treatments-xml/data/03/81/BB/0381BB43B7541672FF72D10D6039A477.xml
2024-06-21 12:22:17 +02:00

1433 lines
174 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="D9805FA503CBB8A537B51B2099757532" ID-CLB-Dataset="29430" ID-DOI="10.11646/zootaxa.4482.1.5" ID-GBIF-Dataset="f998a746-73a0-42cf-8d27-c747bacd94ce" ID-ISSN="1175-5326" ID-Zenodo-Dep="1440508" ID-ZooBank="94310D7F-A204-40D9-A6F2-601987F90D27" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538373868170" checkinUser="plazi" docAuthor="Dole-Olivier, Marie-Jose, Hafid, Hinda &amp; Piscart, Christophe" docDate="2018" docId="0381BB43B7541672FF72D10D6039A477" docLanguage="en" docName="zootaxa.4482.1.5.pdf" docOrigin="Zootaxa 4482 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudoniphargus djemoi Dole-Olivier &amp; Hafid &amp; Piscart 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="137" masterDocId="FFB8C33BB755167EFFE5D4036130A360" masterDocTitle="A new groundwater species of Pseudoniphargus (Amphipoda: Pseudoniphargidae) fromAlgeria" masterLastPageNumber="139" masterPageNumber="125" pageNumber="126" updateTime="1698559220868" updateUser="ExternalLinkService">
<mods:mods id="9BA55EA0A8DCC86D541C0354F4E5C019" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D8F67091D32BDA36782613CC666D425C">
<mods:title id="1D694603250A0A4097FF463BC860B776">A new groundwater species of Pseudoniphargus (Amphipoda: Pseudoniphargidae) fromAlgeria</mods:title>
</mods:titleInfo>
<mods:name id="57DCBCD20A2B4A23A155B36E1BCE46DE" type="personal">
<mods:role id="AED5F4885E1F9DE81A1C212E77EE5340">
<mods:roleTerm id="5E5866C2BBB82230F01EE6BBAA639428">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9BFF8A73EF880228DF1CFB072C3519D9">Dole-Olivier, Marie-Jose</mods:namePart>
</mods:name>
<mods:name id="39C0C33954FCE83A027E9DD5F09A437C" type="personal">
<mods:role id="A303A0109AC2209EFEF033535BE0DFDF">
<mods:roleTerm id="DB00B71B8883995AAF6697E1621EA973">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5682BE54E522274368576815667A0E39">Hafid, Hinda</mods:namePart>
</mods:name>
<mods:name id="E67DDB4F89F3FC09B33E7735F531BCD0" type="personal">
<mods:role id="6C23866B04BB0574DB9EF4ADDB9BBFE3">
<mods:roleTerm id="BA65C81AE7550245C6218F3C72EC098A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DBFD091D0952757751913AE72B6B3080">Piscart, Christophe</mods:namePart>
</mods:name>
<mods:typeOfResource id="F07C28D78CF744BEF0C11CEB874BC925">text</mods:typeOfResource>
<mods:relatedItem id="F5F80F1E6E68BB8D43A18729BD7C34B6" type="host">
<mods:titleInfo id="2050791DC0E1E0053512DAE72E0BF947">
<mods:title id="D2225343835783828A42A0C7AAF0B542">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="E6ED8C4122D93CC1DF96224E59608998">
<mods:date id="C0FBEF98568685C2C673577A98FC65DA">2018</mods:date>
<mods:detail id="79014041FE028BE61668091BEF7FF9E1" type="pubDate">
<mods:number id="A0AD7E5C10ADDCF28B879F0170B830CE">2018-09-17</mods:number>
</mods:detail>
<mods:detail id="4470FD0F790CFFEE7AF92F397BFA0170" type="volume">
<mods:number id="3C46AFDFED2F350EBE1609DBF1481071">4482</mods:number>
</mods:detail>
<mods:detail id="6660E14471F50C1BF3812B3131EF18C3" type="issue">
<mods:number id="19D356433C883D0CDD2AB42F59544F40">1</mods:number>
</mods:detail>
<mods:extent id="FABBBD36126432BEF121C7851C13FE6F" unit="page">
<mods:start id="5C8128291DCF04022A9FEFD1A4DA67D5">125</mods:start>
<mods:end id="8A203D6C69932B75C7BD81A597819059">139</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="9FA7BB35DC4E2ACBE7AF42721C4EFD95">journal article</mods:classification>
<mods:identifier id="1B4AE58098CD6A81EC86FB9176709222" type="CLB-Dataset">29430</mods:identifier>
<mods:identifier id="EFC877831E2513381AC186ED3214A72A" type="DOI">10.11646/zootaxa.4482.1.5</mods:identifier>
<mods:identifier id="ACEEBC605F87520F303F511767AE39E5" type="GBIF-Dataset">f998a746-73a0-42cf-8d27-c747bacd94ce</mods:identifier>
<mods:identifier id="468884F7511075C5935A42BBEB4883BB" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="1D06CAF01404968D4A2D81BE7FF99C33" type="Zenodo-Dep">1440508</mods:identifier>
<mods:identifier id="2E97F372023406C52155808D50DDD3ED" type="ZooBank">94310D7F-A204-40D9-A6F2-601987F90D27</mods:identifier>
</mods:mods>
<treatment id="0381BB43B7541672FF72D10D6039A477" ID-DOI="http://doi.org/10.5281/zenodo.5990681" ID-GBIF-Taxon="148421519" ID-Zenodo-Dep="5990681" LSID="urn:lsid:plazi:treatment:0381BB43B7541672FF72D10D6039A477" httpUri="http://treatment.plazi.org/id/0381BB43B7541672FF72D10D6039A477" lastPageId="12" lastPageNumber="137" pageId="1" pageNumber="126">
<subSubSection id="C33259DEB754167FFF72D10D603AA62A" pageId="1" pageNumber="126" type="nomenclature">
<paragraph id="8B970A55B754167FFF72D10D6318A648" blockId="1.[151,552,1294,1354]" box="[151,552,1294,1320]" pageId="1" pageNumber="126">
<heading id="D0DFBD39B754167FFF72D10D6318A648" bold="true" box="[151,552,1294,1320]" fontSize="11" level="1" pageId="1" pageNumber="126" reason="1">
<emphasis id="B95CD647B754167FFF72D10D6318A648" bold="true" box="[151,552,1294,1320]" pageId="1" pageNumber="126">
<taxonomicName id="4C2871D6B754167FFF72D10D60F3A648" ID-CoL="7ZL2S" authority="Dole-Olivier &amp; Hafid &amp; Piscart, 2018" authorityName="Dole-Olivier &amp; Hafid &amp; Piscart" authorityYear="2018" box="[151,451,1294,1320]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="126" phylum="Arthropoda" rank="species" species="djemoi" status="sp. nov.">
<emphasis id="B95CD647B754167FFF72D10D60F3A648" bold="true" box="[151,451,1294,1320]" italics="true" pageId="1" pageNumber="126">Pseudoniphargus djemoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F6B3CB754167FFE2ED10D6318A648" box="[459,552,1294,1320]" pageId="1" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B970A55B754167FFF72D132603AA62A" blockId="1.[151,552,1294,1354]" box="[151,266,1329,1354]" pageId="1" pageNumber="126">
(
<figureCitation id="131316D0B754167FFF7AD1326031A62A" box="[159,257,1329,1354]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="2.[151,250,1478,1500]" captionStartId-1="8.[151,250,1929,1951]" captionStartId-2="9.[151,250,1919,1941]" captionStartId-3="10.[151,250,1937,1959]" captionStartId-4="11.[151,250,1989,2011]" captionTargetBox-0="[151,1385,822,1412]" captionTargetBox-1="[209,1374,217,1906]" captionTargetBox-2="[163,1357,199,1899]" captionTargetBox-3="[191,1353,205,1908]" captionTargetBox-4="[192,1382,195,1931]" captionTargetId-0="figure@2.[151,1436,777,1454]" captionTargetId-1="figure@8.[178,1402,193,1916]" captionTargetId-2="figure@9.[162,1427,193,1904]" captionTargetId-3="figure@10.[172,1402,193,1919]" captionTargetId-4="figure@11.[152,1435,193,1968]" captionTargetPageId-0="2" captionTargetPageId-1="8" captionTargetPageId-2="9" captionTargetPageId-3="10" captionTargetPageId-4="11" captionText-0="FIGURE 1. Pseudoniphargus djemoi sp. n. Habitus of female 3 (6 mm)." captionText-1="FIGURE 2. Pseudoniphargus djemoi sp. n. A, pereopod 4 of female 1 (6,5 mm); B, pereopod 7 of male 1 (6 mm) with detail of basis (B1), propodus and dactylus (B2); C, pereopod 5 of male 1 with detail of propodus and dactylus (C1); D, pereopod 6 of male 1 with detail of basis (D1), propodus and dactylus (D2); E, Antenna 2 of male 1." captionText-2="FIGURE 3. Pseudoniphargus djemoi sp. n. A, uropod 3 of female 1 (6,5 mm); B, uropod 3 of male 2 (6,5 mm); C, telson of female 1 (6,5 mm); D, telson of male 1 (6 mm); E, uropods 1 and 2 of female 1 (6,5 mm); F,antenna 1 of female 2 (6 mm) with detail of accessory flagellum (F1), and apex of flagellum (F2); G, antenna 1 of male 1 (6 mm) with detail of apex of flagellum (G1) and accessory flagellum (G2); H, uropods 1 and 2 of male 1 (6 mm)." captionText-3="FIGURE 4. Pseudoniphargus djemoi sp. n. A, maxillipeds of male 2 (5,5 mm), with detail of distal part of inner plates (A1); B, left mandible of female 1 (6,5 mm); C, right mandible of female 1 (6,5 mm); DF, epimera 1 to 3; G, labrum of male 2 (5,5 mm); H, labium of male 2 (5,5 mm); I, maxilla 1 of female 1 (6,5 mm)." captionText-4="FIGURE 5. Pseudoniphargus djemoi sp. n., male 1 (6 mm). A, gnathopod 1; B, pereopod 4, C, gnathopod 2." httpUri-0="https://zenodo.org/record/1440510/files/figure.png" httpUri-1="https://zenodo.org/record/1440512/files/figure.png" httpUri-2="https://zenodo.org/record/1440514/files/figure.png" httpUri-3="https://zenodo.org/record/1440516/files/figure.png" httpUri-4="https://zenodo.org/record/1440518/files/figure.png" pageId="1" pageNumber="126">Figs 15</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33259DEB754167CFF72D17A6428A39E" lastPageId="2" lastPageNumber="127" pageId="1" pageNumber="126" type="materials_examined">
<paragraph id="8B970A55B754167FFF72D17A603AA5D2" blockId="1.[151,1437,1401,2002]" pageId="1" pageNumber="126">
<materialsCitation id="3B400008B754167FFF72D17A6092A6BA" ID-GBIF-Occurrence="1913889571" location="The" pageId="1" pageNumber="126" specimenCount="75" specimenCount-female="16" specimenCount-juvenile="17" specimenCount-male="5">
<emphasis id="B95CD647B754167FFF72D17A60B4A6F2" bold="true" box="[151,388,1401,1426]" pageId="1" pageNumber="126">Material examined.</emphasis>
A set of
<specimenCount id="9D2EC1DCB754167FFE1CD17963A5A6F2" box="[505,661,1401,1426]" pageId="1" pageNumber="126" type="generic">37 specimens</specimenCount>
(
<specimenCount id="9D2EC1DCB754167FFD4CD17A63FEA6F2" box="[681,718,1401,1426]" pageId="1" pageNumber="126" type="male">5♂</specimenCount>
,
<specimenCount id="9D2EC1DCB754167FFD38D17A6220A6F2" box="[733,784,1401,1426]" pageId="1" pageNumber="126" type="female">15♀</specimenCount>
,
<specimenCount id="9D2EC1DCB754167FFCC5D179629CA6F2" box="[800,940,1401,1426]" pageId="1" pageNumber="126" type="juvenile">17 juveniles</specimenCount>
&amp; unidentifiable specimens) was used for dissection (maximum size
<quantity id="4CD0A7B0B754167FFE39D19E630CA6D5" box="[476,572,1437,1462]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="1" pageNumber="126" unit="mm" value="6.5">6.5 mm</quantity>
,
<specimenCount id="9D2EC1DCB754167FFDB5D19D6355A6D6" box="[592,613,1438,1462]" pageId="1" pageNumber="126" type="female"></specimenCount>
1).
<location id="8EF75C8EB754167FFD76D19E638FA6D6" LSID="urn:lsid:plazi:treatment:0381BB43B7541672FF72D10D6039A477:8EF75C8EB754167FFD76D19E638FA6D6" box="[659,703,1437,1462]" name="The" pageId="1" pageNumber="126">The</location>
five largest and less damaged specimens were selected for descriptions, as follows
</materialsCitation>
. Syntypes: 2 ♂♂ (♂1:
<quantity id="4CD0A7B0B754167FFD47D1C263D3A6B9" box="[674,739,1473,1498]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="126" unit="mm" value="6.0">6 mm</quantity>
(MNHN-IU-2017-327), ♂2:
<quantity id="4CD0A7B0B754167FFBC9D1C265B2A6B9" box="[1068,1154,1473,1498]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="1" pageNumber="126" unit="mm" value="5.5">5.5 mm</quantity>
(MNHN-IU-2017-328)), and 3 ♀♀♀ (♀1:
<quantity id="4CD0A7B0B754167FFEBED1E66081A69D" box="[347,433,1509,1533]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="1" pageNumber="126" unit="mm" value="6.5">6.5 mm</quantity>
(MNHN-IU-2017-329); ♀2:
<quantity id="4CD0A7B0B754167FFD1FD1E6620BA69D" box="[762,827,1509,1533]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="126" unit="mm" value="6.0">6 mm</quantity>
(MNHN-IU-2017-330);
<materialsCitation id="3B400008B754167FFBB7D1E6603AA5D2" ID-GBIF-Occurrence="1913889570" collectionCode="MNHN, UCBLZ" county="Each" location="Paris." municipality="Museum National d'Histoire Naturelle" pageId="1" pageNumber="126" specimenCount="1" specimenCount-female="1" stateProvince="CEntre">
<specimenCount id="9D2EC1DCB754167FFBB7D1E66556A69D" box="[1106,1126,1509,1533]" pageId="1" pageNumber="126" type="female"></specimenCount>
3:
<quantity id="4CD0A7B0B754167FFB60D1E665F6A69D" box="[1157,1222,1509,1533]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="126" unit="mm" value="6.0">6 mm</quantity>
(MNHN-IU-2017- 331) with non-setose oostegites.
<collectingCounty id="62F672D9B754167FFDF5D20A6378A542" box="[528,584,1545,1570]" pageId="1" pageNumber="126">Each</collectingCounty>
specimen was completely dissected on a single slide and deposited in the collections of the
<collectingMunicipality id="6BF3902FB754167FFEB8D22D623CA526" box="[349,780,1581,1606]" pageId="1" pageNumber="126">Museum National dHistoire Naturelle</collectingMunicipality>
(
<collectionCode id="ED399290B754167FFCFFD22D6243A526" box="[794,883,1582,1606]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="126">MNHN</collectionCode>
) of
<location id="8EF75C8EB754167FFC7AD22E62EFA526" LSID="urn:lsid:plazi:treatment:0381BB43B7541672FF72D10D6039A477:8EF75C8EB754167FFC7AD22E62EFA526" box="[927,991,1581,1606]" county="Each" municipality="Museum National d'Histoire Naturelle" name="Paris." pageId="1" pageNumber="126" stateProvince="CEntre">Paris.</location>
Additional specimens (N= 60) from the same well were deposited in the collections of the
<location id="8EF75C8EB754167FFD35D2526299A50A" LSID="urn:lsid:plazi:treatment:0381BB43B7541672FF72D10D6039A477:8EF75C8EB754167FFD35D2526299A50A" box="[720,937,1617,1642]" county="Each" municipality="Museum National d'Histoire Naturelle" name="University of Lyon" pageId="1" pageNumber="126" stateProvince="CEntre">University of Lyon</location>
,
<collectingRegion id="49ECC4B7B754167FFC51D2516533A50A" box="[948,1027,1618,1642]" country="France" name="Centre" pageId="1" pageNumber="126">CEntre</collectingRegion>
de REssources pour les
<location id="8EF75C8EB754167FFAFDD2526027A5EE" LSID="urn:lsid:plazi:treatment:0381BB43B7541672FF72D10D6039A477:8EF75C8EB754167FFAFDD2526027A5EE" county="Each" municipality="Museum National d'Histoire Naturelle" name="Sciences de l'Evolution" pageId="1" pageNumber="126" stateProvince="CEntre">Sciences de lEvolution</location>
(CERESE, n°
<collectionCode id="ED399290B754167FFE2ED2756317A5EE" box="[459,551,1654,1678]" pageId="1" pageNumber="126">UCBLZ</collectionCode>
2017.4.1;
<collectionCode id="ED399290B754167FFD4CD2756235A5EE" box="[681,773,1654,1678]" pageId="1" pageNumber="126">UCBLZ</collectionCode>
2017.4.2;
<collectionCode id="ED399290B754167FFC63D27562D2A5EE" box="[902,994,1654,1678]" pageId="1" pageNumber="126">UCBLZ</collectionCode>
2017.4.3;
<collectionCode id="ED399290B754167FFB86D275658FA5EE" box="[1123,1215,1654,1678]" pageId="1" pageNumber="126">UCBLZ</collectionCode>
2017.4.4;
<collectionCode id="ED399290B754167FFAA5D27564ACA5EE" box="[1344,1436,1654,1678]" pageId="1" pageNumber="126">UCBLZ</collectionCode>
2017.4.5).
</materialsCitation>
</paragraph>
<paragraph id="8B970A55B754167CFF22D2BE6428A39E" blockId="1.[151,1437,1401,2002]" lastBlockId="2.[151,1437,151,722]" lastPageId="2" lastPageNumber="127" pageId="1" pageNumber="126">
<emphasis id="B95CD647B754167FFF22D2BE63AFA5B6" bold="true" box="[199,671,1725,1750]" pageId="1" pageNumber="126">Type locality and ecological conditions.</emphasis>
The sampled wells were situated in the Tamlouka plain (36°46”N 7°01'46&quot;E), northwest of Oum El Bouaghi, Algeria. This plain essentially consists of quaternary fluvial alluvial filling. The aquifer is oriented southwest-northeast with a thickness varying between
<quantity id="4CD0A7B0B754167FFBA9D3056591A47D" box="[1100,1185,1798,1822]" metricMagnitude="1" metricUnit="m" metricValue="1.05" metricValueMax="1.2" metricValueMin="0.9" pageId="1" pageNumber="126" unit="m" value="10.5" valueMax="12.0" valueMin="9.0">912 m</quantity>
(upstream) and
<quantity id="4CD0A7B0B754167FFAB3D30664ACA47D" box="[1366,1436,1797,1822]" metricMagnitude="0" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="1" pageNumber="126" unit="m" value="6.5" valueMax="8.0" valueMin="5.0">58 m</quantity>
(downstream). Eleven wells out of 16 prospected harbored the new species (
<bibRefCitation id="EFB977A4B754167FFC0FD32965E3A422" box="[1002,1235,1833,1858]" pageId="1" pageNumber="126" refId="ref8620" refString="Merzoug, D., Khiari, A., Ait Boughrous, A. &amp; Boutin, C. (2010) Faune aquatique et qualite de l'eau des puits et sources de la region d'Oum-El-Bouaghi (nord-est Algerien). Hydroecologie Appliquee, 17, 77 - 97. https: // doi. org / 10.1051 / hydro / 2010001" type="journal article">
Merzoug
<emphasis id="B95CD647B754167FFBB2D32965BFA421" box="[1111,1167,1833,1857]" italics="true" pageId="1" pageNumber="126">et al.</emphasis>
2010
</bibRefCitation>
). When present in a well, the species was rather abundant (total abundance 736 specimens, varying from 13 to 305 individuals per well) compared to another co-occurring species belonging to the same genus (
<taxonomicName id="4C2871D6B754167FFBE5D37165B6A4E9" authorityName="Chevreux" authorityYear="1901" box="[1024,1158,1905,1929]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="126" phylum="Arthropoda" rank="species" species="africanus">
<emphasis id="B95CD647B754167FFBE5D37165B6A4E9" box="[1024,1158,1905,1929]" italics="true" pageId="1" pageNumber="126">P. africanus</emphasis>
</taxonomicName>
), which was collected in two wells only (total abundance: 24 specimens). Amphipod diversity of the sampling area included also a new, not yet described genus observed in five wells (96 specimens;
<bibRefCitation id="EFB977A4B754167FFCCFD3B96522A4B2" box="[810,1042,1977,2002]" pageId="1" pageNumber="126" refId="ref8620" refString="Merzoug, D., Khiari, A., Ait Boughrous, A. &amp; Boutin, C. (2010) Faune aquatique et qualite de l'eau des puits et sources de la region d'Oum-El-Bouaghi (nord-est Algerien). Hydroecologie Appliquee, 17, 77 - 97. https: // doi. org / 10.1051 / hydro / 2010001" type="journal article">
Merzoug
<emphasis id="B95CD647B754167FFC73D3B962FEA4B1" box="[918,974,1977,2001]" italics="true" pageId="1" pageNumber="126">et al.</emphasis>
2010
</bibRefCitation>
). Physicochemical conditions were characterized by very high water mineralization (Electric Conductivity ranging from 1344 µS.cm -1 to 2211 µS.cm - 1), temperatures varying from 19°C to 21,7°C and oxygen content from 6,8mg.L -1 to 8,3 mg.L -1. Detailed analyses also demonstrated that most of the wells were polluted by organic contaminants (
<bibRefCitation id="EFB977A4B757167CFBC7D4E5643CA39E" box="[1058,1292,229,254]" pageId="2" pageNumber="127" refId="ref8620" refString="Merzoug, D., Khiari, A., Ait Boughrous, A. &amp; Boutin, C. (2010) Faune aquatique et qualite de l'eau des puits et sources de la region d'Oum-El-Bouaghi (nord-est Algerien). Hydroecologie Appliquee, 17, 77 - 97. https: // doi. org / 10.1051 / hydro / 2010001" type="journal article">
Merzoug
<emphasis id="B95CD647B757167CFB75D4E565F8A39D" box="[1168,1224,229,253]" italics="true" pageId="2" pageNumber="127">et al.</emphasis>
2010
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33259DEB757167CFF22D50A6023A20A" pageId="2" pageNumber="127" type="etymology">
<paragraph id="8B970A55B757167CFF22D50A6023A20A" blockId="2.[151,1437,151,722]" pageId="2" pageNumber="127">
<emphasis id="B95CD647B757167CFF22D50A607EA242" bold="true" box="[199,334,265,290]" pageId="2" pageNumber="127">Etymology.</emphasis>
The specific name
<taxonomicName id="4C2871D6B757167CFDDCD50A63B5A241" authorityName="Dole-Olivier &amp; Hafid &amp; Piscart" authorityYear="2018" box="[569,645,265,289]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="127" phylum="Arthropoda" rank="species" species="djemoi">
<emphasis id="B95CD647B757167CFDDCD50A63B5A241" box="[569,645,265,289]" italics="true" pageId="2" pageNumber="127">djemoi</emphasis>
</taxonomicName>
refers to Prof. Djemoi Merzoug (University Larbi Ben H'idi), who worked on the groundwater invertebrate fauna of the area and discovered the species. It can be treated as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C33259DEB757167CFF22D57665C9A1EA" pageId="2" pageNumber="127" type="diagnosis">
<paragraph id="8B970A55B757167CFF22D57665C9A1EA" blockId="2.[151,1437,151,722]" pageId="2" pageNumber="127">
<emphasis id="B95CD647B757167CFF22D5766071A2EE" bold="true" box="[199,321,373,398]" pageId="2" pageNumber="127">Diagnosis.</emphasis>
Antenna 1 short, with accessory flagellum shorter than first article of main flagellum. Antenna 2 flagellum 6- or 7-articulate, shorter than peduncle articles 4+5 combined. Male gnathopod 1 carpus about as long as corresponding propodus. Coxal plate 4 with posterior excavation. Basis of pereopods 67 without posterodistal lobe. Pereopod 7 propodus not elongated. Uropod 1 without basofacial spine(s). Uropod 3 elongate in both sexes, sexually-dimorphic. Male uropod 3 with peduncle slightly elongate, about 2.54 times as long as broad; exopod strongly elongate and upcurved, about 14.5 times as long as broad. Female uropod 3 peduncle subrectangular, 1.7 times as long as broad; exopod moderately elongate, about 11.1 times as long as broad. Telson short and wide with 2 (
<specimenCount id="9D2EC1DCB757167CFF50D67161FBA1EA" box="[181,203,626,650]" pageId="2" pageNumber="127" type="male"></specimenCount>
) and 3 (
<specimenCount id="9D2EC1DCB757167CFECCD671600BA1EA" box="[297,315,626,650]" pageId="2" pageNumber="127" type="female"></specimenCount>
) spines at each distal corner, and with poorly marked and shallow mediodistal notch.
</paragraph>
</subSubSection>
<subSubSection id="C33259DEB757167AFF22D69662D6A18C" lastPageId="4" lastPageNumber="129" pageId="2" pageNumber="127" type="description">
<paragraph id="8B970A55B757167CFF22D69665CCA1B2" blockId="2.[151,1437,151,722]" pageId="2" pageNumber="127">
<emphasis id="B95CD647B757167CFF22D6966069A1CE" bold="true" box="[199,345,661,686]" pageId="2" pageNumber="127">Description.</emphasis>
Body aspect as in
<figureCitation id="131316D0B757167CFDD1D6966341A1CE" box="[564,625,661,686]" captionStart="FIGURE 1" captionStartId="2.[151,250,1478,1500]" captionTargetBox="[151,1385,822,1412]" captionTargetId="figure@2.[151,1436,777,1454]" captionTargetPageId="2" captionText="FIGURE 1. Pseudoniphargus djemoi sp. n. Habitus of female 3 (6 mm)." httpUri="https://zenodo.org/record/1440510/files/figure.png" pageId="2" pageNumber="127">Fig.1</figureCitation>
; colorless, eyeless. Head lacking rostrum, with broadly rounded lateral plates, anteroventral margin weakly recessed. Pereopodal coxal plates 13 slightly longer than wide.
</paragraph>
<caption id="DF575ADDB757167CFF72D1C5624BA6BC" box="[151,891,1478,1500]" httpUri="https://zenodo.org/record/1440510/files/figure.png" pageId="2" pageNumber="127" startId="2.[151,250,1478,1500]" targetBox="[151,1385,822,1412]" targetPageId="2">
<paragraph id="8B970A55B757167CFF72D1C5624BA6BC" blockId="2.[151,891,1478,1500]" box="[151,891,1478,1500]" pageId="2" pageNumber="127">
<emphasis id="B95CD647B757167CFF72D1C56023A6BC" bold="true" box="[151,275,1478,1500]" pageId="2" pageNumber="127">FIGURE 1.</emphasis>
<taxonomicName id="4C2871D6B757167CFEFCD1C56324A6BC" authority="Dole-Olivier &amp; Hafid &amp; Piscart, 2018" authorityName="Dole-Olivier &amp; Hafid &amp; Piscart" authorityYear="2018" box="[281,532,1478,1500]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="127" phylum="Arthropoda" rank="species" species="djemoi" status="sp. nov.">
<emphasis id="B95CD647B757167CFEFCD1C56324A6BC" box="[281,532,1478,1500]" italics="true" pageId="2" pageNumber="127">Pseudoniphargus djemoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F6B3CB757167CFDFED1C46364A6BC" box="[539,596,1479,1500]" pageId="2" pageNumber="127" rank="species">
<emphasis id="B95CD647B757167CFDFED1C46364A6BC" bold="true" box="[539,596,1479,1500]" pageId="2" pageNumber="127">sp. n.</emphasis>
</taxonomicNameLabel>
Habitus of female 3 (6 mm).
</paragraph>
</caption>
<paragraph id="8B970A55B757167CFF22D20965F2A5D2" blockId="2.[151,1437,1545,2038]" pageId="2" pageNumber="127">
Antenna 1 (
<figureCitation id="131316D0B757167CFEA8D20A60F5A542" box="[333,453,1545,1570]" captionStart="FIGURE 3" captionStartId="9.[151,250,1919,1941]" captionTargetBox="[163,1357,199,1899]" captionTargetId="figure@9.[162,1427,193,1904]" captionTargetPageId="9" captionText="FIGURE 3. Pseudoniphargus djemoi sp. n. A, uropod 3 of female 1 (6,5 mm); B, uropod 3 of male 2 (6,5 mm); C, telson of female 1 (6,5 mm); D, telson of male 1 (6 mm); E, uropods 1 and 2 of female 1 (6,5 mm); F,antenna 1 of female 2 (6 mm) with detail of accessory flagellum (F1), and apex of flagellum (F2); G, antenna 1 of male 1 (6 mm) with detail of apex of flagellum (G1) and accessory flagellum (G2); H, uropods 1 and 2 of male 1 (6 mm)." httpUri="https://zenodo.org/record/1440514/files/figure.png" pageId="2" pageNumber="127">Fig. 3F, G</figureCitation>
): short, less than half body length with fairly slender peduncle; accessory flagellum 2- articulate, shorter than first article of main flagellum (at least half its length); main flagellum composed of 1416 articles; most articles bearing one aesthetasc and 34 setae in apical position. Distal-most aesthetasc longer than terminal article of flagellum. A few flaccid sensorial tufted setae present on third peduncle article in both sexes (
<figureCitation id="131316D0B757167CFF7AD29A61C6A5D2" box="[159,246,1689,1714]" captionStart="FIGURE 3" captionStartId="9.[151,250,1919,1941]" captionTargetBox="[163,1357,199,1899]" captionTargetId="figure@9.[162,1427,193,1904]" captionTargetPageId="9" captionText="FIGURE 3. Pseudoniphargus djemoi sp. n. A, uropod 3 of female 1 (6,5 mm); B, uropod 3 of male 2 (6,5 mm); C, telson of female 1 (6,5 mm); D, telson of male 1 (6 mm); E, uropods 1 and 2 of female 1 (6,5 mm); F,antenna 1 of female 2 (6 mm) with detail of accessory flagellum (F1), and apex of flagellum (F2); G, antenna 1 of male 1 (6 mm) with detail of apex of flagellum (G1) and accessory flagellum (G2); H, uropods 1 and 2 of male 1 (6 mm)." httpUri="https://zenodo.org/record/1440514/files/figure.png" pageId="2" pageNumber="127">Fig. 3F</figureCitation>
<figureCitation id="131316D0B757167CFF13D2996031A5D2" box="[246,257,1690,1714]" captionStart="FIGURE 1" captionStartId="2.[151,250,1478,1500]" captionTargetBox="[151,1385,822,1412]" captionTargetId="figure@2.[151,1436,777,1454]" captionTargetPageId="2" captionText="FIGURE 1. Pseudoniphargus djemoi sp. n. Habitus of female 3 (6 mm)." httpUri="https://zenodo.org/record/1440510/files/figure.png" pageId="2" pageNumber="127">1</figureCitation>
, G
<figureCitation id="131316D0B757167CFEC6D2996001A5D2" box="[291,305,1690,1714]" captionStart="FIGURE 2" captionStartId="8.[151,250,1929,1951]" captionTargetBox="[209,1374,217,1906]" captionTargetId="figure@8.[178,1402,193,1916]" captionTargetPageId="8" captionText="FIGURE 2. Pseudoniphargus djemoi sp. n. A, pereopod 4 of female 1 (6,5 mm); B, pereopod 7 of male 1 (6 mm) with detail of basis (B1), propodus and dactylus (B2); C, pereopod 5 of male 1 with detail of propodus and dactylus (C1); D, pereopod 6 of male 1 with detail of basis (D1), propodus and dactylus (D2); E, Antenna 2 of male 1." httpUri="https://zenodo.org/record/1440512/files/figure.png" pageId="2" pageNumber="127">2</figureCitation>
) and on terminal article of male main flagellum (compare
<figureCitation id="131316D0B757167CFC24D29A6522A5D2" box="[961,1042,1689,1714]" captionStart="FIGURE 3" captionStartId="9.[151,250,1919,1941]" captionTargetBox="[163,1357,199,1899]" captionTargetId="figure@9.[162,1427,193,1904]" captionTargetPageId="9" captionText="FIGURE 3. Pseudoniphargus djemoi sp. n. A, uropod 3 of female 1 (6,5 mm); B, uropod 3 of male 2 (6,5 mm); C, telson of female 1 (6,5 mm); D, telson of male 1 (6 mm); E, uropods 1 and 2 of female 1 (6,5 mm); F,antenna 1 of female 2 (6 mm) with detail of accessory flagellum (F1), and apex of flagellum (F2); G, antenna 1 of male 1 (6 mm) with detail of apex of flagellum (G1) and accessory flagellum (G2); H, uropods 1 and 2 of male 1 (6 mm)." httpUri="https://zenodo.org/record/1440514/files/figure.png" pageId="2" pageNumber="127">Fig. 3F</figureCitation>
<figureCitation id="131316D0B757167CFBF7D2996510A5D2" box="[1042,1056,1690,1714]" captionStart="FIGURE 2" captionStartId="8.[151,250,1929,1951]" captionTargetBox="[209,1374,217,1906]" captionTargetId="figure@8.[178,1402,193,1916]" captionTargetPageId="8" captionText="FIGURE 2. Pseudoniphargus djemoi sp. n. A, pereopod 4 of female 1 (6,5 mm); B, pereopod 7 of male 1 (6 mm) with detail of basis (B1), propodus and dactylus (B2); C, pereopod 5 of male 1 with detail of propodus and dactylus (C1); D, pereopod 6 of male 1 with detail of basis (D1), propodus and dactylus (D2); E, Antenna 2 of male 1." httpUri="https://zenodo.org/record/1440512/files/figure.png" pageId="2" pageNumber="127">2</figureCitation>
vs.
<figureCitation id="131316D0B757167CFBABD29A6599A5D2" box="[1102,1193,1689,1714]" captionStart="FIGURE 3" captionStartId="9.[151,250,1919,1941]" captionTargetBox="[163,1357,199,1899]" captionTargetId="figure@9.[162,1427,193,1904]" captionTargetPageId="9" captionText="FIGURE 3. Pseudoniphargus djemoi sp. n. A, uropod 3 of female 1 (6,5 mm); B, uropod 3 of male 2 (6,5 mm); C, telson of female 1 (6,5 mm); D, telson of male 1 (6 mm); E, uropods 1 and 2 of female 1 (6,5 mm); F,antenna 1 of female 2 (6 mm) with detail of accessory flagellum (F1), and apex of flagellum (F2); G, antenna 1 of male 1 (6 mm) with detail of apex of flagellum (G1) and accessory flagellum (G2); H, uropods 1 and 2 of male 1 (6 mm)." httpUri="https://zenodo.org/record/1440514/files/figure.png" pageId="2" pageNumber="127">Fig. 3G</figureCitation>
<figureCitation id="131316D0B757167CFB4FD2996585A5D2" box="[1194,1205,1690,1714]" captionStart="FIGURE 1" captionStartId="2.[151,250,1478,1500]" captionTargetBox="[151,1385,822,1412]" captionTargetId="figure@2.[151,1436,777,1454]" captionTargetPageId="2" captionText="FIGURE 1. Pseudoniphargus djemoi sp. n. Habitus of female 3 (6 mm)." httpUri="https://zenodo.org/record/1440510/files/figure.png" pageId="2" pageNumber="127">1</figureCitation>
).
</paragraph>
<paragraph id="8B970A55B757167CFF22D2BD60A8A47E" blockId="2.[151,1437,1545,2038]" pageId="2" pageNumber="127">
Antenna 2 (
<figureCitation id="131316D0B757167CFEAFD2BE6092A5B6" box="[330,418,1725,1750]" captionStart="FIGURE 2" captionStartId="8.[151,250,1929,1951]" captionTargetBox="[209,1374,217,1906]" captionTargetId="figure@8.[178,1402,193,1916]" captionTargetPageId="8" captionText="FIGURE 2. Pseudoniphargus djemoi sp. n. A, pereopod 4 of female 1 (6,5 mm); B, pereopod 7 of male 1 (6 mm) with detail of basis (B1), propodus and dactylus (B2); C, pereopod 5 of male 1 with detail of propodus and dactylus (C1); D, pereopod 6 of male 1 with detail of basis (D1), propodus and dactylus (D2); E, Antenna 2 of male 1." httpUri="https://zenodo.org/record/1440512/files/figure.png" pageId="2" pageNumber="127">Fig. 2E</figureCitation>
): Less than half length of antenna 1. As in most
<taxonomicName id="4C2871D6B757167CFC24D2BE65B4A5B5" authorityName="Dole-Olivier &amp; Hafid &amp; Piscart" authorityYear="2018" box="[961,1156,1725,1749]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis id="B95CD647B757167CFC24D2BE65B4A5B5" box="[961,1156,1725,1749]" italics="true" pageId="2" pageNumber="127">Pseudoniphargus</emphasis>
</taxonomicName>
species, peduncle article 4 longest; peduncle articles 35 relative length as: 0.35: 1: 0.82. Flagellum 6- or 7-articulate, shorter than peduncle articles 4+5 combined.
</paragraph>
<paragraph id="8B970A55B757167CFF22D329642BA422" blockId="2.[151,1437,1545,2038]" box="[199,1307,1833,1858]" pageId="2" pageNumber="127">
Upper lip (
<figureCitation id="131316D0B757167CFEA4D32A60A6A422" box="[321,406,1833,1858]" captionStart="FIGURE 4" captionStartId="10.[151,250,1937,1959]" captionTargetBox="[191,1353,205,1908]" captionTargetId="figure@10.[172,1402,193,1919]" captionTargetPageId="10" captionText="FIGURE 4. Pseudoniphargus djemoi sp. n. A, maxillipeds of male 2 (5,5 mm), with detail of distal part of inner plates (A1); B, left mandible of female 1 (6,5 mm); C, right mandible of female 1 (6,5 mm); DF, epimera 1 to 3; G, labrum of male 2 (5,5 mm); H, labium of male 2 (5,5 mm); I, maxilla 1 of female 1 (6,5 mm)." httpUri="https://zenodo.org/record/1440516/files/figure.png" pageId="2" pageNumber="127">Fig.4G</figureCitation>
): Distal margin rounded, bearing two symmetrical tufts of short and fine setules.
</paragraph>
<paragraph id="8B970A55B757167CFF22D34E6216A4E9" blockId="2.[151,1437,1545,2038]" pageId="2" pageNumber="127">
Left mandible (
<figureCitation id="131316D0B757167CFE91D34E60FCA406" box="[372,460,1869,1894]" captionStart="FIGURE 4" captionStartId="10.[151,250,1937,1959]" captionTargetBox="[191,1353,205,1908]" captionTargetId="figure@10.[172,1402,193,1919]" captionTargetPageId="10" captionText="FIGURE 4. Pseudoniphargus djemoi sp. n. A, maxillipeds of male 2 (5,5 mm), with detail of distal part of inner plates (A1); B, left mandible of female 1 (6,5 mm); C, right mandible of female 1 (6,5 mm); DF, epimera 1 to 3; G, labrum of male 2 (5,5 mm); H, labium of male 2 (5,5 mm); I, maxilla 1 of female 1 (6,5 mm)." httpUri="https://zenodo.org/record/1440516/files/figure.png" pageId="2" pageNumber="127">Fig. 4B</figureCitation>
): incisor 5-denticulate,
<emphasis id="B95CD647B757167CFD36D34E6249A405" box="[723,889,1869,1893]" italics="true" pageId="2" pageNumber="127">lacinia mobilis</emphasis>
4-denticulate, spine row composed of 5 pappose setae; molar process well developed, with long molar seta.
</paragraph>
<paragraph id="8B970A55B757167DFF22D39662BEA398" blockId="2.[151,1437,1545,2038]" lastBlockId="3.[151,1437,151,2013]" lastPageId="3" lastPageNumber="128" pageId="2" pageNumber="127">
Right mandible (
<figureCitation id="131316D0B757167CFE66D39660ECA4CE" box="[387,476,1941,1966]" captionStart="FIGURE 4" captionStartId="10.[151,250,1937,1959]" captionTargetBox="[191,1353,205,1908]" captionTargetId="figure@10.[172,1402,193,1919]" captionTargetPageId="10" captionText="FIGURE 4. Pseudoniphargus djemoi sp. n. A, maxillipeds of male 2 (5,5 mm), with detail of distal part of inner plates (A1); B, left mandible of female 1 (6,5 mm); C, right mandible of female 1 (6,5 mm); DF, epimera 1 to 3; G, labrum of male 2 (5,5 mm); H, labium of male 2 (5,5 mm); I, maxilla 1 of female 1 (6,5 mm)." httpUri="https://zenodo.org/record/1440516/files/figure.png" pageId="2" pageNumber="127">Fig. 4C</figureCitation>
): incisor 4-denticulate,
<emphasis id="B95CD647B757167CFD06D3956242A4CD" box="[739,882,1941,1965]" italics="true" pageId="2" pageNumber="127">pars molaris</emphasis>
bifid, each branch with row of numerous (~7 and 4 respectively) small rounded denticles; setal row composed of only 3 pappose setae; molar process as in left counterpart; molar seta lacking. Left and right palps similar, 3-articulated. Article 1 naked; article 2 longer, as long as article 3, bearing 45 simple setae; article 3 with 3 long E-setae, one of them with a row of small and delicate setules proximally; distal most inner margin of article with row of plumose D-setae; 1 simple A-seta and 1 B-seta on dorsal and lateral faces of article as figured (see
<bibRefCitation id="EFB977A4B756167DFD3DD4E362B1A398" box="[728,897,223,248]" pageId="3" pageNumber="128" refId="ref8448" refString="Karaman, G. (1969) XXVII. Beitrag zur kenntnis der Amphipoden. Arten der genera Echinogammarus Stebb. und Chaetogammarus Mart. an der jugoslawischer adriakuste. Glasnik Republickog zavoda zazastitu prirode i Prirodnjacke zbirke u Titogradu, 2, 59 - 84." type="journal article">Karaman 1969</bibRefCitation>
).
</paragraph>
<paragraph id="8B970A55B756167DFF22D506609DA204" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="128">
Lower lip (= Labium) (
<figureCitation id="131316D0B756167DFE34D507631DA27C" box="[465,557,260,285]" captionStart="FIGURE 4" captionStartId="10.[151,250,1937,1959]" captionTargetBox="[191,1353,205,1908]" captionTargetId="figure@10.[172,1402,193,1919]" captionTargetPageId="10" captionText="FIGURE 4. Pseudoniphargus djemoi sp. n. A, maxillipeds of male 2 (5,5 mm), with detail of distal part of inner plates (A1); B, left mandible of female 1 (6,5 mm); C, right mandible of female 1 (6,5 mm); DF, epimera 1 to 3; G, labrum of male 2 (5,5 mm); H, labium of male 2 (5,5 mm); I, maxilla 1 of female 1 (6,5 mm)." httpUri="https://zenodo.org/record/1440516/files/figure.png" pageId="3" pageNumber="128">Fig. 4H</figureCitation>
): inner lobes smooth, weakly developed, fused proximally (only in
<specimenCount id="9D2EC1DCB756167DFACAD5076475A27C" box="[1327,1349,260,284]" pageId="3" pageNumber="128" type="male"></specimenCount>
2), with the two lobes distinctly separated only in the distal part. Outer lobes well developed, with relatively long and narrow lateral processes.
</paragraph>
<paragraph id="8B970A55B756167DFF22D56C62F5A295" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="128">
Maxilla 1 (
<figureCitation id="131316D0B756167DFEADD56C60AAA2E8" box="[328,410,367,392]" captionStart="FIGURE 4" captionStartId="10.[151,250,1937,1959]" captionTargetBox="[191,1353,205,1908]" captionTargetId="figure@10.[172,1402,193,1919]" captionTargetPageId="10" captionText="FIGURE 4. Pseudoniphargus djemoi sp. n. A, maxillipeds of male 2 (5,5 mm), with detail of distal part of inner plates (A1); B, left mandible of female 1 (6,5 mm); C, right mandible of female 1 (6,5 mm); DF, epimera 1 to 3; G, labrum of male 2 (5,5 mm); H, labium of male 2 (5,5 mm); I, maxilla 1 of female 1 (6,5 mm)." httpUri="https://zenodo.org/record/1440516/files/figure.png" pageId="3" pageNumber="128">
Fig.
<date id="FF962C95B756167DFE9AD57360AAA2E8" box="[383,410,368,392]" pageId="3" pageNumber="128">4I</date>
</figureCitation>
): inner plate with 2 long apical plumose setae. Outer plate slightly curved and bearing 7 apical stout spines, most of them bifid, some simple (all bifid in ♂1); inner margin partially covered with few fine setules. Palp 2-articulated, with distal article slightly curved, bearing 68 (depending on specimens) terminal stout setae, and long setules (314, depending on specimens) on outer margin.
</paragraph>
<paragraph id="8B970A55B756167DFF22D5FC6519A100" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="128">Maxilla 2 (not represented): inner plate with numerous (~12) long setae in disto-medial position, with additional fine setules on inner margin. Outer plate with ca. 1015 long apical setae and 5 long curved plumose setae on outer subdistal margin; both groups of setae separated by a short spinule.</paragraph>
<paragraph id="8B970A55B756167DFF22D66F628AA0C5" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="128">
Maxilliped (
<figureCitation id="131316D0B756167DFEB7D66F609EA1E4" box="[338,430,620,645]" captionStart="FIGURE 4" captionStartId="10.[151,250,1937,1959]" captionTargetBox="[191,1353,205,1908]" captionTargetId="figure@10.[172,1402,193,1919]" captionTargetPageId="10" captionText="FIGURE 4. Pseudoniphargus djemoi sp. n. A, maxillipeds of male 2 (5,5 mm), with detail of distal part of inner plates (A1); B, left mandible of female 1 (6,5 mm); C, right mandible of female 1 (6,5 mm); DF, epimera 1 to 3; G, labrum of male 2 (5,5 mm); H, labium of male 2 (5,5 mm); I, maxilla 1 of female 1 (6,5 mm)." httpUri="https://zenodo.org/record/1440516/files/figure.png" pageId="3" pageNumber="128">Fig. 4A</figureCitation>
): inner plate ~
<date id="FF962C95B756167DFDB7D66F635EA1E4" box="[594,622,620,644]" pageId="3" pageNumber="128">4x</date>
longer than wide, distal margin with with 3 plumose and 1 long simple setae, and 2 lanceolate ciliated teeth (
<figureCitation id="131316D0B756167DFDDBD68C63ADA1C8" box="[574,669,655,680]" captionStart="FIGURE 4" captionStartId="10.[151,250,1937,1959]" captionTargetBox="[191,1353,205,1908]" captionTargetId="figure@10.[172,1402,193,1919]" captionTargetPageId="10" captionText="FIGURE 4. Pseudoniphargus djemoi sp. n. A, maxillipeds of male 2 (5,5 mm), with detail of distal part of inner plates (A1); B, left mandible of female 1 (6,5 mm); C, right mandible of female 1 (6,5 mm); DF, epimera 1 to 3; G, labrum of male 2 (5,5 mm); H, labium of male 2 (5,5 mm); I, maxilla 1 of female 1 (6,5 mm)." httpUri="https://zenodo.org/record/1440516/files/figure.png" pageId="3" pageNumber="128">Fig. 4A</figureCitation>
<figureCitation id="131316D0B756167DFD78D6936397A1C8" box="[669,679,656,680]" captionStart="FIGURE 1" captionStartId="2.[151,250,1478,1500]" captionTargetBox="[151,1385,822,1412]" captionTargetId="figure@2.[151,1436,777,1454]" captionTargetPageId="2" captionText="FIGURE 1. Pseudoniphargus djemoi sp. n. Habitus of female 3 (6 mm)." httpUri="https://zenodo.org/record/1440510/files/figure.png" pageId="3" pageNumber="128">1</figureCitation>
); distal part of plate covered with fine setules and bearing 1 lateral seta on outer margin. Outer plate with lateral margin evenly rounded, naked; distomedial margin fringed with~10 knife-like teeth; teeth progressively larger towards distal. Slight variation in form of teeth observed between left and right outer plates, the left bearing more spatule-like teeth. Each teeth row continuing with 12 long simple and 2 plumose curved setae. Additional setae are scattered close to the dental row. First article of palp short, bearing a single distal seta on distomedial angle; article 2 long, bearing numerous setae (~15) on inner margin, some of them bifid. Article 3 slightly expanded distally, with numerous (~10) setae (some of them bifid) on distal margin. Dactylus with single seta on outer margin; unguis shorter than dactylus.
</paragraph>
<paragraph id="8B970A55B756167DFF22D7AC65C5A7C0" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="128">
Gnathopod 1 (
<figureCitation id="131316D0B756167DFE8AD7AC60FDA0A8" box="[367,461,943,968]" captionStart="FIGURE 5" captionStartId="11.[151,250,1989,2011]" captionTargetBox="[192,1382,195,1931]" captionTargetId="figure@11.[152,1435,193,1968]" captionTargetPageId="11" captionText="FIGURE 5. Pseudoniphargus djemoi sp. n., male 1 (6 mm). A, gnathopod 1; B, pereopod 4, C, gnathopod 2." httpUri="https://zenodo.org/record/1440518/files/figure.png" pageId="3" pageNumber="128">Fig. 5A</figureCitation>
): Basis wider from proximal to median region, with a row of 8 very long setae on posterior margin, plus another row of 4 similar setae proximally on lateral surface of article. Ischium and merus of similar length, bearing 2 (ischium) and &lt;10 (merus) simple setae on posterior margin. Carpus length similar to propodus, with 6 or 7 (
<specimenCount id="9D2EC1DCB756167DFE78D01F609FA754" box="[413,431,1052,1076]" pageId="3" pageNumber="128" type="female"></specimenCount>
) rows of 36 long and bifid setae on posterior margin, most of them bearing tiny setules along distal half of their length (only the longest), and 4 setae of variable lenght on anterior margin. Propodus 1,38 times longer than wide with about 11 setae irregularly distributed in 4 clusters over anterior margin of article, 46 strong bifid spines of variable length at palm angle (
<figureCitation id="131316D0B756167DFD3AD0846210A7C0" box="[735,800,1159,1184]" captionStart="FIGURE 5" captionStartId="11.[151,250,1989,2011]" captionTargetBox="[192,1382,195,1931]" captionTargetId="figure@11.[152,1435,193,1968]" captionTargetPageId="11" captionText="FIGURE 5. Pseudoniphargus djemoi sp. n., male 1 (6 mm). A, gnathopod 1; B, pereopod 4, C, gnathopod 2." httpUri="https://zenodo.org/record/1440518/files/figure.png" pageId="3" pageNumber="128">Fig. 5</figureCitation>
A-detail); palm margin weakly armoured.
</paragraph>
<paragraph id="8B970A55B756167DFF22D0AF63D6A635" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="128">
Gnathopod 2 (
<figureCitation id="131316D0B756167DFE82D0AF60F1A7A5" box="[359,449,1196,1221]" captionStart="FIGURE 5" captionStartId="11.[151,250,1989,2011]" captionTargetBox="[192,1382,195,1931]" captionTargetId="figure@11.[152,1435,193,1968]" captionTargetPageId="11" captionText="FIGURE 5. Pseudoniphargus djemoi sp. n., male 1 (6 mm). A, gnathopod 1; B, pereopod 4, C, gnathopod 2." httpUri="https://zenodo.org/record/1440518/files/figure.png" pageId="3" pageNumber="128">Fig. 5C</figureCitation>
): Basis with 7 (
<specimenCount id="9D2EC1DCB756167DFD95D0AF63B8A7A4" box="[624,648,1196,1220]" pageId="3" pageNumber="128" type="male"></specimenCount>
1) or 11 (
<specimenCount id="9D2EC1DCB756167DFD0BD0AF6230A7A4" box="[750,768,1196,1220]" pageId="3" pageNumber="128" type="female"></specimenCount>
) setae on posterior margin and 4 (
<specimenCount id="9D2EC1DCB756167DFB67D0AF65AAA7A4" box="[1154,1178,1196,1220]" pageId="3" pageNumber="128" type="male"></specimenCount>
1) or 910 (
<specimenCount id="9D2EC1DCB756167DFAF8D0AF641FA7A4" box="[1309,1327,1196,1220]" pageId="3" pageNumber="128" type="female"></specimenCount>
) setae on anterior margin. Ischium, merus and carpus all of similar length. Carpus bearing 2 groups of bifid setae on posterior margin and 2 groups of shorter setae on distolateral margin. Propodus 1,5 times longer than wide and more twice longer than carpus; 3 (
<specimenCount id="9D2EC1DCB756167DFE47D11B608AA650" box="[418,442,1304,1328]" pageId="3" pageNumber="128" type="male"></specimenCount>
1) or 4 (
<specimenCount id="9D2EC1DCB756167DFDFBD11B6300A650" box="[542,560,1304,1328]" pageId="3" pageNumber="128" type="female"></specimenCount>
) groups of bifid setae on posterior margin; palmar margin convex, weakly armoured; palm angle with 3 unequal conical spines.
</paragraph>
<paragraph id="8B970A55B756167DFF22D15C6068A6FD" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="128">Pereopod 3 (not figured): similar to pereopod 4, but with coxal plate narrow and high, lacking concave posterior margin.</paragraph>
<paragraph id="8B970A55B756167DFF22D1A4631EA54D" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="128">
Pereopod 4 (
<figureCitation id="131316D0B756167DFEBDD1A46088A6A0" box="[344,440,1447,1472]" captionStart="FIGURE 2" captionStartId="8.[151,250,1929,1951]" captionTargetBox="[209,1374,217,1906]" captionTargetId="figure@8.[178,1402,193,1916]" captionTargetPageId="8" captionText="FIGURE 2. Pseudoniphargus djemoi sp. n. A, pereopod 4 of female 1 (6,5 mm); B, pereopod 7 of male 1 (6 mm) with detail of basis (B1), propodus and dactylus (B2); C, pereopod 5 of male 1 with detail of propodus and dactylus (C1); D, pereopod 6 of male 1 with detail of basis (D1), propodus and dactylus (D2); E, Antenna 2 of male 1." httpUri="https://zenodo.org/record/1440512/files/figure.png" pageId="3" pageNumber="128">Figs 2A</figureCitation>
;
<figureCitation id="131316D0B756167DFE20D1A460D9A6A0" box="[453,489,1447,1472]" captionStart="FIGURE 5" captionStartId="11.[151,250,1989,2011]" captionTargetBox="[192,1382,195,1931]" captionTargetId="figure@11.[152,1435,193,1968]" captionTargetPageId="11" captionText="FIGURE 5. Pseudoniphargus djemoi sp. n., male 1 (6 mm). A, gnathopod 1; B, pereopod 4, C, gnathopod 2." httpUri="https://zenodo.org/record/1440518/files/figure.png" pageId="3" pageNumber="128">5B</figureCitation>
): coxal plate as high as wide, with convex anterior and concave posterior margins, and 5 short setules on ventral border. Basis armed with 5 (
<specimenCount id="9D2EC1DCB756167DFCF9D1CF6202A684" box="[796,818,1484,1508]" pageId="3" pageNumber="128" type="male"></specimenCount>
) or 9 (
<specimenCount id="9D2EC1DCB756167DFC98D1CF62BFA684" box="[893,911,1484,1508]" pageId="3" pageNumber="128" type="female"></specimenCount>
) long setae on posterior margin, 2 (
<specimenCount id="9D2EC1DCB756167DFAF8D1CF6403A684" box="[1309,1331,1484,1508]" pageId="3" pageNumber="128" type="male"></specimenCount>
) or 6 (
<specimenCount id="9D2EC1DCB756167DFA9AD1CF64A1A684" box="[1407,1425,1484,1508]" pageId="3" pageNumber="128" type="female"></specimenCount>
) anterior setae and 1 distal seta. Carpus shorter than both merus and propodus. Dactylus 3035% of propodus length; unguis shorter than dactylus.
</paragraph>
<paragraph id="8B970A55B756167DFF22D23464BEA5F8" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="128">
Pereopod 5 (
<figureCitation id="131316D0B756167DFEB1D23460FFA530" box="[340,463,1591,1616]" captionStart="FIGURE 2" captionStartId="8.[151,250,1929,1951]" captionTargetBox="[209,1374,217,1906]" captionTargetId="figure@8.[178,1402,193,1916]" captionTargetPageId="8" captionText="FIGURE 2. Pseudoniphargus djemoi sp. n. A, pereopod 4 of female 1 (6,5 mm); B, pereopod 7 of male 1 (6 mm) with detail of basis (B1), propodus and dactylus (B2); C, pereopod 5 of male 1 with detail of propodus and dactylus (C1); D, pereopod 6 of male 1 with detail of basis (D1), propodus and dactylus (D2); E, Antenna 2 of male 1." httpUri="https://zenodo.org/record/1440512/files/figure.png" pageId="3" pageNumber="128">Fig. 2C, C</figureCitation>
<figureCitation id="131316D0B756167DFE2AD23B60EAA530" box="[463,474,1592,1616]" captionStart="FIGURE 1" captionStartId="2.[151,250,1478,1500]" captionTargetBox="[151,1385,822,1412]" captionTargetId="figure@2.[151,1436,777,1454]" captionTargetPageId="2" captionText="FIGURE 1. Pseudoniphargus djemoi sp. n. Habitus of female 3 (6 mm)." httpUri="https://zenodo.org/record/1440510/files/figure.png" pageId="3" pageNumber="128">1</figureCitation>
): remarkably shorter than pereopods 6 and 7 (about half as long as their length). Basis large with anterior and posterior margins convex, and with a reduced posterodistal lobe. Merus and carpus similar in length, propodus slightly longer. Dactylus short (34% of propodus) with a short claw (40% of dactylus length).
</paragraph>
<paragraph id="8B970A55B756167DFF22D2A7632AA448" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="128">
Pereopods 67 (
<figureCitation id="131316D0B756167DFE9AD2A760F4A5DC" box="[383,452,1700,1725]" captionStart="FIGURE 2" captionStartId="8.[151,250,1929,1951]" captionTargetBox="[209,1374,217,1906]" captionTargetId="figure@8.[178,1402,193,1916]" captionTargetPageId="8" captionText="FIGURE 2. Pseudoniphargus djemoi sp. n. A, pereopod 4 of female 1 (6,5 mm); B, pereopod 7 of male 1 (6 mm) with detail of basis (B1), propodus and dactylus (B2); C, pereopod 5 of male 1 with detail of propodus and dactylus (C1); D, pereopod 6 of male 1 with detail of basis (D1), propodus and dactylus (D2); E, Antenna 2 of male 1." httpUri="https://zenodo.org/record/1440512/files/figure.png" pageId="3" pageNumber="128">Fig. 2</figureCitation>
B-B2, D-D2): pereopod 7 longer than pereopod 6. Basis with truncated posteroventral angle giving article a trapezoidal aspect. Articles progressively longer from merus to propodus (merus length = 43% of propodus). Pereopod 7 propodus not elongated (less than 7 times as long as broad). Dactylus with a short claw (2530% of dactylus length).
</paragraph>
<paragraph id="8B970A55B756167DFF22D33763ABA410" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="128">
Epimeral plates 13 (
<figureCitation id="131316D0B756167DFE27D337630FA42C" box="[450,575,1844,1869]" captionStart="FIGURE 4" captionStartId="10.[151,250,1937,1959]" captionTargetBox="[191,1353,205,1908]" captionTargetId="figure@10.[172,1402,193,1919]" captionTargetPageId="10" captionText="FIGURE 4. Pseudoniphargus djemoi sp. n. A, maxillipeds of male 2 (5,5 mm), with detail of distal part of inner plates (A1); B, left mandible of female 1 (6,5 mm); C, right mandible of female 1 (6,5 mm); DF, epimera 1 to 3; G, labrum of male 2 (5,5 mm); H, labium of male 2 (5,5 mm); I, maxilla 1 of female 1 (6,5 mm)." httpUri="https://zenodo.org/record/1440516/files/figure.png" pageId="3" pageNumber="128">Fig. 4DF</figureCitation>
): posteroventral angle straight; ventral border of plate 2 slightly rounded; ventral margin armature of plates as: 1-3-3/4).
</paragraph>
<paragraph id="8B970A55B756167DFF22D37F653AA4F4" blockId="3.[151,1437,151,2013]" box="[199,1034,1916,1941]" pageId="3" pageNumber="128">
Metasomites 13 smooth (
<figureCitation id="131316D0B756167DFE15D37F6301A4F4" box="[496,561,1916,1941]" captionStart="FIGURE 1" captionStartId="2.[151,250,1478,1500]" captionTargetBox="[151,1385,822,1412]" captionTargetId="figure@2.[151,1436,777,1454]" captionTargetPageId="2" captionText="FIGURE 1. Pseudoniphargus djemoi sp. n. Habitus of female 3 (6 mm)." httpUri="https://zenodo.org/record/1440510/files/figure.png" pageId="3" pageNumber="128">Fig. 1</figureCitation>
), without denticulated posterodorsal spur.
</paragraph>
<paragraph id="8B970A55B756167DFF22D3A36212A4BD" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="128">Pleopods 13 (not represented): protopods bearing each 2 retinacles. Exopods 9, 8, and 6-articulate, respectively; endopods 7, 6, and 6-articulate, respectively.</paragraph>
<paragraph id="8B970A55B751167AFF22D4946272A27B" blockId="4.[151,1437,151,2035]" pageId="4" pageNumber="129">
Uropod 1 (
<figureCitation id="131316D0B751167AFEA3D49460F2A3D0" box="[326,450,151,176]" captionStart="FIGURE 3" captionStartId="9.[151,250,1919,1941]" captionTargetBox="[163,1357,199,1899]" captionTargetId="figure@9.[162,1427,193,1904]" captionTargetPageId="9" captionText="FIGURE 3. Pseudoniphargus djemoi sp. n. A, uropod 3 of female 1 (6,5 mm); B, uropod 3 of male 2 (6,5 mm); C, telson of female 1 (6,5 mm); D, telson of male 1 (6 mm); E, uropods 1 and 2 of female 1 (6,5 mm); F,antenna 1 of female 2 (6 mm) with detail of accessory flagellum (F1), and apex of flagellum (F2); G, antenna 1 of male 1 (6 mm) with detail of apex of flagellum (G1) and accessory flagellum (G2); H, uropods 1 and 2 of male 1 (6 mm)." httpUri="https://zenodo.org/record/1440514/files/figure.png" pageId="4" pageNumber="129">Fig. 3E, H</figureCitation>
) peduncle longer than both exopod and endopod, basofacial spine lacking; 2 unequal spines on posterolateral angle and single spine on posteromedial angle of article; posterolateral margin with 3(
<specimenCount id="9D2EC1DCB751167AFA91D4BF64BAA3B4" box="[1396,1418,188,212]" pageId="4" pageNumber="129" type="male"></specimenCount>
)- 4(
<specimenCount id="9D2EC1DCB751167AFF57D4E361F3A398" box="[178,195,224,248]" pageId="4" pageNumber="129" type="female"></specimenCount>
) spines, posteromedial margin smooth. Exopodi slightly shorter than endopodi, both articles each with 5 apical spines (2 of them very short) and without marginal armature.
</paragraph>
<paragraph id="8B970A55B751167AFF22D52461D4A203" blockId="4.[151,1437,151,2035]" pageId="4" pageNumber="129">
Uropod 2 (
<figureCitation id="131316D0B751167AFEA0D52460F3A25F" box="[325,451,295,319]" captionStart="FIGURE 3" captionStartId="9.[151,250,1919,1941]" captionTargetBox="[163,1357,199,1899]" captionTargetId="figure@9.[162,1427,193,1904]" captionTargetPageId="9" captionText="FIGURE 3. Pseudoniphargus djemoi sp. n. A, uropod 3 of female 1 (6,5 mm); B, uropod 3 of male 2 (6,5 mm); C, telson of female 1 (6,5 mm); D, telson of male 1 (6 mm); E, uropods 1 and 2 of female 1 (6,5 mm); F,antenna 1 of female 2 (6 mm) with detail of accessory flagellum (F1), and apex of flagellum (F2); G, antenna 1 of male 1 (6 mm) with detail of apex of flagellum (G1) and accessory flagellum (G2); H, uropods 1 and 2 of male 1 (6 mm)." httpUri="https://zenodo.org/record/1440514/files/figure.png" pageId="4" pageNumber="129">Fig. 3E, H</figureCitation>
): peduncle with 1 (♂)-3 (♀) distal spines and 1 lateral spine; rami each with 5 apical spines.
</paragraph>
<paragraph id="8B970A55B751167AFF22D56D60B3A2AE" blockId="4.[151,1437,151,2035]" pageId="4" pageNumber="129">
Male Uropod 3 (
<figureCitation id="131316D0B751167AFE68D56D60DAA2E7" box="[397,490,366,391]" captionStart="FIGURE 3" captionStartId="9.[151,250,1919,1941]" captionTargetBox="[163,1357,199,1899]" captionTargetId="figure@9.[162,1427,193,1904]" captionTargetPageId="9" captionText="FIGURE 3. Pseudoniphargus djemoi sp. n. A, uropod 3 of female 1 (6,5 mm); B, uropod 3 of male 2 (6,5 mm); C, telson of female 1 (6,5 mm); D, telson of male 1 (6 mm); E, uropods 1 and 2 of female 1 (6,5 mm); F,antenna 1 of female 2 (6 mm) with detail of accessory flagellum (F1), and apex of flagellum (F2); G, antenna 1 of male 1 (6 mm) with detail of apex of flagellum (G1) and accessory flagellum (G2); H, uropods 1 and 2 of male 1 (6 mm)." httpUri="https://zenodo.org/record/1440514/files/figure.png" pageId="4" pageNumber="129">Fig. 3B</figureCitation>
): peduncle slightly elongate, about 2.54 times as long as broad; exopod strongly elongate and upcurved, about 14.5 times as long as broad; bearing 5 external and 7 internal group of 14 spines; 4 6 short apical spines.
</paragraph>
<paragraph id="8B970A55B751167AFF22D5DA61D4A15A" blockId="4.[151,1437,151,2035]" pageId="4" pageNumber="129">
Female Uropod 3 (
<figureCitation id="131316D0B751167AFE79D5DA60C4A292" box="[412,500,473,498]" captionStart="FIGURE 3" captionStartId="9.[151,250,1919,1941]" captionTargetBox="[163,1357,199,1899]" captionTargetId="figure@9.[162,1427,193,1904]" captionTargetPageId="9" captionText="FIGURE 3. Pseudoniphargus djemoi sp. n. A, uropod 3 of female 1 (6,5 mm); B, uropod 3 of male 2 (6,5 mm); C, telson of female 1 (6,5 mm); D, telson of male 1 (6 mm); E, uropods 1 and 2 of female 1 (6,5 mm); F,antenna 1 of female 2 (6 mm) with detail of accessory flagellum (F1), and apex of flagellum (F2); G, antenna 1 of male 1 (6 mm) with detail of apex of flagellum (G1) and accessory flagellum (G2); H, uropods 1 and 2 of male 1 (6 mm)." httpUri="https://zenodo.org/record/1440514/files/figure.png" pageId="4" pageNumber="129">Fig. 3A</figureCitation>
) peduncle subrectangular, 1.7 times as long as broad; exopod moderately elongate, about 11.1 times as long as broad; bearing 5 external and 4 internal group of 14 short spines; 46 short apical spines.
</paragraph>
<paragraph id="8B970A55B751167AFF22D64762D6A18C" blockId="4.[151,1437,151,2035]" pageId="4" pageNumber="129">
Telson (
<figureCitation id="131316D0B751167AFEC6D6476092A13D" box="[291,418,580,605]" captionStart="FIGURE 3" captionStartId="9.[151,250,1919,1941]" captionTargetBox="[163,1357,199,1899]" captionTargetId="figure@9.[162,1427,193,1904]" captionTargetPageId="9" captionText="FIGURE 3. Pseudoniphargus djemoi sp. n. A, uropod 3 of female 1 (6,5 mm); B, uropod 3 of male 2 (6,5 mm); C, telson of female 1 (6,5 mm); D, telson of male 1 (6 mm); E, uropods 1 and 2 of female 1 (6,5 mm); F,antenna 1 of female 2 (6 mm) with detail of accessory flagellum (F1), and apex of flagellum (F2); G, antenna 1 of male 1 (6 mm) with detail of apex of flagellum (G1) and accessory flagellum (G2); H, uropods 1 and 2 of male 1 (6 mm)." httpUri="https://zenodo.org/record/1440514/files/figure.png" pageId="4" pageNumber="129">Fig. 3C, D</figureCitation>
): subrectangular, slightly broader than long, shallowly excavate. Cluster of 23 subequal spines subdistally at each side, plus pair of flaccid sensorial tufted seta (plumose setules) on lateral margin; spines shorter than telson itself in both sexes. Sexual dimorphism apparent in telson being larger in the female, with emargination extremely shallow, and three spines at each side compared to only two in the male. Lateral sensorial tufted setae are also in a more distal position in the male than in the female.
</paragraph>
</subSubSection>
<subSubSection id="C33259DEB751167AFF22D6F463A8A62B" pageId="4" pageNumber="129" type="discussion">
<paragraph id="8B970A55B751167AFF22D6F4642FA715" blockId="4.[151,1437,151,2035]" pageId="4" pageNumber="129">
<emphasis id="B95CD647B751167AFF22D6F460EFA070" bold="true" box="[199,479,759,784]" pageId="4" pageNumber="129">Remarks and affinities.</emphasis>
<taxonomicName id="4C2871D6B751167AFE00D6F463CCA06F" authorityName="Dole-Olivier &amp; Hafid &amp; Piscart" authorityYear="2018" box="[485,764,759,783]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="129" phylum="Arthropoda" rank="species" species="djemoi">
<emphasis id="B95CD647B751167AFE00D6F463CCA06F" box="[485,764,759,783]" italics="true" pageId="4" pageNumber="129">Pseudoniphargus djemoi</emphasis>
</taxonomicName>
belongs to a group of species characterized by the common display of a male uropod 3 exopod strongly elongate (more than 10 times as long as broad) but with the corresponding peduncle only moderately elongate (between 2.1 and 3.5 times as long as broad), combined with a pereopod 5 - pereopod 7 basis lacking a strongly overhanging posterodistal lobe in the male (
<bibRefCitation id="EFB977A4B751167AFB4AD76164A1A01B" box="[1199,1425,866,891]" pageId="4" pageNumber="129" refId="ref9184" refString="Stokkan, M., Perez-Fernandez, A., Baena, M. &amp; Jaume, D. (2018) Two new species of Pseudoniphargus (Amphipoda: Pseudoniphargidae) from southern Spain. Zootaxa, 4418 (3), 264 - 280. https: // doi. org / 10.11646 / zootaxa. 4418.3.4" type="journal article">
Stokkan
<emphasis id="B95CD647B751167AFAF1D7606475A01B" box="[1300,1349,867,891]" italics="true" pageId="4" pageNumber="129">et al</emphasis>
. 2018
</bibRefCitation>
). These species include:
<taxonomicName id="4C2871D6B751167AFE43D78463C4A0FF" authority="Notenboom, 1987" authorityName="Notenboom" authorityYear="1987" box="[422,756,902,927]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="129" phylum="Arthropoda" rank="species" species="fragilis">
<emphasis id="B95CD647B751167AFE43D7846327A0FE" box="[422,535,902,926]" italics="true" pageId="4" pageNumber="129">P. fragilis</emphasis>
<bibRefCitation id="EFB977A4B751167AFDC7D78563C4A0FF" box="[546,756,902,927]" pageId="4" pageNumber="129" refId="ref8796" refString="Notenboom, J. (1987) Species of the genus Pseudoniphargus Chevreux, 1901 (Amphipoda) from the Betic Cordillera of southern Spain. Bijdragen tot de Dierkunde, 57 (1), 87 - 150." type="journal article">Notenboom, 1987</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C2871D6B751167AFCD3D784659EA0FF" authority="Notenboom, 1987" authorityName="Notenboom" authorityYear="1987" box="[822,1198,902,927]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="129" phylum="Arthropoda" rank="species" species="nevadensis">
<emphasis id="B95CD647B751167AFCD3D78462E1A0FE" box="[822,977,902,926]" italics="true" pageId="4" pageNumber="129">P. nevadensis</emphasis>
<bibRefCitation id="EFB977A4B751167AFC39D785659EA0FF" box="[988,1198,902,927]" pageId="4" pageNumber="129" refId="ref8796" refString="Notenboom, J. (1987) Species of the genus Pseudoniphargus Chevreux, 1901 (Amphipoda) from the Betic Cordillera of southern Spain. Bijdragen tot de Dierkunde, 57 (1), 87 - 150." type="journal article">Notenboom, 1987</bibRefCitation>
</taxonomicName>
, both from southern
<collectingCountry id="F33F4AC5B751167AFF72D7A961EAA0A3" box="[151,218,938,963]" name="Spain" pageId="4" pageNumber="129">Spain</collectingCountry>
;
<taxonomicName id="4C2871D6B751167AFF0FD7A86303A0A2" authority="Sanchez, 1991" authorityName="Sanchez" authorityYear="1991" box="[234,563,938,963]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="129" phylum="Arthropoda" rank="species" species="associatus">
<emphasis id="B95CD647B751167AFF0FD7A8604EA0A2" box="[234,382,939,962]" italics="true" pageId="4" pageNumber="129">P. associatus</emphasis>
<bibRefCitation id="EFB977A4B751167AFE6CD7A96303A0A2" box="[393,563,938,963]" pageId="4" pageNumber="129" refId="ref8999" refString="Sanchez, E. L. (1991) New records of Pseudoniphargus (Crustacea, Amphipoda) from ground waters of Tenerife and Hierro, with description of new species. Stygologia, 6 (1), 53 - 64." type="journal article">Sánchez, 1991</bibRefCitation>
</taxonomicName>
, from Tenerife (
<collectingRegion id="49ECC4B7B751167AFD10D7A9626DA0A3" box="[757,861,938,963]" country="Spain" name="Canarias" pageId="4" pageNumber="129">Canaries</collectingRegion>
); and
<taxonomicName id="4C2871D6B751167AFC4CD7A86466A0A3" authority="Coineau &amp; Boutin, 1996" authorityName="Coineau &amp; Boutin" authorityYear="1996" box="[937,1366,938,963]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="129" phylum="Arthropoda" rank="species" species="longipes">
<emphasis id="B95CD647B751167AFC4CD7A86517A0A2" box="[937,1063,938,962]" italics="true" pageId="4" pageNumber="129">P. longipes</emphasis>
<bibRefCitation id="EFB977A4B751167AFBD6D7A96466A0A3" box="[1075,1366,938,963]" pageId="4" pageNumber="129" refId="ref8241" refString="Coineau, N. &amp; Boutin, C. (1996) Age and origin of the stygobiontic amphipod Pseudoniphargus (Crustacea) in Morocco, with the description of three new species. Bollettino del Museo Civico di Storia Naturale di Uerona, 20, 503 - 520." type="journal article">Coineau &amp; Boutin, 1996</bibRefCitation>
</taxonomicName>
, from northern
<collectingCountry id="F33F4AC5B751167AFF1AD7CC6054A086" box="[255,356,975,998]" name="Morocco" pageId="4" pageNumber="129">Morocco</collectingCountry>
(see
<bibRefCitation id="EFB977A4B751167AFE41D7CE635DA086" box="[420,621,973,998]" pageId="4" pageNumber="129" refId="ref8796" refString="Notenboom, J. (1987) Species of the genus Pseudoniphargus Chevreux, 1901 (Amphipoda) from the Betic Cordillera of southern Spain. Bijdragen tot de Dierkunde, 57 (1), 87 - 150." type="journal article">Notenboom 1987</bibRefCitation>
;
<bibRefCitation id="EFB977A4B751167AFD9ED7CE6214A086" box="[635,804,973,998]" pageId="4" pageNumber="129" refId="ref8999" refString="Sanchez, E. L. (1991) New records of Pseudoniphargus (Crustacea, Amphipoda) from ground waters of Tenerife and Hierro, with description of new species. Stygologia, 6 (1), 53 - 64." type="journal article">Sánchez, 1991</bibRefCitation>
;
<bibRefCitation id="EFB977A4B751167AFCD7D7CE6579A086" box="[818,1097,973,998]" pageId="4" pageNumber="129" refId="ref8241" refString="Coineau, N. &amp; Boutin, C. (1996) Age and origin of the stygobiontic amphipod Pseudoniphargus (Crustacea) in Morocco, with the description of three new species. Bollettino del Museo Civico di Storia Naturale di Uerona, 20, 503 - 520." type="journal article">Coineau &amp; Boutin 1996</bibRefCitation>
). Of them, only
<taxonomicName id="4C2871D6B751167AFAECD7CC64ACA086" authorityName="Sanchez" authorityYear="1991" box="[1289,1436,975,998]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="129" phylum="Arthropoda" rank="species" species="associatus">
<emphasis id="B95CD647B751167AFAECD7CC64ACA086" box="[1289,1436,975,998]" italics="true" pageId="4" pageNumber="129">P. associatus</emphasis>
</taxonomicName>
shares with the new species the absence of basofacial spine on the peduncle of uropod 1. But it can be readily differentiated from the new taxon on the basis of the much shorter female uropod 3 exopod, up to 6.3 times longer than wide (versus up to 11.1 times longer than wide in
<taxonomicName id="4C2871D6B751167AFCF5D0396249A731" authority="Dole-Olivier &amp; Hafid &amp; Piscart, 2018" authorityName="Dole-Olivier &amp; Hafid &amp; Piscart" authorityYear="2018" box="[784,889,1081,1105]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="129" phylum="Arthropoda" rank="species" species="djemoi" status="sp. nov.">
<emphasis id="B95CD647B751167AFCF5D0396249A731" box="[784,889,1081,1105]" italics="true" pageId="4" pageNumber="129">P. djemoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F6B3CB751167AFC67D03A62EEA732" box="[898,990,1081,1106]" pageId="4" pageNumber="129" rank="species">
<emphasis id="B95CD647B751167AFC67D03A62EEA732" bold="true" box="[898,990,1081,1106]" pageId="4" pageNumber="129">sp. nov.</emphasis>
</taxonomicNameLabel>
), and male gnathopod 1 carpus shorter than the corresponding propodus (versus carpus and propodus about equal length in
<taxonomicName id="4C2871D6B751167AFBA2D05E659EA715" authority="Dole-Olivier &amp; Hafid &amp; Piscart, 2018" authorityName="Dole-Olivier &amp; Hafid &amp; Piscart" authorityYear="2018" box="[1095,1198,1117,1141]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="129" phylum="Arthropoda" rank="species" species="djemoi" status="sp. nov.">
<emphasis id="B95CD647B751167AFBA2D05E659EA715" box="[1095,1198,1117,1141]" italics="true" pageId="4" pageNumber="129">P. djemoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F6B3CB751167AFB50D05F643FA715" box="[1205,1295,1116,1141]" pageId="4" pageNumber="129" rank="species">
<emphasis id="B95CD647B751167AFB50D05F643FA715" bold="true" box="[1205,1295,1116,1141]" pageId="4" pageNumber="129">sp. nov.</emphasis>
</taxonomicNameLabel>
).
</paragraph>
<paragraph id="8B970A55B751167AFF22D0836221A664" blockId="4.[151,1437,151,2035]" pageId="4" pageNumber="129">
Three species of
<taxonomicName id="4C2871D6B751167AFE62D083637AA7F8" authorityName="Dole-Olivier &amp; Hafid &amp; Piscart" authorityYear="2018" box="[391,586,1152,1176]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis id="B95CD647B751167AFE62D083637AA7F8" box="[391,586,1152,1176]" italics="true" pageId="4" pageNumber="129">Pseudoniphargus</emphasis>
</taxonomicName>
are known only from the female:
<taxonomicName id="4C2871D6B751167AFC29D0826422A7F8" authority="Messouli, Messana" authorityName="Messouli, Messana &amp; Yacoubi-Khebiza" authorityYear="2006" box="[972,1298,1152,1177]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="129" phylum="Arthropoda" rank="species" species="duplus">
<emphasis id="B95CD647B751167AFC29D0826503A7F8" box="[972,1075,1152,1176]" italics="true" pageId="4" pageNumber="129">P. duplus</emphasis>
Messouli, Messana
</taxonomicName>
&amp; Yacoubi- Khebiza, 2006, and
<taxonomicName id="4C2871D6B751167AFE9AD0A66224A7DD" authority="Karaman &amp; Ruffo, 1989" authorityName="Karaman &amp; Ruffo" authorityYear="1989" box="[383,788,1188,1213]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="129" phylum="Arthropoda" rank="species" species="italicus">
<emphasis id="B95CD647B751167AFE9AD0A660DFA7DC" box="[383,495,1188,1212]" italics="true" pageId="4" pageNumber="129">P. italicus</emphasis>
<bibRefCitation id="EFB977A4B751167AFE1CD0A66224A7DD" box="[505,788,1188,1213]" pageId="4" pageNumber="129" refId="ref8495" refString="Karaman, G. S. &amp; Ruffo, S. (1989) Tyrrhenogammarus sardous, new genus and species with a description of several new taxa of the genus Pseudoniphargus Chevreux, 1901 from Sicily (Amphipoda, Gammaridea). Animalia, 16, 161 - 192." type="journal article">Karaman &amp; Ruffo, 1989</bibRefCitation>
</taxonomicName>
, both from
<collectingRegion id="49ECC4B7B751167AFC78D0A762D2A7DD" box="[925,994,1188,1213]" country="Italy" name="Sicilia" pageId="4" pageNumber="129">Sicily</collectingRegion>
; and
<taxonomicName id="4C2871D6B751167AFBC7D0A66467A7DD" authority="Stock, 1988" authorityName="Stock" authorityYear="1988" box="[1058,1367,1188,1213]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="129" phylum="Arthropoda" rank="species" species="unispinosus">
<emphasis id="B95CD647B751167AFBC7D0A665F4A7DC" box="[1058,1220,1189,1212]" italics="true" pageId="4" pageNumber="129">P. unispinosus</emphasis>
<bibRefCitation id="EFB977A4B751167AFB28D0A76467A7DD" box="[1229,1367,1188,1213]" pageId="4" pageNumber="129" refId="ref9078" refString="Stock J. H. (1988) The amphipod genus Pseudoniphargus (Crustacea) in the Canary Islands. Bijdragen tot de Dierkunde, 58 (1), 47 - 78." type="journal article">Stock, 1988</bibRefCitation>
</taxonomicName>
, from Tenerife (
<collectingRegion id="49ECC4B7B751167AFEE6D0C4605BA780" box="[259,363,1223,1248]" country="Spain" name="Canarias" pageId="4" pageNumber="129">Canaries</collectingRegion>
). Again, they display a basofacial spine on the peduncle of uropod 1, in sharp contrast with the condition exhibited by the new species described herein.
</paragraph>
<paragraph id="8B970A55B751167AFF22D11363A8A62B" blockId="4.[151,1437,151,2035]" pageId="4" pageNumber="129">
A key to the species of
<taxonomicName id="4C2871D6B751167AFE32D10C63AAA647" authorityName="Dole-Olivier &amp; Hafid &amp; Piscart" authorityYear="2018" box="[471,666,1295,1319]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis id="B95CD647B751167AFE32D10C63AAA647" box="[471,666,1295,1319]" italics="true" pageId="4" pageNumber="129">Pseudoniphargus</emphasis>
</taxonomicName>
present in Africa and the Mediterranean islands, including the new species, is presented below (see also
<tableCitation id="C6AA3FEEB751167AFDD0D13063B7A62B" box="[565,647,1331,1356]" captionStart="TABLE 1" captionStartId="6.[147,230,617,639]" captionText="TABLE 1. (Continued)" pageId="4" pageNumber="129">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33259DEB7511678FF22D1556659A554" lastPageId="6" lastPageNumber="131" pageId="4" pageNumber="129" type="distribution">
<paragraph id="8B970A55B751167AFF22D155647BA5D1" blockId="4.[151,1437,151,2035]" pageId="4" pageNumber="129">
<emphasis id="B95CD647B751167AFF22D15562EBA60F" bold="true" box="[199,987,1366,1391]" pageId="4" pageNumber="129">
Distribution of the genus
<taxonomicName id="4C2871D6B751167AFDE7D15563FCA60F" authorityName="Dole-Olivier &amp; Hafid &amp; Piscart" authorityYear="2018" box="[514,716,1366,1391]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis id="B95CD647B751167AFDE7D15563FCA60F" bold="true" box="[514,716,1366,1391]" italics="true" pageId="4" pageNumber="129">Pseudoniphargus</emphasis>
</taxonomicName>
in continental Africa.
</emphasis>
The genus
<taxonomicName id="4C2871D6B751167AFB83D1546419A60F" authorityName="Dole-Olivier &amp; Hafid &amp; Piscart" authorityYear="2018" box="[1126,1321,1367,1391]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis id="B95CD647B751167AFB83D1546419A60F" box="[1126,1321,1367,1391]" italics="true" pageId="4" pageNumber="129">Pseudoniphargus</emphasis>
</taxonomicName>
is widely distributed around the
<collectingRegion id="49ECC4B7B751167AFE79D17860C7A6F2" box="[412,503,1403,1426]" country="American Samoa" name="Western" pageId="4" pageNumber="129">Western</collectingRegion>
Mediterranean coasts and along the Atlantic coasts of the Iberian Peninsula and
<collectingCountry id="F33F4AC5B751167AFF72D19C61CCA6D6" box="[151,252,1439,1462]" name="Morocco" pageId="4" pageNumber="129">Morocco</collectingCountry>
(see
<bibRefCitation id="EFB977A4B751167AFEDED19D6330A6D7" box="[315,512,1438,1463]" pageId="4" pageNumber="129" refId="ref8836" refString="Notenboom, J. (1988) Phylogenetic relationships and biogeography of the groundwater-dwelling amphipod genus Pseudoniphargus (Crustacea), with emphasis on the Iberian species. Bijdragen tot de Dierkunde, 58 (2), 159 - 204." type="journal article">Notenboom 1988</bibRefCitation>
and references herein). The genus is also very diversified in Mediterranean (e.g. Balearic,
<collectingRegion id="49ECC4B7B751167AFEE0D1C1606DA6BB" box="[261,349,1474,1499]" country="France" name="Corse" pageId="4" pageNumber="129">
<collectingCountry id="F33F4AC5B751167AFEE0D1C1606DA6BB" box="[261,349,1474,1499]" name="France" pageId="4" pageNumber="129">Corsica</collectingCountry>
</collectingRegion>
,
<collectingRegion id="49ECC4B7B751167AFE89D1C1609FA6BB" box="[364,431,1474,1499]" country="Italy" name="Sicilia" pageId="4" pageNumber="129">Sicily</collectingRegion>
) and Atlantic islands (e.g. Açores,
<collectingRegion id="49ECC4B7B751167AFCA8D1C16282A6BB" box="[845,946,1474,1499]" country="Spain" name="Canarias" pageId="4" pageNumber="129">Canaries</collectingRegion>
, Madeira) as far as
<collectingCountry id="F33F4AC5B751167AFB78D1C16433A6BB" box="[1181,1283,1474,1499]" name="Bermuda" pageId="4" pageNumber="129">Bermuda</collectingCountry>
(
<bibRefCitation id="EFB977A4B751167AFAF0D1C161E3A69F" pageId="4" pageNumber="129" refId="ref9112" refString="Stock, J. H., Holsinger, J. R., Sket, B. &amp; Iliffe, T. M. (1986) Two new species of Pseudoniphargus (Amphipoda), in Bermudian groundwaters. Zoologica Scripta, 15 (3), 237 - 249. https: // doi. org / 10.1111 / j. 1463 - 6409.1986. tb 00226. x" type="journal article">
Stock
<emphasis id="B95CD647B751167AFA84D1C064ACA6BA" box="[1377,1436,1474,1498]" italics="true" pageId="4" pageNumber="129">et al.</emphasis>
1986
</bibRefCitation>
). This wide distribution likely results in the ability of
<taxonomicName id="4C2871D6B751167AFCDBD1E56531A69E" authorityName="Dole-Olivier &amp; Hafid &amp; Piscart" authorityYear="2018" box="[830,1025,1510,1534]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis id="B95CD647B751167AFCDBD1E56531A69E" box="[830,1025,1510,1534]" italics="true" pageId="4" pageNumber="129">Pseudoniphargus</emphasis>
</taxonomicName>
species to colonize a wide range of environmental conditions (
<bibRefCitation id="EFB977A4B751167AFE2DD20A63A5A542" box="[456,661,1545,1570]" pageId="4" pageNumber="129" refId="ref8836" refString="Notenboom, J. (1988) Phylogenetic relationships and biogeography of the groundwater-dwelling amphipod genus Pseudoniphargus (Crustacea), with emphasis on the Iberian species. Bijdragen tot de Dierkunde, 58 (2), 159 - 204." type="journal article">Notenboom 1988</bibRefCitation>
). For instance, the genus is known to inhabit a very wide range of salinity (from anchihaline caves in marine coastal areas to purely freshwater springs) as well as a wide range of altitude (from
<quantity id="4CD0A7B0B751167AFEDBD25260FEA509" box="[318,462,1617,1641]" metricMagnitude="2" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="4" pageNumber="129" unit="m" value="500.0" valueMax="1000.0" valueMin="0.0">0 to 1000 m</quantity>
a.s.l. for
<taxonomicName id="4C2871D6B751167AFDA4D252652EA509" authority="Stock, 1980" authorityName="Stock" authorityYear="1980" box="[577,1054,1617,1642]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="129" phylum="Arthropoda" rank="species" species="unisexualis">
<emphasis id="B95CD647B751167AFDA4D25262BBA509" box="[577,907,1617,1641]" italics="true" pageId="4" pageNumber="129">Pseudoniphargus unisexualis</emphasis>
<bibRefCitation id="EFB977A4B751167AFC73D252652EA509" box="[918,1054,1617,1642]" pageId="4" pageNumber="129" refId="ref9042" refString="Stock, J. H. (1980) Regression model evolution as exemplified by the genus Pseudoniphargus (Amphipoda). Bijdragen tot de Dierkunde, 50 (1), 105 - 144." type="journal article">
Stock,
<quantity id="4CD0A7B0B751167AFC03D252652EA509" box="[998,1054,1617,1641]" metricMagnitude="1" metricUnit="m" metricValue="5.0291999999999994" pageId="4" pageNumber="129" unit="in" value="1980.0">1980</quantity>
</bibRefCitation>
</taxonomicName>
in
<collectingCountry id="F33F4AC5B751167AFBADD25265BAA50A" box="[1096,1162,1617,1642]" name="Spain" pageId="4" pageNumber="129">Spain</collectingCountry>
) or temperatures (up to 44°C,
<bibRefCitation id="EFB977A4B751167AFF3AD27760F8A5ED" box="[223,456,1652,1677]" pageId="4" pageNumber="129" refId="ref8683" refString="Messouli, M., Boutin, C., Yacoubi-Khebiza, M. &amp; Coineau, N. (2001) Pseudoniphargus (subterranean crustacean amphipod) from Morocco: systematics, phylogeny and ecological and biogeographic aspects. Proceedings of the 13 th International Congress of Speleology, Brasilia, 15 - 22 July 2001, 391 - 394." type="proceedings paper">
Messouli
<emphasis id="B95CD647B751167AFEA8D27660B5A5ED" box="[333,389,1653,1677]" italics="true" pageId="4" pageNumber="129">et al.</emphasis>
2001
</bibRefCitation>
). Recently, the finding of
<taxonomicName id="4C2871D6B751167AFD09D276629FA5ED" authorityName="Dole-Olivier &amp; Hafid &amp; Piscart" authorityYear="2018" box="[748,943,1653,1677]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis id="B95CD647B751167AFD09D276629FA5ED" box="[748,943,1653,1677]" italics="true" pageId="4" pageNumber="129">Pseudoniphargus</emphasis>
</taxonomicName>
in
<collectingCountry id="F33F4AC5B751167AFC36D2776526A5ED" box="[979,1046,1652,1677]" name="Benin" pageId="4" pageNumber="129">Benin</collectingCountry>
extended significantly the African distribution of the genus from the Maghreb to western Africa and south to the Sahara (
<bibRefCitation id="EFB977A4B751167AFBBAD29B640FA5D1" box="[1119,1343,1688,1713]" pageId="4" pageNumber="129" refId="ref8546" refString="Lagnika, M., Messouli, M., Ibikounle, M., Sakiti, N., Boutin, C. &amp; Coineau, N. (2016) First record of groundwater amphipods (Crustacea) from Benin; range extension of the genus Pseudoniphargus to South of the Sahara, in western Africa. Bulletin de la Societe d'Histoire Naturelle de Toulouse, 152, 21 - 30." type="journal article">
Lagnika
<emphasis id="B95CD647B751167AFB26D29A65CCA5D0" box="[1219,1276,1688,1712]" italics="true" pageId="4" pageNumber="129">et al.</emphasis>
2016
</bibRefCitation>
).
</paragraph>
<paragraph id="8B970A55B751167AFF22D2BE65D2A493" blockId="4.[151,1437,151,2035]" pageId="4" pageNumber="129">
However, the wide distribution of the genus does not reflect the dispersal ability of species, that remains very low. The narrow geographic range of most
<taxonomicName id="4C2871D6B751167AFD74D2E36264A598" authorityName="Dole-Olivier &amp; Hafid &amp; Piscart" authorityYear="2018" box="[657,852,1760,1784]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis id="B95CD647B751167AFD74D2E36264A598" box="[657,852,1760,1784]" italics="true" pageId="4" pageNumber="129">Pseudoniphargus</emphasis>
</taxonomicName>
species, which are highly endemic and frequently known only from one or two locations (
<bibRefCitation id="EFB977A4B751167AFD86D30063D9A47C" box="[611,745,1795,1820]" pageId="4" pageNumber="129" refId="ref9042" refString="Stock, J. H. (1980) Regression model evolution as exemplified by the genus Pseudoniphargus (Amphipoda). Bijdragen tot de Dierkunde, 50 (1), 105 - 144." type="journal article">Stock 1980</bibRefCitation>
;
<bibRefCitation id="EFB977A4B751167AFD12D30062F0A47C" box="[759,960,1795,1820]" pageId="4" pageNumber="129" refId="ref8796" refString="Notenboom, J. (1987) Species of the genus Pseudoniphargus Chevreux, 1901 (Amphipoda) from the Betic Cordillera of southern Spain. Bijdragen tot de Dierkunde, 57 (1), 87 - 150." type="journal article">Notenboom 1987</bibRefCitation>
;
<bibRefCitation id="EFB977A4B751167AFC28D30065D5A47C" box="[973,1253,1795,1820]" pageId="4" pageNumber="129" refId="ref8241" refString="Coineau, N. &amp; Boutin, C. (1996) Age and origin of the stygobiontic amphipod Pseudoniphargus (Crustacea) in Morocco, with the description of three new species. Bollettino del Museo Civico di Storia Naturale di Uerona, 20, 503 - 520." type="journal article">Coineau &amp; Boutin 1996</bibRefCitation>
;
<bibRefCitation id="EFB977A4B751167AFB17D30061E1A420" pageId="4" pageNumber="129" refId="ref8745" refString="Messouli, M., Messana, G. &amp; Yacoubi-Kebhiza, M. (2006) Three new species of Pseudoniphargus (Amphipoda), from the groundwater of three Mediterranean islands, with notes on the Ps. adriaticus. Subterranean Biology, 4, 79 - 101." type="journal article">
Messouli
<emphasis id="B95CD647B751167AFA87D30764ACA47C" box="[1378,1436,1796,1820]" italics="true" pageId="4" pageNumber="129">et al.</emphasis>
2006
</bibRefCitation>
) strengthens this idea. In North Africa, only
<taxonomicName id="4C2871D6B751167AFD28D32B62BDA45F" authorityName="Fakher-el-Abiari, Oulbaz, Messouli &amp; Coineau" authorityYear="1999" box="[717,909,1831,1855]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="129" phylum="Arthropoda" rank="species" species="longiflagellum">
<emphasis id="B95CD647B751167AFD28D32B62BDA45F" box="[717,909,1831,1855]" italics="true" pageId="4" pageNumber="129">P. longiflagellum</emphasis>
</taxonomicName>
(in
<collectingCountry id="F33F4AC5B751167AFC59D32B6513A420" box="[956,1059,1832,1856]" name="Morocco" pageId="4" pageNumber="129">Morocco</collectingCountry>
) and
<taxonomicName id="4C2871D6B751167AFB86D32B65D9A420" authorityName="Chevreux" authorityYear="1901" box="[1123,1257,1832,1856]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="129" phylum="Arthropoda" rank="species" species="africanus">
<emphasis id="B95CD647B751167AFB86D32B65D9A420" box="[1123,1257,1832,1856]" italics="true" pageId="4" pageNumber="129">P. africanus</emphasis>
</taxonomicName>
(in
<collectingCountry id="F33F4AC5B751167AFAFDD324645CA420" box="[1304,1388,1831,1856]" name="Algeria" pageId="4" pageNumber="129">Algeria</collectingCountry>
and
<collectingCountry id="F33F4AC5B751167AFF72D34861DCA404" box="[151,236,1867,1892]" name="Tunisia" pageId="4" pageNumber="129">Tunisia</collectingCountry>
) seem to have a relatively wide distribution (
<figureCitation id="131316D0B751167AFD04D3486215A404" box="[737,805,1867,1892]" captionStart="FIGURE 6" captionStartId="12.[151,250,1256,1278]" captionTargetBox="[157,1399,200,1228]" captionTargetId="figure@12.[150,1436,193,1235]" captionTargetPageId="12" captionText="FIGURE 6. Distribution of African species of Pseudoniphargus species." httpUri="https://zenodo.org/record/1440520/files/figure.png" pageId="4" pageNumber="129">Fig. 6</figureCitation>
). However, very few samples have been collected at this part of the world. Consequently, without more intensive sampling, it is still hazardous to conclude definitively on these aspects, considering also that some misidentifications may have occured. For example, the wide distribution of
<taxonomicName id="4C2871D6B751167AFF51D3B46008A4AF" authorityName="Chevreux" authorityYear="1901" box="[180,312,1975,1999]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="129" phylum="Arthropoda" rank="species" species="africanus">
<emphasis id="B95CD647B751167AFF51D3B46008A4AF" box="[180,312,1975,1999]" italics="true" pageId="4" pageNumber="129">P. africanus</emphasis>
</taxonomicName>
, previously reported to occur in the
<collectingRegion id="49ECC4B7B751167AFD29D3B562B1A4AF" box="[716,897,1974,1999]" country="Spain" name="Illes Baleares" pageId="4" pageNumber="129">Balearic Islands</collectingRegion>
,
<collectingCountry id="F33F4AC5B751167AFC69D3B562D2A4AF" box="[908,994,1974,1999]" name="Croatia" pageId="4" pageNumber="129">Croatia</collectingCountry>
,
<collectingCountry id="F33F4AC5B751167AFC09D3B4650BA4AE" box="[1004,1083,1975,1998]" name="France" pageId="4" pageNumber="129">France</collectingCountry>
,
<collectingCountry id="F33F4AC5B751167AFBA0D3B56549A4AF" box="[1093,1145,1974,1999]" name="Italy" pageId="4" pageNumber="129">Italy</collectingCountry>
,
<collectingRegion id="49ECC4B7B751167AFB61D3B565F7A4AF" box="[1156,1223,1974,1999]" country="Italy" name="Sicilia" pageId="4" pageNumber="129">Sicily</collectingRegion>
, and
<collectingCountry id="F33F4AC5B751167AFAE5D3B564B0A4AF" box="[1280,1408,1974,1999]" name="Serbia" pageId="4" pageNumber="129">Yugoslavia</collectingCountry>
, is now considered to be reduced to
<collectingCountry id="F33F4AC5B751167AFDECD3D96350A493" box="[521,608,2010,2035]" name="Algeria" pageId="4" pageNumber="129">Algeria</collectingCountry>
, from where it was first described (
<bibRefCitation id="EFB977A4B751167AFC0ED3D965E6A493" box="[1003,1238,2010,2035]" pageId="4" pageNumber="129" refId="ref8745" refString="Messouli, M., Messana, G. &amp; Yacoubi-Kebhiza, M. (2006) Three new species of Pseudoniphargus (Amphipoda), from the groundwater of three Mediterranean islands, with notes on the Ps. adriaticus. Subterranean Biology, 4, 79 - 101." type="journal article">
Messouli
<emphasis id="B95CD647B751167AFBBFD3D865A2A492" box="[1114,1170,2010,2034]" italics="true" pageId="4" pageNumber="129">et al.</emphasis>
2006
</bibRefCitation>
).
</paragraph>
<paragraph id="8B970A55B750167BFF76D66661CFA11B" blockId="5.[147,255,613,635]" box="[147,255,613,635]" pageId="5" pageNumber="130">
<emphasis id="B95CD647B750167BFF76D66661CFA11B" bold="true" box="[147,255,613,635]" pageId="5" pageNumber="130">
<tableCitation id="C6AA3FEEB750167BFF76D66661C9A11B" box="[147,249,613,635]" captionStart="TABLE 1" captionStartId="6.[147,230,617,639]" captionText="TABLE 1. (Continued)" pageId="5" pageNumber="130">TABLE 1</tableCitation>
.
</emphasis>
</paragraph>
<paragraph id="8B970A55B750167BFECAD6B067CBA7C5" pageId="5" pageNumber="130">
<table id="F928F8F5B750E981FF45D6846663A7C6" box="[160,1875,647,1190]" gridcols="10" gridrows="6" pageId="5" pageNumber="130">
<tr id="35180817B750E981FF45D6846663A1A6" box="[160,1875,647,710]" gridrow="0" pageId="5" pageNumber="130" rowspan-0="1">
<th id="76C9616BB750E981FECBD6846348A1A6" box="[302,632,647,710]" gridcol="1" gridrow="0" pageId="5" pageNumber="130">
<taxonomicName id="4C2871D6B750167BFECAD6B060FDA1A6" authorityName="Dole-Olivier &amp; Hafid &amp; Piscart" authorityYear="2018" box="[303,461,691,710]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="130" phylum="Arthropoda" rank="genus">
<emphasis id="B95CD647B750167BFECAD6B060FDA1A6" box="[303,461,691,710]" italics="true" pageId="5" pageNumber="130">Pseudoniphargus</emphasis>
</taxonomicName>
species --&gt;
</th>
<th id="76C9616BB750E981FD30D6846544A1A6" box="[725,1140,647,710]" colspan="3" colspanRight="2" gridcol="2" gridrow="0" pageId="5" pageNumber="130">
<emphasis id="B95CD647B750167BFD30D6846526A1FB" bold="true" box="[725,1046,647,667]" pageId="5" pageNumber="130">Species from Algeria and Tunisia</emphasis>
<emphasis id="B95CD647B750167BFD30D6B06544A1A6" box="[725,1140,691,710]" italics="true" pageId="5" pageNumber="130">
<taxonomicName id="4C2871D6B750167BFD30D6B06222A1A6" authorityName="Dole-Olivier &amp; Hafid &amp; Piscart" authorityYear="2018" box="[725,786,691,710]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="djemoi">djemoi</taxonomicName>
<taxonomicName id="4C2871D6B750167BFCB0D6B0629AA1A6" authorityName="Chevreux" authorityYear="1901" box="[853,938,691,710]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="africanus">africanus</taxonomicName>
<taxonomicName id="4C2871D6B750167BFC13D6B06544A1A6" authorityName="Stock" authorityYear="1980" box="[1014,1140,691,710]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="macrotelsonis">macrotelsonis</taxonomicName>
</emphasis>
</th>
<th id="76C9616BB750E981FB40D68464F9A1A6" box="[1189,1481,647,710]" colspan="2" colspanRight="1" gridcol="5" gridrow="0" pageId="5" pageNumber="130">
<emphasis id="B95CD647B750167BFB40D684644BA1FB" bold="true" box="[1189,1403,647,667]" pageId="5" pageNumber="130">Species from Morocco</emphasis>
<emphasis id="B95CD647B750167BFB40D6B064F9A1A5" box="[1189,1481,691,710]" italics="true" pageId="5" pageNumber="130">
<taxonomicName id="4C2871D6B750167BFB40D6B06419A1A6" authorityName="Fakher-el-Abiari, Oulbaz, Messouli &amp; Coineau" authorityYear="1999" box="[1189,1321,691,710]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="longiflagellum">longiflagellum</taxonomicName>
<taxonomicName id="4C2871D6B750167BFABED6B064F9A1A5" authorityName="Boutin &amp; Coineau" authorityYear="1988" box="[1371,1481,691,709]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="maroccanus">maroccanus</taxonomicName>
</emphasis>
</th>
<th id="76C9616BB750E981FA08D684677AA1A6" box="[1517,1610,647,710]" gridcol="7" gridrow="0" pageId="5" pageNumber="130">
<taxonomicName id="4C2871D6B750167BFA08D6B0672FA1A6" authorityName="Coineau &amp; Boutin" authorityYear="1996" box="[1517,1567,691,710]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="ruffoi">
<emphasis id="B95CD647B750167BFA08D6B0672FA1A6" box="[1517,1567,691,710]" italics="true" pageId="5" pageNumber="130">ruffoi</emphasis>
</taxonomicName>
</th>
<th id="76C9616BB750E981F98BD684678BA1A6" box="[1646,1723,647,710]" gridcol="8" gridrow="0" pageId="5" pageNumber="130">
<taxonomicName id="4C2871D6B750167BF98BD6B0678BA1A6" authorityName="Coineau &amp; Boutin" authorityYear="1996" box="[1646,1723,691,710]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="longipes">
<emphasis id="B95CD647B750167BF98BD6B0678BA1A6" box="[1646,1723,691,710]" italics="true" pageId="5" pageNumber="130">longipes</emphasis>
</taxonomicName>
</th>
<th id="76C9616BB750E981F938D6846663A1A6" box="[1757,1875,647,710]" gridcol="9" gridrow="0" pageId="5" pageNumber="130">
<taxonomicName id="4C2871D6B750167BF90DD6B06663A1A5" authorityName="Coineau &amp; Boutin" authorityYear="1996" box="[1768,1875,691,709]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="romanorum">
<emphasis id="B95CD647B750167BF90DD6B06663A1A5" box="[1768,1875,691,709]" italics="true" pageId="5" pageNumber="130">romanorum</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="35180817B750E981FF45D6DD6663A031" box="[160,1875,734,849]" gridrow="1" pageId="5" pageNumber="130">
<th id="76C9616BB750E981FF45D6DD61CDA031" box="[160,253,734,849]" gridcol="0" gridrow="1" pageId="5" pageNumber="130">Antenna 1 Antenna 2</th>
<td id="76C9616BB750E981FECBD6DD6348A031" box="[302,632,734,849]" gridcol="1" gridrow="1" pageId="5" pageNumber="130">flagellum: number of articles accessory flagellum longer than first flagellar article flagellum:number of articles</td>
<td id="76C9616BB750E981FD30D6DD6224A031" box="[725,788,734,849]" gridcol="2" gridrow="1" pageId="5" pageNumber="130">14-15 NO 6</td>
<td id="76C9616BB750E981FCAFD6DD62BBA031" box="[842,907,734,849]" gridcol="3" gridrow="1" pageId="5" pageNumber="130">18 NO 7</td>
<td id="76C9616BB750E981FC0FD6DD6544A031" box="[1002,1140,734,849]" gridcol="4" gridrow="1" pageId="5" pageNumber="130">18 NO 7</td>
<td id="76C9616BB750E981FB40D6DD65D6A031" box="[1189,1254,734,849]" gridcol="5" gridrow="1" pageId="5" pageNumber="130">
<emphasis id="B95CD647B750167BFB40D6DC65FEA031" bold="true" pageId="5" pageNumber="130">32-41 YES 9-13</emphasis>
</td>
<td id="76C9616BB750E981FAAAD6DD64F9A031" box="[1359,1481,734,849]" gridcol="6" gridrow="1" pageId="5" pageNumber="130">
16-18
<emphasis id="B95CD647B750167BFAAAD709644BA07E" bold="true" box="[1359,1403,778,798]" pageId="5" pageNumber="130">YES</emphasis>
6-7
</td>
<td id="76C9616BB750E981FA08D6DD677AA031" box="[1517,1610,734,849]" gridcol="7" gridrow="1" pageId="5" pageNumber="130">
<emphasis id="B95CD647B750167BFA08D6DC6734A031" bold="true" pageId="5" pageNumber="130">23-25 YES (6) 11</emphasis>
</td>
<td id="76C9616BB750E981F98BD6DD678BA031" box="[1646,1723,734,849]" gridcol="8" gridrow="1" pageId="5" pageNumber="130">
<emphasis id="B95CD647B750167BF98BD6DC6792A193" bold="true" box="[1646,1698,735,755]" pageId="5" pageNumber="130">22-25</emphasis>
NO
<emphasis id="B95CD647B750167BF98BD73E6793A031" bold="true" box="[1646,1699,829,849]" pageId="5" pageNumber="130">11-14</emphasis>
</td>
<td id="76C9616BB750E981F938D6DD6663A031" box="[1757,1875,734,849]" gridcol="9" gridrow="1" pageId="5" pageNumber="130">
16-20
<emphasis id="B95CD647B750167BF938D7096604A07E" bold="true" box="[1757,1844,778,798]" pageId="5" pageNumber="130">YES (6)</emphasis>
10
</td>
</tr>
<tr id="35180817B750E981FF45D7646663A01C" box="[160,1875,871,892]" gridrow="2" pageId="5" pageNumber="130">
<th id="76C9616BB750E981FF45D76461CDA01C" box="[160,253,871,892]" gridcol="0" gridrow="2" pageId="5" pageNumber="130">P4</th>
<td id="76C9616BB750E981FECBD7646348A01C" box="[302,632,871,892]" gridcol="1" gridrow="2" pageId="5" pageNumber="130">coxal plate posterior excavation</td>
<td id="76C9616BB750E981FD30D7646224A01C" box="[725,788,871,892]" gridcol="2" gridrow="2" pageId="5" pageNumber="130">++</td>
<td id="76C9616BB750E981FCAFD76462BBA01C" box="[842,907,871,892]" gridcol="3" gridrow="2" pageId="5" pageNumber="130">+</td>
<td id="76C9616BB750E981FC0FD7646544A01C" box="[1002,1140,871,892]" gridcol="4" gridrow="2" pageId="5" pageNumber="130">+</td>
<td id="76C9616BB750E981FB40D76465D6A01C" box="[1189,1254,871,892]" gridcol="5" gridrow="2" pageId="5" pageNumber="130">
<emphasis id="B95CD647B750167BFB55D76B65E6A01C" bold="true" box="[1200,1238,872,892]" pageId="5" pageNumber="130">+++</emphasis>
</td>
<td id="76C9616BB750E981FAAAD76464F9A01C" box="[1359,1481,871,892]" gridcol="6" gridrow="2" pageId="5" pageNumber="130">+</td>
<td id="76C9616BB750E981FA08D764677AA01C" box="[1517,1610,871,892]" gridcol="7" gridrow="2" pageId="5" pageNumber="130">++</td>
<td id="76C9616BB750E981F98BD764678BA01C" box="[1646,1723,871,892]" gridcol="8" gridrow="2" pageId="5" pageNumber="130">++</td>
<td id="76C9616BB750E981F938D7646663A01C" box="[1757,1875,871,892]" gridcol="9" gridrow="2" pageId="5" pageNumber="130">++</td>
</tr>
<tr id="35180817B750E981FF45D7916663A09C" box="[160,1875,914,1020]" gridrow="3" pageId="5" pageNumber="130">
<th id="76C9616BB750E981FF45D79161CDA09C" box="[160,253,914,1020]" gridcol="0" gridrow="3" pageId="5" pageNumber="130">P7</th>
<td id="76C9616BB750E981FECBD7916348A09C" box="[302,632,914,1020]" gridcol="1" gridrow="3" pageId="5" pageNumber="130">basis elongated vs wide basis postero distal lobe Propodus length/width ratio</td>
<td id="76C9616BB750E981FD30D7916224A09C" box="[725,788,914,1020]" gridcol="2" gridrow="3" pageId="5" pageNumber="130">
<emphasis id="B95CD647B750167BFD30D7906224A0B1" bold="true" box="[725,788,915,977]" pageId="5" pageNumber="130">wide absent</emphasis>
6.7
</td>
<td id="76C9616BB750E981FCAFD79162BBA09C" box="[842,907,914,1020]" gridcol="3" gridrow="3" pageId="5" pageNumber="130">
<emphasis id="B95CD647B750167BFCAFD7906247A0C7" bold="true" box="[842,887,915,935]" pageId="5" pageNumber="130">wide</emphasis>
present 7.6
</td>
<td id="76C9616BB750E981FC0FD7916544A09C" box="[1002,1140,914,1020]" gridcol="4" gridrow="3" pageId="5" pageNumber="130">
<emphasis id="B95CD647B750167BFC0ED7906528A0C7" bold="true" box="[1003,1048,915,935]" pageId="5" pageNumber="130">wide</emphasis>
present 7.6
</td>
<td id="76C9616BB750E981FB40D79165D6A09C" box="[1189,1254,914,1020]" gridcol="5" gridrow="3" pageId="5" pageNumber="130">
<emphasis id="B95CD647B750167BFB40D79065E2A0C7" bold="true" box="[1189,1234,915,935]" pageId="5" pageNumber="130">wide</emphasis>
present 8.8
</td>
<td id="76C9616BB750E981FAAAD79164F9A09C" box="[1359,1481,914,1020]" gridcol="6" gridrow="3" pageId="5" pageNumber="130">
elongated
<emphasis id="B95CD647B750167BFAAAD7BE64BEA0B1" bold="true" box="[1359,1422,957,977]" pageId="5" pageNumber="130">absent</emphasis>
♀8.6-ϐ9.4
</td>
<td id="76C9616BB750E981FA08D791677AA09C" box="[1517,1610,914,1020]" gridcol="7" gridrow="3" pageId="5" pageNumber="130">
<emphasis id="B95CD647B750167BFA08D790672AA0C7" bold="true" box="[1517,1562,915,935]" pageId="5" pageNumber="130">wide</emphasis>
present 10.2
</td>
<td id="76C9616BB750E981F98BD791678BA09C" box="[1646,1723,914,1020]" gridcol="8" gridrow="3" pageId="5" pageNumber="130">
<emphasis id="B95CD647B750167BF98BD79067ABA0C7" bold="true" box="[1646,1691,915,935]" pageId="5" pageNumber="130">wide</emphasis>
present
<emphasis id="B95CD647B750167BF98BD7EB67A5A09C" bold="true" box="[1646,1685,1000,1020]" pageId="5" pageNumber="130">15.8</emphasis>
</td>
<td id="76C9616BB750E981F938D7916663A09C" box="[1757,1875,914,1020]" gridcol="9" gridrow="3" pageId="5" pageNumber="130">
<emphasis id="B95CD647B750167BF938D790663AA0C7" bold="true" box="[1757,1802,915,935]" pageId="5" pageNumber="130">wide</emphasis>
present 10.6
</td>
</tr>
<tr id="35180817B750E981FF45D0116663A71C" box="[160,1875,1042,1148]" gridrow="4" pageId="5" pageNumber="130">
<th id="76C9616BB750E981FF45D01161CDA71C" box="[160,253,1042,1148]" gridcol="0" gridrow="4" pageId="5" pageNumber="130">Uropod 1</th>
<td id="76C9616BB750E981FECBD0116348A71C" box="[302,632,1042,1148]" gridcol="1" gridrow="4" pageId="5" pageNumber="130">pedunculus basoventral spine pedunculus inner medial spine lateral spines on rami</td>
<td id="76C9616BB750E981FD30D0116224A71C" box="[725,788,1042,1148]" gridcol="2" gridrow="4" pageId="5" pageNumber="130">
<emphasis id="B95CD647B750167BFD30D01063D1A747" bold="true" box="[725,737,1043,1063]" pageId="5" pageNumber="130">0</emphasis>
0 0
</td>
<td id="76C9616BB750E981FCAFD01162BBA71C" box="[842,907,1042,1148]" gridcol="3" gridrow="4" pageId="5" pageNumber="130">
1 0
<emphasis id="B95CD647B750167BFCAFD06B6258A71C" bold="true" box="[842,872,1128,1148]" pageId="5" pageNumber="130">0-1</emphasis>
</td>
<td id="76C9616BB750E981FC0FD0116544A71C" box="[1002,1140,1042,1148]" gridcol="4" gridrow="4" pageId="5" pageNumber="130">
<emphasis id="B95CD647B750167BFC0ED01062C7A731" bold="true" box="[1003,1015,1043,1105]" pageId="5" pageNumber="130">0 1</emphasis>
0
</td>
<td id="76C9616BB750E981FB40D01165D6A71C" box="[1189,1254,1042,1148]" gridcol="5" gridrow="4" pageId="5" pageNumber="130">
1
<emphasis id="B95CD647B750167BFB40D03E65F3A71C" bold="true" box="[1189,1219,1085,1148]" pageId="5" pageNumber="130">2 0-1</emphasis>
</td>
<td id="76C9616BB750E981FAAAD01164F9A71C" box="[1359,1481,1042,1148]" gridcol="6" gridrow="4" pageId="5" pageNumber="130">
1
<emphasis id="B95CD647B750167BFAAAD03E646AA71C" bold="true" box="[1359,1371,1085,1148]" pageId="5" pageNumber="130">1 1</emphasis>
</td>
<td id="76C9616BB750E981FA08D011677AA71C" box="[1517,1610,1042,1148]" gridcol="7" gridrow="4" pageId="5" pageNumber="130">
1
<emphasis id="B95CD647B750167BFA08D03E673BA71C" bold="true" box="[1517,1547,1085,1148]" pageId="5" pageNumber="130">4 1-2</emphasis>
</td>
<td id="76C9616BB750E981F98BD011678BA71C" box="[1646,1723,1042,1148]" gridcol="8" gridrow="4" pageId="5" pageNumber="130">
1
<emphasis id="B95CD647B750167BF98BD03E6749A71C" bold="true" box="[1646,1657,1085,1148]" pageId="5" pageNumber="130">3 1</emphasis>
</td>
<td id="76C9616BB750E981F938D0116663A71C" box="[1757,1875,1042,1148]" gridcol="9" gridrow="4" pageId="5" pageNumber="130">
1 0
<emphasis id="B95CD647B750167BF938D06B67D8A71C" bold="true" box="[1757,1768,1128,1148]" pageId="5" pageNumber="130">1</emphasis>
</td>
</tr>
<tr id="35180817B750E981FF45D0916663A7C6" box="[160,1875,1170,1190]" gridrow="5" pageId="5" pageNumber="130">
<th id="76C9616BB750E981FF45D09161CDA7C6" box="[160,253,1170,1190]" gridcol="0" gridrow="5" pageId="5" pageNumber="130">Uropod 2</th>
<td id="76C9616BB750E981FECBD0916348A7C6" box="[302,632,1170,1190]" gridcol="1" gridrow="5" pageId="5" pageNumber="130">lateral spines on rami</td>
<td id="76C9616BB750E981FD30D0916224A7C6" box="[725,788,1170,1190]" gridcol="2" gridrow="5" pageId="5" pageNumber="130">0</td>
<td id="76C9616BB750E981FCAFD09162BBA7C6" box="[842,907,1170,1190]" gridcol="3" gridrow="5" pageId="5" pageNumber="130">0</td>
<td id="76C9616BB750E981FC0FD0916544A7C6" box="[1002,1140,1170,1190]" gridcol="4" gridrow="5" pageId="5" pageNumber="130">0</td>
<td id="76C9616BB750E981FB40D09165D6A7C6" box="[1189,1254,1170,1190]" gridcol="5" gridrow="5" pageId="5" pageNumber="130">1-3</td>
<td id="76C9616BB750E981FAAAD09164F9A7C6" box="[1359,1481,1170,1190]" gridcol="6" gridrow="5" pageId="5" pageNumber="130">1</td>
<td id="76C9616BB750E981FA08D091677AA7C6" box="[1517,1610,1170,1190]" gridcol="7" gridrow="5" pageId="5" pageNumber="130">0-1</td>
<td id="76C9616BB750E981F98BD091678BA7C6" box="[1646,1723,1170,1190]" gridcol="8" gridrow="5" pageId="5" pageNumber="130">0-1</td>
<td id="76C9616BB750E981F938D0916663A7C6" box="[1757,1875,1170,1190]" gridcol="9" gridrow="5" pageId="5" pageNumber="130">0-1</td>
</tr>
</table>
</paragraph>
<paragraph id="8B970A55B750167BFF45D0BF6611A7B0" blockId="5.[160,1825,1212,1361]" box="[160,1825,1212,1233]" pageId="5" pageNumber="130">
Uropod 3 pedunculus elongated NO
<emphasis id="B95CD647B750167BFCAFD0BE6527A7B1" bold="true" box="[842,1047,1213,1233]" pageId="5" pageNumber="130">YES YES</emphasis>
slightly slightly NO slightly slightly
</paragraph>
<paragraph id="8B970A55B750167BFEDFD0E46639A79C" blockId="5.[160,1825,1212,1361]" box="[314,1801,1255,1276]" pageId="5" pageNumber="130">
lengh: sexual dimorphism? moderate
<emphasis id="B95CD647B750167BFCAFD0EB6527A79C" bold="true" box="[842,1047,1256,1276]" pageId="5" pageNumber="130">YES YES</emphasis>
? NO NO moderate
<emphasis id="B95CD647B750167BF938D0EB6639A79C" bold="true" box="[1757,1801,1256,1276]" pageId="5" pageNumber="130">YES</emphasis>
</paragraph>
<paragraph id="8B970A55B750167BFECAD11067D8A646" blockId="5.[160,1825,1212,1361]" box="[303,1768,1298,1318]" pageId="5" pageNumber="130">
groups of exopod marginal spines
<emphasis id="B95CD647B750167BFD30D111658CA646" bold="true" box="[725,1212,1298,1318]" pageId="5" pageNumber="130">
9-12 5-
<date id="FF962C95B750167BFCBED111658CA646" box="[859,1212,1298,1318]" pageId="5" pageNumber="130" value="2014-05-12" valueMax="2014-12-12" valueMin="2014-05-12">12 5-12 14</date>
</emphasis>
5-
<date id="FF962C95B750167BFA84D11167B5A645" box="[1377,1669,1298,1318]" pageId="5" pageNumber="130" value="2010-09-06" valueMax="2010-10-06" valueMin="2010-09-06">6 9-10 10</date>
7
</paragraph>
<paragraph id="8B970A55B750167BFECAD13F67D9A631" blockId="5.[160,1825,1212,1361]" box="[303,1769,1340,1361]" pageId="5" pageNumber="130">
number of distal spines on endopod
<emphasis id="B95CD647B750167BFD30D13E6581A631" bold="true" box="[725,1201,1341,1361]" pageId="5" pageNumber="130">1 1 1 1</emphasis>
1-2 2 2
<emphasis id="B95CD647B750167BF938D13E67D9A631" bold="true" box="[1757,1769,1341,1361]" pageId="5" pageNumber="130">1</emphasis>
</paragraph>
<paragraph id="8B970A55B750167BFF45D1646672A69B" pageId="5" pageNumber="130">
<table id="F928F8F5B750E981FF45D1646672A69B" box="[160,1858,1383,1531]" gridcols="11" gridrows="4" pageId="5" pageNumber="130">
<tr id="35180817B750E981FF45D1646672A61B" box="[160,1858,1383,1403]" gridrow="0" pageId="5" pageNumber="130">
<th id="76C9616BB750E981FF45D16461DCA61B" box="[160,236,1383,1403]" gridcol="0" gridrow="0" pageId="5" pageNumber="130">Telson</th>
<th id="76C9616BB750E981FECBD1646357A61B" box="[302,615,1383,1403]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="5" pageNumber="130">distal spines on each lobe</th>
<th id="76C9616BB750E981FD30D1646229A61B" box="[725,793,1383,1403]" gridcol="3" gridrow="0" pageId="5" pageNumber="130">ϐ2, ♀3</th>
<th id="76C9616BB750E981FCAFD16462F1A61B" box="[842,961,1383,1403]" gridcol="4" gridrow="0" pageId="5" pageNumber="130">
<emphasis id="B95CD647B750167BFCAFD1646258A61B" bold="true" box="[842,872,1383,1403]" pageId="5" pageNumber="130">2-4</emphasis>
</th>
<th id="76C9616BB750E981FC0FD1646579A61B" box="[1002,1097,1383,1403]" gridcol="5" gridrow="0" pageId="5" pageNumber="130">
<emphasis id="B95CD647B750167BFC0ED1646539A61B" bold="true" box="[1003,1033,1383,1403]" pageId="5" pageNumber="130">2-4</emphasis>
</th>
<th id="76C9616BB750E981FB40D16465D5A61B" box="[1189,1253,1383,1403]" gridcol="6" gridrow="0" pageId="5" pageNumber="130">
<emphasis id="B95CD647B750167BFB40D1646581A61B" bold="true" box="[1189,1201,1383,1403]" pageId="5" pageNumber="130">3</emphasis>
</th>
<th id="76C9616BB750E981FAAAD16464A4A61B" box="[1359,1428,1383,1403]" gridcol="7" gridrow="0" pageId="5" pageNumber="130">
<emphasis id="B95CD647B750167BFAAAD16464A4A61B" bold="true" box="[1359,1428,1383,1403]" pageId="5" pageNumber="130">64, ♀5</emphasis>
</th>
<th id="76C9616BB750E981FA08D1646717A61B" box="[1517,1575,1383,1403]" gridcol="8" gridrow="0" pageId="5" pageNumber="130">
<emphasis id="B95CD647B750167BFA08D16464C9A61B" bold="true" box="[1517,1529,1383,1403]" pageId="5" pageNumber="130">3</emphasis>
</th>
<th id="76C9616BB750E981F98BD1646798A61B" box="[1646,1704,1383,1403]" gridcol="9" gridrow="0" pageId="5" pageNumber="130">
<emphasis id="B95CD647B750167BF98BD164674AA61B" bold="true" box="[1646,1658,1383,1403]" pageId="5" pageNumber="130">3</emphasis>
</th>
<th id="76C9616BB750E981F938D1646672A61B" box="[1757,1858,1383,1403]" gridcol="10" gridrow="0" pageId="5" pageNumber="130">2</th>
</tr>
<tr id="35180817B750E981FF45D1926672A6C6" box="[160,1858,1425,1446]" gridrow="1" pageId="5" pageNumber="130" rowspan-0="1" rowspan-2="1">
<td id="76C9616BB750E981FECBD19260D4A6C6" box="[302,484,1425,1446]" gridcol="1" gridrow="1" pageId="5" pageNumber="130">shape</td>
<td id="76C9616BB750E981FD30D1926229A6C6" box="[725,793,1425,1446]" gridcol="3" gridrow="1" pageId="5" pageNumber="130">
<emphasis id="B95CD647B750167BFD30D191623FA6C6" bold="true" box="[725,783,1426,1446]" pageId="5" pageNumber="130">broad</emphasis>
</td>
<td id="76C9616BB750E981FCAFD19262F1A6C6" box="[842,961,1425,1446]" gridcol="4" gridrow="1" pageId="5" pageNumber="130">quadrangular</td>
<td id="76C9616BB750E981FC0FD1926579A6C6" box="[1002,1097,1425,1446]" gridcol="5" gridrow="1" pageId="5" pageNumber="130">elongated</td>
<td id="76C9616BB750E981FB40D19265D5A6C6" box="[1189,1253,1425,1446]" gridcol="6" gridrow="1" pageId="5" pageNumber="130">
<emphasis id="B95CD647B750167BFB40D19165EFA6C6" bold="true" box="[1189,1247,1426,1446]" pageId="5" pageNumber="130">broad</emphasis>
</td>
<td id="76C9616BB750E981FAAAD19264A4A6C6" box="[1359,1428,1425,1446]" gridcol="7" gridrow="1" pageId="5" pageNumber="130">
<emphasis id="B95CD647B750167BFAAAD19164B9A6C6" bold="true" box="[1359,1417,1426,1446]" pageId="5" pageNumber="130">broad</emphasis>
</td>
<td id="76C9616BB750E981FA08D1926717A6C6" box="[1517,1575,1425,1446]" gridcol="8" gridrow="1" pageId="5" pageNumber="130">
<emphasis id="B95CD647B750167BFA08D1916717A6C6" bold="true" box="[1517,1575,1426,1446]" pageId="5" pageNumber="130">broad</emphasis>
</td>
<td id="76C9616BB750E981F98BD1926798A6C6" box="[1646,1704,1425,1446]" gridcol="9" gridrow="1" pageId="5" pageNumber="130">
<emphasis id="B95CD647B750167BF98BD1916798A6C6" bold="true" box="[1646,1704,1426,1446]" pageId="5" pageNumber="130">broad</emphasis>
</td>
<td id="76C9616BB750E981F938D1926672A6C6" box="[1757,1858,1425,1446]" gridcol="10" gridrow="1" pageId="5" pageNumber="130">
<emphasis id="B95CD647B750167BF938D1916627A6C6" bold="true" box="[1757,1815,1426,1446]" pageId="5" pageNumber="130">broad</emphasis>
</td>
</tr>
<tr id="35180817B750E981FF45D1BF6672A6B0" box="[160,1858,1468,1488]" gridrow="2" pageId="5" pageNumber="130" rowspan-0="1" rowspan-2="1">
<td id="76C9616BB750E981FECBD1BF60D4A6B0" box="[302,484,1468,1488]" gridcol="1" gridrow="2" pageId="5" pageNumber="130">mediodistal notch</td>
<td id="76C9616BB750E981FD30D1BF6229A6B0" box="[725,793,1468,1488]" gridcol="3" gridrow="2" pageId="5" pageNumber="130">
<emphasis id="B95CD647B750167BFD04D1BF63DEA6B0" bold="true" box="[737,750,1468,1488]" pageId="5" pageNumber="130">+</emphasis>
</td>
<td id="76C9616BB750E981FCAFD1BF62F1A6B0" box="[842,961,1468,1488]" gridcol="4" gridrow="2" pageId="5" pageNumber="130">+++</td>
<td id="76C9616BB750E981FC0FD1BF6579A6B0" box="[1002,1097,1468,1488]" gridcol="5" gridrow="2" pageId="5" pageNumber="130">+++</td>
<td id="76C9616BB750E981FB40D1BF65D5A6B0" box="[1189,1253,1468,1488]" gridcol="6" gridrow="2" pageId="5" pageNumber="130">++</td>
<td id="76C9616BB750E981FAAAD1BF64A4A6B0" box="[1359,1428,1468,1488]" gridcol="7" gridrow="2" pageId="5" pageNumber="130">++</td>
<td id="76C9616BB750E981FA08D1BF6717A6B0" box="[1517,1575,1468,1488]" gridcol="8" gridrow="2" pageId="5" pageNumber="130">
<emphasis id="B95CD647B750167BFA1DD1BF6735A6B0" bold="true" box="[1528,1541,1468,1488]" pageId="5" pageNumber="130">+</emphasis>
</td>
<td id="76C9616BB750E981F98BD1BF6798A6B0" box="[1646,1704,1468,1488]" gridcol="9" gridrow="2" pageId="5" pageNumber="130">++</td>
<td id="76C9616BB750E981F938D1BF6672A6B0" box="[1757,1858,1468,1488]" gridcol="10" gridrow="2" pageId="5" pageNumber="130">++</td>
</tr>
<tr id="35180817B750E981FF45D1E46672A69B" box="[160,1858,1511,1531]" gridrow="3" pageId="5" pageNumber="130">
<th id="76C9616BB750E981FF45D1E461DCA69B" box="[160,236,1511,1531]" gridcol="0" gridrow="3" pageId="5" pageNumber="130">Epimera</th>
<td id="76C9616BB750E981FECBD1E460D4A69B" box="[302,484,1511,1531]" gridcol="1" gridrow="3" pageId="5" pageNumber="130">ventral spines (from</td>
<td id="76C9616BB750E981FE1ED1E46357A69B" box="[507,615,1511,1531]" gridcol="2" gridrow="3" pageId="5" pageNumber="130">plate 1 to 3)</td>
<td id="76C9616BB750E981FD30D1E46229A69B" box="[725,793,1511,1531]" gridcol="3" gridrow="3" pageId="5" pageNumber="130">
<emphasis id="B95CD647B750167BFD30D1E46239A69B" bold="true" box="[725,777,1511,1531]" pageId="5" pageNumber="130">1, 3,3</emphasis>
</td>
<td id="76C9616BB750E981FCAFD1E462F1A69B" box="[842,961,1511,1531]" gridcol="4" gridrow="3" pageId="5" pageNumber="130">0, 1, 1</td>
<td id="76C9616BB750E981FC0FD1E46579A69B" box="[1002,1097,1511,1531]" gridcol="5" gridrow="3" pageId="5" pageNumber="130">0, 1-2, 1-2</td>
<td id="76C9616BB750E981FB40D1E465D5A69B" box="[1189,1253,1511,1531]" gridcol="6" gridrow="3" pageId="5" pageNumber="130">
<emphasis id="B95CD647B750167BFB40D1E465D5A69B" bold="true" box="[1189,1253,1511,1531]" pageId="5" pageNumber="130">2-4,3,3</emphasis>
</td>
<td id="76C9616BB750E981FAAAD1E464A4A69B" box="[1359,1428,1511,1531]" gridcol="7" gridrow="3" pageId="5" pageNumber="130">0, 1,1</td>
<td id="76C9616BB750E981FA08D1E46717A69B" box="[1517,1575,1511,1531]" gridcol="8" gridrow="3" pageId="5" pageNumber="130">
<emphasis id="B95CD647B750167BFA08D1E46710A69B" bold="true" box="[1517,1568,1511,1531]" pageId="5" pageNumber="130">1, 4,4</emphasis>
</td>
<td id="76C9616BB750E981F98BD1E46798A69B" box="[1646,1704,1511,1531]" gridcol="9" gridrow="3" pageId="5" pageNumber="130">
<emphasis id="B95CD647B750167BF98BD1E46796A69B" bold="true" box="[1646,1702,1511,1531]" pageId="5" pageNumber="130">1, 4, 5</emphasis>
</td>
<td id="76C9616BB750E981F938D1E46672A69B" box="[1757,1858,1511,1531]" gridcol="10" gridrow="3" pageId="5" pageNumber="130">(ϐ): 1, 4, 4</td>
</tr>
</table>
</paragraph>
<paragraph id="8B970A55B7501678F9F0D21A6659A554" blockId="5.[1557,1896,1561,1583]" lastBlockId="6.[1557,1897,1566,1588]" lastPageId="6" lastPageNumber="131" pageId="5" pageNumber="130">
<emphasis id="B95CD647B7501678F9F0D21A6659A554" italics="true" lastPageId="6" lastPageNumber="131" pageId="5" pageNumber="130">……continued on the next page ……continued on the next page</emphasis>
</paragraph>
</subSubSection>
<caption id="DF575ADDB7531678FF76D66A604EA11F" box="[147,382,617,639]" pageId="6" pageNumber="131">
<paragraph id="8B970A55B7531678FF76D66A604EA11F" blockId="6.[147,382,617,639]" box="[147,382,617,639]" pageId="6" pageNumber="131">
<emphasis id="B95CD647B7531678FF76D66A61CFA11F" bold="true" box="[147,255,617,639]" pageId="6" pageNumber="131">TABLE 1.</emphasis>
(Continued)
</paragraph>
</caption>
<paragraph id="8B970A55B7531678FEFED6B46673A569" pageId="6" pageNumber="131">
<table id="F928F8F5B753E981FF76D68F667CA569" box="[147,1868,652,1545]" gridcols="10" gridrows="9" pageId="6" pageNumber="131">
<tr id="35180817B753E981FF76D68F667CA1B3" box="[147,1868,652,723]" gridrow="0" pageId="6" pageNumber="131" rowspan-0="1">
<th id="76C9616BB753E981FEFED68F6354A1B3" box="[283,612,652,723]" gridcol="1" gridrow="0" pageId="6" pageNumber="131">
<taxonomicName id="4C2871D6B7531678FEFED6B46089A1AA" authorityName="Dole-Olivier &amp; Hafid &amp; Piscart" authorityYear="2018" box="[283,441,695,714]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis id="B95CD647B7531678FEFED6B46089A1AA" box="[283,441,695,714]" italics="true" pageId="6" pageNumber="131">Pseudoniphargus</emphasis>
</taxonomicName>
species --&gt;
</th>
<th id="76C9616BB753E981FD25D68F656FA1B3" box="[704,1119,652,723]" colspan="3" colspanRight="2" gridcol="2" gridrow="0" pageId="6" pageNumber="131">
<emphasis id="B95CD647B7531678FD25D68F652DA1C0" bold="true" box="[704,1053,652,672]" pageId="6" pageNumber="131">Species from Mediterranean Islands</emphasis>
<emphasis id="B95CD647B7531678FD2AD6C3656BA1B3" box="[719,1115,704,723]" italics="true" pageId="6" pageNumber="131">
<taxonomicName id="4C2871D6B7531678FD2AD6C3621BA1B3" authorityName="S. Karaman" authorityYear="1955" box="[719,811,704,723]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="131" phylum="Arthropoda" rank="species" species="adriaticus">adriaticus</taxonomicName>
<taxonomicName id="4C2871D6B7531678FC9DD6C36280A1B3" authorityName="Jaume" authorityYear="1991" box="[888,944,704,723]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="131" phylum="Arthropoda" rank="species" species="daviui">daviui</taxonomicName>
<taxonomicName id="4C2871D6B7531678FBFAD6C3656BA1B3" authorityName="Messouli, Messana &amp; Yacoubi-Khebiza" authorityYear="2006" box="[1055,1115,704,723]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="131" phylum="Arthropoda" rank="species" species="duplus">duplus</taxonomicName>
</emphasis>
</th>
<th id="76C9616BB753E981FB4BD68F6415A1B3" box="[1198,1317,652,723]" gridcol="5" gridrow="0" pageId="6" pageNumber="131">
<taxonomicName id="4C2871D6B7531678FB2FD6C3643AA1B3" authorityName="Karaman &amp; Ruffo" authorityYear="1989" box="[1226,1290,704,723]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="131" phylum="Arthropoda" rank="species" species="sodalis">
<emphasis id="B95CD647B7531678FB2FD6C3643AA1B3" box="[1226,1290,704,723]" italics="true" pageId="6" pageNumber="131">sodalis</emphasis>
</taxonomicName>
</th>
<th id="76C9616BB753E981FAB9D68F648BA1B3" box="[1372,1467,652,723]" gridcol="6" gridrow="0" pageId="6" pageNumber="131">
<taxonomicName id="4C2871D6B7531678FA85D6C3648BA1B3" authorityName="Pretus" authorityYear="1988" box="[1376,1467,704,723]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="131" phylum="Arthropoda" rank="species" species="mercadali">
<emphasis id="B95CD647B7531678FA85D6C3648BA1B3" box="[1376,1467,704,723]" italics="true" pageId="6" pageNumber="131">mercadali</emphasis>
</taxonomicName>
</th>
<th id="76C9616BB753E981FA07D68F6770A1B3" box="[1506,1600,652,723]" gridcol="7" gridrow="0" pageId="6" pageNumber="131">
<taxonomicName id="4C2871D6B7531678FA07D6C36770A1B3" authorityName="Karaman &amp; Ruffo" authorityYear="1989" box="[1506,1600,704,723]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="131" phylum="Arthropoda" rank="species" species="inconditus">
<emphasis id="B95CD647B7531678FA07D6C36770A1B3" box="[1506,1600,704,723]" italics="true" pageId="6" pageNumber="131">inconditus</emphasis>
</taxonomicName>
</th>
<th id="76C9616BB753E981F9B8D68F67E4A1B3" box="[1629,1748,652,723]" gridcol="8" gridrow="0" pageId="6" pageNumber="131">
<taxonomicName id="4C2871D6B7531678F99DD6C3678AA1B3" authorityName="Karaman &amp; Ruffo" authorityYear="1989" box="[1656,1722,704,723]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="131" phylum="Arthropoda" rank="species" species="italicus">
<emphasis id="B95CD647B7531678F99DD6C3678AA1B3" box="[1656,1722,704,723]" italics="true" pageId="6" pageNumber="131">italicus</emphasis>
</taxonomicName>
</th>
<th id="76C9616BB753E981F8E8D68F667CA1B3" box="[1805,1868,652,723]" gridcol="9" gridrow="0" pageId="6" pageNumber="131">
<taxonomicName id="4C2871D6B7531678F8FED6C2667BA1B3" authorityName="Jaume" authorityYear="1991" box="[1819,1867,705,723]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="131" phylum="Arthropoda" rank="species" species="triasi">
<emphasis id="B95CD647B7531678F8FED6C2667BA1B3" box="[1819,1867,705,723]" italics="true" pageId="6" pageNumber="131">triasi</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="35180817B753E981FF76D6E0667CA03E" box="[147,1868,739,862]" gridrow="1" pageId="6" pageNumber="131">
<th id="76C9616BB753E981FF76D6E061C0A03E" box="[147,240,739,862]" gridcol="0" gridrow="1" pageId="6" pageNumber="131">Antenna 1 Antenna 2</th>
<td id="76C9616BB753E981FEFED6E06354A03E" box="[283,612,739,862]" gridcol="1" gridrow="1" pageId="6" pageNumber="131">flagellum: number of articles accessory flagellum longer than first flagellar article flagellum:number of articles</td>
<td id="76C9616BB753E981FD25D6E06208A03E" box="[704,824,739,862]" gridcol="2" gridrow="1" pageId="6" pageNumber="131">13-15 NO 5-7</td>
<td id="76C9616BB753E981FC91D6E06285A03E" box="[884,949,739,862]" gridcol="3" gridrow="1" pageId="6" pageNumber="131">
12
<emphasis id="B95CD647B7531678FC9BD71F629AA050" bold="true" box="[894,938,796,816]" pageId="6" pageNumber="131">YES</emphasis>
6
</td>
<td id="76C9616BB753E981FBFED6E0656FA03E" box="[1051,1119,739,862]" gridcol="4" gridrow="1" pageId="6" pageNumber="131">
<emphasis id="B95CD647B7531678FBD7D6EF6563A050" bold="true" pageId="6" pageNumber="131">31 YES</emphasis>
6
</td>
<td id="76C9616BB753E981FB4BD6E06415A03E" box="[1198,1317,739,862]" gridcol="5" gridrow="1" pageId="6" pageNumber="131">
<emphasis id="B95CD647B7531678FB3BD6EF65C5A060" bold="true" box="[1246,1269,748,768]" pageId="6" pageNumber="131">22</emphasis>
NO 8
</td>
<td id="76C9616BB753E981FAB9D6E0648BA03E" box="[1372,1467,739,862]" gridcol="6" gridrow="1" pageId="6" pageNumber="131">17 NO 7</td>
<td id="76C9616BB753E981FA07D6E06770A03E" box="[1506,1600,739,862]" gridcol="7" gridrow="1" pageId="6" pageNumber="131">?? NO?</td>
<td id="76C9616BB753E981F9B8D6E067E4A03E" box="[1629,1748,739,862]" gridcol="8" gridrow="1" pageId="6" pageNumber="131">
18
<emphasis id="B95CD647B7531678F966D71F679FA050" bold="true" box="[1667,1711,796,816]" pageId="6" pageNumber="131">YES</emphasis>
7
</td>
<td id="76C9616BB753E981F8E8D6E0667CA03E" box="[1805,1868,739,862]" gridcol="9" gridrow="1" pageId="6" pageNumber="131">15 NO 6</td>
</tr>
<tr id="35180817B753E981FF76D76F667CA0EA" box="[147,1868,876,906]" gridrow="2" pageId="6" pageNumber="131">
<th id="76C9616BB753E981FF76D76F61C0A0EA" box="[147,240,876,906]" gridcol="0" gridrow="2" pageId="6" pageNumber="131">P4</th>
<td id="76C9616BB753E981FEFED76F6354A0EA" box="[283,612,876,906]" gridcol="1" gridrow="2" pageId="6" pageNumber="131">coxal plate posterior excavation</td>
<td id="76C9616BB753E981FD25D76F6208A0EA" box="[704,824,876,906]" gridcol="2" gridrow="2" pageId="6" pageNumber="131">+</td>
<td id="76C9616BB753E981FC91D76F6285A0EA" box="[884,949,876,906]" gridcol="3" gridrow="2" pageId="6" pageNumber="131">+</td>
<td id="76C9616BB753E981FBFED76F656FA0EA" box="[1051,1119,876,906]" gridcol="4" gridrow="2" pageId="6" pageNumber="131">
<emphasis id="B95CD647B7531678FBCAD7756566A0EA" bold="true" box="[1071,1110,886,906]" pageId="6" pageNumber="131">+++</emphasis>
</td>
<td id="76C9616BB753E981FB4BD76F6415A0EA" box="[1198,1317,876,906]" gridcol="5" gridrow="2" pageId="6" pageNumber="131">++</td>
<td id="76C9616BB753E981FAB9D76F648BA0EA" box="[1372,1467,876,906]" gridcol="6" gridrow="2" pageId="6" pageNumber="131">+</td>
<td id="76C9616BB753E981FA07D76F6770A0EA" box="[1506,1600,876,906]" gridcol="7" gridrow="2" pageId="6" pageNumber="131">+</td>
<td id="76C9616BB753E981F9B8D76F67E4A0EA" box="[1629,1748,876,906]" gridcol="8" gridrow="2" pageId="6" pageNumber="131">?</td>
<td id="76C9616BB753E981F8E8D76F667CA0EA" box="[1805,1868,876,906]" gridcol="9" gridrow="2" pageId="6" pageNumber="131">+</td>
</tr>
<tr id="35180817B753E981FF76D794667CA769" box="[147,1868,919,1033]" gridrow="3" pageId="6" pageNumber="131">
<th id="76C9616BB753E981FF76D79461C0A769" box="[147,240,919,1033]" gridcol="0" gridrow="3" pageId="6" pageNumber="131">P7</th>
<td id="76C9616BB753E981FEFED7946354A769" box="[283,612,919,1033]" gridcol="1" gridrow="3" pageId="6" pageNumber="131">basis elongated vs wide basis postero distal lobe Propodus length/width ratio</td>
<td id="76C9616BB753E981FD25D7946208A769" box="[704,824,919,1033]" gridcol="2" gridrow="3" pageId="6" pageNumber="131">elongated present 8.5</td>
<td id="76C9616BB753E981FC91D7946285A769" box="[884,949,919,1033]" gridcol="3" gridrow="3" pageId="6" pageNumber="131">
<emphasis id="B95CD647B7531678FC9BD7A3629BA0D4" bold="true" box="[894,939,928,948]" pageId="6" pageNumber="131">wide</emphasis>
present
<emphasis id="B95CD647B7531678FC6CD7F66290A769" bold="true" box="[905,928,1013,1033]" pageId="6" pageNumber="131">13</emphasis>
</td>
<td id="76C9616BB753E981FBFED794656FA769" box="[1051,1119,919,1033]" gridcol="4" gridrow="3" pageId="6" pageNumber="131">
<emphasis id="B95CD647B7531678FBC3D7A36563A0D4" bold="true" box="[1062,1107,928,948]" pageId="6" pageNumber="131">wide</emphasis>
present
<emphasis id="B95CD647B7531678FBD7D7F66579A769" bold="true" box="[1074,1097,1013,1033]" pageId="6" pageNumber="131">12</emphasis>
</td>
<td id="76C9616BB753E981FB4BD7946415A769" box="[1198,1317,919,1033]" gridcol="5" gridrow="3" pageId="6" pageNumber="131">
elongated
<emphasis id="B95CD647B7531678FB2FD7C86438A0BF" bold="true" box="[1226,1288,971,991]" pageId="6" pageNumber="131">absent</emphasis>
10.5
</td>
<td id="76C9616BB753E981FAB9D794648BA769" box="[1372,1467,919,1033]" gridcol="6" gridrow="3" pageId="6" pageNumber="131">
elongated present
<emphasis id="B95CD647B7531678FA91D7F664ABA769" bold="true" box="[1396,1435,1013,1033]" pageId="6" pageNumber="131">12.3</emphasis>
</td>
<td id="76C9616BB753E981FA07D7946770A769" box="[1506,1600,919,1033]" gridcol="7" gridrow="3" pageId="6" pageNumber="131">elongated present ±9.2</td>
<td id="76C9616BB753E981F9B8D79467E4A769" box="[1629,1748,919,1033]" gridcol="8" gridrow="3" pageId="6" pageNumber="131">elongated present 7,5</td>
<td id="76C9616BB753E981F8E8D794667CA769" box="[1805,1868,919,1033]" gridcol="9" gridrow="3" pageId="6" pageNumber="131">
<emphasis id="B95CD647B7531678F8F3D7A3667CA0BF" bold="true" pageId="6" pageNumber="131">wide absent</emphasis>
10.5
</td>
</tr>
<tr id="35180817B753E981FF76D014667CA7E9" box="[147,1868,1047,1161]" gridrow="4" pageId="6" pageNumber="131">
<th id="76C9616BB753E981FF76D01461C0A7E9" box="[147,240,1047,1161]" gridcol="0" gridrow="4" pageId="6" pageNumber="131">Uropod 1</th>
<td id="76C9616BB753E981FEFED0146354A7E9" box="[283,612,1047,1161]" gridcol="1" gridrow="4" pageId="6" pageNumber="131">pedunculus basoventral spine pedunculus inner medial spine lateral spines on rami</td>
<td id="76C9616BB753E981FD25D0146208A7E9" box="[704,824,1047,1161]" gridcol="2" gridrow="4" pageId="6" pageNumber="131">1 0 0</td>
<td id="76C9616BB753E981FC91D0146285A7E9" box="[884,949,1047,1161]" gridcol="3" gridrow="4" pageId="6" pageNumber="131">
<emphasis id="B95CD647B7531678FC6AD02362AAA754" bold="true" box="[911,922,1056,1076]" pageId="6" pageNumber="131">0</emphasis>
0 0
</td>
<td id="76C9616BB753E981FBFED014656FA7E9" box="[1051,1119,1047,1161]" gridcol="4" gridrow="4" pageId="6" pageNumber="131">
2
<emphasis id="B95CD647B7531678FBD2D048657CA7E9" bold="true" pageId="6" pageNumber="131">1 0-1</emphasis>
</td>
<td id="76C9616BB753E981FB4BD0146415A7E9" box="[1198,1317,1047,1161]" gridcol="5" gridrow="4" pageId="6" pageNumber="131">
<emphasis id="B95CD647B7531678FB01D02365DFA754" bold="true" box="[1252,1263,1056,1076]" pageId="6" pageNumber="131">0</emphasis>
0 0
</td>
<td id="76C9616BB753E981FAB9D014648BA7E9" box="[1372,1467,1047,1161]" gridcol="6" gridrow="4" pageId="6" pageNumber="131">1 1 0</td>
<td id="76C9616BB753E981FA07D0146770A7E9" box="[1506,1600,1047,1161]" gridcol="7" gridrow="4" pageId="6" pageNumber="131">1 0 0</td>
<td id="76C9616BB753E981F9B8D01467E4A7E9" box="[1629,1748,1047,1161]" gridcol="8" gridrow="4" pageId="6" pageNumber="131">1 1 1</td>
<td id="76C9616BB753E981F8E8D014667CA7E9" box="[1805,1868,1047,1161]" gridcol="9" gridrow="4" pageId="6" pageNumber="131">1 1 0</td>
</tr>
<tr id="35180817B753E981FF76D094667CA7D3" box="[147,1868,1175,1203]" gridrow="5" pageId="6" pageNumber="131">
<th id="76C9616BB753E981FF76D09461C0A7D3" box="[147,240,1175,1203]" gridcol="0" gridrow="5" pageId="6" pageNumber="131">Uropod 2</th>
<td id="76C9616BB753E981FEFED0946354A7D3" box="[283,612,1175,1203]" gridcol="1" gridrow="5" pageId="6" pageNumber="131">lateral spines on rami</td>
<td id="76C9616BB753E981FD25D0946208A7D3" box="[704,824,1175,1203]" gridcol="2" gridrow="5" pageId="6" pageNumber="131">not given</td>
<td id="76C9616BB753E981FC91D0946285A7D3" box="[884,949,1175,1203]" gridcol="3" gridrow="5" pageId="6" pageNumber="131">0</td>
<td id="76C9616BB753E981FBFED094656FA7D3" box="[1051,1119,1175,1203]" gridcol="4" gridrow="5" pageId="6" pageNumber="131">0-1</td>
<td id="76C9616BB753E981FB4BD0946415A7D3" box="[1198,1317,1175,1203]" gridcol="5" gridrow="5" pageId="6" pageNumber="131">0</td>
<td id="76C9616BB753E981FAB9D094648BA7D3" box="[1372,1467,1175,1203]" gridcol="6" gridrow="5" pageId="6" pageNumber="131">0</td>
<td id="76C9616BB753E981FA07D0946770A7D3" box="[1506,1600,1175,1203]" gridcol="7" gridrow="5" pageId="6" pageNumber="131">0</td>
<td id="76C9616BB753E981F9B8D09467E4A7D3" box="[1629,1748,1175,1203]" gridcol="8" gridrow="5" pageId="6" pageNumber="131">1</td>
<td id="76C9616BB753E981F8E8D094667CA7D3" box="[1805,1868,1175,1203]" gridcol="9" gridrow="5" pageId="6" pageNumber="131">0</td>
</tr>
<tr id="35180817B753E981FF76D0C2667CA63E" box="[147,1868,1217,1374]" gridrow="6" pageId="6" pageNumber="131">
<th id="76C9616BB753E981FF76D0C261C0A63E" box="[147,240,1217,1374]" gridcol="0" gridrow="6" pageId="6" pageNumber="131">Uropod 3</th>
<td id="76C9616BB753E981FEFED0C26354A63E" box="[283,612,1217,1374]" gridcol="1" gridrow="6" pageId="6" pageNumber="131">pedunculus elongated lengh: sexual dimorphism? groups of exopod marginal spines number of distal spines on endopod</td>
<td id="76C9616BB753E981FD25D0C26208A63E" box="[704,824,1217,1374]" gridcol="2" gridrow="6" pageId="6" pageNumber="131">
<emphasis id="B95CD647B7531678FD02D0C96223A669" bold="true" box="[743,787,1226,1289]" pageId="6" pageNumber="131">YES YES</emphasis>
4-7
<emphasis id="B95CD647B7531678FD12D1496232A63E" bold="true" box="[759,770,1354,1374]" pageId="6" pageNumber="131">1</emphasis>
</td>
<td id="76C9616BB753E981FC91D0C26285A63E" box="[884,949,1217,1374]" gridcol="3" gridrow="6" pageId="6" pageNumber="131">
NO NO 5-6
<emphasis id="B95CD647B7531678FC6AD14962AAA63E" bold="true" box="[911,922,1354,1374]" pageId="6" pageNumber="131">1</emphasis>
</td>
<td id="76C9616BB753E981FBFED0C2656FA63E" box="[1051,1119,1217,1374]" gridcol="4" gridrow="6" pageId="6" pageNumber="131">
slightly? 8
<emphasis id="B95CD647B7531678FBD2D1496572A63E" bold="true" box="[1079,1090,1354,1374]" pageId="6" pageNumber="131">1</emphasis>
</td>
<td id="76C9616BB753E981FB4BD0C26415A63E" box="[1198,1317,1217,1374]" gridcol="5" gridrow="6" pageId="6" pageNumber="131">
<emphasis id="B95CD647B7531678FB31D0C965CEA654" bold="true" box="[1236,1280,1226,1332]" pageId="6" pageNumber="131">YES YES 5-24</emphasis>
2?
</td>
<td id="76C9616BB753E981FAB9D0C2648BA63E" box="[1372,1467,1217,1374]" gridcol="6" gridrow="6" pageId="6" pageNumber="131">
<emphasis id="B95CD647B7531678FA97D0C964AEA669" bold="true" box="[1394,1438,1226,1289]" pageId="6" pageNumber="131">YES YES</emphasis>
6-19
<emphasis id="B95CD647B7531678FA67D14964BDA63E" bold="true" box="[1410,1421,1354,1374]" pageId="6" pageNumber="131">1</emphasis>
</td>
<td id="76C9616BB753E981FA07D0C26770A63E" box="[1506,1600,1217,1374]" gridcol="7" gridrow="6" pageId="6" pageNumber="131">
<emphasis id="B95CD647B7531678FA1ED0C96717A669" bold="true" box="[1531,1575,1226,1289]" pageId="6" pageNumber="131">YES YES</emphasis>
4-6
<emphasis id="B95CD647B7531678F9EED1496726A63E" bold="true" box="[1547,1558,1354,1374]" pageId="6" pageNumber="131">1</emphasis>
</td>
<td id="76C9616BB753E981F9B8D0C267E4A63E" box="[1629,1748,1217,1374]" gridcol="8" gridrow="6" pageId="6" pageNumber="131">
<emphasis id="B95CD647B7531678F966D0C9679FA7BE" bold="true" box="[1667,1711,1226,1246]" pageId="6" pageNumber="131">YES</emphasis>
? 6
<emphasis id="B95CD647B7531678F976D14967AEA63E" bold="true" box="[1683,1694,1354,1374]" pageId="6" pageNumber="131">1</emphasis>
</td>
<td id="76C9616BB753E981F8E8D0C2667CA63E" box="[1805,1868,1217,1374]" gridcol="9" gridrow="6" pageId="6" pageNumber="131">
NO NO 10-11
<emphasis id="B95CD647B7531678F8C2D1496602A63E" bold="true" box="[1831,1842,1354,1374]" pageId="6" pageNumber="131">1</emphasis>
</td>
</tr>
<tr id="35180817B753E981FF76D16F667CA6BE" box="[147,1868,1388,1502]" gridrow="7" pageId="6" pageNumber="131">
<th id="76C9616BB753E981FF76D16F61C0A6BE" box="[147,240,1388,1502]" gridcol="0" gridrow="7" pageId="6" pageNumber="131">Telson</th>
<td id="76C9616BB753E981FEFED16F6354A6BE" box="[283,612,1388,1502]" gridcol="1" gridrow="7" pageId="6" pageNumber="131">distal spines on each lobe shape mediodistal notch</td>
<td id="76C9616BB753E981FD25D16F6208A6BE" box="[704,824,1388,1502]" gridcol="2" gridrow="7" pageId="6" pageNumber="131">2 quadrangular + to ++</td>
<td id="76C9616BB753E981FC91D16F6285A6BE" box="[884,949,1388,1502]" gridcol="3" gridrow="7" pageId="6" pageNumber="131">
1-2
<emphasis id="B95CD647B7531678FC9DD1A36281A6D4" bold="true" box="[888,945,1440,1460]" pageId="6" pageNumber="131">broad</emphasis>
++
</td>
<td id="76C9616BB753E981FBFED16F656FA6BE" box="[1051,1119,1388,1502]" gridcol="4" gridrow="7" pageId="6" pageNumber="131">
2-3
<emphasis id="B95CD647B7531678FBC5D1A36579A6BE" bold="true" box="[1056,1113,1440,1502]" pageId="6" pageNumber="131">broad +</emphasis>
</td>
<td id="76C9616BB753E981FB4BD16F6415A6BE" box="[1198,1317,1388,1502]" gridcol="5" gridrow="7" pageId="6" pageNumber="131">2-3 quadrangular +++</td>
<td id="76C9616BB753E981FAB9D16F648BA6BE" box="[1372,1467,1388,1502]" gridcol="6" gridrow="7" pageId="6" pageNumber="131">
2-3
<emphasis id="B95CD647B7531678FA8ED1A364A4A6BE" bold="true" box="[1387,1444,1440,1502]" pageId="6" pageNumber="131">broad +</emphasis>
</td>
<td id="76C9616BB753E981FA07D16F6770A6BE" box="[1506,1600,1388,1502]" gridcol="7" gridrow="7" pageId="6" pageNumber="131">
<emphasis id="B95CD647B7531678F9EED176672DA6BE" bold="true" pageId="6" pageNumber="131">3 broad +</emphasis>
</td>
<td id="76C9616BB753E981F9B8D16F67E4A6BE" box="[1629,1748,1388,1502]" gridcol="8" gridrow="7" pageId="6" pageNumber="131">3 quadrangular ++</td>
<td id="76C9616BB753E981F8E8D16F667CA6BE" box="[1805,1868,1388,1502]" gridcol="9" gridrow="7" pageId="6" pageNumber="131">
2-3
<emphasis id="B95CD647B7531678F8F5D1A36602A6BE" bold="true" box="[1808,1865,1440,1502]" pageId="6" pageNumber="131">broad 0</emphasis>
</td>
</tr>
<tr id="35180817B753E981FF76D1EF667CA569" box="[147,1868,1516,1545]" gridrow="8" pageId="6" pageNumber="131">
<th id="76C9616BB753E981FF76D1EF61C0A569" box="[147,240,1516,1545]" gridcol="0" gridrow="8" pageId="6" pageNumber="131">Epimera</th>
<td id="76C9616BB753E981FEFED1EF6354A569" box="[283,612,1516,1545]" gridcol="1" gridrow="8" pageId="6" pageNumber="131">ventral spines (from plate 1 to 3)</td>
<td id="76C9616BB753E981FD25D1EF6208A569" box="[704,824,1516,1545]" gridcol="2" gridrow="8" pageId="6" pageNumber="131">0,1,1</td>
<td id="76C9616BB753E981FC91D1EF6285A569" box="[884,949,1516,1545]" gridcol="3" gridrow="8" pageId="6" pageNumber="131">0,2,2</td>
<td id="76C9616BB753E981FBFED1EF656FA569" box="[1051,1119,1516,1545]" gridcol="4" gridrow="8" pageId="6" pageNumber="131">0, 1, 2</td>
<td id="76C9616BB753E981FB4BD1EF6415A569" box="[1198,1317,1516,1545]" gridcol="5" gridrow="8" pageId="6" pageNumber="131">0, 1, 2</td>
<td id="76C9616BB753E981FAB9D1EF648BA569" box="[1372,1467,1516,1545]" gridcol="6" gridrow="8" pageId="6" pageNumber="131">
<emphasis id="B95CD647B7531678FA94D1F664AEA569" bold="true" box="[1393,1438,1525,1545]" pageId="6" pageNumber="131">1,2,4</emphasis>
</td>
<td id="76C9616BB753E981FA07D1EF6770A569" box="[1506,1600,1516,1545]" gridcol="7" gridrow="8" pageId="6" pageNumber="131">
<emphasis id="B95CD647B7531678FA10D1F6671DA569" bold="true" box="[1525,1581,1525,1545]" pageId="6" pageNumber="131">2, 4, 5</emphasis>
</td>
<td id="76C9616BB753E981F9B8D1EF67E4A569" box="[1629,1748,1516,1545]" gridcol="8" gridrow="8" pageId="6" pageNumber="131">0,1,1</td>
<td id="76C9616BB753E981F8E8D1EF667CA569" box="[1805,1868,1516,1545]" gridcol="9" gridrow="8" pageId="6" pageNumber="131">0,3,3</td>
</tr>
</table>
</paragraph>
<caption id="DF575ADDB7521679FF76D636604EA12B" box="[147,382,565,587]" pageId="7" pageNumber="132">
<paragraph id="8B970A55B7521679FF76D636604EA12B" blockId="7.[147,1772,565,775]" box="[147,382,565,587]" pageId="7" pageNumber="132">
<emphasis id="B95CD647B7521679FF76D63661CFA12B" bold="true" box="[147,255,565,587]" pageId="7" pageNumber="132">TABLE 1.</emphasis>
(Continued)
</paragraph>
</caption>
<subSubSection id="C33259DEB7521672FD0ED65B6039A477" lastPageId="12" lastPageNumber="137" pageId="7" pageNumber="132" type="description">
<paragraph id="8B970A55B7521679FD0ED65B679EA10C" blockId="7.[147,1772,565,775]" box="[747,1710,600,620]" pageId="7" pageNumber="132">
<materialsCitation id="3B400008B7521679FD0ED65B679EA10C" ID-GBIF-Occurrence="1913889569" box="[747,1710,600,620]" country="Benin" location="Mediterranean Islands" pageId="7" pageNumber="132" specimenCount="1">
<emphasis id="B95CD647B7521679FD0ED65B679EA10C" bold="true" box="[747,1710,600,620]" pageId="7" pageNumber="132">
Species from
<location id="8EF75C8EB7521679FC89D65B6578A10C" LSID="urn:lsid:plazi:treatment:0381BB43B7541672FF72D10D6039A477:8EF75C8EB7521679FC89D65B6578A10C" box="[876,1096,600,620]" country="Benin" name="Mediterranean Islands" pageId="7" pageNumber="132">Mediterranean Islands</location>
<collectingCountry id="F33F4AC5B7521679F990D65B679EA10C" box="[1653,1710,600,620]" name="Benin" pageId="7" pageNumber="132">Benin</collectingCountry>
</emphasis>
</materialsCitation>
</paragraph>
<paragraph id="8B970A55B7521679FEC3D68067F3A1AB" blockId="7.[147,1772,565,775]" pageId="7" pageNumber="132">
<taxonomicName id="4C2871D6B7521679FEC3D68060F4A1F6" authorityName="Dole-Olivier &amp; Hafid &amp; Piscart" authorityYear="2018" box="[294,452,643,662]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="7" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis id="B95CD647B7521679FEC3D68060F4A1F6" box="[294,452,643,662]" italics="true" pageId="7" pageNumber="132">Pseudoniphargus</emphasis>
</taxonomicName>
species --&gt;
<emphasis id="B95CD647B7521679FD15D68F67DCA1FF" box="[752,1772,652,671]" italics="true" pageId="7" pageNumber="132">
pityusensis
<taxonomicName id="4C2871D6B7521679FC76D68F62DDA1FE" authorityName="Pretus" authorityYear="1990" box="[915,1005,652,670]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="7" pageNumber="132" phylum="Arthropoda" rank="species" species="racovitzai">racovitzai</taxonomicName>
<taxonomicName id="4C2871D6B7521679FBACD68F65ABA1FF" authorityName="Pretus" authorityYear="1990" box="[1097,1179,652,671]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="7" pageNumber="132" phylum="Arthropoda" rank="species" species="pedrerae">pedrerae</taxonomicName>
<taxonomicName id="4C2871D6B7521679FAE2D68F646FA1FF" authorityName="Messouli, Messana &amp; Yacoubi-Khebiza" authorityYear="2006" box="[1287,1375,652,671]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="7" pageNumber="132" phylum="Arthropoda" rank="species" species="planasiae">planasiae</taxonomicName>
<taxonomicName id="4C2871D6B7521679FA20D68F6735A1FF" authorityName="Messouli, Messana &amp; Yacoubi-Khebiza" authorityYear="2006" box="[1477,1541,652,671]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="7" pageNumber="132" phylum="Arthropoda" rank="species" species="obritus">obritus</taxonomicName>
<taxonomicName id="4C2871D6B7521679F968D68F67DCA1FF" authorityName="Lagnika, Messouli, Ibikounle, Sakiti, Boutin &amp; Coineau" authorityYear="2016" box="[1677,1772,652,671]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="7" pageNumber="132" phylum="Arthropoda" rank="species" species="beninensis">beninensis</taxonomicName>
</emphasis>
Antenna 1 flagellum: number of articles 18 12
<emphasis id="B95CD647B7521679FB85D6BB65B3A1AC" bold="true" box="[1120,1155,696,716]" pageId="7" pageNumber="132">&gt;21</emphasis>
<date id="FF962C95B7521679FAFCD6B464C1A1AB" box="[1305,1521,695,715]" pageId="7" pageNumber="132" value="2016-12-16">12-16 16</date>
14
</paragraph>
<paragraph id="8B970A55B7521679FEC3D6D963F1A18D" blockId="7.[147,1772,565,775]" box="[294,705,729,749]" pageId="7" pageNumber="132">accessory flagellum longer than first flagellar</paragraph>
<paragraph id="8B970A55B7521679FCC4D6EB67FDA19B" blockId="7.[147,1772,565,775]" box="[801,1741,743,763]" pageId="7" pageNumber="132">
= NO = NO
<emphasis id="B95CD647B7521679FA2AD6E467FDA19B" bold="true" box="[1487,1741,743,763]" pageId="7" pageNumber="132">YES YES</emphasis>
</paragraph>
<paragraph id="8B970A55B7521679FEC3D6F0606EA067" blockId="7.[147,1772,565,775]" box="[294,350,755,775]" pageId="7" pageNumber="132">article</paragraph>
<paragraph id="8B970A55B7521679FF76D70D67FCA71F" pageId="7" pageNumber="132">
<table id="F928F8F5B752E981FF76D70E67E7A71F" box="[147,1751,781,1151]" gridcols="9" gridrows="5" pageId="7" pageNumber="132">
<tr id="35180817B752E981FF76D70E67E7A04A" box="[147,1751,781,810]" gridrow="0" pageId="7" pageNumber="132">
<th id="76C9616BB752E981FF76D70E61C0A04A" box="[147,240,781,810]" gridcol="0" gridrow="0" pageId="7" pageNumber="132">Antenna 2</th>
<th id="76C9616BB752E981FEC3D70E6376A04A" box="[294,582,781,810]" gridcol="1" gridrow="0" pageId="7" pageNumber="132">flagellum:number of articles</th>
<th id="76C9616BB752E981FD13D70E627EA04A" box="[758,846,781,810]" gridcol="2" gridrow="0" pageId="7" pageNumber="132">6</th>
<th id="76C9616BB752E981FC6AD70E62D7A04A" box="[911,999,781,810]" gridcol="3" gridrow="0" pageId="7" pageNumber="132">6</th>
<th id="76C9616BB752E981FBB4D70E65A2A04A" box="[1105,1170,781,810]" gridcol="4" gridrow="0" pageId="7" pageNumber="132">8</th>
<th id="76C9616BB752E981FAE4D70E641BA04A" box="[1281,1323,781,810]" gridcol="5" gridrow="0" pageId="7" pageNumber="132">7 (</th>
<th id="76C9616BB752E981FAA4D70E6469A04A" box="[1345,1369,781,810]" gridcol="6" gridrow="0" pageId="7" pageNumber="132">)</th>
<th id="76C9616BB752E981FA5CD70E6721A04A" box="[1465,1553,781,810]" gridcol="7" gridrow="0" pageId="7" pageNumber="132">6-7</th>
<th id="76C9616BB752E981F973D70E67E7A04A" box="[1686,1751,781,810]" gridcol="8" gridrow="0" pageId="7" pageNumber="132">8</th>
</tr>
<tr id="35180817B752E981FF76D73B67E7A035" box="[147,1751,824,853]" gridrow="1" pageId="7" pageNumber="132">
<th id="76C9616BB752E981FF76D73B61C0A035" box="[147,240,824,853]" gridcol="0" gridrow="1" pageId="7" pageNumber="132">P4</th>
<td id="76C9616BB752E981FEC3D73B6376A035" box="[294,582,824,853]" gridcol="1" gridrow="1" pageId="7" pageNumber="132">coxal plate posterior excavation</td>
<td id="76C9616BB752E981FD13D73B627EA035" box="[758,846,824,853]" gridcol="2" gridrow="1" pageId="7" pageNumber="132">++</td>
<td id="76C9616BB752E981FC6AD73B62D7A035" box="[911,999,824,853]" gridcol="3" gridrow="1" pageId="7" pageNumber="132">+</td>
<td id="76C9616BB752E981FBB4D73B65A2A035" box="[1105,1170,824,853]" gridcol="4" gridrow="1" pageId="7" pageNumber="132">+</td>
<td id="76C9616BB752E981FAE4D73B6469A035" box="[1281,1369,824,853]" colspan="2" colspanRight="1" gridcol="5" gridrow="1" pageId="7" pageNumber="132">++</td>
<td id="76C9616BB752E981FA5CD73B6721A035" box="[1465,1553,824,853]" gridcol="7" gridrow="1" pageId="7" pageNumber="132">+</td>
<td id="76C9616BB752E981F973D73B67E7A035" box="[1686,1751,824,853]" gridcol="8" gridrow="1" pageId="7" pageNumber="132">++</td>
</tr>
<table id="F928F8F5B752E981FF76D76067E7A6CA" box="[147,1751,867,1450]" gridrows="10" pageId="7" pageNumber="132">
<tr id="35180817B752E981FF76D76067E7A0B5" box="[147,1751,867,981]" gridrow="2" pageId="7" pageNumber="132" rowspan-5="1" rowspan-6="1">
<th id="76C9616BB752E981FF76D76061C0A0B5" box="[147,240,867,981]" gridcol="0" gridrow="2" pageId="7" pageNumber="132">P7</th>
<td id="76C9616BB752E981FEC3D7606376A0B5" box="[294,582,867,981]" gridcol="1" gridrow="2" pageId="7" pageNumber="132">basis elongated vs wide basis postero distal lobe Propodus length/width ratio</td>
<td id="76C9616BB752E981FD13D760627EA0B5" box="[758,846,867,981]" gridcol="2" gridrow="2" pageId="7" pageNumber="132">
elongated
<emphasis id="B95CD647B7521679FCE7D7956206A0B5" bold="true" box="[770,833,918,981]" pageId="7" pageNumber="132">absent 11.3</emphasis>
</td>
<td id="76C9616BB752E981FC6AD76062D7A0B5" box="[911,999,867,981]" gridcol="3" gridrow="2" pageId="7" pageNumber="132">
elongated
<emphasis id="B95CD647B7521679FC7ED79562E9A0CA" bold="true" box="[923,985,918,938]" pageId="7" pageNumber="132">absent</emphasis>
8.6
</td>
<td id="76C9616BB752E981FBB4D76065A2A0B5" box="[1105,1170,867,981]" gridcol="4" gridrow="2" pageId="7" pageNumber="132">
<emphasis id="B95CD647B7521679FBBED76F65B8A0E0" bold="true" box="[1115,1160,876,896]" pageId="7" pageNumber="132">wide</emphasis>
present 10
</td>
elongated present 8.4
<td id="76C9616BB752E981FA5CD7606721A0B5" box="[1465,1553,867,981]" gridcol="7" gridrow="2" pageId="7" pageNumber="132">elongated present 7.5</td>
<td id="76C9616BB752E981F973D76067E7A0B5" box="[1686,1751,867,981]" gridcol="8" gridrow="2" pageId="7" pageNumber="132">
<emphasis id="B95CD647B7521679F944D76F67FEA0E0" bold="true" box="[1697,1742,876,896]" pageId="7" pageNumber="132">wide</emphasis>
present
<emphasis id="B95CD647B7521679F957D7C2678DA0B5" bold="true" box="[1714,1725,961,981]" pageId="7" pageNumber="132">7</emphasis>
</td>
</tr>
<tr id="35180817B752E981FF76D7E167E7A734" box="[147,1751,994,1108]" gridrow="3" pageId="7" pageNumber="132">
<th id="76C9616BB752E981FF76D7E161C0A734" box="[147,240,994,1108]" gridcol="0" gridrow="3" pageId="7" pageNumber="132">Uropod 1</th>
<td id="76C9616BB752E981FEC3D7E16376A734" box="[294,582,994,1108]" gridcol="1" gridrow="3" pageId="7" pageNumber="132">pedunculus basoventral spine pedunculus inner medial spine lateral spines on rami</td>
<td id="76C9616BB752E981FD13D7E1627EA734" box="[758,846,994,1108]" gridcol="2" gridrow="3" pageId="7" pageNumber="132">1 0 0-2</td>
<td id="76C9616BB752E981FC6AD7E162D7A734" box="[911,999,994,1108]" gridcol="3" gridrow="3" pageId="7" pageNumber="132">1 0 0</td>
<td id="76C9616BB752E981FBB4D7E165A2A734" box="[1105,1170,994,1108]" gridcol="4" gridrow="3" pageId="7" pageNumber="132">1 0 0</td>
<td id="76C9616BB752E981FAE4D7E1641BA734" box="[1281,1323,994,1108]" gridcol="5" gridrow="3" pageId="7" pageNumber="132">1 (0 (0 (</td>
<td id="76C9616BB752E981FAA4D7E16469A734" box="[1345,1369,994,1108]" gridcol="6" gridrow="3" pageId="7" pageNumber="132">)))</td>
<td id="76C9616BB752E981FA5CD7E16721A734" box="[1465,1553,994,1108]" gridcol="7" gridrow="3" pageId="7" pageNumber="132">0- 2 0- 1 0- 1</td>
<td id="76C9616BB752E981F973D7E167E7A734" box="[1686,1751,994,1108]" gridcol="8" gridrow="3" pageId="7" pageNumber="132">1 1 1</td>
</tr>
<tr id="35180817B752E981FF76D06167E7A71F" box="[147,1751,1122,1151]" gridrow="4" pageId="7" pageNumber="132">
<th id="76C9616BB752E981FF76D06161C0A71F" box="[147,240,1122,1151]" gridcol="0" gridrow="4" pageId="7" pageNumber="132">Uropod 2</th>
<td id="76C9616BB752E981FEC3D0616376A71F" box="[294,582,1122,1151]" gridcol="1" gridrow="4" pageId="7" pageNumber="132">lateral spines on rami</td>
<td id="76C9616BB752E981FD13D061627EA71F" box="[758,846,1122,1151]" gridcol="2" gridrow="4" pageId="7" pageNumber="132">0</td>
<td id="76C9616BB752E981FC6AD06162D7A71F" box="[911,999,1122,1151]" gridcol="3" gridrow="4" pageId="7" pageNumber="132">0</td>
<td id="76C9616BB752E981FBB4D06165A2A71F" box="[1105,1170,1122,1151]" gridcol="4" gridrow="4" pageId="7" pageNumber="132">0</td>
<td id="76C9616BB752E981FAE4D061641BA71F" box="[1281,1323,1122,1151]" gridcol="5" gridrow="4" pageId="7" pageNumber="132">0 (</td>
<td id="76C9616BB752E981FAA4D0616469A71F" box="[1345,1369,1122,1151]" gridcol="6" gridrow="4" pageId="7" pageNumber="132">)</td>
<td id="76C9616BB752E981FA5CD0616721A71F" box="[1465,1553,1122,1151]" gridcol="7" gridrow="4" pageId="7" pageNumber="132">0- 1</td>
<td id="76C9616BB752E981F973D06167E7A71F" box="[1686,1751,1122,1151]" gridcol="8" gridrow="4" pageId="7" pageNumber="132">1-2</td>
</tr>
<paragraph id="8B970A55B7521679FF76D08E678DA64A" pageId="7" pageNumber="132">
Uropod 3 pedunculus elongated
<emphasis id="B95CD647B7521679FCE9D0956208A7CA" bold="true" box="[780,824,1174,1194]" pageId="7" pageNumber="132">YES</emphasis>
NO NO
<emphasis id="B95CD647B7521679FAF2D09564CBA7CA" bold="true" box="[1303,1531,1174,1194]" pageId="7" pageNumber="132">YES YES</emphasis>
NO lengh: sexual dimorphism?? NO NO
<emphasis id="B95CD647B7521679FAF2D0C264CBA7B5" bold="true" box="[1303,1531,1217,1237]" pageId="7" pageNumber="132">YES YES</emphasis>
? groups of exopod marginal spines 11 1-3 7-8 7-14 8 7 number of distal spines on endopod 1 1 1 1 1 1
</paragraph>
<paragraph id="8B970A55B7521679FF76D13467FEA6B4" pageId="7" pageNumber="132">
<table id="F928F8F5B752E981FF76D13467E3A6B5" box="[147,1747,1335,1493]" gridcols="9" gridrows="4" pageId="7" pageNumber="132">
<tr id="35180817B752E981FF76D13467E3A634" box="[147,1747,1335,1364]" gridrow="0" pageId="7" pageNumber="132">
<th id="76C9616BB752E981FF76D13461EFA634" box="[147,223,1335,1364]" gridcol="0" gridrow="0" pageId="7" pageNumber="132">Telson</th>
<th id="76C9616BB752E981FEC3D13460EBA634" box="[294,475,1335,1364]" gridcol="1" gridrow="0" pageId="7" pageNumber="132">distal spines on each</th>
<th id="76C9616BB752E981FE17D134636EA634" box="[498,606,1335,1364]" gridcol="2" gridrow="0" pageId="7" pageNumber="132">lobe</th>
<th id="76C9616BB752E981FCE0D134620FA634" box="[773,831,1335,1364]" gridcol="3" gridrow="0" pageId="7" pageNumber="132">2</th>
<th id="76C9616BB752E981FC76D13462DDA634" box="[915,1005,1335,1364]" gridcol="4" gridrow="0" pageId="7" pageNumber="132">1-2</th>
<th id="76C9616BB752E981FBB0D13465BFA634" box="[1109,1167,1335,1364]" gridcol="5" gridrow="0" pageId="7" pageNumber="132">2</th>
<th id="76C9616BB752E981FAF5D134647AA634" box="[1296,1354,1335,1364]" gridcol="6" gridrow="0" pageId="7" pageNumber="132">3-4</th>
<th id="76C9616BB752E981FA5CD1346721A634" box="[1465,1553,1335,1364]" gridcol="7" gridrow="0" pageId="7" pageNumber="132">2</th>
<th id="76C9616BB752E981F97FD13467E3A634" box="[1690,1747,1335,1364]" gridcol="8" gridrow="0" pageId="7" pageNumber="132">3</th>
</tr>
<tr id="35180817B752E981FF76D16167E3A61F" box="[147,1747,1378,1407]" gridrow="1" pageId="7" pageNumber="132" rowspan-0="1" rowspan-2="1">
<td id="76C9616BB752E981FEC3D16160EBA61F" box="[294,475,1378,1407]" gridcol="1" gridrow="1" pageId="7" pageNumber="132">shape</td>
<emphasis id="B95CD647B7521679FCE0D168647AA61F" bold="true" box="[773,1354,1387,1407]" pageId="7" pageNumber="132">
<td id="76C9616BB752E981FCE0D161620FA61F" box="[773,831,1378,1407]" gridcol="3" gridrow="1" pageId="7" pageNumber="132">broad</td>
<td id="76C9616BB752E981FC76D16162DDA61F" box="[915,1005,1378,1407]" gridcol="4" gridrow="1" pageId="7" pageNumber="132">broad</td>
<td id="76C9616BB752E981FBB0D16165BFA61F" box="[1109,1167,1378,1407]" gridcol="5" gridrow="1" pageId="7" pageNumber="132">broad</td>
<td id="76C9616BB752E981FAF5D161647AA61F" box="[1296,1354,1378,1407]" gridcol="6" gridrow="1" pageId="7" pageNumber="132">broad</td>
</emphasis>
<td id="76C9616BB752E981FA5CD1616721A61F" box="[1465,1553,1378,1407]" gridcol="7" gridrow="1" pageId="7" pageNumber="132">elongated</td>
<td id="76C9616BB752E981F97FD16167E3A61F" box="[1690,1747,1378,1407]" gridcol="8" gridrow="1" pageId="7" pageNumber="132">
<emphasis id="B95CD647B7521679F97FD16867E3A61F" bold="true" box="[1690,1747,1387,1407]" pageId="7" pageNumber="132">broad</emphasis>
</td>
</tr>
<tr id="35180817B752E981FF76D18E67E3A6CA" box="[147,1747,1421,1450]" gridrow="2" pageId="7" pageNumber="132" rowspan-0="1" rowspan-2="1">
<td id="76C9616BB752E981FEC3D18E60EBA6CA" box="[294,475,1421,1450]" gridcol="1" gridrow="2" pageId="7" pageNumber="132">mediodistal notch</td>
<td id="76C9616BB752E981FCE0D18E620FA6CA" box="[773,831,1421,1450]" gridcol="3" gridrow="2" pageId="7" pageNumber="132">+++</td>
<td id="76C9616BB752E981FC76D18E62DDA6CA" box="[915,1005,1421,1450]" gridcol="4" gridrow="2" pageId="7" pageNumber="132">
<emphasis id="B95CD647B7521679FC76D1956290A6CA" bold="true" box="[915,928,1430,1450]" pageId="7" pageNumber="132">+</emphasis>
/ ++
</td>
<td id="76C9616BB752E981FBB0D18E65BFA6CA" box="[1109,1167,1421,1450]" gridcol="5" gridrow="2" pageId="7" pageNumber="132">
<emphasis id="B95CD647B7521679FB89D1956547A6CA" bold="true" box="[1132,1143,1430,1450]" pageId="7" pageNumber="132">0</emphasis>
</td>
<td id="76C9616BB752E981FAF5D18E647AA6CA" box="[1296,1354,1421,1450]" gridcol="6" gridrow="2" pageId="7" pageNumber="132">++</td>
<td id="76C9616BB752E981FA5CD18E6721A6CA" box="[1465,1553,1421,1450]" gridcol="7" gridrow="2" pageId="7" pageNumber="132">
<emphasis id="B95CD647B7521679FA3BD19564DBA6CA" bold="true" box="[1502,1515,1430,1450]" pageId="7" pageNumber="132">+</emphasis>
</td>
<td id="76C9616BB752E981F97FD18E67E3A6CA" box="[1690,1747,1421,1450]" gridcol="8" gridrow="2" pageId="7" pageNumber="132">+++</td>
</tr>
<tr id="35180817B752E981FF76D1B467E3A6B5" box="[147,1747,1463,1493]" gridrow="3" pageId="7" pageNumber="132">
<th id="76C9616BB752E981FF76D1B461EFA6B5" box="[147,223,1463,1493]" gridcol="0" gridrow="3" pageId="7" pageNumber="132">Epimera</th>
<td id="76C9616BB752E981FEC3D1B460EBA6B5" box="[294,475,1463,1493]" gridcol="1" gridrow="3" pageId="7" pageNumber="132">ventral spines (from</td>
<td id="76C9616BB752E981FE17D1B4636EA6B5" box="[498,606,1463,1493]" gridcol="2" gridrow="3" pageId="7" pageNumber="132">plate 1 to 3)</td>
<td id="76C9616BB752E981FCE0D1B4620FA6B5" box="[773,831,1463,1493]" gridcol="3" gridrow="3" pageId="7" pageNumber="132">
<emphasis id="B95CD647B7521679FCEED1C26209A6B5" bold="true" box="[779,825,1473,1493]" pageId="7" pageNumber="132">0,2,3</emphasis>
</td>
<td id="76C9616BB752E981FC76D1B462DDA6B5" box="[915,1005,1463,1493]" gridcol="4" gridrow="3" pageId="7" pageNumber="132">0, 1, 1</td>
<td id="76C9616BB752E981FBB0D1B465BFA6B5" box="[1109,1167,1463,1493]" gridcol="5" gridrow="3" pageId="7" pageNumber="132">
<emphasis id="B95CD647B7521679FBBDD1C265BBA6B5" bold="true" box="[1112,1163,1473,1493]" pageId="7" pageNumber="132">1, 5,3</emphasis>
</td>
<td id="76C9616BB752E981FAF5D1B4647AA6B5" box="[1296,1354,1463,1493]" gridcol="6" gridrow="3" pageId="7" pageNumber="132">
<emphasis id="B95CD647B7521679FAF3D1C26474A6B5" bold="true" box="[1302,1348,1473,1493]" pageId="7" pageNumber="132">0,4,3</emphasis>
</td>
<td id="76C9616BB752E981FA5CD1B46721A6B5" box="[1465,1553,1463,1493]" gridcol="7" gridrow="3" pageId="7" pageNumber="132">1,2,1</td>
<td id="76C9616BB752E981F97FD1B467E3A6B5" box="[1690,1747,1463,1493]" gridcol="8" gridrow="3" pageId="7" pageNumber="132">1,2,3</td>
</tr>
</table>
</paragraph>
</table>
</table>
</paragraph>
<paragraph id="8B970A55B7521679FF76D1EA62B7A69F" blockId="7.[147,903,1513,1575]" box="[147,903,1513,1535]" pageId="7" pageNumber="132">0: absent; +: poorly marked; ++: marked; +++: strongly marked (V-shaped)</paragraph>
<paragraph id="8B970A55B7521679FF76D212630EA547" blockId="7.[147,903,1513,1575]" box="[147,574,1553,1575]" pageId="7" pageNumber="132">Bold cases indicate apomorphic characters</paragraph>
<caption id="DF575ADDB75D1676FF72D38A62CCA4BD" httpUri="https://zenodo.org/record/1440512/files/figure.png" pageId="8" pageNumber="133" startId="8.[151,250,1929,1951]" targetBox="[209,1374,217,1906]" targetPageId="8">
<paragraph id="8B970A55B75D1676FF72D38A62CCA4BD" blockId="8.[151,1435,1929,2013]" pageId="8" pageNumber="133">
<emphasis id="B95CD647B75D1676FF72D38A6022A4FE" bold="true" box="[151,274,1929,1951]" pageId="8" pageNumber="133">FIGURE 2.</emphasis>
<taxonomicName id="4C2871D6B75D1676FEFDD38A6323A4FF" authority="Dole-Olivier &amp; Hafid &amp; Piscart, 2018" authorityName="Dole-Olivier &amp; Hafid &amp; Piscart" authorityYear="2018" box="[280,531,1929,1951]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="133" phylum="Arthropoda" rank="species" species="djemoi" status="sp. nov.">
<emphasis id="B95CD647B75D1676FEFDD38A6323A4FF" box="[280,531,1929,1951]" italics="true" pageId="8" pageNumber="133">Pseudoniphargus djemoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F6B3CB75D1676FDFDD3896361A4FF" box="[536,593,1930,1951]" pageId="8" pageNumber="133" rank="species">
<emphasis id="B95CD647B75D1676FDFDD3896361A4FF" bold="true" box="[536,593,1930,1951]" pageId="8" pageNumber="133">sp. n.</emphasis>
</taxonomicNameLabel>
A, pereopod 4 of female 1 (6,5 mm); B, pereopod 7 of male 1 (6 mm) with detail of basis (B1), propodus and dactylus (B2); C, pereopod 5 of male 1 with detail of propodus and dactylus (C1); D, pereopod 6 of male 1 with detail of basis (D1), propodus and dactylus (D2); E, Antenna 2 of male 1.
</paragraph>
</caption>
<caption id="DF575ADDB75C1677FF72D37C62B5A491" httpUri="https://zenodo.org/record/1440514/files/figure.png" pageId="9" pageNumber="134" startId="9.[151,250,1919,1941]" targetBox="[163,1357,199,1899]" targetPageId="9">
<paragraph id="8B970A55B75C1677FF72D37C62B5A491" blockId="9.[151,1436,1919,2033]" pageId="9" pageNumber="134">
<emphasis id="B95CD647B75C1677FF72D37C6023A4F4" bold="true" box="[151,275,1919,1941]" pageId="9" pageNumber="134">FIGURE 3.</emphasis>
<taxonomicName id="4C2871D6B75C1677FEFED37C6327A4F5" authority="Dole-Olivier &amp; Hafid &amp; Piscart, 2018" authorityName="Dole-Olivier &amp; Hafid &amp; Piscart" authorityYear="2018" box="[283,535,1919,1941]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="134" phylum="Arthropoda" rank="species" species="djemoi" status="sp. nov.">
<emphasis id="B95CD647B75C1677FEFED37C6327A4F5" box="[283,535,1919,1941]" italics="true" pageId="9" pageNumber="134">Pseudoniphargus djemoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F6B3CB75C1677FDFBD3836369A4F5" box="[542,601,1920,1941]" pageId="9" pageNumber="134" rank="species">
<emphasis id="B95CD647B75C1677FDFBD3836369A4F5" bold="true" box="[542,601,1920,1941]" pageId="9" pageNumber="134">sp. n.</emphasis>
</taxonomicNameLabel>
A, uropod 3 of female 1 (6,5 mm); B, uropod 3 of male 2 (6,5 mm); C, telson of female 1 (6,5 mm); D, telson of male 1 (6 mm); E, uropods 1 and 2 of female 1 (6,5 mm); F,antenna 1 of female 2 (6 mm) with detail of accessory flagellum (F1), and apex of flagellum (F2); G, antenna 1 of male 1 (6 mm) with detail of apex of flagellum (G1) and accessory flagellum (G2); H, uropods 1 and 2 of male 1 (6 mm).
</paragraph>
</caption>
<caption id="DF575ADDB75F1674FF72D392625AA484" httpUri="https://zenodo.org/record/1440516/files/figure.png" pageId="10" pageNumber="135" startId="10.[151,250,1937,1959]" targetBox="[191,1353,205,1908]" targetPageId="10">
<paragraph id="8B970A55B75F1674FF72D392625AA484" blockId="10.[151,1436,1937,2020]" pageId="10" pageNumber="135">
<emphasis id="B95CD647B75F1674FF72D3926022A4C6" bold="true" box="[151,274,1937,1959]" pageId="10" pageNumber="135">FIGURE 4.</emphasis>
<taxonomicName id="4C2871D6B75F1674FEFDD3926322A4C7" authority="Dole-Olivier &amp; Hafid &amp; Piscart, 2018" authorityName="Dole-Olivier &amp; Hafid &amp; Piscart" authorityYear="2018" box="[280,530,1937,1959]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="135" phylum="Arthropoda" rank="species" species="djemoi" status="sp. nov.">
<emphasis id="B95CD647B75F1674FEFDD3926322A4C7" box="[280,530,1937,1959]" italics="true" pageId="10" pageNumber="135">Pseudoniphargus djemoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F6B3CB75F1674FDFDD3926361A4C6" box="[536,593,1937,1958]" pageId="10" pageNumber="135" rank="species">
<emphasis id="B95CD647B75F1674FDFDD3926361A4C6" bold="true" box="[536,593,1937,1958]" pageId="10" pageNumber="135">sp. n.</emphasis>
</taxonomicNameLabel>
A, maxillipeds of male 2 (5,5 mm), with detail of distal part of inner plates (A1); B, left mandible of female 1 (6,5 mm); C, right mandible of female 1 (6,5 mm); DF, epimera 1 to 3; G, labrum of male 2 (5,5 mm); H, labium of male 2 (5,5 mm); I, maxilla 1 of female 1 (6,5 mm).
</paragraph>
</caption>
<caption id="DF575ADDB75E1675FF72D3C665DAA4BB" box="[151,1258,1989,2011]" httpUri="https://zenodo.org/record/1440518/files/figure.png" pageId="11" pageNumber="136" startId="11.[151,250,1989,2011]" targetBox="[192,1382,195,1931]" targetPageId="11">
<paragraph id="8B970A55B75E1675FF72D3C665DAA4BB" blockId="11.[151,1258,1989,2011]" box="[151,1258,1989,2011]" pageId="11" pageNumber="136">
<emphasis id="B95CD647B75E1675FF72D3C66022A4BB" bold="true" box="[151,274,1989,2011]" pageId="11" pageNumber="136">FIGURE 5.</emphasis>
<taxonomicName id="4C2871D6B75E1675FEFCD3C66324A4BB" authority="Dole-Olivier &amp; Hafid &amp; Piscart, 2018" authorityName="Dole-Olivier &amp; Hafid &amp; Piscart" authorityYear="2018" box="[281,532,1989,2011]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="11" pageNumber="136" phylum="Arthropoda" rank="species" species="djemoi" status="sp. nov.">
<emphasis id="B95CD647B75E1675FEFCD3C66324A4BB" box="[281,532,1989,2011]" italics="true" pageId="11" pageNumber="136">Pseudoniphargus djemoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26F6B3CB75E1675FDFED3C56364A4BB" box="[539,596,1990,2011]" pageId="11" pageNumber="136" rank="species">
<emphasis id="B95CD647B75E1675FDFED3C56364A4BB" bold="true" box="[539,596,1990,2011]" pageId="11" pageNumber="136">sp. n.</emphasis>
</taxonomicNameLabel>
, male 1 (6 mm). A, gnathopod 1; B, pereopod 4, C, gnathopod 2.
</paragraph>
</caption>
<caption id="DF575ADDB7591672FF72D0EB624BA79E" box="[151,891,1256,1278]" httpUri="https://zenodo.org/record/1440520/files/figure.png" pageId="12" pageNumber="137" startId="12.[151,250,1256,1278]" targetBox="[157,1399,200,1228]" targetPageId="12">
<paragraph id="8B970A55B7591672FF72D0EB624BA79E" blockId="12.[151,891,1256,1278]" box="[151,891,1256,1278]" pageId="12" pageNumber="137">
<emphasis id="B95CD647B7591672FF72D0EB6022A79D" bold="true" box="[151,274,1256,1278]" pageId="12" pageNumber="137">FIGURE 6.</emphasis>
Distribution of African species of
<taxonomicName id="4C2871D6B7591672FD96D0EB6213A79E" authorityName="Dole-Olivier &amp; Hafid &amp; Piscart" authorityYear="2018" box="[627,803,1256,1278]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis id="B95CD647B7591672FD96D0EB6213A79E" box="[627,803,1256,1278]" italics="true" pageId="12" pageNumber="137">Pseudoniphargus</emphasis>
</taxonomicName>
species.
</paragraph>
</caption>
<paragraph id="8B970A55B7591672FF2AD1296039A477" blockId="12.[151,1437,1322,1815]" pageId="12" pageNumber="137">
The high level of endemism, associated to a high species diversity, is likely to be the result of the successive and independent colonization of inland waters by several marine ancestors in the Late Tertiary between the Eocene and the Pliocene (
<bibRefCitation id="EFB977A4B7591672FE85D17160D4A6EB" box="[352,484,1394,1419]" pageId="12" pageNumber="137" refId="ref9042" refString="Stock, J. H. (1980) Regression model evolution as exemplified by the genus Pseudoniphargus (Amphipoda). Bijdragen tot de Dierkunde, 50 (1), 105 - 144." type="journal article">Stock 1980</bibRefCitation>
;
<bibRefCitation id="EFB977A4B7591672FE0AD1716230A6EB" box="[495,768,1394,1419]" pageId="12" pageNumber="137" refId="ref8241" refString="Coineau, N. &amp; Boutin, C. (1996) Age and origin of the stygobiontic amphipod Pseudoniphargus (Crustacea) in Morocco, with the description of three new species. Bollettino del Museo Civico di Storia Naturale di Uerona, 20, 503 - 520." type="journal article">Coineau &amp; Boutin 1996</bibRefCitation>
). The origin of continental African species is old compared to the species inhabiting the Atlantic and Mediterranean Islands. The colonization of freshwater likely started during the early Miocene (around -20 MYA) from littoral sands of the Tethys.
<taxonomicName id="4C2871D6B7591672FBFED1B86592A6B3" authorityName="Chevreux" authorityYear="1901" box="[1051,1186,1467,1491]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="137" phylum="Arthropoda" rank="species" species="africanus">
<emphasis id="B95CD647B7591672FBFED1B86592A6B3" box="[1051,1186,1467,1491]" italics="true" pageId="12" pageNumber="137">P. africanus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2871D6B7591672FB39D1B864A5A6B3" authorityName="Stock" authorityYear="1980" box="[1244,1429,1467,1491]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="137" phylum="Arthropoda" rank="species" species="macrotelsonis">
<emphasis id="B95CD647B7591672FB39D1B864A5A6B3" box="[1244,1429,1467,1491]" italics="true" pageId="12" pageNumber="137">P. macrotelsonis</emphasis>
</taxonomicName>
, found in
<collectingCountry id="F33F4AC5B7591672FEE7D1DD6066A697" box="[258,342,1502,1527]" name="Algeria" pageId="12" pageNumber="137">Algeria</collectingCountry>
and
<collectingCountry id="F33F4AC5B7591672FE77D1DD60D8A697" box="[402,488,1502,1527]" name="Tunisia" pageId="12" pageNumber="137">Tunisia</collectingCountry>
, were for a long time considered to be the oldest species in North African inland waters (
<bibRefCitation id="EFB977A4B7591672FF14D201633DA57B" box="[241,525,1538,1563]" pageId="12" pageNumber="137" refId="ref8086" refString="Boutin, C. &amp; Coineau, N. (1988) Pseudoniphargus maroccanus n. sp. (subterranean amphipod), the first representative of the genus in Morocco, phylogenetic relationships and paleobiogeography. Crustaceana, 13 (Supplement), 1 - 19." type="journal article">Boutin &amp; Coineau, 1988</bibRefCitation>
) until the recent discovery of
<taxonomicName id="4C2871D6B7591672FC8CD20062CCA57B" authorityName="Lagnika, Messouli, Ibikounle, Sakiti, Boutin &amp; Coineau" authorityYear="2016" box="[873,1020,1539,1563]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="137" phylum="Arthropoda" rank="species" species="beninensis">
<emphasis id="B95CD647B7591672FC8CD20062CCA57B" box="[873,1020,1539,1563]" italics="true" pageId="12" pageNumber="137">P. beninensis</emphasis>
</taxonomicName>
, for which the ancestors could have colonized the northeastern part of
<collectingCountry id="F33F4AC5B7591672FDF0D2256368A55F" box="[533,600,1574,1599]" name="Benin" pageId="12" pageNumber="137">Benin</collectingCountry>
at the beginning of the Eocene (
<bibRefCitation id="EFB977A4B7591672FC24D2256591A55F" box="[961,1185,1574,1599]" pageId="12" pageNumber="137" refId="ref8546" refString="Lagnika, M., Messouli, M., Ibikounle, M., Sakiti, N., Boutin, C. &amp; Coineau, N. (2016) First record of groundwater amphipods (Crustacea) from Benin; range extension of the genus Pseudoniphargus to South of the Sahara, in western Africa. Bulletin de la Societe d'Histoire Naturelle de Toulouse, 152, 21 - 30." type="journal article">
Lagnika
<emphasis id="B95CD647B7591672FBC0D224656DA55E" box="[1061,1117,1574,1598]" italics="true" pageId="12" pageNumber="137">et al.</emphasis>
2016
</bibRefCitation>
). However, the lack of knowledge about the freshwater stygofauna of Africa (
<bibRefCitation id="EFB977A4B7591672FCF1D2496572A503" box="[788,1090,1610,1635]" pageId="12" pageNumber="137" refId="ref9243" refString="Tuekam Kayo, R., Marmonier, P., Zebaze Togouet, S. H., Nola, M. &amp; Piscart, C. (2012) An annotated checklist of freshwater stygobiotic crustaceans of Africa and Madagascar. Crustaceana, 85, 1613 - 1631. https: // doi. org / 10.1163 / 15685403 - 00003134" type="journal article">
Tuekam Kayo
<emphasis id="B95CD647B7591672FC27D24862CEA503" box="[962,1022,1611,1635]" italics="true" pageId="12" pageNumber="137">et al.</emphasis>
2012
</bibRefCitation>
) and the fact that at least ten species from northern
<collectingCountry id="F33F4AC5B7591672FE7BD26C6333A5E6" box="[414,515,1647,1670]" name="Morocco" pageId="12" pageNumber="137">Morocco</collectingCountry>
(
<bibRefCitation id="EFB977A4B7591672FDF3D26D6239A5E7" box="[534,777,1646,1671]" pageId="12" pageNumber="137" refId="ref8683" refString="Messouli, M., Boutin, C., Yacoubi-Khebiza, M. &amp; Coineau, N. (2001) Pseudoniphargus (subterranean crustacean amphipod) from Morocco: systematics, phylogeny and ecological and biogeographic aspects. Proceedings of the 13 th International Congress of Speleology, Brasilia, 15 - 22 July 2001, 391 - 394." type="proceedings paper">
Messouli
<emphasis id="B95CD647B7591672FD6CD26C63F5A5E6" box="[649,709,1646,1670]" italics="true" pageId="12" pageNumber="137">et al.</emphasis>
2001
</bibRefCitation>
) and four from
<collectingCountry id="F33F4AC5B7591672FC2ED26D652FA5E7" box="[971,1055,1646,1671]" name="Algeria" pageId="12" pageNumber="137">Algeria</collectingCountry>
and
<collectingCountry id="F33F4AC5B7591672FBBBD26D6581A5E7" box="[1118,1201,1646,1671]" name="Tunisia" pageId="12" pageNumber="137">Tunisia</collectingCountry>
remain undescribed (
<bibRefCitation id="EFB977A4B7591672FF7BD2916354A5CB" box="[158,612,1682,1707]" pageId="12" pageNumber="137" refId="ref9042" refString="Stock, J. H. (1980) Regression model evolution as exemplified by the genus Pseudoniphargus (Amphipoda). Bijdragen tot de Dierkunde, 50 (1), 105 - 144." type="journal article">Stock 1980, C.P. personal observations</bibRefCitation>
), do not allow us to establish a sound hypothesis on the origin and the geographical distribution of the genus
<taxonomicName id="4C2871D6B7591672FDB6D2B56226A5AE" authorityName="Dole-Olivier &amp; Hafid &amp; Piscart" authorityYear="2018" box="[595,790,1718,1742]" class="Malacostraca" family="Pseudoniphargidae" genus="Pseudoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis id="B95CD647B7591672FDB6D2B56226A5AE" box="[595,790,1718,1742]" italics="true" pageId="12" pageNumber="137">Pseudoniphargus</emphasis>
</taxonomicName>
in Continental Africa. Further investigations using both additional samples and DNA analysis are hence required to disentangle the global and local distribution pattern of the genus.
</paragraph>
</subSubSection>
</treatment>
</document>