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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.470.8548" ID-PMC="PMC4304036" ID-Pensoft-Pub="1313-2970-470-17" ID-Pensoft-UUID="FFDAFF85127CFFB3AA5915611C3A767A" ID-PubMed="25632243" ID-Zenodo-Dep="578680" ID-ZooBank="0E78A6A90B82401199EED5895E7F8A9E" ModsDocID="1313-2970-470-17" checkinTime="1451244867417" checkinUser="pensoft" docAuthor="Breure, Abraham S. H. &amp; Ablett, Jonathan D." docDate="2015" docId="9885B145E3A25370869DD21302386850" docLanguage="en" docName="ZooKeys 470: 17-143" docOrigin="ZooKeys 470" docPubDate="2015-01-12" docSource="http://dx.doi.org/10.3897/zookeys.470.8548" docTitle="Bulimulus (Drymaeus) interruptus subsp. pallidus Preston 1909" docType="treatment" docVersion="3" id="FFDAFF85127CFFB3AA5915611C3A767A" lastPageNumber="58" masterDocId="FFDAFF85127CFFB3AA5915611C3A767A" masterDocTitle="Annotated type catalogue of the Megaspiridae, Orthalicidae, and Simpulopsidae (Mollusca, Gastropoda, Orthalicoidea) in the Natural History Museum, London" masterLastPageNumber="143" masterPageNumber="17" pageNumber="58" updateTime="1668159795410" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Annotated type catalogue of the Megaspiridae, Orthalicidae, and Simpulopsidae (Mollusca, Gastropoda, Orthalicoidea) in the Natural History Museum, London</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Breure, Abraham S. H.</mods:namePart>
<mods:affiliation>Naturalis Biodiversity Center, P. O. Box 9517, Leiden, the Netherlands</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ablett, Jonathan D.</mods:namePart>
<mods:affiliation>Natural History Museum, Division of Higher Invertebrates, London, SW 7 5 BD, UK</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2015</mods:date>
<mods:detail type="pubDate">
<mods:number>2015-01-12</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>470</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>17</mods:start>
<mods:end>143</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.470.8548</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.470.8548</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-470-17</mods:identifier>
<mods:identifier type="ZooBank">0E78A6A90B82401199EED5895E7F8A9E</mods:identifier>
<mods:identifier type="Pensoft-UUID">FFDAFF85127CFFB3AA5915611C3A767A</mods:identifier>
<mods:identifier type="Zenodo-Dep">578680</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="182234793" LSID="urn:lsid:plazi:treatment:9885B145E3A25370869DD21302386850" httpUri="http://treatment.plazi.org/id/9885B145E3A25370869DD21302386850" lastPageNumber="58" pageId="41" pageNumber="58">
<subSubSection pageId="41" pageNumber="58" type="nomenclature">
<paragraph pageId="41" pageNumber="58">
<taxonomicName LSID="9885B145-E3A2-5370-869D-D21302386850" authority="Preston, 1909" authorityName="Preston" authorityYear="1909" class="Gastropoda" family="Orthalicidae" genus="Bulimulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bulimulus (Drymaeus) interruptus subsp. pallidus" order="Stylommatophora" pageId="41" pageNumber="58" phylum="Mollusca" rank="subSpecies" species="interruptus" subGenus="Drymaeus" subSpecies="pallidus">
<pageBreakToken pageId="41" pageNumber="58" start="start">Bulimulus</pageBreakToken>
(Drymaeus) interruptus pallidus Preston, 1909
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="41" pageNumber="58" type="reference_group">
<paragraph pageId="41" pageNumber="58">
<taxonomicName authorityName="Preston" authorityYear="1909" class="Gastropoda" family="Orthalicidae" genus="Bulimulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bulimulus (Drymaeus) interruptus subsp. var. var. pallidus" order="Stylommatophora" pageId="41" pageNumber="58" phylum="Mollusca" rank="variety" species="interruptus" subGenus="Drymaeus" subSpecies="var." variety="pallidus">Bulimulus (Drymaeus) interruptus var. pallidus</taxonomicName>
<bibRefCitation author="Preston, HB" journalOrPublisher="Annals and Magazine of Natural History" pageId="51" pageNumber="68" pagination="507 - 513" publicationUrl="10.1080/00222930908692616" refId="B105" refString="Preston, HB, 1909. New land, freswater and marine shells from South America. Annals and Magazine of Natural History 3: 507 - 513, DOI: 10.1080/00222930908692616" title="New land, freswater and marine shells from South America." url="10.1080/00222930908692616" volume="3" year="1909">Preston 1909</bibRefCitation>
: 511, fig. 2.
</paragraph>
</subSubSection>
<subSubSection pageId="41" pageNumber="58" type="type locality">
<paragraph pageId="41" pageNumber="58">Type locality.</paragraph>
<paragraph pageId="41" pageNumber="58">&quot;Merida, Venezuela&quot;.</paragraph>
</subSubSection>
<subSubSection pageId="41" pageNumber="58" type="label">
<paragraph pageId="41" pageNumber="58">Label.</paragraph>
<paragraph pageId="41" pageNumber="58">&quot;Merida, Venezuela&quot;.</paragraph>
</subSubSection>
<subSubSection pageId="41" pageNumber="58" type="dimensions">
<paragraph pageId="41" pageNumber="58">Dimensions.</paragraph>
<paragraph pageId="41" pageNumber="58">Not given. Figured specimen H 23.5, D 11.9, W 4.3+.</paragraph>
</subSubSection>
<subSubSection pageId="41" pageNumber="58" type="type material">
<paragraph pageId="41" pageNumber="58">Type material.</paragraph>
<paragraph pageId="41" pageNumber="58">NHMUK 1914.4.3.41, lectotype (ex Preston).</paragraph>
</subSubSection>
<subSubSection pageId="41" pageNumber="58" type="remarks">
<paragraph pageId="41" pageNumber="58">Remarks.</paragraph>
<paragraph pageId="41" pageNumber="58">
This varietal name has been treated as unavailable under Art. 45.6 ICZN by
<bibRefCitation author="Breure, ASH" journalOrPublisher="ZooKeys" pageId="46" pageNumber="63" pagination="1 - 367" publicationUrl="10.3897/zookeys.392.6328" refId="B14" refString="Breure, ASH, Ablett, JD, 2014. Annotated type catalogue of the Bulimulidae (Mollusca, Gastropoda, Orthalicoidea) in the Natural History Museum, London. ZooKeys 392: 1 - 367, DOI: 10.3897/zookeys.392.6328" title="Annotated type catalogue of the Bulimulidae (Mollusca, Gastropoda, Orthalicoidea) in the Natural History Museum, London." url="10.3897/zookeys.392.6328" volume="392" year="2014">Breure and Ablett (2014</bibRefCitation>
: 96). Paul Callomon (pers. commun.) has suggested that this should be reconsidered and doubted if a lectotype of the nominal name already existed.
</paragraph>
<paragraph pageId="41" pageNumber="58">
<emphasis bold="true" pageId="41" pageNumber="58">Our opinion is as follows:</emphasis>
</paragraph>
<paragraph pageId="41" pageNumber="58">
Preston undoubtedly had a series of specimens at hand when describing
<taxonomicName authorityName="Preston" authorityYear="1909" class="Gastropoda" family="Bulimulidae" genus="Bulimulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bulimulus (Drymaeus) interruptus" order="Stylommatophora" pageId="41" pageNumber="58" phylum="Mollusca" rank="species" species="interruptus" subGenus="Drymaeus">Bulimulus (Drymaeus) interruptus</taxonomicName>
; both the wording &quot;to be greatly variable&quot; and &quot;its principle forms&quot; are indicative of this.
</paragraph>
<paragraph pageId="41" pageNumber="58">
<bibRefCitation author="Breure, ASH" journalOrPublisher="Zoologische Verhandelingen Leiden" pageId="45" pageNumber="62" pagination="1 - 215" refId="B10" refString="Breure, ASH, 1979. Systematics, phylogeny and zoogeography of Bulimulinae (Mollusca). Zoologische Verhandelingen Leiden 168: 1 - 215" title="Systematics, phylogeny and zoogeography of Bulimulinae (Mollusca)." volume="168" year="1979">Breure (1979</bibRefCitation>
: 120) mentioned this taxon in his listing under
<taxonomicName authority="" authorityName="Albers" authorityYear="1850" class="Gastropoda" family="Bulimulidae" genus="Drymaeus" kingdom="Animalia" lsidName="Drymaeus (Mesembrinus)" order="Stylommatophora" pageId="41" pageNumber="58" phylum="Mollusca" rank="subGenus" subGenus="Mesembrinus">Drymaeus (Mesembrinus)</taxonomicName>
Albers, 1850, and stated &quot;HT BMNH 1914.4.3.38&quot; [referring to a single specimen, thus qualifying Art. 74.6.1.2 jo. 74.3]; Art. 74.6 ICZN rules this statement as a lectotype designation. It should be noted, however, that Breure did not list the &quot;var.
<taxonomicName class="Gastropoda" family="Bulimulidae" genus="Drymaeus" kingdom="Animalia" lsidName="var. pallidus" order="Stylommatophora" pageId="41" pageNumber="58" phylum="Mollusca" rank="variety" species="interruptus" variety="pallidus">
<emphasis italics="true" pageId="41" pageNumber="58">pallidus</emphasis>
</taxonomicName>
&quot; of Preston in his paper (cf. point d below).
</paragraph>
<paragraph pageId="41" pageNumber="58">
<normalizedToken originalValue="Köhler">Koehler</normalizedToken>
(1997) also concluded that this taxon was described from several specimens but said &quot;A holotype has not been designated. Therefore, the present specimen [ZMB 59597] is a syntype. Consequently, the specimen in the BMNH referred to as holotype by
<bibRefCitation author="Breure, ASH" journalOrPublisher="Zoologische Verhandelingen Leiden" pageId="45" pageNumber="62" pagination="1 - 215" refId="B10" refString="Breure, ASH, 1979. Systematics, phylogeny and zoogeography of Bulimulinae (Mollusca). Zoologische Verhandelingen Leiden 168: 1 - 215" title="Systematics, phylogeny and zoogeography of Bulimulinae (Mollusca)." volume="168" year="1979">Breure (1979)</bibRefCitation>
is a syntype&quot;. Overlooking, as explained in the previous item, the lectotype designation under Art. 74.6 (see previous point).
</paragraph>
<paragraph pageId="41" pageNumber="58">
Reconsidering the (un)availability under Art. 45.6, it is important to note that var.
<taxonomicName class="Gastropoda" family="Bulimulidae" genus="Drymaeus" kingdom="Animalia" lsidName="var. pallidus" order="Stylommatophora" pageId="41" pageNumber="58" phylum="Mollusca" rank="variety" species="interruptus" variety="pallidus">
<emphasis italics="true" pageId="41" pageNumber="58">pallidus</emphasis>
</taxonomicName>
was proposed before 1961 and has to be treated as subspecific (see the contributions of Steve Lingafelter and Doug Yanega on the Taxacom listing, http://to.ly/zFZO). It may be noted that the only reference to this taxon after
<normalizedToken originalValue="Prestons">Preston's</normalizedToken>
publication is in
<bibRefCitation author="Baker, HB" journalOrPublisher="Occasional Papers of the Museum of Zoology, University of Michigan" pageId="45" pageNumber="62" pagination="1 - 49" refId="B5" refString="Baker, HB, 1926. The Mollusca collected by the University of Michigan-Williamson expedition in Venezuela, IV. Occasional Papers of the Museum of Zoology, University of Michigan 167: 1 - 49" title="The Mollusca collected by the University of Michigan-Williamson expedition in Venezuela, IV." volume="167" year="1926">Baker (1926</bibRefCitation>
: 44), who regarded it as a synonym of
<taxonomicName authorityName="L.Pfeiffer" authorityYear="1848" class="Gastropoda" family="Bulimulidae" genus="Bulimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bulimus granadensis" order="Pulmonata" pageId="41" pageNumber="58" phylum="Mollusca" rank="species" species="granadensis">
<emphasis italics="true" pageId="41" pageNumber="58">Bulimus granadensis</emphasis>
</taxonomicName>
Pfeiffer, 1848 (see also
<bibRefCitation author="Richardson, CL" journalOrPublisher="Tryonia" pageId="51" pageNumber="68" refId="B114" refString="Richardson, CL, 1995. Bulimulidae: catalog of species. Tryonia 28" title="Bulimulidae: catalog of species." volume="28" year="1995">Richardson 1995</bibRefCitation>
: 133).
</paragraph>
<paragraph pageId="41" pageNumber="58">
While
<taxonomicName authorityName="Preston" authorityYear="1909" class="Gastropoda" family="Bulimulidae" genus="Bulimulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bulimulus (Drymaeus) interruptus subsp. var. var. pallidus" order="Stylommatophora" pageId="41" pageNumber="58" phylum="Mollusca" rank="variety" species="interruptus" subGenus="Drymaeus" subSpecies="var." variety="pallidus">Bulimulus (Drymaeus) interruptus var. pallidus</taxonomicName>
is an available name, we concur with
<bibRefCitation author="Baker, HB" journalOrPublisher="Occasional Papers of the Museum of Zoology, University of Michigan" pageId="45" pageNumber="62" pagination="1 - 49" refId="B5" refString="Baker, HB, 1926. The Mollusca collected by the University of Michigan-Williamson expedition in Venezuela, IV. Occasional Papers of the Museum of Zoology, University of Michigan 167: 1 - 49" title="The Mollusca collected by the University of Michigan-Williamson expedition in Venezuela, IV." volume="167" year="1926">Baker (1926)</bibRefCitation>
and
<bibRefCitation author="Richardson, CL" journalOrPublisher="Tryonia" pageId="51" pageNumber="68" refId="B114" refString="Richardson, CL, 1995. Bulimulidae: catalog of species. Tryonia 28" title="Bulimulidae: catalog of species." volume="28" year="1995">Richardson (1995)</bibRefCitation>
to consider this taxon as a synonym of the nominal form.
</paragraph>
<paragraph pageId="41" pageNumber="58">
The specimen of var.
<taxonomicName authorityName="Preston" authorityYear="1909" class="Gastropoda" family="Bulimulidae" genus="Bulimulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="var. pallidus" order="Stylommatophora" pageId="41" pageNumber="58" phylum="Mollusca" rank="variety" species="interruptus" variety="pallidus">
<emphasis italics="true" pageId="41" pageNumber="58">pallidus</emphasis>
</taxonomicName>
in the NHMUK is now designated lectotype (
<emphasis bold="true" pageId="41" pageNumber="58">design. n.</emphasis>
) to fixate this synonymisation. The text in
<bibRefCitation author="Breure, ASH" journalOrPublisher="ZooKeys" pageId="46" pageNumber="63" pagination="1 - 367" publicationUrl="10.3897/zookeys.392.6328" refId="B14" refString="Breure, ASH, Ablett, JD, 2014. Annotated type catalogue of the Bulimulidae (Mollusca, Gastropoda, Orthalicoidea) in the Natural History Museum, London. ZooKeys 392: 1 - 367, DOI: 10.3897/zookeys.392.6328" title="Annotated type catalogue of the Bulimulidae (Mollusca, Gastropoda, Orthalicoidea) in the Natural History Museum, London." url="10.3897/zookeys.392.6328" volume="392" year="2014">Breure and Ablett (2014</bibRefCitation>
: 96), under 'Type
<normalizedToken originalValue="material">material'</normalizedToken>
, should be corrected as follows: &quot;NHMUK 1914.3.38, lectotype; 1914.4.3.39-40, 42-43, four paralectotypes&quot;.
</paragraph>
</subSubSection>
<subSubSection pageId="41" pageNumber="58" type="current systematic position">
<paragraph pageId="41" pageNumber="58">Current systematic position.</paragraph>
<paragraph pageId="41" pageNumber="58">
<taxonomicName authorityName="Tryon" authorityYear="1867" class="Gastropoda" family="Bulimulidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pulmonata" pageId="41" pageNumber="58" phylum="Mollusca" rank="family">Bulimulidae</taxonomicName>
,
<taxonomicName baseAuthorityName="Preston" baseAuthorityYear="1909" class="Gastropoda" family="Bulimulidae" genus="Drymaeus" kingdom="Animalia" lsidName="Drymaeus (Mesembrinus) interruptus" order="Stylommatophora" pageId="41" pageNumber="58" phylum="Mollusca" rank="species" species="interruptus" subGenus="Mesembrinus">Drymaeus (Mesembrinus) interruptus</taxonomicName>
(Preston, 1909).
</paragraph>
</subSubSection>
</treatment>
</document>