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<document id="7B7AB63E3832A041D886459D26A522CE" ID-CLB-Dataset="9218" ID-DOI="10.11646/zootaxa.4896.2.7" ID-GBIF-Dataset="7fbdd6a3-8010-4616-965d-241515af8423" ID-ISSN="1175-5326" ID-Zenodo-Dep="4381842" ID-ZooBank="71E90DA6-7595-4010-A849-0EBA20D15921" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1608543618368" checkinUser="plazi" docAuthor="Bota-Sierra, Cornelio Andrés &amp; Novelo-Gutiérrez, Rodolfo" docDate="2020" docId="03A087FA2319FF86FF3D64CDFD9BBABC" docLanguage="en" docName="zootaxa.4896.2.7.pdf" docOrigin="Zootaxa 4896 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Epigomphus brillantina Bota-Sierra &amp; Novelo-Gutiérrez 2020, sp. nov." docType="treatment" docVersion="10" lastPageNumber="275" masterDocId="FF99FF82231FFF8CFFAA6245FF94BE1D" masterDocTitle="Two new species of Colombian Epigomphus (Odonata: Gomphidae)" masterLastPageNumber="276" masterPageNumber="265" pageNumber="271" updateTime="1698847844051" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="7FFC569A73DE0537BB89A5C8CD3E8657" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="2FA75D598328F2521B01622051390FA1">
<mods:title id="8725B047E711CBBF50CBD662E276BF04">Two new species of Colombian Epigomphus (Odonata: Gomphidae)</mods:title>
</mods:titleInfo>
<mods:name id="4FABA89CD508DCF76B303C7085F372AE" type="personal">
<mods:role id="19BDB61A5C4F8F62DEC625A88E8100F2">
<mods:roleTerm id="87EDC0CAB71A16738E12F24BFDC48AAA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="73F7A6DA63CCFF96CBD6BAA1956FC72D">Bota-Sierra, Cornelio Andrés</mods:namePart>
<mods:affiliation id="B7B5A26213DECC78F03D2FC0A5F40787">Instituto de Ecología, A. C. Red de Biodiversidad y Sistemática. Carretera Antigua a Coatepec 351, El Haya, 91070 Xalapa, Veracruz, MEXICO. &amp; Grupo de Entomología Universidad de Antioquia (GEUA), Universidad de Antioquia, Medellín, Antioquia, COLOMBIA.</mods:affiliation>
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<mods:role id="F4A3D6012CE642AACA400DEB2C7A8A31">
<mods:roleTerm id="0B53B431C7FA7C9A94BCC1083C0DD616">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="93D04E72EF7A495A72064D89045DBCC4">Novelo-Gutiérrez, Rodolfo</mods:namePart>
<mods:affiliation id="A350DFC250E9B0ACCBA88DD0A677E3C7">Instituto de Ecología, A. C. Red de Biodiversidad y Sistemática. Carretera Antigua a Coatepec 351, El Haya, 91070 Xalapa, Veracruz, MEXICO. &amp; rodolfo. novelo @ inecol. mx; http: // orcid. org / 0000 - 0003 - 3163 - 2820</mods:affiliation>
<mods:nameIdentifier id="0DB31D54051A258EFE25AE7E8F312C4A" type="email">rodolfo.novelo@inecol.mx</mods:nameIdentifier>
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<mods:title id="5E4B81C22A3E4B1B5AAFEEDD911C5203">Zootaxa</mods:title>
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<mods:part id="428D02245D4FDD858E8D9289F94D723E">
<mods:date id="33447FE217C773E6F8D1C5AE3477C1D1">2020</mods:date>
<mods:detail id="0875DE4A28953FAE791B100C982A4EE4" type="pubDate">
<mods:number id="72207D75A82B29FF6B08B5DF89C7EFDC">2020-12-21</mods:number>
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<mods:detail id="F5CB335CBEF927F76BD2F6CFAC4B98C1" type="volume">
<mods:number id="429CFAF7C5406481EF63D4A1E93BC4FC">4896</mods:number>
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<mods:number id="4FBDA745B86541EDA98659FAF50B7F3A">2</mods:number>
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<mods:classification id="DFCB29390376CAF2D18CBDE6300E10D1">journal article</mods:classification>
<mods:identifier id="5769E31D1678754C6D9711CE98C4DEC3" type="CLB-Dataset">9218</mods:identifier>
<mods:identifier id="6CB8772CF57013B31E589FA6258049F9" type="DOI">10.11646/zootaxa.4896.2.7</mods:identifier>
<mods:identifier id="3B95FA4FD9411ADFF6C4F9515C160F10" type="GBIF-Dataset">7fbdd6a3-8010-4616-965d-241515af8423</mods:identifier>
<mods:identifier id="C15CA00A974875C46E7B663F3B3B8020" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03A087FA2319FF86FF3D64CDFD9BBABC" ID-DOI="http://doi.org/10.5281/zenodo.4382789" ID-GBIF-Taxon="176234988" ID-Zenodo-Dep="4382789" LSID="urn:lsid:plazi:treatment:03A087FA2319FF86FF3D64CDFD9BBABC" httpUri="http://treatment.plazi.org/id/03A087FA2319FF86FF3D64CDFD9BBABC" lastPageId="10" lastPageNumber="275" pageId="6" pageNumber="271">
<subSubSection id="C31365672319FF8AFF3D64CDFD89B8BF" box="[151,541,1672,1699]" pageId="6" pageNumber="271" type="nomenclature">
<paragraph id="8BB636EC2319FF8AFF3D64CDFD89B8BF" blockId="6.[151,541,1672,1735]" box="[151,541,1672,1699]" pageId="6" pageNumber="271">
<heading id="D0FE81802319FF8AFF3D64CDFD89B8BF" bold="true" box="[151,541,1672,1699]" fontSize="11" level="1" pageId="6" pageNumber="271" reason="1">
<emphasis id="B97DEAFE2319FF8AFF3D64CDFD89B8BF" bold="true" box="[151,541,1672,1699]" pageId="6" pageNumber="271">
<taxonomicName id="4C094D6F2319FF8AFF3D64CDFE2DB8BE" ID-CoL="9X2ML" authority="Bota-Sierra &amp; Novelo-Gutiérrez, 2020" authorityName="Bota-Sierra &amp; Novelo-Gutiérrez" authorityYear="2020" box="[151,441,1672,1699]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="271" phylum="Arthropoda" rank="species" species="brillantina" status="sp. nov.">
<emphasis id="B97DEAFE2319FF8AFF3D64CDFE2DB8BE" bold="true" box="[151,441,1672,1699]" italics="true" pageId="6" pageNumber="271">Epigomphus brillantina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57852319FF8AFE6A64CDFD89B8BF" box="[448,541,1672,1698]" pageId="6" pageNumber="271" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31365672319FF8AFF3D64E9FE86B8DA" box="[151,274,1708,1735]" pageId="6" pageNumber="271" type="description">
<paragraph id="8BB636EC2319FF8AFF3D64E9FE86B8DA" blockId="6.[151,541,1672,1735]" box="[151,274,1708,1735]" pageId="6" pageNumber="271">
(
<figureCitation id="13322A692319FF8AFF3564E9FE9EB8DA" box="[159,266,1708,1735]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStartId-0="7.[151,250,1724,1748]" captionStartId-1="8.[151,250,1913,1937]" captionStartId-2="9.[151,250,1573,1597]" captionTargetBox-0="[296,1292,237,1673]" captionTargetBox-1="[201,1387,381,1885]" captionTargetBox-2="[156,1426,181,1549]" captionTargetId-0="figure-54@7.[296,1292,237,1701]" captionTargetId-1="figure-153@8.[201,1387,381,1889]" captionTargetId-2="figure-17@9.[156,1430,181,1549]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionText-0="FIGURE 6. Epigomphus brillantina habitus: a, teneral specimen live in the field (photograph by Camilo Flórez); b, male holotype." captionText-1="FIGURE 7. Epigomphus brillantina: male paratype dorsal view of head; male holotype: b, lateral view of S2; c, ventral view of S2. a. h.: anterior hamuli; a. l.: anterior lamina; cor: cornua; hrn: horns; l. l. p.: lateral lobe slender projection; p. h.: posterior hamuli; prep: prepuce; s1: first segment of vesica spermalis; s3: third segment of vesica spermalis." captionText-2="FIGURE 8. Epigomphus brillantina male paratype S10 and caudal appendages: a, dorsal view; b ventral view; c, posterior view; d, lateral view. ap: apical process; da. e. p.: dorsoapical external process; da. i. p.: dorsoapical internal process; m. p.: medial process; s. d.: small denticles." figureDoi-0="http://doi.org/10.5281/zenodo.4381866" figureDoi-1="http://doi.org/10.5281/zenodo.4381872" figureDoi-2="http://doi.org/10.5281/zenodo.4381874" httpUri-0="https://zenodo.org/record/4381866/files/figure.png" httpUri-1="https://zenodo.org/record/4381872/files/figure.png" httpUri-2="https://zenodo.org/record/4381874/files/figure.png" pageId="6" pageNumber="271">Figs. 6–8</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31365672319FF8AFF3D64B1FC4CB94B" pageId="6" pageNumber="271" type="materials_examined">
<paragraph id="8BB636EC2319FF8AFF3D64B1FC4CB94B" blockId="6.[151,1436,1780,2023]" pageId="6" pageNumber="271">
<emphasis id="B97DEAFE2319FF8AFF3D64B1FE10B913" bold="true" box="[151,388,1780,1806]" pageId="6" pageNumber="271">Material examined:</emphasis>
<materialsCitation id="3B613CB12319FF8AFE2664B0FDC9B94A" ID-GBIF-Occurrence="3392786301" collectingDate="2014-05-28" collectionCode="CEUA" collectorName="Leg &amp; C. Florez &amp; C. Bota." country="Colombia" county="Remedios Municipality" elevation="558" location="Finca La Brillantina" municipality="La Cruz Township" pageId="6" pageNumber="271" specimenCode="CEUA 81502" specimenCount="3" specimenCount-male="2" stateProvince="Antioquia Department" typeStatus="holotype">
Two adult males (
<specimenCount id="9D0FFD652319FF8AFDF364B1FD00B90D" box="[601,660,1780,1808]" pageId="6" pageNumber="271" type="male">2 ♂♂</specimenCount>
).
<collectionCode id="ED18AE292319FF8AFD1964B1FC90B913" box="[691,772,1780,1806]" pageId="6" pageNumber="271">CEUA</collectionCode>
:
<emphasis id="B97DEAFE2319FF8AFCA564B1FCE8B913" bold="true" box="[783,892,1780,1806]" pageId="6" pageNumber="271">
<typeStatus id="54B2884E2319FF8AFCA564B1FCE8B913" box="[783,892,1780,1806]" pageId="6" pageNumber="271" type="holotype">Holotype</typeStatus>
</emphasis>
.
<specimenCount id="9D0FFD652319FF8AFC2164B0FC09B90D" box="[907,925,1781,1808]" pageId="6" pageNumber="271" type="male">♂</specimenCount>
,
<specimenCode id="DBAF9E972319FF8AFC1A64B1FBD8B912" box="[944,1100,1780,1807]" collectionCode="CEUA" pageId="6" pageNumber="271">CEUA 81502</specimenCode>
,
<date id="FFB7102C2319FF8AFBF264B1FB67B913" box="[1112,1267,1780,1807]" pageId="6" pageNumber="271" value="2014-05-28">
<collectingDate id="EFF3E9C42319FF8AFBF264B1FB67B913" box="[1112,1267,1780,1807]" pageId="6" pageNumber="271" value="2014-05-28">28 May 2014</collectingDate>
</date>
,
<collectingCountry id="F31E767C2319FF8AFAAA64B1FA0CB913" box="[1280,1432,1780,1806]" name="Colombia" pageId="6" pageNumber="271">COLOMBIA</collectingCountry>
,
<collectingRegion id="49CDF80E2319FF8AFF3D655DFE02B92E" box="[151,406,1816,1843]" country="Colombia" name="Antioquia" pageId="6" pageNumber="271">Antioquia Department</collectingRegion>
,
<collectingCounty id="62D74E602319FF8AFE09655DFD39B92F" box="[419,685,1816,1842]" pageId="6" pageNumber="271">Remedios Municipality</collectingCounty>
,
<collectingMunicipality id="6BD2AC962319FF8AFD13655CFC1AB92F" box="[697,910,1816,1843]" pageId="6" pageNumber="271">La Cruz Township</collectingMunicipality>
,
<location id="8ED660372319FF8AFC30655DFBEBB92F" LSID="urn:lsid:plazi:treatment:03A087FA2319FF86FF3D64CDFD9BBABC:8ED660372319FF8AFC30655DFBEBB92F" box="[922,1151,1816,1843]" country="Colombia" county="Remedios Municipality" municipality="La Cruz Township" name="Finca La Brillantina" pageId="6" pageNumber="271" stateProvince="Antioquia Department">Finca La Brillantina</location>
<geoCoordinate id="EE3D502B2319FF8AFB2D655DFB69B92E" box="[1159,1277,1816,1843]" pageId="6" pageNumber="271">6.881105°</geoCoordinate>
,
<geoCoordinate id="EE3D502B2319FF8AFAA3655DFA0CB92E" box="[1289,1432,1816,1843]" pageId="6" pageNumber="271">-74.569295°</geoCoordinate>
;
<elevation id="0024D1DF2319FF8AFF3D6579FE8FB94B" box="[151,283,1852,1879]" metricMagnitude="2" metricUnit="m" metricValue="5.58" pageId="6" pageNumber="271" unit="m" value="558.0">
<quantity id="4CF19B092319FF8AFF3D6579FF4AB94A" box="[151,222,1852,1879]" metricMagnitude="2" metricUnit="m" metricValue="5.58" pageId="6" pageNumber="271" unit="m" value="558.0">558 m</quantity>
a.s.l.
</elevation>
,
<collectorName id="26FC533A2319FF8AFE8D6578FEC3B94A" box="[295,343,1853,1879]" pageId="6" pageNumber="271">Leg</collectorName>
:
<collectorName id="26FC533A2319FF8AFECB6579FE5EB94B" box="[353,458,1852,1878]" pageId="6" pageNumber="271">C. Flórez</collectorName>
and
<collectorName id="26FC533A2319FF8AFDAA6579FDC9B94A" box="[512,605,1852,1879]" pageId="6" pageNumber="271">C. Bota.</collectorName>
</materialsCitation>
<materialsCitation id="3B613CB12319FF8AFDCE6579FC4CB94B" ID-GBIF-Occurrence="3003801302" box="[612,984,1852,1880]" collectingDate="2014-05-28" collectionCode="CEUA" collectorName="C. Florez &amp; C. Bota." country="Colombia" county="Remedios Municipality" elevation="558" location="Finca La Brillantina" municipality="La Cruz Township" pageId="6" pageNumber="271" specimenCode="CEUA 81502" specimenCount="1" specimenCount-male="1" stateProvince="Antioquia" typeStatus="paratype">
<emphasis id="B97DEAFE2319FF8AFDCE6579FD45B94B" bold="true" box="[612,721,1852,1878]" pageId="6" pageNumber="271">
<typeStatus id="54B2884E2319FF8AFDCE6579FD45B94B" box="[612,721,1852,1878]" pageId="6" pageNumber="271" type="paratype">Paratype</typeStatus>
</emphasis>
,
<specimenCount id="9D0FFD652319FF8AFD746579FC9CB945" box="[734,776,1852,1880]" pageId="6" pageNumber="271" type="male">1 ♂</specimenCount>
same as holotype.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31365672319FF8AFF6D6525FB43B9FB" pageId="6" pageNumber="271" type="etymology">
<paragraph id="8BB636EC2319FF8AFF6D6525FB43B9FB" blockId="6.[151,1436,1780,2023]" pageId="6" pageNumber="271">
<emphasis id="B97DEAFE2319FF8AFF6D6525FEC6B967" bold="true" box="[199,338,1888,1914]" pageId="6" pageNumber="271">Etymology:</emphasis>
<taxonomicName id="4C094D6F2319FF8AFEF36525FE45B967" authorityName="Bota-Sierra &amp; Novelo-Gutiérrez" authorityYear="2020" box="[345,465,1888,1914]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="271" phylum="Arthropoda" rank="species" species="brillantina">
<emphasis id="B97DEAFE2319FF8AFEF36525FE45B967" box="[345,465,1888,1914]" italics="true" pageId="6" pageNumber="271">brillantina</emphasis>
</taxonomicName>
(to be treated as an indeclinable noun) is the word in Spanish for brilliantine, referring to something shiny. It is the name of the forest island protected and preserved by Don Gustavo Campuzano and his family for the last 30 years, which is a refugee to many species of this region’s flora and fauna, including several endemic and threatened
<taxonomicName id="4C094D6F2319FF8AFE026589FD93B9FB" authorityName="Fabricius" authorityYear="1793" box="[424,519,1996,2022]" class="Insecta" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="271" phylum="Arthropoda" rank="order">Odonata</taxonomicName>
species and the only location so far known for this new species.
</paragraph>
</subSubSection>
<subSubSection id="C31365672318FF86FF6D62D3FCB7BC98" lastPageId="10" lastPageNumber="275" pageId="7" pageNumber="272" type="description">
<paragraph id="8BB636EC2318FF8BFF6D62D3FA1EBEC8" blockId="7.[151,1436,150,213]" pageId="7" pageNumber="272">
<emphasis id="B97DEAFE2318FF8BFF6D62D3FE49BEAC" bold="true" box="[199,477,150,177]" pageId="7" pageNumber="272">
Description of
<typeStatus id="54B2884E2318FF8BFEDC62D2FE49BEAC" box="[374,477,151,177]" pageId="7" pageNumber="272" type="holotype">holotype</typeStatus>
</emphasis>
.
<emphasis id="B97DEAFE2318FF8BFE4162D2FDBCBEAC" box="[491,552,151,177]" italics="true" pageId="7" pageNumber="272">Head</emphasis>
brown, frons with two yellow greenish stripes separated by a small brown portion on the middle, clypeus with dorsolateral green spots. Occiput with two small diagonal scars (
<figureCitation id="13322A692318FF8BFB7162FEFABCBEC8" box="[1243,1320,187,213]" captionStart="FIGURE 6" captionStartId="7.[151,250,1724,1748]" captionTargetBox="[296,1292,237,1673]" captionTargetId="figure-54@7.[296,1292,237,1701]" captionTargetPageId="7" captionText="FIGURE 6. Epigomphus brillantina habitus: a, teneral specimen live in the field (photograph by Camilo Flórez); b, male holotype." figureDoi="http://doi.org/10.5281/zenodo.4381866" httpUri="https://zenodo.org/record/4381866/files/figure.png" pageId="7" pageNumber="272">Figs. 6</figureCitation>
a–b, 7a).
</paragraph>
<caption id="DF7666642318FF8BFF3D64F9FF4AB8E4" ID-DOI="http://doi.org/10.5281/zenodo.4381866" ID-Zenodo-Dep="4381866" httpUri="https://zenodo.org/record/4381866/files/figure.png" pageId="7" pageNumber="272" startId="7.[151,250,1724,1748]" targetBox="[296,1292,237,1673]" targetPageId="7">
<paragraph id="8BB636EC2318FF8BFF3D64F9FF4AB8E4" blockId="7.[151,1436,1724,1785]" pageId="7" pageNumber="272">
<emphasis id="B97DEAFE2318FF8BFF3D64F9FE80B8C9" bold="true" box="[151,276,1724,1748]" pageId="7" pageNumber="272">FIGURE 6.</emphasis>
<taxonomicName id="4C094D6F2318FF8BFEB164F8FD99B8C8" authorityName="Bota-Sierra &amp; Novelo-Gutiérrez" authorityYear="2020" box="[283,525,1725,1749]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="272" phylum="Arthropoda" rank="species" species="brillantina">
<emphasis id="B97DEAFE2318FF8BFEB164F8FD99B8C8" box="[283,525,1725,1749]" italics="true" pageId="7" pageNumber="272">Epigomphus brillantina</emphasis>
</taxonomicName>
habitus: a, teneral specimen live in the field (photograph by Camilo Flórez); b, male holotype.
</paragraph>
</caption>
<paragraph id="8BB636EC2318FF84FF6D6562FB17BF78" blockId="7.[151,1437,1831,2037]" lastBlockId="8.[151,1436,151,357]" lastPageId="8" lastPageNumber="273" pageId="7" pageNumber="272">
<emphasis id="B97DEAFE2318FF8BFF6D6562FE8CB95C" box="[199,280,1831,1857]" italics="true" pageId="7" pageNumber="272">Thorax</emphasis>
. Brown, but yellow as follows: pronotum except the posterior lobe, a longitudinal line on mesepisternum and a transverse line on its anterior border, second pale antehumeral stripe reduced to a single, small, posterior spot, a longitudinal line on mesepimeron, a stripe on metepisternum which apex converges with the completely yellow metepimeron, antealar crest with yellow spots (
<figureCitation id="13322A692318FF8BFD7265D6FCB2B9B3" box="[728,806,1939,1966]" captionStart="FIGURE 6" captionStartId="7.[151,250,1724,1748]" captionTargetBox="[296,1292,237,1673]" captionTargetId="figure-54@7.[296,1292,237,1701]" captionTargetPageId="7" captionText="FIGURE 6. Epigomphus brillantina habitus: a, teneral specimen live in the field (photograph by Camilo Flórez); b, male holotype." figureDoi="http://doi.org/10.5281/zenodo.4381866" httpUri="https://zenodo.org/record/4381866/files/figure.png" pageId="7" pageNumber="272">Figs. 6</figureCitation>
a–b). Legs brown except distal portion of femora, tibiae, and tarsi which are black. Metafemora approximately 1 ½ the length of mesofemora (
<figureCitation id="13322A692318FF8BFBF165F2FB3DB9CF" box="[1115,1193,1975,2002]" captionStart="FIGURE 6" captionStartId="7.[151,250,1724,1748]" captionTargetBox="[296,1292,237,1673]" captionTargetId="figure-54@7.[296,1292,237,1701]" captionTargetPageId="7" captionText="FIGURE 6. Epigomphus brillantina habitus: a, teneral specimen live in the field (photograph by Camilo Flórez); b, male holotype." figureDoi="http://doi.org/10.5281/zenodo.4381866" httpUri="https://zenodo.org/record/4381866/files/figure.png" pageId="7" pageNumber="272">Figs. 6</figureCitation>
a–b). All femora with external row of spines. Tibiae with long, slender spines on external border, except protibiae with a row of ten scalelike spines on external side. Tarsal claws brown with subapical tooth.
<emphasis id="B97DEAFE2317FF84FC0E62D2FC7CBEAD" box="[932,1000,151,176]" italics="true" pageId="8" pageNumber="273">Wings</emphasis>
. Hyaline, black veins and brown Pt. Pt covering 9 ½ cells in FW, 7 ½ in right and 8 ½ cells in left HW. Left FW with 17 Ax, right FW with 18, FWs with 15 Px; left HW with 14 Ax, right HW with 13 Ax; left HW with 12 Px, right HW with 13 Px; supratriangles, triangles and subtriangles without crossveins. Second reinforced antenodal in FWs 6th, in the right HW the 6th and in the left the 7th (
<figureCitation id="13322A692317FF84FF596362FED1BF5C" box="[243,325,295,321]" captionStart="FIGURE 4" captionStartId="4.[151,250,1872,1896]" captionTargetBox="[148,1439,525,1834]" captionTargetId="figure-190@4.[148,1439,525,1849]" captionTargetPageId="4" captionText="FIGURE 4. Epigomphus rufus male paratype abdomen: caudal appendages: a, lateral view; c, dorsal view. b, S2 lateral view; d, S2 ventral view. a. h.: anterior hamuli; a. l.: anterior lamina; a. t.: apical tooth; p. h.: posterior hamuli; s1: first segment of vesica spermalis; sa. t.: subapical tooth." figureDoi="http://doi.org/10.5281/zenodo.4381858" httpUri="https://zenodo.org/record/4381858/files/figure.png" pageId="8" pageNumber="273">Fig. 4a</figureCitation>
). Cubito-anal crossveins in addition to inner side of subtriangle,
<quantity id="4CF19B092317FF84FBB26362FBD7BF5C" box="[1048,1091,295,321]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="8" pageNumber="273" unit="in" value="3.0">3 in</quantity>
FWs,
<quantity id="4CF19B092317FF84FB266362FB23BF5C" box="[1164,1207,295,321]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="8" pageNumber="273" unit="in" value="2.0">2 in</quantity>
HWs. Crossveins in space between sectors of arculus and point of branching of RP on HW
<quantity id="4CF19B092317FF84FC1A630EFC4FBF78" box="[944,987,331,357]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="8" pageNumber="273" unit="in" value="3.0">3 in</quantity>
left,
<quantity id="4CF19B092317FF84FBBF630EFBD4BF78" box="[1045,1088,331,357]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="8" pageNumber="273" unit="in" value="4.0">4 in</quantity>
right.
</paragraph>
<caption id="DF7666642317FF84FF3D653CFBE9B9C4" ID-DOI="http://doi.org/10.5281/zenodo.4381872" ID-Zenodo-Dep="4381872" httpUri="https://zenodo.org/record/4381872/files/figure.png" pageId="8" pageNumber="273" startId="8.[151,250,1913,1937]" targetBox="[201,1387,381,1885]" targetPageId="8">
<paragraph id="8BB636EC2317FF84FF3D653CFBE9B9C4" blockId="8.[151,1436,1913,2010]" pageId="8" pageNumber="273">
<emphasis id="B97DEAFE2317FF84FF3D653CFE80B98F" bold="true" box="[151,276,1913,1938]" pageId="8" pageNumber="273">FIGURE 7.</emphasis>
<emphasis id="B97DEAFE2317FF84FEB1653FFD81B98F" box="[283,533,1914,1938]" italics="true" pageId="8" pageNumber="273">
<taxonomicName id="4C094D6F2317FF84FEB1653FFD9BB98F" authorityName="Bota-Sierra &amp; Novelo-Gutiérrez" authorityYear="2020" box="[283,527,1914,1938]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="273" phylum="Arthropoda" rank="species" species="brillantina">Epigomphus brillantina</taxonomicName>
:
</emphasis>
male paratype dorsal view of head; male holotype: b, lateral view of S2; c, ventral view of S2. a. h.: anterior hamuli; a. l.: anterior lamina; cor: cornua; hrn: horns; l. l. p.: lateral lobe slender projection; p. h.: posterior hamuli; prep: prepuce; s1: first segment of vesica spermalis; s3: third segment of vesica spermalis.
</paragraph>
</caption>
<caption id="DF7666642316FF85FF3D6460FD85B898" ID-DOI="http://doi.org/10.5281/zenodo.4381874" ID-Zenodo-Dep="4381874" httpUri="https://zenodo.org/record/4381874/files/figure.png" pageId="9" pageNumber="274" startId="9.[151,250,1573,1597]" targetBox="[156,1426,181,1549]" targetPageId="9">
<paragraph id="8BB636EC2316FF85FF3D6460FD85B898" blockId="9.[151,1436,1573,1669]" pageId="9" pageNumber="274">
<emphasis id="B97DEAFE2316FF85FF3D6460FE81B820" bold="true" box="[151,277,1573,1597]" pageId="9" pageNumber="274">FIGURE 8.</emphasis>
<taxonomicName id="4C094D6F2316FF85FEB46460FD85B820" authorityName="Bota-Sierra &amp; Novelo-Gutiérrez" authorityYear="2020" box="[286,529,1573,1597]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="274" phylum="Arthropoda" rank="species" species="brillantina">
<emphasis id="B97DEAFE2316FF85FEB46460FD85B820" box="[286,529,1573,1597]" italics="true" pageId="9" pageNumber="274">Epigomphus brillantina</emphasis>
</taxonomicName>
male paratype S10 and caudal appendages: a, dorsal view; b ventral view; c, posterior view; d, lateral view. ap: apical process; da. e. p.: dorsoapical external process; da. i. p.: dorsoapical internal process; m. p.: medial process; s. d.: small denticles.
</paragraph>
</caption>
<paragraph id="8BB636EC2316FF86FF6D64F6FAFCBFCF" blockId="9.[151,1437,1715,2030]" lastBlockId="10.[151,1437,151,1185]" lastPageId="10" lastPageNumber="275" pageId="9" pageNumber="274">
<emphasis id="B97DEAFE2316FF85FF6D64F6FEA5B8D0" box="[199,305,1715,1741]" italics="true" pageId="9" pageNumber="274">Abdomen</emphasis>
. Club-like, becoming thinner on S3 and widening again on S8, S10 is the widest segment (
<figureCitation id="13322A692316FF85FABD64F6FAF7B8D3" box="[1303,1379,1715,1742]" captionStart="FIGURE 6" captionStartId="7.[151,250,1724,1748]" captionTargetBox="[296,1292,237,1673]" captionTargetId="figure-54@7.[296,1292,237,1701]" captionTargetPageId="7" captionText="FIGURE 6. Epigomphus brillantina habitus: a, teneral specimen live in the field (photograph by Camilo Flórez); b, male holotype." figureDoi="http://doi.org/10.5281/zenodo.4381866" httpUri="https://zenodo.org/record/4381866/files/figure.png" pageId="9" pageNumber="274">Figs. 6</figureCitation>
a–b). Dark brown but S1 yellow except posterodorsal portion, S2 auricles and lateroventral portion yellow, S3 with an anterolateral yellow spot and a thin yellow posterolateral line, S4–6 with anterior yellow rings, S7 anterior ¾ yellow, S8 ventrolateral margin yellow (
<figureCitation id="13322A692316FF85FD98655AFDEAB927" box="[562,638,1823,1850]" captionStart="FIGURE 6" captionStartId="7.[151,250,1724,1748]" captionTargetBox="[296,1292,237,1673]" captionTargetId="figure-54@7.[296,1292,237,1701]" captionTargetPageId="7" captionText="FIGURE 6. Epigomphus brillantina habitus: a, teneral specimen live in the field (photograph by Camilo Flórez); b, male holotype." figureDoi="http://doi.org/10.5281/zenodo.4381866" httpUri="https://zenodo.org/record/4381866/files/figure.png" pageId="9" pageNumber="274">Figs. 6</figureCitation>
a–b). Auricles rounded and dorsally swollen with a posterobasal process bearing rows of three black teeth directed ventrally (
<figureCitation id="13322A692316FF85FD526506FCD2B943" box="[760,838,1859,1886]" captionStart="FIGURE 7" captionStartId="8.[151,250,1913,1937]" captionTargetBox="[201,1387,381,1885]" captionTargetId="figure-153@8.[201,1387,381,1889]" captionTargetPageId="8" captionText="FIGURE 7. Epigomphus brillantina: male paratype dorsal view of head; male holotype: b, lateral view of S2; c, ventral view of S2. a. h.: anterior hamuli; a. l.: anterior lamina; cor: cornua; hrn: horns; l. l. p.: lateral lobe slender projection; p. h.: posterior hamuli; prep: prepuce; s1: first segment of vesica spermalis; s3: third segment of vesica spermalis." figureDoi="http://doi.org/10.5281/zenodo.4381872" httpUri="https://zenodo.org/record/4381872/files/figure.png" pageId="9" pageNumber="274">Fig 7c</figureCitation>
). Components of genital fossa dark brown, anterior lamina posterior border sinuose in ventral view with a medial concavity (
<figureCitation id="13322A692316FF85FC746522FBBDB99F" box="[990,1065,1895,1922]" captionStart="FIGURE 7" captionStartId="8.[151,250,1913,1937]" captionTargetBox="[201,1387,381,1885]" captionTargetId="figure-153@8.[201,1387,381,1889]" captionTargetPageId="8" captionText="FIGURE 7. Epigomphus brillantina: male paratype dorsal view of head; male holotype: b, lateral view of S2; c, ventral view of S2. a. h.: anterior hamuli; a. l.: anterior lamina; cor: cornua; hrn: horns; l. l. p.: lateral lobe slender projection; p. h.: posterior hamuli; prep: prepuce; s1: first segment of vesica spermalis; s3: third segment of vesica spermalis." figureDoi="http://doi.org/10.5281/zenodo.4381872" httpUri="https://zenodo.org/record/4381872/files/figure.png" pageId="9" pageNumber="274">Fig 7c</figureCitation>
); anterior hamuli petiolate, widening towards apical portion, ending in a rounded point (
<figureCitation id="13322A692316FF85FC8865CEFCF8B9BB" box="[802,876,1931,1958]" captionStart="FIGURE 7" captionStartId="8.[151,250,1913,1937]" captionTargetBox="[201,1387,381,1885]" captionTargetId="figure-153@8.[201,1387,381,1889]" captionTargetPageId="8" captionText="FIGURE 7. Epigomphus brillantina: male paratype dorsal view of head; male holotype: b, lateral view of S2; c, ventral view of S2. a. h.: anterior hamuli; a. l.: anterior lamina; cor: cornua; hrn: horns; l. l. p.: lateral lobe slender projection; p. h.: posterior hamuli; prep: prepuce; s1: first segment of vesica spermalis; s3: third segment of vesica spermalis." figureDoi="http://doi.org/10.5281/zenodo.4381872" httpUri="https://zenodo.org/record/4381872/files/figure.png" pageId="9" pageNumber="274">Fig 7c</figureCitation>
); posterior hamuli long, approximately twice the length of anterior hamuli, slightly curved backwards in lateral view (
<figureCitation id="13322A692316FF85FC6B65EAFB9AB9D7" box="[961,1038,1967,1994]" captionStart="FIGURE 7" captionStartId="8.[151,250,1913,1937]" captionTargetBox="[201,1387,381,1885]" captionTargetId="figure-153@8.[201,1387,381,1889]" captionTargetPageId="8" captionText="FIGURE 7. Epigomphus brillantina: male paratype dorsal view of head; male holotype: b, lateral view of S2; c, ventral view of S2. a. h.: anterior hamuli; a. l.: anterior lamina; cor: cornua; hrn: horns; l. l. p.: lateral lobe slender projection; p. h.: posterior hamuli; prep: prepuce; s1: first segment of vesica spermalis; s3: third segment of vesica spermalis." figureDoi="http://doi.org/10.5281/zenodo.4381872" httpUri="https://zenodo.org/record/4381872/files/figure.png" pageId="9" pageNumber="274">Fig 7b</figureCitation>
), its mesial side covered by black tubercles (
<figureCitation id="13322A692316FF85FEBB6596FECAB9F3" box="[273,350,2003,2030]" captionStart="FIGURE 7" captionStartId="8.[151,250,1913,1937]" captionTargetBox="[201,1387,381,1885]" captionTargetId="figure-153@8.[201,1387,381,1889]" captionTargetPageId="8" captionText="FIGURE 7. Epigomphus brillantina: male paratype dorsal view of head; male holotype: b, lateral view of S2; c, ventral view of S2. a. h.: anterior hamuli; a. l.: anterior lamina; cor: cornua; hrn: horns; l. l. p.: lateral lobe slender projection; p. h.: posterior hamuli; prep: prepuce; s1: first segment of vesica spermalis; s3: third segment of vesica spermalis." figureDoi="http://doi.org/10.5281/zenodo.4381872" httpUri="https://zenodo.org/record/4381872/files/figure.png" pageId="9" pageNumber="274">Fig 7c</figureCitation>
).
<emphasis id="B97DEAFE2316FF85FEDE6591FDA5B9F0" box="[372,561,2003,2029]" italics="true" pageId="9" pageNumber="274">Vesica spermalis</emphasis>
(
<figureCitation id="13322A692316FF85FDE86596FD1BB9F0" box="[578,655,2003,2029]" captionStart="FIGURE 7" captionStartId="8.[151,250,1913,1937]" captionTargetBox="[201,1387,381,1885]" captionTargetId="figure-153@8.[201,1387,381,1889]" captionTargetPageId="8" captionText="FIGURE 7. Epigomphus brillantina: male paratype dorsal view of head; male holotype: b, lateral view of S2; c, ventral view of S2. a. h.: anterior hamuli; a. l.: anterior lamina; cor: cornua; hrn: horns; l. l. p.: lateral lobe slender projection; p. h.: posterior hamuli; prep: prepuce; s1: first segment of vesica spermalis; s3: third segment of vesica spermalis." figureDoi="http://doi.org/10.5281/zenodo.4381872" httpUri="https://zenodo.org/record/4381872/files/figure.png" pageId="9" pageNumber="274">Figs. 7</figureCitation>
b–c, 8a): segment 1 concave in ventral view, in lateral view with a ventroapical rounded process directed distally; segment 3 widening towards its distal portion, two ventro apical horns directed distally; segment 4, prepuce rounded, lateral lobe strongly bulging dorsally, with a slender projection directed posteriorly and parallel to cornua; cornua short. Cercus longer than epiproct and S10 (
<figureCitation id="13322A692315FF86FB65629AFA8ABEE7" box="[1231,1310,223,250]" captionStart="FIGURE 8" captionStartId="9.[151,250,1573,1597]" captionTargetBox="[156,1426,181,1549]" captionTargetId="figure-17@9.[156,1430,181,1549]" captionTargetPageId="9" captionText="FIGURE 8. Epigomphus brillantina male paratype S10 and caudal appendages: a, dorsal view; b ventral view; c, posterior view; d, lateral view. ap: apical process; da. e. p.: dorsoapical external process; da. i. p.: dorsoapical internal process; m. p.: medial process; s. d.: small denticles." figureDoi="http://doi.org/10.5281/zenodo.4381874" httpUri="https://zenodo.org/record/4381874/files/figure.png" pageId="10" pageNumber="275">Figs. 8</figureCitation>
e–f). Cerci concave mesially, in lateral view, with a dorsoapical pointed mesial process which turns downwards (
<figureCitation id="13322A692315FF86FAC86346FF30BF5F" captionStart="FIGURE 8" captionStartId="9.[151,250,1573,1597]" captionTargetBox="[156,1426,181,1549]" captionTargetId="figure-17@9.[156,1430,181,1549]" captionTargetPageId="9" captionText="FIGURE 8. Epigomphus brillantina male paratype S10 and caudal appendages: a, dorsal view; b ventral view; c, posterior view; d, lateral view. ap: apical process; da. e. p.: dorsoapical external process; da. i. p.: dorsoapical internal process; m. p.: medial process; s. d.: small denticles." figureDoi="http://doi.org/10.5281/zenodo.4381874" httpUri="https://zenodo.org/record/4381874/files/figure.png" pageId="10" pageNumber="275">Figs. 8</figureCitation>
e–f), the ventroapical edge has a series of four small denticles (
<figureCitation id="13322A692315FF86FC206362FC7FBF5C" box="[906,1003,295,321]" captionStart="FIGURE 8" captionStartId="9.[151,250,1573,1597]" captionTargetBox="[156,1426,181,1549]" captionTargetId="figure-17@9.[156,1430,181,1549]" captionTargetPageId="9" captionText="FIGURE 8. Epigomphus brillantina male paratype S10 and caudal appendages: a, dorsal view; b ventral view; c, posterior view; d, lateral view. ap: apical process; da. e. p.: dorsoapical external process; da. i. p.: dorsoapical internal process; m. p.: medial process; s. d.: small denticles." figureDoi="http://doi.org/10.5281/zenodo.4381874" httpUri="https://zenodo.org/record/4381874/files/figure.png" pageId="10" pageNumber="275">Figs. 8c</figureCitation>
, e–f), in posterior and dorsal views a rounded dorsoapical external process is observed (
<figureCitation id="13322A692315FF86FD47630EFCA9BF7B" box="[749,829,331,358]" captionStart="FIGURE 8" captionStartId="9.[151,250,1573,1597]" captionTargetBox="[156,1426,181,1549]" captionTargetId="figure-17@9.[156,1430,181,1549]" captionTargetPageId="9" captionText="FIGURE 8. Epigomphus brillantina male paratype S10 and caudal appendages: a, dorsal view; b ventral view; c, posterior view; d, lateral view. ap: apical process; da. e. p.: dorsoapical external process; da. i. p.: dorsoapical internal process; m. p.: medial process; s. d.: small denticles." figureDoi="http://doi.org/10.5281/zenodo.4381874" httpUri="https://zenodo.org/record/4381874/files/figure.png" pageId="10" pageNumber="275">Figs. 8</figureCitation>
b–c). Epiproct divided in two lateroapical branches (
<figureCitation id="13322A692315FF86FF0A632AFF64BF97" box="[160,240,367,394]" captionStart="FIGURE 8" captionStartId="9.[151,250,1573,1597]" captionTargetBox="[156,1426,181,1549]" captionTargetId="figure-17@9.[156,1430,181,1549]" captionTargetPageId="9" captionText="FIGURE 8. Epigomphus brillantina male paratype S10 and caudal appendages: a, dorsal view; b ventral view; c, posterior view; d, lateral view. ap: apical process; da. e. p.: dorsoapical external process; da. i. p.: dorsoapical internal process; m. p.: medial process; s. d.: small denticles." figureDoi="http://doi.org/10.5281/zenodo.4381874" httpUri="https://zenodo.org/record/4381874/files/figure.png" pageId="10" pageNumber="275">Figs. 8</figureCitation>
b–d), each of them with a rounded tubercle on tip and an elevated medial process which is easily observed in posterior view (
<figureCitation id="13322A692315FF86FE6C63D6FD83BFB3" box="[454,535,403,430]" captionStart="FIGURE 8" captionStartId="9.[151,250,1573,1597]" captionTargetBox="[156,1426,181,1549]" captionTargetId="figure-17@9.[156,1430,181,1549]" captionTargetPageId="9" captionText="FIGURE 8. Epigomphus brillantina male paratype S10 and caudal appendages: a, dorsal view; b ventral view; c, posterior view; d, lateral view. ap: apical process; da. e. p.: dorsoapical external process; da. i. p.: dorsoapical internal process; m. p.: medial process; s. d.: small denticles." figureDoi="http://doi.org/10.5281/zenodo.4381874" httpUri="https://zenodo.org/record/4381874/files/figure.png" pageId="10" pageNumber="275">Figs. 8</figureCitation>
b–c); the lateroapical branches converge in a medial oval notch which is not easily observed in dorsal view since it is invaginated under the lateroapical processes of epiproct (
<figureCitation id="13322A692315FF86FB4B63F2FACFBFCF" box="[1249,1371,439,466]" captionStart="FIGURE 8" captionStartId="9.[151,250,1573,1597]" captionTargetBox="[156,1426,181,1549]" captionTargetId="figure-17@9.[156,1430,181,1549]" captionTargetPageId="9" captionText="FIGURE 8. Epigomphus brillantina male paratype S10 and caudal appendages: a, dorsal view; b ventral view; c, posterior view; d, lateral view. ap: apical process; da. e. p.: dorsoapical external process; da. i. p.: dorsoapical internal process; m. p.: medial process; s. d.: small denticles." figureDoi="http://doi.org/10.5281/zenodo.4381874" httpUri="https://zenodo.org/record/4381874/files/figure.png" pageId="10" pageNumber="275">Figs. 8b, d</figureCitation>
).
</paragraph>
<paragraph id="8BB636EC2315FF86FF6D639EFCB7BFE8" blockId="10.[151,1437,151,1185]" box="[199,803,474,501]" pageId="10" pageNumber="275">
<emphasis id="B97DEAFE2315FF86FF6D639EFEE2BFE8" bold="true" box="[199,374,475,501]" pageId="10" pageNumber="275">Measurements</emphasis>
(mm): TL 57, Ab 44, FW 37, HW 35.
</paragraph>
<paragraph id="8BB636EC2315FF86FF6D63BAFAE7BC7C" blockId="10.[151,1437,151,1185]" pageId="10" pageNumber="275">
<emphasis id="B97DEAFE2315FF86FF6D63BAFE25BC05" bold="true" box="[199,433,510,537]" pageId="10" pageNumber="275">
<typeStatus id="54B2884E2315FF86FF6D63BAFEA0BC04" box="[199,308,511,537]" pageId="10" pageNumber="275" type="paratype">Paratype</typeStatus>
variation:
</emphasis>
Pt. Pt covering 8 cells in FW, 9 ½ HW. Ax in FW 16–17, Px in FW 12; Ax, in HW 13, Px in HW 13. Second reinforced antenodal in FWs 6–7th; (
<figureCitation id="13322A692315FF86FD556066FCC4BC20" box="[767,848,547,573]" captionStart="FIGURE 4" captionStartId="4.[151,250,1872,1896]" captionTargetBox="[148,1439,525,1834]" captionTargetId="figure-190@4.[148,1439,525,1849]" captionTargetPageId="4" captionText="FIGURE 4. Epigomphus rufus male paratype abdomen: caudal appendages: a, lateral view; c, dorsal view. b, S2 lateral view; d, S2 ventral view. a. h.: anterior hamuli; a. l.: anterior lamina; a. t.: apical tooth; p. h.: posterior hamuli; s1: first segment of vesica spermalis; sa. t.: subapical tooth." figureDoi="http://doi.org/10.5281/zenodo.4381858" httpUri="https://zenodo.org/record/4381858/files/figure.png" pageId="10" pageNumber="275">Fig. 4a</figureCitation>
). Two cubito-anal crossveins in addition to inner side of subtriangle in FW. Five crossveins in space between sectors of arculus and point of branching of RP on HW.
</paragraph>
<paragraph id="8BB636EC2315FF86FF6D602EFCB7BC98" blockId="10.[151,1437,151,1185]" box="[199,803,618,645]" pageId="10" pageNumber="275">
<emphasis id="B97DEAFE2315FF86FF6D602EFEE2BC98" bold="true" box="[199,374,619,645]" pageId="10" pageNumber="275">Measurements</emphasis>
(mm): TL 56, Ab 43, FW 36, HW 34.
</paragraph>
</subSubSection>
<subSubSection id="C31365672315FF86FF6D60CBFD8FBA28" pageId="10" pageNumber="275" type="diagnosis">
<paragraph id="8BB636EC2315FF86FF6D60CBFD8FBA28" blockId="10.[151,1437,151,1185]" pageId="10" pageNumber="275">
<emphasis id="B97DEAFE2315FF86FF6D60CBFED6BCB5" bold="true" box="[199,322,654,680]" pageId="10" pageNumber="275">Diagnosis.</emphasis>
<taxonomicName id="4C094D6F2315FF86FEE060CAFDC2BCB4" authorityName="Bota-Sierra &amp; Novelo-Gutiérrez" authorityYear="2020" box="[330,598,655,681]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="275" phylum="Arthropoda" rank="species" species="brillantina">
<emphasis id="B97DEAFE2315FF86FEE060CAFDC2BCB4" box="[330,598,655,681]" italics="true" pageId="10" pageNumber="275">Epigomphus brillantina</emphasis>
</taxonomicName>
shares with a group of 16 species, including most of the South American and some Central American species, its second pale antehumeral stripe reduced to a single small posterior spot (
<tableCitation id="C68B03572315FF86FF356092FF67BCEC" box="[159,243,727,753]" captionStart="TABLE 1" captionStartId="6.[151,239,440,464]" captionTargetPageId="6" captionText="TABLE 1. Epigomphus species with second pale antehumeral stripe reduced to a single small posterior spot, their distribution, and male cercus length compared with S10 length." httpUri="http://table.plazi.org/id/DF7666642319FF8AFF3D63FDFC91BFE9" pageId="10" pageNumber="275" tableUuid="DF7666642319FF8AFF3D63FDFC91BFE9">Table 1</tableCitation>
). Among them, the males of eleven species share with
<taxonomicName id="4C094D6F2315FF86FCC96092FC6FBCEC" authorityName="Bota-Sierra &amp; Novelo-Gutiérrez" authorityYear="2020" box="[867,1019,727,753]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="275" phylum="Arthropoda" rank="species" species="brillantina">
<emphasis id="B97DEAFE2315FF86FCC96092FC6FBCEC" box="[867,1019,727,753]" italics="true" pageId="10" pageNumber="275">E. brillantina</emphasis>
</taxonomicName>
cercus length longer than S10 (
<figureCitation id="13322A692315FF86FAC86092FF30BD0B" captionStart="FIGURE 8" captionStartId="9.[151,250,1573,1597]" captionTargetBox="[156,1426,181,1549]" captionTargetId="figure-17@9.[156,1430,181,1549]" captionTargetPageId="9" captionText="FIGURE 8. Epigomphus brillantina male paratype S10 and caudal appendages: a, dorsal view; b ventral view; c, posterior view; d, lateral view. ap: apical process; da. e. p.: dorsoapical external process; da. i. p.: dorsoapical internal process; m. p.: medial process; s. d.: small denticles." figureDoi="http://doi.org/10.5281/zenodo.4381874" httpUri="https://zenodo.org/record/4381874/files/figure.png" pageId="10" pageNumber="275">Figs. 8</figureCitation>
e–f). Within this group,
<taxonomicName id="4C094D6F2315FF86FE1F60BEFDD8BD08" authorityName="Bota-Sierra &amp; Novelo-Gutiérrez" authorityYear="2020" box="[437,588,763,789]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="275" phylum="Arthropoda" rank="species" species="brillantina">
<emphasis id="B97DEAFE2315FF86FE1F60BEFDD8BD08" box="[437,588,763,789]" italics="true" pageId="10" pageNumber="275">E. brillantina</emphasis>
</taxonomicName>
and
<taxonomicName id="4C094D6F2315FF86FD2960BEFC8ABD08" authorityName="Calvert" authorityYear="1903" box="[643,798,763,789]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="275" phylum="Arthropoda" rank="species" species="quadracies">
<emphasis id="B97DEAFE2315FF86FD2960BEFC8ABD08" box="[643,798,763,789]" italics="true" pageId="10" pageNumber="275">E. quadracies</emphasis>
</taxonomicName>
are the only two species that share cercus with a pointed dorsoapical process curved downward easily observed in lateral view (
<figureCitation id="13322A692315FF86FC1F615AFB95BD27" box="[949,1025,799,826]" captionStart="FIGURE 8" captionStartId="9.[151,250,1573,1597]" captionTargetBox="[156,1426,181,1549]" captionTargetId="figure-17@9.[156,1430,181,1549]" captionTargetPageId="9" captionText="FIGURE 8. Epigomphus brillantina male paratype S10 and caudal appendages: a, dorsal view; b ventral view; c, posterior view; d, lateral view. ap: apical process; da. e. p.: dorsoapical external process; da. i. p.: dorsoapical internal process; m. p.: medial process; s. d.: small denticles." figureDoi="http://doi.org/10.5281/zenodo.4381874" httpUri="https://zenodo.org/record/4381874/files/figure.png" pageId="10" pageNumber="275">Figs. 8</figureCitation>
e–f), but
<taxonomicName id="4C094D6F2315FF86FBC1615AFA97BD24" authorityName="Bota-Sierra &amp; Novelo-Gutiérrez" authorityYear="2020" box="[1131,1283,799,825]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="275" phylum="Arthropoda" rank="species" species="brillantina">
<emphasis id="B97DEAFE2315FF86FBC1615AFA97BD24" box="[1131,1283,799,825]" italics="true" pageId="10" pageNumber="275">E. brillantina</emphasis>
</taxonomicName>
can be easily distinguished from
<taxonomicName id="4C094D6F2315FF86FED96106FD9BBD40" authorityName="Calvert" authorityYear="1903" box="[371,527,835,861]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="275" phylum="Arthropoda" rank="species" species="quadracies">
<emphasis id="B97DEAFE2315FF86FED96106FD9BBD40" box="[371,527,835,861]" italics="true" pageId="10" pageNumber="275">E. quadracies</emphasis>
</taxonomicName>
by the shape of its epiproct which has a rounded apical tubercle on each lateroapical process while in
<taxonomicName id="4C094D6F2315FF86FE086122FDABBD9C" authorityName="Calvert" authorityYear="1903" box="[418,575,871,897]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="275" phylum="Arthropoda" rank="species" species="quadracies">
<emphasis id="B97DEAFE2315FF86FE086122FDABBD9C" box="[418,575,871,897]" italics="true" pageId="10" pageNumber="275">E. quadracies</emphasis>
</taxonomicName>
two apical points are observed (
<bibRefCitation id="EF984B1D2315FF86FC1A6122FBD3BD9C" author="Calvert, P. P." box="[944,1095,871,897]" pageId="10" pageNumber="275" pagination="323 - 354" refId="ref5110" refString="Calvert, P. P. (1920) The Costa Rican species of Epigomphus and their mutual mating adaptations (Odonata). Transactions of the American Entomological Society, 46, 323 - 354." type="journal article" year="1920">Calvert 1920</bibRefCitation>
;
<bibRefCitation id="EF984B1D2315FF86FBFE6122FB6EBD9F" author="Kennedy, C. H." box="[1108,1274,871,898]" pageId="10" pageNumber="275" pagination="662 - 666" refId="ref5390" refString="Kennedy, C. H. (1946) Epigomphus subquadrices, a New Dragonfly (Odonata: Gomphidae) from Panama, with notes on E. quadrices and Eugomphus N. Subgen. Annals of the Entomological Society of America, 39, 662 - 666. https: // doi. org / 10.1093 / aesa / 39.4.662" type="journal article" year="1946">Kennedy 1946</bibRefCitation>
) (
<figureCitation id="13322A692315FF86FAB96122FAF4BD9F" box="[1299,1376,871,898]" captionStart="FIGURE 8" captionStartId="9.[151,250,1573,1597]" captionTargetBox="[156,1426,181,1549]" captionTargetId="figure-17@9.[156,1430,181,1549]" captionTargetPageId="9" captionText="FIGURE 8. Epigomphus brillantina male paratype S10 and caudal appendages: a, dorsal view; b ventral view; c, posterior view; d, lateral view. ap: apical process; da. e. p.: dorsoapical external process; da. i. p.: dorsoapical internal process; m. p.: medial process; s. d.: small denticles." figureDoi="http://doi.org/10.5281/zenodo.4381874" httpUri="https://zenodo.org/record/4381874/files/figure.png" pageId="10" pageNumber="275">Figs. 8</figureCitation>
b–d). Also, the median portion of the posterior border of epiproct has a quadrangular notch in
<taxonomicName id="4C094D6F2315FF86FB3361CEFAA3BDB8" authorityName="Calvert" authorityYear="1903" box="[1177,1335,907,933]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="275" phylum="Arthropoda" rank="species" species="quadracies">
<emphasis id="B97DEAFE2315FF86FB3361CEFAA3BDB8" box="[1177,1335,907,933]" italics="true" pageId="10" pageNumber="275">E. quadracies</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF984B1D2315FF86FAE261CEFF46BDD4" author="Calvert, P. P." pageId="10" pageNumber="275" pagination="323 - 354" refId="ref5110" refString="Calvert, P. P. (1920) The Costa Rican species of Epigomphus and their mutual mating adaptations (Odonata). Transactions of the American Entomological Society, 46, 323 - 354." type="journal article" year="1920">Calvert 1920</bibRefCitation>
;
<bibRefCitation id="EF984B1D2315FF86FF7461EAFE16BDD7" author="Kennedy, C. H." box="[222,386,943,970]" pageId="10" pageNumber="275" pagination="662 - 666" refId="ref5390" refString="Kennedy, C. H. (1946) Epigomphus subquadrices, a New Dragonfly (Odonata: Gomphidae) from Panama, with notes on E. quadrices and Eugomphus N. Subgen. Annals of the Entomological Society of America, 39, 662 - 666. https: // doi. org / 10.1093 / aesa / 39.4.662" type="journal article" year="1946">Kennedy 1946</bibRefCitation>
) that can be observed both in ventral and dorsal view, while in
<taxonomicName id="4C094D6F2315FF86FBE861EAFB4DBDD4" authorityName="Bota-Sierra &amp; Novelo-Gutiérrez" authorityYear="2020" box="[1090,1241,943,969]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="275" phylum="Arthropoda" rank="species" species="brillantina">
<emphasis id="B97DEAFE2315FF86FBE861EAFB4DBDD4" box="[1090,1241,943,969]" italics="true" pageId="10" pageNumber="275">E. brillantina</emphasis>
</taxonomicName>
this notch is oval and is not easily observed in dorsal view since it is invaginated under the lateroapical processes of epiproct (
<figureCitation id="13322A692315FF86FACE6196FF30BA0F" captionStart="FIGURE 8" captionStartId="9.[151,250,1573,1597]" captionTargetBox="[156,1426,181,1549]" captionTargetId="figure-17@9.[156,1430,181,1549]" captionTargetPageId="9" captionText="FIGURE 8. Epigomphus brillantina male paratype S10 and caudal appendages: a, dorsal view; b ventral view; c, posterior view; d, lateral view. ap: apical process; da. e. p.: dorsoapical external process; da. i. p.: dorsoapical internal process; m. p.: medial process; s. d.: small denticles." figureDoi="http://doi.org/10.5281/zenodo.4381874" httpUri="https://zenodo.org/record/4381874/files/figure.png" pageId="10" pageNumber="275">Figs. 8</figureCitation>
b–d). Furthermore, the dorsal slender projection of the vesica spermalis fourth segment is longer in
<taxonomicName id="4C094D6F2315FF86FAAF61B2FA08BA0C" authorityName="Bota-Sierra &amp; Novelo-Gutiérrez" authorityYear="2020" box="[1285,1436,1015,1041]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="275" phylum="Arthropoda" rank="species" species="brillantina">
<emphasis id="B97DEAFE2315FF86FAAF61B2FA08BA0C" box="[1285,1436,1015,1041]" italics="true" pageId="10" pageNumber="275">E. brillantina</emphasis>
</taxonomicName>
compared to that of
<taxonomicName id="4C094D6F2315FF86FED3665EFD80BA28" authorityName="Calvert" authorityYear="1903" box="[377,532,1051,1077]" class="Insecta" family="Gomphidae" genus="Epigomphus" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="275" phylum="Arthropoda" rank="species" species="quadracies">
<emphasis id="B97DEAFE2315FF86FED3665EFD80BA28" box="[377,532,1051,1077]" italics="true" pageId="10" pageNumber="275">E. quadracies</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C31365672315FF86FF6D667BFD9BBABC" pageId="10" pageNumber="275" type="distribution">
<paragraph id="8BB636EC2315FF86FF6D667BFD9BBABC" blockId="10.[151,1437,151,1185]" pageId="10" pageNumber="275">
<emphasis id="B97DEAFE2315FF86FF6D667BFE60BA45" bold="true" box="[199,500,1086,1113]" pageId="10" pageNumber="275">Distribution and habitat:</emphasis>
This species is so far only known to occur in one locality in the eastern foothills of the Colombian Central Cordillera, in the
<collectingRegion id="49CDF80E2315FF86FDC86626FD4ABA60" box="[610,734,1123,1149]" country="Colombia" name="Magdalena" pageId="10" pageNumber="275">Magdalena</collectingRegion>
basin (
<figureCitation id="13322A692315FF86FC9A6626FCE0BA60" box="[816,884,1122,1149]" captionStart="FIGURE 5" captionStartId="5.[151,250,1632,1656]" captionTargetBox="[185,1406,746,1601]" captionTargetId="figure-280@5.[180,1406,741,1609]" captionTargetPageId="5" captionText="FIGURE 5. Map with the type localities of Epigomphus rufus and E. brillantina." figureDoi="http://doi.org/10.5281/zenodo.4381862" httpUri="https://zenodo.org/record/4381862/files/figure.png" pageId="10" pageNumber="275">Fig. 5</figureCitation>
). The specimens were found in the interior of the forest close to first order streams.
</paragraph>
</subSubSection>
</treatment>
</document>
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