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<document ID-DOI="http://doi.org/10.5281/zenodo.6787471" ID-GBIF-Dataset="b35950e1-8ed6-4e16-a7d0-b1fd10add089" ID-Zenodo-Dep="6787471" ID-ZooBank="460A71F5-9390-4956-B2AE-287382C7AD7A" checkinTime="1656686604254" checkinUser="felipe" docAuthor="Löbl, Ivan" docDate="2022" docId="A22EF609FFC64006FC4CF986FAC7D2CC" docLanguage="en" docName="JpnJSystEntomol.28.1.85-88.pdf" docOrigin="Japanese Journal of Systematic Entomology 28 (1)" docTitle="Scaphisoma uenoi Löbl 2022, sp. nov." docType="treatment" docVersion="3" lastPageNumber="87" masterDocId="5E178E71FFC64004FFA9FFE7FFA4D116" masterDocTitle="On ‘ Scaphisoma tricolor Heller’ from Japan and Errata to the Descriptions of Scaphisoma konvickai Löbl and Toxidium hartmanni Löbl (Coleoptera: Staphylinidae: Scaphidiinae)" masterLastPageNumber="88" masterPageNumber="85" pageNumber="85" updateTime="1656687639904" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>On ‘ Scaphisoma tricolor Heller’ from Japan and Errata to the Descriptions of Scaphisoma konvickai Löbl and Toxidium hartmanni Löbl (Coleoptera: Staphylinidae: Scaphidiinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Löbl, Ivan</mods:namePart>
<mods:affiliation>Muséum d’histoire naturelle, Case postale 6434, CH- 1211 Genève 6, Switzerland</mods:affiliation>
<mods:nameIdentifier type="email">ivan.lobl@bluewin.ch</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Japanese Journal of Systematic Entomology</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-06-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>28</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>85</mods:start>
<mods:end>88</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.6787471</mods:identifier>
<mods:identifier type="GBIF-Dataset">b35950e1-8ed6-4e16-a7d0-b1fd10add089</mods:identifier>
<mods:identifier type="Zenodo-Dep">6787471</mods:identifier>
<mods:identifier type="ZooBank">460A71F5-9390-4956-B2AE-287382C7AD7A</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6787434" ID-Zenodo-Dep="6787434" LSID="urn:lsid:plazi:treatment:A22EF609FFC64006FC4CF986FAC7D2CC" httpUri="http://treatment.plazi.org/id/A22EF609FFC64006FC4CF986FAC7D2CC" lastPageId="2" lastPageNumber="87" pageId="0" pageNumber="85">
<subSubSection box="[997,1307,1633,1660]" pageId="0" pageNumber="85" type="nomenclature">
<paragraph blockId="0.[997,1307,1633,1688]" box="[997,1307,1633,1660]" pageId="0" pageNumber="85">
<heading box="[997,1307,1633,1660]" centered="true" fontSize="11" level="2" pageId="0" pageNumber="85" reason="1">
<taxonomicName authority="Löbl, 2022" authorityName="Löbl" authorityYear="2022" box="[997,1212,1633,1660]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="uenoi" status="sp. nov.">
<emphasis bold="true" box="[997,1212,1633,1660]" italics="true" pageId="0" pageNumber="85">Scaphisoma uenoi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1219,1307,1634,1660]" pageId="0" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[1101,1203,1666,1688]" pageId="0" pageNumber="85" type="description">
<paragraph blockId="0.[997,1307,1633,1688]" box="[1101,1203,1666,1688]" pageId="0" pageNumber="85">
<heading box="[1101,1203,1666,1688]" centered="true" fontSize="9" level="0" pageId="0" pageNumber="85" reason="4">
(
<figureCitation box="[1108,1195,1666,1688]" captionStart="Figs 1–4" captionStartId="1.[1076,1118,1342,1364]" captionTargetBox="[157,1054,197,1491]" captionTargetId="figure-265@1.[151,1064,189,1493]" captionTargetPageId="1" captionText="Figs 1–4. Aedeagus of Scaphisoma uenoi sp. nov.; 1. Median lobe, dorsal view; 2. Median lobe with internal sac, lateral view; 3. Parameres, ventral view; 4. Internal sac, dorsal view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6787481" httpUri="https://zenodo.org/record/6787481/files/figure.png" pageId="0" pageNumber="85">Figs 1–4</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection lastPageId="1" lastPageNumber="86" pageId="0" pageNumber="85" type="materials_examined">
<paragraph blockId="0.[845,1460,1730,2041]" lastBlockId="1.[151,768,1536,2041]" lastPageId="1" lastPageNumber="86" pageId="0" pageNumber="85">
<materialsCitation collectingDate="1991-07-16" collectionCode="EUMJ" collectorName="T. Ueno" country="Japan" location="Is. Hahajima Mt. Kuwanoki" municipality="Ogasawara" pageId="0" pageNumber="85" specimenCount="1" typeStatus="holotype">
<emphasis box="[883,1025,1730,1752]" italics="true" pageId="0" pageNumber="85">Type material</emphasis>
.
<typeStatus box="[1040,1134,1730,1752]" pageId="0" pageNumber="85">Holotype</typeStatus>
[
<collectingMunicipality box="[1149,1267,1730,1752]" pageId="0" pageNumber="85">Ogasawara</collectingMunicipality>
]
<location LSID="urn:lsid:plazi:treatment:A22EF609FFC64006FC4CF986FAC7D2CC:2F5811C4FFC64004FAABF924FC12D7EE" country="Japan" municipality="Ogasawara" name="Is. Hahajima Mt. Kuwanoki" pageId="0" pageNumber="85">Is. Hahajima Mt. Kuwanoki</location>
<date box="[959,1076,1762,1784]" pageId="0" pageNumber="85" value="1991-07-16">
<collectingDate box="[959,1076,1762,1784]" pageId="0" pageNumber="85" value="1991-07-16">16.vii.1991</collectingDate>
</date>
<collectorName box="[1084,1166,1763,1784]" pageId="0" pageNumber="85">T. Ueno</collectorName>
leg. (
<collectionCode box="[1229,1301,1763,1785]" collectionName="Japan, Matsuyama, Ehime University" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="0" pageNumber="85">EUMJ</collectionCode>
)
</materialsCitation>
.
<materialsCitation collectingDate="1991-07-16" collectionCode="EUMJ" collectorName="T. Ueno" country="Japan" location="Is. Hahajima Mt. Kuwanoki" municipality="Ogasawara" pageId="0" pageNumber="85" specimenCount="16" specimenCount-female="4" specimenCount-male="12" typeStatus="paratype">
<typeStatus box="[1320,1422,1763,1784]" pageId="0" pageNumber="85">Paratypes</typeStatus>
.
<specimenCount pageId="0" pageNumber="85" type="male">12 males</specimenCount>
,
<specimenCount box="[916,1014,1795,1817]" pageId="0" pageNumber="85" type="female">4 females</specimenCount>
, with the same data as the holotype
</materialsCitation>
;
<materialsCitation collectingDate="1969-07-15" collectorName="T. Nakane" country="Japan" location="Hahajima Bonin Is." pageId="0" pageNumber="85" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount box="[1385,1455,1795,1817]" pageId="0" pageNumber="85" type="male">1 male</specimenCount>
,
<location LSID="urn:lsid:plazi:treatment:A22EF609FFC64006FC4CF986FAC7D2CC:2F5811C4FFC64004FCE7F8C4FBB5D62E" box="[846,1041,1827,1849]" country="Italy" name="Hahajima Bonin Is." pageId="0" pageNumber="85">Hahajima Bonin Is.</location>
<date box="[1048,1168,1827,1849]" pageId="0" pageNumber="85" value="1969-07-15">
<collectingDate box="[1048,1168,1827,1849]" pageId="0" pageNumber="85" value="1969-07-15">15.VII.1969</collectingDate>
</date>
<collectorName box="[1175,1280,1827,1849]" pageId="0" pageNumber="85">T. Nakane</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="1976-06-02" collectorName="T. Nakane" country="Japan" location="Bonin" municipality="Okimura Hahajima" pageId="0" pageNumber="85" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount box="[1292,1361,1827,1849]" pageId="0" pageNumber="85" type="male">1 male</specimenCount>
,
<collectingMunicipality pageId="0" pageNumber="85">Okimura Hahajima</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:A22EF609FFC64006FC4CF986FAC7D2CC:2F5811C4FFC64004FC6BF8A4FBA4D64F" box="[962,1024,1859,1881]" country="Italy" municipality="Okimura Hahajima" name="Bonin" pageId="0" pageNumber="85">Bonin</location>
<date box="[1034,1132,1859,1881]" pageId="0" pageNumber="85" value="1976-06-02">
<collectingDate box="[1034,1132,1859,1881]" pageId="0" pageNumber="85" value="1976-06-02">2.vi.1976</collectingDate>
</date>
<collectorName box="[1140,1252,1859,1881]" pageId="0" pageNumber="85">T. Nakane</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="1986-07-01" collectingDateMax="1986-07-08" collectingDateMin="1986-07-01" collectorName="H. Makihara" country="Japan" location="Ogasawara" pageId="0" pageNumber="85" specimenCount="1" specimenCount-male="1" stateProvince="Haha-jima" typeStatus="paratype">
<specimenCount box="[1265,1338,1859,1881]" pageId="0" pageNumber="85" type="male">1 male</specimenCount>
,
<collectingRegion box="[1352,1460,1859,1881]" pageId="0" pageNumber="85">Haha-jima</collectingRegion>
<location LSID="urn:lsid:plazi:treatment:A22EF609FFC64006FC4CF986FAC7D2CC:2F5811C4FFC64004FCE4F884FC18D66E" box="[845,956,1891,1912]" country="Italy" name="Ogasawara" pageId="0" pageNumber="85" stateProvince="Haha-jima">Ogasawara</location>
<date box="[965,1100,1891,1913]" pageId="0" pageNumber="85" value="1986-07-01" valueMax="1986-07-08" valueMin="1986-07-01">
<collectingDate box="[965,1100,1891,1913]" pageId="0" pageNumber="85" value="1986-07-01" valueMax="1986-07-08" valueMin="1986-07-01">1–8.VII.1986</collectingDate>
</date>
<collectorName box="[1109,1240,1891,1913]" pageId="0" pageNumber="85">H. Makihara</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="1968-08-20" collectorName="T. Nakane" country="Japan" location="Bonin" municipality="Chichijima" pageId="0" pageNumber="85" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<specimenCount box="[1253,1335,1891,1913]" pageId="0" pageNumber="85" type="male">2 males</specimenCount>
,
<collectingMunicipality box="[1347,1458,1891,1913]" pageId="0" pageNumber="85">Chichijima</collectingMunicipality>
<location LSID="urn:lsid:plazi:treatment:A22EF609FFC64006FC4CF986FAC7D2CC:2F5811C4FFC64004FCE4F864FC28D68F" box="[845,908,1923,1945]" country="Italy" municipality="Chichijima" name="Bonin" pageId="0" pageNumber="85">Bonin</location>
Is.
<date box="[941,1081,1923,1945]" pageId="0" pageNumber="85" value="1968-08-20">
<collectingDate box="[941,1081,1923,1945]" pageId="0" pageNumber="85" value="1968-08-20">20.VIII.1968</collectingDate>
</date>
<collectorName box="[1091,1204,1923,1945]" pageId="0" pageNumber="85">T. Nakane</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="1974-05-03" collectorName="H. Yamada" country="Japan" location="Ogasawara Islands" municipality="Is. Chichijima" pageId="0" pageNumber="85" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount box="[1218,1293,1923,1945]" pageId="0" pageNumber="85" type="male">1 male</specimenCount>
,
<collectingMunicipality box="[1307,1458,1923,1945]" pageId="0" pageNumber="85">Is. Chichijima</collectingMunicipality>
<location LSID="urn:lsid:plazi:treatment:A22EF609FFC64006FC4CF986FAC7D2CC:2F5811C4FFC64004FCE4F844FBABD6AF" box="[845,1039,1955,1977]" country="Italy" municipality="Is. Chichijima" name="Ogasawara Islands" pageId="0" pageNumber="85">Ogasawara Islands</location>
<date box="[1048,1176,1955,1977]" pageId="0" pageNumber="85" value="1974-05-03">
<collectingDate box="[1048,1176,1955,1977]" pageId="0" pageNumber="85" value="1974-05-03">3 May 1974</collectingDate>
</date>
<collectorName box="[1184,1299,1955,1977]" pageId="0" pageNumber="85">H. Yamada</collectorName>
leg.
</materialsCitation>
;
<materialsCitation collectingDate="1969-07-15" collectorName="T. Nakane" country="Japan" location="Bonin Is." municipality="Chichijima" pageId="0" pageNumber="85" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount box="[1362,1455,1955,1977]" pageId="0" pageNumber="85" type="female">1 female</specimenCount>
,
<collectingMunicipality box="[845,954,1987,2009]" pageId="0" pageNumber="85">Chichijima</collectingMunicipality>
<location LSID="urn:lsid:plazi:treatment:A22EF609FFC64006FC4CF986FAC7D2CC:2F5811C4FFC64004FC69F824FBBFD6CF" box="[960,1051,1987,2009]" country="Italy" municipality="Chichijima" name="Bonin Is." pageId="0" pageNumber="85">Bonin Is.</location>
<date box="[1056,1176,1987,2009]" pageId="0" pageNumber="85" value="1969-07-15">
<collectingDate box="[1056,1176,1987,2009]" pageId="0" pageNumber="85" value="1969-07-15">15.VII.1969</collectingDate>
</date>
<collectorName box="[1182,1286,1987,2009]" pageId="0" pageNumber="85">T. Nakane</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="1974-05-09" collectingDateMax="1974-05-10" collectingDateMin="1974-05-09" collectorName="Y. Hori" country="Japan" location="Ogasawara" pageId="0" pageNumber="85" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Chichi Jima" typeStatus="paratype">
<specimenCount box="[1296,1368,1987,2009]" pageId="0" pageNumber="85" type="male">1 male</specimenCount>
,
<specimenCount box="[1372,1455,1987,2009]" pageId="0" pageNumber="85" type="female">1 female</specimenCount>
,
<collectingRegion box="[845,975,2019,2041]" pageId="0" pageNumber="85">Chichi Jima</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:A22EF609FFC64006FC4CF986FAC7D2CC:2F5811C4FFC64004FC75F803FBEBD6EF" box="[988,1103,2020,2041]" country="Italy" name="Ogasawara" pageId="0" pageNumber="85" stateProvince="Chichi Jima">Ogasawara</location>
<date box="[1113,1294,2019,2041]" pageId="0" pageNumber="85" value="1974-05-09" valueMax="1974-05-10" valueMin="1974-05-09">
<collectingDate box="[1113,1294,2019,2041]" pageId="0" pageNumber="85" value="1974-05-09" valueMax="1974-05-10" valueMin="1974-05-09">9. and 10.V.1974</collectingDate>
</date>
<collectorName box="[1303,1381,2019,2041]" pageId="0" pageNumber="85">Y. Hori</collectorName>
leg.
</materialsCitation>
;
<materialsCitation collectingDate="1987-10-22" collectorName="M. Sakai" country="Japan" lastPageId="1" lastPageNumber="86" location="Hiji" municipality="Ryukyu" pageId="0" pageNumber="85" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Okinawa" typeStatus="paratype">
<specimenCount lastPageId="1" lastPageNumber="86" pageId="0" pageNumber="85" type="male">1 male</specimenCount>
,
<specimenCount box="[212,300,1537,1559]" pageId="1" pageNumber="86" type="female">1 female</specimenCount>
, [
<collectingRegion box="[316,411,1537,1559]" pageId="1" pageNumber="86">Okinawa</collectingRegion>
:
<collectingMunicipality box="[421,501,1536,1558]" pageId="1" pageNumber="86">Ryukyu</collectingMunicipality>
]
<location LSID="urn:lsid:plazi:treatment:A22EF609FFC64006FC4CF986FAC7D2CC:2F5811C4FFC74005FDA8F9E6FD83D701" box="[513,551,1537,1559]" country="Italy" municipality="Ryukyu" name="Hiji" pageId="1" pageNumber="86" stateProvince="Okinawa">Hiji</location>
–
<location LSID="urn:lsid:plazi:treatment:A22EF609FFC64006FC4CF986FAC7D2CC:2F5811C4FFC74005FDE8F9E6FD59D701" box="[577,765,1537,1559]" country="Italy" municipality="Ryukyu" name="Hiji Fall Kunigami" pageId="1" pageNumber="86" stateProvince="Okinawa">Hiji Fall Kunigami</location>
<date box="[151,254,1569,1590]" pageId="1" pageNumber="86" value="1987-10-22">
<collectingDate box="[151,254,1569,1590]" pageId="1" pageNumber="86" value="1987-10-22">22.X.1987</collectingDate>
</date>
<collectorName box="[261,349,1569,1591]" pageId="1" pageNumber="86">M. Sakai</collectorName>
leg.
</materialsCitation>
;
<materialsCitation collectingDate="1987-11-01" collectingDateMax="1987-11-03" collectingDateMin="1987-11-01" collectorName="T. Ueno" country="Japan" location="Is. Tokashiki" municipality="Ryukyus" pageId="1" pageNumber="86" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Okinawa" typeStatus="paratype">
<specimenCount box="[404,473,1569,1591]" pageId="1" pageNumber="86" type="male">1 male</specimenCount>
,
<specimenCount box="[483,570,1569,1591]" pageId="1" pageNumber="86" type="female">1 female</specimenCount>
,
<collectingCountry box="[580,640,1569,1590]" name="Japan" pageId="1" pageNumber="86">Japan</collectingCountry>
:
<collectingMunicipality box="[649,735,1569,1591]" pageId="1" pageNumber="86">Ryukyus</collectingMunicipality>
<location LSID="urn:lsid:plazi:treatment:A22EF609FFC64006FC4CF986FAC7D2CC:2F5811C4FFC74005FD4CF9C6FF5CD741" country="Japan" municipality="Ryukyus" name="Is. Tokashiki" pageId="1" pageNumber="86" stateProvince="Okinawa">Is. Tokashiki</location>
<collectingRegion box="[256,345,1601,1623]" country="Japan" name="Okinawa" pageId="1" pageNumber="86">Okinawa</collectingRegion>
Pref.
<date box="[408,525,1601,1623]" pageId="1" pageNumber="86" value="1987-11-01" valueMax="1987-11-03" valueMin="1987-11-01">
<collectingDate box="[408,525,1601,1623]" pageId="1" pageNumber="86" value="1987-11-01" valueMax="1987-11-03" valueMin="1987-11-01">1–3.xi.1987</collectingDate>
</date>
<collectorName box="[532,613,1601,1622]" pageId="1" pageNumber="86">T. Ueno</collectorName>
leg.
</materialsCitation>
;
<materialsCitation collectingDate="1987-09-27" collectionCode="EUMJ, MHNG" collectorName="T. Ueno" country="Japan" location="Nago Is." municipality="Ryukyus" pageId="1" pageNumber="86" specimenCount="1" specimenCount-female="1" stateProvince="Okinawa" typeStatus="paratype">
<specimenCount box="[672,761,1601,1623]" pageId="1" pageNumber="86" type="female">1 female</specimenCount>
,
<collectingCountry box="[153,215,1634,1655]" name="Japan" pageId="1" pageNumber="86">Japan</collectingCountry>
:
<collectingMunicipality box="[227,316,1633,1655]" pageId="1" pageNumber="86">Ryukyus</collectingMunicipality>
<location LSID="urn:lsid:plazi:treatment:A22EF609FFC64006FC4CF986FAC7D2CC:2F5811C4FFC74005FEEDF985FE39D761" box="[324,413,1634,1655]" country="Japan" municipality="Ryukyus" name="Nago Is." pageId="1" pageNumber="86" stateProvince="Okinawa">Nago Is.</location>
<collectingRegion box="[421,512,1633,1655]" country="Japan" name="Okinawa" pageId="1" pageNumber="86">Okinawa</collectingRegion>
<date box="[521,630,1633,1655]" pageId="1" pageNumber="86" value="1987-09-27">
<collectingDate box="[521,630,1633,1655]" pageId="1" pageNumber="86" value="1987-09-27">27.ix.1987</collectingDate>
</date>
<collectorName box="[638,721,1634,1655]" pageId="1" pageNumber="86">T. Ueno</collectorName>
leg. (
<collectionCode box="[158,226,1665,1686]" collectionName="Japan, Matsuyama, Ehime University" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="1" pageNumber="86">EUMJ</collectionCode>
,
<collectionCode box="[236,316,1665,1687]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="1" pageNumber="86" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.6787481" ID-Zenodo-Dep="6787481" httpUri="https://zenodo.org/record/6787481/files/figure.png" pageId="1" pageNumber="86" startId="1.[1076,1118,1342,1364]" targetBox="[157,1054,197,1491]" targetPageId="1">
<paragraph blockId="1.[1076,1435,1342,1497]" pageId="1" pageNumber="86">
Figs 1–4. Aedeagus of
<taxonomicName authority="Löbl, 2022" authorityName="Löbl" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="86" phylum="Arthropoda" rank="species" species="uenoi" status="sp. nov.">
<emphasis italics="true" pageId="1" pageNumber="86">Scaphisoma uenoi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1135,1211,1369,1390]" pageId="1" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
; 1. Median lobe, dorsal view; 2. Median lobe with internal sac, lateral view; 3. Parameres, ventral view; 4. Internal sac, dorsal view. Scale bars = 0.2 mm.
</paragraph>
</caption>
<subSubSection lastPageId="2" lastPageNumber="87" pageId="1" pageNumber="86" type="description">
<paragraph blockId="1.[151,768,1536,2041]" lastBlockId="1.[818,1433,1537,2041]" pageId="1" pageNumber="86">
<emphasis box="[189,304,1698,1719]" italics="true" pageId="1" pageNumber="86">Description</emphasis>
. Male. Length
<quantity box="[453,597,1697,1719]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.1" metricValueMin="1.9" pageId="1" pageNumber="86" unit="mm" value="2.0" valueMax="2.1" valueMin="1.9">1.90–2.10 mm</quantity>
, width
<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.4649999999999999" metricValueMax="1.55" metricValueMin="1.38" pageId="1" pageNumber="86" unit="mm" value="1.4649999999999999" valueMax="1.55" valueMin="1.38">1.38–1.55 mm</quantity>
. Head, pronotum and hypomere light, reddish-brown. Pronotum along basal margin narrow darkened, sometimes slightly darkened on median or basomedian area. Antennae and mouth-parts yellowish. Exposed scutellum dark brown to blackish. Elytra brown to blackish on anterior two thirds, yellowish or ochraceous on apical fourth to third. Mesanepisterna, metaventrite, metanepisterna, metepimera and ventrites I to III brown, following ventrites lighter, usually yellowish, similar to exposed tergites and apices of elytra, or somewhat darker. Coxae, femora and tibiae about as light as abdominal apex, tarsi still slightly lighter. Antennae long, antennomere
<collectionCode box="[961,978,1569,1590]" pageId="1" pageNumber="86">II</collectionCode>
about 3 times as long as antennomere III. Antennomeres IV to
<collectionCode box="[1025,1051,1601,1622]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="1" pageNumber="86">VI</collectionCode>
narrow, similar in width, antennomere V longer than antennomeres IV or
<collectionCode box="[1167,1196,1634,1655]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="1" pageNumber="86">VI</collectionCode>
. Antennomeres VII and IX–XI about twice as wide as antennomere
<collectionCode box="[1290,1319,1666,1687]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="1" pageNumber="86">VI</collectionCode>
, similar in length.
</paragraph>
<paragraph blockId="1.[818,1433,1537,2041]" pageId="1" pageNumber="86">Pronotum lacking microsculpture, with evenly arcuate lateral margins. Lateral margin carinae exposed in dorsal view. Lateral margin striae impunctate. Discal punctation irregular, very fine, with puncture intervals usually 2 to 5 times as large as puncture diameters. Exposed apex of scutellum triangular, with acute tip.</paragraph>
<paragraph blockId="1.[818,1433,1537,2041]" lastBlockId="2.[178,795,193,2046]" lastPageId="2" lastPageNumber="87" pageId="1" pageNumber="86">Elytra lacking microsculpture, weakly narrowed apically, with lateral margins nearly evenly arcuate. Lateral margin carinae throughout exposed. Lateral margin striae punctate except near apices, sutural margin not raised. Sutural striae shallow, starting well posterior of level of scutellar tip, weakly converging apically, adsutural areas flat, with each with single puncture row. Apical margins truncate, inner apical angle slightly posterior level of outer apical angles, lacking obvious crenulation or denticles. Prevailing punctation on darkly coloured surface coarse and rather dense, well delimited, with puncture intervals mostly about as large to three times as large as puncture diameters; punctation along bases and lateral margins finer yet well delimited, than on light apical areas denser, much shallower und poorly delimited.</paragraph>
<paragraph blockId="2.[178,795,193,2046]" pageId="2" pageNumber="87">
Hypomera and mesanepisterna extremely finely punctate, lacking microsculpture, with distinct pubescence. Mesepimera about 4 times as long as wide and as long as intervals to mesocoxae. Metaventrite bearing strigulate microsculpture on most of mesal areas extended onto part of lateral areas, most of lateral areas without microsculpture; middle of metaventrite convex, with narrow area anterior metacoxal process flattened or slightly impressed; prevailing punctuation on metaventrite very fine and sparse; two small, dense patches of rather coarse punctures present on apicomedian area; with fine punctures forming two admesal rows converging anteriorly, forming inverted V-shaped pattern and delimiting narrow area bearing oblique to longitudinal microstriae; dense antecoxal rows of coarse punctures present. Submesocoxal areas narrow, about
<quantity box="[178,322,964,986]" metricMagnitude="-5" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="2" pageNumber="87" unit="mm" value="0.035" valueMax="0.04" valueMin="0.03">0.03–0.04 mm</quantity>
long, about as long as sixth to fifth of shortest intervals to metacoxae; submesocoxal lines parallel to the mesocoxal margins, coarsely punctured. Metanepisterna slightly convex, impressed below margin of metaventrite, narrowed anteriad, lacking microsculpture, with inner margin nearly straight; punctation very fine and sparse, as that on lateral parts of metaventrite. Metepimera bearing striate microsculpture.
</paragraph>
<paragraph blockId="2.[178,795,193,2046]" pageId="2" pageNumber="87">
Legs with protibae and metatibiae straight and narrow. Mesotibiae thickened and bent. Tarsomeres I of prolegs and mesolegs strongly widened, wider than apices of respective tibiae. Tarsomeres
<collectionCode box="[398,416,1318,1339]" pageId="2" pageNumber="87">II</collectionCode>
of prolegs moderately widened. Tarsomeres
<collectionCode box="[297,313,1350,1371]" pageId="2" pageNumber="87">II</collectionCode>
of mesolegs nearly as wide as tarsomeres I.
</paragraph>
<paragraph blockId="2.[178,795,193,2046]" pageId="2" pageNumber="87">
Abdomen with striate microsculpture. Ventrite I with narrow basomedian patch of coarse punctures, prevailing punctures very fine and sparse, as on rest of exposed abdomen. Submetacoxal areas about
<quantity box="[544,631,1478,1500]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="87" unit="mm" value="0.05">0.05 mm</quantity>
long, as long as fourth of interval between them and apical margin of ventrite I; submetacoxal lines convex, punctate. Apex of ventrite
<collectionCode box="[767,793,1543,1564]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="2" pageNumber="87">VI</collectionCode>
expanded to form large mesal lobe about
<quantity box="[594,683,1574,1596]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="87" unit="mm" value="0.25">0.25 mm</quantity>
long, with rounded margin. Aedeagus as
<figureCitation box="[475,616,1607,1629]" captionStart="Figs 1–4" captionStartId="1.[1076,1118,1342,1364]" captionTargetBox="[157,1054,197,1491]" captionTargetId="figure-265@1.[151,1064,189,1493]" captionTargetPageId="1" captionText="Figs 1–4. Aedeagus of Scaphisoma uenoi sp. nov.; 1. Median lobe, dorsal view; 2. Median lobe with internal sac, lateral view; 3. Parameres, ventral view; 4. Internal sac, dorsal view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6787481" httpUri="https://zenodo.org/record/6787481/files/figure.png" pageId="2" pageNumber="87">
Figs 1–4,
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long.
</paragraph>
<paragraph blockId="2.[178,795,193,2046]" pageId="2" pageNumber="87">
Female. Head and pronotum brown to blackish, not or only somewhat lighter than elytra. Elytra with apical margins rounded, finely crenulate near inner angles, inner apical angles situated distinctly posterior of level of outer apical angles. Mesotibiae straight, not thickened. Tarsomeres I and
<collectionCode box="[178,194,1800,1821]" pageId="2" pageNumber="87">II</collectionCode>
not widened. Ventrite
<collectionCode box="[418,444,1800,1821]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="2" pageNumber="87">VI</collectionCode>
not expanded apically. Gonocoxite cylindric, evenly narrow posterior basal section and bent outward, with distinct subapical seta. Gonostyle with one short and one very long apical seta.
</paragraph>
</subSubSection>
<subSubSection box="[215,744,1928,1950]" pageId="2" pageNumber="87" type="distribution">
<paragraph blockId="2.[178,795,193,2046]" box="[215,744,1928,1950]" pageId="2" pageNumber="87">
Distribution.
<collectingCountry box="[346,406,1928,1949]" name="Japan" pageId="2" pageNumber="87">Japan</collectingCountry>
: Ogasawara and
<collectingRegion box="[572,661,1928,1950]" country="Japan" name="Okinawa" pageId="2" pageNumber="87">Okinawa</collectingRegion>
Islands.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="87" type="etymology">
<paragraph blockId="2.[178,795,193,2046]" pageId="2" pageNumber="87">
<emphasis box="[215,322,1960,1981]" italics="true" pageId="2" pageNumber="87">Etymology</emphasis>
. The species is named after the Mr. Teruhisa Ueno (Kyushu University), who collected the bulk of the studied specimens.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="87" type="discussion">
<paragraph blockId="2.[845,1461,193,986]" pageId="2" pageNumber="87">
<emphasis box="[883,969,193,214]" italics="true" pageId="2" pageNumber="87">Remarks</emphasis>
. The
<taxonomicName authorityName=", Therefore, Lobl" authorityYear="2018" box="[1025,1125,193,215]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="87" phylum="Arthropoda" rank="species" species="tricolor">
<emphasis box="[1025,1037,193,214]" italics="true" pageId="2" pageNumber="87">S</emphasis>
.
<emphasis box="[1050,1125,193,214]" italics="true" pageId="2" pageNumber="87">tricolor</emphasis>
</taxonomicName>
group, defined by
<bibRefCitation author="Lobl, I. &amp; R. Ogawa" journalOrPublisher="Linzer biologische Beitrage" pageId="2" pageNumber="87" pagination="1339 - 1492" part="48" refId="ref2973" refString="Lobl, I. &amp; R. Ogawa, 2016. On the Scaphisomatini (Coleoptera, Staphylinidae, Scaphidiinae) of the Philippines, IV: the genera Sapitia Achard and Scaphisoma Leach. Linzer biologische Beitrage, 48 (2): 1339 - 1492." title="On the Scaphisomatini (Coleoptera, Staphylinidae, Scaphidiinae) of the Philippines, IV: the genera Sapitia Achard and Scaphisoma Leach" type="journal article" year="2016">Löbl &amp; Ogawa (2016)</bibRefCitation>
, comprises at present 24 species. The characters of three additional species known in females only suggest also their possible close relationship to
<taxonomicName authorityName=", Therefore, Lobl" authorityYear="2018" box="[1263,1369,289,311]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="87" phylum="Arthropoda" rank="species" species="tricolor">
<emphasis box="[1263,1275,290,311]" italics="true" pageId="2" pageNumber="87">S</emphasis>
.
<emphasis box="[1291,1369,289,310]" italics="true" pageId="2" pageNumber="87">tricolor</emphasis>
</taxonomicName>
. Among these species, the males of the following six exhibit similar, conspicuous colour pattern, with the pronotum different sexually (see Löbl &amp; Ogawa, 20216):
<taxonomicName authority="Heller" authorityName="Heller" authorityYear="1917" box="[1247,1427,386,408]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="87" phylum="Arthropoda" rank="species" species="tricolor">
<emphasis box="[1247,1259,386,407]" italics="true" pageId="2" pageNumber="87">S</emphasis>
.
<emphasis box="[1275,1353,386,407]" italics="true" pageId="2" pageNumber="87">tricolor</emphasis>
Heller
</taxonomicName>
,
<taxonomicName authority="Lobl" authorityName="Lobl" authorityYear="1982" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="87" phylum="Arthropoda" rank="species" species="chujoi">
<emphasis box="[1440,1452,386,407]" italics="true" pageId="2" pageNumber="87">S</emphasis>
.
<emphasis box="[845,907,418,439]" italics="true" pageId="2" pageNumber="87">chujoi</emphasis>
Löbl
</taxonomicName>
,
<taxonomicName authority="Lobl" authorityName="Lobl" authorityYear="2015" box="[976,1129,418,440]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="87" phylum="Arthropoda" rank="species" species="tarsale">
<emphasis box="[976,988,418,439]" italics="true" pageId="2" pageNumber="87">S</emphasis>
.
<emphasis box="[1003,1072,418,439]" italics="true" pageId="2" pageNumber="87">tarsale</emphasis>
Löbl
</taxonomicName>
,
<taxonomicName authority="Lobl &amp; Ogawa" authorityName="Lobl &amp; Ogawa, 2016:" authorityYear="1435" box="[1141,1455,418,440]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="87" phylum="Arthropoda" rank="species" species="tricoloratum">
<emphasis box="[1141,1153,418,439]" italics="true" pageId="2" pageNumber="87">S</emphasis>
.
<emphasis box="[1167,1292,418,439]" italics="true" pageId="2" pageNumber="87">tricoloratum</emphasis>
Löbl &amp; Ogawa
</taxonomicName>
,
<taxonomicName authority="Lobl &amp; Ogawa" authorityName="Lobl &amp; Ogawa, 2016:" authorityYear="1436" box="[845,1178,450,472]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="87" phylum="Arthropoda" rank="species" species="tricolorinotum">
<emphasis box="[845,857,450,471]" italics="true" pageId="2" pageNumber="87">S</emphasis>
.
<emphasis box="[872,1017,450,471]" italics="true" pageId="2" pageNumber="87">tricolorinotum</emphasis>
Löbl &amp; Ogawa
</taxonomicName>
and
<taxonomicName authority="Löbl, 2022" authorityName="Löbl" authorityYear="2022" box="[1231,1313,450,472]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="87" phylum="Arthropoda" rank="species" species="uenoi" status="sp. nov.">
<emphasis box="[1231,1243,450,471]" italics="true" pageId="2" pageNumber="87">S</emphasis>
.
<emphasis box="[1258,1313,451,472]" italics="true" pageId="2" pageNumber="87">uenoi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1321,1401,451,472]" pageId="2" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
They share also most of the external characters, while their aedeagi provide reliable diagnostic features. The new species may be distinguished from
<taxonomicName authorityName="Heller" authorityYear="1917" box="[1073,1173,546,568]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="87" phylum="Arthropoda" rank="species" species="tricolor">
<emphasis box="[1073,1083,547,568]" italics="true" pageId="2" pageNumber="87">S</emphasis>
.
<emphasis box="[1098,1173,546,567]" italics="true" pageId="2" pageNumber="87">tricolor</emphasis>
</taxonomicName>
by the apical process of the median lobe with sinuate dorsal and ventral branches and the internal sac lacking tuberculate medial piece.
<taxonomicName authorityName="Löbl" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="87" phylum="Arthropoda" rank="species" species="uenoi">
<emphasis italics="true" pageId="2" pageNumber="87">Scaphisoma uenoi</emphasis>
</taxonomicName>
is distinguished from the Philippine
<taxonomicName authorityName="Lobl &amp; Ogawa, 2016:" authorityYear="1435" box="[1300,1459,643,664]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="87" phylum="Arthropoda" rank="species" species="tricoloratum">
<emphasis box="[1300,1312,643,664]" italics="true" pageId="2" pageNumber="87">S</emphasis>
.
<emphasis box="[1328,1459,643,664]" italics="true" pageId="2" pageNumber="87">tricoloratum</emphasis>
</taxonomicName>
by the strongly expanded parameral laminae and from
<taxonomicName authorityName="Lobl &amp; Ogawa, 2016:" authorityYear="1436" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="87" phylum="Arthropoda" rank="species" species="tricolorinotum">
<emphasis box="[1440,1452,675,696]" italics="true" pageId="2" pageNumber="87">S</emphasis>
.
<emphasis box="[845,996,707,728]" italics="true" pageId="2" pageNumber="87">tricolorinotum</emphasis>
</taxonomicName>
by the ventral branch of the apical process not gradually narrowed and not curved in dorsal view.
<taxonomicName authorityName="Lobl" authorityYear="1982" box="[845,1044,771,792]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="87" phylum="Arthropoda" rank="species" species="chujoi">
<emphasis box="[845,1044,771,792]" italics="true" pageId="2" pageNumber="87">Scaphisoma chujoi</emphasis>
</taxonomicName>
differs strongly from all these species by the weakly inflexed ventral branch of the apical process, and from
<taxonomicName authorityName="Löbl" authorityYear="2022" box="[945,1027,836,857]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="87" phylum="Arthropoda" rank="species" species="uenoi">
<emphasis box="[945,957,836,857]" italics="true" pageId="2" pageNumber="87">S</emphasis>
.
<emphasis box="[972,1027,836,857]" italics="true" pageId="2" pageNumber="87">uenoi</emphasis>
</taxonomicName>
also by the parameres less expanded. The median lobe and parameres of
<taxonomicName authorityName="Lobl" authorityYear="2015" box="[1175,1276,868,889]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="87" phylum="Arthropoda" rank="species" species="tarsale">
<emphasis box="[1175,1187,868,889]" italics="true" pageId="2" pageNumber="87">S</emphasis>
.
<emphasis box="[1204,1276,868,889]" italics="true" pageId="2" pageNumber="87">tarsale</emphasis>
</taxonomicName>
are quite similar to those of
<taxonomicName authorityName="Löbl" authorityYear="2022" box="[968,1054,900,921]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="87" phylum="Arthropoda" rank="species" species="uenoi">
<emphasis box="[968,980,900,921]" italics="true" pageId="2" pageNumber="87">S</emphasis>
.
<emphasis box="[997,1054,900,921]" italics="true" pageId="2" pageNumber="87">uenoi</emphasis>
</taxonomicName>
, however the structure of the internal sac differs significantly, notably by the transverse proximal sclerite and the absence of mesal sclerotized structure.
</paragraph>
</subSubSection>
</treatment>
</document>
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