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<mods:title id="967B503AC524D9496908A4AEC7790B68">Lectotype designation, redescription, and synonymy of Homonyx cupreus Guérin-Méneville, 1839 (Coleoptera: Scarabaeidae: Rutelinae: Rutelini)</mods:title>
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<mods:namePart id="02D1C28D6C963C9F65964709FD71BD95">Mondaca, José</mods:namePart>
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<treatment id="A2429413E773C368BEE0F1AD0EFE6B9E" ID-DOI="http://doi.org/10.5281/zenodo.6524016" ID-GBIF-Taxon="195194959" ID-Zenodo-Dep="6524016" LSID="urn:lsid:plazi:treatment:A2429413E773C368BEE0F1AD0EFE6B9E" httpUri="http://treatment.plazi.org/id/A2429413E773C368BEE0F1AD0EFE6B9E" lastPageId="4" lastPageNumber="411" pageId="1" pageNumber="408">
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<paragraph id="2A542505E773C36DBEE0F1AD0F8F695A" blockId="1.[151,676,1770,1833]" box="[151,676,1770,1797]" pageId="1" pageNumber="408">
<heading id="711C9269E773C36DBEE0F1AD0F8F695A" bold="true" box="[151,676,1770,1797]" fontSize="11" level="1" pageId="1" pageNumber="408" reason="1">
<taxonomicName id="EDEB5E86E773C36DBEE0F1AD0F8F695A" authority="Guerin-Meneville, 1839" authorityName="Guerin-Meneville" authorityYear="1839" box="[151,676,1770,1797]" class="Insecta" family="Rutelidae" genus="Homonyx" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="408" phylum="Arthropoda" rank="species" species="cupreus">
<emphasis id="189FF917E773C36DBEE0F1AD0F8F695A" bold="true" box="[151,676,1770,1797]" pageId="1" pageNumber="408">
<emphasis id="189FF917E773C36DBEE0F1AD0C5E695A" bold="true" box="[151,373,1771,1797]" italics="true" pageId="1" pageNumber="408">Homonyx cupreus</emphasis>
<bibRefCitation id="4E7A58F4E773C36DBF0AF1AC0F8F695A" author="Guerin-Meneville, F. E." box="[381,676,1770,1797]" pageId="1" pageNumber="408" pagination="295 - 305" refId="ref3525" refString="Guerin-Meneville, F. E. (1839) Description de quelques Coleopteres des cotes du detroit de Magellan. Revue Zoologique, par la Societe Cuvierienne, 1839, 295 - 305." type="journal article" year="1839">Guérin-Méneville, 1839</bibRefCitation>
</emphasis>
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<paragraph id="2A542505E773C36DBEE0F0490C336976" blockId="1.[151,676,1770,1833]" box="[151,280,1807,1833]" pageId="1" pageNumber="408">
(
<figureCitation id="B2D03980E773C36DBED7F0490C3B6976" box="[160,272,1807,1833]" captionStart="FIGURE 1" captionStartId="3.[151,250,1687,1711]" captionTargetBox="[169,1421,429,1654]" captionTargetId="figure-123@3.[156,1431,417,1664]" captionTargetPageId="3" captionText="FIGURE 1. Homonyx cupreus Guérin-Méneville, 1839. A, Lectotype female of Homonyx cupreus; B, labels, mouthparts and ventrite of A; C, female from Minas Gerais, Brazil; D, labels of C; E, lectotype of Homonyx chalceus bahianus Ohaus, 1913 (= H. cupreus); F, labels, mouthparts and aedeagus of E; G, aedeagus, lateral and caudal views of E; H, male from Bahia, Brazil; I, labels of H; J, aedeagus, lateral and caudal views of H." figureDoi="http://doi.org/10.5281/zenodo.6524018" httpUri="https://zenodo.org/record/6524018/files/figure.png" pageId="1" pageNumber="408">Fig. 1AJ</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="62F1768EE773C36DBEE0F01209E269F6" pageId="1" pageNumber="408" type="reference_group">
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<treatmentCitation id="AB4A0314E773C36DBEE0F0120F466933" author="Guerin-Meneville, F. E." box="[151,621,1876,1900]" page="300" pageId="1" pageNumber="408" year="1839">
<taxonomicName id="EDEB5E86E773C36DBEE0F0120F466933" authority="Guerin-Meneville 1839: 300" authorityName="Guerin-Meneville" authorityPageNumber="300" authorityYear="1839" box="[151,621,1876,1900]" class="Insecta" family="Rutelidae" genus="Homonyx" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="408" phylum="Arthropoda" rank="species" species="cupreus">
<emphasis id="189FF917E773C36DBEE0F0120C606934" box="[151,331,1876,1899]" italics="true" pageId="1" pageNumber="408">Homonyx cupreus</emphasis>
<bibRefCitation id="4E7A58F4E773C36DBF26F0120F466933" author="Guerin-Meneville, F. E." box="[337,621,1876,1900]" pageId="1" pageNumber="408" pagination="295 - 305" refId="ref3525" refString="Guerin-Meneville, F. E. (1839) Description de quelques Coleopteres des cotes du detroit de Magellan. Revue Zoologique, par la Societe Cuvierienne, 1839, 295 - 305." type="journal article" year="1839">Guérin-Méneville 1839:300</bibRefCitation>
</taxonomicName>
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(original description);
<treatmentCitation id="AB4A0314E773C36DBD38F012090D6933" author="Burmeister, H. C. C." box="[847,1062,1876,1900]" page="391" pageId="1" pageNumber="408" year="1844">
<bibRefCitation id="4E7A58F4E773C36DBD38F012090D6933" author="Burmeister, H. C. C." box="[847,1062,1876,1900]" pageId="1" pageNumber="408" refId="ref3482" refString="Burmeister, H. C. C. (1844) Handbuch der Entomologie (Coleoptera Lamellicornia Anthobia et Phyllophaga Systellochela). Vol. 4. Part 1. T. C. F. Enslin, Berlin, 588 pp." type="book" year="1844">Burmeister 1844: 391</bibRefCitation>
</treatmentCitation>
(catalog);
<treatmentCitation id="AB4A0314E773C36DBAF9F012080C6933" author="Solier, A. J. J." box="[1166,1319,1875,1900]" page="94" pageId="1" pageNumber="408" year="1851">
<bibRefCitation id="4E7A58F4E773C36DBAF9F012080C6933" author="Solier, A. J. J." box="[1166,1319,1875,1900]" pageId="1" pageNumber="408" pagination="1 - 285" refId="ref3971" refString="Solier, A. J. J. (1851) Zoologia: Coleopteros. In: Gay, C. (Ed.), Historia fisica y politica de Chile segun documentos adquiridos en esta republica durante doce anos de residencia en ella Paris. Zoologia, 5, 1 - 285." type="journal article" year="1851">Solier 1851: 94</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="AB4A0314E773C36DBB59F0120C1B69D5" author="Lacordaire, J. T." page="361" pageId="1" pageNumber="408" year="1855">
<bibRefCitation id="4E7A58F4E773C36DBB59F0120C1B69D5" author="Lacordaire, J. T." pageId="1" pageNumber="408" refId="ref3612" refString="Lacordaire, J. T. (1855) Histoire Naturelle des Insectes. Genera des Coleopteres 3. Librairie Encyclopedique de Roret, Paris, 594 pp." type="book" year="1855">Lacordaire 1855: 361</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="AB4A0314E773C36DBF4EF0340CF369D5" author="Reed, E. C." box="[313,472,1906,1930]" page="287" pageId="1" pageNumber="408" year="1876">
<bibRefCitation id="4E7A58F4E773C36DBF4EF0340CF369D5" author="Reed, E. C." box="[313,472,1906,1930]" pageId="1" pageNumber="408" pagination="274 - 295" refId="ref3938" refString="Reed, E. C. (1876) Catalogo de los coleopteros de Chile (segunda parte). Anales de la Universidad de Chile, 48, 274 - 295." type="journal article" year="1876">Reed 1876: 287</bibRefCitation>
</treatmentCitation>
(catalog);
<treatmentCitation id="AB4A0314E773C36DBC32F0340FC869D5" author="Ohaus, F." box="[581,739,1906,1930]" page="71" pageId="1" pageNumber="408" year="1934">
<bibRefCitation id="4E7A58F4E773C36DBC32F0340FC869D5" author="Ohaus, F." box="[581,739,1906,1930]" pageId="1" pageNumber="408" pagination="1 - 172" refId="ref3876" refString="Ohaus, F. (1934) Coleoptera lamellicornia, Fam. Scarabaeidae, Subfam. Rutelinae. T. 1: Tribus Rutelini. Genera Insectorum, 199 A, 1 - 172." type="journal article" year="1934">Ohaus 1934: 71</bibRefCitation>
</treatmentCitation>
(catalog);
<treatmentCitation id="AB4A0314E773C36DBD27F034091069D5" author="Blackwelder, R. E." box="[848,1083,1906,1930]" page="237" pageId="1" pageNumber="408" year="1944">
<bibRefCitation id="4E7A58F4E773C36DBD27F034091069D5" author="Blackwelder, R. E." box="[848,1083,1906,1930]" pageId="1" pageNumber="408" pagination="189 - 341" refId="ref3439" refString="Blackwelder, R. E. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. United States National Museum Bulletin, 185 (2), 189 - 341." type="journal article" year="1944">Blackwelder 1944: 237</bibRefCitation>
</treatmentCitation>
(checklist); Machatschke 1970: 19 (checklist);
<treatmentCitation id="AB4A0314E773C36DBF48F0D70CE669F6" author="Soula, M." box="[319,461,1937,1961]" page="9" pageId="1" pageNumber="408" year="2010">
<bibRefCitation id="4E7A58F4E773C36DBF48F0D70CE669F6" author="Soula, M." box="[319,461,1937,1961]" pageId="1" pageNumber="408" refId="ref4026" refString="Soula, M. (2010) Les Coleopteres du Nouveau Monde. Vol. 4. Rutelini 4. Revision des Pelidnotina 4. Une revision des genres Catoclastus, Homonyx. Retour sur quelques especes ou groupe d'especes de Pelidnota. Addenda 2010. (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot; Pelidnotina &quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale. AECFT, Saintry, 66 pp." type="book" year="2010">Soula 2010: 9</bibRefCitation>
</treatmentCitation>
(revision);
<treatmentCitation id="AB4A0314E773C36DBC35F0D70FD069F6" author="Krajcik, M." box="[578,763,1937,1961]" page="129" pageId="1" pageNumber="408" year="2012">
<bibRefCitation id="4E7A58F4E773C36DBC35F0D70FD069F6" author="Krajcik, M." box="[578,763,1937,1961]" pageId="1" pageNumber="408" pagination="1 - 278" refId="ref3588" refString="Krajcik, M. (2012) Checklist of the world Scarabaeoidea. Animma. X, Supplement 5, 1 - 278." type="journal article" year="2012">Krajčík 2012: 129</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="AB4A0314E773C36DBD0EF0D7094A69F6" author="Moore, M. R. &amp; Jameson, M. L. &amp; Garner, B. H. &amp; Audibert, C. &amp; Smith, A. B. T. &amp; Seidel, M." box="[889,1121,1937,1961]" httpUri="http://treatment.plazi.org/id/7FDC71955FD752FCA24B82A601EC5459" page="119" pageId="1" pageNumber="408" year="2017">
<bibRefCitation id="4E7A58F4E773C36DBD0EF0D7094A69F6" author="Moore, M. R. &amp; Jameson, M. L. &amp; Garner, B. H. &amp; Audibert, C. &amp; Smith, A. B. T. &amp; Seidel, M." box="[889,1121,1937,1961]" pageId="1" pageNumber="408" pagination="1 - 349" refId="ref3719" refString="Moore, M. R., Jameson, M. L., Garner, B. H., Audibert, C., Smith, A. B. T. &amp; Seidel, M. (2017) Synopsis of the pelidnotine scarabs (Coleoptera, Scarabaeidae, Rutelinae, Rutelini) and annotated catalogue of the species and subspecies. ZooKeys, 666, 1 - 349. https: // doi. org / 10.3897 / zookeys. 666.9191" type="journal article" year="2017">
Moore
<emphasis id="189FF917E773C36DBDB4F0D40EDB69F6" box="[963,1008,1937,1961]" italics="true" pageId="1" pageNumber="408">et al</emphasis>
. 2017: 119
</bibRefCitation>
</treatmentCitation>
(catalog).
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</paragraph>
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<paragraph id="2A542505E773C36EBEE0F0F60F386E92" blockId="1.[151,1436,1875,2023]" lastBlockId="2.[151,1436,150,573]" lastPageId="2" lastPageNumber="409" pageId="1" pageNumber="408">
Type locality:
<collectingCountry id="52FC6595E773C36DBF51F0F60C4F6997" box="[294,356,1968,1992]" name="Brazil" pageId="1" pageNumber="408">Brazil</collectingCountry>
(
<emphasis id="189FF917E773C36DBF06F0F60CBB6998" box="[369,400,1968,1991]" italics="true" pageId="1" pageNumber="408">i.e.</emphasis>
, localities within the known range of the taxon following Recommendation 76A.1.4 of the
<bibRefCitation id="4E7A58F4E773C36DBB6CF0F60F9269B9" author="International Commission on Zoological Nomenclature" pageId="1" pageNumber="408" refId="ref3559" refString="International Commission on Zoological Nomenclature (1999) International Code of Zoological Nomenclature. 4 th Edition. International Trust for Zoological Nomenclature, London, 306 pp." type="book" year="1999">International Commission on Zoological Nomenclature 1999</bibRefCitation>
). The original
<typeStatus id="F5509BA7E773C36DBD23F0890EAB69B8" box="[852,896,1999,2023]" pageId="1" pageNumber="408">type</typeStatus>
locality of “côtes de détroit de Magellan” (Guérin- Méneville 1839) is erroneous and is here corrected (following Recommendation 76A.2 of the
<bibRefCitation id="4E7A58F4E770C36EBA01F7D10F236E92" author="International Commission on Zoological Nomenclature" pageId="2" pageNumber="409" refId="ref3559" refString="International Commission on Zoological Nomenclature (1999) International Code of Zoological Nomenclature. 4 th Edition. International Trust for Zoological Nomenclature, London, 306 pp." type="book" year="1999">International Commission on Zoological Nomenclature 1999</bibRefCitation>
).
</paragraph>
<paragraph id="2A542505E770C36EBEE0F7920CB26FDA" blockId="2.[151,1436,150,573]" pageId="2" pageNumber="409">
<materialsCitation id="9A832F58E770C36EBEE0F7920CB26FDA" ID-GBIF-Occurrence="3763209305" collectionCode="NHMUK" collectorName="Det. J. Mondaca E." determinerName="Det. J. Mondaca" pageId="2" pageNumber="409" specimenCount="1" specimenCount-female="1" typeStatus="lectotype">
<typeStatus id="F5509BA7E770C36EBEE0F7920DD76EB3" box="[151,252,212,236]" pageId="2" pageNumber="409" type="lectotype">Lectotype</typeStatus>
female (present designation) at
<collectionCode id="4CFABDC0E770C36EBC37F7920F896EB4" box="[576,674,212,235]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="2" pageNumber="409" type="Museum">NHMUK</collectionCode>
, labeled (
<figureCitation id="B2D03980E770C36EBD74F7920E7A6EB3" box="[771,849,212,236]" captionStart="FIGURE 1" captionStartId="3.[151,250,1687,1711]" captionTargetBox="[169,1421,429,1654]" captionTargetId="figure-123@3.[156,1431,417,1664]" captionTargetPageId="3" captionText="FIGURE 1. Homonyx cupreus Guérin-Méneville, 1839. A, Lectotype female of Homonyx cupreus; B, labels, mouthparts and ventrite of A; C, female from Minas Gerais, Brazil; D, labels of C; E, lectotype of Homonyx chalceus bahianus Ohaus, 1913 (= H. cupreus); F, labels, mouthparts and aedeagus of E; G, aedeagus, lateral and caudal views of E; H, male from Bahia, Brazil; I, labels of H; J, aedeagus, lateral and caudal views of H." figureDoi="http://doi.org/10.5281/zenodo.6524018" httpUri="https://zenodo.org/record/6524018/files/figure.png" pageId="2" pageNumber="409">Fig. 1B</figureCitation>
): a) “526”, b) “Magellan” [handwritten on green label], c) “
<emphasis id="189FF917E770C36EBEA5F7B50C0D6F55" box="[210,294,243,266]" italics="true" pageId="2" pageNumber="409">Omonyx</emphasis>
/
<taxonomicName id="EDEB5E86E770C36EBF4CF7B50CA16F55" authorityName="Guerin-Meneville" authorityYear="1839" box="[315,394,243,266]" class="Insecta" family="Rutelidae" genus="Homonyx" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="409" phylum="Arthropoda" rank="species" species="cupreus">
<emphasis id="189FF917E770C36EBF4CF7B50CA16F55" box="[315,394,243,266]" italics="true" pageId="2" pageNumber="409">cupreus</emphasis>
</taxonomicName>
. guer. / Rev. Zool. (Type) / Magellan.” [handwritten on white label], d) “
<taxonomicName id="EDEB5E86E770C36EBA03F7B509FE6F55" authorityName="Guerin-Meneville" authorityYear="1839" box="[1140,1237,243,266]" class="Insecta" family="Rutelidae" genus="Homonyx" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="409" phylum="Arthropoda" rank="genus">
<emphasis id="189FF917E770C36EBA03F7B509FE6F55" box="[1140,1237,243,266]" italics="true" pageId="2" pageNumber="409">Homonyx</emphasis>
</taxonomicName>
/
<taxonomicName id="EDEB5E86E770C36EBA9DF7B508126F55" authorityName="Guerin-Meneville" authorityYear="1839" box="[1258,1337,243,266]" class="Insecta" family="Rutelidae" genus="Homonyx" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="409" phylum="Arthropoda" rank="species" species="cupreus">
<emphasis id="189FF917E770C36EBA9DF7B508126F55" box="[1258,1337,243,266]" italics="true" pageId="2" pageNumber="409">cupreus</emphasis>
</taxonomicName>
/ Guerin- Typus / Magellania” [handwritten on white label], e) “So named / in Reiches / Collection / C.W.” [typeset on white label], f) “Type” [typeset on white circle label, red margin], g) Mouthparts glued on transparent card, h) Sixth ventrite glued on cardboard, I) “
<typeStatus id="F5509BA7E770C36EBF2CF6090CC56F38" box="[347,494,335,359]" pageId="2" pageNumber="409" type="lectotype">LECTOTYPE</typeStatus>
<specimenCount id="3CEDEE8CE770C36EBF82F6090F236F39" box="[501,520,335,358]" pageId="2" pageNumber="409" type="female"></specimenCount>
/
<taxonomicName id="EDEB5E86E770C36EBC6AF6090FFF6F39" authorityName="Guerin-Meneville" authorityYear="1839" box="[541,724,335,358]" class="Insecta" family="Rutelidae" genus="Homonyx" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="409" phylum="Arthropoda" rank="species" species="cupreus">
<emphasis id="189FF917E770C36EBC6AF6090FFF6F39" box="[541,724,335,358]" italics="true" pageId="2" pageNumber="409">Homonyx cupreus</emphasis>
</taxonomicName>
/
<bibRefCitation id="4E7A58F4E770C36EBC9EF6090EF46F38" author="Guerin-Meneville, F. E." box="[745,991,335,359]" pageId="2" pageNumber="409" pagination="295 - 305" refId="ref3525" refString="Guerin-Meneville, F. E. (1839) Description de quelques Coleopteres des cotes du detroit de Magellan. Revue Zoologique, par la Societe Cuvierienne, 1839, 295 - 305." type="journal article" year="1839">Guérin-Méneville, 1839</bibRefCitation>
/
<collectorName id="871E40D3E770C36EBD83F60909926F39" box="[1012,1209,335,359]" pageId="2" pageNumber="409">
<determinerName id="DF88C9C1E770C36EBD83F60909B06F38" box="[1012,1179,335,359]" pageId="2" pageNumber="409">Det. J. Mondaca</determinerName>
E.
</collectorName>
2022” [typeset on red label, black margin].
</materialsCitation>
</paragraph>
<paragraph id="2A542505E770C36EBEE0F6CA0CB26F9C" blockId="2.[151,1436,150,573]" pageId="2" pageNumber="409">
<emphasis id="189FF917E770C36EBEE0F6CA0C1C6FFB" bold="true" box="[151,311,396,420]" pageId="2" pageNumber="409">New synonym.</emphasis>
<taxonomicName id="EDEB5E86E770C36EBF37F6CB0FD86FFB" authority="Ohaus 1913: 495 - 496" authorityName="Ohaus" authorityPageNumber="495 - 496" authorityYear="1913" box="[320,755,396,420]" class="Insecta" family="Rutelidae" genus="Homonyx" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="409" phylum="Arthropoda" rank="species" species="bahianus">
<emphasis id="189FF917E770C36EBF37F6CB0F2D6FFB" box="[320,518,396,420]" italics="true" pageId="2" pageNumber="409">Homonyx bahianus</emphasis>
<bibRefCitation id="4E7A58F4E770C36EBC78F6CA0FD86FFB" author="Ohaus, F" box="[527,755,396,420]" pageId="2" pageNumber="409" pagination="487 - 511" refId="ref3804" refString="Ohaus, F (1913) XI. Beitrag zur Kenntnis der Ruteliden. (Col.). Deutsche Entomologische Zeitschrift, 1913, 487 - 511." type="journal article" year="1913">Ohaus 1913: 495496</bibRefCitation>
</taxonomicName>
[originally
<taxonomicName id="EDEB5E86E770C36EBD18F6CB09656FFB" authorityName="Ohaus" authorityYear="1913" box="[879,1102,396,420]" class="Insecta" family="Rutelidae" genus="Homonyx" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="409" phylum="Arthropoda" rank="subSpecies" species="chalceus" subSpecies="bahianus">
<emphasis id="189FF917E770C36EBD18F6CB09656FFB" box="[879,1102,396,420]" italics="true" pageId="2" pageNumber="409">H. chalceus bahianus</emphasis>
</taxonomicName>
].
<materialsCitation id="9A832F58E770C36EBA12F6CA0CBE6F9C" ID-GBIF-Occurrence="3763209306" country="Brazil" location="San Antonio da Barra" pageId="2" pageNumber="409" specimenCount="1" stateProvince="Bahia">
<typeStatus id="F5509BA7E770C36EBA12F6CA09BC6FFB" box="[1125,1175,396,420]" pageId="2" pageNumber="409">Type</typeStatus>
locality:
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,
<collectingRegion id="E82FEBE7E770C36EBF7BF6ED0C616F9C" box="[268,330,427,451]" country="Brazil" name="Bahia" pageId="2" pageNumber="409">Bahia</collectingRegion>
,
<collectingCountry id="52FC6595E770C36EBF23F6ED0CBE6F9C" box="[340,405,427,451]" name="Brazil" pageId="2" pageNumber="409">Brazil</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="2A542505E770C36EBEE0F68C0E436C62" blockId="2.[151,1436,150,573]" pageId="2" pageNumber="409">
<materialsCitation id="9A832F58E770C36EBEE0F68C0E436C62" ID-GBIF-Occurrence="3763209307" collectionCode="ZMHB" collectorName="Ch. Pujol" determinerName="Soula" location="Antonio da Barra" pageId="2" pageNumber="409" specimenCount="2" stateProvince="Bahia" typeStatus="lectotype">
<typeStatus id="F5509BA7E770C36EBEE0F68C0DD76FBD" box="[151,252,458,482]" pageId="2" pageNumber="409" type="lectotype">Lectotype</typeStatus>
male at
<collectionCode id="4CFABDC0E770C36EBF21F68C0CB46FBE" box="[342,415,458,481]" pageId="2" pageNumber="409">ZMHB</collectionCode>
(designated by
<bibRefCitation id="4E7A58F4E770C36EBC33F68F0F906FBE" author="Soula, M." box="[580,699,457,481]" pageId="2" pageNumber="409" refId="ref4026" refString="Soula, M. (2010) Les Coleopteres du Nouveau Monde. Vol. 4. Rutelini 4. Revision des Pelidnotina 4. Une revision des genres Catoclastus, Homonyx. Retour sur quelques especes ou groupe d'especes de Pelidnota. Addenda 2010. (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot; Pelidnotina &quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale. AECFT, Saintry, 66 pp." type="book" year="2010">Soula 2010</bibRefCitation>
), labeled (
<figureCitation id="B2D03980E770C36EBD51F68F0E586FBE" box="[806,883,457,481]" captionStart="FIGURE 1" captionStartId="3.[151,250,1687,1711]" captionTargetBox="[169,1421,429,1654]" captionTargetId="figure-123@3.[156,1431,417,1664]" captionTargetPageId="3" captionText="FIGURE 1. Homonyx cupreus Guérin-Méneville, 1839. A, Lectotype female of Homonyx cupreus; B, labels, mouthparts and ventrite of A; C, female from Minas Gerais, Brazil; D, labels of C; E, lectotype of Homonyx chalceus bahianus Ohaus, 1913 (= H. cupreus); F, labels, mouthparts and aedeagus of E; G, aedeagus, lateral and caudal views of E; H, male from Bahia, Brazil; I, labels of H; J, aedeagus, lateral and caudal views of H." figureDoi="http://doi.org/10.5281/zenodo.6524018" httpUri="https://zenodo.org/record/6524018/files/figure.png" pageId="2" pageNumber="409">Fig. 1F</figureCitation>
): a) “Typus!” [typeset on red label], b) “S
<location id="2F3473DEE770C36EBB5DF68F0DD46FA0" LSID="urn:lsid:plazi:treatment:A2429413E773C368BEE0F1AD0EFE6B9E:2F3473DEE770C36EBB5DF68F0DD46FA0" name="Antonio da Barra" pageId="2" pageNumber="409" stateProvince="Bahia">Antonio da Barra</location>
/ Prov. de
<collectingRegion id="E82FEBE7E770C36EBF1BF6AE0C8C6C5F" box="[364,423,488,512]" country="Brazil" name="Bahia" pageId="2" pageNumber="409">Bahia</collectingRegion>
/
<collectorName id="871E40D3E770C36EBFCCF6AE0F306C5F" box="[443,539,488,512]" pageId="2" pageNumber="409">Ch. Pujol</collectorName>
1890” [typeset on beige label], c) “
<taxonomicName id="EDEB5E86E770C36EBDF4F6AF09546C5F" authority="Ohs." authorityName="Ohaus" authorityYear="1913" box="[899,1151,488,512]" class="Insecta" family="Rutelidae" genus="Homonyx" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="409" phylum="Arthropoda" rank="species" species="bahianus">
<emphasis id="189FF917E770C36EBDF4F6AF09636C5F" box="[899,1096,488,512]" italics="true" pageId="2" pageNumber="409">Homonyx bahianus</emphasis>
Ohs.
</taxonomicName>
” [handwritten on red label], d) “
<typeStatus id="F5509BA7E770C36EBE9BF5410C7F6C40" box="[236,340,519,543]" pageId="2" pageNumber="409" type="lectotype">Lectotype</typeStatus>
2010 /
<taxonomicName id="EDEB5E86E770C36EBFD6F5410F2D6C40" authorityName="Guerin-Meneville" authorityYear="1839" box="[417,518,519,543]" class="Insecta" family="Rutelidae" genus="Homonyx" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="409" phylum="Arthropoda" rank="genus">Homonyx</taxonomicName>
/
<taxonomicName id="EDEB5E86E770C36EBC6DF5410FB16C40" authority="Oh" authorityName="Ohaus" authorityYear="1913" box="[538,666,519,543]" class="Insecta" family="Rutelidae" genus="Homonyx" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="409" phylum="Arthropoda" rank="species" species="bahianus">bahianus Oh</taxonomicName>
/
<determinerName id="DF88C9C1E770C36EBCD9F5410FC36C40" box="[686,744,519,543]" pageId="2" pageNumber="409">Soula</determinerName>
det” [typeset and handwritten on red label], e) Mouthparts glued on beige cardboard, f) Male genitalia glued on beige cardboard point.
</materialsCitation>
</paragraph>
<paragraph id="2A542505E770C36EBEE0F52D0F156DDD" blockId="2.[151,1437,619,2014]" pageId="2" pageNumber="409">
<emphasis id="189FF917E770C36EBEE0F52D0F2E6CDA" bold="true" box="[151,517,619,645]" pageId="2" pageNumber="409">Additional material examined.</emphasis>
<materialsCitation id="9A832F58E770C36EBC7AF52D0CEE6C91" ID-GBIF-Occurrence="3763209308" collectingDate="2013-11-05" collectionCode="NHMUK, R" collectorName="E.F. Nascimento, J.R. Carvalho, E. Menezes, F. Bravo, D. Cordeiro" country="Brazil" elevation="24493" latitude="-13.55" location="Iacu Fazenda Sossego" longLatPrecision="1290" longitude="-44.25" pageId="2" pageNumber="409" specimenCount="1" specimenCount-male="1" stateProvince="Bahia">
<collectorName id="871E40D3E770C36EBC7AF52D0F686CDA" box="[525,579,619,645]" pageId="2" pageNumber="409">Nine</collectorName>
specimens from:
<emphasis id="189FF917E770C36EBD7AF52D09306CDA" bold="true" box="[781,1051,619,645]" pageId="2" pageNumber="409">
<collectingCountry id="52FC6595E770C36EBD7AF52D0E516CDA" box="[781,890,619,645]" name="Brazil" pageId="2" pageNumber="409">BRAZIL</collectingCountry>
,
<collectingRegion id="E82FEBE7E770C36EBDF0F52D093D6CDA" box="[903,1046,619,645]" country="Brazil" name="Bahia" pageId="2" pageNumber="409">Bahia State</collectingRegion>
:
</emphasis>
<collectorName id="871E40D3E770C36EBA54F52D094C6CDA" box="[1059,1127,619,645]" pageId="2" pageNumber="409">Bahia</collectorName>
,
<collectorName id="871E40D3E770C36EBA04F52A080D6CDA" box="[1139,1318,619,646]" pageId="2" pageNumber="409">Frey Collection</collectorName>
, 1905-100 (
<specimenCount id="3CEDEE8CE770C36EBED5F5D60DDB6CF6" box="[162,240,655,682]" pageId="2" pageNumber="409" type="male">1 male</specimenCount>
,
<collectionCode id="4CFABDC0E770C36EBE8BF5D60C406CF5" box="[252,363,656,682]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="2" pageNumber="409" type="Museum">NHMUK</collectionCode>
);
<location id="2F3473DEE770C36EBF08F5D60F506CF5" LSID="urn:lsid:plazi:treatment:A2429413E773C368BEE0F1AD0EFE6B9E:2F3473DEE770C36EBF08F5D60F506CF5" box="[383,635,655,682]" country="Brazil" name="Iacu Fazenda Sossego" pageId="2" pageNumber="409" stateProvince="Bahia">Iaçú Fazenda Sossego</location>
,
<date id="5E5503C5E770C36EBCF0F5C90E2B6CF5" box="[647,768,655,682]" pageId="2" pageNumber="409" value="2013-11-05">
<collectingDate id="4E11FA2DE770C36EBCF0F5C90E2B6CF5" box="[647,768,655,682]" pageId="2" pageNumber="409" value="2013-11-05">5-XI-2013</collectingDate>
</date>
,
<collectorName id="871E40D3E770C36EBD7BF5D60EE16CF6" box="[780,970,655,682]" pageId="2" pageNumber="409">E.F. Nascimento</collectorName>
, J.
<collectionCode id="4CFABDC0E770C36EBD91F5D60ED66CF5" box="[998,1021,656,682]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="409">R</collectionCode>
.
<collectorName id="871E40D3E770C36EBA7EF5C909596CF6" box="[1033,1138,655,681]" pageId="2" pageNumber="409">Carvalho</collectorName>
, E.
<collectorName id="871E40D3E770C36EBAE8F5D6082E6CF5" box="[1183,1285,656,682]" pageId="2" pageNumber="409">Menezes</collectorName>
, F.
<collectorName id="871E40D3E770C36EBB59F5D6085E6CF5" box="[1326,1397,656,682]" pageId="2" pageNumber="409">Bravo</collectorName>
, D.
<collectorName id="871E40D3E770C36EBEE0F5F50DD06C92" box="[151,251,691,717]" pageId="2" pageNumber="409">Cordeiro</collectorName>
(
<specimenCount id="3CEDEE8CE770C36EBF7AF5F20C716C92" box="[269,346,691,718]" pageId="2" pageNumber="409" type="male">1 male</specimenCount>
,
<collectionCode id="4CFABDC0E770C36EBF11F5F20C926C91" box="[358,441,692,718]" pageId="2" pageNumber="409">CEMT</collectionCode>
);
</materialsCitation>
<materialsCitation id="9A832F58E770C36EBFBAF5F20F3C6CAD" ID-GBIF-Occurrence="3763209302" collectingDate="2013-11-05" collectionCode="NHMUK, R" collectorName="F.B. &amp; T.Z. &amp; A.N. &amp; J.R. &amp; A." country="Brazil" elevation="24493" latitude="-13.55" location="Iacu Fazenda Sossego" longLatPrecision="1290" longitude="-44.25" pageId="2" pageNumber="409" specimenCount="1" specimenCount-female="2" specimenCount-male="1" stateProvince="Bahia">
<location id="2F3473DEE770C36EBFBAF5F20E236C92" LSID="urn:lsid:plazi:treatment:A2429413E773C368BEE0F1AD0EFE6B9E:2F3473DEE770C36EBFBAF5F20E236C92" box="[461,776,691,718]" country="Brazil" latitude="-13.55" longLatPrecision="1290" longitude="-44.25" name="Sao Felix do Coribe-Coribe" pageId="2" pageNumber="409" stateProvince="Bahia">São Félix do Coribe-Coribe</location>
, km
<quantity id="ED1388E0E770C36EBD48F5F20E986C91" box="[831,947,692,718]" metricMagnitude="4" metricUnit="m" metricValue="2.4493" pageId="2" pageNumber="409" unit="m" value="24493.0">
<elevation id="A1C6C236E770C36EBD48F5F20E986C91" box="[831,947,692,718]" metricMagnitude="4" metricUnit="m" metricValue="2.4493" pageId="2" pageNumber="409" unit="m" value="24493.0">24, 493 m</elevation>
</quantity>
,
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,
<geoCoordinate id="4FDF43C2E770C36EBA5DF5F509BB6C91" box="[1066,1168,691,718]" degrees="44" direction="west" minutes="15" orientation="longitude" pageId="2" pageNumber="409" precision="925" value="-44.25">44°15W</geoCoordinate>
,
<date id="5E5503C5E770C36EBAEAF5F208346C91" box="[1181,1311,692,718]" pageId="2" pageNumber="409" value="2007-12-07">
<collectingDate id="4E11FA2DE770C36EBAEAF5F208346C91" box="[1181,1311,692,718]" pageId="2" pageNumber="409" value="2007-12-07">7-XII-2007</collectingDate>
</date>
,
<collectorName id="871E40D3E770C36EBB5CF5F208706C91" box="[1323,1371,692,718]" name="F. Bravo" pageId="2" pageNumber="409">F.B.</collectorName>
,
<collectionCode id="4CFABDC0E770C36EBB10F5F208506C91" box="[1383,1403,692,718]" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="409">T</collectionCode>
.Z., A.N., J.
<collectionCode id="4CFABDC0E770C36EBE98F59E0C2C6CAD" box="[239,263,728,754]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="409">R</collectionCode>
., A. (
<specimenCount id="3CEDEE8CE770C36EBF30F59E0C806CAE" box="[327,427,727,754]" pageId="2" pageNumber="409" type="female">1 female</specimenCount>
,
<collectionCode id="4CFABDC0E770C36EBFCFF59E0F206CAD" box="[440,523,728,754]" pageId="2" pageNumber="409">CEMT</collectionCode>
);
</materialsCitation>
<materialsCitation id="9A832F58E770C36EBC57F5910CF16D49" ID-GBIF-Occurrence="3763209303" collectingDate="2013-11-05" collectionCode="NHMUK, R" collectorName="W. Grossi &amp; Rafael Parizotto" country="Brazil" elevation="24493" latitude="-13.55" location="Iacu Fazenda Sossego" longLatPrecision="1290" longitude="-44.25" pageId="2" pageNumber="409" specimenCount="1" specimenCount-female="2" specimenCount-male="1" stateProvince="Bahia">
<location id="2F3473DEE770C36EBC57F5910F9C6CAE" LSID="urn:lsid:plazi:treatment:A2429413E773C368BEE0F1AD0EFE6B9E:2F3473DEE770C36EBC57F5910F9C6CAE" box="[544,695,727,753]" country="Brazil" latitude="-15.4775" longLatPrecision="21" longitude="-40.915836" name="Encruzilhada" pageId="2" pageNumber="409" stateProvince="Bahia">Encruzilhada</location>
,
<date id="5E5503C5E770C36EBCB4F59E0E786CAD" box="[707,851,728,754]" pageId="2" pageNumber="409" value="2007-12-10">
<collectingDate id="4E11FA2DE770C36EBCB4F59E0E786CAD" box="[707,851,728,754]" pageId="2" pageNumber="409" value="2007-12-10">
10-XII-
<quantity id="ED1388E0E770C36EBD6FF59E0E786CAD" box="[792,851,728,754]" metricMagnitude="6" metricUnit="m" metricValue="2.0076" pageId="2" pageNumber="409" unit="m" value="2007600.0">2007</quantity>
</collectingDate>
</date>
, 600 m,
<geoCoordinate id="4FDF43C2E770C36EBDCDF591096F6CAD" box="[954,1092,727,754]" degrees="15" direction="south" minutes="28" orientation="latitude" pageId="2" pageNumber="409" precision="15" seconds="39" value="-15.4775">15°2839S</geoCoordinate>
,
<geoCoordinate id="4FDF43C2E770C36EBA25F59109CE6CAD" box="[1106,1253,727,754]" degrees="40" direction="west" minutes="54" orientation="longitude" pageId="2" pageNumber="409" precision="15" seconds="57" value="-40.915836">40°5457W</geoCoordinate>
,
<collectorName id="871E40D3E770C36EBA84F59108146CAE" box="[1267,1343,727,753]" pageId="2" pageNumber="409">Grossi</collectorName>
,
<collectorName id="871E40D3E770C36EBB3BF59108B36CAE" box="[1356,1432,727,753]" pageId="2" pageNumber="409">Rafael</collectorName>
,
<collectorName id="871E40D3E770C36EBEE0F5BD0DD76D4A" box="[151,252,763,789]" pageId="2" pageNumber="409">Parizotto</collectorName>
(
<specimenCount id="3CEDEE8CE770C36EBF78F5BA0C586D4A" box="[271,371,763,790]" pageId="2" pageNumber="409" type="female">1 female</specimenCount>
,
<collectionCode id="4CFABDC0E770C36EBF08F5BA0CF96D49" box="[383,466,764,790]" pageId="2" pageNumber="409">CEMT</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="9A832F58E770C36EBF92F5BD0C176D65" ID-GBIF-Occurrence="3763209304" collectingDate="2012-12" collectionCode="R" collectorName="P. C. Grossi &amp; E. J. Grossi" country="Brazil" location="Fazenda Faceiro" municipality="Aguas Vermelhas" pageId="2" pageNumber="409" specimenCount="1" stateProvince="Minas Gerais">
<emphasis id="189FF917E770C36EBF92F5BD0FFE6D4A" bold="true" box="[485,725,763,789]" pageId="2" pageNumber="409">
<collectingRegion id="E82FEBE7E770C36EBF92F5BD0FFB6D4A" box="[485,720,763,789]" country="Brazil" name="Minas Gerais" pageId="2" pageNumber="409">Minas Gerais State</collectingRegion>
:
</emphasis>
<collectingMunicipality id="CA30BF7FE770C36EBCAAF5BB0E8E6D4A" box="[733,933,763,792]" pageId="2" pageNumber="409">Águas Vermelhas</collectingMunicipality>
,
<location id="2F3473DEE770C36EBDC6F5BD09466D4A" LSID="urn:lsid:plazi:treatment:A2429413E773C368BEE0F1AD0EFE6B9E:2F3473DEE770C36EBDC6F5BD09466D4A" box="[945,1133,763,789]" country="Brazil" municipality="Aguas Vermelhas" name="Fazenda Faceiro" pageId="2" pageNumber="409" stateProvince="Minas Gerais">Fazenda Faceiro</location>
,
<date id="5E5503C5E770C36EBA0EF5BD09F26D4A" box="[1145,1241,763,789]" pageId="2" pageNumber="409" value="2012-12">
<collectingDate id="4E11FA2DE770C36EBA0EF5BD09F26D4A" box="[1145,1241,763,789]" pageId="2" pageNumber="409" value="2012-12">xii.2012</collectingDate>
</date>
,
<collectorName id="871E40D3E770C36EBA92F5BA084D6D4A" box="[1253,1382,763,790]" pageId="2" pageNumber="409">P.C. Grossi</collectorName>
,
<collectorName id="871E40D3E770C36EBB05F5BA0DC86D66" pageId="2" pageNumber="409">E.J. Grossi</collectorName>
, G.A.
<collectionCode id="4CFABDC0E770C36EBF51F4660C176D65" box="[294,316,800,826]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="409">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="9A832F58E770C36EBF30F4590F7E6D01" ID-GBIF-Occurrence="3763209301" collectingDate="1997-12" collectionCode="NHMUK, CEMT" collectorName="F.Z. Vaz de Mello &amp; A.M. Bello" country="Brazil" county="Aguas Vermelhas" elevation="2007600" latitude="-15.4775" location="Itaobim" longLatPrecision="21" longitude="-40.915836" municipality="Vaz de Mello" pageId="2" pageNumber="409" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Minas Gerais">
<collectorName id="871E40D3E770C36EBF30F4590C916D65" box="[327,442,799,826]" pageId="2" pageNumber="409">Melo, J.A</collectorName>
.
<collectorName id="871E40D3E770C36EBFB0F4590F386D66" box="[455,531,799,825]" pageId="2" pageNumber="409">Rafael</collectorName>
, at light (
<specimenCount id="3CEDEE8CE770C36EBC08F4660FDB6D66" box="[639,752,799,826]" pageId="2" pageNumber="409" type="female">2 females</specimenCount>
,
<collectionCode id="4CFABDC0E770C36EBC8BF4660E406D65" box="[764,875,800,826]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="2" pageNumber="409" type="Museum">NHMUK</collectionCode>
);
<collectingCounty id="C3355D89E770C36EBD08F467096C6D66" box="[895,1095,799,828]" pageId="2" pageNumber="409">Águas Vermelhas</collectingCounty>
,
<date id="5E5503C5E770C36EBA24F46609966D65" box="[1107,1213,800,826]" pageId="2" pageNumber="409" value="1997-12">
<collectingDate id="4E11FA2DE770C36EBA24F46609966D65" box="[1107,1213,800,826]" pageId="2" pageNumber="409" value="1997-12">XII-1997</collectingDate>
</date>
, F.Z.
<collectingMunicipality id="CA30BF7FE770C36EBA89F46608B36D66" box="[1278,1432,799,826]" pageId="2" pageNumber="409">Vaz de Mello</collectingMunicipality>
, A.M.
<collectorName id="871E40D3E770C36EBEADF4050C3C6D02" box="[218,279,835,861]" pageId="2" pageNumber="409">Bello</collectorName>
(
<specimenCount id="3CEDEE8CE770C36EBF5DF4020C536D02" box="[298,376,835,862]" pageId="2" pageNumber="409" type="male">1 male</specimenCount>
,
<specimenCount id="3CEDEE8CE770C36EBFF2F4020CC16D02" box="[389,490,835,862]" pageId="2" pageNumber="409" type="female">1 female</specimenCount>
,
<collectionCode id="4CFABDC0E770C36EBF81F4020F626D01" box="[502,585,836,862]" pageId="2" pageNumber="409">CEMT</collectionCode>
);
</materialsCitation>
<materialsCitation id="9A832F58E770C36EBC2AF4050F116DDD" ID-GBIF-Occurrence="3763209309" collectingDate="1997-12" collectionCode="NHMUK, CEMT" collectorName="P. C. Grossi &amp; J. A. Rafael &amp; D. Parizotto" country="Brazil" county="Aguas Vermelhas" elevation="2007600" latitude="-15.4775" location="Itaobim" longLatPrecision="21" longitude="-40.915836" municipality="Vaz de Mello" pageId="2" pageNumber="409" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Minas Gerais">
<location id="2F3473DEE770C36EBC2AF4050F9C6D02" LSID="urn:lsid:plazi:treatment:A2429413E773C368BEE0F1AD0EFE6B9E:2F3473DEE770C36EBC2AF4050F9C6D02" box="[605,695,835,861]" country="Brazil" county="Aguas Vermelhas" latitude="-15.4775" longLatPrecision="21" longitude="-40.915836" municipality="Vaz de Mello" name="Itaobim" pageId="2" pageNumber="409" stateProvince="Minas Gerais">Itaobim</location>
,
<date id="5E5503C5E770C36EBCB4F4020E6D6D01" box="[707,838,836,862]" pageId="2" pageNumber="409" value="2007-12-09">
9-XII-
<quantity id="ED1388E0E770C36EBD7CF4020E6D6D01" box="[779,838,836,862]" metricMagnitude="6" metricUnit="m" metricValue="2.0076" pageId="2" pageNumber="409" unit="m" value="2007600.0">
<elevation id="A1C6C236E770C36EBD7CF4020E6D6D01" box="[779,838,836,862]" metricMagnitude="6" metricUnit="m" metricValue="2.0076" pageId="2" pageNumber="409" unit="m" value="2007600.0">2007</elevation>
</quantity>
</date>
, 600 m,
<geoCoordinate id="4FDF43C2E770C36EBDDEF405091F6D01" box="[937,1076,835,862]" degrees="15" direction="south" minutes="28" orientation="latitude" pageId="2" pageNumber="409" precision="15" seconds="39" value="-15.4775">15°2839S</geoCoordinate>
,
<geoCoordinate id="4FDF43C2E770C36EBA36F40509FF6D01" box="[1089,1236,835,862]" degrees="40" direction="west" minutes="54" orientation="longitude" pageId="2" pageNumber="409" precision="15" seconds="57" value="-40.915836">40°5457W</geoCoordinate>
,
<collectorName id="871E40D3E770C36EBA96F40208486D02" box="[1249,1379,835,862]" pageId="2" pageNumber="409">P.C. Grossi</collectorName>
,
<collectorName id="871E40D3E770C36EBB18F4020DC86DDE" pageId="2" pageNumber="409">J.A. Rafael</collectorName>
,
<collectorName id="871E40D3E770C36EBE99F42E0C5E6DDE" box="[238,373,871,898]" pageId="2" pageNumber="409">D. Parizotto</collectorName>
(
<specimenCount id="3CEDEE8CE770C36EBFFFF42E0CFF6DDE" box="[392,468,871,898]" pageId="2" pageNumber="409" type="male">1 male</specimenCount>
,
<collectionCode id="4CFABDC0E770C36EBFA8F42E0F196DDD" box="[479,562,872,898]" pageId="2" pageNumber="409">CEMT</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="62F1768EE770C36FBEB0F4CD0FE96FD6" lastPageId="3" lastPageNumber="410" pageId="2" pageNumber="409" type="description">
<paragraph id="2A542505E770C36FBEB0F4CD0FE96FD6" blockId="2.[151,1437,619,2014]" lastBlockId="3.[151,1436,151,394]" lastPageId="3" lastPageNumber="410" pageId="2" pageNumber="409">
<emphasis id="189FF917E770C36EBEB0F4CD0C5E6DFA" bold="true" box="[199,373,907,933]" pageId="2" pageNumber="409">Redescription.</emphasis>
<emphasis id="189FF917E770C36EBF0AF4CA0CA06DFA" box="[381,395,908,933]" italics="true" pageId="2" pageNumber="409">n</emphasis>
=
<specimenCount id="3CEDEE8CE770C36EBFDDF4CD0F316DFA" box="[426,538,907,933]" pageId="2" pageNumber="409" type="female">6 females</specimenCount>
,
<specimenCount id="3CEDEE8CE770C36EBC51F4CA0FAA6DFA" box="[550,641,907,934]" pageId="2" pageNumber="409" type="male">4 males</specimenCount>
. 19.021.0 mm (females), 12.018.0 mm (males); width 9.010.0 mm (females), 8.011.0 mm (males).
<emphasis id="189FF917E770C36EBC65F4E90F776D95" bold="true" box="[530,604,943,970]" pageId="2" pageNumber="409">
<emphasis id="189FF917E770C36EBC65F4E90F786D95" bold="true" box="[530,595,943,970]" italics="true" pageId="2" pageNumber="409">Color</emphasis>
:
</emphasis>
Dorsal surface coppery red or coppery brown, shiny with metallic green reflections on head, pronotum and legs (
<figureCitation id="B2D03980E770C36EBC2DF4950FDC6DB1" box="[602,759,979,1006]" captionStart="FIGURE 1" captionStartId="3.[151,250,1687,1711]" captionTargetBox="[169,1421,429,1654]" captionTargetId="figure-123@3.[156,1431,417,1664]" captionTargetPageId="3" captionText="FIGURE 1. Homonyx cupreus Guérin-Méneville, 1839. A, Lectotype female of Homonyx cupreus; B, labels, mouthparts and ventrite of A; C, female from Minas Gerais, Brazil; D, labels of C; E, lectotype of Homonyx chalceus bahianus Ohaus, 1913 (= H. cupreus); F, labels, mouthparts and aedeagus of E; G, aedeagus, lateral and caudal views of E; H, male from Bahia, Brazil; I, labels of H; J, aedeagus, lateral and caudal views of H." figureDoi="http://doi.org/10.5281/zenodo.6524018" httpUri="https://zenodo.org/record/6524018/files/figure.png" pageId="2" pageNumber="409">Fig. 1A, C, H</figureCitation>
).
<emphasis id="189FF917E770C36EBD7DF4950E706DB1" bold="true" box="[778,859,979,1006]" italics="true" pageId="2" pageNumber="409">Shape:</emphasis>
Elongate oval, parallel-sided, diverging posteriorly towards end of body; dorsal surface convex, glabrous, venter almost flat, sparsely setose; elytral apex rounded.
<emphasis id="189FF917E770C36EBEE0F35D0DF46A69" bold="true" box="[151,223,1051,1078]" italics="true" pageId="2" pageNumber="409">Head:</emphasis>
Clypeus subtrapezoidal, rounded apically, anterior margin reflexed vertically; dorsal surface slightly convex and glabrous, densely punctate, and transversely striate; punctures small, some confluent. Frontoclypeal suture not impressed, weakly indicated laterally. Frons convex in lateral view, with small, dense punctures. Eyes small, rounded, interocular width equals 5.0 transverse eye diameters, ventrally globose; eye canthus flattened, not carinate. Mandibles exposed beyond clypeal margins, with apex bidentate and laterally with 2 large, rounded, reflexed teeth; molar area wider than long, surface strongly striate. Labrum horizontal, projected beyond clypeal margin, apex bilobed, medially emarginate and setose at margins. Maxilla strongly dentate, with 6 teeth; galea not fused, with moderately long setae; maxillary palps with 4 palpomeres; palpomere 1 shorter than 2; 2 longer than 3, 4 fusiform, longer than 23 together. Labium subquadrate with basal margins rounded, apex emarginate, concave in anterior half and medially sulcate posteriorly; labial palps short, with 3 palpomeres; palpomere 1 shorter than 2; 3 fusiform, longer than 12 together. Antenna with 10 antennomeres in both sexes, apical 3 antennomeres forming club (
<figureCitation id="B2D03980E770C36EBB18F2C50DF56B9D" captionStart="FIGURE 1" captionStartId="3.[151,250,1687,1711]" captionTargetBox="[169,1421,429,1654]" captionTargetId="figure-123@3.[156,1431,417,1664]" captionTargetPageId="3" captionText="FIGURE 1. Homonyx cupreus Guérin-Méneville, 1839. A, Lectotype female of Homonyx cupreus; B, labels, mouthparts and ventrite of A; C, female from Minas Gerais, Brazil; D, labels of C; E, lectotype of Homonyx chalceus bahianus Ohaus, 1913 (= H. cupreus); F, labels, mouthparts and aedeagus of E; G, aedeagus, lateral and caudal views of E; H, male from Bahia, Brazil; I, labels of H; J, aedeagus, lateral and caudal views of H." figureDoi="http://doi.org/10.5281/zenodo.6524018" httpUri="https://zenodo.org/record/6524018/files/figure.png" pageId="2" pageNumber="409">Fig. 1A, C</figureCitation>
); scape claviform; pedicel submoniliform; antennomere 3 subcylindrical; 45 subquadrate or moniliform; 6 cupiform; 6 discoidal; club with 3 antennomeres subequal or slightly longer than funicule.
<emphasis id="189FF917E770C36EBADDF28A08036BB9" bold="true" box="[1194,1320,1484,1510]" pageId="2" pageNumber="409">
<emphasis id="189FF917E770C36EBADDF28A08346BB9" bold="true" box="[1194,1311,1484,1510]" italics="true" pageId="2" pageNumber="409">Pronotum</emphasis>
:
</emphasis>
Widest at base, apical angles acute, basal angles obtuse or acute; dorsal surface glabrous, finely and densely punctate, laterally with a deep circular depression or fovea present only in females; punctures small in size; marginal bead complete, not interrupted basomedially, not setose basolaterally; posterior margin sinuate laterally and basomedially round.
<emphasis id="189FF917E770C36EBEE0F11D0C7F6829" bold="true" box="[151,340,1627,1654]" italics="true" pageId="2" pageNumber="409">Scutellar shield:</emphasis>
Subtriangular, wider than long, apex rounded; surface finely punctate, glabrous.
<emphasis id="189FF917E770C36EBA95F11D08196829" bold="true" box="[1250,1330,1627,1654]" italics="true" pageId="2" pageNumber="409">Elytra:</emphasis>
Together longer than broad, with dorsal surface striate and punctate, glabrous; elytral suture apically angled; humeral umbone with prominent tubercle.
<emphasis id="189FF917E770C36EBFCDF1E20F2568E1" bold="true" box="[442,526,1700,1726]" italics="true" pageId="2" pageNumber="409">Venter:</emphasis>
Punctures sparse, small to moderate. Prosternum small, subtriangular, scarcely projecting between procoxae. Mesoventrite without anterior process. Metaventrite glabrous, sparsely and finely punctate.
<emphasis id="189FF917E770C36EBF74F1AD0C516959" bold="true" box="[259,378,1771,1798]" italics="true" pageId="2" pageNumber="409">Abdomen:</emphasis>
Ventrite 1 short, medially subtriangular; 24 subequal in width; 5 wider than previous ventrites; propygidium shorter than 5 with the apex slightly emarginate in male, straight in female; surface finely, sparsely punctate, with setiferous punctures in transverse rows on posterior margins, laterally with fine transverse striae; pygidium subtriangular, wider than long; surface convex, transversally striate, setose at apex; lateral and caudal margins with complete marginal bead.
<emphasis id="189FF917E770C36EBC31F03A0FAD69C9" bold="true" box="[582,646,1916,1942]" italics="true" pageId="2" pageNumber="409">Legs:</emphasis>
Protibia with 3 external teeth unequally separated, acute distally; distal tooth elongate, externally curved, medial tooth large, basal tooth small; subapical spur variable in length, straight or slightly curved. Protarsomeres subtriangular, gradually enlarged distally. Pro-, meso-, and metatarsal onychia each with 1 seta. Meso- and metatibiae flat, with contiguous, subequal apical spurs, slightly curved and somewhat flat, with round or acute apices; apical border of mesotibia with numerous spinules, spinules also appear on external surface forming oblique carinae; metatibia without spinules at apical border, and notched. Protarsus of male with large, unequal claws; internal claw thicker, greatly recurved, simple, not toothed, apex acute or round; outer claw long, slightly curved, with apex acute; meso- and metatarsal claws similar in size, simple, not toothed. Female claws simple.
<emphasis id="189FF917E771C36FBE85F60C0C8E6F3A" bold="true" box="[242,421,330,357]" italics="true" pageId="3" pageNumber="410">Male genitalia:</emphasis>
Aedeagus with parameres fused dorsoventrally; apices rounded and divergent externally (frontal view), curved in lateral view (
<figureCitation id="B2D03980E771C36FBC34F6290F9F6FD6" box="[579,692,367,393]" captionStart="FIGURE 1" captionStartId="3.[151,250,1687,1711]" captionTargetBox="[169,1421,429,1654]" captionTargetId="figure-123@3.[156,1431,417,1664]" captionTargetPageId="3" captionText="FIGURE 1. Homonyx cupreus Guérin-Méneville, 1839. A, Lectotype female of Homonyx cupreus; B, labels, mouthparts and ventrite of A; C, female from Minas Gerais, Brazil; D, labels of C; E, lectotype of Homonyx chalceus bahianus Ohaus, 1913 (= H. cupreus); F, labels, mouthparts and aedeagus of E; G, aedeagus, lateral and caudal views of E; H, male from Bahia, Brazil; I, labels of H; J, aedeagus, lateral and caudal views of H." figureDoi="http://doi.org/10.5281/zenodo.6524018" httpUri="https://zenodo.org/record/6524018/files/figure.png" pageId="3" pageNumber="410">Fig. 1G, J</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="7E94758DE771C36FBEE0F1D10FF76944" ID-DOI="http://doi.org/10.5281/zenodo.6524018" ID-Zenodo-Dep="6524018" httpUri="https://zenodo.org/record/6524018/files/figure.png" pageId="3" pageNumber="410" startId="3.[151,250,1687,1711]" targetBox="[169,1421,429,1654]" targetPageId="3">
<paragraph id="2A542505E771C36FBEE0F1D10FF76944" blockId="3.[151,1437,1687,1820]" pageId="3" pageNumber="410">
<emphasis id="189FF917E771C36FBEE0F1D10C3F68F0" bold="true" box="[151,276,1687,1711]" pageId="3" pageNumber="410">FIGURE 1.</emphasis>
<taxonomicName id="EDEB5E86E771C36FBF6CF1DE0FF868EF" authority="Guerin-Meneville, 1839" authorityName="Guerin-Meneville" authorityYear="1839" box="[283,723,1687,1712]" class="Insecta" family="Rutelidae" genus="Homonyx" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="410" phylum="Arthropoda" rank="species" species="cupreus">
<emphasis id="189FF917E771C36FBF6CF1DE0CF868F0" box="[283,467,1688,1711]" italics="true" pageId="3" pageNumber="410">Homonyx cupreus</emphasis>
<bibRefCitation id="4E7A58F4E771C36FBFADF1D10FF868EF" author="Guerin-Meneville, F. E." box="[474,723,1687,1712]" pageId="3" pageNumber="410" pagination="295 - 305" refId="ref3525" refString="Guerin-Meneville, F. E. (1839) Description de quelques Coleopteres des cotes du detroit de Magellan. Revue Zoologique, par la Societe Cuvierienne, 1839, 295 - 305." type="journal article" year="1839">Guérin-Méneville, 1839</bibRefCitation>
</taxonomicName>
.
<emphasis id="189FF917E771C36FBCABF1D10FC568F0" bold="true" box="[732,750,1687,1711]" pageId="3" pageNumber="410">A</emphasis>
, Lectotype female of
<taxonomicName id="EDEB5E86E771C36FBDA6F1DE09A268F0" authorityName="Guerin-Meneville" authorityYear="1839" box="[977,1161,1688,1711]" class="Insecta" family="Rutelidae" genus="Homonyx" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="410" phylum="Arthropoda" rank="species" species="cupreus">
<emphasis id="189FF917E771C36FBDA6F1DE09A268F0" box="[977,1161,1688,1711]" italics="true" pageId="3" pageNumber="410">Homonyx cupreus</emphasis>
</taxonomicName>
;
<emphasis id="189FF917E771C36FBAE0F1DE098368EF" bold="true" box="[1175,1192,1688,1712]" pageId="3" pageNumber="410">B</emphasis>
, labels, mouthparts and ventrite of A;
<emphasis id="189FF917E771C36FBF53F1FD0C1D688C" bold="true" box="[292,310,1723,1747]" pageId="3" pageNumber="410">C</emphasis>
, female from Minas Gerais, Brazil;
<emphasis id="189FF917E771C36FBCD4F1FA0F9E688B" bold="true" box="[675,693,1724,1748]" pageId="3" pageNumber="410">D</emphasis>
, labels of C;
<emphasis id="189FF917E771C36FBD4BF1FA0E66688B" bold="true" box="[828,845,1724,1748]" pageId="3" pageNumber="410">E</emphasis>
, lectotype of
<taxonomicName id="EDEB5E86E771C36FBDA0F1FA08AB688B" authority="Ohaus, 1913" authorityName="Ohaus" authorityYear="1913" box="[983,1408,1723,1748]" class="Insecta" family="Rutelidae" genus="Homonyx" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="410" phylum="Arthropoda" rank="subSpecies" species="chalceus" subSpecies="bahianus">
<emphasis id="189FF917E771C36FBDA0F1FA09D1688C" box="[983,1274,1723,1747]" italics="true" pageId="3" pageNumber="410">Homonyx chalceus bahianus</emphasis>
<bibRefCitation id="4E7A58F4E771C36FBB77F1FD08AB688B" author="Ohaus, F" box="[1280,1408,1723,1748]" pageId="3" pageNumber="410" pagination="487 - 511" refId="ref3804" refString="Ohaus, F (1913) XI. Beitrag zur Kenntnis der Ruteliden. (Col.). Deutsche Entomologische Zeitschrift, 1913, 487 - 511." type="journal article" year="1913">Ohaus, 1913</bibRefCitation>
</taxonomicName>
(=
<taxonomicName id="EDEB5E86E771C36FBEE0F1A60C2E68A8" authorityName="Guerin-Meneville" authorityYear="1839" box="[151,261,1760,1783]" class="Insecta" family="Rutelidae" genus="Homonyx" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="410" phylum="Arthropoda" rank="species" species="cupreus">
<emphasis id="189FF917E771C36FBEE0F1A60C2E68A8" box="[151,261,1760,1783]" italics="true" pageId="3" pageNumber="410">H. cupreus</emphasis>
</taxonomicName>
);
<emphasis id="189FF917E771C36FBF6BF1A60C0068A7" bold="true" box="[284,299,1760,1784]" pageId="3" pageNumber="410">F</emphasis>
, labels, mouthparts and aedeagus of E;
<emphasis id="189FF917E771C36FBCB7F1990FFF68A8" bold="true" box="[704,724,1759,1783]" pageId="3" pageNumber="410">G</emphasis>
, aedeagus, lateral and caudal views of E;
<emphasis id="189FF917E771C36FBA08F1A609B868A7" bold="true" box="[1151,1171,1760,1784]" pageId="3" pageNumber="410">H</emphasis>
, male from Bahia, Brazil;
<emphasis id="189FF917E771C36FBEE0F0420D8A6943" bold="true" box="[151,161,1796,1820]" pageId="3" pageNumber="410">I</emphasis>
, labels of H;
<emphasis id="189FF917E771C36FBF5DF0420C1C6943" bold="true" box="[298,311,1796,1820]" pageId="3" pageNumber="410">J</emphasis>
, aedeagus, lateral and caudal views of H.
</paragraph>
</caption>
<subSubSection id="62F1768EE771C368BEB0F00C0C186F3A" lastPageId="4" lastPageNumber="411" pageId="3" pageNumber="410" type="discussion">
<paragraph id="2A542505E771C36FBEB0F00C0C1D69AB" blockId="3.[151,1437,1866,2036]" pageId="3" pageNumber="410">
<emphasis id="189FF917E771C36FBEB0F00C0C10693B" bold="true" box="[199,315,1866,1892]" pageId="3" pageNumber="410">Remarks.</emphasis>
Based on external morphology and color pattern of the adults, the
<typeStatus id="F5509BA7E771C36FBA5EF00C09BB693B" box="[1065,1168,1866,1892]" pageId="3" pageNumber="410" type="lectotype">lectotype</typeStatus>
female of
<taxonomicName id="EDEB5E86E771C36FBB7EF00C08A8693C" authorityName="Guerin-Meneville" authorityYear="1839" box="[1289,1411,1866,1891]" class="Insecta" family="Rutelidae" genus="Homonyx" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="410" phylum="Arthropoda" rank="species" species="cupreus">
<emphasis id="189FF917E771C36FBB7EF00C08A8693C" box="[1289,1411,1866,1891]" italics="true" pageId="3" pageNumber="410">H. cupreus</emphasis>
</taxonomicName>
is conspecific with the females of a species of
<taxonomicName id="EDEB5E86E771C36FBCFFF0280FD869D8" authorityName="Guerin-Meneville" authorityYear="1839" box="[648,755,1902,1927]" class="Insecta" family="Rutelidae" genus="Homonyx" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="410" phylum="Arthropoda" rank="genus">
<emphasis id="189FF917E771C36FBCFFF0280FD869D8" box="[648,755,1902,1927]" italics="true" pageId="3" pageNumber="410">Homonyx</emphasis>
</taxonomicName>
that inhabits eastern
<collectingCountry id="52FC6595E771C36FBD94F028090C69D7" box="[995,1063,1902,1928]" name="Brazil" pageId="3" pageNumber="410">Brazil</collectingCountry>
(
<collectingRegion id="E82FEBE7E771C36FBA41F028095269D7" box="[1078,1145,1902,1928]" country="Brazil" name="Bahia" pageId="3" pageNumber="410">Bahia</collectingRegion>
and
<collectingRegion id="E82FEBE7E771C36FBAC7F028086D69D7" box="[1200,1350,1902,1928]" country="Brazil" name="Minas Gerais" pageId="3" pageNumber="410">Minas Gerais</collectingRegion>
states). The male genitalia of this last species is nearly identical to the male genitalia of the
<typeStatus id="F5509BA7E771C36FBA4CF0D4098969F3" box="[1083,1186,1938,1964]" pageId="3" pageNumber="410" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="EDEB5E86E771C36FBAB1F0D408B669F3" authority="Ohaus" authorityName="Ohaus" authorityYear="1913" box="[1222,1437,1938,1964]" class="Insecta" family="Rutelidae" genus="Homonyx" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="410" phylum="Arthropoda" rank="species" species="bahianus">
<emphasis id="189FF917E771C36FBAB1F0D4086569F3" box="[1222,1358,1938,1964]" italics="true" pageId="3" pageNumber="410">H. bahianus</emphasis>
Ohaus
</taxonomicName>
(=
<taxonomicName id="EDEB5E86E771C36FBECEF0F00C86698F" authorityName="Ohaus" authorityYear="1913" box="[185,429,1974,2000]" class="Insecta" family="Rutelidae" genus="Homonyx" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="410" phylum="Arthropoda" rank="subSpecies" species="chalceus" subSpecies="bahianus">
<emphasis id="189FF917E771C36FBECEF0F00C86698F" box="[185,429,1974,2000]" italics="true" pageId="3" pageNumber="410">H. chalceus bahianus</emphasis>
</taxonomicName>
). Based on this evidence, I establish that
<taxonomicName id="EDEB5E86E771C36FBDFFF0F00957698F" authorityName="Ohaus" authorityYear="1913" box="[904,1148,1974,2000]" class="Insecta" family="Rutelidae" genus="Homonyx" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="410" phylum="Arthropoda" rank="subSpecies" species="chalceus" subSpecies="bahianus">
<emphasis id="189FF917E771C36FBDFFF0F00957698F" box="[904,1148,1974,2000]" italics="true" pageId="3" pageNumber="410">H. chalceus bahianus</emphasis>
</taxonomicName>
is a new junior synonym of
<taxonomicName id="EDEB5E86E771C36FBEC1F09C0C1B69AC" authorityName="Guerin-Meneville" authorityYear="1839" box="[182,304,2010,2035]" class="Insecta" family="Rutelidae" genus="Homonyx" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="410" phylum="Arthropoda" rank="species" species="cupreus">
<emphasis id="189FF917E771C36FBEC1F09C0C1B69AC" box="[182,304,2010,2035]" italics="true" pageId="3" pageNumber="410">H. cupreus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="2A542505E776C368BEB0F7D10C186F3A" blockId="4.[151,1437,151,393]" pageId="4" pageNumber="411">
There appears to be only small differences in body color, size and density of the elytral punctation and shape of the parameres between
<taxonomicName id="EDEB5E86E776C368BFBBF7FD0F616E8B" authorityName="Guerin-Meneville" authorityYear="1839" box="[460,586,187,212]" class="Insecta" family="Rutelidae" genus="Homonyx" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="411" phylum="Arthropoda" rank="species" species="cupreus">
<emphasis id="189FF917E776C368BFBBF7FD0F616E8B" box="[460,586,187,212]" italics="true" pageId="4" pageNumber="411">H. cupreus</emphasis>
</taxonomicName>
from
<collectingCountry id="52FC6595E776C368BCE1F7FD0FF16E8A" box="[662,730,187,213]" name="Brazil" pageId="4" pageNumber="411">Brazil</collectingCountry>
and
<taxonomicName id="EDEB5E86E776C368BD6FF7FD094C6E8A" authority="Blanchard, 1851" authorityName="Blanchard" authorityYear="1851" box="[792,1127,186,213]" class="Insecta" family="Rutelidae" genus="Homonyx" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="411" phylum="Arthropoda" rank="species" species="chalceus">
<emphasis id="189FF917E776C368BD6FF7FD0E8B6E8A" box="[792,928,187,213]" italics="true" pageId="4" pageNumber="411">H. chalceus</emphasis>
Blanchard, 1851
</taxonomicName>
from northern and central
<collectingCountry id="52FC6595E776C368BEE0F7990C216EA6" box="[151,266,223,249]" name="Argentina" pageId="4" pageNumber="411">Argentina</collectingCountry>
. These characters are generally highly variable between species in other
<taxonomicName id="EDEB5E86E776C368BA4BF79909BD6EA6" authorityName="MacLeay" authorityYear="1819" box="[1084,1174,223,249]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="412" phylum="Arthropoda" rank="tribe" tribe="Rutelini">Rutelini</taxonomicName>
genera (
<emphasis id="189FF917E776C368BA8EF799080A6EA7" box="[1273,1313,223,248]" italics="true" pageId="4" pageNumber="411">e.g.</emphasis>
,
<taxonomicName id="EDEB5E86E776C368BB58F7990C6E6F42" authority="MacLeay, 1819" authorityName="MacLeay" authorityYear="1819" class="Insecta" family="Rutelidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="411" phylum="Arthropoda" rank="genus">
<emphasis id="189FF917E776C368BB58F79908B76EA6" box="[1327,1436,223,249]" italics="true" pageId="4" pageNumber="411">Pelidnota</emphasis>
MacLeay, 1819
</taxonomicName>
and
<taxonomicName id="EDEB5E86E776C368BF0AF6450FBE6F42" authority="Solier, 1851" authorityName="Solier" authorityYear="1851" box="[381,661,258,285]" class="Insecta" family="Rutelidae" genus="Catoclastus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="411" phylum="Arthropoda" rank="genus">
<emphasis id="189FF917E776C368BF0AF6450F2A6F42" box="[381,513,259,285]" italics="true" pageId="4" pageNumber="411">Catoclastus</emphasis>
<bibRefCitation id="4E7A58F4E776C368BC7FF6450FBE6F42" author="Solier, A. J. J." box="[520,661,258,285]" pageId="4" pageNumber="411" pagination="1 - 285" refId="ref3971" refString="Solier, A. J. J. (1851) Zoologia: Coleopteros. In: Gay, C. (Ed.), Historia fisica y politica de Chile segun documentos adquiridos en esta republica durante doce anos de residencia en ella Paris. Zoologia, 5, 1 - 285." type="journal article" year="1851">Solier, 1851</bibRefCitation>
</taxonomicName>
). It is likely that
<taxonomicName id="EDEB5E86E776C368BD23F6450EE56F43" authorityName="Guerin-Meneville" authorityYear="1839" box="[852,974,259,284]" class="Insecta" family="Rutelidae" genus="Homonyx" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="411" phylum="Arthropoda" rank="species" species="cupreus">
<emphasis id="189FF917E776C368BD23F6450EE56F43" box="[852,974,259,284]" italics="true" pageId="4" pageNumber="411">H. cupreus</emphasis>
</taxonomicName>
and
<taxonomicName id="EDEB5E86E776C368BA72F64509A16F42" authorityName="Blanchard" authorityYear="1851" box="[1029,1162,259,285]" class="Insecta" family="Rutelidae" genus="Homonyx" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="411" phylum="Arthropoda" rank="species" species="chalceus">
<emphasis id="189FF917E776C368BA72F64509A16F42" box="[1029,1162,259,285]" italics="true" pageId="4" pageNumber="411">H. chalceus</emphasis>
</taxonomicName>
are conspecific, but this hypothesis needs to be studied further using
<typeStatus id="F5509BA7E776C368BCF9F6610F956F1E" box="[654,702,295,321]" pageId="4" pageNumber="411">type</typeStatus>
specimens and large series of specimens before the synonymy is made official.
</paragraph>
</subSubSection>
<subSubSection id="62F1768EE776C368BEB0F6280EA66FD6" box="[199,909,366,393]" pageId="4" pageNumber="411" type="distribution">
<paragraph id="2A542505E776C368BEB0F6280EA66FD6" blockId="4.[151,1437,151,393]" box="[199,909,366,393]" pageId="4" pageNumber="411">
<emphasis id="189FF917E776C368BEB0F6280C4A6FD7" bold="true" box="[199,353,366,392]" pageId="4" pageNumber="411">Distribution.</emphasis>
<collectingCountry id="52FC6595E776C368BF10F6290C856FD6" box="[359,430,367,393]" name="Brazil" pageId="4" pageNumber="411">Brazil</collectingCountry>
, states of
<collectingRegion id="E82FEBE7E776C368BC6AF6290F756FD6" box="[541,606,367,393]" country="Brazil" name="Bahia" pageId="4" pageNumber="411">Bahia</collectingRegion>
and
<collectingRegion id="E82FEBE7E776C368BCE2F6290E006FD6" box="[661,811,367,393]" country="Brazil" name="Minas Gerais" pageId="4" pageNumber="411">Minas Gerais</collectingRegion>
(
<figureCitation id="B2D03980E776C368BD4CF6290EAB6FD6" box="[827,896,367,393]" captionStart="FIGURE 2" captionStartId="4.[151,250,1233,1257]" captionTargetBox="[158,1429,414,1207]" captionTargetId="figure-134@4.[158,1429,414,1207]" captionTargetPageId="4" captionText="FIGURE 2. Distribution records of Homonyx cupreus Guérin-Méneville, 1839 in Brazil (yellow circles)." figureDoi="http://doi.org/10.5281/zenodo.6524022" httpUri="https://zenodo.org/record/6524022/files/figure.png" pageId="4" pageNumber="411">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="7E94758DE776C368BEE0F39709E96AB6" ID-DOI="http://doi.org/10.5281/zenodo.6524022" ID-Zenodo-Dep="6524022" box="[151,1218,1233,1257]" httpUri="https://zenodo.org/record/6524022/files/figure.png" pageId="4" pageNumber="411" startId="4.[151,250,1233,1257]" targetBox="[158,1429,414,1207]" targetPageId="4">
<paragraph id="2A542505E776C368BEE0F39709E96AB6" blockId="4.[151,1218,1233,1257]" box="[151,1218,1233,1257]" pageId="4" pageNumber="411">
<emphasis id="189FF917E776C368BEE0F3970C386AB6" bold="true" box="[151,275,1233,1257]" pageId="4" pageNumber="411">FIGURE 2.</emphasis>
Distribution records of
<taxonomicName id="EDEB5E86E776C368BC71F3940E926AB6" authority="Guerin-Meneville, 1839" authorityName="Guerin-Meneville" authorityYear="1839" box="[518,953,1233,1257]" class="Insecta" family="Rutelidae" genus="Homonyx" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="411" phylum="Arthropoda" rank="species" species="cupreus">
<emphasis id="189FF917E776C368BC71F3940F966AB6" box="[518,701,1234,1257]" italics="true" pageId="4" pageNumber="411">Homonyx cupreus</emphasis>
<bibRefCitation id="4E7A58F4E776C368BCB4F3970E926AB6" author="Guerin-Meneville, F. E." box="[707,953,1233,1257]" pageId="4" pageNumber="411" pagination="295 - 305" refId="ref3525" refString="Guerin-Meneville, F. E. (1839) Description de quelques Coleopteres des cotes du detroit de Magellan. Revue Zoologique, par la Societe Cuvierienne, 1839, 295 - 305." type="journal article" year="1839">Guérin-Méneville, 1839</bibRefCitation>
</taxonomicName>
in Brazil (yellow circles).
</paragraph>
</caption>
<subSubSection id="62F1768EE776C368BEB0F25E0EFE6B9E" pageId="4" pageNumber="411" type="discussion">
<paragraph id="2A542505E776C368BEB0F25E0EFE6B9E" blockId="4.[151,1436,1303,1474]" pageId="4" pageNumber="411">
<emphasis id="189FF917E776C368BEB0F25E0C106B6D" bold="true" box="[199,315,1304,1330]" pageId="4" pageNumber="411">Remarks.</emphasis>
The erroneous record from
<collectingCountry id="52FC6595E776C368BC01F2510F986B6E" box="[630,691,1303,1329]" name="Chile" pageId="4" pageNumber="411">Chile</collectingCountry>
was treated previously by
<bibRefCitation id="4E7A58F4E776C368BDA8F25E09966B6D" author="Moore, M. R. &amp; Jameson, M. L. &amp; Garner, B. H. &amp; Audibert, C. &amp; Smith, A. B. T. &amp; Seidel, M." box="[991,1213,1303,1330]" pageId="4" pageNumber="411" pagination="1 - 349" refId="ref3719" refString="Moore, M. R., Jameson, M. L., Garner, B. H., Audibert, C., Smith, A. B. T. &amp; Seidel, M. (2017) Synopsis of the pelidnotine scarabs (Coleoptera, Scarabaeidae, Rutelinae, Rutelini) and annotated catalogue of the species and subspecies. ZooKeys, 666, 1 - 349. https: // doi. org / 10.3897 / zookeys. 666.9191" type="journal article" year="2017">
Moore
<emphasis id="189FF917E776C368BA47F25E094A6B6E" box="[1072,1121,1303,1329]" italics="true" pageId="4" pageNumber="411">et al</emphasis>
. (2017)
</bibRefCitation>
. Now, it is probable that the
<typeStatus id="F5509BA7E776C368BE87F27D0C7C6B0A" box="[240,343,1339,1365]" pageId="4" pageNumber="411" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="EDEB5E86E776C368BF0CF27A0CDE6B0A" authorityName="Guerin-Meneville" authorityYear="1839" box="[379,501,1340,1365]" class="Insecta" family="Rutelidae" genus="Homonyx" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="411" phylum="Arthropoda" rank="species" species="cupreus">
<emphasis id="189FF917E776C368BF0CF27A0CDE6B0A" box="[379,501,1340,1365]" italics="true" pageId="4" pageNumber="411">H. cupreus</emphasis>
</taxonomicName>
comes from
<collectingRegion id="E82FEBE7E776C368BCF1F27D0E0C6B0A" box="[646,807,1339,1365]" country="Brazil" name="Rio de Janeiro" pageId="4" pageNumber="411">Rio de Janeiro</collectingRegion>
(
<collectingCountry id="52FC6595E776C368BD42F27D0E546B09" box="[821,895,1339,1366]" name="Brazil" pageId="4" pageNumber="411">Brazil</collectingCountry>
), a locality that was explored in
<date id="5E5503C5E776C368BA91F27D08B76B0A" box="[1254,1436,1339,1365]" pageId="4" pageNumber="411" value="1826-09">September 1826</date>
by Alcide dOrbigny during his travels in South America. The state of
<collectingRegion id="E82FEBE7E776C368BDC2F21909726B26" box="[949,1113,1375,1401]" country="Brazil" name="Rio de Janeiro" pageId="4" pageNumber="411">Rio de Janeiro</collectingRegion>
is next to the Brazilian state of
<collectingRegion id="E82FEBE7E776C368BEC1F2C50C7A6BC2" box="[182,337,1411,1437]" country="Brazil" name="Minas Gerais" pageId="4" pageNumber="411">Minas Gerais</collectingRegion>
. It is possible that the distribution of
<taxonomicName id="EDEB5E86E776C368BC82F2C20E5B6BC2" authorityName="Guerin-Meneville" authorityYear="1839" box="[757,880,1412,1437]" class="Insecta" family="Rutelidae" genus="Homonyx" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="411" phylum="Arthropoda" rank="species" species="cupreus">
<emphasis id="189FF917E776C368BC82F2C20E5B6BC2" box="[757,880,1412,1437]" italics="true" pageId="4" pageNumber="411">H. cupreus</emphasis>
</taxonomicName>
is more extensive than currently now known and may cover territories located both in the northeast and southeast of
<collectingCountry id="52FC6595E776C368BDFEF2E10EFA6B9E" box="[905,977,1447,1473]" name="Brazil" pageId="4" pageNumber="411">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>