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120 lines
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<document ID-DOI="http://dx.doi.org/10.3897/ijm.5.1899" ID-GBIF-Dataset="91d53148-40a1-4b34-a990-f5a6d3ca50c9" ID-GBIF-Taxon="154473563" ID-Pensoft-Pub="1875-2543--27" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1875-2543--27" ModsDocOrigin="International Journal of Myriapodology " ModsDocTitle="The curious genitalic structures of Tancitares michoacanus (Polydesmida, Rhachodesmidae)" checkinTime="1553125527789" checkinUser="pensoft" docAuthor="Hoffman, Richard L." docDate="2011" docId="9885BCD52CD609CC2C3A2462E0B921F4" docLanguage="en" docName="InternJourMyriapod 5: 27-33" docOrigin="International Journal of Myriapodology 5" docSource="http://dx.doi.org/10.3897/ijm.5.1899" docTitle="Tancitares Chamberlin" docType="treatment" docVersion="3" lastPageId="2" lastPageNumber="29" masterDocId="4168FFC57F7AFFB5FF89AD2FFFE9FFA2" masterDocTitle="The curious genitalic structures of Tancitares michoacanus (Polydesmida, Rhachodesmidae)" masterLastPageNumber="33" masterPageNumber="27" pageId="1" pageNumber="28" updateTime="1643555422598" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The curious genitalic structures of Tancitares michoacanus (Polydesmida, Rhachodesmidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hoffman, Richard L.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>International Journal of Myriapodology</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2011</mods:date>
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<mods:detail type="volume">
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<mods:number>5</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>27</mods:start>
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<mods:end>33</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/ijm.5.1899</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/ijm.5.1899</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1875-2543--27</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="154473563" LSID="urn:lsid:plazi:treatment:9885BCD52CD609CC2C3A2462E0B921F4" httpUri="http://treatment.plazi.org/id/9885BCD52CD609CC2C3A2462E0B921F4" lastPageId="2" lastPageNumber="29" pageId="1" pageNumber="28">
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<subSubSection pageId="1" pageNumber="28" type="nomenclature">
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<paragraph pageId="1" pageNumber="28">
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Genus
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<taxonomicName LSID="http://species-id.net/wiki/Tancitares" authority="Chamberlin" class="Diplopoda" family="Rhachodesmidae" genus="Tancitares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tancitares" order="Polydesmida" pageId="1" pageNumber="28" phylum="Arthropoda" rank="genus">Tancitares Chamberlin</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="28" type="reference_group">
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<paragraph pageId="1" pageNumber="28">
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<taxonomicName class="Diplopoda" family="Rhachodesmidae" genus="Tancitares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tancitares" order="Polydesmida" pageId="1" pageNumber="28" phylum="Arthropoda" rank="genus">Tancitares</taxonomicName>
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Chamberlin, 1942, Proc. Biol. Soc. Washington 55: 58. - Loomis, 1968, U. S. Nat. Mus. Bull. 266: 41. - Hoffman, 1999, Va. Mus. Nat. Hist. Spec. Publ. 8: 429.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="28" type="type species">
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<paragraph pageId="1" pageNumber="28">Type species.</paragraph>
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<paragraph pageId="1" pageNumber="28">
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<taxonomicName class="Diplopoda" family="Rhachodesmidae" genus="Tancitares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tancitares michoacanus" order="Polydesmida" pageId="1" pageNumber="28" phylum="Arthropoda" rank="species" species="michoacanus">Tancitares michoacanus</taxonomicName>
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Chamberlin, by original designation and monotypy.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="28" type="diagnosis">
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<paragraph pageId="1" pageNumber="28">Diagnosis.</paragraph>
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<paragraph pageId="1" pageNumber="28">Males of the type species are distinguished from all other rhachodesmids by the occurrence of digitiform accessory processes on the coxae of the second pair of legs (Fig. 3), and by the reduction of the soleniferous element of the gonotelopodite to a vestigial basal remnant, with hypertrophy of the remaining telopodite (Fig. 7). Females may be recognized by the elongated tubular form of the female genitalia and modified pleurosternal region of the 3rd segment (Fig. 10).</paragraph>
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<caption pageId="1" pageNumber="28">
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<paragraph pageId="1" pageNumber="28">
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Figures 1-4.
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<taxonomicName class="Diplopoda" family="Rhachodesmidae" genus="Tancitares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tancitares michoacanus" order="Polydesmida" pageId="1" pageNumber="28" phylum="Arthropoda" rank="species" species="michoacanus">Tancitares michoacanus</taxonomicName>
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,structural details. 1 Head and metaterga of segments 1-5, dorsal aspect 2 Segments 16-20, dorsal aspect, showing peritremata ending at ozopores 3 Coxae of 2nd pair of legs of male, ventral aspect, showing large digitiform processes directed anteriad 4 Gonopods in situ, ventral aspect, small numbers represent the approximate extent of segments 5-7, leg bases not indicated. Drawings from male topoparatype.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="28" type="range">
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<paragraph pageId="1" pageNumber="28">Range.</paragraph>
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<paragraph pageId="1" pageNumber="28">
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The genus is known only from the western part of the Mexican transverse volcanic belt in
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<normalizedToken originalValue="Michoacán">Michoacan</normalizedToken>
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and Colima.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="2" lastPageNumber="29" pageId="1" pageNumber="28" type="affinities">
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<paragraph pageId="1" pageNumber="28">Affinities.</paragraph>
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<paragraph pageId="1" pageNumber="28">
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Among the ranks of established rhachodesmid genera, only one is a possible close relative of
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<taxonomicName class="Diplopoda" family="Rhachodesmidae" genus="Tancitares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tancitares" order="Polydesmida" pageId="1" pageNumber="28" phylum="Arthropoda" rank="genus">Tancitares</taxonomicName>
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: the rather poorly-known genus
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<taxonomicName class="Diplopoda" family="Rhachodesmidae" genus="Pararhachistes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pararhachistes" order="Polydesmida" pageId="1" pageNumber="28" phylum="Arthropoda" rank="genus">Pararhachistes</taxonomicName>
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Pocock (1909) which was proposed for two species (
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<taxonomicName class="Diplopoda" family="Rhachodesmidae" genus="Pararhachistes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pararhachistes elevatus" order="Polydesmida" pageId="1" pageNumber="28" phylum="Arthropoda" rank="species" species="elevatus">Pararhachistes elevatus</taxonomicName>
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and
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<taxonomicName class="Diplopoda" family="Rhachodesmidae" genus="Pararhachistes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pararhachistes vertebratus" order="Polydesmida" pageId="1" pageNumber="28" phylum="Arthropoda" rank="species" species="vertebratus">Pararhachistes vertebratus</taxonomicName>
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) from the coastal mountains of Guerrero.
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<normalizedToken originalValue="Pocock’s">Pocock's</normalizedToken>
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illustrations (1909: pls. 12, 13) portray a general similarity in body form, the large size of the gonopod aperture, and small secondary processes on the male 2nd coxae. The figure of the gonopod of elevatus (reproduced here as Fig. 8), indicates a slender, sinuous process just distad of the basal fossa that might be the homologue of the solenomere in
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<taxonomicName class="Diplopoda" family="Rhachodesmidae" genus="Tancitares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tancitares" order="Polydesmida" pageId="1" pageNumber="28" phylum="Arthropoda" rank="genus">Tancitares</taxonomicName>
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. Unfortunately the type material of
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<normalizedToken originalValue="Pocock’s">Pocock's</normalizedToken>
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two species was not found during several personal searches of the British Museum myriapod collection, and may be presumed lost, denying the opportunity for an examination with higher magnification. Eventually topotypic material may become available. A third nominal species,
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<taxonomicName class="Diplopoda" family="Rhachodesmidae" genus="Pararhachistes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pararhachistes amblus" order="Polydesmida" pageId="1" pageNumber="28" phylum="Arthropoda" rank="species" species="amblus">Pararhachistes amblus</taxonomicName>
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<bibRefCitation author="Chamberlin, RV" journalOrPublisher="Bulletin of the University of Utah" pageId="6" pageNumber="33" pagination="3 - 19" title="On centipeds and millipeds from Mexican caves." volume="33" year="1942 b">Chamberlin (1942b)</bibRefCitation>
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is known to me from a male paratype. This species is not congeneric with those named by Pocock, differing inter alia in that the efferent duct extends to the apex of the telopodite and is not carried on a slender basal solenomere. Lastly, two other species (galeanae and potosinus) named in
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<taxonomicName class="Diplopoda" family="Rhachodesmidae" genus="Pararhachistes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pararhachistes" order="Polydesmida" pageId="1" pageNumber="28" phylum="Arthropoda" rank="genus">Pararhachistes</taxonomicName>
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by
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<bibRefCitation author="Chamberlin, RV" journalOrPublisher="Bulletin of the University of Utah," pageId="6" pageNumber="33" pagination="3 - 103" title="On Mexican millipeds." volume="34" year="1943">Chamberlin (1943)</bibRefCitation>
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, were based on female specimens which cannot be assigned to any genus with present knowledge of this family. Their geographic origin (San Luis
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<normalizedToken originalValue="Potosí">Potosi</normalizedToken>
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and Nuevo
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<normalizedToken originalValue="León">Leon</normalizedToken>
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) argues against their placement in either
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<taxonomicName class="Diplopoda" family="Rhachodesmidae" genus="Pararhachistes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pararhachistes" order="Polydesmida" pageId="1" pageNumber="28" phylum="Arthropoda" rank="genus">Pararhachistes</taxonomicName>
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or
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<taxonomicName class="Diplopoda" family="Rhachodesmidae" genus="Tancitares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tancitares" order="Polydesmida" pageId="1" pageNumber="28" phylum="Arthropoda" rank="genus">Tancitares</taxonomicName>
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.
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</paragraph>
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<caption pageId="2" pageNumber="29">
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<paragraph pageId="2" pageNumber="29">
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<pageBreakToken pageId="2" pageNumber="29" start="start">Figures</pageBreakToken>
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5-8.
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<taxonomicName class="Diplopoda" family="Rhachodesmidae" genus="Tancitares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tancitares michoacanus" order="Polydesmida" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="michoacanus">Tancitares michoacanus</taxonomicName>
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, gonopod structure. 5 Left gonopod, approximate mesal aspect 6 Apex of telopodite in a different aspect to show entire curvature of subterminal process 7 Telopodite of right gonopod, specimen from Colima, drawn from cleared microscope preparation to show details of basal region, somewhat enlarged over scale of Fig. 5. Figures 5-6 from male topoparatype 8
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<taxonomicName class="Diplopoda" family="Rhachodesmidae" genus="Pararhachistes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pararhachistes elevatus" order="Polydesmida" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="elevatus">Pararhachistes elevatus</taxonomicName>
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Pocock, mesal aspect of left gonopod, for comparison with Fig. 5. Enlarged and redrawn from Fig. 7c, Plate 13, in Pocock 1909.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |