treatments-xml/data/CF/46/87/CF46879EFFB3FF8AFF5B706EFC592CBA.xml
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<document id="67F2BD1067C27290CEF6F96702C5F887" ID-CLB-Dataset="82886" ID-DOI="10.1080/00222933.2021.2017499" ID-GBIF-Dataset="5cd01cfe-52bb-4092-97b2-804c3c7d145d" ID-ISSN="1464-5262" ID-Zenodo-Dep="6758313" ID-ZooBank="D766FC5E-CEC8-4184-BE90-C25350B90612" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1656311158681" checkinUser="plazi" docAuthor="Costa, Leidiane O., Duarte, Paulo R. M., Iannuzzi, Luciana &amp; Grossi, Paschoal C." docDate="2022" docId="CF46879EFFB3FF8AFF5B706EFC592CBA" docLanguage="en" docName="JNATHIST.56.1-4.191-225.pdf" docOrigin="Journal of Natural History 56 (1 - 4)" docSource="http://dx.doi.org/10.1080/00222933.2021.2017499" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Gibboryctes gracilicornis Dupuis and Dechambre 2008" docType="treatment" docVersion="7" lastPageNumber="216" masterDocId="337FFFE6FFAAFF90FFF57271FF852A10" masterDocTitle="Taxonomic revision and notes on natural history of the enigmatic beetle genus Gibboryctes Endrödi (Coleoptera: Melolonthidae: Dynastinae)" masterLastPageNumber="225" masterPageNumber="191" pageNumber="215" updateTime="1699352275692" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2213D1507A17D846F8C8CF0877D8C365">Taxonomic revision and notes on natural history of the enigmatic beetle genus Gibboryctes Endrödi (Coleoptera: Melolonthidae: Dynastinae)</mods:title>
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<mods:namePart id="8C2043F96B98F3C6098657800118993E">Costa, Leidiane O.</mods:namePart>
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<mods:namePart id="521C558153BC963F3FD4BA3A7A5B0A41">Iannuzzi, Luciana</mods:namePart>
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<treatment id="CF46879EFFB3FF8AFF5B706EFC592CBA" ID-DOI="http://doi.org/10.5281/zenodo.6761932" ID-GBIF-Taxon="196400129" ID-Zenodo-Dep="6761932" LSID="urn:lsid:plazi:treatment:CF46879EFFB3FF8AFF5B706EFC592CBA" httpUri="http://treatment.plazi.org/id/CF46879EFFB3FF8AFF5B706EFC592CBA" lastPageId="26" lastPageNumber="216" pageId="25" pageNumber="215">
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<paragraph id="47503688FFB3FF89FF5B706EFBF02829" blockId="25.[174,1141,543,569]" box="[174,1141,543,569]" pageId="25" pageNumber="215">
<heading id="1C1881E4FFB3FF89FF5B706EFBF02829" bold="true" box="[174,1141,543,569]" centered="true" fontSize="11" level="2" pageId="25" pageNumber="215" reason="3">
<treatmentCitationGroup id="67FF11A6FFB3FF89FF5B706EFBF02829" box="[174,1141,543,569]" pageId="25" pageNumber="215">
<emphasis id="759BEA9AFFB3FF89FF5B706EFBF02829" bold="true" box="[174,1141,543,569]" pageId="25" pageNumber="215">
<treatmentCitation id="C64E1099FFB3FF89FF5B706EFC052829" author="Dupuis F &amp; Dechambre RP" box="[174,896,543,569]" page="22" pageId="25" pageNumber="215" year="2008">
<taxonomicName id="80EF4D0BFFB3FF89FF5B706EFC052829" authority="Dupuis and Dechambre 2008: 22" authorityName="Dupuis and Dechambre" authorityPageNumber="22" authorityYear="2008" box="[174,896,543,569]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="215" phylum="Arthropoda" rank="species" species="gracilicornis">
Gibboryctes gracilicornis
<bibRefCitation id="237E4B79FFB3FF89FE1A706EFC052829" author="Dupuis F &amp; Dechambre RP" box="[495,896,543,569]" pageId="25" pageNumber="215" pagination="7 - 25" refId="ref13302" refString="Dupuis F, Dechambre RP. 2008. Les Heterogomphus du groupe pauson (Perty, 1930) (Coleoptera, Dynastinae). Coleopteres. 14: 7 - 25." type="journal article" year="2008">
<emphasis id="759BEA9AFFB3FF89FE1A706EFC052829" bold="true" box="[495,896,543,569]" italics="true" pageId="25" pageNumber="215">Dupuis and Dechambre 2008: 22</emphasis>
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</taxonomicName>
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<emphasis id="759BEA9AFFB3FF89FC7D706EFBF02829" bold="true" box="[904,1141,543,569]" italics="true" pageId="25" pageNumber="215">(new combination).</emphasis>
</emphasis>
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</heading>
</paragraph>
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<subSubSection id="0FF56503FFB3FF89FF557021FDD228BF" pageId="25" pageNumber="215" type="materials_examined">
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<emphasis id="759BEA9AFFB3FF89FF557021FEC1287A" bold="true" box="[160,324,592,618]" italics="true" pageId="25" pageNumber="215">
<typeStatus id="9854882AFFB3FF89FF557021FF5D287A" box="[160,216,592,618]" pageId="25" pageNumber="215">Type</typeStatus>
material
</emphasis>
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<paragraph id="47503688FFB3FF89FF557002FDD228BF" blockId="25.[160,1155,592,687]" pageId="25" pageNumber="215">
<taxonomicName id="80EF4D0BFFB3FF89FF557002FDD3289C" authority="Prell, 1912" authorityName="Prell" authorityYear="1912" box="[160,598,627,652]" class="Insecta" family="Dynastidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="215" phylum="Arthropoda" rank="species" species="gracilicornis">
<emphasis id="759BEA9AFFB3FF89FF557002FE5D289C" box="[160,472,627,652]" italics="true" pageId="25" pageNumber="215">Heterogomphus gracilicornis</emphasis>
<bibRefCitation id="237E4B79FFB3FF89FE157002FDD3289C" author="Prell H" box="[480,598,627,652]" pageId="25" pageNumber="215" pagination="93 - 176" refId="ref13871" refString="Prell H. 1912. Revision des Dynastinen-Genus Heterogomphus Burm. Mem Soc Entomol Belg. 20: 93 - 176." type="journal article" year="1912">Prell, 1912</bibRefCitation>
</taxonomicName>
,
<typeStatus id="9854882AFFB3FF89FD917002FD4D289C" box="[612,712,627,652]" pageId="25" pageNumber="215" type="holotype">holotype</typeStatus>
not examined.
<taxonomicName id="80EF4D0BFFB3FF89FC887002FEB928BF" authority="Endrodi, 1978" authorityName="Endrodi" authorityYear="1978" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="215" phylum="Arthropoda" rank="species" species="acuminatus">
<emphasis id="759BEA9AFFB3FF89FC887002FB06289C" box="[893,1155,627,652]" italics="true" pageId="25" pageNumber="215">Gibboryctes acuminatus</emphasis>
<bibRefCitation id="237E4B79FFB3FF89FF5570E7FEB928BF" author="Endrodi S" box="[160,316,662,687]" pageId="25" pageNumber="215" pagination="79 - 82" refId="ref13423" refString="Endrodi S. 1978. Neue Dynastinen aus Amerika (Coleoptera, Melolonthidae). Mitt Zool Mus Berlin. 54: 79 - 82. doi: 10.1002 / mmnz. 19780540103." type="journal article" year="1978">Endrödi, 1978</bibRefCitation>
</taxonomicName>
,
<typeStatus id="9854882AFFB3FF89FEBD70E7FE2928BF" box="[328,428,662,687]" pageId="25" pageNumber="215" type="holotype">holotype</typeStatus>
not examined.
</paragraph>
</subSubSection>
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<emphasis id="759BEA9AFFB3FF89FF5570AAFEFC28E5" bold="true" box="[160,377,731,757]" italics="true" pageId="25" pageNumber="215">Non-type material</emphasis>
</paragraph>
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<paragraph id="47503688FFB3FF89FF55708EFBA9294D" blockId="25.[160,1156,731,861]" pageId="25" pageNumber="215">
<materialsCitation id="F7873CD5FFB3FF89FF55708EFC0B2908" ID-GBIF-Occurrence="3819562313" box="[160,910,767,792]" collectingDate="2005-12" collectionCode="BCRC" collectorName="La Paz" country="Argentina" location="Mendoza" pageId="25" pageNumber="215" specimenCount="2" specimenCount-male="2" stateProvince="Mendoza">
<collectingCountry id="3FF87618FFB3FF89FF55708EFEA22908" box="[160,295,767,792]" name="Argentina" pageId="25" pageNumber="215">ARGENTINA</collectingCountry>
:
<collectingRegion id="852BF86AFFB3FF89FEC1708EFE1B2908" box="[308,414,767,792]" country="Argentina" name="Mendoza" pageId="25" pageNumber="215">Mendoza</collectingRegion>
,
<collectorName id="EA1A535EFFB3FF89FE59708EFE7D2908" box="[428,504,767,792]" pageId="25" pageNumber="215">La Paz</collectorName>
,
<date id="33511048FFB3FF89FDF3708EFD3E2908" box="[518,699,767,792]" pageId="25" pageNumber="215" value="2005-12">
<collectingDate id="2315E9A0FFB3FF89FDF3708EFD3E2908" box="[518,699,767,792]" pageId="25" pageNumber="215" value="2005-12">December 2005</collectingDate>
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<specimenCount id="51E9FD01FFB3FF89FD28708EFCB32908" box="[733,822,767,792]" pageId="25" pageNumber="215" type="male">2 males</specimenCount>
(
<collectionCode id="21FEAE4DFFB3FF89FCB3708EFC0C2908" box="[838,905,767,792]" country="Taiwan" httpUri="http://grbio.org/cool/cv7g-ikqh" name="Bioresource Collection and Research Center" pageId="25" pageNumber="215">BCRC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="F7873CD5FFB3FF89FC69708EFF35294D" ID-GBIF-Occurrence="3819562326" collectingDate="1962-11" collectingDateMax="2005-01-25" collectingDateMin="1962-11" collectionCode="BCRC" collectorName="El Quebrado &amp; Frias" country="Argentina" location="Choya" pageId="25" pageNumber="215" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Santiago del Estero">
<collectingRegion id="852BF86AFFB3FF89FC69708EFB052908" box="[924,1152,767,792]" country="Argentina" name="Santiago del Estero" pageId="25" pageNumber="215">Santiago Del Estero</collectingRegion>
:
<location id="42306053FFB3FF89FF557150FF6C292A" LSID="urn:lsid:plazi:treatment:CF46879EFFB3FF8AFF5B706EFC592CBA:42306053FFB3FF89FF557150FF6C292A" box="[160,233,801,826]" country="Argentina" name="Choya" pageId="25" pageNumber="215" stateProvince="Santiago del Estero">Choya</location>
,
<collectorName id="EA1A535EFFB3FF89FF067150FE07292A" box="[243,386,801,826]" pageId="25" pageNumber="215">El Quebrado</collectorName>
,
<date id="33511048FFB3FF89FE7E7150FDBA292A" box="[395,575,801,826]" pageId="25" pageNumber="215" value="1962-11">
<collectingDate id="2315E9A0FFB3FF89FE7E7150FDBA292A" box="[395,575,801,826]" pageId="25" pageNumber="215" value="1962-11">November 1962</collectingDate>
</date>
<specimenCount id="51E9FD01FFB3FF89FDAF7150FD36292A" box="[602,691,801,826]" pageId="25" pageNumber="215" type="male">2 males</specimenCount>
,
<specimenCount id="51E9FD01FFB3FF89FD487150FCAD292A" box="[701,808,801,826]" pageId="25" pageNumber="215" type="female">2 females</specimenCount>
(
<collectionCode id="21FEAE4DFFB3FF89FCC07150FCFD292A" box="[821,888,801,826]" country="Taiwan" httpUri="http://grbio.org/cool/cv7g-ikqh" name="Bioresource Collection and Research Center" pageId="25" pageNumber="215">BCRC</collectionCode>
);
<collectorName id="EA1A535EFFB3FF89FC727150FC38292A" box="[903,957,801,826]" pageId="25" pageNumber="215">Frias</collectorName>
,
<date id="33511048FFB3FF89FC327150FB05292A" box="[967,1152,801,826]" pageId="25" pageNumber="215" value="2005-01-25">
<collectingDate id="2315E9A0FFB3FF89FC327150FB05292A" box="[967,1152,801,826]" pageId="25" pageNumber="215" value="2005-01-25">25 January 2005</collectingDate>
</date>
, E
</materialsCitation>
.
<materialsCitation id="F7873CD5FFB3FF89FF4F7135FD77294D" ID-GBIF-Occurrence="3819562328" box="[186,754,836,861]" collectingDate="2007-12" collectionCode="CERPE" collectorName="Pinto" country="Argentina" location="Abadie" pageId="25" pageNumber="215" specimenCount="1" specimenCount-male="1">
<location id="42306053FFB3FF89FF4F7135FE8C294D" LSID="urn:lsid:plazi:treatment:CF46879EFFB3FF8AFF5B706EFC592CBA:42306053FFB3FF89FF4F7135FE8C294D" box="[186,265,836,861]" country="Argentina" name="Abadie" pageId="25" pageNumber="215">Abadie</location>
<specimenCount id="51E9FD01FFB3FF89FED07135FEF4294D" box="[293,369,836,861]" pageId="25" pageNumber="215" type="male">1 male</specimenCount>
(
<collectionCode id="21FEAE4DFFB3FF89FE8B7135FE49294D" box="[382,460,836,861]" pageId="25" pageNumber="215">CERPE</collectionCode>
);
<collectorName id="EA1A535EFFB3FF89FE2E7135FD9D294D" box="[475,536,836,861]" pageId="25" pageNumber="215">Pinto</collectorName>
,
<date id="33511048FFB3FF89FDD67135FD52294D" box="[547,727,836,861]" pageId="25" pageNumber="215" value="2007-12">
<collectingDate id="2315E9A0FFB3FF89FDD67135FD52294D" box="[547,727,836,861]" pageId="25" pageNumber="215" value="2007-12">December 2007</collectingDate>
</date>
, E
</materialsCitation>
.
<materialsCitation id="F7873CD5FFB3FF89FD097135FBAD294D" ID-GBIF-Occurrence="3819562321" box="[764,1064,836,861]" collectionCode="CERPE" country="Argentina" location="Abadie" pageId="25" pageNumber="215" specimenCount="1" specimenCount-female="1">
<location id="42306053FFB3FF89FD097135FCCE294D" LSID="urn:lsid:plazi:treatment:CF46879EFFB3FF8AFF5B706EFC592CBA:42306053FFB3FF89FD097135FCCE294D" box="[764,843,836,861]" country="Argentina" name="Abadie" pageId="25" pageNumber="215">Abadie</location>
<specimenCount id="51E9FD01FFB3FF89FC927135FC4D294D" box="[871,968,836,861]" pageId="25" pageNumber="215" type="female">1 female</specimenCount>
(
<collectionCode id="21FEAE4DFFB3FF89FC207135FBA6294D" box="[981,1059,836,861]" pageId="25" pageNumber="215">CERPE</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0FF56503FFB3FF8AFF5571F8FD492F42" lastPageId="26" lastPageNumber="216" pageId="25" pageNumber="215" type="description">
<paragraph id="47503688FFB3FF89FF5571F8FE0F29B3" blockId="25.[160,1156,905,1728]" box="[160,394,905,931]" pageId="25" pageNumber="215">
<emphasis id="759BEA9AFFB3FF89FF5571F8FE0F29B3" bold="true" box="[160,394,905,931]" pageId="25" pageNumber="215">
Male
<emphasis id="759BEA9AFFB3FF89FF1671F8FE0F29B3" bold="true" box="[227,394,905,931]" italics="true" pageId="25" pageNumber="215">
(
<figureCitation id="DFD42A0DFFB3FF89FF1E71F8FECE29B3" box="[235,331,905,931]" captionStart="Figure 4" captionStartId="7.[160,225,1747,1770]" captionTargetBox="[171,1108,143,1711]" captionTargetId="figure-3@7.[166,1149,142,1715]" captionTargetPageId="7" captionText="Figure 4. Heterogomphus gracilicornis Prell, 1912. (a,b) Male habitus in dorsal and lateral views, respectively; (c,d) female habitus in dorsal and lateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.6758325" httpUri="https://zenodo.org/record/6758325/files/figure.png" pageId="25" pageNumber="215">Figure 4</figureCitation>
(a,b))
</emphasis>
</emphasis>
</paragraph>
<paragraph id="47503688FFB3FF8AFF5571DDFC2729F7" blockId="25.[160,1156,905,1728]" lastBlockId="26.[160,1156,142,999]" lastPageId="26" lastPageNumber="216" pageId="25" pageNumber="215">
<emphasis id="759BEA9AFFB3FF89FF5571DDFEC329D6" bold="true" box="[160,326,940,966]" italics="true" pageId="25" pageNumber="215">Redescription.</emphasis>
<emphasis id="759BEA9AFFB3FF89FEA371DCFE2E29D6" bold="true" box="[342,427,941,966]" pageId="25" pageNumber="215">Length</emphasis>
:
<quantity id="80179B6DFFB3FF89FE4D71DCFDDF29D6" box="[440,602,941,966]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" metricValueMax="2.63" metricValueMin="2.57" pageId="25" pageNumber="215" unit="mm" value="26.0" valueMax="26.3" valueMin="25.7">25.726.3 mm</quantity>
.
<emphasis id="759BEA9AFFB3FF89FD9171DCFD2829D6" bold="true" box="[612,685,941,966]" pageId="25" pageNumber="215">Width</emphasis>
: 12.0
<quantity id="80179B6DFFB3FF89FD0071DCFCD929D6" box="[757,860,941,966]" metricMagnitude="-2" metricUnit="m" metricValue="1.24" pageId="25" pageNumber="215" unit="mm" value="12.4">12.4 mm</quantity>
.
<emphasis id="759BEA9AFFB3FF89FC9371DCFC3329D6" bold="true" box="[870,950,941,966]" pageId="25" pageNumber="215">Colour</emphasis>
: Head, pronotum, legs and venter black; elytra reddish brown.
<emphasis id="759BEA9AFFB3FF89FD6971BEFD5E29F8" bold="true" box="[668,731,975,1000]" pageId="25" pageNumber="215">Head</emphasis>
: Subtriangular, provided with a long, curved backward horn at middle in lateral view. Clypeus triangular, acuminate anteriorly, densely punctate, moderately setose. Horn wide at basis (covering the clypeus in dorsal view), gradually narrowing towards apex; posterior surface weakly lobed at middle in lateral view; surface moderately punctate; punctures oval, scattered about 3 diameters of punctures. Frons smooth; interocular width equals 3.3 times transverse eye diameters. Ocular cantus triangular, with acute outer corners; surface rugose, scarcely setose on outer corners.
<emphasis id="759BEA9AFFB3FF89FEB376B3FE562ECB" bold="true" box="[326,467,1218,1243]" pageId="25" pageNumber="215">Mouthparts</emphasis>
: Labrum transverse, subrectangular, with a fringe of bristles on apical margin (
<figureCitation id="DFD42A0DFFB3FF89FE857694FE4A2EEE" box="[368,463,1253,1278]" captionStart="Figure 6" captionStartId="9.[160,225,1090,1113]" captionTargetBox="[166,1145,146,1050]" captionTargetId="figure-188@9.[166,1149,142,1058]" captionTargetPageId="9" captionText="Figure 6. Labrum in dorsal view. (a) Gibboryctes szelenyii Endrödi; (b) Gibboryctes ebeninus sp. nov.; (c) Gibboryctes endroedii sp. nov.; (d) Heterogomphus gracilicornis Prell; (e) Tetragus waldenfelsi (Endrödi)." figureDoi="http://doi.org/10.5281/zenodo.6758331" httpUri="https://zenodo.org/record/6758331/files/figure.png" pageId="25" pageNumber="215">Figure 6</figureCitation>
(d)). Mandibles with 2 lobed teeth raised laterally to clypeus; teeth deeply separated by a V-shaped notch; subapical tooth wider compared to apical tooth; molar area striated (
<figureCitation id="DFD42A0DFFB3FF89FE11775BFDC22F53" box="[484,583,1322,1347]" captionStart="Figure 7" captionStartId="10.[160,225,1156,1179]" captionTargetBox="[169,1149,148,1090]" captionTargetId="figure-156@10.[166,1149,142,1125]" captionTargetPageId="10" captionText="Figure 7. Right mandible in dorsal view. (a) Gibboryctes szelenyii Endrödi; (b) Gibboryctes ebeninus sp. nov.; (c) Gibboryctes endroedii sp. nov.; (d) Heterogomphus gracilicornis Prell; (e) Tetragus waldenfelsi (Endrödi). Red arrows indicate the apical tooth in (ae), striate molar in (d)." figureDoi="http://doi.org/10.5281/zenodo.6758333" httpUri="https://zenodo.org/record/6758333/files/figure.png" pageId="25" pageNumber="215">Figure 7</figureCitation>
(d)). Maxillary galea triangular, with 3 triangular, pointed teeth at inner margin (1 apical, 1 medial, 1 basal) (
<figureCitation id="DFD42A0DFFB3FF89FCA1773CFC312F76" box="[852,948,1357,1382]" captionStart="Figure 8" captionStartId="11.[160,225,1161,1184]" captionTargetBox="[167,1127,143,1126]" captionTargetId="figure-61@11.[166,1149,142,1129]" captionTargetPageId="11" captionText="Figure 8. Right maxilla in ventral view. (a) Gibboryctes szelenyii Endrödi; (b) Gibboryctes ebeninus sp. nov.; (c) Gibboryctes endroedii sp. nov.; (d) Heterogomphus gracilicornis Prell; (e) Tetragus waldenfelsi (Endrödi). Red arrows indicating triangular galea in (a), elongate maxillary palpomere II in (b), lobed inner margin of galea and stipe with triangular lobe in (c), pointed and triangular teeth on galea in (d), scythe-shape d teeth and rounded inner margin of galea in (e)." figureDoi="http://doi.org/10.5281/zenodo.6758335" httpUri="https://zenodo.org/record/6758335/files/figure.png" pageId="25" pageNumber="215">Figure 8</figureCitation>
(d)). Papomeres I IV combined as long as maxilla in length. Labium subtriangular, slightly rounded laterally on basal half, weakly bilobed at apex; surface densely rugopunctate; punctures large, deep and coalescent; sides covered with innumerous erect bristles, disc glabrous.
<emphasis id="759BEA9AFFB3FF89FF5577A6FE912FE0" bold="true" box="[160,276,1495,1520]" pageId="25" pageNumber="215">Antennae</emphasis>
: Provided with 10 segments; club oval, subequal in length to anteromeres IIVII combined.
<emphasis id="759BEA9AFFB3FF89FE94778BFE302C03" bold="true" box="[353,437,1530,1555]" pageId="25" pageNumber="215">Thorax</emphasis>
: Pronotum subpentagonal in dorsal view, with sinuous lateral margins in lateral view; anterior margin with a deep, V-shaped notch at middle; posterior margin sinuous, with no border; posterior discal area with a strong, apically bifurcate horn in dorsal view (
<figureCitation id="DFD42A0DFFB3FF89FE547413FD812C6B" box="[417,516,1634,1659]" captionStart="Figure 9" captionStartId="12.[160,225,1146,1169]" captionTargetBox="[184,1146,142,1069]" captionTargetId="figure-59@12.[166,1149,142,1114]" captionTargetPageId="12" captionText="Figure 9. Detail of head, pronotum and prothoracic legs of males in dorsal view: (a) Gibboryctes szelenyii (Endrödi); (b) Gibboryctes endroedii sp. nov.; (c) Heterogomphus gracilicornis Prell; (d) Tetragus waldenfelsi (Endrödi)." figureDoi="http://doi.org/10.5281/zenodo.6758337" httpUri="https://zenodo.org/record/6758337/files/figure.png" pageId="25" pageNumber="215">Figure 9</figureCitation>
(c)); horn curved frontward, wide at basis, narrowing towards apex; anterior discal surface smooth; anterolateral area strongly rugose (aureola oposita), that extends posteriorly in 2 regions (on horn basis, on posterior corners) separated by a finely punctate area; dorsal surface of horn moderately covered with shallow punctures (separated by about 3 diameter of punctures), becoming finely punctate on posterior area; posterior margin coarsely rugose. Scutellar plate parabolic, densely covered with large, deep, setigerous, irregularly scattered punctures. Elytral surface weakly covered with small, shallow, irregularly scattered punctures; sutural stria well marked; discal and lateral striae barely marked; epipleuron separated from elytral lateral edge by a deep groove on anterior area in dorsal view, becoming close posteriorly.
<emphasis id="759BEA9AFFB0FF8AFEDF73F0FEE42B8A" bold="true" box="[298,353,385,410]" pageId="26" pageNumber="216">Legs</emphasis>
: Protarsi with sickle-shaped claws; tarsomere V clavate, as short as tarsomeres IIIV combined; tarsomeres IIV beaker-shaped, subequal in length. Protibiae with 4 teeth on outer margins (1 apical, 2 medial, 1 basal); apical tooth curved; medial teeth slightly narrow, elongate (1.47 times longer than width). Mesotarsi with tarsomeres IIV beaker-shaped, gradually decreasing in size. Mesotibial inner surface densely covered with long bristles; outer surface with 2 transverse (1 basal, 1 medial), crenulate carinae; apex truncated; carinae and apex covered with setae. Metatarsi similar to mesotarsi. Metatibiae wide apically compared with the mesotibiae.
<emphasis id="759BEA9AFFB0FF8AFC5A7002FBA0289C" bold="true" box="[943,1061,627,652]" pageId="26" pageNumber="216">Abdomen</emphasis>
: Tergite VIII transverse, strongly convex in lateral view, densely covered with setigerous punctures on sides and anteriorly; discal area glabrous. Ventrites IIV densely rugopunctate and setose on sides, weakly puncate and glabrous on disc; ventrite VI emarginate at middle of posterior margin, bordered with setae posteriorly, weakly punctate on sides, smooth on disc.
<emphasis id="759BEA9AFFB0FF8AFEF37150FED2292A" bold="true" box="[262,343,801,826]" pageId="26" pageNumber="216">Venter</emphasis>
: Prosternal process subtriangular, glabrous. Mesoventrite completely, densely setose. Metaventrite smooth longitudinally on disc.
<emphasis id="759BEA9AFFB0FF8AFC947132FC5D294C" bold="true" box="[865,984,835,860]" pageId="26" pageNumber="216">Aedeagus</emphasis>
: Parameres, in dorsal view, symmetrical, smooth, oval, rounded at basal half, gradually convergent towards apical half (
<figureCitation id="DFD42A0DFFB0FF8AFE7E71F8FE7D29B2" box="[395,504,905,930]" captionStart="Figure 10" captionStartId="13.[160,226,1164,1187]" captionTargetBox="[169,1145,145,1132]" captionTargetId="figure-89@13.[166,1149,142,1133]" captionTargetPageId="13" captionText="Figure 10. Aedeagus with parameres in dorsal and lateral views, respectively. (a,b) Gibboryctes szelenyii Endrödi; (c,d) Gibboryctes endroedii sp. nov.; (e,f) Heterogomphus gracilicornis Prell; (g,h) Tetragus waldenfelsi (Endrödi)." figureDoi="http://doi.org/10.5281/zenodo.6758339" httpUri="https://zenodo.org/record/6758339/files/figure.png" pageId="26" pageNumber="216">Figure 10</figureCitation>
(e)). Parameres, in lateral view, shorter (1.8 times shorter than phalobasis), subtriangular, dorsoventrally wide at basal half, becoming gradually flattened towards apical half (
<figureCitation id="DFD42A0DFFB0FF8AFE0071BFFDE729F7" box="[501,610,974,999]" captionStart="Figure 10" captionStartId="13.[160,226,1164,1187]" captionTargetBox="[169,1145,145,1132]" captionTargetId="figure-89@13.[166,1149,142,1133]" captionTargetPageId="13" captionText="Figure 10. Aedeagus with parameres in dorsal and lateral views, respectively. (a,b) Gibboryctes szelenyii Endrödi; (c,d) Gibboryctes endroedii sp. nov.; (e,f) Heterogomphus gracilicornis Prell; (g,h) Tetragus waldenfelsi (Endrödi)." figureDoi="http://doi.org/10.5281/zenodo.6758339" httpUri="https://zenodo.org/record/6758339/files/figure.png" pageId="26" pageNumber="216">Figure 10</figureCitation>
(f)), ventrally without carina.
</paragraph>
<paragraph id="47503688FFB0FF8AFF557652FE212E2D" blockId="26.[160,1156,1059,1362]" box="[160,420,1059,1085]" pageId="26" pageNumber="216">
<emphasis id="759BEA9AFFB0FF8AFF557652FE212E2D" bold="true" box="[160,420,1059,1085]" pageId="26" pageNumber="216">
Female
<emphasis id="759BEA9AFFB0FF8AFEF57652FE212E2D" bold="true" box="[256,420,1059,1085]" italics="true" pageId="26" pageNumber="216">
(
<figureCitation id="DFD42A0DFFB0FF8AFEFC7652FEED2E2D" box="[265,360,1059,1085]" captionStart="Figure 4" captionStartId="7.[160,225,1747,1770]" captionTargetBox="[171,1108,143,1711]" captionTargetId="figure-3@7.[166,1149,142,1715]" captionTargetPageId="7" captionText="Figure 4. Heterogomphus gracilicornis Prell, 1912. (a,b) Male habitus in dorsal and lateral views, respectively; (c,d) female habitus in dorsal and lateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.6758325" httpUri="https://zenodo.org/record/6758325/files/figure.png" pageId="26" pageNumber="216">Figure 4</figureCitation>
(c,d))
</emphasis>
</emphasis>
</paragraph>
<paragraph id="47503688FFB0FF8AFF557634FD492F42" blockId="26.[160,1156,1059,1362]" pageId="26" pageNumber="216">
<emphasis id="759BEA9AFFB0FF8AFF557634FEC32E4F" bold="true" box="[160,326,1093,1119]" italics="true" pageId="26" pageNumber="216">Redescription.</emphasis>
<emphasis id="759BEA9AFFB0FF8AFEA37637FE2F2E4F" bold="true" box="[342,426,1094,1119]" pageId="26" pageNumber="216">Length</emphasis>
: 26.0
<quantity id="80179B6DFFB0FF8AFE047637FDD02E4F" box="[497,597,1094,1119]" metricMagnitude="-2" metricUnit="m" metricValue="2.66" pageId="26" pageNumber="216" unit="mm" value="26.6">26.6 mm</quantity>
.
<emphasis id="759BEA9AFFB0FF8AFDAB7637FD232E4F" bold="true" box="[606,678,1094,1119]" pageId="26" pageNumber="216">Width</emphasis>
: 12.713.0 mm. Quite distinct from male in the following aspects:
<emphasis id="759BEA9AFFB0FF8AFE5E7618FE6F2E92" bold="true" box="[427,490,1129,1154]" pageId="26" pageNumber="216">Head</emphasis>
: Surface with a short, transverse tubercle (
<figureCitation id="DFD42A0DFFB0FF8AFC1C7618FBD22E92" box="[1001,1111,1129,1154]" captionStart="Figure 11" captionStartId="14.[160,225,1441,1464]" captionTargetBox="[168,1146,144,1407]" captionTargetId="figure-42@14.[166,1149,142,1410]" captionTargetPageId="14" captionText="Figure 11. Detail of head, pronotum and prothoracic legs of females in dorsal view: (a) Gibboryctes szelenyii Endrödi;(b) Gibboryctes ebeninus sp. nov.; (c) Gibboryctes endroedi sp.nov.; (d) Heterogomphus gracilicornis Prell; (e) Tetragus waldenfelsi (Endrödi). Arrow indicates punctate frons in (b)." figureDoi="http://doi.org/10.5281/zenodo.6758341" httpUri="https://zenodo.org/record/6758341/files/figure.png" pageId="26" pageNumber="216">Figure 11</figureCitation>
(d)); ocular canthus wide, with rounded anterior margin.
<emphasis id="759BEA9AFFB0FF8AFCF376FDFCDF2EB5" bold="true" box="[774,858,1164,1189]" pageId="26" pageNumber="216">Thorax</emphasis>
: Pronotum quadrangular, with no horn, with a weak notch at middle of anterior margin (
<figureCitation id="DFD42A0DFFB0FF8AFC7776DFFC6B2ED7" box="[898,1006,1198,1223]" captionStart="Figure 11" captionStartId="14.[160,225,1441,1464]" captionTargetBox="[168,1146,144,1407]" captionTargetId="figure-42@14.[166,1149,142,1410]" captionTargetPageId="14" captionText="Figure 11. Detail of head, pronotum and prothoracic legs of females in dorsal view: (a) Gibboryctes szelenyii Endrödi;(b) Gibboryctes ebeninus sp. nov.; (c) Gibboryctes endroedi sp.nov.; (d) Heterogomphus gracilicornis Prell; (e) Tetragus waldenfelsi (Endrödi). Arrow indicates punctate frons in (b)." figureDoi="http://doi.org/10.5281/zenodo.6758341" httpUri="https://zenodo.org/record/6758341/files/figure.png" pageId="26" pageNumber="216">Figure 11</figureCitation>
(d)); pronotal surface densely covered with large, deep, ocellate, coalescent punctures (
<figureCitation id="DFD42A0DFFB0FF8AFC1976A0FBDD2EFA" box="[1004,1112,1233,1258]" captionStart="Figure 11" captionStartId="14.[160,225,1441,1464]" captionTargetBox="[168,1146,144,1407]" captionTargetId="figure-42@14.[166,1149,142,1410]" captionTargetPageId="14" captionText="Figure 11. Detail of head, pronotum and prothoracic legs of females in dorsal view: (a) Gibboryctes szelenyii Endrödi;(b) Gibboryctes ebeninus sp. nov.; (c) Gibboryctes endroedi sp.nov.; (d) Heterogomphus gracilicornis Prell; (e) Tetragus waldenfelsi (Endrödi). Arrow indicates punctate frons in (b)." figureDoi="http://doi.org/10.5281/zenodo.6758341" httpUri="https://zenodo.org/record/6758341/files/figure.png" pageId="26" pageNumber="216">Figure 11</figureCitation>
(d)); elytral surface densely punctate.
<emphasis id="759BEA9AFFB0FF8AFDC77685FDEF2F1D" bold="true" box="[562,618,1268,1293]" pageId="26" pageNumber="216">Legs</emphasis>
: Protibial dorsal surface rugose; outer teeth rounded at apex (
<figureCitation id="DFD42A0DFFB0FF8AFE987767FE522F3F" box="[365,471,1302,1327]" captionStart="Figure 11" captionStartId="14.[160,225,1441,1464]" captionTargetBox="[168,1146,144,1407]" captionTargetId="figure-42@14.[166,1149,142,1410]" captionTargetPageId="14" captionText="Figure 11. Detail of head, pronotum and prothoracic legs of females in dorsal view: (a) Gibboryctes szelenyii Endrödi;(b) Gibboryctes ebeninus sp. nov.; (c) Gibboryctes endroedi sp.nov.; (d) Heterogomphus gracilicornis Prell; (e) Tetragus waldenfelsi (Endrödi). Arrow indicates punctate frons in (b)." figureDoi="http://doi.org/10.5281/zenodo.6758341" httpUri="https://zenodo.org/record/6758341/files/figure.png" pageId="26" pageNumber="216">Figure 11</figureCitation>
(d)).
<emphasis id="759BEA9AFFB0FF8AFDF97767FD042F3F" bold="true" box="[524,641,1302,1327]" pageId="26" pageNumber="216">Abdomen</emphasis>
: Tergite VIII flattened in lateral view; ventrite VI subtriangular, with complete posterior margin.
</paragraph>
</subSubSection>
<subSubSection id="0FF56503FFB0FF8AFF5577FCFE7B2C22" pageId="26" pageNumber="216" type="discussion">
<paragraph id="47503688FFB0FF8AFF5577FCFE832FB7" blockId="26.[160,1155,1421,1586]" box="[160,262,1421,1447]" pageId="26" pageNumber="216">
<emphasis id="759BEA9AFFB0FF8AFF5577FCFE832FB7" bold="true" box="[160,262,1421,1447]" italics="true" pageId="26" pageNumber="216">Remarks</emphasis>
</paragraph>
<paragraph id="47503688FFB0FF8AFF5577C0FE7B2C22" blockId="26.[160,1155,1421,1586]" pageId="26" pageNumber="216">
It was not possible to access the
<typeStatus id="9854882AFFB0FF8AFDFB77C0FDC52FDA" box="[526,576,1457,1482]" pageId="26" pageNumber="216">type</typeStatus>
specimens of
<taxonomicName id="80EF4D0BFFB0FF8AFD1577C0FCFA2FDA" box="[736,895,1457,1482]" class="Insecta" family="Dynastidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="216" phylum="Arthropoda" rank="species" species="gracilicornis">
<emphasis id="759BEA9AFFB0FF8AFD1577C0FCFA2FDA" box="[736,895,1457,1482]" italics="true" pageId="26" pageNumber="216">H. gracilicornis</emphasis>
</taxonomicName>
and
<taxonomicName id="80EF4D0BFFB0FF8AFC4277C0FBD72FDA" authorityName="Endrodi" authorityYear="1978" box="[951,1106,1457,1482]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="216" phylum="Arthropoda" rank="species" species="acuminatus">
<emphasis id="759BEA9AFFB0FF8AFC4277C0FBD72FDA" box="[951,1106,1457,1482]" italics="true" pageId="26" pageNumber="216">G. acuminatus</emphasis>
</taxonomicName>
. For this reason, the additional specimens were identified through the descriptions, highresolution images, and information from the
<typeStatus id="9854882AFFB0FF8AFD437787FD6D2C1F" box="[694,744,1526,1551]" pageId="26" pageNumber="216">type</typeStatus>
labels of both species available in
<bibRefCitation id="237E4B79FFB0FF8AFF557468FE7F2C22" author="Dupuis F &amp; Dechambre RP" box="[160,506,1561,1586]" pageId="26" pageNumber="216" pagination="7 - 25" refId="ref13302" refString="Dupuis F, Dechambre RP. 2008. Les Heterogomphus du groupe pauson (Perty, 1930) (Coleoptera, Dynastinae). Coleopteres. 14: 7 - 25." type="journal article" year="2008">Dupuis and Dechambre (2008)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0FF56503FFB0FF8AFF55741CFC592CBA" pageId="26" pageNumber="216" type="distribution">
<paragraph id="47503688FFB0FF8AFF55741CFDD92C97" blockId="26.[160,988,1645,1706]" box="[160,604,1645,1671]" pageId="26" pageNumber="216">
<emphasis id="759BEA9AFFB0FF8AFF55741CFDD92C97" bold="true" box="[160,604,1645,1671]" pageId="26" pageNumber="216">
Geographic distribution
<emphasis id="759BEA9AFFB0FF8AFE39741CFDD92C97" bold="true" box="[460,604,1645,1671]" italics="true" pageId="26" pageNumber="216">
(
<figureCitation id="DFD42A0DFFB0FF8AFE20741CFDD12C97" box="[469,596,1645,1671]" captionStart="Figure 14" captionStartId="17.[160,227,1388,1411]" captionTargetBox="[168,1147,145,1355]" captionTargetId="figure-89@17.[166,1149,142,1357]" captionTargetPageId="17" captionText="Figure 14. Geographic distribution maps. (a) Tetragus species; (b) Gibboryctes species; (c) Heterogomphus gracilicornis Prell." figureDoi="http://doi.org/10.5281/zenodo.6758347" httpUri="https://zenodo.org/record/6758347/files/figure.png" pageId="26" pageNumber="216">Figure 14C</figureCitation>
)
</emphasis>
</emphasis>
</paragraph>
<paragraph id="47503688FFB0FF8AFF5574E0FC592CBA" blockId="26.[160,988,1645,1706]" box="[160,988,1681,1706]" pageId="26" pageNumber="216">
<materialsCitation id="F7873CD5FFB0FF8AFF5574E0FC5D2CBA" ID-GBIF-Occurrence="3819562314" box="[160,984,1681,1706]" country="Argentina" location="Mendoza" pageId="26" pageNumber="216" specimenCount="1" stateProvince="Mendoza">
Only known from
<collectingCountry id="3FF87618FFB0FF8AFE9974E0FE5E2CBA" box="[364,475,1681,1706]" name="Argentina" pageId="26" pageNumber="216">Argentina</collectingCountry>
(
<collectingRegion id="852BF86AFFB0FF8AFE1D74E0FDD12CBA" box="[488,596,1681,1706]" country="Argentina" name="Mendoza" pageId="26" pageNumber="216">Mendoza</collectingRegion>
,
<collectingRegion id="852BF86AFFB0FF8AFDAA74E0FCBD2CBA" box="[607,824,1681,1706]" country="Argentina" name="Santiago del Estero" pageId="26" pageNumber="216">Santiago del Estero</collectingRegion>
[new record])
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>