319 lines
33 KiB
XML
319 lines
33 KiB
XML
<document id="63E735CFA80426166188468222061D86" ID-CLB-Dataset="45364" ID-DOI="10.11646/zootaxa.3826.1.3" ID-GBIF-Dataset="6f4b1535-6c6b-4446-aa26-b1ba83ad9126" ID-ISSN="1175-5326" ID-Zenodo-Dep="228537" ID-ZooBank="53549D58-8F38-47B5-879F-0245E900C131" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460444968431" checkinUser="plazi" docAuthor="Merle, Didier, Pacaud, Jean-Michel, Métais, Grégoire, Bartolini, Annachiara, Lashari, Rafiq A., Brohi, Imdad A., Solangi, Sarfraz H., Marivaux, Laurent & Welcomme, Jean-Loup" docDate="2014" docId="03A087E00E44FFBDFF7FFAD2FB2BFD03" docLanguage="en" docName="zt03826p138.pdf" docOrigin="Zootaxa 3826 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lyrischapa blanfordi Vredenburg 1923, comb. nov." docType="treatment" docVersion="9" lastPageNumber="116" masterDocId="FF99FF980E49FFB2FFE8FFEEFFDAFFAB" masterDocTitle="Volutidae (Mollusca: Gastropoda) of the Lakhra Formation (Earliest Eocene, Sindh, Pakistan): systematics, biostratigraphy and paleobiogeography" masterLastPageNumber="138" masterPageNumber="101" pageNumber="114" updateTime="1698333283847" updateUser="ExternalLinkService">
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<mods:title id="EB5BC411F68785EF130FC97111D9EBE5">Volutidae (Mollusca: Gastropoda) of the Lakhra Formation (Earliest Eocene, Sindh, Pakistan): systematics, biostratigraphy and paleobiogeography</mods:title>
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<mods:namePart id="46E14D10EF7DF2833A661238BCA4DFD3">Merle, Didier</mods:namePart>
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<mods:namePart id="9540CD8F614BF073D4F832D7732DECF8">Pacaud, Jean-Michel</mods:namePart>
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<mods:namePart id="3AEE3326226DA68511F50BA0065EB807">Métais, Grégoire</mods:namePart>
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<mods:namePart id="C7D572ABF993254A44D4DB0648055D98">Bartolini, Annachiara</mods:namePart>
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<mods:namePart id="6FF03DC318CF94020D857A51CEA14B34">Lashari, Rafiq A.</mods:namePart>
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<mods:namePart id="D8E3E97975BA767D25B2DB50662AC14F">Brohi, Imdad A.</mods:namePart>
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<mods:title id="5DEFDD870B7AF766AF4001E81CBBB2F1">Zootaxa</mods:title>
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<mods:date id="0F21FF42FB1341AD346FA28CA5D72CE5">2014</mods:date>
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<mods:number id="D81E68349E4CE75D0DB50DAB49432B92">3826</mods:number>
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<treatment id="03A087E00E44FFBDFF7FFAD2FB2BFD03" ID-DOI="http://doi.org/10.5281/zenodo.6137824" ID-GBIF-Taxon="119509279" ID-Zenodo-Dep="6137824" LSID="urn:lsid:plazi:treatment:03A087E00E44FFBDFF7FFAD2FB2BFD03" httpUri="http://treatment.plazi.org/id/03A087E00E44FFBDFF7FFAD2FB2BFD03" lastPageId="15" lastPageNumber="116" pageId="13" pageNumber="114">
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<paragraph id="8BB636F60E44FFBFFF7FFAD2FCC6FAFD" blockId="13.[151,796,1340,1399]" box="[151,796,1340,1366]" pageId="13" pageNumber="114">
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<heading id="D0FE819A0E44FFBFFF7FFAD2FCC6FAFD" bold="true" box="[151,796,1340,1366]" fontSize="11" level="1" pageId="13" pageNumber="114" reason="1">
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<emphasis id="B97DEAE40E44FFBFFF7FFAD2FCC6FAFD" bold="true" box="[151,796,1340,1366]" pageId="13" pageNumber="114">
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<taxonomicName id="4C094D750E44FFBFFF7FFAD2FD55FAFD" authority="Vredenburg, 1923" authorityName="Vredenburg" authorityYear="1923" box="[151,655,1340,1366]" class="Gastropoda" family="Volutidae" genus="Lyrischapa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="114" phylum="Mollusca" rank="species" species="blanfordi" status="comb. nov.">
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<emphasis id="B97DEAE40E44FFBFFF7FFAD2FE4EFAFD" bold="true" box="[151,404,1340,1366]" italics="true" pageId="13" pageNumber="114">Lyrischapa blanfordi</emphasis>
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(
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<bibRefCitation id="EF984B070E44FFBFFE4CFAD2FD5DFAFD" author="Vredenburg" box="[420,647,1340,1366]" pageId="13" pageNumber="114" refString="Vredenburg, E. W. (1923) Indian Tertiary Gastropoda, IV. Olividae, Harpidae, Marginellidae, Volutidae and Mitridae, with comparative diagnosis of new species. Records of the Geological Survey of India, 54, 243 - 276." type="journal article" year="1923">Vredenburg, 1923</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel id="A24E579F0E44FFBFFD7EFAD2FCC6FAFD" box="[662,796,1340,1366]" pageId="13" pageNumber="114">comb. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BB636F60E44FFBFFF7FFAB0FEC7FADC" blockId="13.[151,796,1340,1399]" box="[151,285,1374,1399]" pageId="13" pageNumber="114">
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(
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<figureCitation id="13322A730E44FFBFFF77FAB0FECFFADC" box="[159,277,1374,1399]" captionStart="FIGURE 6. A – F" captionStartId="14.[151,250,1914,1936]" captionTargetBox="[216,1374,193,1894]" captionTargetId="figure@14.[216,1374,193,1896]" captionTargetPageId="14" captionText="FIGURE 6. A – F. Lyrischapa blanfordi (Vredenburg, 1923), A – C (CPAG. RAN. I. 17, ventral (A), dorsal (B) and apical (C) views, stn 4), D – E (CPAG. RAN. I. 16, ventral (D) and dorsal (E) views, stn 4), F (CPAG. RAN. I. 15, ventral view, stn 4); G – I. Lyrischapa brevispira sp. nov., holotype, CPAG. RAN. I. 18, ventral (G), dorsal (H) and apical (I) views, stn 4. Scale bars: A – H = 10 mm." httpUri="https://zenodo.org/record/228543/files/figure.png" pageId="13" pageNumber="114">Fig. 6A–F</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C313657D0E44FFBFFF7FFA48FD57FA51" pageId="13" pageNumber="114" type="reference_group">
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<paragraph id="8BB636F60E44FFBFFF7FFA48FDE1FA17" blockId="13.[151,1310,1446,1530]" box="[151,571,1446,1468]" pageId="13" pageNumber="114">
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<treatmentCitationGroup id="AB1911D80E44FFBFFF7FFA48FDE1FA17" box="[151,571,1446,1468]" pageId="13" pageNumber="114">
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<taxonomicName id="4C094D750E44FFBFFF7FFA48FDECFA17" authority="Vredenburg, 1923: 255" authorityName="Vredenburg" authorityPageNumber="255" authorityYear="1923" box="[151,566,1446,1468]" class="Gastropoda" family="Volutidae" genus="Athleta" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="114" phylum="Mollusca" rank="species" species="blanfordi">
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<emphasis id="B97DEAE40E44FFBFFF7FFA48FE99FA17" box="[151,323,1446,1468]" italics="true" pageId="13" pageNumber="114">Athleta blanfordi</emphasis>
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<treatmentCitation id="0AA810E70E44FFBFFEA2FA48FDECFA17" author="Vredenburg" box="[330,566,1446,1468]" page="255" pageId="13" pageNumber="114" year="1923">
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<bibRefCitation id="EF984B070E44FFBFFEA2FA48FDD9FA17" author="Vredenburg" box="[330,515,1446,1468]" pageId="13" pageNumber="114" refString="Vredenburg, E. W. (1923) Indian Tertiary Gastropoda, IV. Olividae, Harpidae, Marginellidae, Volutidae and Mitridae, with comparative diagnosis of new species. Records of the Geological Survey of India, 54, 243 - 276." type="journal article" year="1923">Vredenburg, 1923</bibRefCitation>
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: 255
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</treatmentCitation>
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||
</taxonomicName>
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||
.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BB636F60E44FFBFFF7FFA2BFD57FA51" blockId="13.[151,1310,1446,1530]" pageId="13" pageNumber="114">
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<treatmentCitationGroup id="AB1911D80E44FFBFFF7FFA2BFD57FA51" pageId="13" pageNumber="114">
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<taxonomicName id="4C094D750E44FFBFFF7FFA2BFEE2FA70" box="[151,312,1477,1499]" class="Gastropoda" family="Volutidae" genus="Aulicina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="114" phylum="Mollusca" rank="species" species="haimei">
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<emphasis id="B97DEAE40E44FFBFFF7FFA2BFEE2FA70" box="[151,312,1477,1499]" italics="true" pageId="13" pageNumber="114">Aulicina haimei</emphasis>
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</taxonomicName>
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(non
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<taxonomicName id="4C094D750E44FFBFFE9BFA2BFCCBFA70" authority="d'Archiac & Haime, 1853" authorityName="d'Archiac & Haime" authorityYear="1853" box="[371,785,1477,1499]" class="Gastropoda" family="Volutidae" genus="Voluta" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="114" phylum="Mollusca" rank="species" species="haimei">
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<emphasis id="B97DEAE40E44FFBFFE9BFA2BFE25FA70" box="[371,511,1477,1499]" italics="true" pageId="13" pageNumber="114">Voluta haimei</emphasis>
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d’Archiac & Haime, 1853
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</taxonomicName>
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).
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<treatmentCitation id="0AA810E70E44FFBFFCCAFA2BFB85FA70" author="Cossmann" box="[802,1119,1477,1499]" page="25" pageId="13" pageNumber="114" year="1909">
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<bibRefCitation id="EF984B070E44FFBFFCCAFA2BFBE0FA70" author="Cossmann" box="[802,1082,1477,1499]" pageId="13" pageNumber="114" refString="Cossmann, M. & Pissarro, G. (1909) The Mollusca of the Ranikot Series. Part 1, Cephalopoda and Gastropoda. Memoirs of the Geological Survey of India, Palaeontologia Indica, New Series, 3 (1), 1 - 83." type="journal article" year="1909">Cossmann & Pissarro 1909</bibRefCitation>
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: 25
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</treatmentCitation>
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, pl. 2, fig. 27 only.
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<taxonomicName id="4C094D750E44FFBFFF7FFA0AFE99FA51" box="[151,323,1508,1530]" class="Gastropoda" family="Volutidae" genus="Athleta" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="114" phylum="Mollusca" rank="species" species="blanfordi">
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<emphasis id="B97DEAE40E44FFBFFF7FFA0AFE99FA51" box="[151,323,1508,1530]" italics="true" pageId="13" pageNumber="114">Athleta blanfordi</emphasis>
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</taxonomicName>
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. Cotter
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<emphasis id="B97DEAE40E44FFBFFE7EFA0BFE70FA51" box="[406,426,1509,1530]" italics="true" pageId="13" pageNumber="114">in</emphasis>
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||
<treatmentCitation id="0AA810E70E44FFBFFE47FA0AFD53FA51" author="Vredenburg" box="[431,649,1508,1530]" page="36" pageId="13" pageNumber="114" year="1928">
|
||
<bibRefCitation id="EF984B070E44FFBFFE47FA0AFDB9FA51" author="Vredenburg" box="[431,611,1508,1530]" pageId="13" pageNumber="114" refString="Vredenburg, E. W. (1928) A supplement to the Mollusca of the Ranikot Series. Memoirs of the Geological Survey of India, Palaeontologia Indica, New Series, 10 (4), 1 - 25." type="journal article" year="1928">Vredenburg 1928</bibRefCitation>
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: 36
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</treatmentCitation>
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||
.
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</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="C313657D0E44FFBFFF7FF9C8FC5BF860" pageId="13" pageNumber="114" type="description">
|
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<paragraph id="8BB636F60E44FFBFFF7FF9C8FC5BF860" blockId="13.[151,1437,1574,2031]" pageId="13" pageNumber="114">
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<emphasis id="B97DEAE40E44FFBFFF7FF9C8FEF3F994" bold="true" box="[151,297,1574,1599]" pageId="13" pageNumber="114">Description.</emphasis>
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Shell biconic, elongate, H 68–85, D
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<quantity id="4CF19B130E44FFBFFD2DF9C8FC9AF995" box="[709,832,1574,1599]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="13" pageNumber="114" unit="mm" value="40.0" valueMax="50.0" valueMin="30.0">30–50 mm</quantity>
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. Protoconch smooth, globose and slightly deviated, of two whorls, H 6, D
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<quantity id="4CF19B130E44FFBFFE9DF9A5FE1EF9C9" box="[373,452,1611,1635]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="114" unit="mm" value="10.0">10 mm</quantity>
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when complete. Protoconch/teleoconch transition not obvious. Teleoconch of 3 whorls. Spire low, occupying 6% of total shell height. Spire whorls almost flat, separated by shallow suture undulating between shoulder spines of preceding whorl. Last whorl elongate, rather conical. Spiral sculpture on spire of obsolete threads, visible only on first teleoconch whorl. On juvenile specimens of one whorl, last whorl sculptured by very fine threads, covering abapical three quarters of shell, disappearing on succeeding whorls. Axial sculpture of only spiny shoulder costae. Shoulder spines oriented adapically on last whorl. First whorl: 11 costae, becoming more prominent on second half of whorl. Second whorl: 8–9 costae; third whorl: 8–9 costae. No anal sulcus. Aperture narrow, occupying 89% of total height, 18% of diameter. Inner lip straight, columella with six strong folds succeeded posteriorly by two weaker folds; inner lip callus narrow. Outer lip thin, not thickened externally. Siphonal canal short, slightly curved to right towards aperture; siphonal notch weak; siphonal fasciole low, weak, but delineated by carina spreading dorsally on gerontic specimens.
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</paragraph>
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</subSubSection>
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<subSubSection id="C313657D0E44FFBFFF2FF838FAE7F844" box="[199,1341,2006,2031]" pageId="13" pageNumber="114" type="materials_examined">
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<paragraph id="8BB636F60E44FFBFFF2FF838FAE7F844" blockId="13.[151,1437,1574,2031]" box="[199,1341,2006,2031]" pageId="13" pageNumber="114">
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<emphasis id="B97DEAE40E44FFBFFF2FF838FEECF844" bold="true" box="[199,310,2006,2031]" pageId="13" pageNumber="114">Material.</emphasis>
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28 spm (stn 4: 3 spm
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<collectionCode id="ED18AE330E44FFBFFDC6F838FDA5F844" box="[558,639,2006,2031]" pageId="13" pageNumber="114">CPAG</collectionCode>
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.RAN.
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<date id="FFB710360E44FFBFFD2EF838FCCFF844" box="[710,789,2006,2031]" pageId="13" pageNumber="114" value="1915-01-17">I.15–17</date>
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, cast
|
||
<collectionCode id="ED18AE330E44FFBFFCBBF838FC76F844" LSID="urn:lsid:biocol.org:col:34988" box="[851,940,2006,2031]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="13" pageNumber="114">MNHN</collectionCode>
|
||
.F.
|
||
<accessionNumber id="945AAB150E44FFBFFC2AF838FBC4F844" box="[962,1054,2006,2031]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A50353" pageId="13" pageNumber="114" type="EnaNcbi">A50353</accessionNumber>
|
||
–55 and 25 spm
|
||
<collectionCode id="ED18AE330E44FFBFFB3FF839FAE8F844" LSID="urn:lsid:biocol.org:col:34988" box="[1239,1330,2007,2031]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="13" pageNumber="114">MNHN</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF76667E0E47FFBCFF7FF894FF20F840" httpUri="https://zenodo.org/record/228543/files/figure.png" pageId="14" pageNumber="115" targetBox="[216,1374,193,1894]" targetPageId="14">
|
||
<paragraph id="8BB636F60E47FFBCFF7FF894FF20F840" blockId="14.[151,1436,1914,2027]" pageId="14" pageNumber="115">
|
||
<emphasis id="B97DEAE40E47FFBCFF7FF894FE8BF83B" bold="true" box="[151,337,1914,1936]" pageId="14" pageNumber="115">FIGURE 6. A–F.</emphasis>
|
||
<taxonomicName id="4C094D750E47FFBCFEB3F894FCD3F83B" authority="Vredenburg, 1923" authorityName="Vredenburg" authorityYear="1923" box="[347,777,1914,1936]" class="Gastropoda" family="Volutidae" genus="Lyrischapa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="115" phylum="Mollusca" rank="species" species="blanfordi">
|
||
<emphasis id="B97DEAE40E47FFBCFEB3F894FDE9F83B" box="[347,563,1914,1936]" italics="true" pageId="14" pageNumber="115">Lyrischapa blanfordi</emphasis>
|
||
(Vredenburg, 1923)
|
||
</taxonomicName>
|
||
, A–C (CPAG.RAN.I.17, ventral (A), dorsal (B) and apical (C) views, stn 4), D–E (CPAG.RAN.I.16, ventral (D) and dorsal (E) views, stn 4), F (CPAG.RAN.I.15, ventral view, stn 4);
|
||
<emphasis id="B97DEAE40E47FFBCFA83F876FA46F805" bold="true" box="[1387,1436,1944,1966]" pageId="14" pageNumber="115">G–I.</emphasis>
|
||
<taxonomicName id="4C094D750E47FFBCFF7FF859FEACF866" box="[151,374,1975,1997]" class="Gastropoda" family="Volutidae" genus="Lyrischapa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="115" phylum="Mollusca" rank="species" species="brevispira" status="sp. nov.">
|
||
<emphasis id="B97DEAE40E47FFBCFF7FF859FEACF866" box="[151,374,1975,1997]" italics="true" pageId="14" pageNumber="115">Lyrischapa brevispira</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24E579F0E47FFBCFE94F859FE17F866" box="[380,461,1975,1997]" pageId="14" pageNumber="115" rank="species">
|
||
<emphasis id="B97DEAE40E47FFBCFE94F859FE17F866" bold="true" box="[380,461,1975,1997]" pageId="14" pageNumber="115">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype, CPAG.RAN.I.18, ventral (G), dorsal (H) and apical (I) views, stn 4. Scale bars: A–H = 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C313657D0E46FFBDFF2FFF79FB2BFD03" pageId="15" pageNumber="116" type="discussion">
|
||
<paragraph id="8BB636F60E46FFBDFF2FFF79FB34FD2F" blockId="15.[151,1436,151,680]" pageId="15" pageNumber="116">
|
||
<emphasis id="B97DEAE40E46FFBDFF2FFF79FE95FF1B" bold="true" box="[199,335,151,176]" pageId="15" pageNumber="116">Comments.</emphasis>
|
||
Although
|
||
<bibRefCitation id="EF984B070E46FFBDFE21FF76FCCBFF1B" author="Cossmann" box="[457,785,151,176]" pageId="15" pageNumber="116" refString="Cossmann, M. & Pissarro, G. (1909) The Mollusca of the Ranikot Series. Part 1, Cephalopoda and Gastropoda. Memoirs of the Geological Survey of India, Palaeontologia Indica, New Series, 3 (1), 1 - 83." type="journal article" year="1909">Cossmann & Pissarro (1909)</bibRefCitation>
|
||
confounded specimens of
|
||
<taxonomicName id="4C094D750E46FFBDFBABFF77FB74FF1B" box="[1091,1198,151,176]" class="Gastropoda" family="Volutidae" genus="Voluta" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="116" phylum="Mollusca" rank="species" species="haimei">
|
||
<emphasis id="B97DEAE40E46FFBDFBABFF77FB74FF1B" box="[1091,1198,151,176]" italics="true" pageId="15" pageNumber="116">L. haimei</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="4C094D750E46FFBDFB18FF77FAACFF1B" box="[1264,1398,151,176]" class="Gastropoda" family="Volutidae" genus="Lyrischapa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="116" phylum="Mollusca" rank="species" species="blanfordi">
|
||
<emphasis id="B97DEAE40E46FFBDFB18FF77FAACFF1B" box="[1264,1398,151,176]" italics="true" pageId="15" pageNumber="116">L. blanfordi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C094D750E46FFBDFA6DFF77FF24FF7E" class="Gastropoda" family="Volutidae" genus="Lyrischapa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="116" phylum="Mollusca" rank="species" species="blanfordi">
|
||
<emphasis id="B97DEAE40E46FFBDFA6DFF77FF24FF7E" italics="true" pageId="15" pageNumber="116">L. blanfordi</emphasis>
|
||
</taxonomicName>
|
||
was originally described in the genus
|
||
<taxonomicName id="4C094D750E46FFBDFD46FF52FD25FF7E" box="[686,767,188,213]" class="Gastropoda" family="Volutidae" genus="Athleta" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="116" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97DEAE40E46FFBDFD46FF52FD25FF7E" box="[686,767,188,213]" italics="true" pageId="15" pageNumber="116">Athleta</emphasis>
|
||
</taxonomicName>
|
||
and later
|
||
<bibRefCitation id="EF984B070E46FFBDFC98FF52FBCBFF7F" author="Givens" box="[880,1041,188,213]" pageId="15" pageNumber="116" refString="Givens, C. R. (1991) Old world Tethyan occurrences of Lyrischapa (Gastropoda: Volutidae) and biogeographic implications. Journal of Paleontology, 65, 661 - 670." type="journal article" year="1991">Givens (1991)</bibRefCitation>
|
||
did not discuss it in his revision of the genus
|
||
<taxonomicName id="4C094D750E46FFBDFEF9FF31FE54FF53" box="[273,398,223,248]" class="Gastropoda" family="Volutidae" genus="Lyrischapa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="116" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97DEAE40E46FFBDFEF9FF31FE54FF53" box="[273,398,223,248]" italics="true" pageId="15" pageNumber="116">Lyrischapa</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="EF984B070E46FFBDFE49FF31FDA0FF53" author="Vredenburg" box="[417,634,223,248]" pageId="15" pageNumber="116" refString="Vredenburg, E. W. (1923) Indian Tertiary Gastropoda, IV. Olividae, Harpidae, Marginellidae, Volutidae and Mitridae, with comparative diagnosis of new species. Records of the Geological Survey of India, 54, 243 - 276." type="journal article" year="1923">Vredenburg (1923)</bibRefCitation>
|
||
compared
|
||
<taxonomicName id="4C094D750E46FFBDFCE8FF0FFC53FF53" box="[768,905,223,248]" class="Gastropoda" family="Volutidae" genus="Lyrischapa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="116" phylum="Mollusca" rank="species" species="blanfordi">
|
||
<emphasis id="B97DEAE40E46FFBDFCE8FF0FFC53FF53" box="[768,905,223,248]" italics="true" pageId="15" pageNumber="116">L. blanfordi</emphasis>
|
||
</taxonomicName>
|
||
with two Bartonian species from the Anglo- Parisian Basin:
|
||
<taxonomicName id="4C094D750E46FFBDFEAEFEEAFDC7FEB6" box="[326,541,260,285]" class="Gastropoda" family="Volutidae" genus="Athleta" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="116" phylum="Mollusca" rank="species" species="athleta" subGenus="Athleta">
|
||
<emphasis id="B97DEAE40E46FFBDFEAEFEEAFDC7FEB6" box="[326,541,260,285]" italics="true" pageId="15" pageNumber="116">Athleta (A.) athleta</emphasis>
|
||
</taxonomicName>
|
||
(Solander
|
||
<emphasis id="B97DEAE40E46FFBDFD70FEEBFD74FEB7" box="[664,686,261,284]" italics="true" pageId="15" pageNumber="116">in</emphasis>
|
||
Brander, 1766) and
|
||
<taxonomicName id="4C094D750E46FFBDFC7CFEEAFA94FEB6" authority="Deshayes, 1835" authorityName="Deshayes" authorityYear="1835" box="[916,1358,260,285]" class="Gastropoda" family="Volutidae" genus="Athleta" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="116" phylum="Mollusca" rank="species" species="strombiformis" subGenus="Athleta">
|
||
<emphasis id="B97DEAE40E46FFBDFC7CFEEAFB5AFEB6" box="[916,1152,260,285]" italics="true" pageId="15" pageNumber="116">A. (A.) strombiformis</emphasis>
|
||
(Deshayes, 1835)
|
||
</taxonomicName>
|
||
. These species exhibit a biconic shape and have shoulder spines, so generating a superficial resemblance with
|
||
<taxonomicName id="4C094D750E46FFBDFAF9FEC7FA4FFEEB" box="[1297,1429,295,320]" class="Gastropoda" family="Volutidae" genus="Lyrischapa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="116" phylum="Mollusca" rank="species" species="blanfordi">
|
||
<emphasis id="B97DEAE40E46FFBDFAF9FEC7FA4FFEEB" box="[1297,1429,295,320]" italics="true" pageId="15" pageNumber="116">L. blanfordi</emphasis>
|
||
</taxonomicName>
|
||
, but they differ by their early teleoconch whorls and by their columellar folds. In
|
||
<taxonomicName id="4C094D750E46FFBDFB8CFEA2FAD6FECE" box="[1124,1292,332,357]" class="Gastropoda" family="Volutidae" genus="Athleta" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="116" phylum="Mollusca" rank="species" species="athleta" subGenus="Athleta">
|
||
<emphasis id="B97DEAE40E46FFBDFB8CFEA2FAD6FECE" box="[1124,1292,332,357]" italics="true" pageId="15" pageNumber="116">A. (A.) athleta</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C094D750E46FFBDFAA5FEA2FEE9FE23" class="Gastropoda" family="Volutidae" genus="Athleta" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="116" phylum="Mollusca" rank="species" species="strombiformis" subGenus="Athleta">
|
||
<emphasis id="B97DEAE40E46FFBDFAA5FEA2FEE9FE23" italics="true" pageId="15" pageNumber="116">A. (A.) strombiformis</emphasis>
|
||
</taxonomicName>
|
||
, the early teleoconch whorls have two main spiral rows of nodules (subsutural and shoulder rows) and one or two abapical rows. This kind of sculpture is typical of Paleogene
|
||
<taxonomicName id="4C094D750E46FFBDFBF2FE7AFB02FE06" box="[1050,1240,404,429]" class="Gastropoda" family="Volutidae" genus="Athleta" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="116" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97DEAE40E46FFBDFBF2FE7AFB02FE06" box="[1050,1240,404,429]" italics="true" pageId="15" pageNumber="116">Athleta (Athleta)</emphasis>
|
||
</taxonomicName>
|
||
species and it is present in other subgenera such as
|
||
<emphasis id="B97DEAE40E46FFBDFDF4FE56FDEFFE64" box="[540,565,440,463]" italics="true" pageId="15" pageNumber="116">A.</emphasis>
|
||
(
|
||
<emphasis id="B97DEAE40E46FFBDFDACFE59FD08FE7B" box="[580,722,439,464]" italics="true" pageId="15" pageNumber="116">
|
||
<taxonomicName id="4C094D750E46FFBDFDACFE59FD10FE7B" box="[580,714,439,464]" class="Gastropoda" family="Volutidae" genus="Volutospina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="116" phylum="Mollusca" rank="genus">Volutospina</taxonomicName>
|
||
)
|
||
</emphasis>
|
||
,
|
||
<emphasis id="B97DEAE40E46FFBDFD08FE56FC4BFE7B" box="[736,913,439,464]" italics="true" pageId="15" pageNumber="116">A. (Volutopupa)</emphasis>
|
||
an
|
||
<emphasis id="B97DEAE40E46FFBDFC5BFE59FB5CFE7B" box="[947,1158,439,464]" italics="true" pageId="15" pageNumber="116">
|
||
d A. (
|
||
<taxonomicName id="4C094D750E46FFBDFC07FE59FBA4FE7B" box="[1007,1150,439,464]" class="Gastropoda" family="Volutidae" genus="Volutocorbis" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="116" phylum="Mollusca" rank="genus">Volutocorbis</taxonomicName>
|
||
)
|
||
</emphasis>
|
||
. However, it is absent in
|
||
<taxonomicName id="4C094D750E46FFBDFF7FFE32FECEFE5E" box="[151,276,476,501]" class="Gastropoda" family="Volutidae" genus="Lyrischapa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="116" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97DEAE40E46FFBDFF7FFE32FECEFE5E" box="[151,276,476,501]" italics="true" pageId="15" pageNumber="116">Lyrischapa</emphasis>
|
||
</taxonomicName>
|
||
, which has only a single shoulder row during the ontogenetic development of the shell.
|
||
<taxonomicName id="4C094D750E46FFBDFB13FE32FA46FE5E" box="[1275,1436,476,501]" class="Gastropoda" family="Volutidae" genus="Athleta" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="116" phylum="Mollusca" rank="species" species="athleta" subGenus="Athleta">
|
||
<emphasis id="B97DEAE40E46FFBDFB13FE32FA46FE5E" box="[1275,1436,476,501]" italics="true" pageId="15" pageNumber="116">A. (A.) athleta</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C094D750E46FFBDFF2EFDEEFE68FDB3" box="[198,434,511,536]" class="Gastropoda" family="Volutidae" genus="Athleta" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="116" phylum="Mollusca" rank="species" species="strombiformis" subGenus="Athleta">
|
||
<emphasis id="B97DEAE40E46FFBDFF2EFDEEFE68FDB3" box="[198,434,511,536]" italics="true" pageId="15" pageNumber="116">A. (A.) strombiformis</emphasis>
|
||
</taxonomicName>
|
||
also bear three strong columellar folds and several weaker folds adapically, whereas in
|
||
<taxonomicName id="4C094D750E46FFBDFA6DFDEFFF24FD96" class="Gastropoda" family="Volutidae" genus="Lyrischapa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="116" phylum="Mollusca" rank="species" species="blanfordi">
|
||
<emphasis id="B97DEAE40E46FFBDFA6DFDEFFF24FD96" italics="true" pageId="15" pageNumber="116">L. blanfordi</emphasis>
|
||
</taxonomicName>
|
||
the number of folds varies from 6 to 8. In
|
||
<taxonomicName id="4C094D750E46FFBDFD30FDCBFC86FD96" box="[728,860,548,573]" class="Gastropoda" family="Volutidae" genus="Lyrischapa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="116" phylum="Mollusca" rank="species" species="blanfordi">
|
||
<emphasis id="B97DEAE40E46FFBDFD30FDCBFC86FD96" box="[728,860,548,573]" italics="true" pageId="15" pageNumber="116">L. blanfordi</emphasis>
|
||
</taxonomicName>
|
||
, as in other members of
|
||
<taxonomicName id="4C094D750E46FFBDFB98FDCAFB37FD96" box="[1136,1261,548,573]" class="Gastropoda" family="Volutidae" genus="Lyrischapa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="116" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97DEAE40E46FFBDFB98FDCAFB37FD96" box="[1136,1261,548,573]" italics="true" pageId="15" pageNumber="116">Lyrischapa</emphasis>
|
||
</taxonomicName>
|
||
, these folds are stronger in most specimens than those observed in the two Bartonian
|
||
<taxonomicName id="4C094D750E46FFBDFC44FDA9FC27FDCB" box="[940,1021,583,608]" class="Gastropoda" family="Volutidae" genus="Athleta" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="116" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97DEAE40E46FFBDFC44FDA9FC27FDCB" box="[940,1021,583,608]" italics="true" pageId="15" pageNumber="116">Athleta</emphasis>
|
||
</taxonomicName>
|
||
species and other Paleogene
|
||
<taxonomicName id="4C094D750E46FFBDFAA3FDA9FA46FDCB" box="[1355,1436,583,608]" class="Gastropoda" family="Volutidae" genus="Athleta" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="116" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97DEAE40E46FFBDFAA3FDA9FA46FDCB" box="[1355,1436,583,608]" italics="true" pageId="15" pageNumber="116">Athleta</emphasis>
|
||
</taxonomicName>
|
||
species. Therefore, it is with little uncertainty that we transfer
|
||
<taxonomicName id="4C094D750E46FFBDFCA7FD83FC0EFD2E" box="[847,980,620,645]" class="Gastropoda" family="Volutidae" genus="Lyrischapa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="116" phylum="Mollusca" rank="species" species="blanfordi">
|
||
<emphasis id="B97DEAE40E46FFBDFCA7FD83FC0EFD2E" box="[847,980,620,645]" italics="true" pageId="15" pageNumber="116">L. blanfordi</emphasis>
|
||
</taxonomicName>
|
||
to the genus
|
||
<taxonomicName id="4C094D750E46FFBDFB82FD82FB3DFD2E" box="[1130,1255,620,645]" class="Gastropoda" family="Volutidae" genus="Lyrischapa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="116" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97DEAE40E46FFBDFB82FD82FB3DFD2E" box="[1130,1255,620,645]" italics="true" pageId="15" pageNumber="116">Lyrischapa</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB636F60E46FFBDFF2FFD61FB2BFD03" blockId="15.[151,1436,151,680]" box="[199,1265,655,680]" pageId="15" pageNumber="116">
|
||
<emphasis id="B97DEAE40E46FFBDFF2FFD61FE62FD03" bold="true" box="[199,440,655,680]" pageId="15" pageNumber="116">Stratigraphic range.</emphasis>
|
||
Lakhra Formation: Jhirak (
|
||
<bibRefCitation id="EF984B070E46FFBDFD04FD61FC69FD03" author="Vredenburg" box="[748,947,655,680]" pageId="15" pageNumber="116" refString="Vredenburg, E. W. (1923) Indian Tertiary Gastropoda, IV. Olividae, Harpidae, Marginellidae, Volutidae and Mitridae, with comparative diagnosis of new species. Records of the Geological Survey of India, 54, 243 - 276." type="journal article" year="1923">Vredenburg 1923</bibRefCitation>
|
||
), Lakhra Dome (this paper).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |