1001 lines
125 KiB
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1001 lines
125 KiB
XML
<document id="67B85388B7A3E3424ECBA6A8DEFD6A17" ID-DOI="10.5281/zenodo.11376950" ID-ISSN="1525-9153" ID-Zenodo-Dep="11376950" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1716906473563" checkinUser="felipe" docAuthor="Venegas, Pablo J., Echevarría, Lourdes Y., Lobos, Simón E., Sales Nunes, Pedro M., Abstract. - Based, Omar Torres-Carvajal & The, Peru." docDate="2016" docId="0385EC6CF366FE1FFC8BFE26FC2BE9E6" docLanguage="en" docName="AmphibReptileCons.10.1.e121.pdf" docOrigin="Amphibian & Reptile Conservation (e 121) 10 (1)" docStyle="DocumentStyle:D97478CDCB68366E5D4AECD282BFFB6F.2:AmphibReptileConse.2015-.journal_article.0cover" docStyleId="D97478CDCB68366E5D4AECD282BFFB6F" docStyleName="AmphibReptileConse.2015-.journal_article.0cover" docStyleVersion="2" docTitle="Pholidobolus ulisesi Venegas & Echevarría & Lobos & Sales Nunes & Abstract.-Based & The 2016, sp. nov." docType="treatment" docUuid="283DAECE-3FD5-496D-963B-A4E8E4DC8CA7" docUuidSource="ZooBank" docVersion="3" lastPageNumber="27" masterDocId="FFBC9414F367FE19FF9CFFF7FFF9ED07" masterDocTitle="A new species of Andean microteiid lizard (Gymnophthalmidae: Cercosaurinae: Pholidobolus) from Peru, with comments on P. vertebralis" masterLastPageNumber="33" masterPageNumber="21" pageNumber="22" updateTime="1716946738442" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="32A5109882FADEB4262BFDFBDCACDA75">A new species of Andean microteiid lizard (Gymnophthalmidae: Cercosaurinae: Pholidobolus) from Peru, with comments on P. vertebralis</mods:title>
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<mods:namePart id="AB227290B1772956DC2923D84B95220D">Venegas, Pablo J.</mods:namePart>
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<mods:affiliation id="7BB13EDC6BE7B9C196503F17EAEB9E6C">División de Herpetología-Centro de Ornitología y Biodiversidad (CORBIDI), Santa Rita N ˚ 105 36 Of. 202, Urb. Huertos de San Antonio, Surco, Lima, PERÚ</mods:affiliation>
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<mods:namePart id="B3DFBF539206471D8FE589A424A89FC2">Echevarría, Lourdes Y.</mods:namePart>
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<mods:affiliation id="DB86FD2EBDEDBD6F649BF0CE5E859093">Laboratório de Sistemática de Vertebrados, Pontifícia Universidade Católica do Rio Grande do Sul, Porto Alegre, RS, BRAZIL & División de Herpetología-Centro de Ornitología y Biodiversidad (CORBIDI), Santa Rita N ˚ 105 36 Of. 202, Urb. Huertos de San Antonio, Surco, Lima, PERÚ</mods:affiliation>
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<mods:namePart id="8CDD3C59BD9652138E3AD31F13048BD3">Lobos, Simón E.</mods:namePart>
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<mods:affiliation id="1C28444F4E4836F11C13F34070CD00B9">Museo de Zoología, Escuela de Biología, Pontificia Universidad Católica del Ecuador, Avenida 12 de Octubre 1076 y Roca, Apartado 17 - 01 - 2184, Quito, ECUADOR</mods:affiliation>
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<mods:namePart id="552DD44BCE7D37C2B2D1B39FAF24C116">Sales Nunes, Pedro M.</mods:namePart>
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<mods:affiliation id="540A266B94DD78191B1549D114834DDE">Universidade Federal de Pernambuco, Centro de Biociências, Departamento de Zoologia, Av. Professor Moraes Rego, s / n. Cidade Universitária CEP 50670 - 901, Recife, PE, BRAZIL</mods:affiliation>
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<mods:namePart id="AD784B2E5A158951C7EEB383BD36C929">Abstract. - Based, Omar Torres-Carvajal</mods:namePart>
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<mods:namePart id="D32D8776E4122297C26680AC41CBF2F9">The, Peru.</mods:namePart>
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<mods:title id="8C4583683ACC29821880B1F6E7C6BE18">Amphibian & Reptile Conservation</mods:title>
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<mods:date id="47703F121110B3B28B71E5B3E2D33DDB">2016</mods:date>
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<mods:title id="85D4DD26946CC2A062E98AADCD7A49A6">e 121</mods:title>
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<mods:number id="9B1A35B6D34AA6CE772FC49C103CFD03">2016-07-30</mods:number>
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<mods:number id="8E1798E39D232AF8C9D6484FF79D5359">10</mods:number>
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<mods:identifier id="D3857137F2A8463F29E3A0C792F99098" type="DOI">10.5281/zenodo.11376950</mods:identifier>
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<taxonomicName id="4C2C26F9F366FE18FC8BFE26FBE3ECED" ID-CoL="8GQ6S" authority="Venegas & Echevarría & Lobos & Sales Nunes & Abstract.-Based & The, 2016" authorityName="Venegas & Echevarría & Lobos & Sales Nunes & Abstract.-Based & The" authorityYear="2016" box="[791,1050,465,490]" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="1" pageNumber="22" phylum="Chordata" rank="species" species="ulisesi" status="sp. nov.">
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<emphasis id="B9588168F366FE18FC8BFE26FBE3ECED" bold="true" box="[791,1050,465,490]" italics="true" pageId="1" pageNumber="22">Pholidobolus ulisesi</emphasis>
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.
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<taxonomicName id="4C2C26F9F366FE18FC8BFD7FFBEFEF98" authorityName="O'Shaughnessy" authorityYear="1879" box="[791,1046,647,671]" class="Squamata" family="Gymnophthalmidae" genus="Cercosaura" kingdom="Animalia" pageId="1" pageNumber="22" phylum="Chordata" rank="species" species="vertebralis">
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<emphasis id="B9588168F366FE18FC8BFD7FFBEFEF98" box="[791,1046,647,671]" italics="true" pageId="1" pageNumber="22">Cercosaura vertebralis</emphasis>
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—
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<bibRefCitation id="EFBD208BF366FE18FBADFD7FFAD8EF98" author="Doan TM & Cusi JC" box="[1073,1313,647,671]" pageId="1" pageNumber="22" refId="ref7015" refString="Doan TM, Cusi JC. 2014. Geographic distribution of Cercosaura vertebralis O'Shaughnessy, 1879 (Reptilia: Squamata: Gymnophthalmidae) and the status of Cercosaura ampuedai (Lancini, 1968). Check List 10: 1,195 - 1,200." type="journal volume" year="2014">Doan and Cusi 2014</bibRefCitation>
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(part):
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.
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<emphasis id="B9588168F366FE18FC8BFD11FC5CEFF9" box="[791,933,742,766]" italics="true" pageId="1" pageNumber="22">Pholidobolus</emphasis>
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sp.
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—
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<treatmentCitation id="0A8D7B6BF366FE18FC7CFD11FAC2EFF9" author="Torres-Carvajal O & Lobos SE & Venegas PJ" box="[992,1339,742,766]" page="286" pageId="1" pageNumber="22" year="2015">
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<bibRefCitation id="EFBD208BF366FE18FC7CFD11FAC2EFF9" author="Torres-Carvajal O & Lobos SE & Venegas PJ" box="[992,1339,742,766]" pageId="1" pageNumber="22" pagination="281 - 288" refId="ref7796" refString="Torres-Carvajal O, Lobos SE, Venegas PJ. 2015. Phylogeny of Neotropical Cercosaura (Squamata: Gymnophthalmidae) lizards. Molecular Phylogenetics and Evolution 93: 281 - 288." type="journal article" year="2015">Torres-Carvajal et al. 2015: 286</bibRefCitation>
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.
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<taxonomicName id="4C2C26F9F366FE18FC8BFCD2FC31EE3A" box="[791,968,805,829]" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="1" pageNumber="22" phylum="Chordata" rank="species" species="undetermined">
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<emphasis id="B9588168F366FE18FC8BFCD2FC5CEE3A" box="[791,933,805,829]" italics="true" pageId="1" pageNumber="22">Pholidobolus</emphasis>
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sp.
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—
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<treatmentCitation id="0A8D7B6BF366FE18FC7EFCD2FACEEE3A" author="Torres-Carvajal O & Lobos SE & Venegas PJ & Chavez G & Aguirre-Penafiel V & Zurita D & Echevarria LY" box="[994,1335,805,829]" page="70" pageId="1" pageNumber="22" year="2016">
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<bibRefCitation id="EFBD208BF366FE18FC7EFCD2FACEEE3A" author="Torres-Carvajal O & Lobos SE & Venegas PJ & Chavez G & Aguirre-Penafiel V & Zurita D & Echevarria LY" box="[994,1335,805,829]" pageId="1" pageNumber="22" pagination="63 - 75" refId="ref7828" refString="Torres-Carvajal O, Lobos SE, Venegas PJ, Chavez G, Aguirre-Penafiel V, Zurita D, Echevarria LY. 2016. Phylogeny and biogeography of the most diverse clade of South American gymnophthalmid lizards (Squamata, Gymnophthalmidae, Cercosaurinae). Molecular Phylogenetics and Evolution 99: 63 - 75." type="journal article" year="2016">Torres-Carvajal et al. 2016: 70</bibRefCitation>
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(
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).
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|
||
<materialsCitation id="3B445727F366FE18FC8BFC73FC3BE919" collectingDate="2013-03-07" collectionCode="CORBIDI" collectorName="de Huamantanga & P. J. Venegas." country="Peru" county="Jaen province" elevation="2" latitude="-5.663358" location="Bosque de Huamantanga" longLatPrecision="1" longitude="-78.943275" municipality="Huabal district" pageId="1" pageNumber="22" specimenCode="CORBIDI 12734" specimenCount="1" stateProvince="Cajamarca" typeStatus="holotype">
|
||
<emphasis id="B9588168F366FE18FC8BFC73FC68EE9A" bold="true" box="[791,913,900,925]" pageId="1" pageNumber="22">
|
||
<typeStatus id="5497E3D8F366FE18FC8BFC73FC77EE9A" box="[791,910,900,925]" pageId="1" pageNumber="22" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCode id="DB8AF501F366FE18FC3BFC72FB95EE9A" box="[935,1132,901,925]" collectionCode="CORBIDI" country="Peru" httpUri="http://grbio.org/cool/g3tc-zhv7" name="Centro de Ornitologia y Biodiversidad" pageId="1" pageNumber="22">CORBIDI 12734</specimenCode>
|
||
, an adult male from
|
||
<location id="8EF30BA1F366FE18FC8BFC51FBD0EEB9" LSID="urn:lsid:plazi:treatment:0385EC6CF366FE1FFC8BFE26FC2BE9E6:8EF30BA1F366FE18FC8BFC51FBD0EEB9" box="[791,1065,933,958]" country="Peru" county="Jaen province" latitude="-5.663358" longLatPrecision="1" longitude="-78.943275" municipality="Huabal district" name="Bosque de Huamantanga" pageId="1" pageNumber="22" stateProvince="Cajamarca">
|
||
Bosque
|
||
<collectorName id="26D938ACF366FE18FCEDFC52FBD0EEB9" box="[881,1065,933,958]" pageId="1" pageNumber="22">de Huamantanga</collectorName>
|
||
</location>
|
||
(
|
||
<geoCoordinate id="EE183BBDF366FE18FBA0FC52FB24EEBA" box="[1084,1245,933,957]" degrees="5" direction="south" minutes="39" orientation="latitude" pageId="1" pageNumber="22" precision="1" seconds="48.09" value="-5.663358">5°39’48.09’’ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE183BBDF366FE18FB71FC52FCCDEEDA" degrees="78" direction="west" minutes="56" orientation="longitude" pageId="1" pageNumber="22" precision="1" seconds="35.8" value="-78.943275">78°56’35.8’’ W</geoCoordinate>
|
||
), at
|
||
<quantity id="4CD4F09FF366FE18FCFCFC32FC4EEEDA" box="[864,951,965,989]" metricMagnitude="0" metricUnit="m" metricValue="2.211" pageId="1" pageNumber="22" unit="m" value="2.211">
|
||
<elevation id="0001BA49F366FE18FCFCFC32FC4EEEDA" box="[864,951,965,989]" metricMagnitude="0" metricUnit="m" metricValue="2.211" pageId="1" pageNumber="22" unit="m" value="2.211">2,211 m</elevation>
|
||
</quantity>
|
||
elevation,
|
||
<collectingMunicipality id="6BF7C700F366FE18FBB1FC32FB37EEDA" box="[1069,1230,965,989]" pageId="1" pageNumber="22">Huabal district</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="62F225F6F366FE18FB44FC32FA96EEDA" box="[1240,1391,965,989]" pageId="1" pageNumber="22">Jaén province</collectingCounty>
|
||
,
|
||
<collectingRegion id="49E89398F366FE18FC8BFC12FC70EEFA" box="[791,905,997,1021]" country="Peru" name="Cajamarca" pageId="1" pageNumber="22">Cajamarca</collectingRegion>
|
||
department,
|
||
<collectingCountry id="F33B1DEAF366FE18FB89FC11FBB0EEFA" box="[1045,1097,998,1021]" name="Peru" pageId="1" pageNumber="22">Peru</collectingCountry>
|
||
, collected on
|
||
<date id="FF927BBAF366FE18FB41FC11FA8AEEFA" box="[1245,1395,997,1021]" pageId="1" pageNumber="22" value="2013-03-07">
|
||
<collectingDate id="EFD68252F366FE18FB41FC11FA8AEEFA" box="[1245,1395,997,1021]" pageId="1" pageNumber="22" value="2013-03-07">7 March 2013</collectingDate>
|
||
</date>
|
||
by
|
||
<collectorName id="26D938ACF366FE18FCA4FBF1FC3BE919" box="[824,962,1030,1054]" pageId="1" pageNumber="22">P.J. Venegas.</collectorName>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8B935D7AF366FE18FC8BFBB3FAFFE8BA" blockId="1.[789,1395,1092,1469]" pageId="1" pageNumber="22">
|
||
<materialsCitation id="3B445727F366FE18FC8BFBB3FAD1E95A" box="[791,1320,1092,1117]" collectionCode="CORBIDI" pageId="1" pageNumber="22" specimenCount="46" specimenCount-juvenile="46" typeStatus="paratype">
|
||
<emphasis id="B9588168F366FE18FC8BFBB3FC2FE95A" bold="true" box="[791,982,1092,1117]" pageId="1" pageNumber="22">
|
||
<typeStatus id="5497E3D8F366FE18FC8BFBB3FC6FE95A" box="[791,918,1092,1117]" pageId="1" pageNumber="22" type="paratype">Paratypes</typeStatus>
|
||
(17):
|
||
</emphasis>
|
||
<collectionCode id="ED3DC5BFF366FE18FC42FBB2FBB5E95A" box="[990,1100,1093,1117]" country="Peru" httpUri="http://grbio.org/cool/g3tc-zhv7" name="Centro de Ornitologia y Biodiversidad" pageId="1" pageNumber="22">CORBIDI</collectionCode>
|
||
12740–
|
||
<specimenCount id="9D2A96F3F366FE18FB3FFBB2FAD1E95A" box="[1187,1320,1093,1117]" count="46" pageId="1" pageNumber="22" type="juvenile">46 juveniles</specimenCount>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B445727F366FE18FAAFFBB2FB75E97A" collectionCode="CORBIDI" pageId="1" pageNumber="22" specimenCode="CORBIDI 12735-36, 12739" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="DB8AF501F366FE18FAAFFBB2FC39E97A" collectionCode="CORBIDI" country="Peru" httpUri="http://grbio.org/cool/g3tc-zhv7" name="Centro de Ornitologia y Biodiversidad" pageId="1" pageNumber="22">CORBIDI 12735–36</specimenCode>
|
||
,
|
||
<specimenCode id="DB8AF501F366FE18FC55FB92FBF5E97A" box="[969,1036,1125,1149]" collectionCode="CORBIDI" country="Peru" httpUri="http://grbio.org/cool/g3tc-zhv7" name="Centro de Ornitologia y Biodiversidad" pageId="1" pageNumber="22">12739</specimenCode>
|
||
adult males
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B445727F366FE18FB09FB92FB00E99A" collectionCode="CORBIDI" pageId="1" pageNumber="22" specimenCount="38" specimenCount-adult="38" typeStatus="paratype">
|
||
<collectionCode id="ED3DC5BFF366FE18FB09FB92FAFAE97A" box="[1173,1283,1125,1149]" country="Peru" httpUri="http://grbio.org/cool/g3tc-zhv7" name="Centro de Ornitologia y Biodiversidad" pageId="1" pageNumber="22">CORBIDI</collectionCode>
|
||
12737–
|
||
<specimenCount id="9D2A96F3F366FE18FAC5FB92FCB3E99A" count="38" pageId="1" pageNumber="22" type="adult">38 adult</specimenCount>
|
||
females, all collected with the
|
||
<typeStatus id="5497E3D8F366FE18FB04FB72FB00E99A" box="[1176,1273,1157,1181]" pageId="1" pageNumber="22" type="holotype">holotype</typeStatus>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B445727F366FE18FA9FFB72FBBDE83A" collectingDate="2008-08-24" collectionCode="CORBIDI" collectorName="El Chaupe & M. Dobiey" country="Peru" elevation="2" latitude="-5.2356" location="Namballe district" longLatPrecision="1" longitude="-79.09905" municipality="San Ignacio province" pageId="1" pageNumber="22" specimenCode="CORBIDI 00871-73" specimenCount="1" stateProvince="Cajamarca" typeStatus="paratype">
|
||
<specimenCode id="DB8AF501F366FE18FA9FFB72FC7DE9BA" collectionCode="CORBIDI" country="Peru" httpUri="http://grbio.org/cool/g3tc-zhv7" name="Centro de Ornitologia y Biodiversidad" pageId="1" pageNumber="22">CORBIDI 00871–73</specimenCode>
|
||
, an adult female, an adult male and a juvenile, respectively, from
|
||
<collectorName id="26D938ACF366FE18FC47FB32FBB3E9DA" box="[987,1098,1221,1245]" pageId="1" pageNumber="22">El Chaupe</collectorName>
|
||
(
|
||
<geoCoordinate id="EE183BBDF366FE18FBC4FB32FB1DE9DA" box="[1112,1252,1221,1245]" degrees="5" direction="south" minutes="14" orientation="latitude" pageId="1" pageNumber="22" precision="1" seconds="8.16" value="-5.2356">5°14´8.16” S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE183BBDF366FE18FB73FB32FCCAE9FA" degrees="79" direction="west" minutes="5" orientation="longitude" pageId="1" pageNumber="22" precision="1" seconds="56.58" value="-79.09905">79°5´56.58” W</geoCoordinate>
|
||
), at
|
||
<quantity id="4CD4F09FF366FE18FCC3FB12FC4EE9FA" box="[863,951,1253,1277]" metricMagnitude="0" metricUnit="m" metricValue="2.016" pageId="1" pageNumber="22" unit="m" value="2.016">
|
||
<elevation id="0001BA49F366FE18FCC3FB12FC4EE9FA" box="[863,951,1253,1277]" metricMagnitude="0" metricUnit="m" metricValue="2.016" pageId="1" pageNumber="22" unit="m" value="2.016">2,016 m</elevation>
|
||
</quantity>
|
||
elevation,
|
||
<location id="8EF30BA1F366FE18FBB1FB12FB11E9FA" LSID="urn:lsid:plazi:treatment:0385EC6CF366FE1FFC8BFE26FC2BE9E6:8EF30BA1F366FE18FBB1FB12FB11E9FA" box="[1069,1256,1253,1277]" country="Peru" latitude="-5.2356" longLatPrecision="1" longitude="-79.09905" municipality="San Ignacio province" name="Namballe district" pageId="1" pageNumber="22" stateProvince="Cajamarca">Namballe district</location>
|
||
,
|
||
<collectingMunicipality id="6BF7C700F366FE18FB6FFB12FC8EE81A" pageId="1" pageNumber="22">San Ignacio province</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="49E89398F366FE18FC18FAF2FC0FE81A" box="[900,1014,1285,1309]" country="Peru" name="Cajamarca" pageId="1" pageNumber="22">Cajamarca</collectingRegion>
|
||
department,
|
||
<collectingCountry id="F33B1DEAF366FE18FB1BFAF1FB42E81A" box="[1159,1211,1286,1309]" name="Peru" pageId="1" pageNumber="22">Peru</collectingCountry>
|
||
, collected by
|
||
<collectorName id="26D938ACF366FE18FAC8FAF1FC9FE83A" pageId="1" pageNumber="22">M. Dobiey</collectorName>
|
||
on
|
||
<date id="FF927BBAF366FE18FC0EFAD2FBBDE83A" box="[914,1092,1317,1341]" pageId="1" pageNumber="22" value="2008-08-24">
|
||
<collectingDate id="EFD68252F366FE18FC0EFAD2FBBDE83A" box="[914,1092,1317,1341]" pageId="1" pageNumber="22" value="2008-08-24">24 August 2008</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B445727F366FE18FBCDFAD2FC6DE85A" collectionCode="CORBIDI" country="Peru" pageId="1" pageNumber="22" specimenCode="CORBIDI 14889" specimenCount="1" stateProvince="Cajamarca" typeStatus="paratype">
|
||
<specimenCode id="DB8AF501F366FE18FBCDFAD2FAF4E83A" box="[1105,1293,1317,1341]" collectionCode="CORBIDI" country="Peru" httpUri="http://grbio.org/cool/g3tc-zhv7" name="Centro de Ornitologia y Biodiversidad" pageId="1" pageNumber="22">CORBIDI 14889</specimenCode>
|
||
, an adult female, and
|
||
</materialsCitation>
|
||
<materialsCitation id="3B445727F366FE18FC07FAB2FAFBE8BA" collectingDate="2014-09-26" collectionCode="CORBIDI" collectorName="K. Garcia" country="Peru" elevation="3" latitude="-5.753015" location="San Felipe de Jaen" longLatPrecision="1" longitude="-79.23885" municipality="Jaen province" pageId="1" pageNumber="22" specimenCode="CORBIDI 14896" specimenCount="1" stateProvince="Cajamarca" typeStatus="paratype">
|
||
<specimenCode id="DB8AF501F366FE18FC07FAB2FBADE85A" box="[923,1108,1349,1373]" collectionCode="CORBIDI" country="Peru" httpUri="http://grbio.org/cool/g3tc-zhv7" name="Centro de Ornitologia y Biodiversidad" pageId="1" pageNumber="22">CORBIDI 14896</specimenCode>
|
||
, a juvenile, from
|
||
<location id="8EF30BA1F366FE18FA8DFAB2FC7CE87A" LSID="urn:lsid:plazi:treatment:0385EC6CF366FE1FFC8BFE26FC2BE9E6:8EF30BA1F366FE18FA8DFAB2FC7CE87A" country="Peru" latitude="-5.753015" longLatPrecision="1" longitude="-79.23885" municipality="Jaen province" name="San Felipe de Jaen" pageId="1" pageNumber="22" stateProvince="Cajamarca">San Felipe de Jaén</location>
|
||
(
|
||
<geoCoordinate id="EE183BBDF366FE18FC08FA92FBC4E87A" box="[916,1085,1381,1405]" degrees="5" direction="south" minutes="45" orientation="latitude" pageId="1" pageNumber="22" precision="1" seconds="10.854" value="-5.753015">5°45’10.854” S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE183BBDF366FE18FBD5FA92FAF3E87A" box="[1097,1290,1381,1405]" degrees="79" direction="west" minutes="14" orientation="longitude" pageId="1" pageNumber="22" precision="1" seconds="19.881" value="-79.23885">79°14’19.881” W</geoCoordinate>
|
||
), at
|
||
<quantity id="4CD4F09FF366FE18FAABFA92FCD5E89A" metricMagnitude="0" metricUnit="m" metricValue="2.641" pageId="1" pageNumber="22" unit="m" value="2.641">
|
||
<elevation id="0001BA49F366FE18FAABFA92FCD5E89A" metricMagnitude="0" metricUnit="m" metricValue="2.641" pageId="1" pageNumber="22" unit="m" value="2.641">2,641 m</elevation>
|
||
</quantity>
|
||
elevation,
|
||
<collectingMunicipality id="6BF7C700F366FE18FC3EFA72FBC1E89A" box="[930,1080,1413,1437]" pageId="1" pageNumber="22">Jaén province</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="49E89398F366FE18FBDFFA72FB4CE89A" box="[1091,1205,1413,1437]" country="Peru" name="Cajamarca" pageId="1" pageNumber="22">Cajamarca</collectingRegion>
|
||
department,
|
||
<collectingCountry id="F33B1DEAF366FE18FADEFA71FA8AE89A" box="[1346,1395,1414,1437]" name="Peru" pageId="1" pageNumber="22">Peru</collectingCountry>
|
||
collected by
|
||
<collectorName id="26D938ACF366FE18FC03FA51FBFEE8BA" box="[927,1031,1445,1469]" pageId="1" pageNumber="22">K. Garcia</collectorName>
|
||
on
|
||
<date id="FF927BBAF366FE18FBB3FA52FAFBE8BA" box="[1071,1282,1445,1469]" pageId="1" pageNumber="22" value="2014-09-26">
|
||
<collectingDate id="EFD68252F366FE18FBB3FA52FAFBE8BA" box="[1071,1282,1445,1469]" pageId="1" pageNumber="22" value="2014-09-26">26 September 2014</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B935D7AF366FE18FC8BFA1EFAD2EB64" blockId="1.[791,1395,1513,1635]" pageId="1" pageNumber="22">
|
||
<emphasis id="B9588168F366FE18FC8BFA1EFB9DEB05" bold="true" box="[791,1124,1513,1538]" pageId="1" pageNumber="22">Photo voucher specimen:</emphasis>
|
||
<materialsCitation id="3B445727F366FE18FBF3FA1CFADEEB64" collectingDate="2007-05-25" collectorName="P. J. Venegas" country="Peru" elevation="2" latitude="-6.0572724" location="Ferrenafe province" longLatPrecision="1" longitude="-79.26677" pageId="1" pageNumber="22" specimenCount="1" stateProvince="Lambayeque">
|
||
Cañaris (
|
||
<geoCoordinate id="EE183BBDF366FE18FB4FFA1DFA97EB05" box="[1235,1390,1514,1538]" degrees="6" direction="south" minutes="03" orientation="latitude" pageId="1" pageNumber="22" precision="1" seconds="26.18" value="-6.0572724">6°03´26.18´´S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE183BBDF366FE18FC8BF9FDFC33EB25" box="[791,970,1546,1570]" degrees="79" direction="west" minutes="16" orientation="longitude" pageId="1" pageNumber="22" precision="1" seconds="00.35" value="-79.26677">79°16´00.35´´W</geoCoordinate>
|
||
), at
|
||
<quantity id="4CD4F09FF366FE18FC65F9FCFBABEB25" box="[1017,1106,1547,1571]" metricMagnitude="0" metricUnit="m" metricValue="2.318" pageId="1" pageNumber="22" unit="m" value="2.318">
|
||
<elevation id="0001BA49F366FE18FC65F9FCFBABEB25" box="[1017,1106,1547,1571]" metricMagnitude="0" metricUnit="m" metricValue="2.318" pageId="1" pageNumber="22" unit="m" value="2.318">2,318 m</elevation>
|
||
</quantity>
|
||
elevation,
|
||
<location id="8EF30BA1F366FE18FB57F9FCFCBFEB44" LSID="urn:lsid:plazi:treatment:0385EC6CF366FE1FFC8BFE26FC2BE9E6:8EF30BA1F366FE18FB57F9FCFCBFEB44" country="Peru" latitude="-6.0572724" longLatPrecision="1" longitude="-79.26677" name="Ferrenafe province" pageId="1" pageNumber="22" stateProvince="Lambayeque">Ferreñafe province</location>
|
||
,
|
||
<collectingRegion id="49E89398F366FE18FCCAF9DCFC19EB44" box="[854,992,1579,1603]" country="Peru" name="Lambayeque" pageId="1" pageNumber="22">Lambayeque</collectingRegion>
|
||
department,
|
||
<collectingCountry id="F33B1DEAF366FE18FBEBF9DCFB52EB45" box="[1143,1195,1579,1602]" name="Peru" pageId="1" pageNumber="22">Peru</collectingCountry>
|
||
, captured and released by
|
||
<collectorName id="26D938ACF366FE18FC1DF9BCFBFDEB64" box="[897,1028,1611,1635]" pageId="1" pageNumber="22">P.J. Venegas</collectorName>
|
||
on
|
||
<date id="FF927BBAF366FE18FBB0F9BDFB43EB64" box="[1068,1210,1610,1635]" pageId="1" pageNumber="22" value="2007-05-25">
|
||
<collectingDate id="EFD68252F366FE18FBB0F9BDFB43EB64" box="[1068,1210,1610,1635]" pageId="1" pageNumber="22" value="2007-05-25">25 May 2007</collectingDate>
|
||
</date>
|
||
(
|
||
<figureCitation id="131741FFF366FE18FB55F9BCFAE6EB64" box="[1225,1311,1611,1635]" captionStart="Fig" captionStartId="4.[141,176,1558,1580]" captionTargetBox="[241,1295,847,1548]" captionTargetId="figure-198@4.[241,1295,847,1550]" captionTargetPageId="4" captionText="Fig. 3. Four individuals of Pholidobolus ulisesi sp. nov. in life. (A) holotype (CORBIDI 12734); (B) adult female (CORBIDI 12737); (C) juvenile (CORBIDI 12744); (D) adult male from Cañaris (photo voucher). Photographs by PJV." figureDoi="http://doi.org/10.5281/zenodo.11376956" httpUri="https://zenodo.org/record/11376956/files/figure.png" pageId="1" pageNumber="22">Fig. 3D</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3360EF1F366FE1BFC8BF97BFD5AEA40" lastPageId="2" lastPageNumber="23" pageId="1" pageNumber="22" type="diagnosis">
|
||
<paragraph id="8B935D7AF366FE1BFC8BF97BFD75E80F" blockId="1.[787,1399,1676,1925]" lastBlockId="2.[138,751,1232,1288]" lastPageId="2" lastPageNumber="23" pageId="1" pageNumber="22">
|
||
<emphasis id="B9588168F366FE18FC8BF97BFC66EBA2" bold="true" box="[791,927,1676,1701]" pageId="1" pageNumber="22">Diagnosis:</emphasis>
|
||
<taxonomicName id="4C2C26F9F366FE18FC3AF97AFB84EBA2" baseAuthorityName="Peters" baseAuthorityYear="1863" box="[934,1149,1677,1701]" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="1" pageNumber="22" phylum="Chordata" rank="species" species="affinis">
|
||
<emphasis id="B9588168F366FE18FC3AF97AFB84EBA2" box="[934,1149,1677,1701]" italics="true" pageId="1" pageNumber="22">Pholidobolus affinis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2C26F9F366FE18FB17F979FB11EBA2" baseAuthorityName="Uzzell" baseAuthorityYear="1973" box="[1163,1256,1677,1701]" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="1" pageNumber="22" phylum="Chordata" rank="species" species="dicrus">
|
||
<emphasis id="B9588168F366FE18FB17F979FB11EBA2" box="[1163,1256,1677,1701]" italics="true" pageId="1" pageNumber="22">P. dicrus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="131741FFF366FE18FB6AF97AFAB4EBA2" box="[1270,1357,1677,1701]" captionStart="Fig" captionStartId="5.[141,176,1564,1586]" captionTargetBox="[141,745,144,1554]" captionTargetId="figure-585@5.[141,745,144,1554]" captionTargetPageId="5" captionText="Fig. 4. Four species of Pholidobolus. (A) adult female of P. dicrus (QCAZ 5304); (B) adult male of P. hillisi (QCAZ 4999); (C) a juvenile of P. vertebralis (QCAZ 5082); (D) adult female of P.sp. from La Granja (CORBIDI 1678). Photographs by: (A) and (B) Santiago R. Ron, (C) OTC, and (D) PJV." figureDoi="http://doi.org/10.5281/zenodo.11376960" httpUri="https://zenodo.org/record/11376960/files/figure.png" pageId="1" pageNumber="22">Fig. 4A</figureCitation>
|
||
),
|
||
<taxonomicName id="4C2C26F9F366FE18FAC3F979FCA9EBC2" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="1" pageNumber="22" phylum="Chordata" rank="species" species="hillisi">
|
||
<emphasis id="B9588168F366FE18FAC3F979FCA9EBC2" italics="true" pageId="1" pageNumber="22">P. hillisi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="131741FFF366FE18FCF8F95AFC43EBC2" box="[868,954,1709,1733]" captionStart="Fig" captionStartId="5.[141,176,1564,1586]" captionTargetBox="[141,745,144,1554]" captionTargetId="figure-585@5.[141,745,144,1554]" captionTargetPageId="5" captionText="Fig. 4. Four species of Pholidobolus. (A) adult female of P. dicrus (QCAZ 5304); (B) adult male of P. hillisi (QCAZ 4999); (C) a juvenile of P. vertebralis (QCAZ 5082); (D) adult female of P.sp. from La Granja (CORBIDI 1678). Photographs by: (A) and (B) Santiago R. Ron, (C) OTC, and (D) PJV." figureDoi="http://doi.org/10.5281/zenodo.11376960" httpUri="https://zenodo.org/record/11376960/files/figure.png" pageId="1" pageNumber="22">Fig. 4B</figureCitation>
|
||
),
|
||
<taxonomicName id="4C2C26F9F366FE18FC4EF959FB91EBC2" authorityName="Montanucci" authorityYear="1973" box="[978,1128,1709,1733]" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="1" pageNumber="22" phylum="Chordata" rank="species" species="prefrontalis">
|
||
<emphasis id="B9588168F366FE18FC4EF959FB91EBC2" box="[978,1128,1709,1733]" italics="true" pageId="1" pageNumber="22">P. prefrontalis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C2C26F9F366FE18FB35F959FAC0EBC2" authorityName="O'Shaughnessy" authorityYear="1879" box="[1193,1337,1709,1733]" class="Squamata" family="Gymnophthalmidae" genus="Cercosaura" kingdom="Animalia" pageId="1" pageNumber="22" phylum="Chordata" rank="species" species="vertebralis">
|
||
<emphasis id="B9588168F366FE18FB35F959FAC0EBC2" box="[1193,1337,1709,1733]" italics="true" pageId="1" pageNumber="22">P. vertebralis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="131741FFF366FE18FADBF95AFCC0EBE2" captionStart="Fig" captionStartId="5.[141,176,1564,1586]" captionTargetBox="[141,745,144,1554]" captionTargetId="figure-585@5.[141,745,144,1554]" captionTargetPageId="5" captionText="Fig. 4. Four species of Pholidobolus. (A) adult female of P. dicrus (QCAZ 5304); (B) adult male of P. hillisi (QCAZ 4999); (C) a juvenile of P. vertebralis (QCAZ 5082); (D) adult female of P.sp. from La Granja (CORBIDI 1678). Photographs by: (A) and (B) Santiago R. Ron, (C) OTC, and (D) PJV." figureDoi="http://doi.org/10.5281/zenodo.11376960" httpUri="https://zenodo.org/record/11376960/files/figure.png" pageId="1" pageNumber="22">Fig. 4C</figureCitation>
|
||
) differ from the new species in having prefrontal scales.
|
||
<taxonomicName id="4C2C26F9F366FE18FCFEF91AFBABEA02" baseAuthorityName="Peters" baseAuthorityYear="1863" box="[866,1106,1773,1797]" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="1" pageNumber="22" phylum="Chordata" rank="species" species="montium">
|
||
<emphasis id="B9588168F366FE18FCFEF91AFBABEA02" box="[866,1106,1773,1797]" italics="true" pageId="1" pageNumber="22">Pholidobolus montium</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2C26F9F366FE18FB1DF919FAF3EA02" authorityName="Montanucci" authorityYear="1973" box="[1153,1290,1773,1797]" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="1" pageNumber="22" phylum="Chordata" rank="species" species="macbrydei">
|
||
<emphasis id="B9588168F366FE18FB1DF919FAF3EA02" box="[1153,1290,1773,1797]" italics="true" pageId="1" pageNumber="22">P. macbrydei</emphasis>
|
||
</taxonomicName>
|
||
have striated and quadrangular dorsal scales (strongly keeled and hexagonal in
|
||
<taxonomicName id="4C2C26F9F366FE18FC3EF8D9FC06EA42" authorityName="Venegas & Echevarría & Lobos & Sales Nunes & Abstract.-Based & The" authorityYear="2016" box="[930,1023,1837,1861]" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="1" pageNumber="22" phylum="Chordata" rank="species" species="ulisesi">
|
||
<emphasis id="B9588168F366FE18FC3EF8D9FC06EA42" box="[930,1023,1837,1861]" italics="true" pageId="1" pageNumber="22">P. ulisesi</emphasis>
|
||
</taxonomicName>
|
||
), and lack the conspicuous narrow, pale brown, vertebral stripe present in
|
||
<taxonomicName id="4C2C26F9F366FE18FB2FF8B9FAE8EA62" authorityName="Venegas & Echevarría & Lobos & Sales Nunes & Abstract.-Based & The" authorityYear="2016" box="[1203,1297,1869,1893]" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="1" pageNumber="22" phylum="Chordata" rank="species" species="ulisesi">
|
||
<emphasis id="B9588168F366FE18FB2FF8B9FAE8EA62" box="[1203,1297,1869,1893]" italics="true" pageId="1" pageNumber="22">P. ulisesi</emphasis>
|
||
</taxonomicName>
|
||
. In addition, the new species has fewer dorsal scales (28–31,
|
||
<emphasis id="B9588168F366FE18FAFBF899FA8AEA82" box="[1383,1395,1902,1925]" italics="true" pageId="1" pageNumber="22">x</emphasis>
|
||
= 29.75) than
|
||
<taxonomicName id="4C2C26F9F365FE1BFEB5FB26FE7EE9EF" baseAuthorityName="Peters" baseAuthorityYear="1863" box="[297,391,1232,1256]" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="2" pageNumber="23" phylum="Chordata" rank="species" species="affinis">
|
||
<emphasis id="B9588168F365FE1BFEB5FB26FE7EE9EF" box="[297,391,1232,1256]" italics="true" pageId="2" pageNumber="23">P. affinis</emphasis>
|
||
</taxonomicName>
|
||
(45–55),
|
||
<taxonomicName id="4C2C26F9F365FE1BFE6FFB26FD95E9EF" baseAuthorityName="Peters" baseAuthorityYear="1863" box="[499,620,1233,1256]" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="2" pageNumber="23" phylum="Chordata" rank="species" species="montium">
|
||
<emphasis id="B9588168F365FE1BFE6FFB26FD95E9EF" box="[499,620,1233,1256]" italics="true" pageId="2" pageNumber="23">P. montium</emphasis>
|
||
</taxonomicName>
|
||
(35–50),
|
||
<taxonomicName id="4C2C26F9F365FE1BFD4BFB26FEFFE80F" authorityName="Montanucci" authorityYear="1973" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="2" pageNumber="23" phylum="Chordata" rank="species" species="prefrontalis">
|
||
<emphasis id="B9588168F365FE1BFD4BFB26FEFFE80F" italics="true" pageId="2" pageNumber="23">P. prefrontalis</emphasis>
|
||
</taxonomicName>
|
||
(37–46), and
|
||
<taxonomicName id="4C2C26F9F365FE1BFE03FB06FDD3E80F" authorityName="Montanucci" authorityYear="1973" box="[415,554,1264,1288]" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="2" pageNumber="23" phylum="Chordata" rank="species" species="macbrydei">
|
||
<emphasis id="B9588168F365FE1BFE03FB06FDD3E80F" box="[415,554,1264,1288]" italics="true" pageId="2" pageNumber="23">P. macbrydei</emphasis>
|
||
</taxonomicName>
|
||
(31–43).
|
||
</paragraph>
|
||
<caption id="DF530DF2F365FE1BFF11FB81FE5FE9AF" ID-DOI="http://doi.org/10.5281/zenodo.11376952" ID-Zenodo-Dep="11376952" httpUri="https://zenodo.org/record/11376952/files/figure.png" pageId="2" pageNumber="23" startId="2.[141,176,1142,1164]" targetBox="[305,1234,168,1113]" targetPageId="2" targetType="figure">
|
||
<paragraph id="8B935D7AF365FE1BFF11FB81FE5FE9AF" blockId="2.[141,1395,1142,1193]" pageId="2" pageNumber="23">
|
||
<emphasis id="B9588168F365FE1BFF11FB81FF37E98C" bold="true" box="[141,206,1142,1164]" pageId="2" pageNumber="23">Fig. 1.</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="4C2C26F9F365FE1BFECDFB81FDECE98C" authority="Venegas & Echevarría & Lobos & Sales Nunes & Abstract.-Based & The, 2016" authorityName="Venegas & Echevarría & Lobos & Sales Nunes & Abstract.-Based & The" authorityYear="2016" box="[337,533,1142,1163]" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="2" pageNumber="23" phylum="Chordata" rank="species" species="ulisesi" status="sp. nov.">
|
||
<emphasis id="B9588168F365FE1BFECDFB81FDECE98C" box="[337,533,1142,1163]" italics="true" pageId="2" pageNumber="23">Pholidobolus ulisesi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B3C13F365FE1BFD80FB81FD91E98C" box="[540,616,1142,1163]" pageId="2" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(CORBIDI 12734; male, SVL = 45.5 mm) in dorsal (
|
||
<emphasis id="B9588168F365FE1BFBEAFB81FB6EE98C" bold="true" box="[1142,1175,1142,1163]" pageId="2" pageNumber="23">top</emphasis>
|
||
) and ventral (
|
||
<emphasis id="B9588168F365FE1BFABDFB81FA93E98C" bold="true" box="[1313,1386,1142,1163]" pageId="2" pageNumber="23">bottom</emphasis>
|
||
) views. Photographs by OTC.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B935D7AF365FE1BFF11FAD9FD5AEA40" blockId="2.[140,747,1326,1863]" pageId="2" pageNumber="23">
|
||
<emphasis id="B9588168F365FE1BFF11FAD9FE9EE84F" bold="true" box="[141,359,1326,1352]" pageId="2" pageNumber="23">Characterization:</emphasis>
|
||
(1) Two or three supraoculars, anteriormost larger than others; (2) prefrontals absent; (3) femoral pores absent in both sexes; (4) two to six opaque lower eyelid scales; (5) scales on dorsal surface of neck striated, becoming strongly keeled between forelimbs and tail; (6) two or three rows of lateral granules at midbody; (7) lateral body fold present; (8) usually two rows of keeled ventrolateral scales on each side; (9) dorsum dark brown with a distinct pale brown middorsal stripe, slender at midbody, becoming grayish brown towards the tail; (10) labial stripe white becoming cream or pale brown along ventrolateral region; (11) sides of body dark brown; (12) cream stripe along forearm; (13) a distinct diagonal white bar with dark brown edges on each side of the mandible, extending from sixth infralabial to proximal pregular; (14) orange spots on sides of body, usually above forelimb and the base of tail in adult males.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3360EF1F365FE1FFF11F89AFE4DEFEF" lastPageId="6" lastPageNumber="27" pageId="2" pageNumber="23" type="description">
|
||
<paragraph id="8B935D7AF365FE1AFF11F89AFE29EA41" blockId="2.[141,745,1901,1958]" lastBlockId="3.[140,747,1294,1958]" lastPageId="3" lastPageNumber="24" pageId="2" pageNumber="23">
|
||
<emphasis id="B9588168F365FE1BFF11F89AFE2DEA81" bold="true" box="[141,468,1901,1926]" pageId="2" pageNumber="23">
|
||
Description of
|
||
<typeStatus id="5497E3D8F365FE1BFEC0F89AFE29EA81" box="[348,464,1901,1926]" pageId="2" pageNumber="23" type="holotype">holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
Adult male (
|
||
<collectionCode id="ED3DC5BFF365FE1BFDE6F899FD10EA81" box="[634,745,1902,1926]" country="Peru" httpUri="http://grbio.org/cool/g3tc-zhv7" name="Centro de Ornitologia y Biodiversidad" pageId="2" pageNumber="23">CORBIDI</collectionCode>
|
||
12734;
|
||
<figureCitation id="131741FFF365FE1BFF7DF879FEABEAA1" box="[225,338,1934,1958]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="2.[141,176,1142,1164]" captionStartId-1="3.[141,176,1180,1202]" captionStartId-2="4.[141,176,1558,1580]" captionTargetBox-0="[305,1234,168,1113]" captionTargetBox-1="[141,723,169,1166]" captionTargetBox-2="[241,1295,847,1548]" captionTargetId-0="figure-398@2.[285,1252,141,1144]" captionTargetId-1="figure-713@3.[141,745,144,1173]" captionTargetId-2="figure-198@4.[241,1295,847,1550]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="Fig. 1. Holotype of Pholidobolus ulisesi sp. nov. (CORBIDI 12734; male, SVL = 45.5 mm) in dorsal (top) and ventral (bottom) views. Photographs by OTC." captionText-1="Fig. 2. Head of the holotype of Pholidobolus ulisesi sp. nov. (CORBIDI 12734) in dorsal (A), ventral (B), and lateral (C) views. Photographs by OTC." captionText-2="Fig. 3. Four individuals of Pholidobolus ulisesi sp. nov. in life. (A) holotype (CORBIDI 12734); (B) adult female (CORBIDI 12737); (C) juvenile (CORBIDI 12744); (D) adult male from Cañaris (photo voucher). Photographs by PJV." figureDoi-0="http://doi.org/10.5281/zenodo.11376952" figureDoi-1="http://doi.org/10.5281/zenodo.11376954" figureDoi-2="http://doi.org/10.5281/zenodo.11376956" httpUri-0="https://zenodo.org/record/11376952/files/figure.png" httpUri-1="https://zenodo.org/record/11376954/files/figure.png" httpUri-2="https://zenodo.org/record/11376956/files/figure.png" pageId="2" pageNumber="23">Fig. 1–3A</figureCitation>
|
||
); SVL 45.
|
||
<quantity id="4CD4F09FF365FE1BFE59F879FDF1EAA1" box="[453,520,1934,1958]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="23" unit="mm" value="5.0">5 mm</quantity>
|
||
; TL
|
||
<quantity id="4CD4F09FF365FE1BFDA1F879FD62EAA1" box="[573,667,1934,1958]" metricMagnitude="-1" metricUnit="m" metricValue="1.04" pageId="2" pageNumber="23" unit="mm" value="104.0">104 mm</quantity>
|
||
; dorsal and lateral head scales juxtaposed, finely wrinkled; rostral hexagonal, 2.03 times as wide as high; frontonasal quadrangular, wider than long, longer than frontal, laterally in contact with nasal, loreal, and first superciliary; prefrontals absent; frontal pentagonal, longer than wide, slightly wider anteriorly, in contact with frontonasal and supraocular I on each side; frontoparietals hexagonal, longer than wide, with medial suture, each in contact laterally with supraoculars I and II; interparietal roughly heptagonal, its lateral borders parallel to each other; parietals slightly smaller than interparietal, pentagonal and positioned anterolaterally to interparietal, each in contact anteriorly with supraocular II and dorsalmost postocular; postparietals three, medial scale smaller than laterals; supralabials seven, fourth longest and below the center of eye; infralabials five, fourth below the center of eye; temporals enlarged, irregularly pentagonal or hexagonal, juxtaposed, finely wrinkled; two finely wrinkled supratemporals, dorsal conspicuously larger than ventral one; nasal divided, irregularly tetragonal, longer than wide, in contact with rostral anteriorly, first and second supralabials ventrally, frontonasal dorsally, loreal posterodorsally and frenocular posteroventrally; nostril on ventral aspect of nasal, directed lateroposteriorly, piercing nasal suture; loreal rectangular; frenocular enlarged, in contact with nasal, separating loreal from supralabials; supraoculars two, with the first being the largest; four elongate superciliaries, first one enlarged, in contact with loreal; palpebral disk divided into two pigmented scales; suboculars three, elongated and similar in size; three postoculars, ventral one smaller than the others; ear opening vertically oval, without denticulate margins; tympanum recessed into a shallow auditory meatus; mental semicircular, wider than long; postmental pentagonal, slightly wider than long, followed posteriorly by three pairs of genials, the anterior two in contact medially and the posterior one separated by postgenials; all genials in contact with infralabials; gulars imbricate, smooth, widened in two longitudinal rows; gular fold incomplete; posterior row of gulars (collar) with two enlarged scales medially, larger than the anterior gulars.
|
||
</paragraph>
|
||
<caption id="DF530DF2F364FE1AFF11FB6BFE5FE9EC" ID-DOI="http://doi.org/10.5281/zenodo.11376954" ID-Zenodo-Dep="11376954" httpUri="https://zenodo.org/record/11376954/files/figure.png" pageId="3" pageNumber="24" startId="3.[141,176,1180,1202]" targetBox="[141,723,169,1166]" targetPageId="3" targetType="figure">
|
||
<paragraph id="8B935D7AF364FE1AFF11FB6BFE5FE9EC" blockId="3.[141,746,1180,1259]" pageId="3" pageNumber="24">
|
||
<emphasis id="B9588168F364FE1AFF11FB6BFF36E9B6" bold="true" box="[141,207,1180,1202]" pageId="3" pageNumber="24">Fig. 2.</emphasis>
|
||
Head of the holotype of
|
||
<taxonomicName id="4C2C26F9F364FE1AFE53FB6BFD6DE9B6" authority="Venegas & Echevarría & Lobos & Sales Nunes & Abstract.-Based & The, 2016" authorityName="Venegas & Echevarría & Lobos & Sales Nunes & Abstract.-Based & The" authorityYear="2016" box="[463,660,1180,1201]" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="3" pageNumber="24" phylum="Chordata" rank="species" species="ulisesi" status="sp. nov.">
|
||
<emphasis id="B9588168F364FE1AFE53FB6BFD6DE9B6" box="[463,660,1180,1201]" italics="true" pageId="3" pageNumber="24">Pholidobolus ulisesi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B3C13F364FE1AFD01FB6BFD13E9B6" box="[669,746,1180,1201]" pageId="3" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(CORBIDI 12734) in dorsal (
|
||
<emphasis id="B9588168F364FE1AFE29FB4FFE3FE9CA" bold="true" box="[437,454,1208,1229]" pageId="3" pageNumber="24">A</emphasis>
|
||
), ventral (
|
||
<emphasis id="B9588168F364FE1AFDADFB4EFDB8E9C9" bold="true" box="[561,577,1209,1230]" pageId="3" pageNumber="24">B</emphasis>
|
||
), and lateral (
|
||
<emphasis id="B9588168F364FE1AFD4CFB4FFD18E9C9" bold="true" box="[720,737,1208,1230]" pageId="3" pageNumber="24">C</emphasis>
|
||
) views. Photographs by OTC.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B935D7AF364FE1AFF31F8B9FABAE96F" blockId="3.[140,747,1294,1958]" lastBlockId="3.[790,1397,144,1256]" pageId="3" pageNumber="24">
|
||
Scales on nape similar in size to dorsals, except for the anteriormost that are widened; scales on sides of neck small and granular; dorsal scales elongated, imbricate, arranged in transverse rows; dorsal scales on nape striated, becoming progressively keeled from forelimbs to tail; number of dorsal scales between occipital and posterior margin of hind limbs 30; dorsal scale rows in a transverse line at midbody 19; dorsals separated from ventrals by two longitudinal rows of large keeled scales on each side; longitudinal fold between fore and hind limbs present; ventrals smooth, wider than long, arranged in 21 transverse rows between collar fold and preanals; six ventral scales in a transverse row at midbody; subcaudals smooth; limbs overlap when adpressed against body; axillary region composed of granular scales; scales on dorsal surface of forelimb striated, imbricate; scales on ventral surface of forearm small and imbricate, those on ventral surface of arm granular; two thick, smooth thenar scales; supradigitals (left/right) 3/3 on finger I, 6/6 on II, 8/8 on III, 9/9 on IV, 6/6 on
|
||
<collectionCode id="ED3DC5BFF364FE1AFBCEFD66FB9EEFAF" box="[1106,1127,657,680]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="24" type="Museum">V</collectionCode>
|
||
; supradigitals 3/3 on toe I, 6/6 on II, 10/9 on III, 12/11 on IV, 8/8 on
|
||
<collectionCode id="ED3DC5BFF364FE1AFB70FD46FAF8EFCF" box="[1260,1281,689,712]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="24" type="Museum">V</collectionCode>
|
||
; subdigital lamellae of forelimb single, 6/6 on finger I, 11/12 on II, 15/16 on III, 16/16 on IV, 9/8 on
|
||
<collectionCode id="ED3DC5BFF364FE1AFB1AFD06FB62EE0F" box="[1158,1179,753,776]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="24" type="Museum">V</collectionCode>
|
||
; subdigital lamellae on toes I and II single, on toe III paired on the middle, on toe IV paired except for a few ones, on toe
|
||
<collectionCode id="ED3DC5BFF364FE1AFA8BFCC6FAD3EE4F" box="[1303,1322,817,840]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="24" type="Museum">V</collectionCode>
|
||
paired at the base; number of subdigital lamellae (pairs when applicable) 6/6 on toe I, 10/11 on II, 16/17 on III, 21/21 on IV, 12/12 on
|
||
<collectionCode id="ED3DC5BFF364FE1AFC5DFC66FC2FEEAF" box="[961,982,913,936]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="24" type="Museum">V</collectionCode>
|
||
; groin region with small keeled, imbricate scales; scales on dorsal surface of hind limbs keeled and imbricate; scales on ventral surface of hind limbs smooth; scales on posterior surface of thighs granular and on shanks striated and imbricate; femoral pores absent; preanal pores absent; cloacal plate paired, bordered by two scales anteriorly, smaller than cloacal scales.
|
||
</paragraph>
|
||
<paragraph id="8B935D7AF364FE1AFCABFB87FBF8E9EF" blockId="3.[790,1397,144,1256]" pageId="3" pageNumber="24">
|
||
Additional measurements (mm) and proportions of the
|
||
<typeStatus id="5497E3D8F364FE1AFCDDFB67FC5AE9AF" box="[833,931,1168,1192]" pageId="3" pageNumber="24" type="holotype">holotype</typeStatus>
|
||
: HL 9.91; HW 6.95; ShL 3.9; AGD 25.6; TL/SVL 2.05; HL/SVL 0.21; HW/SVL 0.15; ShL/SVL 0.08; AGD/SVL 0.56.
|
||
</paragraph>
|
||
<paragraph id="8B935D7AF364FE1AFC8BFAF9FBE3EBEF" blockId="3.[789,1397,1294,1768]" pageId="3" pageNumber="24">
|
||
<emphasis id="B9588168F364FE1AFC8BFAF9FACEE820" bold="true" box="[791,1335,1294,1320]" pageId="3" pageNumber="24">
|
||
Coloration in preservative (
|
||
<figureCitation id="131741FFF364FE1AFBE5FAF9FB2BE820" box="[1145,1234,1294,1319]" captionStart="Fig" captionStartId="2.[141,176,1142,1164]" captionTargetBox="[305,1234,168,1113]" captionTargetId="figure-398@2.[285,1252,141,1144]" captionTargetPageId="2" captionText="Fig. 1. Holotype of Pholidobolus ulisesi sp. nov. (CORBIDI 12734; male, SVL = 45.5 mm) in dorsal (top) and ventral (bottom) views. Photographs by OTC." figureDoi="http://doi.org/10.5281/zenodo.11376952" httpUri="https://zenodo.org/record/11376952/files/figure.png" pageId="3" pageNumber="24">Figs. 1</figureCitation>
|
||
and
|
||
<figureCitation id="131741FFF364FE1AFA8BFAF9FAD0E820" box="[1303,1321,1294,1319]" captionStart="Fig" captionStartId="3.[141,176,1180,1202]" captionTargetBox="[141,723,169,1166]" captionTargetId="figure-713@3.[141,745,144,1173]" captionTargetPageId="3" captionText="Fig. 2. Head of the holotype of Pholidobolus ulisesi sp. nov. (CORBIDI 12734) in dorsal (A), ventral (B), and lateral (C) views. Photographs by OTC." figureDoi="http://doi.org/10.5281/zenodo.11376954" httpUri="https://zenodo.org/record/11376954/files/figure.png" pageId="3" pageNumber="24">2</figureCitation>
|
||
):
|
||
</emphasis>
|
||
Dorsum dark brown with a grayish brown vertebral stripe that is four scales broad at midbody, and extends from occiput onto tail; vertebral stripe wide anteriorly becoming slightly slender at midbody; dorsal surface of head brown, sides of head and body dark brown; two bright cream spots on each side above insertion of forelimbs; light stripe extending ventrolaterally from lips to insertion of hind limbs, white on lips and grayish brown along the body; a distinct diagonal white bar with dark edges on each side of the mandible, extending from the sixth infralabial onto the proximal pregular; dorsal surface of limbs dark brown with a cream stripe along the arms; gular region pale gray, chest and venter dark gray; ventral surface of tail dark gray.
|
||
</paragraph>
|
||
<paragraph id="8B935D7AF364FE1AFC8BF8F9FB1DEAA0" blockId="3.[790,1396,1806,1959]" pageId="3" pageNumber="24">
|
||
<emphasis id="B9588168F364FE1AFC8BF8F9FAF8EA20" bold="true" box="[791,1281,1806,1832]" pageId="3" pageNumber="24">
|
||
Coloration of
|
||
<typeStatus id="5497E3D8F364FE1AFC5AF8F9FBCCEA20" box="[966,1077,1806,1831]" pageId="3" pageNumber="24" type="holotype">holotype</typeStatus>
|
||
in life (
|
||
<figureCitation id="131741FFF364FE1AFB08F8F9FB0AEA20" box="[1172,1267,1806,1831]" captionStart="Fig" captionStartId="4.[141,176,1558,1580]" captionTargetBox="[241,1295,847,1548]" captionTargetId="figure-198@4.[241,1295,847,1550]" captionTargetPageId="4" captionText="Fig. 3. Four individuals of Pholidobolus ulisesi sp. nov. in life. (A) holotype (CORBIDI 12734); (B) adult female (CORBIDI 12737); (C) juvenile (CORBIDI 12744); (D) adult male from Cañaris (photo voucher). Photographs by PJV." figureDoi="http://doi.org/10.5281/zenodo.11376956" httpUri="https://zenodo.org/record/11376956/files/figure.png" pageId="3" pageNumber="24">Fig. 3A</figureCitation>
|
||
):
|
||
</emphasis>
|
||
Similar to that in preservative, but the bright cream spots on each side above forelimbs are replaced by two black ocelli with red centers, and the sides of the base of the tail have scattered red flecks. The iris is light brown.
|
||
</paragraph>
|
||
<caption id="DF530DF2F363FE1DFF11FF67FB86EDC4" ID-Table-UUID="DF530DF2F363FE1DFF11FF67FB86EDC4" httpUri="http://table.plazi.org/id/DF530DF2F363FE1DFF11FF67FB86EDC4" pageId="4" pageNumber="25" startId="4.[141,197,144,165]" targetBox="[152,1357,213,831]" targetIsTable="true" targetPageId="4" targetType="table">
|
||
<paragraph id="8B935D7AF363FE1DFF11FF67FB86EDC4" blockId="4.[141,1394,144,195]" pageId="4" pageNumber="25">
|
||
<emphasis id="B9588168F363FE1DFF11FF67FF26EDA2" bold="true" box="[141,223,144,165]" pageId="4" pageNumber="25">Table 1.</emphasis>
|
||
Squamation characters of
|
||
<taxonomicName id="4C2C26F9F363FE1DFE79FF67FD51EDA2" authorityName="Venegas & Echevarría & Lobos & Sales Nunes & Abstract.-Based & The" authorityYear="2016" box="[485,680,144,165]" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="4" pageNumber="25" phylum="Chordata" rank="species" species="ulisesi">
|
||
<emphasis id="B9588168F363FE1DFE79FF67FD51EDA2" box="[485,680,144,165]" italics="true" pageId="4" pageNumber="25">Pholidobolus ulisesi</emphasis>
|
||
</taxonomicName>
|
||
. Range, followed by mean ± standard deviation, is given for quantitative characters (if applicable). *Includes adults of both sexes and
|
||
<specimenCount id="9D2A96F3F363FE1DFD45FF5AFCBDEDC4" box="[729,836,173,195]" count="10" pageId="4" pageNumber="25" type="juvenile">10 juvenile</specimenCount>
|
||
specimens of undetermined sex.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B935D7AF363FE1DFC2AFF22FADBEE39" pageId="4" pageNumber="25">
|
||
<table id="F92CAFDAF36301E6FF04FF22FAB4EE38" box="[152,1357,213,831]" gridcols="4" gridrows="10" pageId="4" pageNumber="25">
|
||
<tr id="351C5F38F36301E6FF04FF22FAB4EDEE" box="[152,1357,213,233]" gridrow="0" pageId="4" pageNumber="25" rowspan-0="1" rowspan-1="1" rowspan-3="1">
|
||
<th id="76CD3644F36301E6FC2AFF22FB7CEDEE" box="[950,1157,213,233]" gridcol="2" gridrow="0" pageId="4" pageNumber="25">
|
||
<taxonomicName id="4C2C26F9F363FE1DFC2AFF22FB7CEDEE" authority="Characters" authorityName="Characters" box="[950,1157,213,233]" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="4" pageNumber="25" phylum="Chordata" rank="species" species="ulisesi">
|
||
<emphasis id="B9588168F363FE1DFC2AFF22FB7CEDEE" bold="true" box="[950,1157,213,233]" italics="true" pageId="4" pageNumber="25">Pholidobolus ulisesi</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
</tr>
|
||
<tr id="351C5F38F36301E6FF04FF03FAB4EC20" box="[152,1357,244,295]" gridrow="1" pageId="4" pageNumber="25">
|
||
<th id="76CD3644F36301E6FF04FF03FD54EC20" box="[152,685,244,295]" gridcol="0" gridrow="1" pageId="4" pageNumber="25">
|
||
<emphasis id="B9588168F363FE1DFF04FF03FEFEEC0F" bold="true" box="[152,263,244,264]" pageId="4" pageNumber="25">Characters</emphasis>
|
||
</th>
|
||
<td id="76CD3644F36301E6FC9AFF03FC8AEC20" box="[774,883,244,295]" gridcol="1" gridrow="1" pageId="4" pageNumber="25">
|
||
<emphasis id="B9588168F363FE1DFC83FF0DFCACEC20" bold="true" box="[799,858,250,295]" pageId="4" pageNumber="25">
|
||
Males
|
||
<emphasis id="B9588168F363FE1DFCB8FEE4FCC8EC20" bold="true" box="[804,817,275,295]" italics="true" pageId="4" pageNumber="25">n</emphasis>
|
||
= 5
|
||
</emphasis>
|
||
</td>
|
||
<td id="76CD3644F36301E6FC2AFF03FB7CEC20" box="[950,1157,244,295]" gridcol="2" gridrow="1" pageId="4" pageNumber="25">
|
||
<emphasis id="B9588168F363FE1DFC69FF0DFBC0EC20" bold="true" box="[1013,1098,250,295]" pageId="4" pageNumber="25">
|
||
Females
|
||
<emphasis id="B9588168F363FE1DFB9BFEE4FBEDEC20" bold="true" box="[1031,1044,275,295]" italics="true" pageId="4" pageNumber="25">n</emphasis>
|
||
= 4
|
||
</emphasis>
|
||
</td>
|
||
<td id="76CD3644F36301E6FB29FF03FAB4EC20" box="[1205,1357,244,295]" gridcol="3" gridrow="1" pageId="4" pageNumber="25">
|
||
<emphasis id="B9588168F363FE1DFB29FF0DFAD9EC20" bold="true" pageId="4" pageNumber="25">
|
||
All specimens*
|
||
<emphasis id="B9588168F363FE1DFB7FFEE4FB09EC20" bold="true" box="[1251,1264,275,295]" italics="true" pageId="4" pageNumber="25">n</emphasis>
|
||
= 19
|
||
</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr id="351C5F38F36301E6FF04FECFFAB4EC6C" box="[152,1357,312,363]" gridrow="2" pageId="4" pageNumber="25">
|
||
<th id="76CD3644F36301E6FF04FECFFD54EC6C" box="[152,685,312,363]" gridcol="0" gridrow="2" pageId="4" pageNumber="25">Dorsal scales between occipital and posterior margin of hind limb</th>
|
||
<td id="76CD3644F36301E6FC9AFECFFC8AEC6C" box="[774,883,312,363]" gridcol="1" gridrow="2" pageId="4" pageNumber="25">28–31 29.6 ± 1.14</td>
|
||
<td id="76CD3644F36301E6FC2AFECFFB7CEC6C" box="[950,1157,312,363]" gridcol="2" gridrow="2" pageId="4" pageNumber="25">29–32 30.6 ± 1.14</td>
|
||
<td id="76CD3644F36301E6FB29FECFFAB4EC6C" box="[1205,1357,312,363]" gridcol="3" gridrow="2" pageId="4" pageNumber="25">28–32 30.05 ± 1.13</td>
|
||
</tr>
|
||
<tr id="351C5F38F36301E6FF04FE8CFAB4ECA9" box="[152,1357,379,430]" gridrow="3" pageId="4" pageNumber="25">
|
||
<th id="76CD3644F36301E6FF04FE8CFD54ECA9" box="[152,685,379,430]" gridcol="0" gridrow="3" pageId="4" pageNumber="25">Dorsal scale rows in a transverse line at midbody</th>
|
||
<td id="76CD3644F36301E6FC9AFE8CFC8AECA9" box="[774,883,379,430]" gridcol="1" gridrow="3" pageId="4" pageNumber="25">19–22 20.4 ± 1.14</td>
|
||
<td id="76CD3644F36301E6FC2AFE8CFB7CECA9" box="[950,1157,379,430]" gridcol="2" gridrow="3" pageId="4" pageNumber="25">18–21 19.6 ± 1.34</td>
|
||
<td id="76CD3644F36301E6FB29FE8CFAB4ECA9" box="[1205,1357,379,430]" gridcol="3" gridrow="3" pageId="4" pageNumber="25">17–22 20.05 ± 1.43</td>
|
||
</tr>
|
||
<tr id="351C5F38F36301E6FF04FE49FAB4ECF6" box="[152,1357,446,497]" gridrow="4" pageId="4" pageNumber="25">
|
||
<th id="76CD3644F36301E6FF04FE49FD54ECF6" box="[152,685,446,497]" gridcol="0" gridrow="4" pageId="4" pageNumber="25">Ventral scales between collar fold and preanals</th>
|
||
<td id="76CD3644F36301E6FC9AFE49FC8AECF6" box="[774,883,446,497]" gridcol="1" gridrow="4" pageId="4" pageNumber="25">20–21 20.75 ± 0.5</td>
|
||
<td id="76CD3644F36301E6FC2AFE49FB7CECF6" box="[950,1157,446,497]" gridcol="2" gridrow="4" pageId="4" pageNumber="25">20–23 20.8 ± 1.3</td>
|
||
<td id="76CD3644F36301E6FB29FE49FAB4ECF6" box="[1205,1357,446,497]" gridcol="3" gridrow="4" pageId="4" pageNumber="25">20–23 21.06 ± 0.87</td>
|
||
</tr>
|
||
<tr id="351C5F38F36301E6FF04FDF6FAB4EF34" box="[152,1357,513,563]" gridrow="5" pageId="4" pageNumber="25">
|
||
<th id="76CD3644F36301E6FF04FDF6FD54EF34" box="[152,685,513,563]" gridcol="0" gridrow="5" pageId="4" pageNumber="25">Ventral scale rows in a transverse line at midbody</th>
|
||
<td id="76CD3644F36301E6FC9AFDF6FC8AEF34" box="[774,883,513,563]" gridcol="1" gridrow="5" pageId="4" pageNumber="25">6</td>
|
||
<td id="76CD3644F36301E6FC2AFDF6FB7CEF34" box="[950,1157,513,563]" gridcol="2" gridrow="5" pageId="4" pageNumber="25">6–8 6.8 ± 1.1</td>
|
||
<td id="76CD3644F36301E6FB29FDF6FAB4EF34" box="[1205,1357,513,563]" gridcol="3" gridrow="5" pageId="4" pageNumber="25">6–8 6.56 ± 0.92</td>
|
||
</tr>
|
||
<tr id="351C5F38F36301E6FF04FDB4FAB4EF71" box="[152,1357,579,630]" gridrow="6" pageId="4" pageNumber="25">
|
||
<th id="76CD3644F36301E6FF04FDB4FD54EF71" box="[152,685,579,630]" gridcol="0" gridrow="6" pageId="4" pageNumber="25">Subdigital lamellae on Finger IV</th>
|
||
<td id="76CD3644F36301E6FC9AFDB4FC8AEF71" box="[774,883,579,630]" gridcol="1" gridrow="6" pageId="4" pageNumber="25">15–16 15.6 ± 0.55</td>
|
||
<td id="76CD3644F36301E6FC2AFDB4FB7CEF71" box="[950,1157,579,630]" gridcol="2" gridrow="6" pageId="4" pageNumber="25">15–17 15.8 ± 0.84</td>
|
||
<td id="76CD3644F36301E6FB29FDB4FAB4EF71" box="[1205,1357,579,630]" gridcol="3" gridrow="6" pageId="4" pageNumber="25">11–18 15.05 ± 1.65</td>
|
||
</tr>
|
||
<tr id="351C5F38F36301E6FF04FD71FAB4EFBE" box="[152,1357,646,697]" gridrow="7" pageId="4" pageNumber="25">
|
||
<th id="76CD3644F36301E6FF04FD71FD54EFBE" box="[152,685,646,697]" gridcol="0" gridrow="7" pageId="4" pageNumber="25">Subdigital lamellae on Toe IV</th>
|
||
<td id="76CD3644F36301E6FC9AFD71FC8AEFBE" box="[774,883,646,697]" gridcol="1" gridrow="7" pageId="4" pageNumber="25">20–21 20.4 ± 0.55</td>
|
||
<td id="76CD3644F36301E6FC2AFD71FB7CEFBE" box="[950,1157,646,697]" gridcol="2" gridrow="7" pageId="4" pageNumber="25">18–22 20.6 ± 1.52</td>
|
||
<td id="76CD3644F36301E6FB29FD71FAB4EFBE" box="[1205,1357,646,697]" gridcol="3" gridrow="7" pageId="4" pageNumber="25">15–22 19.32 ± 1.95</td>
|
||
</tr>
|
||
<tr id="351C5F38F36301E6FF04FD3EFAB4EFD8" box="[152,1357,713,735]" gridrow="8" pageId="4" pageNumber="25">
|
||
<th id="76CD3644F36301E6FF04FD3EFD54EFD8" box="[152,685,713,735]" gridcol="0" gridrow="8" pageId="4" pageNumber="25">Maximum SVL</th>
|
||
<td id="76CD3644F36301E6FC9AFD3EFC8AEFD8" box="[774,883,713,735]" gridcol="1" gridrow="8" pageId="4" pageNumber="25">45.52</td>
|
||
<td id="76CD3644F36301E6FC2AFD3EFB7CEFD8" box="[950,1157,713,735]" gridcol="2" gridrow="8" pageId="4" pageNumber="25">57.46</td>
|
||
<td id="76CD3644F36301E6FB29FD3EFAB4EFD8" box="[1205,1357,713,735]" gridcol="3" gridrow="8" pageId="4" pageNumber="25">57.46</td>
|
||
</tr>
|
||
<tr id="351C5F38F36301E6FF04FD18FAB4EE38" box="[152,1357,751,831]" gridrow="9" pageId="4" pageNumber="25">
|
||
<th id="76CD3644F36301E6FF04FD18FD54EE38" box="[152,685,751,831]" gridcol="0" gridrow="9" pageId="4" pageNumber="25">TL/SVL</th>
|
||
<td id="76CD3644F36301E6FC9AFD18FC8AEE38" box="[774,883,751,831]" gridcol="1" gridrow="9" pageId="4" pageNumber="25">
|
||
1.92–2.28 2.12 ± 0.18 (
|
||
<emphasis id="B9588168F363FE1DFCBFFCDEFCD6EE39" box="[803,815,809,830]" italics="true" pageId="4" pageNumber="25">n</emphasis>
|
||
= 3)
|
||
</td>
|
||
<td id="76CD3644F36301E6FC2AFD18FB7CEE38" box="[950,1157,751,831]" gridcol="2" gridrow="9" pageId="4" pageNumber="25">
|
||
1.83–2.17 2.05 ± 0.19 (
|
||
<emphasis id="B9588168F363FE1DFB9BFCDEFBEAEE39" box="[1031,1043,809,830]" italics="true" pageId="4" pageNumber="25">n</emphasis>
|
||
= 3)
|
||
</td>
|
||
<td id="76CD3644F36301E6FB29FD18FAB4EE38" box="[1205,1357,751,831]" gridcol="3" gridrow="9" pageId="4" pageNumber="25">
|
||
1.83–2.28 2.05 ± 0.18 (
|
||
<emphasis id="B9588168F363FE1DFB74FCDEFB0DEE39" box="[1256,1268,809,830]" italics="true" pageId="4" pageNumber="25">n</emphasis>
|
||
= 7)
|
||
</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="DF530DF2F363FE1DFF11F9E1FB58EB4E" ID-DOI="http://doi.org/10.5281/zenodo.11376956" ID-Zenodo-Dep="11376956" httpUri="https://zenodo.org/record/11376956/files/figure.png" pageId="4" pageNumber="25" startId="4.[141,176,1558,1580]" targetBox="[241,1295,847,1548]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8B935D7AF363FE1DFF11F9E1FB58EB4E" blockId="4.[141,1394,1558,1609]" pageId="4" pageNumber="25">
|
||
<emphasis id="B9588168F363FE1DFF11F9E1FF29EB2C" bold="true" box="[141,208,1558,1580]" pageId="4" pageNumber="25">Fig. 3.</emphasis>
|
||
Four individuals of
|
||
<taxonomicName id="4C2C26F9F363FE1DFE03F9E1FD9CEB2C" authority="Venegas & Echevarría & Lobos & Sales Nunes & Abstract.-Based & The, 2016" authorityName="Venegas & Echevarría & Lobos & Sales Nunes & Abstract.-Based & The" authorityYear="2016" box="[415,613,1558,1579]" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="4" pageNumber="25" phylum="Chordata" rank="species" species="ulisesi" status="sp. nov.">
|
||
<emphasis id="B9588168F363FE1DFE03F9E1FD9CEB2C" box="[415,613,1558,1579]" italics="true" pageId="4" pageNumber="25">Pholidobolus ulisesi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B3C13F363FE1DFDF2F9E0FD45EB2B" box="[622,700,1559,1580]" pageId="4" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
|
||
in life. (
|
||
<emphasis id="B9588168F363FE1DFC8AF9E1FCDEEB2C" bold="true" box="[790,807,1558,1579]" pageId="4" pageNumber="25">A</emphasis>
|
||
) holotype (CORBIDI 12734); (
|
||
<emphasis id="B9588168F363FE1DFBF8F9E1FB8DEB2C" bold="true" box="[1124,1140,1558,1579]" pageId="4" pageNumber="25">B</emphasis>
|
||
) adult female (CORBIDI 12737); (
|
||
<emphasis id="B9588168F363FE1DFF7AF9C4FF0EEB4E" bold="true" box="[230,247,1587,1609]" pageId="4" pageNumber="25">C</emphasis>
|
||
) juvenile (CORBIDI 12744); (
|
||
<emphasis id="B9588168F363FE1DFDBEF9C4FDCAEB4F" bold="true" box="[546,563,1587,1608]" pageId="4" pageNumber="25">D</emphasis>
|
||
) adult male from Cañaris (photo voucher). Photographs by PJV.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B935D7AF363FE1DFF11F9A9FB6AEA91" blockId="4.[141,747,1630,1975]" lastBlockId="4.[790,1397,1630,1942]" pageId="4" pageNumber="25">
|
||
<emphasis id="B9588168F363FE1DFF11F9A9FEFEEB70" bold="true" box="[141,263,1630,1655]" pageId="4" pageNumber="25">Variation:</emphasis>
|
||
Variation in measurements and scutellation of
|
||
<taxonomicName id="4C2C26F9F363FE1DFF30F988FE7EEB90" authorityName="Venegas & Echevarría & Lobos & Sales Nunes & Abstract.-Based & The" authorityYear="2016" box="[172,391,1663,1687]" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="4" pageNumber="25" phylum="Chordata" rank="species" species="ulisesi">
|
||
<emphasis id="B9588168F363FE1DFF30F988FE7EEB90" box="[172,391,1663,1687]" italics="true" pageId="4" pageNumber="25">Pholidobolus ulisesi</emphasis>
|
||
</taxonomicName>
|
||
is presented in
|
||
<tableCitation id="C6AE68C1F363FE1DFDAAF988FD71EB90" box="[566,648,1663,1687]" captionStart="Table 1" captionStartId="4.[141,197,144,165]" captionTargetId="graphics-242@4.[141,1393,502,569]" captionTargetPageId="4" captionText="Table 1. Squamation characters of Pholidobolus ulisesi. Range, followed by mean ± standard deviation, is given for quantitative" httpUri="http://table.plazi.org/id/DF530DF2F363FE1DFF11FF67FB86EDC4" pageId="4" pageNumber="25" tableUuid="DF530DF2F363FE1DFF11FF67FB86EDC4">Table 1</tableCitation>
|
||
. Usually two supraoculars, 2/3 (left/right) in specimen
|
||
<specimenCode id="DB8AF501F363FE1DFD0DF968FF00EBD0" collectionCode="CORBIDI" country="Peru" httpUri="http://grbio.org/cool/g3tc-zhv7" name="Centro de Ornitologia y Biodiversidad" pageId="4" pageNumber="25">CORBIDI 12742</specimenCode>
|
||
; superciliaries usually four, 3/
|
||
<quantity id="4CD4F09FF363FE1DFDDAF948FD8BEBD0" box="[582,626,1727,1751]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="4" pageNumber="25" unit="in" value="4.0">4 in</quantity>
|
||
<specimenCode id="DB8AF501F363FE1DFDE0F948FF2BEBF0" collectionCode="CORBIDI" country="Peru" httpUri="http://grbio.org/cool/g3tc-zhv7" name="Centro de Ornitologia y Biodiversidad" pageId="4" pageNumber="25">CORBIDI 12749</specimenCode>
|
||
,
|
||
<specimenCode id="DB8AF501F363FE1DFF7EF928FF16EBF0" box="[226,239,1759,1783]" collectionCode="CORBIDI" country="Peru" httpUri="http://grbio.org/cool/g3tc-zhv7" name="Centro de Ornitologia y Biodiversidad" pageId="4" pageNumber="25">6</specimenCode>
|
||
/
|
||
<quantity id="4CD4F09FF363FE1DFF64F928FEDDEBF0" box="[248,292,1759,1783]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="4" pageNumber="25" unit="in" value="5.0">5 in</quantity>
|
||
<specimenCode id="DB8AF501F363FE1DFEB3F928FE14EBF0" box="[303,493,1759,1783]" collectionCode="CORBIDI" country="Peru" httpUri="http://grbio.org/cool/g3tc-zhv7" name="Centro de Ornitologia y Biodiversidad" pageId="4" pageNumber="25">CORBIDI 00873</specimenCode>
|
||
, and 5/
|
||
<quantity id="4CD4F09FF363FE1DFDD8F928FD88EBF0" box="[580,625,1759,1783]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="4" pageNumber="25" unit="in" value="5.0">5 in</quantity>
|
||
<specimenCode id="DB8AF501F363FE1DFDE0F928FF2AEA10" collectionCode="CORBIDI" country="Peru" httpUri="http://grbio.org/cool/g3tc-zhv7" name="Centro de Ornitologia y Biodiversidad" pageId="4" pageNumber="25">CORBIDI 00872</specimenCode>
|
||
; little intrusive scales present on each side, in the posterior angle of frontonasal in
|
||
<specimenCount id="9D2A96F3F363FE1DFE73F8E8FD63EA30" box="[495,666,1823,1847]" count="3" pageId="4" pageNumber="25" type="generic">three specimens</specimenCount>
|
||
(
|
||
<specimenCode id="DB8AF501F363FE1DFD35F8E8FEE8EA50" collectionCode="CORBIDI" country="Peru" httpUri="http://grbio.org/cool/g3tc-zhv7" name="Centro de Ornitologia y Biodiversidad" pageId="4" pageNumber="25">CORBIDI 12735</specimenCode>
|
||
,
|
||
<specimenCode id="DB8AF501F363FE1DFE81F8C8FE9BEA50" box="[285,354,1855,1879]" collectionCode="CORBIDI" country="Peru" httpUri="http://grbio.org/cool/g3tc-zhv7" name="Centro de Ornitologia y Biodiversidad" pageId="4" pageNumber="25">12741</specimenCode>
|
||
,
|
||
<specimenCode id="DB8AF501F363FE1DFEF3F8C8FE4FEA50" box="[367,438,1855,1879]" collectionCode="CORBIDI" country="Peru" httpUri="http://grbio.org/cool/g3tc-zhv7" name="Centro de Ornitologia y Biodiversidad" pageId="4" pageNumber="25">12744</specimenCode>
|
||
); usually seven supralabials, 7/
|
||
<quantity id="4CD4F09FF363FE1DFF38F8A8FF2BEA70" box="[164,210,1887,1911]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="4" pageNumber="25" unit="in" value="6.0">6 in</quantity>
|
||
<specimenCode id="DB8AF501F363FE1DFF42F8A8FE65EA70" box="[222,412,1887,1911]" collectionCode="CORBIDI" country="Peru" httpUri="http://grbio.org/cool/g3tc-zhv7" name="Centro de Ornitologia y Biodiversidad" pageId="4" pageNumber="25">CORBIDI 00871</specimenCode>
|
||
,
|
||
<specimenCode id="DB8AF501F363FE1DFE31F8A8FE09EA70" box="[429,496,1887,1911]" collectionCode="CORBIDI" country="Peru" httpUri="http://grbio.org/cool/g3tc-zhv7" name="Centro de Ornitologia y Biodiversidad" pageId="4" pageNumber="25">12738</specimenCode>
|
||
and 6/
|
||
<quantity id="4CD4F09FF363FE1DFDDFF8A8FD88EA70" box="[579,625,1887,1911]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="4" pageNumber="25" unit="in" value="6.0">6 in</quantity>
|
||
<specimenCode id="DB8AF501F363FE1DFDE1F8A8FF05EA90" collectionCode="CORBIDI" country="Peru" httpUri="http://grbio.org/cool/g3tc-zhv7" name="Centro de Ornitologia y Biodiversidad" pageId="4" pageNumber="25">CORBIDI 12742–43</specimenCode>
|
||
; infralabials usually six, 5/
|
||
<quantity id="4CD4F09FF363FE1DFDA0F888FD94EA90" box="[572,621,1919,1943]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="4" pageNumber="25" unit="in" value="5.0">5 in</quantity>
|
||
<specimenCode id="DB8AF501F363FE1DFDE0F888FF2BEAB0" collectionCode="CORBIDI" country="Peru" httpUri="http://grbio.org/cool/g3tc-zhv7" name="Centro de Ornitologia y Biodiversidad" pageId="4" pageNumber="25">CORBIDI 12738</specimenCode>
|
||
,
|
||
<specimenCode id="DB8AF501F363FE1DFF7EF868FEDEEAB0" box="[226,295,1951,1975]" collectionCode="CORBIDI" country="Peru" httpUri="http://grbio.org/cool/g3tc-zhv7" name="Centro de Ornitologia y Biodiversidad" pageId="4" pageNumber="25">12740</specimenCode>
|
||
,
|
||
<specimenCode id="DB8AF501F363FE1DFEABF868FE85EAB0" box="[311,380,1951,1975]" collectionCode="CORBIDI" country="Peru" httpUri="http://grbio.org/cool/g3tc-zhv7" name="Centro de Ornitologia y Biodiversidad" pageId="4" pageNumber="25">12742</specimenCode>
|
||
,
|
||
<specimenCode id="DB8AF501F363FE1DFE17F868FE61EAB0" box="[395,408,1951,1975]" collectionCode="CORBIDI" country="Peru" httpUri="http://grbio.org/cool/g3tc-zhv7" name="Centro de Ornitologia y Biodiversidad" pageId="4" pageNumber="25">6</specimenCode>
|
||
/
|
||
<quantity id="4CD4F09FF363FE1DFE3DF868FE37EAB0" box="[417,462,1951,1975]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="4" pageNumber="25" unit="in" value="5.0">5 in</quantity>
|
||
<specimenCode id="DB8AF501F363FE1DFE45F868FD6EEAB0" box="[473,663,1951,1975]" collectionCode="CORBIDI" country="Peru" httpUri="http://grbio.org/cool/g3tc-zhv7" name="Centro de Ornitologia y Biodiversidad" pageId="4" pageNumber="25">CORBIDI 00873</specimenCode>
|
||
,
|
||
<specimenCode id="DB8AF501F363FE1DFD3BF868FD13EAB0" box="[679,746,1951,1975]" collectionCode="CORBIDI" country="Peru" httpUri="http://grbio.org/cool/g3tc-zhv7" name="Centro de Ornitologia y Biodiversidad" pageId="4" pageNumber="25">12744</specimenCode>
|
||
and 5/
|
||
<quantity id="4CD4F09FF363FE1DFCC7F9A9FC7CEB71" box="[859,901,1630,1654]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="4" pageNumber="25" unit="in" value="6.0">6 in</quantity>
|
||
<specimenCode id="DB8AF501F363FE1DFC11F9A9FBB0EB71" box="[909,1097,1630,1654]" collectionCode="CORBIDI" country="Peru" httpUri="http://grbio.org/cool/g3tc-zhv7" name="Centro de Ornitologia y Biodiversidad" pageId="4" pageNumber="25">CORBIDI 12735</specimenCode>
|
||
,
|
||
<specimenCode id="DB8AF501F363FE1DFBC9F9A9FB62EB71" box="[1109,1179,1630,1654]" collectionCode="CORBIDI" country="Peru" httpUri="http://grbio.org/cool/g3tc-zhv7" name="Centro de Ornitologia y Biodiversidad" pageId="4" pageNumber="25">12743</specimenCode>
|
||
. Rows of ventrolateral keeled scales vary from two rows in
|
||
<specimenCount id="9D2A96F3F363FE1DFB53F989FA8AEB91" box="[1231,1395,1662,1686]" count="9" pageId="4" pageNumber="25" type="generic">nine specimens</specimenCount>
|
||
(56% of the
|
||
<typeStatus id="5497E3D8F363FE1DFC3EF968FC29EBB0" box="[930,976,1695,1719]" pageId="4" pageNumber="25">type</typeStatus>
|
||
series), one row on each side in
|
||
<specimenCount id="9D2A96F3F363FE1DFAA2F969FC71EBD1" count="3" pageId="4" pageNumber="25" type="generic">three specimens</specimenCount>
|
||
(
|
||
<specimenCode id="DB8AF501F363FE1DFC04F949FBACEBD1" box="[920,1109,1726,1750]" collectionCode="CORBIDI" country="Peru" httpUri="http://grbio.org/cool/g3tc-zhv7" name="Centro de Ornitologia y Biodiversidad" pageId="4" pageNumber="25">CORBIDI 00872</specimenCode>
|
||
,
|
||
<specimenCode id="DB8AF501F363FE1DFBFFF949FB51EBD1" box="[1123,1192,1726,1750]" collectionCode="CORBIDI" country="Peru" httpUri="http://grbio.org/cool/g3tc-zhv7" name="Centro de Ornitologia y Biodiversidad" pageId="4" pageNumber="25">12741</specimenCode>
|
||
, and 12745), three rows on each side in
|
||
<specimenCount id="9D2A96F3F363FE1DFB9FF928FB61EBF1" box="[1027,1176,1758,1782]" count="1" pageId="4" pageNumber="25" type="generic">one specimen</specimenCount>
|
||
(
|
||
<specimenCode id="DB8AF501F363FE1DFB34F929FA9FEBF1" box="[1192,1382,1758,1782]" collectionCode="CORBIDI" country="Peru" httpUri="http://grbio.org/cool/g3tc-zhv7" name="Centro de Ornitologia y Biodiversidad" pageId="4" pageNumber="25">CORBIDI 12739</specimenCode>
|
||
), and absent in
|
||
<specimenCount id="9D2A96F3F363FE1DFC27F908FBDCEA11" box="[955,1061,1790,1814]" count="2" pageId="4" pageNumber="25" type="adult">two adult</specimenCount>
|
||
specimens (
|
||
<specimenCode id="DB8AF501F363FE1DFB28F909FA8AEA11" box="[1204,1395,1790,1814]" collectionCode="CORBIDI" country="Peru" httpUri="http://grbio.org/cool/g3tc-zhv7" name="Centro de Ornitologia y Biodiversidad" pageId="4" pageNumber="25">CORBIDI 00871</specimenCode>
|
||
and
|
||
<specimenCode id="DB8AF501F363FE1DFCDBF8E9FBFCEA31" box="[839,1029,1822,1846]" collectionCode="CORBIDI" country="Peru" httpUri="http://grbio.org/cool/g3tc-zhv7" name="Centro de Ornitologia y Biodiversidad" pageId="4" pageNumber="25">CORBIDI 00873</specimenCode>
|
||
). Usually two scales on posterior cloacal plate, only
|
||
<specimenCount id="9D2A96F3F363FE1DFC43F8C8FB85EA51" box="[991,1148,1854,1878]" count="2" pageId="4" pageNumber="25" type="generic">two specimens</specimenCount>
|
||
(
|
||
<specimenCode id="DB8AF501F363FE1DFB15F8C9FA92EA51" box="[1161,1387,1854,1878]" collectionCode="CORBIDI" country="Peru" httpUri="http://grbio.org/cool/g3tc-zhv7" name="Centro de Ornitologia y Biodiversidad" pageId="4" pageNumber="25">CORBIDI 12737–38</specimenCode>
|
||
) have three scales, and two
|
||
<typeStatus id="5497E3D8F363FE1DFBDFF8A9FB0BEA71" box="[1091,1266,1886,1910]" isOtherTypeStatus="true" pageId="4" pageNumber="25">other specimens</typeStatus>
|
||
(
|
||
<specimenCode id="DB8AF501F363FE1DFA9FF8A9FCA3EA91" collectionCode="CORBIDI" country="Peru" httpUri="http://grbio.org/cool/g3tc-zhv7" name="Centro de Ornitologia y Biodiversidad" pageId="4" pageNumber="25">CORBIDI 00871</specimenCode>
|
||
and 00873) have four scales.
|
||
</paragraph>
|
||
<paragraph id="8B935D7AF362FE1CFC8BFF67FCA0EC2F" blockId="5.[791,1395,144,296]" pageId="5" pageNumber="26">
|
||
(maximum SVL 57.
|
||
<quantity id="4CD4F09FF362FE1CFC70FF67FBD7EDAF" box="[1004,1070,144,168]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="26" unit="mm" value="4.0">4 mm</quantity>
|
||
,
|
||
<emphasis id="B9588168F362FE1CFBA6FF66FBBEEDAF" box="[1082,1095,145,168]" italics="true" pageId="5" pageNumber="26">n</emphasis>
|
||
= 4) than males (maximum SVL 45.
|
||
<quantity id="4CD4F09FF362FE1CFCE8FF47FC4EEDCF" box="[884,951,176,200]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="26" unit="mm" value="5.0">5 mm</quantity>
|
||
,
|
||
<emphasis id="B9588168F362FE1CFC58FF46FC28EDCF" box="[964,977,177,200]" italics="true" pageId="5" pageNumber="26">n</emphasis>
|
||
= 5). Juvenile
|
||
<collectionCode id="ED3DC5BFF362FE1CFBE0FF47FB13EDCF" box="[1148,1258,176,200]" country="Peru" httpUri="http://grbio.org/cool/g3tc-zhv7" name="Centro de Ornitologia y Biodiversidad" pageId="5" pageNumber="26">CORBIDI</collectionCode>
|
||
12743 (
|
||
<figureCitation id="131741FFF362FE1CFADBFF47FCCFEDEF" captionStart="Fig" captionStartId="4.[141,176,1558,1580]" captionTargetBox="[241,1295,847,1548]" captionTargetId="figure-198@4.[241,1295,847,1550]" captionTargetPageId="4" captionText="Fig. 3. Four individuals of Pholidobolus ulisesi sp. nov. in life. (A) holotype (CORBIDI 12734); (B) adult female (CORBIDI 12737); (C) juvenile (CORBIDI 12744); (D) adult male from Cañaris (photo voucher). Photographs by PJV." figureDoi="http://doi.org/10.5281/zenodo.11376956" httpUri="https://zenodo.org/record/11376956/files/figure.png" pageId="5" pageNumber="26">Fig. 3C</figureCitation>
|
||
) differs from adults in having a fragmented dirty cream stripe along the flanks above the ventrolateral stripe.
|
||
</paragraph>
|
||
<caption id="DF530DF2F362FE1CFF11F9EBFD71EBA2" ID-DOI="http://doi.org/10.5281/zenodo.11376960" ID-Zenodo-Dep="11376960" httpUri="https://zenodo.org/record/11376960/files/figure.png" pageId="5" pageNumber="26" startId="5.[141,176,1564,1586]" targetBox="[141,745,144,1554]" targetPageId="5" targetType="figure">
|
||
<paragraph id="8B935D7AF362FE1CFF11F9EBFD71EBA2" blockId="5.[138,749,1564,1701]" pageId="5" pageNumber="26">
|
||
<emphasis id="B9588168F362FE1CFF11F9EBFF29EB36" bold="true" box="[141,208,1564,1586]" pageId="5" pageNumber="26">Fig. 4.</emphasis>
|
||
Four species of
|
||
<taxonomicName id="4C2C26F9F362FE1CFEE0F9EBFE05EB36" box="[380,508,1564,1585]" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="5" pageNumber="26" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9588168F362FE1CFEE0F9EBFE05EB36" box="[380,508,1564,1585]" italics="true" pageId="5" pageNumber="26">Pholidobolus</emphasis>
|
||
</taxonomicName>
|
||
. (
|
||
<emphasis id="B9588168F362FE1CFD88F9EBFDDCEB36" bold="true" box="[532,549,1564,1585]" pageId="5" pageNumber="26">A</emphasis>
|
||
) adult female of
|
||
<taxonomicName id="4C2C26F9F362FE1CFD44F9EBFF3FEB49" baseAuthorityName="Uzzell" baseAuthorityYear="1973" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="5" pageNumber="26" phylum="Chordata" rank="species" species="dicrus">
|
||
<emphasis id="B9588168F362FE1CFD44F9EBFF3FEB49" italics="true" pageId="5" pageNumber="26">P. dicrus</emphasis>
|
||
</taxonomicName>
|
||
(QCAZ 5304); (
|
||
<emphasis id="B9588168F362FE1CFEF5F9CEFE80EB49" bold="true" box="[361,377,1593,1614]" pageId="5" pageNumber="26">B</emphasis>
|
||
) adult male of
|
||
<taxonomicName id="4C2C26F9F362FE1CFD94F9CEFDAFEB49" box="[520,598,1593,1614]" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="5" pageNumber="26" phylum="Chordata" rank="species" species="hillisi">
|
||
<emphasis id="B9588168F362FE1CFD94F9CEFDAFEB49" box="[520,598,1593,1614]" italics="true" pageId="5" pageNumber="26">P. hillisi</emphasis>
|
||
</taxonomicName>
|
||
(QCAZ 4999); (
|
||
<emphasis id="B9588168F362FE1CFF09F9A2FF5FEB6C" bold="true" box="[149,166,1621,1643]" pageId="5" pageNumber="26">C</emphasis>
|
||
) a juvenile of
|
||
<taxonomicName id="4C2C26F9F362FE1CFEAEF9A1FE48EB6D" authorityName="O'Shaughnessy" authorityYear="1879" box="[306,433,1621,1643]" class="Squamata" family="Gymnophthalmidae" genus="Cercosaura" kingdom="Animalia" pageId="5" pageNumber="26" phylum="Chordata" rank="species" species="vertebralis">
|
||
<emphasis id="B9588168F362FE1CFEAEF9A1FE48EB6D" box="[306,433,1621,1643]" italics="true" pageId="5" pageNumber="26">P. vertebralis</emphasis>
|
||
</taxonomicName>
|
||
(QCAZ 5082); (
|
||
<emphasis id="B9588168F362FE1CFDCFF9A1FD9DEB6C" bold="true" box="[595,612,1622,1643]" pageId="5" pageNumber="26">D</emphasis>
|
||
) adult female of
|
||
<emphasis id="B9588168F362FE1CFF36F984FF39EB8F" box="[170,192,1651,1672]" italics="true" pageId="5" pageNumber="26">P.</emphasis>
|
||
sp. from La Granja (CORBIDI 1678). Photographs by: (A) and (B) Santiago R. Ron, (C) OTC, and (D) PJV.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B935D7AF362FE1CFF31F93AFB82EE60" blockId="5.[141,746,1741,1926]" lastBlockId="5.[787,1399,334,1927]" pageId="5" pageNumber="26">
|
||
Males can be distinguished from females by having the contacted margins of rostral and mental distinctly dark brown or black (indistinct or not contrasting in females), and by the presence of red or orange spots above the insertion of forelimbs and on the sides of the base of tail (absent in females;
|
||
<figureCitation id="131741FFF362FE1CFE33F89AFDFDEA81" box="[431,516,1901,1926]" captionStart="Fig" captionStartId="4.[141,176,1558,1580]" captionTargetBox="[241,1295,847,1548]" captionTargetId="figure-198@4.[241,1295,847,1550]" captionTargetPageId="4" captionText="Fig. 3. Four individuals of Pholidobolus ulisesi sp. nov. in life. (A) holotype (CORBIDI 12734); (B) adult female (CORBIDI 12737); (C) juvenile (CORBIDI 12744); (D) adult male from Cañaris (photo voucher). Photographs by PJV." figureDoi="http://doi.org/10.5281/zenodo.11376956" httpUri="https://zenodo.org/record/11376956/files/figure.png" pageId="5" pageNumber="26">Fig. 3B</figureCitation>
|
||
). Females are longer
|
||
<emphasis id="B9588168F362FE1CFC8BFEB9FBABEC60" bold="true" box="[791,1106,334,359]" pageId="5" pageNumber="26">Hemipenial morphology:</emphasis>
|
||
The left hemipenis of the
|
||
<typeStatus id="5497E3D8F362FE1CFC8BFE98FC8DEC80" box="[791,884,367,391]" pageId="5" pageNumber="26" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C2C26F9F362FE1CFC3EFE98FB78EC80" authorityName="Venegas & Echevarría & Lobos & Sales Nunes & Abstract.-Based & The" authorityYear="2016" box="[930,1153,367,391]" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="5" pageNumber="26" phylum="Chordata" rank="species" species="ulisesi">
|
||
<emphasis id="B9588168F362FE1CFC3EFE98FB78EC80" box="[930,1153,367,391]" italics="true" pageId="5" pageNumber="26">Pholidobolus ulisesi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="131741FFF362FE1CFB09FE98FB23EC80" box="[1173,1242,367,391]" captionStart="Fig" captionStartId="6.[141,176,1912,1934]" captionTargetBox="[231,1304,1307,1897]" captionTargetId="figure-614@6.[231,1304,1307,1897]" captionTargetPageId="6" captionText="Fig. 5. Left hemipenis of Pholidobolus ulisesi sp. nov. (CORBIDI 12734 - holotype) in sulcate (left), lateral (middle), and asulcate (right) views. Photographs by PMSN." figureDoi="http://doi.org/10.5281/zenodo.11376962" httpUri="https://zenodo.org/record/11376962/files/figure.png" pageId="5" pageNumber="26">Fig. 5</figureCitation>
|
||
) was everted during preservation and prepared posteriorly. The organ extends along approximately eight millimeters in length. The lobes of the organ are partially everted and the hemipenis is fully expanded. The hemipenial body is roughly conical in shape, with the basis distinctly thinner than the rest of the organ, and bears two small lobes with apical folds in the apex. The sulcus spermaticus is central in position, originating at the base of the organ, and proceeding in a straight line towards the lobes. The sulcus is broader in the region of the lobular crotch, where it is divided by a small fleshy fold; its branches lie on the medial region of the lobes, and end in their tips among folds. The sulcate face of the hemipenial body presents two nude areas parallel to the sulcus spermaticus that run along the entire hemipenial body.
|
||
</paragraph>
|
||
<paragraph id="8B935D7AF362FE1CFCABFC98FA94E8C0" blockId="5.[787,1399,334,1927]" pageId="5" pageNumber="26">
|
||
The lateral and asulcate faces of the hemipenis are ornamented with a series of roughly equidistant flounces with calcareous spinules. Twenty-three rows of flounces extend along the body of the organ. There are four proximal rows restricted to a central position on the basal asulcate face of the hemipenis, all of them are roughly chevron-shaped. The four proximal flounces on the sides are diagonally positioned; the third to fifth flounces are separated from a complementary flounce positioned on the asulcate face and oriented in an inverse diagonal. The subsequent flounces towards the lobes cross the sides of the organ from the sulcate to the asulcate face, forming chevrons with vertices in the central region of each side pointing towards the basis of the organ. These chevron-shaped rows become reduced in size progressively towards the hemipenial apex. Similar to the description of the hemipenis of
|
||
<taxonomicName id="4C2C26F9F362FE1CFC63FA87FB05E880" authorityName="O'Shaughnessy" authorityYear="1879" box="[1023,1276,1391,1415]" class="Squamata" family="Gymnophthalmidae" genus="Cercosaura" kingdom="Animalia" pageId="5" pageNumber="26" phylum="Chordata" rank="species" species="vertebralis">
|
||
<emphasis id="B9588168F362FE1CFC63FA87FB05E880" box="[1023,1276,1391,1415]" italics="true" pageId="5" pageNumber="26">Cercosaura vertebralis</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFBD208BF362FE1CFAB1FA98FC99E8A0" author="Uzzell T." pageId="5" pageNumber="26" pagination="1 - 67" refId="ref7912" refString="Uzzell T. 1973. A revision of the genus Prionodactylus with a new genus for P. leucostictus and notes on the genus Euspondylus (Sauria, Teiidae). Postilla 159: 1 - 67." type="journal article" year="1973">Uzzell (1973)</bibRefCitation>
|
||
, the five distalmost lateral flounces of the hemipenis have an enlarged tooth in the vertex of the chevrons.
|
||
</paragraph>
|
||
<paragraph id="8B935D7AF362FE1CFCABFA38FADDEA00" blockId="5.[787,1399,334,1927]" pageId="5" pageNumber="26">
|
||
The lateral flounces are separated in two groups by a nude area in the central asulcate face that increases in size in the apical region, becoming Y-shaped. The region between the asulcate and lateral sides are marked by a conspicuous unevenness forming a distinctive bulge, which is also present in other species of the
|
||
<emphasis id="B9588168F362FE1CFB71F998FCC5EBA0" italics="true" pageId="5" pageNumber="26">Macropholidus</emphasis>
|
||
+
|
||
<taxonomicName id="4C2C26F9F362FE1CFCFAF978FC0DEBA0" box="[870,1012,1679,1703]" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="5" pageNumber="26" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9588168F362FE1CFCFAF978FC0DEBA0" box="[870,1012,1679,1703]" italics="true" pageId="5" pageNumber="26">Pholidobolus</emphasis>
|
||
</taxonomicName>
|
||
clade (
|
||
<taxonomicName id="4C2C26F9F362FE1CFBCEF978FA95EBA0" authorityName="Parker" authorityYear="1930" box="[1106,1388,1679,1703]" class="Squamata" family="Gymnophthalmidae" genus="Macropholidus" kingdom="Animalia" pageId="5" pageNumber="26" phylum="Chordata" rank="species" species="annectens">
|
||
<emphasis id="B9588168F362FE1CFBCEF978FA95EBA0" box="[1106,1388,1679,1703]" italics="true" pageId="5" pageNumber="26">Macropholidus annectens</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2C26F9F362FE1CFC8BF947FC27EBC0" authorityName="Reeder" authorityYear="1996" box="[791,990,1711,1735]" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="5" pageNumber="26" phylum="Chordata" rank="species" species="huancabambae">
|
||
<emphasis id="B9588168F362FE1CFC8BF947FC27EBC0" box="[791,990,1711,1735]" italics="true" pageId="5" pageNumber="26">M. huancabambae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2C26F9F362FE1CFC71F947FB93EBC0" authorityName="Noble" authorityYear="1921" box="[1005,1130,1711,1735]" class="Squamata" family="Gymnophthalmidae" genus="Macropholidus" kingdom="Animalia" pageId="5" pageNumber="26" phylum="Chordata" rank="species" species="ruthveni">
|
||
<emphasis id="B9588168F362FE1CFC71F947FB93EBC0" box="[1005,1130,1711,1735]" italics="true" pageId="5" pageNumber="26">M. ruthveni</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2C26F9F362FE1CFBE4F958FAA9EBC0" baseAuthorityName="Peters" baseAuthorityYear="1863" box="[1144,1360,1711,1735]" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="5" pageNumber="26" phylum="Chordata" rank="species" species="affinis">
|
||
<emphasis id="B9588168F362FE1CFBE4F958FAA9EBC0" box="[1144,1360,1711,1735]" italics="true" pageId="5" pageNumber="26">Pholidobolus affinis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2C26F9F362FE1CFAC3F947FCA9EBE0" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="5" pageNumber="26" phylum="Chordata" rank="species" species="hillisi">
|
||
<emphasis id="B9588168F362FE1CFAC3F947FCA9EBE0" italics="true" pageId="5" pageNumber="26">P. hillisi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2C26F9F362FE1CFCFCF927FC13EBE0" authorityName="Montanucci" authorityYear="1973" box="[864,1002,1743,1767]" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="5" pageNumber="26" phylum="Chordata" rank="species" species="macbrydei">
|
||
<emphasis id="B9588168F362FE1CFCFCF927FC13EBE0" box="[864,1002,1743,1767]" italics="true" pageId="5" pageNumber="26">P. macbrydei</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2C26F9F362FE1CFC6AF927FB94EBE0" baseAuthorityName="Peters" baseAuthorityYear="1863" box="[1014,1133,1744,1767]" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="5" pageNumber="26" phylum="Chordata" rank="species" species="montium">
|
||
<emphasis id="B9588168F362FE1CFC6AF927FB94EBE0" box="[1014,1133,1744,1767]" italics="true" pageId="5" pageNumber="26">P. montium</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2C26F9F362FE1CFBE5F927FAF6EBE0" authorityName="Montanucci" authorityYear="1973" box="[1145,1295,1743,1767]" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="5" pageNumber="26" phylum="Chordata" rank="species" species="prefrontalis">
|
||
<emphasis id="B9588168F362FE1CFBE5F927FAF6EBE0" box="[1145,1295,1743,1767]" italics="true" pageId="5" pageNumber="26">P. prefrontalis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B9588168F362FE1CFA87F927FCACEA00" italics="true" pageId="5" pageNumber="26">P. vertebralis</emphasis>
|
||
;
|
||
<bibRefCitation id="EFBD208BF362FE1CFCFFF907FC15EA00" author="Nunes PMS" box="[867,1004,1775,1799]" pageId="5" pageNumber="26" refId="ref7392" refString="Nunes PMS. 2011. Morfologia hemipeniana dos lagartos microteideos e suas implicacoes nas relacoes filo- geneticas da familia Gymnophthalmidae (Teioidea: Squamata). Ph. D. Thesis, Departamento de Zoologia, Universidade de Sao Paulo, Sao Paulo, Brazil. 137 p." type="book" year="2011">Nunes, 2011</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFBD208BF362FE1CFC6AF918FAE0EA00" author="Torres-Carvajal O & Venegas PJ & Lobos SE & Mafla-Endara P & Sales Nunes PM" box="[1014,1305,1775,1799]" pageId="5" pageNumber="26" pagination="76 - 88" refId="ref7748" refString="Torres-Carvajal O, Venegas PJ, Lobos SE, Mafla-Endara P, Sales Nunes PM. 2014. A new species of Pholidobolus (Squamata: Gymnophthalmidae) from the Andes of southern Ecuador. Amphibian & Reptile Conservation 8 (1): 76 - 88." type="journal article" year="2014">Torres-Carvajal et al. 2014</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B935D7AF362FE1FFCABF8F8FE4DEFEF" blockId="5.[787,1399,334,1927]" lastBlockId="6.[141,747,144,744]" lastPageId="6" lastPageNumber="27" pageId="5" pageNumber="26">
|
||
The hemipenis of the
|
||
<typeStatus id="5497E3D8F362FE1CFBBFF8F8FB79EA20" box="[1059,1152,1807,1831]" pageId="5" pageNumber="26" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C2C26F9F362FE1CFB3BF8E7FAFFEA20" authorityName="Venegas & Echevarría & Lobos & Sales Nunes & Abstract.-Based & The" authorityYear="2016" box="[1191,1286,1807,1831]" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="5" pageNumber="26" phylum="Chordata" rank="species" species="ulisesi">
|
||
<emphasis id="B9588168F362FE1CFB3BF8E7FAFFEA20" box="[1191,1286,1807,1831]" italics="true" pageId="5" pageNumber="26">P. ulisesi</emphasis>
|
||
</taxonomicName>
|
||
described herein (
|
||
<figureCitation id="131741FFF362FE1CFCF7F8D8FC54EA40" box="[875,941,1839,1863]" captionStart="Fig" captionStartId="6.[141,176,1912,1934]" captionTargetBox="[231,1304,1307,1897]" captionTargetId="figure-614@6.[231,1304,1307,1897]" captionTargetPageId="6" captionText="Fig. 5. Left hemipenis of Pholidobolus ulisesi sp. nov. (CORBIDI 12734 - holotype) in sulcate (left), lateral (middle), and asulcate (right) views. Photographs by PMSN." figureDoi="http://doi.org/10.5281/zenodo.11376962" httpUri="https://zenodo.org/record/11376962/files/figure.png" pageId="5" pageNumber="26">Fig. 5</figureCitation>
|
||
) is broadly congruent with the illustrated by
|
||
<bibRefCitation id="EFBD208BF362FE1CFCA7F8A7FBC8EA60" author="Doan TM & Cusi JC" box="[827,1073,1871,1895]" pageId="5" pageNumber="26" refId="ref7015" refString="Doan TM, Cusi JC. 2014. Geographic distribution of Cercosaura vertebralis O'Shaughnessy, 1879 (Reptilia: Squamata: Gymnophthalmidae) and the status of Cercosaura ampuedai (Lancini, 1968). Check List 10: 1,195 - 1,200." type="journal volume" year="2014">Doan and Cusi (2014)</bibRefCitation>
|
||
for a specimen of
|
||
<taxonomicName id="4C2C26F9F362FE1CFA96F8A7FA95EA60" authorityName="Venegas & Echevarría & Lobos & Sales Nunes & Abstract.-Based & The" authorityYear="2016" box="[1290,1388,1871,1895]" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="5" pageNumber="26" phylum="Chordata" rank="species" species="ulisesi">
|
||
<emphasis id="B9588168F362FE1CFA96F8A7FA95EA60" box="[1290,1388,1871,1895]" italics="true" pageId="5" pageNumber="26">P. ulisesi</emphasis>
|
||
</taxonomicName>
|
||
, considered by them as
|
||
<taxonomicName id="4C2C26F9F362FE1CFB8BF887FB51EA80" authorityName="O'Shaughnessy" authorityYear="1879" box="[1047,1192,1903,1927]" class="Squamata" family="Gymnophthalmidae" genus="Cercosaura" kingdom="Animalia" pageId="5" pageNumber="26" phylum="Chordata" rank="species" species="vertebralis">
|
||
<emphasis id="B9588168F362FE1CFB8BF887FB51EA80" box="[1047,1192,1903,1927]" italics="true" pageId="5" pageNumber="26">P. vertebralis</emphasis>
|
||
</taxonomicName>
|
||
(see “Discussion” hereafter). Although
|
||
<bibRefCitation id="EFBD208BF361FE1FFEE9FF66FD97EDAF" author="Doan TM & Cusi JC" box="[373,622,144,168]" pageId="6" pageNumber="27" refId="ref7015" refString="Doan TM, Cusi JC. 2014. Geographic distribution of Cercosaura vertebralis O'Shaughnessy, 1879 (Reptilia: Squamata: Gymnophthalmidae) and the status of Cercosaura ampuedai (Lancini, 1968). Check List 10: 1,195 - 1,200." type="journal volume" year="2014">Doan and Cusi (2014)</bibRefCitation>
|
||
reported a reduced count of flounces ornamenting the organ (14 versus
|
||
<quantity id="4CD4F09FF361FE1FFF41FF27FEE1EDEF" box="[221,280,208,232]" metricMagnitude="-1" metricUnit="m" metricValue="5.842" pageId="6" pageNumber="27" unit="in" value="23.0">23 in</quantity>
|
||
the
|
||
<typeStatus id="5497E3D8F361FE1FFED3FF27FE55EDEF" box="[335,428,208,232]" pageId="6" pageNumber="27" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C2C26F9F361FE1FFE46FF26FDC4EDEF" authorityName="Venegas & Echevarría & Lobos & Sales Nunes & Abstract.-Based & The" authorityYear="2016" box="[474,573,208,232]" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="6" pageNumber="27" phylum="Chordata" rank="species" species="ulisesi">
|
||
<emphasis id="B9588168F361FE1FFE46FF26FDC4EDEF" box="[474,573,208,232]" italics="true" pageId="6" pageNumber="27">P. ulisesi</emphasis>
|
||
</taxonomicName>
|
||
), their
|
||
<figureCitation id="131741FFF361FE1FFD08FF27FD10EDEF" box="[660,745,208,232]" captionStart="Fig" captionStartId="6.[141,176,1912,1934]" captionTargetBox="[231,1304,1307,1897]" captionTargetId="figure-614@6.[231,1304,1307,1897]" captionTargetPageId="6" captionText="Fig. 5. Left hemipenis of Pholidobolus ulisesi sp. nov. (CORBIDI 12734 - holotype) in sulcate (left), lateral (middle), and asulcate (right) views. Photographs by PMSN." figureDoi="http://doi.org/10.5281/zenodo.11376962" httpUri="https://zenodo.org/record/11376962/files/figure.png" pageId="6" pageNumber="27">Fig. 5B</figureCitation>
|
||
clearly shows at least 18 visible flounces ornamenting the hemipenis sides, plus other flounces not countable due the positioning of the organ and the lack of focus in some areas of the hemipenis photograph. Similar to the hemipenis of
|
||
<taxonomicName id="4C2C26F9F361FE1FFED2FE86FE57EC8F" authorityName="Venegas & Echevarría & Lobos & Sales Nunes & Abstract.-Based & The" authorityYear="2016" box="[334,430,368,392]" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="6" pageNumber="27" phylum="Chordata" rank="species" species="ulisesi">
|
||
<emphasis id="B9588168F361FE1FFED2FE86FE57EC8F" box="[334,430,368,392]" italics="true" pageId="6" pageNumber="27">P. ulisesi</emphasis>
|
||
</taxonomicName>
|
||
described by
|
||
<bibRefCitation id="EFBD208BF361FE1FFDDBFE86FF20ECAF" author="Doan TM & Cusi JC" pageId="6" pageNumber="27" refId="ref7015" refString="Doan TM, Cusi JC. 2014. Geographic distribution of Cercosaura vertebralis O'Shaughnessy, 1879 (Reptilia: Squamata: Gymnophthalmidae) and the status of Cercosaura ampuedai (Lancini, 1968). Check List 10: 1,195 - 1,200." type="journal volume" year="2014">Doan and Cusi (2014)</bibRefCitation>
|
||
, but contrasting with the hemipenis of
|
||
<emphasis id="B9588168F361FE1FFD12FE66FF32ECCF" italics="true" pageId="6" pageNumber="27">P. vertebralis</emphasis>
|
||
illustrated by
|
||
<bibRefCitation id="EFBD208BF361FE1FFEF7FE47FF04ECEF" author="Hernandez-Ruz EJ & Bernal-Gonzalez CA" pageId="6" pageNumber="27" pagination="48 - 57" refId="ref7204" refString="Hernandez-Ruz EJ, Bernal-Gonzalez CA. 2011. Variacion morfologica en Cercosaura vertebralis (Sauria: Gymnophthalmidae) en Colombia. Ingenerias & Am- azonia 4: 48 - 57." type="journal article" year="2011">Hernández-Ruz and Bernal-Gonzalez (2011)</bibRefCitation>
|
||
for a specimen from
|
||
<collectingRegion id="49E89398F361FE1FFE68FE27FDB8ECEF" box="[500,577,464,488]" country="Colombia" name="Narino" pageId="6" pageNumber="27">Nariño</collectingRegion>
|
||
,
|
||
<collectingCountry id="F33B1DEAF361FE1FFDD3FE27FD42ECEF" box="[591,699,464,488]" name="Colombia" pageId="6" pageNumber="27">Colombia</collectingCountry>
|
||
, the hemipenis of the
|
||
<typeStatus id="5497E3D8F361FE1FFED6FE07FE5EEF0F" box="[330,423,496,520]" pageId="6" pageNumber="27" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C2C26F9F361FE1FFE53FE06FDD7EF0F" authorityName="Venegas & Echevarría & Lobos & Sales Nunes & Abstract.-Based & The" authorityYear="2016" box="[463,558,496,520]" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="6" pageNumber="27" phylum="Chordata" rank="species" species="ulisesi">
|
||
<emphasis id="B9588168F361FE1FFE53FE06FDD7EF0F" box="[463,558,496,520]" italics="true" pageId="6" pageNumber="27">P. ulisesi</emphasis>
|
||
</taxonomicName>
|
||
presents the four flounces in basal position at the asulcate face separated from the other flounces ornamenting the hemipenis laterally. In the drawing presented by
|
||
<bibRefCitation id="EFBD208BF361FE1FFE68FDA7FD13EF6F" author="Hernandez-Ruz EJ" box="[500,746,592,616]" pageId="6" pageNumber="27" pagination="1 - 14" refId="ref7154" refString="Hernandez-Ruz EJ. 2005. Taxonomic and biological notes on Cercosaura ampuedai (Lancini, 1968) (Squamata: Gymnophalmidae) in the eastern slope of the Cordillera Oriental of Colombia. Publicacoes Avulsas do Instituto Pau Brasil de Historia Natural 08 - 09: 1 - 14." type="book chapter" year="2005">Hernandez-Ruz (2005)</bibRefCitation>
|
||
for
|
||
<taxonomicName id="4C2C26F9F361FE1FFF2FFD86FE59EF8F" box="[179,416,624,648]" class="Squamata" family="Gymnophthalmidae" genus="Cercosaura" kingdom="Animalia" pageId="6" pageNumber="27" phylum="Chordata" rank="species" species="ampuedai">
|
||
<emphasis id="B9588168F361FE1FFF2FFD86FE59EF8F" box="[179,416,624,648]" italics="true" pageId="6" pageNumber="27">Cercosaura ampuedai</emphasis>
|
||
</taxonomicName>
|
||
(synonym of
|
||
<taxonomicName id="4C2C26F9F361FE1FFDA8FD86FD3AEF8F" authorityName="O'Shaughnessy" authorityYear="1879" box="[564,707,624,648]" class="Squamata" family="Gymnophthalmidae" genus="Cercosaura" kingdom="Animalia" pageId="6" pageNumber="27" phylum="Chordata" rank="species" species="vertebralis">
|
||
<emphasis id="B9588168F361FE1FFDA8FD86FD3AEF8F" box="[564,707,624,648]" italics="true" pageId="6" pageNumber="27">P. vertebralis</emphasis>
|
||
</taxonomicName>
|
||
according to
|
||
<bibRefCitation id="EFBD208BF361FE1FFE98FD66FE0CEFAF" author="Doan TM & Cusi JC" box="[260,501,656,680]" pageId="6" pageNumber="27" refId="ref7015" refString="Doan TM, Cusi JC. 2014. Geographic distribution of Cercosaura vertebralis O'Shaughnessy, 1879 (Reptilia: Squamata: Gymnophthalmidae) and the status of Cercosaura ampuedai (Lancini, 1968). Check List 10: 1,195 - 1,200." type="journal volume" year="2014">Doan and Cusi [2014]</bibRefCitation>
|
||
) such flounces are not visible, probably due the distally misplaced tie made during hemipenial preparation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3360EF1F361FE1FFF11FCF9FAE1EF2F" pageId="6" pageNumber="27" type="distribution">
|
||
<paragraph id="8B935D7AF361FE1FFF11FCF9FE9AE980" blockId="6.[140,746,782,1287]" pageId="6" pageNumber="27">
|
||
<emphasis id="B9588168F361FE1FFF11FCF9FD10EE20" bold="true" box="[141,745,782,807]" pageId="6" pageNumber="27">Distribution and natural history observations:</emphasis>
|
||
<taxonomicName id="4C2C26F9F361FE1FFF11FCD8FE9EEE40" authorityName="Venegas & Echevarría & Lobos & Sales Nunes & Abstract.-Based & The" authorityYear="2016" box="[141,359,815,839]" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="6" pageNumber="27" phylum="Chordata" rank="species" species="ulisesi">
|
||
<emphasis id="B9588168F361FE1FFF11FCD8FE9EEE40" box="[141,359,815,839]" italics="true" pageId="6" pageNumber="27">Pholidobolus ulisesi</emphasis>
|
||
</taxonomicName>
|
||
is known from five localities at elevations of
|
||
<quantity id="4CD4F09FF361FE1FFF60FCB8FE65EE60" box="[252,412,847,871]" metricMagnitude="0" metricUnit="m" metricValue="2.0999999999999996" metricValueMax="2.3" metricValueMin="1.9" pageId="6" pageNumber="27" unit="m" value="2.0999999999999996" valueMax="2.3" valueMin="1.9">
|
||
<geoCoordinate id="EE183BBDF361FE1FFF60FCB8FECEEE60" box="[252,311,847,871]" degrees="1,900" direction="north" orientation="latitude" pageId="6" pageNumber="27" precision="55" value="1.9">1,900</geoCoordinate>
|
||
<geoCoordinate id="EE183BBDF361FE1FFEABFCB8FE78EE60" box="[311,385,847,871]" degrees="2,300" direction="west" orientation="longitude" pageId="6" pageNumber="27" precision="55" value="-2.3">–2,300</geoCoordinate>
|
||
m
|
||
</quantity>
|
||
in
|
||
<collectingRegion id="49E89398F361FE1FFE20FCB8FDD7EE60" box="[444,558,847,871]" country="Peru" name="Cajamarca" pageId="6" pageNumber="27">Cajamarca</collectingRegion>
|
||
and
|
||
<collectingRegion id="49E89398F361FE1FFDFCFCB8FD13EE60" box="[608,746,847,871]" country="Peru" name="Lambayeque" pageId="6" pageNumber="27">Lambayeque</collectingRegion>
|
||
departments, northern
|
||
<collectingCountry id="F33B1DEAF361FE1FFEE3FC87FE49EE80" box="[383,432,880,903]" name="Peru" pageId="6" pageNumber="27">Peru</collectingCountry>
|
||
(
|
||
<figureCitation id="131741FFF361FE1FFE5CFC98FDFBEE80" box="[448,514,879,903]" captionStart="Fig" captionStartId="7.[141,176,1387,1409]" captionTargetBox="[270,1266,164,1374]" captionTargetId="figure-16@7.[268,1268,147,1393]" captionTargetPageId="7" captionText="Fig. 6. Distribution of Pholidobolus in Ecuador and Peru (circles): P. affinis (green); P. dicrus (black); P. hillisi (purple); P. macbry- dei (blue); P. montium (gray); P. prefrontalis (brown); P. ulisesi sp. nov. (red); P. vertebralis (orange); and P.sp. (pink). Localities for P. ulisesi are: (1) Bosque de Huamantanga (type locality); (2) El Chaupe; (3) Estación Biológica Chichilapa in the Santuario Nacional Tabaconas Namballe, taken from Doan and Cusi (2014); (4) San Felipe de Jaén; (5) Cañaris; (6) Quebrada La Iraca (near La Granja village); and (7) Quebrada Checos (near La Granja village) taken from Doan and Cusi (2014)." figureDoi="http://doi.org/10.5281/zenodo.11376964" httpUri="https://zenodo.org/record/11376964/files/figure.png" pageId="6" pageNumber="27">Fig. 6</figureCitation>
|
||
). All recorded localities lie within the Huancabamba depression, a region where the relatively low altitude of the Andean mountains causes fragmentation of montane habitats, and the northern extreme of the Central Andes at Cordillera Occidental in northern
|
||
<collectingCountry id="F33B1DEAF361FE1FFEEAFBE7FE53E920" box="[374,426,1040,1063]" name="Peru" pageId="6" pageNumber="27">Peru</collectingCountry>
|
||
. According to the terrestrial ecorregions of the world by
|
||
<bibRefCitation id="EFBD208BF361FE1FFE24FBD8FD78E940" author="Olson DM & Dinerstein E & Wikramanayake ED & Burgess ND & Powell GVN & Underwood EC & D'amico JA & Itoua I & Strand HE & Morrison JC & Loucks CJ & Allnutt TF & Ricketts TH & Kura Y & Lamoreux JF & Wettengel WW & Hedao P & Kassem KR" box="[440,641,1071,1095]" pageId="6" pageNumber="27" pagination="933 - 938" refId="ref7502" refString="Olson DM, Dinerstein E, Wikramanayake ED, Burgess ND, Powell GVN, Underwood EC, D'amico JA, Itoua I, Strand HE, Morrison JC, Loucks CJ, Allnutt TF, Ricketts TH, Kura Y, Lamoreux JF, Wettengel WW, Hedao P, Kassem KR. 2001. Terrestrial ecoregion of the world: A new map of life on earth. BioScience 51: 933 - 938." type="journal article" year="2001">Olson et al. (2001)</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C2C26F9F361FE1FFD17FBC7FD10E940" authorityName="Venegas & Echevarría & Lobos & Sales Nunes & Abstract.-Based & The" authorityYear="2016" box="[651,745,1071,1095]" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="6" pageNumber="27" phylum="Chordata" rank="species" species="ulisesi">
|
||
<emphasis id="B9588168F361FE1FFD17FBC7FD10E940" box="[651,745,1071,1095]" italics="true" pageId="6" pageNumber="27">P. ulisesi</emphasis>
|
||
</taxonomicName>
|
||
occurs within Eastern Cordillera real montane forest and Marañón dry forest.
|
||
</paragraph>
|
||
<paragraph id="8B935D7AF361FE1FFF31FB78FAE1EF2F" blockId="6.[140,746,782,1287]" lastBlockId="6.[787,1399,144,552]" pageId="6" pageNumber="27">
|
||
<taxonomicName id="4C2C26F9F361FE1FFF31FB78FE7FE9A0" authorityName="Venegas & Echevarría & Lobos & Sales Nunes & Abstract.-Based & The" authorityYear="2016" box="[173,390,1167,1191]" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="6" pageNumber="27" phylum="Chordata" rank="species" species="ulisesi">
|
||
<emphasis id="B9588168F361FE1FFF31FB78FE7FE9A0" box="[173,390,1167,1191]" italics="true" pageId="6" pageNumber="27">Pholidobolus ulisesi</emphasis>
|
||
</taxonomicName>
|
||
was found during the day in sunny and cloudy conditions in secondary montane forest, in the edges of primary montane forest and recently opened areas for cattle ranching, as well as in small plantations of bean and coffee. In the open cattle-ranching areas,
|
||
<taxonomicName id="4C2C26F9F361FE1FFAC3FF66FCAEEDCF" authorityName="Venegas & Echevarría & Lobos & Sales Nunes & Abstract.-Based & The" authorityYear="2016" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="6" pageNumber="27" phylum="Chordata" rank="species" species="ulisesi">
|
||
<emphasis id="B9588168F361FE1FFAC3FF66FCAEEDCF" italics="true" pageId="6" pageNumber="27">P. ulisesi</emphasis>
|
||
</taxonomicName>
|
||
was found moving on fallen trees or hiding under trunks; in secondary montane forest, the lizards were found foraging within herbaceous vegetation and running through the patches of grass. They were especially abundant in coffee and bean plantations, where they were observed running through the herbaceous vegetation and hiding in leaf litter. Sympatric squamate reptiles collected with
|
||
<taxonomicName id="4C2C26F9F361FE1FFCEEFE66FC2BECAF" authorityName="Venegas & Echevarría & Lobos & Sales Nunes & Abstract.-Based & The" authorityYear="2016" box="[882,978,400,424]" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="6" pageNumber="27" phylum="Chordata" rank="species" species="ulisesi">
|
||
<emphasis id="B9588168F361FE1FFCEEFE66FC2BECAF" box="[882,978,400,424]" italics="true" pageId="6" pageNumber="27">P. ulisesi</emphasis>
|
||
</taxonomicName>
|
||
were
|
||
<taxonomicName id="4C2C26F9F361FE1FFB8AFE67FB0AECAF" authorityName="Roze" authorityYear="1952" box="[1046,1267,400,424]" class="Squamata" family="Colubridae" genus="Chironius" kingdom="Animalia" pageId="6" pageNumber="27" phylum="Chordata" rank="species" species="monticola">
|
||
<emphasis id="B9588168F361FE1FFB8AFE67FB0AECAF" box="[1046,1267,400,424]" italics="true" pageId="6" pageNumber="27">Chironius monticola</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2C26F9F361FE1FFAB6FE66FC89ECCF" baseAuthorityName="Boettger" baseAuthorityYear="1898" class="Squamata" family="Colubridae" genus="Dipsas" kingdom="Animalia" pageId="6" pageNumber="27" phylum="Chordata" rank="species" species="peruana">
|
||
<emphasis id="B9588168F361FE1FFAB6FE66FC89ECCF" italics="true" pageId="6" pageNumber="27">Dipsas peruana</emphasis>
|
||
</taxonomicName>
|
||
at El Chaupe and Huamantanga, and
|
||
<taxonomicName id="4C2C26F9F361FE1FFA96FE47FC7BECEF" authorityName="Roze" authorityYear="1952" class="Squamata" family="Colubridae" genus="Chironius" kingdom="Animalia" pageId="6" pageNumber="27" phylum="Chordata" rank="species" species="monticola">
|
||
<emphasis id="B9588168F361FE1FFA96FE47FC7BECEF" italics="true" pageId="6" pageNumber="27">Chironius monticola</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2C26F9F361FE1FFC09FE26FBC4ECEF" baseAuthorityName="Orejas-Miranda" baseAuthorityYear="1964" box="[917,1085,465,488]" class="Squamata" family="Leptotyphlopidae" genus="Epictia" kingdom="Animalia" pageId="6" pageNumber="27" phylum="Chordata" rank="species" species="teaguei">
|
||
<emphasis id="B9588168F361FE1FFC09FE26FBC4ECEF" box="[917,1085,465,488]" italics="true" pageId="6" pageNumber="27">Epictia teaguei</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2C26F9F361FE1FFBCDFE27FA95ECEF" baseAuthorityName="Tschudi" baseAuthorityYear="1845" box="[1105,1388,464,488]" class="Squamata" family="Colubridae" genus="Erythrolamprus" kingdom="Animalia" pageId="6" pageNumber="27" phylum="Chordata" rank="species" species="taeniurus">
|
||
<emphasis id="B9588168F361FE1FFBCDFE27FA95ECEF" box="[1105,1388,464,488]" italics="true" pageId="6" pageNumber="27">Erythrolamprus taeniurus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2C26F9F361FE1FFC8BFE06FC06EF0F" authorityName="Schmidt" authorityYear="1936" box="[791,1023,497,520]" class="Squamata" family="Elapidae" genus="Micrurus" kingdom="Animalia" pageId="6" pageNumber="27" phylum="Chordata" rank="species" species="peruvianus">
|
||
<emphasis id="B9588168F361FE1FFC8BFE06FC06EF0F" box="[791,1023,497,520]" italics="true" pageId="6" pageNumber="27">Micrurus peruvianus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2C26F9F361FE1FFB88FE06FB1AEF0F" authorityName="Venegas, Echevarria & Alvarez" authorityYear="2014" box="[1044,1251,496,520]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="6" pageNumber="27" phylum="Chordata" rank="species" species="arndti">
|
||
<emphasis id="B9588168F361FE1FFB88FE06FB1AEF0F" box="[1044,1251,496,520]" italics="true" pageId="6" pageNumber="27">Stenocercus arndti</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B9588168F361FE1FFB65FE06FC92EF2F" italics="true" pageId="6" pageNumber="27">S. huancabambae</emphasis>
|
||
, and
|
||
<taxonomicName id="4C2C26F9F361FE1FFC3AFDE6FBD0EF2F" authorityName="Cadle" authorityYear="1998" box="[934,1065,529,552]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="6" pageNumber="27" phylum="Chordata" rank="species" species="stigmosus">
|
||
<emphasis id="B9588168F361FE1FFC3AFDE6FBD0EF2F" box="[934,1065,529,552]" italics="true" pageId="6" pageNumber="27">S. stigmosus</emphasis>
|
||
</taxonomicName>
|
||
at Quebrada La Iraca.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3360EF1F361FE1FFC8BFDBEFBD8EE25" pageId="6" pageNumber="27" type="etymology">
|
||
<paragraph id="8B935D7AF361FE1FFC8BFDBEFBD8EE25" blockId="6.[790,1397,585,802]" pageId="6" pageNumber="27">
|
||
<emphasis id="B9588168F361FE1FFC8BFDBEFC51EF65" bold="true" box="[791,936,585,610]" pageId="6" pageNumber="27">Etymology:</emphasis>
|
||
The specific epithet
|
||
<taxonomicName id="4C2C26F9F361FE1FFB11FDBDFB24EF65" authorityName="Venegas & Echevarría & Lobos & Sales Nunes & Abstract.-Based & The" authorityYear="2016" box="[1165,1245,586,610]" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="6" pageNumber="27" phylum="Chordata" rank="species" species="ulisesi">
|
||
“
|
||
<emphasis id="B9588168F361FE1FFB05FDBDFB24EF65" box="[1177,1245,586,610]" italics="true" pageId="6" pageNumber="27">ulisesi</emphasis>
|
||
</taxonomicName>
|
||
” is a noun in the genitive case and a patronym for Ulises Gamonal Guevara, for his significant contribution to the archaeology of
|
||
<collectingRegion id="49E89398F361FE1FFCFDFD5DFC2AEFC5" box="[865,979,682,706]" country="Peru" name="Cajamarca" pageId="6" pageNumber="27">Cajamarca</collectingRegion>
|
||
in northwestern
|
||
<collectingCountry id="F33B1DEAF361FE1FFB18FD5DFB41EFC6" box="[1156,1208,682,705]" name="Peru" pageId="6" pageNumber="27">Peru</collectingCountry>
|
||
. One of his major contributions is the discovery of the>6,000-year-old Faical cave paintings in San Ignacio, declared as Cultural Patrimony of the Nation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3360EF1F361FE1FFC8BFCBFFC2BE9E6" pageId="6" pageNumber="27" type="discussion">
|
||
<paragraph id="8B935D7AF361FE1FFC8BFCBFFC2BE9E6" blockId="6.[790,1396,840,1249]" pageId="6" pageNumber="27">
|
||
<emphasis id="B9588168F361FE1FFC8BFCBFFC69EE66" bold="true" box="[791,912,840,865]" pageId="6" pageNumber="27">Remarks:</emphasis>
|
||
In a molecular phylogeny of
|
||
<taxonomicName id="4C2C26F9F361FE1FFB55FCBEFABFEE67" authorityName="Wagler" authorityYear="1830" box="[1225,1350,841,864]" class="Squamata" family="Gymnophthalmidae" genus="Cercosaura" kingdom="Animalia" pageId="6" pageNumber="27" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9588168F361FE1FFB55FCBEFABFEE67" box="[1225,1350,841,864]" italics="true" pageId="6" pageNumber="27">Cercosaura</emphasis>
|
||
</taxonomicName>
|
||
and related taxa,
|
||
<bibRefCitation id="EFBD208BF361FE1FFC3FFC9EFB20EE86" author="Torres-Carvajal O & Lobos SE & Venegas PJ" box="[931,1241,873,897]" pageId="6" pageNumber="27" pagination="281 - 288" refId="ref7796" refString="Torres-Carvajal O, Lobos SE, Venegas PJ. 2015. Phylogeny of Neotropical Cercosaura (Squamata: Gymnophthalmidae) lizards. Molecular Phylogenetics and Evolution 93: 281 - 288." type="journal article" year="2015">Torres-Carvajal et al. (2015)</bibRefCitation>
|
||
showed, with high support, that
|
||
<taxonomicName id="4C2C26F9F361FE1FFC72FC7EFB37EEA6" authorityName="Venegas & Echevarría & Lobos & Sales Nunes & Abstract.-Based & The" authorityYear="2016" box="[1006,1230,905,929]" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="6" pageNumber="27" phylum="Chordata" rank="species" species="ulisesi">
|
||
<emphasis id="B9588168F361FE1FFC72FC7EFB37EEA6" box="[1006,1230,905,929]" italics="true" pageId="6" pageNumber="27">Pholidobolus ulisesi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C2C26F9F361FE1FFB78FC7EFCCCEEC7" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="6" pageNumber="27" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="B9588168F361FE1FFB78FC7EFA8BEEA6" box="[1252,1394,905,929]" italics="true" pageId="6" pageNumber="27">Pholidobolus</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
in their paper) and
|
||
<taxonomicName id="4C2C26F9F361FE1FFB87FC5DFB8EEEC6" box="[1051,1143,937,961]" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="6" pageNumber="27" phylum="Chordata" rank="species" species="hillisi">
|
||
<emphasis id="B9588168F361FE1FFB87FC5DFB8EEEC6" box="[1051,1143,937,961]" italics="true" pageId="6" pageNumber="27">P. hillisi</emphasis>
|
||
</taxonomicName>
|
||
are sister species. Together they form a clade sister to all other species of
|
||
<taxonomicName id="4C2C26F9F361FE1FFC8BFC1EFC5CE906" box="[791,933,1001,1025]" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="6" pageNumber="27" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9588168F361FE1FFC8BFC1EFC5CE906" box="[791,933,1001,1025]" italics="true" pageId="6" pageNumber="27">Pholidobolus</emphasis>
|
||
</taxonomicName>
|
||
. In addition, these authors found that both “
|
||
<taxonomicName id="4C2C26F9F361FE1FFCBFFBFEFBD1E926" authorityName="O'Shaughnessy" authorityYear="1879" box="[803,1064,1033,1057]" class="Squamata" family="Gymnophthalmidae" genus="Cercosaura" kingdom="Animalia" pageId="6" pageNumber="27" phylum="Chordata" rank="species" species="vertebralis">
|
||
<emphasis id="B9588168F361FE1FFCBFFBFEFC59E927" box="[803,928,1033,1056]" italics="true" pageId="6" pageNumber="27">Cercosaura</emphasis>
|
||
”
|
||
<emphasis id="B9588168F361FE1FFC28FBFEFBD1E926" box="[948,1064,1033,1057]" italics="true" pageId="6" pageNumber="27">vertebralis</emphasis>
|
||
</taxonomicName>
|
||
and “
|
||
<taxonomicName id="4C2C26F9F361FE1FFBF0FBFEFB10E927" authorityName="Wagler" authorityYear="1830" box="[1132,1257,1033,1056]" class="Squamata" family="Gymnophthalmidae" genus="Cercosaura" kingdom="Animalia" pageId="6" pageNumber="27" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9588168F361FE1FFBF0FBFEFB10E927" box="[1132,1257,1033,1056]" italics="true" pageId="6" pageNumber="27">Cercosaura</emphasis>
|
||
</taxonomicName>
|
||
”
|
||
<emphasis id="B9588168F361FE1FFB62FBFEFACFE926" box="[1278,1334,1033,1057]" italics="true" pageId="6" pageNumber="27">dicra</emphasis>
|
||
were nested within
|
||
<taxonomicName id="4C2C26F9F361FE1FFC2CFBDEFBC4E946" box="[944,1085,1065,1089]" class="Squamata" family="Gymnophthalmidae" genus="Pholidobolus" kingdom="Animalia" pageId="6" pageNumber="27" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9588168F361FE1FFC2CFBDEFBC4E946" box="[944,1085,1065,1089]" italics="true" pageId="6" pageNumber="27">Pholidobolus</emphasis>
|
||
</taxonomicName>
|
||
, and were therefore referred to this genus (
|
||
<bibRefCitation id="EFBD208BF361FE1FFC2FFBBEFB24E966" author="Torres-Carvajal O & Lobos SE & Venegas PJ" box="[947,1245,1097,1121]" pageId="6" pageNumber="27" pagination="281 - 288" refId="ref7796" refString="Torres-Carvajal O, Lobos SE, Venegas PJ. 2015. Phylogeny of Neotropical Cercosaura (Squamata: Gymnophthalmidae) lizards. Molecular Phylogenetics and Evolution 93: 281 - 288." type="journal article" year="2015">Torres-Carvajal et al. 2015</bibRefCitation>
|
||
). An identical topology can be observed in a recent molecular phylogeny of the clade Cercosaurinae by
|
||
<bibRefCitation id="EFBD208BF361FE1FFB13FB7EFC9AE9C6" author="Torres-Carvajal O & Lobos SE & Venegas PJ & Chavez G & Aguirre-Penafiel V & Zurita D & Echevarria LY" pageId="6" pageNumber="27" pagination="63 - 75" refId="ref7828" refString="Torres-Carvajal O, Lobos SE, Venegas PJ, Chavez G, Aguirre-Penafiel V, Zurita D, Echevarria LY. 2016. Phylogeny and biogeography of the most diverse clade of South American gymnophthalmid lizards (Squamata, Gymnophthalmidae, Cercosaurinae). Molecular Phylogenetics and Evolution 99: 63 - 75." type="journal article" year="2016">Torres-Carvajal et al. (2016)</bibRefCitation>
|
||
. Therefore, we adopt this taxonomic change in the discussion below.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |