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<document id="E5BED6E80C1CCECDBE23CF8DA6EEE4A0" ID-DOI="10.11646/zootaxa.4755.2.12" ID-GBIF-Dataset="6b11c9bd-eae8-4973-bd53-5ebe9b658b1d" ID-ISSN="1175-5334" ID-Zenodo-Dep="3733721" ID-ZooBank="urn:lsid:zoobank.org:pub:375D7830-D90A-4D91-905F-786A1215DFED" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1585597547394" checkinUser="carolina" docAuthor="Liu, Chen &amp; Wang, Shuxia" docDate="2020" docId="0381830DFFB6FFA7FF0BFE8901EEB7C4" docLanguage="en" docName="zootaxa.4755.2.12.pdf.imf" docOrigin="Zootaxa 4755 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Promalactis serrulata Liu &amp; Wang 2020, sp. nov." docType="treatment" docVersion="9" lastPageNumber="394" masterDocId="FFB8FB75FFB2FFA3FF9CFFC4006BB223" masterDocTitle="Genus Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) from Laos, with descriptions of five new species" masterLastPageNumber="398" masterPageNumber="390" pageNumber="394" updateTime="1698733430596" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="02117665530A95E23BBBAAFC08651881">Genus Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) from Laos, with descriptions of five new species</mods:title>
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<mods:namePart id="47FA9064ADF85B8347A8008A30D6A839">Liu, Chen</mods:namePart>
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<mods:namePart id="C885D7311B757A56E94BC23246FF6D8B">Wang, Shuxia</mods:namePart>
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<mods:date id="5B0ECD0AD1321AC39DB4091BC27F7E48">2020</mods:date>
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<mods:number id="35DA18E6D809647A07B66C4AB9741082">2020-03-24</mods:number>
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<mods:number id="EB5A092F3AA44C9FAF190C298076EC4A">4755</mods:number>
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<treatment id="0381830DFFB6FFA7FF0BFE8901EEB7C4" ID-DOI="http://doi.org/10.5281/zenodo.3812481" ID-GBIF-Taxon="163392069" ID-Zenodo-Dep="3812481" LSID="urn:lsid:plazi:treatment:0381830DFFB6FFA7FF0BFE8901EEB7C4" httpUri="http://treatment.plazi.org/id/0381830DFFB6FFA7FF0BFE8901EEB7C4" lastPageNumber="394" pageId="4" pageNumber="394">
<subSubSection id="C3326190FFB6FFA7FF0BFE890231B344" box="[151,602,333,360]" pageId="4" pageNumber="394" type="nomenclature">
<paragraph id="8B97321BFFB6FFA7FF0BFE890231B344" blockId="4.[151,602,333,395]" box="[151,602,333,360]" pageId="4" pageNumber="394">
<heading id="D0DF8577FFB6FFA7FF0BFE890231B344" bold="true" box="[151,602,333,360]" fontSize="10" level="2" pageId="4" pageNumber="394" reason="2">
<emphasis id="B95CEE09FFB6FFA7FF0BFE890231B344" bold="true" box="[151,602,333,360]" pageId="4" pageNumber="394">
<taxonomicName id="4C284998FFB6FFA7FF0BFE890184B344" authority="Wang" authorityName="Liu &amp; Wang" authorityYear="2020" box="[151,495,333,360]" class="Insecta" family="Oecophoridae" genus="Promalactis" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="394" phylum="Arthropoda" rank="species" species="serrulata" status="sp. nov.">
<emphasis id="B95CEE09FFB6FFA7FF0BFE8901F6B34B" bold="true" box="[151,413,333,360]" italics="true" pageId="4" pageNumber="394">Promalactis serrulata</emphasis>
Wang
</taxonomicName>
,
<taxonomicNameLabel id="A26F5372FFB6FFA7FE60FE890231B344" box="[508,602,333,359]" pageId="4" pageNumber="394" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3326190FFB6FFA7FF0BFEB5014AB3A8" box="[151,289,368,395]" pageId="4" pageNumber="394" type="description">
<paragraph id="8B97321BFFB6FFA7FF0BFEB5014AB3A8" blockId="4.[151,602,333,395]" box="[151,289,368,395]" pageId="4" pageNumber="394">
(
<figureCitation id="13132E9EFFB6FFA7FF03FEB50084B3A8" box="[159,239,368,395]" captionStart="FIGURES 1‒8" captionStartId="5.[151,264,1881,1906]" captionTargetBox="[179,1435,179,1854]" captionTargetId="figure@5.[151,1435,179,1854]" captionTargetPageId="5" captionText="FIGURES 1‒8. Adults of Promalactis spp. 1, P. unidentalis, holotype ♂; 2, P. tenuiclavifera, holotype ♂; 3, P. introflexa, holotype ♂; 4, P. bolikhamsaiana, holotype ♂; 5, P. serrulata, holotype ♂; 6, P. carinata Du, Li et Wang, 2011; 7, P. flavidia Kim, 2018; 8, P. apicuncata Wang, 2018. Scales=2 mm." figureDoi="http://doi.org/10.5281/zenodo.3733723" httpUri="https://zenodo.org/record/3733723/files/figure.png" pageId="4" pageNumber="394">Figs. 5</figureCitation>
,
<figureCitation id="13132E9EFFB6FFA7FF67FEB50173B3A8" box="[251,280,369,395]" captionStart="FIGURES 9‒14" captionStartId="6.[151,264,1898,1923]" captionTargetBox="[248,1351,190,1853]" captionTargetId="figure@6.[226,1361,181,1872]" captionTargetPageId="6" captionText="FIGURES 9‒14. Genitalia of Promalactis spp. 9, P. unidentalis, holotype ♂, slide No. LC19247; 10, P. tenuiclavifera, holotype ♂, slide No. LC19241; 11, P. introflexa, holotype ♂, slide No. LC19242; 12a, P. introflexa, paratype ♀, slide No. LC19302, 12b, enlarged signum, slide No. LC19246; 13, P. bolikhamsaiana, holotype ♂, slide No. LC19262; 14, P. serrulata, holotype ♂, slide No. LC19250. Scales=0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3733725" httpUri="https://zenodo.org/record/3733725/files/figure.png" pageId="4" pageNumber="394">14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3326190FFB6FFA7FF0BFE7D0394B3D4" pageId="4" pageNumber="394" type="materials_examined">
<paragraph id="8B97321BFFB6FFA7FF0BFE7D0394B3D4" blockId="4.[151,1437,440,1511]" pageId="4" pageNumber="394">
<emphasis id="B95CEE09FFB6FFA7FF0BFE7D012FB3F0" bold="true" box="[151,324,440,467]" pageId="4" pageNumber="394">Type material.</emphasis>
<materialsCitation id="3B403846FFB6FFA7FED6FE7D0394B3D4" ID-GBIF-Occurrence="2592567757" collectingDate="2019-08-03" collectorName="YS Bae &amp; DJ Lee &amp; No. LC" country="Laos" elevation="272" latitude="18.4" location="Phou Khao Khouay National Protected Area" longLatPrecision="765" longitude="103.09" pageId="4" pageNumber="394" specimenCount="1" specimenCount-male="1" stateProvince="Bolikhamsai" typeStatus="holotype">
<typeStatus id="54938CB9FFB6FFA7FED6FE7D01D9B3F0" box="[330,434,441,467]" pageId="4" pageNumber="394" type="holotype">Holotype</typeStatus>
<specimenCount id="9D2EF992FFB6FFA7FE25FE7D01BBB3F0" box="[441,464,441,467]" pageId="4" pageNumber="394" type="male">♂</specimenCount>
,
<location id="8EF764C0FFB6FFA7FE47FE7D03A3B3F0" LSID="urn:lsid:plazi:treatment:0381830DFFB6FFA7FF0BFE8901EEB7C4:8EF764C0FFB6FFA7FE47FE7D03A3B3F0" box="[475,968,441,467]" country="Laos" latitude="18.4" longLatPrecision="765" longitude="103.09" name="Phou Khao Khouay National Protected Area" pageId="4" pageNumber="394" stateProvince="Bolikhamsai">Phou Khao Khouay National Protected Area</location>
(
<geoCoordinate id="EE1C54DCFFB6FFA7FC4BFE7D0452B3F0" box="[983,1081,441,467]" degrees="18.40" direction="north" orientation="latitude" pageId="4" pageNumber="394" precision="555" value="18.4">18.40°N</geoCoordinate>
,
<geoCoordinate id="EE1C54DCFFB6FFA7FBD8FE7D04DAB3F0" box="[1092,1201,441,467]" degrees="103.09" direction="east" orientation="longitude" pageId="4" pageNumber="394" precision="555" value="103.09">103.09°E</geoCoordinate>
),
<collectingRegion id="49ECFCF9FFB6FFA7FB58FE7D053FB3F0" box="[1220,1364,441,467]" country="Laos" name="Bolikhamxai" pageId="4" pageNumber="394">Bolikhamsai</collectingRegion>
,
<collectingCountry id="F33F728BFFB6FFA7FAC3FE7D05F3B3F0" box="[1375,1432,441,467]" name="Laos" pageId="4" pageNumber="394">Laos</collectingCountry>
,
<quantity id="4CD09FFEFFB6FFA7FF0BFE19008BB3D4" box="[151,224,477,503]" metricMagnitude="2" metricUnit="m" metricValue="2.7199999999999998" pageId="4" pageNumber="394" unit="m" value="272.0">
<elevation id="0005D528FFB6FFA7FF0BFE19008BB3D4" box="[151,224,477,503]" metricMagnitude="2" metricUnit="m" metricValue="2.7199999999999998" pageId="4" pageNumber="394" unit="m" value="272.0">272 m</elevation>
</quantity>
,
<date id="FF9614DBFFB6FFA7FF70FE190119B3D4" box="[236,370,477,503]" pageId="4" pageNumber="394" value="2019-08-03">
<collectingDate id="EFD2ED33FFB6FFA7FF70FE190119B3D4" box="[236,370,477,503]" pageId="4" pageNumber="394" value="2019-08-03">3.VIII.2019</collectingDate>
</date>
, coll.
<collectorName id="26DD57CDFFB6FFA7FE29FE190265B3D4" box="[437,526,477,503]" pageId="4" pageNumber="394">YS Bae</collectorName>
, MJ Qi &amp;
<collectorName id="26DD57CDFFB6FFA7FD18FE1902BFB3D4" box="[644,724,477,503]" pageId="4" pageNumber="394">DJ Lee</collectorName>
<emphasis id="B95CEE09FFB6FFA7FD47FE1A0378B3D4" box="[731,787,477,503]" italics="true" pageId="4" pageNumber="394">et al.</emphasis>
, slide
<collectorName id="26DD57CDFFB6FFA7FCC1FE1903D8B3D4" box="[861,947,476,503]" pageId="4" pageNumber="394">No. LC</collectorName>
19250.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3326190FFB6FFA7FF5BFDC40457B0A4" pageId="4" pageNumber="394" type="diagnosis">
<paragraph id="8B97321BFFB6FFA7FF5BFDC40457B0A4" blockId="4.[151,1437,440,1511]" pageId="4" pageNumber="394">
<emphasis id="B95CEE09FFB6FFA7FF5BFDC40129B038" bold="true" box="[199,322,512,539]" pageId="4" pageNumber="394">Diagnosis.</emphasis>
This new species is similar to
<taxonomicName id="4C284998FFB6FFA7FD07FDC603CDB038" authority="Wang, 2018" authorityName="Wang" authorityYear="2018" box="[667,934,513,539]" class="Insecta" family="Oecophoridae" genus="Promalactis" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="394" phylum="Arthropoda" rank="species" species="folivalva">
<emphasis id="B95CEE09FFB6FFA7FD07FDC60373B038" box="[667,792,513,539]" italics="true" pageId="4" pageNumber="394">P. folivalva</emphasis>
<bibRefCitation id="EFB94FEAFFB6FFA7FC83FDC503CDB038" author="Wang" box="[799,934,513,539]" firstAuthor="Wang" pageId="4" pageNumber="394" pagination="151 - 160" refId="ref5063" refString="Wang, S. X. &amp; Jia, Y. Y. (2018) Review of the genus Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) III. The trilineata species group, with descriptions of four new species. Zootaxa, 4433 (1), 151 - 160. https: // doi. org / 10.11646 / zootaxa. 4433.1.9" type="journal article" year="2018">
Wang,
<quantity id="4CD09FFEFFB6FFA7FCF2FDC503CDB038" box="[878,934,513,539]" metricMagnitude="1" metricUnit="m" metricValue="5.125719999999999" pageId="4" pageNumber="394" unit="in" value="2018.0">2018</quantity>
</bibRefCitation>
</taxonomicName>
in the male genitalia. It can be distinguished from
<taxonomicName id="4C284998FFB6FFA7FF4AFDE2013FB01C" authorityName="Wang" authorityYear="2018" box="[214,340,549,575]" class="Insecta" family="Oecophoridae" genus="Promalactis" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="394" phylum="Arthropoda" rank="species" species="folivalva">
<emphasis id="B95CEE09FFB6FFA7FF4AFDE2013FB01C" box="[214,340,549,575]" italics="true" pageId="4" pageNumber="394">P. folivalva</emphasis>
</taxonomicName>
by the costa of the valva serrate distally, the juxta without distinct lateral arms and the aedeagus with a nail-like cornutus; whereas in
<taxonomicName id="4C284998FFB6FFA7FDA0FD8E02D0B040" authorityName="Wang" authorityYear="2018" box="[572,699,585,611]" class="Insecta" family="Oecophoridae" genus="Promalactis" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="394" phylum="Arthropoda" rank="species" species="folivalva">
<emphasis id="B95CEE09FFB6FFA7FDA0FD8E02D0B040" box="[572,699,585,611]" italics="true" pageId="4" pageNumber="394">P. folivalva</emphasis>
</taxonomicName>
, the costa of the valva is smooth, the juxta has long lateral arms, and the aedeagus has a long cornutus bearing 3‒4 apical spines (
<bibRefCitation id="EFB94FEAFFB6FFA7FCF4FDA9045AB0A4" author="Jia &amp; Wang" box="[872,1073,621,647]" firstAuthor="Jia" pageId="4" pageNumber="394" pagination="151 - 160" refId="ref5063" refString="Wang, S. X. &amp; Jia, Y. Y. (2018) Review of the genus Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) III. The trilineata species group, with descriptions of four new species. Zootaxa, 4433 (1), 151 - 160. https: // doi. org / 10.11646 / zootaxa. 4433.1.9" type="journal article" year="2018">Jia &amp; Wang 2018</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3326190FFB6FFA7FF5BFD5401FAB758" pageId="4" pageNumber="394" type="description">
<paragraph id="8B97321BFFB6FFA7FF5BFD540281B088" blockId="4.[151,1437,440,1511]" box="[199,746,656,683]" pageId="4" pageNumber="394">
<emphasis id="B95CEE09FFB6FFA7FF5BFD540132B088" bold="true" box="[199,345,656,683]" pageId="4" pageNumber="394">Description.</emphasis>
Adult (
<figureCitation id="13132E9EFFB6FFA7FE33FD550198B088" box="[431,499,656,683]" captionStart="FIGURES 1‒8" captionStartId="5.[151,264,1881,1906]" captionTargetBox="[179,1435,179,1854]" captionTargetId="figure@5.[151,1435,179,1854]" captionTargetPageId="5" captionText="FIGURES 1‒8. Adults of Promalactis spp. 1, P. unidentalis, holotype ♂; 2, P. tenuiclavifera, holotype ♂; 3, P. introflexa, holotype ♂; 4, P. bolikhamsaiana, holotype ♂; 5, P. serrulata, holotype ♂; 6, P. carinata Du, Li et Wang, 2011; 7, P. flavidia Kim, 2018; 8, P. apicuncata Wang, 2018. Scales=2 mm." figureDoi="http://doi.org/10.5281/zenodo.3733723" httpUri="https://zenodo.org/record/3733723/files/figure.png" pageId="4" pageNumber="394">Fig. 5</figureCitation>
). Wingspan 10.0 mm.
</paragraph>
<paragraph id="8B97321BFFB6FFA7FF5BFD710184B118" blockId="4.[151,1437,440,1511]" pageId="4" pageNumber="394">
<emphasis id="B95CEE09FFB6FFA7FF5BFD71016FB0EC" box="[199,260,693,719]" italics="true" pageId="4" pageNumber="394">Head</emphasis>
with vertex cream white, frons silvery grey, occiput rust brown. Labial palpus with second segment pale ochreous yellow on outer surface, orange yellow on inner surface, with a white dot before apex; third segment black tinged with pale ochreous yellow, white at apex. Antenna with scape cream white; flagellum black annulated with cream white on dorsal surface.
</paragraph>
<paragraph id="8B97321BFFB6FFA7FF5BFC810389B614" blockId="4.[151,1437,440,1511]" pageId="4" pageNumber="394">
<emphasis id="B95CEE09FFB6FFA7FF5BFC810173B17C" box="[199,280,837,863]" italics="true" pageId="4" pageNumber="394">Thorax</emphasis>
and tegula rust brown, thorax with cream white spot distally. Forewing rust brown, darker between basal and antemedian fasciae; markings white edged with black scales: costal spot at distal 1/3, inverted sub-triangular, extending to above lower margin of cell before lower angle; basal fascia slender, from above base of fold oblique outwardly to dorsum; antemedian fascia from before basal 1/4 of costal margin to basal 3/5 of dorsum, oblique outward, parallel with basal fascia, evenly wide; dorsal streak from before end of fold oblique outward to below costal spot, straight; apical spot ovate, with dense black scales on inner margin; cilia greyish brown mixed with yellowish brown except white on extension of apical spot. Hindwing and cilia brown.
</paragraph>
<paragraph id="8B97321BFFB6FFA7FF5BFB8504CFB774" blockId="4.[151,1437,440,1511]" pageId="4" pageNumber="394">
<emphasis id="B95CEE09FFB6FFA7FF5BFB850105B678" box="[199,366,1089,1115]" italics="true" pageId="4" pageNumber="394">Male genitalia</emphasis>
(
<figureCitation id="13132E9EFFB6FFA7FEE2FB8501B8B678" box="[382,467,1089,1115]" captionStart="FIGURES 9‒14" captionStartId="6.[151,264,1898,1923]" captionTargetBox="[248,1351,190,1853]" captionTargetId="figure@6.[226,1361,181,1872]" captionTargetPageId="6" captionText="FIGURES 9‒14. Genitalia of Promalactis spp. 9, P. unidentalis, holotype ♂, slide No. LC19247; 10, P. tenuiclavifera, holotype ♂, slide No. LC19241; 11, P. introflexa, holotype ♂, slide No. LC19242; 12a, P. introflexa, paratype ♀, slide No. LC19302, 12b, enlarged signum, slide No. LC19246; 13, P. bolikhamsaiana, holotype ♂, slide No. LC19262; 14, P. serrulata, holotype ♂, slide No. LC19250. Scales=0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3733725" httpUri="https://zenodo.org/record/3733725/files/figure.png" pageId="4" pageNumber="394">Fig. 14</figureCitation>
). Uncus wide at base, narrowed to pointed apex, with sparse setae laterally. Gnathos shorter than uncus; gnathos plate sub-triangular, narrowed to rounded apex; lateral arm shorter, wide. Tegumen bifurcate from posterior 1/3; lateral arm narrowed anteriorly. Valva foliate, widened medially, narrowed from middle to rounded apex, with a short strong apical spine; costa arched, serrate distally; sacculus narrow, setose distally. Saccus shorter than uncus, straight on posterior margin, wide at base, narrowed to posterior 1/3, then abruptly narrowed to pointed apex. Juxta sub-rectangular, concave at middle on posterior margin, forming two triangular posterolateral lobes pointed at apex; basal lobe inverted sub-triangular, small, less than 1/3 length of saccus. Aedeagus slightly longer than valva, uniformly strong; cornutus strong, nail-like, about 1/6 length of aedeagus.
</paragraph>
<paragraph id="8B97321BFFB6FFA7FF5BFAA501FAB758" blockId="4.[151,1437,440,1511]" box="[199,401,1377,1403]" pageId="4" pageNumber="394">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="C3326190FFB6FFA7FF5BFA400227B7BC" box="[199,588,1412,1439]" pageId="4" pageNumber="394" type="distribution">
<paragraph id="8B97321BFFB6FFA7FF5BFA400227B7BC" blockId="4.[151,1437,440,1511]" box="[199,588,1412,1439]" pageId="4" pageNumber="394">
<emphasis id="B95CEE09FFB6FFA7FF5BFA400132B7BC" bold="true" box="[199,345,1412,1439]" pageId="4" pageNumber="394">Distribution</emphasis>
.
<collectingCountry id="F33F728BFFB6FFA7FEFBFA4101F6B7BC" box="[359,413,1413,1439]" name="Laos" pageId="4" pageNumber="394">Laos</collectingCountry>
(
<collectingRegion id="49ECFCF9FFB6FFA7FE30FA410250B7BC" box="[428,571,1413,1439]" country="Laos" name="Bolikhamxai" pageId="4" pageNumber="394">Bolikhamsai</collectingRegion>
<emphasis id="B95CEE09FFB6FFA7FDA0FA40022EB7BC" bold="true" box="[572,581,1412,1439]" pageId="4" pageNumber="394">)</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C3326190FFB6FFA7FF5BFA6D01EEB7C4" pageId="4" pageNumber="394" type="etymology">
<paragraph id="8B97321BFFB6FFA7FF5BFA6D01EEB7C4" blockId="4.[151,1437,440,1511]" pageId="4" pageNumber="394">
<emphasis id="B95CEE09FFB6FFA7FF5BFA6D0125B7E0" bold="true" box="[199,334,1449,1475]" pageId="4" pageNumber="394">Etymology.</emphasis>
The epithet of this new species is derived from the Latin
<emphasis id="B95CEE09FFB6FFA7FC78FA6D043DB7E0" box="[996,1110,1449,1475]" italics="true" pageId="4" pageNumber="394">serrulatus</emphasis>
, referring to the costa of the valva serrate distally.
</paragraph>
</subSubSection>
</treatment>
</document>
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