" ID-GBIF-Dataset="b184dfcc-9b5f-4abb-b401-0f45f76006f6" checkinTime="1450029366780" checkinUser="donat" docAuthor="Dijkstra, Klaas-Douwe B., Kipping, Jens &amp; Mézière, Nicolas" docDate="2015" docId="03A25264CA15FFFAEF37FB524211FE1F" docLanguage="en" docName="Odonatologica_44-4-low_res.pdf.imf" docOrigin="Odonatologica 44 (4)" docTitle="Umma gumma Dijkstra, Meziere &amp; Kipping, sp. nov." docType="treatment" docVersion="27" lastPageNumber="460" masterDocId="FF9B2A1CCA19FFEAEE22FFED4105FFB2" masterDocTitle="Sixty new dragonfly and damselfly species from Africa (Odonata)" masterLastPageNumber="678" masterPageNumber="447" pageNumber="457" updateTime="1695001031402" updateUser="admin">
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<mods:title id="244E2770834830F358DF95937ACF839A">Sixty new dragonfly and damselfly species from Africa (Odonata)</mods:title>
</mods:titleInfo>
<mods:name id="05486848974D84D780D813B60B08E128" type="personal">
<mods:role id="9AEC4316257ED1B0944AA8E9E9CADFD5">
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<mods:namePart id="5BC3C4E02E723B0CEC4C572288E1D693">Dijkstra, Klaas-Douwe B.</mods:namePart>
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<mods:namePart id="BDAA10D7671502E6693D3CC694B6B36B">Kipping, Jens</mods:namePart>
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<mods:namePart id="02DF0DD687AF4AE2F4C24585767F1827">Mézière, Nicolas</mods:namePart>
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<mods:title id="C1C484C56A25A504738377C71F71352B">Odonatologica</mods:title>
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<mods:part id="BF6874E07A4806131DDF66C3461C883D">
<mods:date id="EEE7F4DB7F1A77C80EFDD7278B0C0A90">2015</mods:date>
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<mods:number id="4CDF5BE80C33E37C70856DE8D35C1C64">44</mods:number>
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<mods:identifier id="544E9045338BF5B11E7BF3CDF23E8BEF" type="DOI">10.5281/zenodo.35388</mods:identifier>
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<treatment id="03A25264CA15FFFAEF37FB524211FE1F" ID-DOI="http://doi.org/10.5281/zenodo.5640190" ID-GBIF-Taxon="114923403" ID-Zenodo-Dep="5640190" LSID="urn:lsid:zoobank.org:act:03A25264-CA15-FFFA-EF37-FB524211FE1F" httpUri="http://treatment.plazi.org/id/03A25264CA15FFFAEF37FB524211FE1F" lastPageId="16" lastPageNumber="460" pageId="12" pageNumber="457">
<subSubSection id="C311B0F9CA15FFE6EF37FB5243C8FB4F" pageId="12" pageNumber="457" type="nomenclature">
<paragraph id="8BB4E372CA15FFE6EF37FB5243C8FB4F" pageId="12" pageNumber="457">
<heading id="D0FC541ECA15FFE6EF37FB5243C8FB4F" bold="true" centered="true" fontSize="11" level="2" pageId="12" pageNumber="457" reason="4">
<emphasis id="B97F3F60CA15FFE6EF37FB524290FB6B" bold="true" box="[277,917,1213,1242]" pageId="12" pageNumber="457">
<taxonomicName id="4C0B98F1CA15FFE6EF37FB524218FB68" LSID-ZBK="urn:lsid:zoobank.org:act:03A25264-CA15-FFFA-EF37-FB524211FE1F" authority="Dijkstra, Meziere &amp; Kipping" authorityName="Dijkstra, Meziere &amp; Kipping" box="[277,797,1213,1242]" class="Insecta" family="Calopterygidae" genus="Umma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="457" phylum="Arthropoda" rank="species" species="gumma" status="sp. nov.">
<emphasis id="B97F3F60CA15FFE6EF37FB5240CEFB68" bold="true" box="[277,459,1215,1242]" italics="true" pageId="12" pageNumber="457">Umma gumma</emphasis>
Dijkstra, Mézière &amp; Kipping
</taxonomicName>
<taxonomicNameLabel id="A24C821BCA15FFE6ED01FB524285FB6B" box="[803,896,1215,1241]" pageId="12" pageNumber="457" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<emphasis id="B97F3F60CA15FFE6EFFFFB0C43C8FB4F" bold="true" box="[477,717,1249,1277]" pageId="12" pageNumber="457">Robust Sparklewing</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CA15FFE6EFACFAE1401AFAE6" pageId="12" pageNumber="457" type="description">
<paragraph id="8BB4E372CA15FFE6EFACFAE14219FA95" box="[398,796,1291,1319]" pageId="12" pageNumber="457">
<heading id="D0FC541ECA15FFE6EFACFAE14219FA95" box="[398,796,1291,1319]" centered="true" fontSize="11" level="4" pageId="12" pageNumber="457" reason="4">
<figureCitation id="1330FFF7CA15FFE6EFF4FAE64331FA94" box="[470,564,1291,1318]" captionStart="Photo 1" captionStartId="16.[159,225,1096,1120]" captionTargetBox="[159,1035,459,1055]" captionTargetId="figure@16.[159,1038,459,1058]" captionTargetPageId="16" captionText="Photo 1. Umma gumma, male; Moyabi, Gabon. Photo: JK (24 - ix- 2013)" httpUri="http://dx.doi.org/10.5281/zenodo.35441" pageId="12" pageNumber="457">(Type Photo 1</figureCitation>
,
<figureCitation id="1330FFF7CA15FFE6EC63FAE643CCFA94" box="[577,713,1291,1318]" captionStart-0="Photo 1" captionStart-1="Photo 2" captionStartId-0="16.[159,225,1096,1120]" captionStartId-1="16.[159,225,1353,1378]" captionTargetBox-0="[159,1035,459,1055]" captionTargetId-0="figure@16.[159,1038,459,1058]" captionTargetPageId-0="16" captionText-0="Photo 1. Umma gumma, male; Moyabi, Gabon. Photo: JK (24 - ix- 2013)" captionText-1="Photo 2. Sandy forest stream near Moyabi, Gabon. Habitat of Umma gumma, Paragomphus dispar and Trithemis legrandi. Photo: JK (24 - ix- 2013)" httpUri-0="http://dx.doi.org/10.5281/zenodo.35441" httpUri-1="http://dx.doi.org/10.5281/zenodo.35444" pageId="12" pageNumber="457">Photos 12</figureCitation>
,
<figureCitation id="1330FFF7CA15FFE6ECF6FAE14214FA95" box="[724,785,1292,1319]" captionStart="Figure 1" captionStartId="15.[751,820,1352,1377]" captionTargetBox="[159,708,1067,1550]" captionTargetId="figure@15.[156,716,1049,1550]" captionTargetPageId="15" captionText="Figure 1. Male cerci in dorsal view (left) and paraprocts in ventral view with dotted outline of cerci (right) of Umma gumma sp. nov. and U. longistigma." httpUri="http://dx.doi.org/10.5281/zenodo.35442" pageId="12" pageNumber="457">Fig. 1</figureCitation>
)
</heading>
</paragraph>
<paragraph id="8BB4E372CA15FFE6EEBDFAD7401AFAE6" box="[159,287,1338,1364]" pageId="12" pageNumber="457">
<heading id="D0FC541ECA15FFE6EEBDFAD7401AFAE6" bold="true" box="[159,287,1338,1364]" fontSize="11" level="3" pageId="12" pageNumber="457" reason="6">
<emphasis id="B97F3F60CA15FFE6EEBDFAD7401AFAE6" bold="true" box="[159,287,1338,1364]" pageId="12" pageNumber="457">Taxonomy</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CA15FFE7EEBDFAB243F2FE8F" lastPageId="13" lastPageNumber="458" pageId="12" pageNumber="457" type="discussion">
<paragraph id="8BB4E372CA15FFE7EEBDFAB243F2FE8F" lastPageId="13" lastPageNumber="458" pageId="12" pageNumber="457">
<bibRefCitation id="BB1F7819006E6B932CCE16B3F04DCF6B" author="Pinhey E. C. G." journalOrPublisher="Arnoldia, Rhodesia" pagination="1 - 11" part="4" refId="ref68752" refString="Pinhey E. C. G. 1969. On the genus Umma Kirby (Odonata). Arnoldia, Rhodesia, 4: 1 - 11" title="On the genus Umma Kirby (Odonata)" type="journal article" year="1969">Pinhey (1969)</bibRefCitation>
remarked that the variability of
<taxonomicName id="4C0B98F1CA15FFE6ECE8FA8D4509FACB" authority="Selys, 1869" authorityName="Selys" authorityYear="1869" box="[714,1036,1375,1401]" class="Insecta" family="Calopterygidae" genus="Umma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="457" phylum="Arthropoda" rank="species" species="longistigma">
<emphasis id="B97F3F60CA15FFE6ECE8FA8D426AFACB" box="[714,879,1375,1401]" italics="true" pageId="12" pageNumber="457">U. longistigma</emphasis>
(Selys, 1869)
</taxonomicName>
might conceal unnamed taxa.
<bibRefCitation id="631016C74A1E13559FBD0CCB4825C0EA" author="Vick G. S." journalOrPublisher="Odonatologica" pagination="87 - 98" part="27" refId="ref69194" refString="Vick G. S. 1999. A checklist of the Odonata of the south-west province of Cameroon, with the description of Phyllogomphus corbetae spec. nov. (Anisoptera: Gomphidae). Odonatologica 27: 87 - 98" title="A checklist of the Odonata of the south-west province of Cameroon, with the description of Phyllogomphus corbetae spec. nov. (Anisoptera: Gomphidae)" type="journal article" year="1999">Vick (1999)</bibRefCitation>
noted that two forms with different paraprocts occur together in Cameroon, the typical one having more slender paraprocts, as confirmed by the
<typeStatus id="54B05DD0CA15FFE6EC49FA2243D7FA5B" box="[619,722,1487,1513]" pageId="12" pageNumber="457">holotype</typeStatus>
of
<taxonomicName id="4C0B98F1CA15FFE6ECD7FA3D429DFA5B" box="[757,920,1487,1513]" class="Insecta" family="Calopterygidae" genus="Umma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="457" phylum="Arthropoda" rank="species" species="longistigma">
<emphasis id="B97F3F60CA15FFE6ECD7FA3D429DFA5B" box="[757,920,1487,1513]" italics="true" pageId="12" pageNumber="457">U. longistigma</emphasis>
</taxonomicName>
from that nation in
<collectionCode id="ED1A7BB7CA15FFE6EF2DFA194048F9BD" box="[271,333,1524,1551]" collectionName="Belgium, Brussels, Institut Royal des Sciences Naturelles de Belgique" pageId="12" pageNumber="457">ISNB</collectionCode>
(
<figureCitation id="1330FFF7CA15FFE6EF7CFA19409AF9BD" box="[350,415,1524,1551]" captionStart="Figure 1" captionStartId="15.[751,820,1352,1377]" captionTargetBox="[159,708,1067,1550]" captionTargetId="figure@15.[156,716,1049,1550]" captionTargetPageId="15" captionText="Figure 1. Male cerci in dorsal view (left) and paraprocts in ventral view with dotted outline of cerci (right) of Umma gumma sp. nov. and U. longistigma." httpUri="http://dx.doi.org/10.5281/zenodo.35442" pageId="12" pageNumber="457">Fig. 1</figureCitation>
). Both forms overlap widely across central Africa, are easily separated by other characters as well, and differ genetically where they co-occur in Gabon, thus representing good species.
</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CA14FFE7EEBDFE83406DFE3A" box="[159,360,366,392]" pageId="13" pageNumber="458" type="description">
<paragraph id="8BB4E372CA14FFE7EEBDFE83406DFE3A" box="[159,360,366,392]" pageId="13" pageNumber="458">
<heading id="D0FC541ECA14FFE7EEBDFE83406DFE3A" bold="true" box="[159,360,366,392]" fontSize="11" level="3" pageId="13" pageNumber="458" reason="6">
<emphasis id="B97F3F60CA14FFE7EEBDFE83406DFE3A" bold="true" box="[159,360,366,392]" pageId="13" pageNumber="458">Material studied</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CA14FFE4EEBDFE794509FE4A" lastPageId="14" lastPageNumber="459" pageId="13" pageNumber="458" type="materials_examined">
<paragraph id="8BB4E372CA14FFE7EEBDFE794509FE4A" pageId="13" pageNumber="458">
<materialsCitation id="3B63E92FCA14FFE7EEBDFE79450DFE4A" ID-GBIF-Occurrence="3396361331" collectingDate="2012-09-28" collectionCode="RMNH" collectorName="N. Mézière" country="Gabon" elevation="424 m" httpUri="http://www.boldsystems.org/index.php/Public_RecordView?processid=NSPAF547-15" latitude="-0.8403" location="Alanga- Aboumi road, Moyol, gravelly and sandy forest river (2 3 m wide, 10 50 cm deep)" longLatPrecision="7" longitude="13.9385" pageId="13" pageNumber="458" specimenCode="RMNH.INS.554425" specimenCount="1" specimenCount-male="1" stateProvince="Haut-Ogooué Province" typeStatus="holotype">
<emphasis id="B97F3F60CA14FFE7EEBDFE79400AFE1F" bold="true" box="[159,271,404,429]" pageId="13" pageNumber="458">
<typeStatus id="54B05DD0CA14FFE7EEBDFE79400AFE1F" box="[159,271,404,429]" pageId="13" pageNumber="458">Holotype</typeStatus>
</emphasis>
<emphasis id="B97F3F60CA14FFE7EF3AFE7F4029FE1E" bold="true" box="[280,300,402,428]" pageId="13" pageNumber="458">
<specimenCount id="CA060521064A98B38E855BBF3C8D2F9F" type="male"></specimenCount>
</emphasis>
<emphasis id="B97F3F60CA14FFE7EF0EFE794036FE1F" bold="true" box="[300,307,404,429]" pageId="13" pageNumber="458">.</emphasis>
<specimenCode id="14C7B13B6220EFBA1AC5F4C6A8B65FE9">RMNH.INS.554425</specimenCode>
,
<collectingCountry id="F31CA3E2CA14FFE7EC10FE7F4387FE1F" box="[562,642,402,429]" name="Gabon" pageId="13" pageNumber="458" value="Gabon">Gabon</collectingCountry>
,
<collectingRegion id="49CF2D90CA14FFE7ECB2FE7F42A7FE1F" box="[656,930,402,429]" pageId="13" pageNumber="458">Haut-Ogooué Province</collectingRegion>
,
<location id="4F4A3267EB8CBB273D6BB084B95658F4" LSID="urn:lsid:plazi:treatment:03A25264CA15FFFAEF37FB524211FE1F:4F4A3267EB8CBB273D6BB084B95658F4" country="Gabon" latitude="-0.8403" longLatPrecision="7" longitude="13.9385" name="Alanga- Aboumi road, Moyol, gravelly and sandy forest river (2 3 m wide, 10 50 cm deep)" stateProvince="Haut-Ogooué Province">
Alanga- Aboumi road, Moyol, gravelly and sandy forest river (
<quantity id="4CF34E97CA14FFE7ED24FE554255FE60" box="[774,848,440,466]" metricMagnitude="0" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="13" pageNumber="458" unit="m" value="2.5" valueMax="3.0" valueMin="2.0">23 m</quantity>
wide,
<quantity id="4CF34E97CA14FFE7EDB8FE55450EFE60" box="[922,1035,440,466]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="13" pageNumber="458" unit="cm" value="30.0" valueMax="50.0" valueMin="10.0">1050 cm</quantity>
deep)
</location>
,
<quantity id="4CF34E97CA14FFE7EECEFE334036FE4A" box="[236,307,478,504]" metricMagnitude="2" metricUnit="m" metricValue="4.24" pageId="13" pageNumber="458" unit="m" value="424.0">
<elevation id="C0EFE9322CD747781CE3558AD5E4F73B" box="[236,307,478,504]" metricMagnitude="2" metricUnit="m" metricValue="4.24" pageId="13" pageNumber="458" unit="m" value="424.0">424 m</elevation>
</quantity>
a.s.l. (
<geoCoordinate id="EE3F85B5CA14FFE7EF5AFE3040D8FE4A" box="[376,477,477,504]" direction="south" orientation="latitude" pageId="13" pageNumber="458" precision="5" value="-0.8403">0.8403 ° S</geoCoordinate>
<geoCoordinate id="EE3F85B5CA14FFE7EFC1FE30435EFE4A" box="[483,603,477,504]" direction="east" orientation="longitude" pageId="13" pageNumber="458" precision="5" value="13.9385">13.9385 ° E</geoCoordinate>
),
<date id="FFB5C5B2CA14FFE7EC4FFE3043E9FE4A" box="[621,748,477,504]" pageId="13" pageNumber="458" value="2012-09-28">
<collectingDate id="08B6CE6AA3DE398B6B857B6F19F91BE8" box="[621,748,477,504]" pageId="13" pageNumber="458" value="2012-09-28">28 -ix- 2012</collectingDate>
</date>
, leg.
<collectorName id="0892E61F14C15C1212F9CC720FC04992">N. Mézière</collectorName>
,
<collectionCode id="ED1A7BB7CA14FFE7ED8FFE33450DFE4A" box="[941,1032,478,504]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" pageId="13" pageNumber="458">RMNH</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB4E372CA14FFE4EEBDFDE94509FE4A" lastPageId="14" lastPageNumber="459" pageId="13" pageNumber="458">
<emphasis id="B97F3F60CA14FFE7EEBDFDE94071FDAF" bold="true" box="[159,372,515,541]" pageId="13" pageNumber="458">Further material.</emphasis>
<materialsCitation id="3B63E92FCA14FFE7EF5FFDEE406DFD00" ID-GBIF-Occurrence="1224547681" collectingDate="2008-06-16" collectionCode="RMNH" collectorName="D. B. Dijkstra, J. Kipping &amp;, K. Schütte" country="Cameroon" elevation="85 m" latitude="2.394" location="26 km E of Campo, Campo-Maan National Park (buffer zone), Bitandé River 1 km E of Nkoélon, rocky rainforest river and adjacent roadsides" longLatPrecision="7" longitude="10.054" pageId="13" pageNumber="458" specimenCount="1" stateProvince="South Province">
<collectingCountry id="F31CA3E2CA14FFE7EF5FFDEE4323FDAF" box="[381,550,515,541]" name="Cameroon" pageId="13" pageNumber="458" value="Cameroon">CAMEROON</collectingCountry>
(
<collectingRegion id="49CF2D90CA14FFE7EC1BFDEF43F1FDAF" box="[569,756,514,541]" pageId="13" pageNumber="458">South Province</collectingRegion>
):
<specimenCount id="9D0D28FBCA14FFE7ED2AFDEE422FFDAE" box="[776,810,514,541]" pageId="13" pageNumber="458" type="generic">1 ♂</specimenCount>
,
<location id="7A24BB412945EAD8C3934E48198E2549" LSID="urn:lsid:plazi:treatment:03A25264CA15FFFAEF37FB524211FE1F:7A24BB412945EAD8C3934E48198E2549" country="Cameroon" latitude="2.394" longLatPrecision="7" longitude="10.054" name="26 km E of Campo, Campo-Maan National Park (buffer zone), Bitandé River 1 km E of Nkoélon, rocky rainforest river and adjacent roadsides" stateProvince="South Province">
<quantity id="4CF34E97CA14FFE7ED18FDEE4283FDAF" box="[826,902,514,541]" metricMagnitude="4" metricUnit="m" metricValue="2.6" pageId="13" pageNumber="458" unit="km" value="26.0">26 km</quantity>
E of Campo, Campo-Maan National Park (buffer zone), Bitandé River 1 km E of Nkoélon, rocky rainforest river and adjacent roadsides
</location>
,
<quantity id="4CF34E97CA14FFE7ED71FDA342C9FDDA" box="[851,972,590,616]" metricMagnitude="1" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="13" pageNumber="458" unit="m" value="85.0" valueMax="100.0" valueMin="70.0">
<elevation id="71F130C6F1F178D64C9C9EE6BFB998F7" box="[851,972,590,616]" metricMagnitude="1" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="13" pageNumber="458" unit="m" value="85.0" valueMax="100.0" valueMin="70.0">70100 m</elevation>
</quantity>
a.s.l. (
<geoCoordinate id="EE3F85B5CA14FFE7EE85FD9F4013FD3F" box="[167,278,626,653]" direction="north" orientation="latitude" pageId="13" pageNumber="458" precision="5" value="2.394">2.3940 ° N</geoCoordinate>
<geoCoordinate id="EE3F85B5CA14FFE7EF03FD9F409EFD3F" box="[289,411,626,653]" direction="east" orientation="longitude" pageId="13" pageNumber="458" precision="5" value="10.054">10.0540 ° E</geoCoordinate>
),
<date id="FFB5C5B2CA14FFE7EF96FD9F4330FD3F" box="[436,565,626,653]" pageId="13" pageNumber="458" value="2008-06-16">
<collectingDate id="5606356E1DD6CE23E2995274579A941C" box="[436,565,626,653]" pageId="13" pageNumber="458" value="2008-06-16">16 -vi- 2008</collectingDate>
</date>
, leg. K.-
<collectorName id="0D152713FE135AB4B87CDF48B8CDA7D5">D.B. Dijkstra</collectorName>
,
<collectorName id="6B07F293AA061AF390CDC4F4F599FB3B">J. Kipping &amp;</collectorName>
<collectorName id="1FFA3D4FD9350B212BCCDB724B985518">K. Schütte</collectorName>
,
<collectionCode id="ED1A7BB7CA14FFE7EF2EFD74406DFD00" box="[268,360,665,690]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" pageId="13" pageNumber="458">RMNH</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B63E92FCA14FFE7EF5EFD75432DFD4F" ID-GBIF-Occurrence="1224547683" collectingDate="2010-11-25" collectingDateMax="2010-11-26" collectingDateMin="2010-11-25" collectionCode="RMNH" collectorName="P. H. Lambret" country="Republic of the Congo" elevation="50 m" latitude="-3.7951" location="Conkouati National Park" longLatPrecision="7" longitude="11.3859" municipality="Region de Koulilou" pageId="13" pageNumber="458" specimenCount="1">
<collectingCountry id="F31CA3E2CA14FFE7EF5EFD7543B4FD00" box="[380,689,664,690]" name="Republic of the Congo" pageId="13" pageNumber="458" value="Republic of the Congo">CONGO-BRAZZAVILLE</collectingCountry>
(
<collectingMunicipality id="AEBC253247ED869DB77DF759C18837EA">Region de Koulilou</collectingMunicipality>
):
<specimenCount id="9D0D28FBCA14FFE7EDC1FD754500FD00" box="[995,1029,664,690]" pageId="13" pageNumber="458" type="generic">1 ♂</specimenCount>
,
<location id="2FB7645EBB902FEAA6B0502D275B4D43" LSID="urn:lsid:plazi:treatment:03A25264CA15FFFAEF37FB524211FE1F:2FB7645EBB902FEAA6B0502D275B4D43" country="Republic of the Congo" latitude="-3.7951" longLatPrecision="7" longitude="11.3859" municipality="Region de Koulilou" name="Conkouati National Park">Conkouati National Park</location>
, forest,
<quantity id="4CF34E97CA14FFE7EC3DFD53435CFD6A" box="[543,601,702,728]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="458" unit="m" value="50.0">
<elevation id="E54BE560FB973E9D93FB55C16D709D74" box="[543,601,702,728]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="458" unit="m" value="50.0">50 m</elevation>
</quantity>
a.s.l. (
<geoCoordinate id="EE3F85B5CA14FFE7EC83FD50420DFD6A" box="[673,776,701,728]" direction="south" orientation="latitude" pageId="13" pageNumber="458" precision="5" value="-3.7951">3.7951 ° S</geoCoordinate>
<geoCoordinate id="EE3F85B5CA14FFE7ED2CFD50428DFD6A" box="[782,904,701,728]" direction="east" orientation="longitude" pageId="13" pageNumber="458" precision="5" value="11.3859">11.3859 ° E</geoCoordinate>
),
<date id="FFB5C5B2CA14FFE7EDB9FD5341E1FD4F" pageId="13" pageNumber="458" value="2010-11-25" valueMax="2010-11-26" valueMin="2010-11-25">
<collectingDate id="E6A914F85451501115A5E1AB8807508F" pageId="13" pageNumber="458" value="2010-11-25" valueMax="2010-11-26" valueMin="2010-11-25">2526 -xi- - 2010</collectingDate>
</date>
, leg.
<collectorName id="66147E82784817F3A787F1D153EBF5C2">P.H. Lambret</collectorName>
,
<collectionCode id="ED1A7BB7CA14FFE7EFEEFD0E432DFD4F" box="[460,552,739,765]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" pageId="13" pageNumber="458">RMNH</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B63E92FCA14FFE7EC16FD0E424BFCDF" ID-GBIF-Occurrence="1224547695" collectingDate="2010-05-04" collectionCode="RMNH" collectorName="D. B. Dijkstra" country="Democratic Republic of the Congo" elevation="420 m" latitude="0.83" location="Lilanda and Baombo streams at and upstream of Lilanda and Yafake villages" longLatPrecision="785" longitude="24.34" pageId="13" pageNumber="458" specimenCount="1" stateProvince="Province Orientale">
<collectingCountry id="F31CA3E2CA14FFE7EC16FD0E423CFD4F" box="[564,825,739,765]" name="Democratic Republic of the Congo" pageId="13" pageNumber="458" value="Democratic Republic of the Congo">CONGO-KINSHASA</collectingCountry>
(
<collectingRegion id="49CF2D90CA14FFE7ED68FD0F41C8FC90" pageId="13" pageNumber="458">Province Orientale</collectingRegion>
):
<specimenCount id="9D0D28FBCA14FFE7EEC3FCE54006FC90" box="[225,259,776,802]" pageId="13" pageNumber="458" type="generic">1 ♂</specimenCount>
,
<location id="579F7FE1D5668DA70FE7590431488DFF" LSID="urn:lsid:plazi:treatment:03A25264CA15FFFAEF37FB524211FE1F:579F7FE1D5668DA70FE7590431488DFF" country="Democratic Republic of the Congo" latitude="0.83" longLatPrecision="785" longitude="24.34" name="Lilanda and Baombo streams at and upstream of Lilanda and Yafake villages" stateProvince="Province Orientale">Lilanda and Baombo streams at and upstream of Lilanda and Yafake villages</location>
, sandy clear and blackwater streams and river,
<quantity id="4CF34E97CA14FFE7ED73FCC342D7FCFA" box="[849,978,814,840]" metricMagnitude="2" metricUnit="m" metricValue="4.2" metricValueMax="4.5" metricValueMin="3.9" pageId="13" pageNumber="458" unit="m" value="420.0" valueMax="450.0" valueMin="390.0">
<elevation id="7D5C220AB2C97378E43A12B149B8D6FB" box="[849,978,814,840]" metricMagnitude="2" metricUnit="m" metricValue="4.2" metricValueMax="4.5" metricValueMin="3.9" pageId="13" pageNumber="458" unit="m" value="420.0" valueMax="450.0" valueMin="390.0">390450 m</elevation>
</quantity>
a.s.l. (
<geoCoordinate id="EE3F85B5CA14FFE7EE85FCBF41FFFCDF" box="[167,250,850,877]" direction="north" orientation="latitude" pageId="13" pageNumber="458" precision="555" value="0.83">0.83 ° N</geoCoordinate>
<geoCoordinate id="EE3F85B5CA14FFE7EF23FCBF405AFCDF" box="[257,351,850,877]" direction="east" orientation="longitude" pageId="13" pageNumber="458" precision="555" value="24.34">24.34 ° E</geoCoordinate>
),
<date id="FFB5C5B2CA14FFE7EF51FCBE40E9FCDF" box="[371,492,851,877]" pageId="13" pageNumber="458" value="2010-05-04">
<collectingDate id="1E016323AC4754A412E426ACB68B9380" box="[371,492,851,877]" pageId="13" pageNumber="458" value="2010-05-04">04-v- 2010</collectingDate>
</date>
, leg. K.-
<collectorName id="EAFA7F2D60B1573A96EB4A02FC3ACE16">D.B. Dijkstra</collectorName>
,
<collectionCode id="ED1A7BB7CA14FFE7ECD0FCBE424BFCDF" box="[754,846,851,877]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" pageId="13" pageNumber="458">RMNH</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B63E92FCA14FFE7ED7AFCBE4030FC6F" ID-GBIF-Occurrence="1224547691" collectingDate="2004-11-01" collectingDateMax="2004-11-06" collectingDateMin="2004-11-01" collectionCode="RMNH" collectorName="D. B. Dijkstra" country="Democratic Republic of the Congo" elevation="387.5 m" latitude="1.275" location="NW of Lokutu, between Lukomete and Lunua River" longLatPrecision="78" longitude="23.425" pageId="13" pageNumber="458" specimenCount="1" stateProvince="Province Orientale">
<specimenCount id="9D0D28FBCA14FFE7ED7AFCBE427FFCDE" box="[856,890,850,877]" pageId="13" pageNumber="458" type="generic">1 ♂</specimenCount>
,
<quantity id="4CF34E97CA14FFE7EDAAFCBE42D7FCDF" box="[904,978,850,877]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="458" unit="km" value="20.0">
<locationDeviation id="DAFA044C67BE7EC6BA9BA9EE900F52FD" box="[904,978,850,877]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="458" unit="km" value="20.0">20 km</locationDeviation>
</quantity>
<location id="FDFABF9EE4B13CDA8E208DD85D09DE7B" LSID="urn:lsid:plazi:treatment:03A25264CA15FFFAEF37FB524211FE1F:FDFABF9EE4B13CDA8E208DD85D09DE7B" country="Democratic Republic of the Congo" latitude="1.275" longLatPrecision="78" longitude="23.425" name="NW of Lokutu, between Lukomete and Lunua River" stateProvince="Province Orientale">NW of Lokutu, between Lukomete and Lunua River</location>
, forest streams and clearings,
<quantity id="4CF34E97CA14FFE7EEFEFC734059FC0A" box="[220,348,926,952]" metricMagnitude="2" metricUnit="m" metricValue="3.875" metricValueMax="4.0" metricValueMin="3.75" pageId="13" pageNumber="458" unit="m" value="387.5" valueMax="400.0" valueMin="375.0">
<elevation id="8FA7CFA54AB36C647712BD6A7CE8BC7F" box="[220,348,926,952]" metricMagnitude="2" metricUnit="m" metricValue="3.875" metricValueMax="4.0" metricValueMin="3.75" pageId="13" pageNumber="458" unit="m" value="387.5" valueMax="400.0" valueMin="375.0">375400 m</elevation>
</quantity>
a.s.l. (
<geoCoordinate id="EE3F85B5CA14FFE7EF86FC704300FC0A" box="[420,517,925,952]" direction="north" orientation="latitude" pageId="13" pageNumber="458" precision="55" value="1.275">1.275 ° N</geoCoordinate>
<geoCoordinate id="EE3F85B5CA14FFE7EC29FC704372FC0A" box="[523,631,925,952]" direction="east" orientation="longitude" pageId="13" pageNumber="458" precision="55" value="23.425">23.425 ° E</geoCoordinate>
),
<date id="FFB5C5B2CA14FFE7ECA8FC704232FC0A" box="[650,823,925,952]" pageId="13" pageNumber="458" value="2004-11-01" valueMax="2004-11-06" valueMin="2004-11-01">
<collectingDate id="CD4F694827DC14DFFDF5D7BA9A79D698" box="[650,823,925,952]" pageId="13" pageNumber="458" value="2004-11-01" valueMax="2004-11-06" valueMin="2004-11-01">0106-xi- 2004</collectingDate>
</date>
, leg. K.-
<collectorName id="54391C6A0BF6924E3EB40889B20D6D42">D.B. Dijkstra</collectorName>
,
<collectionCode id="ED1A7BB7CA14FFE7EEFBFC2E4030FC6F" box="[217,309,963,989]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" pageId="13" pageNumber="458">RMNH</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B63E92FCA14FFE7EF63FC2E4372FB9A" ID-GBIF-Occurrence="1224547692" collectingDate="2004-11-01" collectionCode="RMNH" collectorName="D. B. Dijkstra" country="Democratic Republic of the Congo" elevation="4" latitude="1.0866" location="Lokutu" longLatPrecision="7" longitude="23.5368" municipality="Lokutu" pageId="13" pageNumber="458" specimenCount="1" stateProvince="Province Orientale">
<specimenCount id="9D0D28FBCA14FFE7EF63FC2E4066FC6E" box="[321,355,962,989]" pageId="13" pageNumber="458" type="generic">1 ♂</specimenCount>
,
<quantity id="4CF34E97CA14FFE7EF50FC2E40BBFC6F" metricMagnitude="4" metricUnit="m" metricValue="2.0" unit="km" value="20.0">
<locationDeviation id="0D76A50BDFEFFF15A103C8942527F97F" box="[370,446,962,989]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="458" unit="km" value="20.0">20 km</locationDeviation>
</quantity>
NW of
<collectingMunicipality id="EC2596094306FCA2BDC395FB5661F3D2" country="Democratic Republic of the Congo" latitude="1.0866" longLatPrecision="7" longitude="23.5368" name="Lokutu" stateProvince="Province Orientale">Lokutu</collectingMunicipality>
, large shallow, sandy, clear stream (
<quantity id="25749E9A2326BADC72D7D7CAA318EA47" box="[169,245,1000,1026]" metricMagnitude="0" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="13" pageNumber="458" unit="m" value="4.0" valueMax="5.0" valueMin="3.0">35 m</quantity>
wide) in disturbed forest,
<elevation id="B74CCA5701E246AC82C12F2FCBBD2924" metricMagnitude="2" metricUnit="m" metricValue="4.1" unit="m" value="410.0">
<quantity id="60796866CA2C62F866154FFB534C0954" metricMagnitude="2" metricUnit="m" metricValue="4.1" unit="m" value="410.0">410 m</quantity>
</elevation>
a.s.l. (
<geoCoordinate id="EE3F85B5CA14FFE7ECF8FC05424CFBB0" box="[730,841,1000,1026]" direction="north" orientation="latitude" pageId="13" pageNumber="458" precision="5" value="1.0866">1.0866 ° N</geoCoordinate>
<geoCoordinate id="EE3F85B5CA14FFE7ED76FC0542CBFBB0" box="[852,974,1000,1026]" direction="east" orientation="longitude" pageId="13" pageNumber="458" precision="5" value="23.5368">23.5368 ° E</geoCoordinate>
),
<collectingDate id="12F6106525E8BAB67C3062D1AC495FB5" value="2004-11-01">
<date id="C5484DFE300E000F6D6D4092B68286A8" value="2004-11-01">01- xi- 2004</date>
</collectingDate>
, leg. K.-
<collectorName id="80275CCF1BCA87BBA7D7FCCF44F91EB0">D.B. Dijkstra</collectorName>
,
<collectionCode id="ED1A7BB7CA14FFE7EC39FBE34372FB9A" box="[539,631,1038,1064]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" pageId="13" pageNumber="458">RMNH</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B63E92FCA14FFE7ECA7FBE34376FB2A" ID-GBIF-Occurrence="1224547682" collectingDate="2010-06-08" collectionCode="RMNH" collectorName="D. B. Dijkstra" country="Democratic Republic of the Congo" elevation="413" latitude="0.294" location="Ubundu Road, Yoko Forest, Mokonoka Stream" longLatPrecision="7" longitude="25.2888" pageId="13" pageNumber="458" specimenCount="3" stateProvince="Province Orientale">
<specimenCount id="9D0D28FBCA14FFE7ECA7FBE343A2FB95" box="[645,679,1037,1064]" pageId="13" pageNumber="458" type="generic">3 ♂</specimenCount>
,
<location id="6E08C3008D431933483C9CE5BD300CBA" LSID="urn:lsid:plazi:treatment:03A25264CA15FFFAEF37FB524211FE1F:6E08C3008D431933483C9CE5BD300CBA" country="Democratic Republic of the Congo" latitude="0.294" longLatPrecision="7" longitude="25.2888" name="Ubundu Road, Yoko Forest, Mokonoka Stream" stateProvince="Province Orientale">Ubundu Road, Yoko Forest, Mokonoka Stream</location>
, rock-bottomed (and some sand, gravel, leaflitter) stream and small tributaries in rainforest,
<quantity id="4CF34E97CA14FFE7EC60FBB5438BFBC0" box="[578,654,1112,1138]" metricMagnitude="2" metricUnit="m" metricValue="4.13" pageId="13" pageNumber="458" unit="m" value="413.0">
<elevation id="A6EB5EC89911B4FA7941EC3928F8218B" box="[578,654,1112,1138]" metricMagnitude="2" metricUnit="m" metricValue="4.13" pageId="13" pageNumber="458" unit="m" value="413.0">413 m</elevation>
</quantity>
a.s.l. (
<geoCoordinate id="EE3F85B5CA14FFE7ECFEFBB5424EFBC0" box="[732,843,1112,1138]" direction="north" orientation="latitude" pageId="13" pageNumber="458" precision="5" value="0.294">0.2940 ° N</geoCoordinate>
<geoCoordinate id="EE3F85B5CA14FFE7ED77FBB542CAFBC0" box="[853,975,1112,1138]" direction="east" orientation="longitude" pageId="13" pageNumber="458" precision="5" value="25.2888">25.2888 ° E</geoCoordinate>
),
<collectingDate id="5BB45E29A40807BF5C300A49AA515E29" value="2010-06-08">
<date id="56BBEB50108EEAA0D5B3A1E496D48D17" value="2010-06-08">08- vi- 2010</date>
</collectingDate>
, leg. K.-
<collectorName id="29BC4CAED7E0F85198B859F6C2B25281">D.B. Dijkstra</collectorName>
,
<collectionCode id="ED1A7BB7CA14FFE7EC35FB934376FB2A" box="[535,627,1150,1176]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" pageId="13" pageNumber="458">RMNH</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B63E92FCA14FFE7ECA3FB934044FA9F" ID-GBIF-Occurrence="3396361338" collectingDate="2012-12-28" collectionCode="RMNH" collectorName="N. Mézière" country="Gabon" elevation="316 m" httpUri="http://www.boldsystems.org/index.php/Public_RecordView?processid=NSPAF498-15" latitude="-0.5596" location="Kougouleu-Medouneu road, Monts de Cristal, Assegone-Essala" longLatPrecision="7" longitude="10.2362" pageId="13" pageNumber="458" specimenCode="RMNH.INS.554425" specimenCount="1" stateProvince="Estuaire Province">
<collectingCountry id="F31CA3E2CA14FFE7ECA3FB9343ECFB2A" box="[641,745,1150,1176]" name="Gabon" pageId="13" pageNumber="458" value="Gabon">GABON</collectingCountry>
(
<collectingRegion id="49CF2D90CA14FFE7ECD9FB9042D6FB2A" box="[763,979,1149,1176]" pageId="13" pageNumber="458">Estuaire Province</collectingRegion>
):
<specimenCount id="9D0D28FBCA14FFE7EDC8FB934509FB25" box="[1002,1036,1149,1176]" pageId="13" pageNumber="458" type="generic">1 ♂</specimenCount>
(
<specimenCode id="A02F2D6024AA1EBC57DCDB9C49C06D45">RMNH.INS.508732</specimenCode>
),
<location id="09CF3043DA057C381DDB813BBE39532F" LSID="urn:lsid:plazi:treatment:03A25264CA15FFFAEF37FB524211FE1F:09CF3043DA057C381DDB813BBE39532F" country="Gabon" latitude="-0.5596" longLatPrecision="7" longitude="10.2362" name="Kougouleu-Medouneu road, Monts de Cristal, Assegone-Essala" stateProvince="Estuaire Province">Kougouleu-Medouneu road, Monts de Cristal, Assegone-Essala</location>
, mountain stream with rapids and small falls (
<quantity id="4CF34E97CA14FFE7ED55FB2542C6FB50" box="[887,963,1224,1250]" metricMagnitude="0" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="13" pageNumber="458" unit="m" value="3.0" valueMax="4.0" valueMin="2.0">24 m</quantity>
wide,
<quantity id="4CF34E97CA14FFE7EEBDFB0341EEFABA" box="[159,235,1262,1288]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="458" unit="cm" value="50.0">50 cm</quantity>
deep),
<quantity id="4CF34E97CA14FFE7EF68FB03409DFABA" box="[330,408,1262,1288]" metricMagnitude="2" metricUnit="m" metricValue="3.16" pageId="13" pageNumber="458" unit="m" value="316.0">
<elevation id="803A10742D0350EF280A29C2E614BA8C" box="[330,408,1262,1288]" metricMagnitude="2" metricUnit="m" metricValue="3.16" pageId="13" pageNumber="458" unit="m" value="316.0">316 m</elevation>
</quantity>
a.s.l. (
<geoCoordinate id="EE3F85B5CA14FFE7EFCBFB004355FABA" box="[489,592,1261,1288]" direction="south" orientation="latitude" pageId="13" pageNumber="458" precision="5" value="-0.5596">0.5596 ° S</geoCoordinate>
<geoCoordinate id="EE3F85B5CA14FFE7EC7EFB0043D3FABA" box="[604,726,1261,1288]" direction="east" orientation="longitude" pageId="13" pageNumber="458" precision="5" value="10.2362">10.2362 ° E</geoCoordinate>
),
<date id="FFB5C5B2CA14FFE7ECCCFB004272FABA" box="[750,887,1261,1288]" pageId="13" pageNumber="458" value="2012-12-28">
<collectingDate id="8D9867D13FBD2C94339C305CF2CC881E" box="[750,887,1261,1288]" pageId="13" pageNumber="458" value="2012-12-28">28 -xii- 2012</collectingDate>
</date>
, leg
<collectorName id="E1C8853A7FA496D4D26ABCE09EF12A56">N. Mézière</collectorName>
,
<collectionCode id="ED1A7BB7CA14FFE7EEC7FAFE4044FA9F" box="[229,321,1299,1325]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" pageId="13" pageNumber="458">RMNH</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B63E92FCA14FFE7EF6EFAFE426BFAE0" ID-GBIF-Occurrence="1224547706" collectingDate="2009-02-21" collectionCode="RMNH" collectorName="N. Mézière" country="Gabon" elevation="618 m" latitude="-1.6643" location="Ndjima" longLatPrecision="7" longitude="13.0958" pageId="13" pageNumber="458" specimenCount="1" stateProvince="Haut-Ogooué Province">
<collectingCountry id="F31CA3E2CA14FFE7EF6EFAFE40B1FA9F" box="[332,436,1299,1325]" name="Gabon" pageId="13" pageNumber="458" value="Gabon">GABON</collectingCountry>
(
<collectingRegion id="49CF2D90CA14FFE7EFE1FAFF43D9FA9F" box="[451,732,1298,1325]" pageId="13" pageNumber="458">Haut-Ogooué Province</collectingRegion>
):
<specimenCount id="9D0D28FBCA14FFE7ECCDFAFE4214FA9E" box="[751,785,1298,1325]" pageId="13" pageNumber="458" type="generic">1 ♂</specimenCount>
,
<location id="DFF492676F7BEC91B2A4424AF9D96FDB" LSID="urn:lsid:plazi:treatment:03A25264CA15FFFAEF37FB524211FE1F:DFF492676F7BEC91B2A4424AF9D96FDB" country="Gabon" latitude="-1.6643" longLatPrecision="7" longitude="13.0958" name="Ndjima" stateProvince="Haut-Ogooué Province">Ndjima</location>
,
<quantity id="4CF34E97CA14FFE7EDA5FAFE42D4FA9F" box="[903,977,1299,1325]" metricMagnitude="2" metricUnit="m" metricValue="6.18" pageId="13" pageNumber="458" unit="m" value="618.0">
<elevation id="09A9164C32154EFC7E7CD851D693566D" box="[903,977,1299,1325]" metricMagnitude="2" metricUnit="m" metricValue="6.18" pageId="13" pageNumber="458" unit="m" value="618.0">618 m</elevation>
</quantity>
a.s.l. (
<geoCoordinate id="EE3F85B5CA14FFE7EE85FAD5400BFAE0" box="[167,270,1336,1362]" direction="south" orientation="latitude" pageId="13" pageNumber="458" precision="5" value="-1.6643">1.6643 ° S</geoCoordinate>
<geoCoordinate id="EE3F85B5CA14FFE7EF3EFAD54093FAE0" box="[284,406,1336,1362]" direction="east" orientation="longitude" pageId="13" pageNumber="458" precision="5" value="13.0958">13.0958 ° E</geoCoordinate>
),
<date id="FFB5C5B2CA14FFE7EF93FAD54329FAE0" box="[433,556,1336,1362]" pageId="13" pageNumber="458" value="2009-02-21">
<collectingDate id="8C5121166AB3C17B38C35D0A13098F13" box="[433,556,1336,1362]" pageId="13" pageNumber="458" value="2009-02-21">21 -ii- 2009</collectingDate>
</date>
, leg.
<collectorName id="7634AEBE4828D3663B7B8BA42C56B4A7">N. Mézière</collectorName>
,
<collectionCode id="ED1A7BB7CA14FFE7ED30FAD4426BFAE0" box="[786,878,1337,1362]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" pageId="13" pageNumber="458">RMNH</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B63E92FCA14FFE7ED5DFAD541FEFA70" ID-GBIF-Occurrence="1224547726" collectingDate="2009-02-18" collectionCode="RMNH" collectorName="N. Mézière" country="Gabon" elevation="507 m" latitude="-1.3998" longLatPrecision="7" longitude="13.192" pageId="13" pageNumber="458" specimenCount="1" stateProvince="Haut-ogooue Province">
<specimenCount id="9D0D28FBCA14FFE7ED5DFAD542A4FAE0" box="[895,929,1336,1362]" pageId="13" pageNumber="458" type="generic">1 ♂</specimenCount>
, Plateau dOkouma, road to Vuku, Mounana, gravel-bottomed stream (
<quantity id="4CF34E97CA14FFE7ED5EFAB042C7FACA" box="[892,962,1373,1400]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="13" pageNumber="458" unit="m" value="1.5" valueMax="2.0" valueMin="1.0">12 m</quantity>
wide) in grassland,
<quantity id="4CF34E97CA14FFE7EF19FA6E4080FA2F" box="[315,389,1411,1437]" metricMagnitude="2" metricUnit="m" metricValue="5.07" pageId="13" pageNumber="458" unit="m" value="507.0">
<elevation id="57DA0D26CFA572983D597534D54E71AB" box="[315,389,1411,1437]" metricMagnitude="2" metricUnit="m" metricValue="5.07" pageId="13" pageNumber="458" unit="m" value="507.0">507 m</elevation>
</quantity>
a.s.l. (
<geoCoordinate id="EE3F85B5CA14FFE7EFEDFA6F4333FA2F" box="[463,566,1410,1437]" direction="south" orientation="latitude" pageId="13" pageNumber="458" precision="5" value="-1.3998">1.3998 ° S</geoCoordinate>
<geoCoordinate id="EE3F85B5CA14FFE7EC1CFA6F43BDFA2F" box="[574,696,1410,1437]" direction="east" orientation="longitude" pageId="13" pageNumber="458" precision="5" value="13.192">13.1920 ° E</geoCoordinate>
),
<date id="FFB5C5B2CA14FFE7ECEEFA6F4242FA2F" box="[716,839,1410,1437]" pageId="13" pageNumber="458" value="2009-02-18">
<collectingDate id="56F252DBBC3FC3D6034308968440127F" box="[716,839,1410,1437]" pageId="13" pageNumber="458" value="2009-02-18">18 -ii- 2009</collectingDate>
</date>
, leg.
<collectorName id="F7A0335488271E05A2BB96672CE840DC">N. Mézière</collectorName>
,
<collectionCode id="ED1A7BB7CA14FFE7EEBDFA4441FEFA70" box="[159,251,1449,1474]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" pageId="13" pageNumber="458">RMNH</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B63E92FCA14FFE4EF24FA45434DFEAA" ID-GBIF-Occurrence="3396361376" collectingDate="2012-03-27" collectionCode="RMNH" collectorName="N. Mézière" country="Gabon" elevation="483 m" httpUri="http://www.boldsystems.org/index.php/Public_RecordView?processid=NSPAF375-15" lastPageId="14" lastPageNumber="459" latitude="-0.6476" location="righthand road before “ Africa No 1 ” radio station in the direction of Franceville, Moyabi" longLatPrecision="7" longitude="13.68" pageId="13" pageNumber="458" specimenCode="RMNH.INS.554425" specimenCount="1" stateProvince="Haut-ogooue Province">
<specimenCount id="9D0D28FBCA14FFE7EF24FA45402DFA70" box="[262,296,1448,1474]" pageId="13" pageNumber="458" type="generic">1 ♂</specimenCount>
(
<specimenCode id="2147BDDF749585367A407E099B8FE5B1">RMNH.INS.506164</specimenCode>
),
<location id="23C4D409A56D428B44A470C726706E4B" LSID="urn:lsid:plazi:treatment:03A25264CA15FFFAEF37FB524211FE1F:23C4D409A56D428B44A470C726706E4B" country="Gabon" latitude="-0.6476" longLatPrecision="7" longitude="13.68" name="righthand road before “ Africa No 1 ” radio station in the direction of Franceville, Moyabi" stateProvince="Haut-ogooue Province">righthand road before “Africa No 1 ” radio station in the direction of Franceville, Moyabi</location>
, sandy forest river (
<quantity id="4CF34E97CA14FFE7EDFEFA204509FA5A" box="[988,1036,1485,1512]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="13" pageNumber="458" unit="m" value="4.0">4 m</quantity>
wide,&gt;
<quantity id="4CF34E97CA14FFE7EED4FA1E403AF9BF" box="[246,319,1523,1549]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="458" unit="cm" value="50.0">50 cm</quantity>
deep) bordered by ponds,
<quantity id="4CF34E97CA14FFE7ECA2FA1E43CFF9BF" box="[640,714,1523,1549]" metricMagnitude="2" metricUnit="m" metricValue="4.83" pageId="13" pageNumber="458" unit="m" value="483.0">
<elevation id="C21C64B0A58C75E8EC55F801AA3EF1D3" box="[640,714,1523,1549]" metricMagnitude="2" metricUnit="m" metricValue="4.83" pageId="13" pageNumber="458" unit="m" value="483.0">483 m</elevation>
</quantity>
a.s.l. (
<geoCoordinate id="EE3F85B5CA14FFE7ED34FA1F4278F9BF" box="[790,893,1522,1549]" direction="south" orientation="latitude" pageId="13" pageNumber="458" precision="5" value="-0.6476">0.6476 ° S</geoCoordinate>
<geoCoordinate id="EE3F85B5CA14FFE7EDA7FA1F42FAF9BF" box="[901,1023,1522,1549]" direction="east" orientation="longitude" pageId="13" pageNumber="458" precision="5" value="13.68">13.6800° E</geoCoordinate>
),
<date id="FFB5C5B2CA17FFE4EEBDFF104027FEAA" box="[159,290,253,280]" pageId="14" pageNumber="459" value="2012-03-27">
<collectingDate id="87A2B896E8D6BE7D039B2B7606E7F425" box="[159,290,253,280]" pageId="14" pageNumber="459" value="2012-03-27">27 -iii- 2012</collectingDate>
</date>
, leg.
<collectorName id="EC8A5D9820F1572306C7C49B918FC2F0">N. Mézière</collectorName>
,
<collectionCode id="ED1A7BB7CA17FFE4EFCEFF13434DFEAA" box="[492,584,254,280]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" pageId="14" pageNumber="459">RMNH</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B63E92FCA17FFE4EC71FF1342DDFED0" ID-GBIF-Occurrence="1224547710" collectingDate="2012-11-04" collectionCode="RMNH" collectorName="N. Mézière" country="Gabon" elevation="474 m" lastPageId="14" lastPageNumber="459" latitude="-1.6877" location="righthand road before “ Africa No 1 ” radio station in the direction of Franceville, Moyabi" longLatPrecision="7" longitude="13.3113" pageId="14" pageNumber="459" specimenCode="RMNH.INS.554425" specimenCount="1" stateProvince="Haut-ogooue Province">
<specimenCount id="9D0D28FBCA17FFE4EC71FF134370FEA5" box="[595,629,253,280]" pageId="14" pageNumber="459" type="generic">1 ♂</specimenCount>
(
<specimenCode id="0EF87D6D79B4438FCD5CFC0D30B7AA38">RMNH.INS.554468</specimenCode>
), same locality, small gravelly and sandy forest stream bordered by swamp and springs,
<quantity id="4CF34E97CA17FFE4EEBDFEA541EDFED0" box="[159,232,328,354]" metricMagnitude="2" metricUnit="m" metricValue="4.74" pageId="14" pageNumber="459" unit="m" value="474.0">
<elevation id="0581ACFF124C6B52CF25DE0C179B7D1F" box="[159,232,328,354]" metricMagnitude="2" metricUnit="m" metricValue="4.74" pageId="14" pageNumber="459" unit="m" value="474.0">474 m</elevation>
</quantity>
a.s.l. (
<geoCoordinate id="EE3F85B5CA17FFE4EF11FEA5409FFED0" box="[307,410,328,354]" direction="south" orientation="latitude" pageId="14" pageNumber="459" precision="5" value="-1.6877">1.6877 °S</geoCoordinate>
<geoCoordinate id="EE3F85B5CA17FFE4EF83FEA5431EFED0" box="[417,539,328,354]" direction="east" orientation="longitude" pageId="14" pageNumber="459" precision="5" value="13.3113">13.3113 °E</geoCoordinate>
),
<date id="FFB5C5B2CA17FFE4EC0DFEA543B4FED0" box="[559,689,328,354]" pageId="14" pageNumber="459" value="2012-11-04">
<collectingDate id="1016815B4815BFB42245031855C5300D" box="[559,689,328,354]" pageId="14" pageNumber="459" value="2012-11-04">04-xi- 2012</collectingDate>
</date>
, leg.
<collectorName id="2C767728A826C9645736D995AFD48121">N. Mézière</collectorName>
,
<collectionCode id="ED1A7BB7CA17FFE4ED5EFEA442DDFED0" box="[892,984,329,354]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" pageId="14" pageNumber="459">RMNH</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B63E92FCA17FFE4EDC1FEA54062FE60" ID-GBIF-Occurrence="1224547736" collectingDate="2013-09-24" collectionCode="CJKL" collectorName="J. Kipping" country="Gabon" elevation="398 m" latitude="-1.7065" location="Moyabi, concession Rougier, 17 km SSE of Moanda" longLatPrecision="7" longitude="13.2558" pageId="14" pageNumber="459" specimenCount="8" stateProvince="Haut-ogooue Province">
<specimenCount id="9D0D28FBCA17FFE4EDC1FEA54500FED0" box="[995,1029,328,354]" pageId="14" pageNumber="459" type="generic">8 ♂</specimenCount>
,
<location id="D300F9C39C913AF497BFED8E7677EC02" LSID="urn:lsid:plazi:treatment:03A25264CA15FFFAEF37FB524211FE1F:D300F9C39C913AF497BFED8E7677EC02" country="Gabon" latitude="-1.7065" longLatPrecision="7" longitude="13.2558" name="Moyabi, concession Rougier, 17 km SSE of Moanda" stateProvince="Haut-ogooue Province">
Moyabi, concession Rougier,
<quantity id="4CF34E97CA17FFE4EFDAFE834346FE3A" box="[504,579,365,392]" metricMagnitude="4" metricUnit="m" metricValue="1.7" pageId="14" pageNumber="459" unit="km" value="17.0">17 km</quantity>
SSE of Moanda
</location>
, large sandy stream in dense forest (
<figureCitation id="1330FFF7CA17FFE4EF1DFE7F40A5FE1F" box="[319,416,402,429]" captionStart="Photo 2" captionStartId="16.[159,225,1353,1378]" captionText="Photo 2. Sandy forest stream near Moyabi, Gabon. Habitat of Umma gumma, Paragomphus dispar and Trithemis legrandi. Photo: JK (24 - ix- 2013)" httpUri="http://dx.doi.org/10.5281/zenodo.35442" pageId="14" pageNumber="459">Photo 2</figureCitation>
),
<quantity id="4CF34E97CA17FFE4EF9BFE7E4300FE1F" box="[441,517,403,429]" metricMagnitude="2" metricUnit="m" metricValue="3.9799999999999995" pageId="14" pageNumber="459" unit="m" value="398.0">
<elevation id="E10B96E42A576531ADDFDB82509765F8" box="[441,517,403,429]" metricMagnitude="2" metricUnit="m" metricValue="3.9799999999999995" pageId="14" pageNumber="459" unit="m" value="398.0">398 m</elevation>
</quantity>
a.s.l. (
<geoCoordinate id="EE3F85B5CA17FFE4EC71FE7F43BFFE1F" box="[595,698,402,429]" direction="south" orientation="latitude" pageId="14" pageNumber="459" precision="5" value="-1.7065">1.7065 °S</geoCoordinate>
<geoCoordinate id="EE3F85B5CA17FFE4ECE6FE7F4238FE1F" box="[708,829,402,429]" direction="east" orientation="longitude" pageId="14" pageNumber="459" precision="5" value="13.2558">13.2558 °E</geoCoordinate>
),
<date id="FFB5C5B2CA17FFE4ED76FE7F42D0FE1F" box="[852,981,402,429]" pageId="14" pageNumber="459" value="2013-09-24">
<collectingDate id="F5A0ABD85510AA48D7A7DB4C2DBF9F64" box="[852,981,402,429]" pageId="14" pageNumber="459" value="2013-09-24">24 -ix- 2013</collectingDate>
</date>
, leg.
<collectorName id="C5B91953FD4A4CCA8D10101FD5AFC783">J. Kipping</collectorName>
,
<collectionCode id="ED1A7BB7CA17FFE4EF00FE554062FE60" box="[290,359,440,466]" collectionName="CJKL" pageId="14" pageNumber="459">CJKL</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B63E92FCA17FFE4EF53FE5542DDFE60" ID-GBIF-Occurrence="1224547705" box="[369,984,440,466]" collectingDate="2013-09-24" collectionCode="CAGF" collectorName="A. Günther" country="Gabon" elevation="398 m" latitude="-1.7065" location="Moyabi, concession Rougier, 17 km SSE of Moanda" longLatPrecision="7" longitude="13.2558" pageId="14" pageNumber="459" specimenCount="1" specimenCount-male="1" stateProvince="Haut-ogooue Province">
<specimenCount id="9D0D28FBCA17FFE4EF53FE554096FE60" box="[369,403,440,466]" pageId="14" pageNumber="459" type="generic">1 ♂</specimenCount>
, same locality, same date, leg.
<collectorName id="2CE9CAD44E9637749CBA0A8FCA260F89">A. Günther</collectorName>
,
<collectionCode id="ED1A7BB7CA17FFE4EDAEFE5542DDFE60" box="[908,984,440,466]" collectionName="CAGF" pageId="14" pageNumber="459">CAGF</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B63E92FCA17FFE4EDC1FE5540C4FE4A" ID-GBIF-Occurrence="1224547717" box="[369,984,440,466]" collectingDate="2013-09-17" collectionCode="CAGF" collectorName="A. Günther" country="Gabon" elevation="398 m" latitude="-1.7065" location="Moyabi, concession Rougier, 17 km SSE of Moanda" longLatPrecision="7" longitude="13.2558" pageId="14" pageNumber="459" specimenCount="1" stateProvince="Haut-ogooue Province">
<specimenCount id="9D0D28FBCA17FFE4EDC1FE554500FE60" box="[995,1029,440,466]" pageId="14" pageNumber="459" type="generic">1 ♂</specimenCount>
, same locality,
<date id="FFB5C5B2CA17FFE4EF62FE3040C4FE4A" box="[320,449,477,504]" pageId="14" pageNumber="459" value="2013-09-17">
<collectingDate id="FF7B1E9C124D9ADA44AC29B76FE26A15" box="[320,449,477,504]" pageId="14" pageNumber="459" value="2013-09-17">17 -ix-2013</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation id="3B63E92FCA17FFE4EFE9FE33450DFE4A" ID-GBIF-Occurrence="1224547718" box="[459,1032,477,504]" collectingDate="2013-10-01" collectionCode="CJKL" collectorName="J. Kipping" country="Gabon" elevation="398 m" latitude="-1.7065" location="Moyabi, concession Rougier, 17 km SSE of Moanda" longLatPrecision="7" longitude="13.2558" pageId="14" pageNumber="459" specimenCount="1" specimenCount-male="1" stateProvince="Haut-ogooue Province">
<specimenCount id="9D0D28FBCA17FFE4EFE9FE3340E8FE45" box="[459,493,477,504]" pageId="14" pageNumber="459" type="generic">1♂</specimenCount>
same locality,
<date id="FFB5C5B2CA17FFE4ECB6FE334208FE4A" box="[660,781,478,504]" pageId="14" pageNumber="459" value="2013-10-01">
<collectingDate id="E3067B387C0B7DDE2BFCC86DB3D243EB" box="[660,781,478,504]" pageId="14" pageNumber="459" value="2013-10-01">01-x- 2013</collectingDate>
</date>
, leg.
<collectorName id="B2759FA444D79DFD6273638CA17C3313">J. Kipping</collectorName>
,
<collectionCode id="ED1A7BB7CA17FFE4EDE6FE33450DFE4A" box="[964,1032,478,504]" collectionName="CJKL" pageId="14" pageNumber="459">CJKL</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CA17FFE4EEBDFDC54317FD6A" pageId="14" pageNumber="459" type="diagnosis">
<paragraph id="8BB4E372CA17FFE4EEBDFDC5400CFDF1" box="[159,265,552,579]" pageId="14" pageNumber="459">
<emphasis id="B97F3F60CA17FFE4EEBDFDC5400CFDF1" bold="true" box="[159,265,552,579]" pageId="14" pageNumber="459">Genetics</emphasis>
</paragraph>
<paragraph id="8BB4E372CA17FFE4EEBDFDA340ACFD3F" pageId="14" pageNumber="459">
Four unique haplotypes (n= 4) nearest to but distinct from eleven of true
<taxonomicName id="4C0B98F1CA17FFE4EEBDFD9F4046FD3F" box="[159,323,626,653]" class="Insecta" family="Calopterygidae" genus="Umma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="459" phylum="Arthropoda" rank="species" species="longistigma">
<emphasis id="B97F3F60CA17FFE4EEBDFD9F4046FD3F" box="[159,323,626,653]" italics="true" pageId="14" pageNumber="459">U.longistigma</emphasis>
</taxonomicName>
(n = 14).
</paragraph>
<paragraph id="8BB4E372CA17FFE4EEBDFD534317FD6A" box="[159,530,702,728]" pageId="14" pageNumber="459">
<heading id="D0FC541ECA17FFE4EEBDFD534317FD6A" bold="true" box="[159,530,702,728]" fontSize="11" level="3" pageId="14" pageNumber="459" reason="6">
<emphasis id="B97F3F60CA17FFE4EEBDFD534317FD6A" bold="true" box="[159,530,702,728]" pageId="14" pageNumber="459">Male morphological diagnosis</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CA17FFE5EEBDFD0F450EFD6A" lastPageId="15" lastPageNumber="460" pageId="14" pageNumber="459" type="discussion">
<paragraph id="8BB4E372CA17FFE5EEBDFD0F450EFD6A" lastPageId="15" lastPageNumber="460" pageId="14" pageNumber="459">
Similar to sympatric sister-species
<taxonomicName id="4C0B98F1CA17FFE4EC09FD0E43C9FD4F" box="[555,716,738,765]" class="Insecta" family="Calopterygidae" genus="Umma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="459" phylum="Arthropoda" rank="species" species="longistigma">
<emphasis id="B97F3F60CA17FFE4EC09FD0E43C9FD4F" box="[555,716,738,765]" italics="true" pageId="14" pageNumber="459">U. longistigma</emphasis>
</taxonomicName>
by the combination of (a) a uniformly green to blue metallic head, thorax and abdomen, but dark brown to black legs and poststernum; (b) the largely pale rather than dark mandibular bases; (c) the sparse whitish hairs on the poststernum, rather than a dense patch of long dark hairs; (d) 610 cross-veins in the Fw quadrilateral cell; (e)the large Pt with an acute proximal corner, 2.7 3.2 mm; (f) the absence of a clump of thick bristles near the tips of the cerci; and (g)paraprocts that are almost as long as the cerci (
<figureCitation id="1330FFF7CA16FFE5EC2AFECF434CFE8F" box="[520,585,290,317]" captionStart="Figure 1" captionStartId="15.[751,820,1352,1377]" captionTargetBox="[159,708,1067,1550]" captionTargetId="figure@15.[156,716,1049,1550]" captionTargetPageId="15" captionText="Figure 1. Male cerci in dorsal view (left) and paraprocts in ventral view with dotted outline of cerci (right) of Umma gumma sp. nov. and U. longistigma." httpUri="http://dx.doi.org/10.5281/zenodo.35441" pageId="15" pageNumber="460">Fig. 1</figureCitation>
). However, is (1)larger with Hw 31.5 35.0 mm (mean 33.6; n = 15) rather than 30.033.0 mm (31.9; n = 20), with notably more robust build; (2) has a largely pale anterior face to each basal antennal segment, which is entirely dark in most
<taxonomicName id="4C0B98F1CA16FFE5ECC0FE7E4280FE1F" box="[738,901,402,429]" class="Insecta" family="Calopterygidae" genus="Umma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="460" phylum="Arthropoda" rank="species" species="longistigma">
<emphasis id="B97F3F60CA16FFE5ECC0FE7E4280FE1F" box="[738,901,402,429]" italics="true" pageId="15" pageNumber="460">U. longistigma</emphasis>
</taxonomicName>
specimens, although about a third have indistinct or partial markings; (3)no expansion on the bend of the peniss lateral lobes, but their tips are more widened instead; (4)wide-based cerci that do not expand notably distally and end in unmodified rounded or squarish tips, while in
<taxonomicName id="4C0B98F1CA16FFE5ECACFDC4422BFDF1" box="[654,814,552,579]" class="Insecta" family="Calopterygidae" genus="Umma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="460" phylum="Arthropoda" rank="species" species="longistigma">
<emphasis id="B97F3F60CA16FFE5ECACFDC4422BFDF1" box="[654,814,552,579]" italics="true" pageId="15" pageNumber="460">U. longistigma</emphasis>
</taxonomicName>
the internal flanges widen notably towards the tips, which bear a transverse ridge and hollow, with often a subapical notch or tooth where the flange and ridge meet; and (5) paraprocts that are broad throughout with square-cut tips bearing an inward-directed tooth, rather than slender and tapering to rounded hooked tips (
<figureCitation id="1330FFF7CA16FFE5ED9EFD5042FFFD6A" box="[956,1018,701,728]" captionStart="Figure 1" captionStartId="15.[751,820,1352,1377]" captionTargetBox="[159,708,1067,1550]" captionTargetId="figure@15.[156,716,1049,1550]" captionTargetPageId="15" captionText="Figure 1. Male cerci in dorsal view (left) and paraprocts in ventral view with dotted outline of cerci (right) of Umma gumma sp. nov. and U. longistigma." httpUri="http://dx.doi.org/10.5281/zenodo.35441" pageId="15" pageNumber="460">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF74B3FACA17FFE4EEBDFA1E421DF9BE" box="[159,792,1523,1548]" httpUri="http://dx.doi.org/10.5281/zenodo.35441" pageId="14" pageNumber="459" targetBox="[158,1035,991,1480]" targetPageId="14">
<paragraph id="8BB4E372CA17FFE4EEBDFA1E421DF9BE" box="[159,792,1523,1548]" pageId="14" pageNumber="459">
<emphasis id="B97F3F60CA17FFE4EEBDFA1E4033F9BE" bold="true" box="[159,310,1523,1548]" pageId="14" pageNumber="459">Type Photo 1.</emphasis>
Holotype of
<taxonomicName id="4C0B98F1CA17FFE4EFE0FA1E4361F9B9" box="[450,612,1523,1547]" class="Insecta" family="Calopterygidae" genus="Umma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="459" phylum="Arthropoda" rank="species" species="gumma" status="sp. nov.">
<emphasis id="B97F3F60CA17FFE4EFE0FA1E4361F9B9" box="[450,612,1523,1547]" italics="true" pageId="14" pageNumber="459">Umma gumma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C821BCA17FFE4EC48FA1E43C5F9BE" box="[618,704,1523,1548]" pageId="14" pageNumber="459" rank="species">sp. nov.</taxonomicNameLabel>
, RMNH.
</paragraph>
</caption>
<subSubSection id="C311B0F9CA16FFE5EEBDFCE44081FC0A" pageId="15" pageNumber="460" type="etymology">
<paragraph id="8BB4E372CA16FFE5EEBDFCE44021FC91" box="[159,292,777,803]" pageId="15" pageNumber="460">
<emphasis id="B97F3F60CA16FFE5EEBDFCE44021FC91" bold="true" box="[159,292,777,803]" pageId="15" pageNumber="460">Etymology</emphasis>
</paragraph>
<paragraph id="8BB4E372CA16FFE5EEBDFCC34081FC0A" pageId="15" pageNumber="460">
Name refers to the classic 1969 Pink Floyd album »Ummagumma« (noun in apposition).
<heading id="D0FC541ECA16FFE5EEBDFC734081FC0A" bold="true" box="[159,388,926,952]" fontSize="11" level="3" pageId="15" pageNumber="460" reason="6">
<emphasis id="B97F3F60CA16FFE5EEBDFC734081FC0A" bold="true" box="[159,388,926,952]" pageId="15" pageNumber="460">Range and ecology</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CA16FFFAEEBDFC2F4211FE1F" lastPageId="16" lastPageNumber="461" pageId="15" pageNumber="460" type="discussion">
<paragraph id="8BB4E372CA16FFFAEEBDFC2F4211FE1F" lastPageId="16" lastPageNumber="461" pageId="15" pageNumber="460">
Three
<taxonomicName id="4C0B98F1CA16FFE5EECFFC2E40D4FC6F" authority="Kirby, 1890" authorityName="Kirby" authorityYear="1890" box="[237,465,962,989]" class="Insecta" family="Calopterygidae" genus="Umma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="460" phylum="Arthropoda" rank="genus">
<emphasis id="B97F3F60CA16FFE5EECFFC2E403EFC6F" box="[237,315,963,989]" italics="true" pageId="15" pageNumber="460">Umma</emphasis>
<bibRefCitation id="3F625E3B08D8B726A253E5853DA066EB" author="Kirby W. F." journalOrPublisher="With an appendix of fossil species. Gurney &amp; Jackson, London" refId="ref68008" refString="Kirby W. F. 1890. A synonymic catalogue of Neuroptera Odonata, or dragon-flies. With an appendix of fossil species. Gurney &amp; Jackson, London" title="A synonymic catalogue of Neuroptera Odonata, or dragon-flies" type="book" year="1890">Kirby, 1890</bibRefCitation>
</taxonomicName>
species overlap widely in central Africas rainforests:
<taxonomicName id="4C0B98F1CA16FFE5EED8FC044344FBB0" authority="Selys, 1879" authorityName="Selys" authorityYear="1879" box="[250,577,1000,1027]" class="Insecta" family="Calopterygidae" genus="Umma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="460" phylum="Arthropoda" rank="species" species="mesostigma">
<emphasis id="B97F3F60CA16FFE5EED8FC0440A7FBB1" box="[250,418,1001,1027]" italics="true" pageId="15" pageNumber="460">U. mesostigma</emphasis>
(Selys, 1879)
</taxonomicName>
including the probably synonymous
<taxonomicName id="4C0B98F1CA09FFFAEEBDFF1040C9FEAA" authority="Forster, 1916" authorityName="Forster" authorityYear="1916" box="[159,460,253,280]" class="Insecta" family="Calopterygidae" genus="Umma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="461" phylum="Arthropoda" rank="species" species="saphirina">
<emphasis id="B97F3F60CA09FFFAEEBDFF104029FEAA" box="[159,300,253,280]" italics="true" pageId="16" pageNumber="461">U.saphirina</emphasis>
Förster, 1916
</taxonomicName>
is found closest to the source on often gravelly sections, while
<taxonomicName id="4C0B98F1CA09FFFAEF76FECE40FFFE8F" box="[340,506,290,317]" class="Insecta" family="Calopterygidae" genus="Umma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="461" phylum="Arthropoda" rank="species" species="longistigma">
<emphasis id="B97F3F60CA09FFFAEF76FECE40FFFE8F" box="[340,506,290,317]" italics="true" pageId="16" pageNumber="461">U. longistigma</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0B98F1CA09FFFAEC15FECE43B4FE8F" box="[567,689,291,317]" class="Insecta" family="Calopterygidae" genus="Umma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="461" phylum="Arthropoda" rank="species" species="gumma">
<emphasis id="B97F3F60CA09FFFAEC15FECE43B4FE8F" box="[567,689,291,317]" italics="true" pageId="16" pageNumber="461">U. gumma</emphasis>
</taxonomicName>
appear to co-occur on larger and often sandy streams (
<figureCitation id="1330FFF7CA09FFFAEFECFEA5432AFED0" box="[462,559,328,354]" captionStart="Photo 2" captionStartId="16.[159,225,1353,1378]" captionText="Photo 2. Sandy forest stream near Moyabi, Gabon. Habitat of Umma gumma, Paragomphus dispar and Trithemis legrandi. Photo: JK (24 - ix- 2013)" httpUri="http://dx.doi.org/10.5281/zenodo.35444" pageId="16" pageNumber="461">Photo 2</figureCitation>
). The new species has been recorded up to
<quantity id="4CF34E97CA09FFFAEE9FFE834000FE3A" box="[189,261,366,392]" metricMagnitude="2" metricUnit="m" metricValue="6.2" pageId="16" pageNumber="461" unit="m" value="620.0">620 m</quantity>
a.s.l. and favours less shady sections, occurs at lower densities, and comes down to the water only with brighter sunshine.
</paragraph>
</subSubSection>
</treatment>
</document>