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<document id="33FB7694A94C4C3B13D26BBC09AFD699" ID-CLB-Dataset="39076" ID-DOI="10.11646/zootaxa.4111.2.1" ID-GBIF-Dataset="ee60b4f8-1cce-4e63-89e6-126b89d45e64" ID-ISSN="1175-5326" ID-Zenodo-Dep="257706" ID-ZooBank="0208587B-598C-4202-A4BA-8FDECD2C93B9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1463119531886" checkinUser="plazi" docAuthor="Anker, Arthur &amp; Pachelle, Paulo P. G." docDate="2016" docId="895FDE15FF069E75FF5AFEB1FE3BD5E9" docLanguage="en" docName="zootaxa.4111.2.1.pdf.imf" docOrigin="Zootaxa 4111 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Axianassa darrylfelderi Anker &amp; Lazarus 2015" docType="treatment" docVersion="13" lastPageNumber="110" masterDocId="7566A66DFF0E9E7CFFCDFF96FFCBD251" masterDocTitle="Mud-shrimps of the genus Axianassa Schmitt, 1924 from Panama, with description of two new species (Decapoda: Gebiidea: Laomediidae)" masterLastPageNumber="125" masterPageNumber="101" pageNumber="109" updateTime="1698662496277" updateUser="ExternalLinkService">
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<mods:title id="A938CF45B5B742BDDBDCC2D19795CD6E">Mud-shrimps of the genus Axianassa Schmitt, 1924 from Panama, with description of two new species (Decapoda: Gebiidea: Laomediidae)</mods:title>
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<mods:namePart id="A1C5E1113CA85C5EE2C1BF21D19FF90A">Anker, Arthur</mods:namePart>
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<mods:namePart id="94A3F79AC60D894AF90C4DED4E6BDD86">Pachelle, Paulo P. G.</mods:namePart>
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<mods:date id="FF7E0101DBC260445FF606FFF8588CE5">2016</mods:date>
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<treatment id="895FDE15FF069E75FF5AFEB1FE3BD5E9" ID-DOI="http://doi.org/10.5281/zenodo.6062664" ID-GBIF-Taxon="120677865" ID-Zenodo-Dep="6062664" LSID="urn:lsid:plazi:treatment:895FDE15FF069E75FF5AFEB1FE3BD5E9" httpUri="http://treatment.plazi.org/id/895FDE15FF069E75FF5AFEB1FE3BD5E9" lastPageId="9" lastPageNumber="110" pageId="8" pageNumber="109">
<subSubSection id="49EC3C88FF069E74FF5AFEB1FF20D335" pageId="8" pageNumber="109" type="nomenclature">
<paragraph id="01496F03FF069E74FF5AFEB1FD0CD311" blockId="8.[151,711,295,357]" box="[151,711,295,320]" pageId="8" pageNumber="109">
<heading id="5A01D86FFF069E74FF5AFEB1FD0CD311" bold="true" box="[151,711,295,320]" fontSize="10" level="2" pageId="8" pageNumber="109" reason="2">
<taxonomicName id="C6F61480FF069E74FF5AFEB1FD0CD311" ID-CoL="bbb4befe-6b46-402a-bf6c-0feeb661b947" authority="Anker &amp; Lazarus, 2015" authorityName="Anker &amp; Lazarus" authorityYear="2015" box="[151,711,295,320]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="109" phylum="Arthropoda" rank="species" species="darrylfelderi">
<emphasis id="3382B311FF069E74FF5AFEB1FD0CD311" bold="true" box="[151,711,295,320]" pageId="8" pageNumber="109">
<emphasis id="3382B311FF069E74FF5AFEB1FE6DD311" bold="true" box="[151,422,295,320]" italics="true" pageId="8" pageNumber="109">Axianassa darrylfelderi</emphasis>
<bibRefCitation id="656712F2FF069E74FE60FEB1FD0CD311" author="Anker" box="[429,711,295,320]" pageId="8" pageNumber="109" refString="Anker, A. &amp; Lazarus, J. F. (2015) Description of two new associated infaunal decapod crustaceans (Axianassidae and Alpheidae) from the tropical eastern Pacific. Papeis Avulsos de Zoologia, 55, 115 - 129. http: // dx. doi. org / 10.1590 / 0031 - 1049.2015.55.08" type="journal article" year="2015">Anker &amp; Lazarus, 2015</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="01496F03FF069E74FF5AFEDAFF20D335" blockId="8.[151,711,295,357]" box="[151,235,332,357]" pageId="8" pageNumber="109">
(
<figureCitation id="99CD7386FF069E74FF52FEDAFF28D335" box="[159,227,332,357]" captionStart="FIGURE 7" captionStartId="8.[151,244,1798,1820]" captionTargetBox="[151,1436,690,1776]" captionTargetId="figure@8.[151,1436,690,1777]" captionTargetPageId="8" captionText="FIGURE 7. Axianassa darrylfelderi Anker &amp; Lazarus, 2015: A – C—male (cl 8.9 mm) from Azuero Peninsula, Panama (MZUSP 34081); A—dorsal view; B—detached major cheliped, mesial view; C—detached minor cheliped, mesial view; D, E—ovigerous female (cl 9.0 mm) from the same locality (MZUSP 34489), in dorsal (D) and lateral (E) views." httpUri="https://zenodo.org/record/257713/files/figure.png" pageId="8" pageNumber="109">Fig. 7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="49EC3C88FF069E74FF5AFE02FCDFD3FB" box="[151,788,404,426]" pageId="8" pageNumber="109" type="reference_group">
<paragraph id="01496F03FF069E74FF5AFE02FCDFD3FB" blockId="8.[151,788,404,426]" box="[151,788,404,426]" pageId="8" pageNumber="109">
<treatmentCitationGroup id="21E6482DFF069E74FF5AFE02FCDFD3FB" box="[151,788,404,426]" pageId="8" pageNumber="109">
<taxonomicName id="C6F61480FF069E74FF5AFE02FD61D3FB" ID-CoL="bbb4befe-6b46-402a-bf6c-0feeb661b947" authority="Anker &amp; Lazarus 2015: 116" authorityName="Anker &amp; Lazarus" authorityPageNumber="116" authorityYear="2015" box="[151,682,404,426]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="109" phylum="Arthropoda" rank="species" species="darrylfelderi">
<emphasis id="3382B311FF069E74FF5AFE02FE4DD3FB" box="[151,390,404,426]" italics="true" pageId="8" pageNumber="109">Axianassa darrylfelderi</emphasis>
<treatmentCitation id="80574912FF069E74FE40FE02FD61D3FB" author="Anker" box="[397,682,404,426]" page="116" pageId="8" pageNumber="109" year="2015">
<bibRefCitation id="656712F2FF069E74FE40FE02FDB2D3FB" author="Anker" box="[397,633,404,426]" pageId="8" pageNumber="109" refString="Anker, A. &amp; Lazarus, J. F. (2015) Description of two new associated infaunal decapod crustaceans (Axianassidae and Alpheidae) from the tropical eastern Pacific. Papeis Avulsos de Zoologia, 55, 115 - 129. http: // dx. doi. org / 10.1590 / 0031 - 1049.2015.55.08" type="journal article" year="2015">Anker &amp; Lazarus 2015</bibRefCitation>
: 116
</treatmentCitation>
</taxonomicName>
, figs. 1–4.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="49EC3C88FF069E74FF5AFE40FBF9D00A" pageId="8" pageNumber="109" type="materials_examined">
<paragraph id="01496F03FF069E74FF5AFE40FBF9D00A" blockId="8.[151,1436,470,639]" pageId="8" pageNumber="109">
<emphasis id="3382B311FF069E74FF5AFE40FEB7D3BE" bold="true" box="[151,380,470,495]" pageId="8" pageNumber="109">Material examined</emphasis>
.
<specimenCount id="17F0A48AFF069E74FE41FE41FE11D3BE" box="[396,474,470,495]" pageId="8" pageNumber="109" type="male">1 male</specimenCount>
(cl
<quantity id="C60EC2E6FF069E74FDC6FE41FDA3D3BE" box="[523,616,471,495]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="8" pageNumber="109" unit="mm" value="8.9">8.9 mm</quantity>
),
<collectionCode id="67E7F7C6FF069E74FDB0FE41FD11D3BE" box="[637,730,471,495]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="8" pageNumber="109">MZUSP</collectionCode>
34081, Pacific coast of
<collectingCountry id="79E12F93FF069E74FC3AFE41FB98D3BF" box="[1015,1107,471,494]" name="Panama" pageId="8" pageNumber="109">Panama</collectingCountry>
, Playa El Agallito, intertidal sand-mud flat, in burrow, coll. A. Anker, J.F. Lazarus, T. Kaji,
<date id="754849C3FF069E74FC43FE6CFBC1D042" box="[910,1034,506,531]" pageId="8" pageNumber="109" value="2015-03-22">22.iii.2015</date>
; 1 ovigerous female (cl 9.0 mm),
<collectionCode id="67E7F7C6FF069E74FF5AFD89FF3FD066" box="[151,244,543,567]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="8" pageNumber="109">MZUSP</collectionCode>
34489, Pacific coast of
<collectingCountry id="79E12F93FF069E74FDD7FD89FDBDD067" box="[538,630,543,566]" name="Panama" pageId="8" pageNumber="109">Panama</collectingCountry>
, Playa El Agallito, intertidal sand-mud flat, in burrow together with alpheid shrimp (
<taxonomicName id="C6F61480FF069E74FE83FDD5FE1CD00A" box="[334,471,579,603]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="109" phylum="Arthropoda" rank="genus">
<emphasis id="3382B311FF069E74FE83FDD5FE1CD00A" box="[334,471,579,603]" italics="true" pageId="8" pageNumber="109">Leptalpheus</emphasis>
</taxonomicName>
sp.), coll. A. Anker, J.F. Lazarus, T. Kaji,
<date id="754849C3FF069E74FC7FFDD4FBE4D00A" box="[946,1071,578,603]" pageId="8" pageNumber="109" value="2015-03-22">22.iii.2015</date>
.
</paragraph>
</subSubSection>
<subSubSection id="49EC3C88FF069E75FF0AFDF0FC2CD285" lastPageId="9" lastPageNumber="110" pageId="8" pageNumber="109" type="description">
<paragraph id="01496F03FF069E74FF0AFDF0FACAD02F" blockId="8.[151,1436,470,639]" box="[199,1281,614,639]" pageId="8" pageNumber="109">
<emphasis id="3382B311FF069E74FF0AFDF0FE9AD02E" bold="true" box="[199,337,614,639]" pageId="8" pageNumber="109">Description</emphasis>
. See
<bibRefCitation id="656712F2FF069E74FE42FDF0FD6DD02E" author="Anker" box="[399,678,614,639]" pageId="8" pageNumber="109" refString="Anker, A. &amp; Lazarus, J. F. (2015) Description of two new associated infaunal decapod crustaceans (Axianassidae and Alpheidae) from the tropical eastern Pacific. Papeis Avulsos de Zoologia, 55, 115 - 129. http: // dx. doi. org / 10.1590 / 0031 - 1049.2015.55.08" type="journal article" year="2015">Anker &amp; Lazarus (2015)</bibRefCitation>
; additional colour photographs are provided in
<figureCitation id="99CD7386FF069E74FB74FDF0FB36D02F" box="[1209,1277,614,639]" captionStart="FIGURE 7" captionStartId="8.[151,244,1798,1820]" captionTargetBox="[151,1436,690,1776]" captionTargetId="figure@8.[151,1436,690,1777]" captionTargetPageId="8" captionText="FIGURE 7. Axianassa darrylfelderi Anker &amp; Lazarus, 2015: A – C—male (cl 8.9 mm) from Azuero Peninsula, Panama (MZUSP 34081); A—dorsal view; B—detached major cheliped, mesial view; C—detached minor cheliped, mesial view; D, E—ovigerous female (cl 9.0 mm) from the same locality (MZUSP 34489), in dorsal (D) and lateral (E) views." httpUri="https://zenodo.org/record/257713/files/figure.png" pageId="8" pageNumber="109">Fig. 7</figureCitation>
.
</paragraph>
<caption id="55893F8BFF069E74FF5AF890FBE6D507" httpUri="https://zenodo.org/record/257713/files/figure.png" pageId="8" pageNumber="109" targetBox="[151,1436,690,1776]" targetPageId="8">
<paragraph id="01496F03FF069E74FF5AF890FBE6D507" blockId="8.[151,1436,1798,1878]" pageId="8" pageNumber="109">
<emphasis id="3382B311FF069E74FF5AF890FEC2D54D" bold="true" box="[151,265,1798,1820]" pageId="8" pageNumber="109">FIGURE 7</emphasis>
.
<taxonomicName id="C6F61480FF069E74FED5F890FD3ED54A" authority="Anker &amp; Lazarus, 2015" authorityName="Anker &amp; Lazarus" authorityYear="2015" box="[280,757,1798,1819]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="109" phylum="Arthropoda" rank="species" species="darrylfelderi">
<emphasis id="3382B311FF069E74FED5F890FE35D54A" box="[280,510,1798,1819]" italics="true" pageId="8" pageNumber="109">Axianassa darrylfelderi</emphasis>
Anker &amp; Lazarus, 2015
</taxonomicName>
: A–C—male (cl 8.9 mm) from
<taxonomicName id="C6F61480FF069E74FBFEF890FBB2D54A" box="[1075,1145,1798,1819]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="116" phylum="Arthropoda" rank="species" species="azuero">Azuero</taxonomicName>
Peninsula, Panama (MZUSP 34081); A—dorsal view; B—detached major cheliped, mesial view; C—detached minor cheliped, mesial view; D, E—ovigerous female (cl 9.0 mm) from the same locality (MZUSP 34489), in dorsal (D) and lateral (E) views.
</paragraph>
</caption>
<paragraph id="01496F03FF079E75FF0AFF01FC2CD285" blockId="9.[151,1437,151,1977]" pageId="9" pageNumber="110">
<emphasis id="3382B311FF079E75FF0AFF01FE93D2E1" bold="true" box="[199,344,151,176]" pageId="9" pageNumber="110">Colouration</emphasis>
. Pale pink to more intense reddish pink, depending on state of chromatophore contraction; inner organs yellow or orange (
<figureCitation id="99CD7386FF079E75FE78FF2AFE32D285" box="[437,505,188,213]" captionStart="FIGURE 7" captionStartId="8.[151,244,1798,1820]" captionTargetBox="[151,1436,690,1776]" captionTargetId="figure@8.[151,1436,690,1777]" captionTargetPageId="8" captionText="FIGURE 7. Axianassa darrylfelderi Anker &amp; Lazarus, 2015: A – C—male (cl 8.9 mm) from Azuero Peninsula, Panama (MZUSP 34081); A—dorsal view; B—detached major cheliped, mesial view; C—detached minor cheliped, mesial view; D, E—ovigerous female (cl 9.0 mm) from the same locality (MZUSP 34489), in dorsal (D) and lateral (E) views." httpUri="https://zenodo.org/record/257713/files/figure.png" pageId="9" pageNumber="110">Fig. 7</figureCitation>
; see also
<bibRefCitation id="656712F2FF079E75FDA8FF2AFCA1D284" author="Anker" box="[613,874,188,213]" pageId="9" pageNumber="110" refString="Anker, A. &amp; Lazarus, J. F. (2015) Description of two new associated infaunal decapod crustaceans (Axianassidae and Alpheidae) from the tropical eastern Pacific. Papeis Avulsos de Zoologia, 55, 115 - 129. http: // dx. doi. org / 10.1590 / 0031 - 1049.2015.55.08" type="journal article" year="2015">Anker &amp; Lazarus 2015</bibRefCitation>
: figs. 3, 4).
</paragraph>
</subSubSection>
<subSubSection id="49EC3C88FF079E75FF0AFF49FBBFD334" pageId="9" pageNumber="110" type="distribution">
<paragraph id="01496F03FF079E75FF0AFF49FD9DD34C" blockId="9.[151,1437,151,1977]" pageId="9" pageNumber="110">
<emphasis id="3382B311FF079E75FF0AFF49FE92D2A9" bold="true" box="[199,345,223,248]" pageId="9" pageNumber="110">Distribution</emphasis>
. Eastern Pacific:
<collectingCountry id="79E12F93FF079E75FDE8FF49FD58D2A9" box="[549,659,223,248]" name="Colombia" pageId="9" pageNumber="110">Colombia</collectingCountry>
(Bahía Málaga) and
<collectingCountry id="79E12F93FF079E75FC44FF76FC29D2A6" box="[905,994,224,247]" name="Panama" pageId="9" pageNumber="110">Panama</collectingCountry>
(Playa El Agallito,
<taxonomicName id="C6F61480FF079E75FB07FF76FAD6D2A9" box="[1226,1309,224,248]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="116" phylum="Arthropoda" rank="species" species="azuero">Azuero</taxonomicName>
Peninsula) (
<bibRefCitation id="656712F2FF079E75FF53FE92FD82D34C" author="Anker" box="[158,585,260,285]" pageId="9" pageNumber="110" refString="Anker, A. &amp; Lazarus, J. F. (2015) Description of two new associated infaunal decapod crustaceans (Axianassidae and Alpheidae) from the tropical eastern Pacific. Papeis Avulsos de Zoologia, 55, 115 - 129. http: // dx. doi. org / 10.1590 / 0031 - 1049.2015.55.08" type="journal article" year="2015">Anker &amp; Lazarus 2015; present study</bibRefCitation>
).
</paragraph>
<paragraph id="01496F03FF079E75FF0AFEB1FBBFD334" blockId="9.[151,1437,151,1977]" pageId="9" pageNumber="110">
<emphasis id="3382B311FF079E75FF0AFEB1FEE0D311" bold="true" box="[199,299,295,320]" pageId="9" pageNumber="110">Ecology.</emphasis>
Intertidal and shallow subtidal (less than
<quantity id="C60EC2E6FF079E75FD32FEB1FC8BD311" box="[767,832,295,320]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="110" unit="m" value="0.5">0.5 m</quantity>
) mud flats, sometimes near mangroves; in burrows in mud, sometimes with commensal alpheid shrimps,
<taxonomicName id="C6F61480FF079E75FD18FEDAFC95D335" box="[725,862,332,356]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="genus">
<emphasis id="3382B311FF079E75FD18FEDAFC95D335" box="[725,862,332,356]" italics="true" pageId="9" pageNumber="110">Leptalpheus</emphasis>
</taxonomicName>
sp. (A. Anker, in study).
</paragraph>
</subSubSection>
<subSubSection id="49EC3C88FF079E75FF0AFEF9FE3BD5E9" pageId="9" pageNumber="110" type="discussion">
<paragraph id="01496F03FF079E75FF0AFEF9FB79D3A4" blockId="9.[151,1437,151,1977]" pageId="9" pageNumber="110">
<emphasis id="3382B311FF079E75FF0AFEF9FEFFD3D9" bold="true" box="[199,308,367,392]" pageId="9" pageNumber="110">Remarks</emphasis>
. The recently described
<taxonomicName id="C6F61480FF079E75FD81FEE6FC9DD3D9" box="[588,854,368,392]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="darrylfelderi">
<emphasis id="3382B311FF079E75FD81FEE6FC9DD3D9" box="[588,854,368,392]" italics="true" pageId="9" pageNumber="110">Axianassa darrylfelderi</emphasis>
</taxonomicName>
was previously only known from the
<typeStatus id="DE4DD1A1FF079E75FAC5FEE6FAF3D3D9" box="[1288,1336,368,392]" pageId="9" pageNumber="110">type</typeStatus>
material from Bahía Málaga,
<collectingCountry id="79E12F93FF079E75FE4EFE02FE39D3FC" box="[387,498,404,429]" name="Colombia" pageId="9" pageNumber="110">Colombia</collectingCountry>
(
<bibRefCitation id="656712F2FF079E75FDCCFE02FCC6D3FC" author="Anker" box="[513,781,404,429]" pageId="9" pageNumber="110" refString="Anker, A. &amp; Lazarus, J. F. (2015) Description of two new associated infaunal decapod crustaceans (Axianassidae and Alpheidae) from the tropical eastern Pacific. Papeis Avulsos de Zoologia, 55, 115 - 129. http: // dx. doi. org / 10.1590 / 0031 - 1049.2015.55.08" type="journal article" year="2015">Anker &amp; Lazarus 2015</bibRefCitation>
). The two specimens collected on the eastern coastline of Panama’s
<taxonomicName id="C6F61480FF079E75FEC1FE2EFE94D381" box="[268,351,440,464]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="116" phylum="Arthropoda" rank="species" species="azuero">Azuero</taxonomicName>
Peninsula represent only the second record of
<taxonomicName id="C6F61480FF079E75FC4CFE2EFBF9D381" box="[897,1074,440,464]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="darrylfelderi">
<emphasis id="3382B311FF079E75FC4CFE2EFBF9D381" box="[897,1074,440,464]" italics="true" pageId="9" pageNumber="110">A. darrylfelderi</emphasis>
</taxonomicName>
and also the first record of the species for
<collectingCountry id="79E12F93FF079E75FEDBFE4BFEB9D3A5" box="[278,370,477,500]" name="Panama" pageId="9" pageNumber="110">Panama</collectingCountry>
. They match the
<typeStatus id="DE4DD1A1FF079E75FDFFFE4BFDA9D3A4" box="[562,610,477,501]" pageId="9" pageNumber="110">type</typeStatus>
material from
<collectingCountry id="79E12F93FF079E75FCC7FE4AFCB3D3A4" box="[778,888,476,501]" name="Colombia" pageId="9" pageNumber="110">Colombia</collectingCountry>
in all diagnostic characters.
</paragraph>
<paragraph id="01496F03FF079E75FF0AFE69FB64D0A1" blockId="9.[151,1437,151,1977]" pageId="9" pageNumber="110">
<bibRefCitation id="656712F2FF079E75FF0AFE69FE16D049" author="Anker" box="[199,477,511,536]" pageId="9" pageNumber="110" refString="Anker, A. &amp; Lazarus, J. F. (2015) Description of two new associated infaunal decapod crustaceans (Axianassidae and Alpheidae) from the tropical eastern Pacific. Papeis Avulsos de Zoologia, 55, 115 - 129. http: // dx. doi. org / 10.1590 / 0031 - 1049.2015.55.08" type="journal article" year="2015">Anker &amp; Lazarus (2015)</bibRefCitation>
used three characters to separate
<taxonomicName id="C6F61480FF079E75FC95FD96FBCCD049" box="[856,1031,512,536]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="darrylfelderi">
<emphasis id="3382B311FF079E75FC95FD96FBCCD049" box="[856,1031,512,536]" italics="true" pageId="9" pageNumber="110">A. darrylfelderi</emphasis>
</taxonomicName>
from its presumed western Atlantic sister species,
<taxonomicName id="C6F61480FF079E75FE8DFDB2FCD2D06D" authority="Rodrigues &amp; Shimizu, 1992" authorityName="Rodrigues &amp; Shimizu" authorityYear="1992" box="[320,793,548,573]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="3382B311FF079E75FE8DFDB2FE0ED06D" box="[320,453,548,572]" italics="true" pageId="9" pageNumber="110">A. australis</emphasis>
<bibRefCitation id="656712F2FF079E75FE1DFDB2FCD2D06D" author="Rodrigues" box="[464,793,548,573]" pageId="9" pageNumber="110" refString="Rodrigues, S. A. &amp; Shimizu, R. M. (1992) Description of a new Axianassa (Crustacea: Decapoda: Thalassinidea) from Brazil, and its first larval stage. Proceedings of the Biological Society of Washington, 105, 317 - 323. http: // biodiversitylibrary. org / page / 35607490" type="journal article" year="1992">Rodrigues &amp; Shimizu, 1992</bibRefCitation>
</taxonomicName>
, based on the description and illustrations provided by
<bibRefCitation id="656712F2FF079E75FF5AFDD1FE15D031" author="Rodrigues" box="[151,478,583,608]" pageId="9" pageNumber="110" refString="Rodrigues, S. A. &amp; Shimizu, R. M. (1992) Description of a new Axianassa (Crustacea: Decapoda: Thalassinidea) from Brazil, and its first larval stage. Proceedings of the Biological Society of Washington, 105, 317 - 323. http: // biodiversitylibrary. org / page / 35607490" type="journal article" year="1992">Rodrigues &amp; Shimizu (1992)</bibRefCitation>
. These characters are the presence or absence of teeth on the distolateral margin of the uropodal exopod (two or three in
<taxonomicName id="C6F61480FF079E75FDE0FDFAFD2BD0D5" box="[557,736,620,644]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="darrylfelderi">
<emphasis id="3382B311FF079E75FDE0FDFAFD2BD0D5" box="[557,736,620,644]" italics="true" pageId="9" pageNumber="110">A. darrylfelderi</emphasis>
</taxonomicName>
vs. none in
<taxonomicName id="C6F61480FF079E75FCB1FDFAFBC9D0D5" box="[892,1026,620,644]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="3382B311FF079E75FCB1FDFAFBC9D0D5" box="[892,1026,620,644]" italics="true" pageId="9" pageNumber="110">A. australis</emphasis>
</taxonomicName>
), the length of the antennal acicle (relatively longer in
<taxonomicName id="C6F61480FF079E75FEB6FD06FDE3D0F9" box="[379,552,656,680]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="darrylfelderi">
<emphasis id="3382B311FF079E75FEB6FD06FDE3D0F9" box="[379,552,656,680]" italics="true" pageId="9" pageNumber="110">A. darrylfelderi</emphasis>
</taxonomicName>
compared to
<taxonomicName id="C6F61480FF079E75FD0DFD06FC8AD0F9" box="[704,833,656,680]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="3382B311FF079E75FD0DFD06FC8AD0F9" box="[704,833,656,680]" italics="true" pageId="9" pageNumber="110">A. australis</emphasis>
</taxonomicName>
) and the armature of the cutting edges of the pollex in the male minor cheliped (with three strong double teeth, in addition to numerous smaller teeth in
<taxonomicName id="C6F61480FF079E75FB21FD22FA57D09D" box="[1260,1436,692,716]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="darrylfelderi">
<emphasis id="3382B311FF079E75FB21FD22FA57D09D" box="[1260,1436,692,716]" italics="true" pageId="9" pageNumber="110">A. darrylfelderi</emphasis>
</taxonomicName>
vs. with one proximal double or triple tooth and two simple, more distal teeth in
<taxonomicName id="C6F61480FF079E75FBD0FD4EFB55D0A1" box="[1053,1182,728,752]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="3382B311FF079E75FBD0FD4EFB55D0A1" box="[1053,1182,728,752]" italics="true" pageId="9" pageNumber="110">A. australis</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="01496F03FF079E75FF0AFD6AFC78D6F1" blockId="9.[151,1437,151,1977]" pageId="9" pageNumber="110">
A thorough examination of the Panamanian specimens of
<taxonomicName id="C6F61480FF079E75FC9EFD6AFBCAD145" box="[851,1025,764,788]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="darrylfelderi">
<emphasis id="3382B311FF079E75FC9EFD6AFBCAD145" box="[851,1025,764,788]" italics="true" pageId="9" pageNumber="110">A. darrylfelderi</emphasis>
</taxonomicName>
revealed the presence of five (male) and one to three (female) teeth and a small movable spinule or spiniform seta (in both male and female), the latter obscured by the adjacent plumose setae, on the distolateral margin of the uropodal exopod. This observation prompted a re-examination of the male
<typeStatus id="DE4DD1A1FF079E75FD90FCFEFD76D1D1" box="[605,701,872,896]" pageId="9" pageNumber="110" type="paratype">paratype</typeStatus>
from
<collectingCountry id="79E12F93FF079E75FCC8FCF1FCB8D1D1" box="[773,883,871,896]" name="Colombia" pageId="9" pageNumber="110">Colombia</collectingCountry>
(MZUSP 33014), which was confirmed to have only one tooth and one spiniform seta, the latter having been either overlooked (being concealed by the adjacent thick plumose setae) or illustrated as a fixed tooth in
<bibRefCitation id="656712F2FF079E75FD29FC39FB9CD199" author="Anker" box="[740,1111,943,968]" pageId="9" pageNumber="110" refString="Anker, A. &amp; Lazarus, J. F. (2015) Description of two new associated infaunal decapod crustaceans (Axianassidae and Alpheidae) from the tropical eastern Pacific. Papeis Avulsos de Zoologia, 55, 115 - 129. http: // dx. doi. org / 10.1590 / 0031 - 1049.2015.55.08" type="journal article" year="2015">Anker &amp; Lazarus (2015: fig. 1K)</bibRefCitation>
. Four Brazilian specimens of
<taxonomicName id="C6F61480FF079E75FF5AFC42FED7D1BD" box="[151,284,980,1004]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="3382B311FF079E75FF5AFC42FED7D1BD" box="[151,284,980,1004]" italics="true" pageId="9" pageNumber="110">A. australis</emphasis>
</taxonomicName>
, viz. the male
<typeStatus id="DE4DD1A1FF079E75FE02FC42FDFAD1BC" box="[463,561,980,1005]" pageId="9" pageNumber="110" type="holotype">holotype</typeStatus>
from Bahia (MZUSP 10584), the male
<typeStatus id="DE4DD1A1FF079E75FBC7FC43FBA0D1BC" box="[1034,1131,981,1005]" pageId="9" pageNumber="110" type="paratype">paratype</typeStatus>
from São Paulo (MZUSP 11105), a male from Ceará (MZUSP 32611), and an ovigerous female from Bahia (MZUSP 32131), were also examined. Each of them was found to have one to three fixed teeth and one spiniform seta on the uropodal exopod, which suggests that the description “rami without spines” and the illustration of the uropod of
<taxonomicName id="C6F61480FF079E75FB3BFBD6FD58D62D" authority="in Rodrigues &amp; Shimizu" authorityName="in Rodrigues &amp; Shimizu" authorityYear="1992" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="3382B311FF079E75FB3BFBD6FAB0D609" box="[1270,1403,1088,1112]" italics="true" pageId="9" pageNumber="110">A. australis</emphasis>
in
<bibRefCitation id="656712F2FF079E75FF5AFBF2FD58D62D" author="Rodrigues" box="[151,659,1124,1149]" pageId="9" pageNumber="110" refString="Rodrigues, S. A. &amp; Shimizu, R. M. (1992) Description of a new Axianassa (Crustacea: Decapoda: Thalassinidea) from Brazil, and its first larval stage. Proceedings of the Biological Society of Washington, 105, 317 - 323. http: // biodiversitylibrary. org / page / 35607490" type="journal article" year="1992" yearSuffix="p">Rodrigues &amp; Shimizu (1992: p. 319, fig. 20)</bibRefCitation>
</taxonomicName>
are incorrect. Thus, both species have at least one fixed tooth and a small spiniform seta on the distolateral margin of the uropodal exopod.
</paragraph>
<paragraph id="01496F03FF079E75FF0AFB3AFAD2D7CC" blockId="9.[151,1437,151,1977]" pageId="9" pageNumber="110">
The antennal acicle was illustrated as relatively short, not reaching 0.25 length of the fourth article, in the original description of
<taxonomicName id="C6F61480FF079E75FE51FB46FDD4D6B9" box="[412,543,1232,1256]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="3382B311FF079E75FE51FB46FDD4D6B9" box="[412,543,1232,1256]" italics="true" pageId="9" pageNumber="110">A. australis</emphasis>
</taxonomicName>
by
<bibRefCitation id="656712F2FF079E75FD80FB59FC23D6B9" author="Rodrigues" box="[589,1000,1231,1256]" pageId="9" pageNumber="110" refString="Rodrigues, S. A. &amp; Shimizu, R. M. (1992) Description of a new Axianassa (Crustacea: Decapoda: Thalassinidea) from Brazil, and its first larval stage. Proceedings of the Biological Society of Washington, 105, 317 - 323. http: // biodiversitylibrary. org / page / 35607490" type="journal article" year="1992">Rodrigues &amp; Shimizu (1992: fig. 3)</bibRefCitation>
. Our examination of the
<typeStatus id="DE4DD1A1FF079E75FAC4FB59FAA0D6B9" box="[1289,1387,1231,1256]" pageId="9" pageNumber="110" type="holotype">holotype</typeStatus>
and two other Brazilian specimens of
<taxonomicName id="C6F61480FF079E75FDDEFB62FD5FD75D" box="[531,660,1268,1292]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="3382B311FF079E75FDDEFB62FD5FD75D" box="[531,660,1268,1292]" italics="true" pageId="9" pageNumber="110">A. australis</emphasis>
</taxonomicName>
shows that the acicle tip indeed reaches at most to about 0.25 length of the fourth article, thus being somewhat shorter than the acicle of
<taxonomicName id="C6F61480FF079E75FC5EFA8EFB8AD761" box="[915,1089,1304,1328]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="darrylfelderi">
<emphasis id="3382B311FF079E75FC5EFA8EFB8AD761" box="[915,1089,1304,1328]" italics="true" pageId="9" pageNumber="110">A. darrylfelderi</emphasis>
</taxonomicName>
, which is closer to 0.35 length of the fourth article (
<bibRefCitation id="656712F2FF079E75FEB0FAAAFD4ED704" author="Anker" box="[381,645,1340,1365]" pageId="9" pageNumber="110" refString="Anker, A. &amp; Lazarus, J. F. (2015) Description of two new associated infaunal decapod crustaceans (Axianassidae and Alpheidae) from the tropical eastern Pacific. Papeis Avulsos de Zoologia, 55, 115 - 129. http: // dx. doi. org / 10.1590 / 0031 - 1049.2015.55.08" type="journal article" year="2015">Anker &amp; Lazarus 2015</bibRefCitation>
). In contrast, the armature on the cutting edge of the pollex of the male minor cheliped does not seem to be a reliable character to separate
<taxonomicName id="C6F61480FF079E75FC40FAF6FBF0D729" box="[909,1083,1376,1400]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="darrylfelderi">
<emphasis id="3382B311FF079E75FC40FAF6FBF0D729" box="[909,1083,1376,1400]" italics="true" pageId="9" pageNumber="110">A. darrylfelderi</emphasis>
</taxonomicName>
from
<taxonomicName id="C6F61480FF079E75FB4CFAF6FAC8D729" box="[1153,1283,1376,1400]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="3382B311FF079E75FB4CFAF6FAC8D729" box="[1153,1283,1376,1400]" italics="true" pageId="9" pageNumber="110">A. australis</emphasis>
</taxonomicName>
, because two strong double teeth are present at least in one Brazilian specimen of the latter species (MZUSP 32611).
</paragraph>
<paragraph id="01496F03FF079E75FF0AFA31FE3BD5E9" blockId="9.[151,1437,151,1977]" pageId="9" pageNumber="110">
During our search for other morphological characters distinguishing
<taxonomicName id="C6F61480FF079E75FC2AFA3EFB52D791" box="[999,1177,1448,1472]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="darrylfelderi">
<emphasis id="3382B311FF079E75FC2AFA3EFB52D791" box="[999,1177,1448,1472]" italics="true" pageId="9" pageNumber="110">A. darrylfelderi</emphasis>
</taxonomicName>
from
<taxonomicName id="C6F61480FF079E75FB29FA3EFAA2D791" box="[1252,1385,1448,1472]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="3382B311FF079E75FB29FA3EFAA2D791" box="[1252,1385,1448,1472]" italics="true" pageId="9" pageNumber="110">A. australis</emphasis>
</taxonomicName>
, we noted that the former species is variable in the number of teeth on the dorsomesial surface of the third maxilliped coxa. Three specimens, viz. the ovigerous female from
<collectingCountry id="79E12F93FF079E75FD32FA67FC93D459" box="[767,856,1521,1544]" name="Panama" pageId="9" pageNumber="110">Panama</collectingCountry>
(MZUSP 34489), as well as the
<typeStatus id="DE4DD1A1FF079E75FB0FFA79FAEFD459" box="[1218,1316,1519,1544]" pageId="9" pageNumber="110" type="holotype">holotype</typeStatus>
(cf.
<bibRefCitation id="656712F2FF079E75FA9BFA79FE9BD47C" author="Anker" pageId="9" pageNumber="110" refString="Anker, A. &amp; Lazarus, J. F. (2015) Description of two new associated infaunal decapod crustaceans (Axianassidae and Alpheidae) from the tropical eastern Pacific. Papeis Avulsos de Zoologia, 55, 115 - 129. http: // dx. doi. org / 10.1590 / 0031 - 1049.2015.55.08" type="journal article" year="2015">Anker &amp; Lazarus 2015</bibRefCitation>
: fig. 1H, I) and
<typeStatus id="DE4DD1A1FF079E75FDC9F983FDAFD47C" box="[516,612,1557,1581]" pageId="9" pageNumber="110" type="paratype">paratype</typeStatus>
(MZUSP 33014) from
<collectingCountry id="79E12F93FF079E75FCA3F982FC16D47C" box="[878,989,1556,1581]" name="Colombia" pageId="9" pageNumber="110">Colombia</collectingCountry>
have two small adjacent teeth, whereas one male from
<collectingCountry id="79E12F93FF079E75FE84F9AFFE69D401" box="[329,418,1593,1616]" name="Panama" pageId="9" pageNumber="110">Panama</collectingCountry>
(MZUSP 34081) has only one larger tooth, as in
<taxonomicName id="C6F61480FF079E75FC28F9AEFBA2D401" box="[997,1129,1592,1616]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="3382B311FF079E75FC28F9AEFBA2D401" box="[997,1129,1592,1616]" italics="true" pageId="9" pageNumber="110">A. australis</emphasis>
</taxonomicName>
(cf.
<bibRefCitation id="656712F2FF079E75FB6EF9A1FF18D425" author="Rodrigues" pageId="9" pageNumber="110" refString="Rodrigues, S. A. &amp; Shimizu, R. M. (1992) Description of a new Axianassa (Crustacea: Decapoda: Thalassinidea) from Brazil, and its first larval stage. Proceedings of the Biological Society of Washington, 105, 317 - 323. http: // biodiversitylibrary. org / page / 35607490" type="journal article" year="1992">Rodrigues &amp; Shimizu 1992</bibRefCitation>
: fig. 9). In males of
<taxonomicName id="C6F61480FF079E75FE02F9CAFD49D425" box="[463,642,1628,1652]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="darrylfelderi">
<emphasis id="3382B311FF079E75FE02F9CAFD49D425" box="[463,642,1628,1652]" italics="true" pageId="9" pageNumber="110">A. darrylfelderi</emphasis>
</taxonomicName>
, the lateral surface of the third to fifth pleura is covered by a fine pubescence composed of numerous long setae (
<bibRefCitation id="656712F2FF079E75FD0DF9E9FBFCD4C9" author="Anker" box="[704,1079,1663,1688]" pageId="9" pageNumber="110" refString="Anker, A. &amp; Lazarus, J. F. (2015) Description of two new associated infaunal decapod crustaceans (Axianassidae and Alpheidae) from the tropical eastern Pacific. Papeis Avulsos de Zoologia, 55, 115 - 129. http: // dx. doi. org / 10.1590 / 0031 - 1049.2015.55.08" type="journal article" year="2015">Anker &amp; Lazarus 2015: fig. 4A</bibRefCitation>
). In males of
<taxonomicName id="C6F61480FF079E75FB10F916FAA9D4C9" box="[1245,1378,1664,1688]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="3382B311FF079E75FB10F916FAA9D4C9" box="[1245,1378,1664,1688]" italics="true" pageId="9" pageNumber="110">A. australis</emphasis>
</taxonomicName>
, this pubescence appears to be much denser, especially on the fourth and fifth pleura (
<bibRefCitation id="656712F2FF079E75FBE0F932FF6FD4B1" author="Rodrigues" pageId="9" pageNumber="110" refString="Rodrigues, S. A. &amp; Shimizu, R. M. (1992) Description of a new Axianassa (Crustacea: Decapoda: Thalassinidea) from Brazil, and its first larval stage. Proceedings of the Biological Society of Washington, 105, 317 - 323. http: // biodiversitylibrary. org / page / 35607490" type="journal article" year="1992">Rodrigues &amp; Shimizu 1992: fig. 1</bibRefCitation>
), sometimes obscuring the pleural margins. The palm-finger height-length ratio of the male major cheliped appears to be higher in
<taxonomicName id="C6F61480FF079E75FE66F97AFD97D555" box="[427,604,1772,1796]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="darrylfelderi">
<emphasis id="3382B311FF079E75FE66F97AFD97D555" box="[427,604,1772,1796]" italics="true" pageId="9" pageNumber="110">A. darrylfelderi</emphasis>
</taxonomicName>
than in
<taxonomicName id="C6F61480FF079E75FD72F97AFC88D555" box="[703,835,1772,1796]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="3382B311FF079E75FD72F97AFC88D555" box="[703,835,1772,1796]" italics="true" pageId="9" pageNumber="110">A. australis</emphasis>
</taxonomicName>
, but this may only be true for very large males (cf.
<bibRefCitation id="656712F2FF079E75FF5AF899FE56D579" author="Anker" box="[151,413,1807,1832]" pageId="9" pageNumber="110" refString="Anker, A. &amp; Lazarus, J. F. (2015) Description of two new associated infaunal decapod crustaceans (Axianassidae and Alpheidae) from the tropical eastern Pacific. Papeis Avulsos de Zoologia, 55, 115 - 129. http: // dx. doi. org / 10.1590 / 0031 - 1049.2015.55.08" type="journal article" year="2015">Anker &amp; Lazarus 2015</bibRefCitation>
: fig. 2A, B;
<bibRefCitation id="656712F2FF079E75FDEAF899FC94D579" author="Rodrigues" box="[551,863,1807,1832]" pageId="9" pageNumber="110" refString="Rodrigues, S. A. &amp; Shimizu, R. M. (1992) Description of a new Axianassa (Crustacea: Decapoda: Thalassinidea) from Brazil, and its first larval stage. Proceedings of the Biological Society of Washington, 105, 317 - 323. http: // biodiversitylibrary. org / page / 35607490" type="journal article" year="1992">Rodrigues &amp; Shimizu 1992</bibRefCitation>
: fig. 12). In summary, the extent of variation of the aforementioned characters cannot be evaluated based on the currently available material. Nevertheless three of them are tentatively included in the key below as morphological differences between these two geographically separated cryptic taxa. Molecular analyses, e.g., of the barcoding segment of COI, are highly desirable to confirm the validity of
<taxonomicName id="C6F61480FF079E75FEF6F836FE22D5E9" box="[315,489,1952,1976]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="darrylfelderi">
<emphasis id="3382B311FF079E75FEF6F836FE22D5E9" box="[315,489,1952,1976]" italics="true" pageId="9" pageNumber="110">A. darrylfelderi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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