248 lines
22 KiB
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248 lines
22 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.3790166" ID-GBIF-Dataset="ca35c50f-b0c5-4032-b06d-f71b00710c8d" ID-GBIF-Taxon="163751298" ID-ISSN="1313–2970" ID-Zenodo-Dep="3790166" ID-ZooBank="C26E1A82-0DD4-48EF-865C-9D8AA788B739" approvalRequired="188" approvalRequired_for_taxonomicNames="59" approvalRequired_for_textStreams="129" checkinTime="1587990637238" checkinUser="plazi" docAuthor="Crabo, Lars & Lafontaine, Donald" docDate="2009" docId="75513F417B30FF94FF02E95390D9FDD9" docLanguage="en" docName="ZK_article_2110.pdf" docOrigin="ZooKeys 30 (30)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@648ded35" docStyleName="zookeys.2008.journal_article" docTitle="Lasionycta phoca Crabo & Lafontaine, 2009, sp. n." docType="treatment" docVersion="5" lastPageId="66" lastPageNumber="66" masterDocId="896847397B71FFD6FF8EE829912DFFF8" masterDocTitle="A Revision of Lasionycta Aurivillius (Lepidoptera: Noctuidae) for North America and notes on Eurasian species, with descriptions of 17 new species, 6 new subspecies, a new genus, and two new species of Tricholita Grote" masterLastPageNumber="156" masterPageNumber="1" pageId="65" pageNumber="66" updateTime="1643569142744" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A Revision of Lasionycta Aurivillius (Lepidoptera: Noctuidae) for North America and notes on Eurasian species, with descriptions of 17 new species, 6 new subspecies, a new genus, and two new species of Tricholita Grote</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Crabo, Lars</mods:namePart>
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<mods:affiliation>Washington State University, Bellingham, United States of America</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lafontaine, Donald</mods:namePart>
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<mods:affiliation>Agriculture and Agri-Food Canada, Ottawa, Canada</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2009</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2009-12-18</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>30</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>30</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>156</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.3897/zookeys.30.308</mods:identifier>
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<mods:identifier type="GBIF-Dataset">ca35c50f-b0c5-4032-b06d-f71b00710c8d</mods:identifier>
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<mods:identifier type="ISSN">1313–2970</mods:identifier>
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<mods:identifier type="Zenodo-Dep">576576</mods:identifier>
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<mods:identifier type="ZooBank">C26E1A82-0DD4-48EF-865C-9D8AA788B739</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3790166" ID-GBIF-Taxon="163751298" ID-Zenodo-Dep="3790166" LSID="urn:lsid:plazi:treatment:75513F417B30FF94FF02E95390D9FDD9" httpUri="http://treatment.plazi.org/id/75513F417B30FF94FF02E95390D9FDD9" lastPageId="66" lastPageNumber="66" pageId="65" pageNumber="66">
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<subSubSection pageId="65" pageNumber="66" type="nomenclature">
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<paragraph blockId="65.[140,474,377,404]" box="[140,474,377,404]" pageId="65" pageNumber="66">
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<heading box="[140,474,377,404]" fontSize="11" level="2" pageId="65" pageNumber="66" reason="3">
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<emphasis bold="true" box="[140,474,377,404]" pageId="65" pageNumber="66">
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<taxonomicName box="[140,345,377,404]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="phoca">
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<emphasis bold="true" box="[140,345,377,404]" italics="true" pageId="65" pageNumber="66">Lasionycta phoca</emphasis>
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</taxonomicName>
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sub-group
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</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="65.[140,1108,448,1672]" pageId="65" pageNumber="66">
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The
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<taxonomicName box="[190,282,448,475]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="phoca">
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<emphasis box="[190,206,449,474]" italics="true" pageId="65" pageNumber="66">L</emphasis>
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.
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<emphasis box="[220,282,448,474]" italics="true" pageId="65" pageNumber="66">phoca</emphasis>
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</taxonomicName>
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sub-group contains seven mostly medium-size (expanse
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<quantity box="[919,1047,448,475]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" metricValueMax="3.6" metricValueMin="3.0" pageId="65" pageNumber="66" unit="mm" value="33.0" valueMax="36.0" valueMin="30.0">30–36 mm</quantity>
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) species from alpine or subarctic habitats. Most occur in western North America, with two in the Northeast. Th e forewing is gray to brown gray with dark basal, antemedial, and postmedial lines; the orbicular and reniform spots are weakly defined, evident mostly due to pale filling, and the claviform spot is absent; in several species the forewing is mottled with white, blue gray, or yellow. Th e dorsal hindwing is typically dark with a pale fringe. Th e ventral hindwing, often useful for diagnosis, is white to pale gray (rarely brownish gray) with dark chevron- or arrowhead-shaped discal spot, postmedial line, and marginal band. A small species from the Sierra
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<collectingRegion box="[785,870,730,756]" country="United States of America" name="Nevada" pageId="65" pageNumber="66">Nevada</collectingRegion>
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(
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<taxonomicName authority="Crabo & Lafontaine, 2009" authorityName="Crabo & Lafontaine" authorityYear="2009" box="[887,1068,731,756]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="mono" status="sp. n.">
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<emphasis box="[887,1068,731,756]" italics="true" pageId="65" pageNumber="66">Lasionycta mono</emphasis>
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</taxonomicName>
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<taxonomicNameLabel pageId="65" pageNumber="66" rank="species">sp. n.</taxonomicNameLabel>
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) with a hindwing resembling species in the
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<taxonomicName box="[681,834,765,792]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="staudingeri">
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<emphasis box="[681,697,766,791]" italics="true" pageId="65" pageNumber="66">L</emphasis>
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.
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<emphasis box="[714,834,765,791]" italics="true" pageId="65" pageNumber="66">staudingeri</emphasis>
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</taxonomicName>
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sub-group is associated with this sub-group by the shape of the valve.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66" type="description">
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<paragraph blockId="65.[140,1108,448,1672]" pageId="65" pageNumber="66">
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The male valve is elongate with a weak to moderate narrowing at the base of the cucullus. The cucullus is fairly stout and similar to those of the
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<taxonomicName baseAuthorityName="Staudinger" baseAuthorityYear="1857" box="[850,985,871,897]" class="Insecta" family="Noctuidae" genus="Lasiestra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="leucocycla">
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<emphasis box="[850,866,871,896]" italics="true" pageId="65" pageNumber="66">L</emphasis>
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.
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<emphasis box="[881,985,871,897]" italics="true" pageId="65" pageNumber="66">leucocycla</emphasis>
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</taxonomicName>
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sub-group in shape. The corona is variable, simple with partial double row near the apex in most species, but comprised of several rows in two species. Th e digitus is cylindrical. The female genitalia are similar to those of the
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<taxonomicName baseAuthorityName="Staudinger" baseAuthorityYear="1857" box="[626,761,976,1003]" class="Insecta" family="Noctuidae" genus="Lasiestra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="leucocycla">
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<emphasis box="[626,642,977,1002]" italics="true" pageId="65" pageNumber="66">L</emphasis>
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.
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<emphasis box="[657,761,976,1002]" italics="true" pageId="65" pageNumber="66">leucocycla</emphasis>
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</taxonomicName>
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sub-group. Th e corpus bursae is ovoid with a 50 % medial constriction and an appendix bursae of similar size to the corpus bursae. The male antennae are biserrate with triangular individual segments 1.5–2.1× as wide as the central shaft.
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</paragraph>
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<paragraph blockId="65.[140,1108,448,1672]" pageId="65" pageNumber="66">
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Species in this sub-group are among the most diffi cult to identify in the genus. The genitalia are only helpful in a few of the species and the male antennae are generally indistinguishable. Most species resemble each other and several are variable to the point where individuals approach other species in appearance. For this reason, a definitive diagnosis is easiest when series can be examined. Identification is simplified by narrowing the possibilities by locality since no more than three species occur in a region (see Table 1). Th e habitus, especially the ventral hindwing pattern, can then be used for definitive diagnosis. Species status was determined by lack of evidence of intergradation in areas of sympatry together with genital and DNA differences in most cases. Assessment of species limits was most difficult in
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<taxonomicName box="[769,904,1434,1460]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="uniformis">
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<emphasis box="[769,785,1435,1460]" italics="true" pageId="65" pageNumber="66">L</emphasis>
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.
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<emphasis box="[800,904,1434,1460]" italics="true" pageId="65" pageNumber="66">uniformis</emphasis>
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</taxonomicName>
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, a variable widely distributed species with many local forms and several CO1 haplotypes. The rationale for treating it as one species are discussed under
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<taxonomicName box="[685,819,1504,1531]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="uniformis">
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<emphasis box="[685,701,1505,1530]" italics="true" pageId="65" pageNumber="66">L</emphasis>
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.
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<emphasis box="[715,819,1504,1530]" italics="true" pageId="65" pageNumber="66">uniformis</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph blockId="65.[140,1108,448,1672]" lastBlockId="66.[140,1108,167,545]" lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66">
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The CO1 DNA sequences of all
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<taxonomicName box="[546,608,1539,1565]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="phoca">
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<emphasis box="[546,608,1539,1565]" italics="true" pageId="65" pageNumber="66">phoca</emphasis>
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</taxonomicName>
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sub-group species except
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<taxonomicName authorityName="Crabo & Lafontaine" authorityYear="2009" box="[895,986,1540,1566]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="caesia">
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<emphasis box="[895,911,1540,1565]" italics="true" pageId="65" pageNumber="66">L</emphasis>
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.
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<emphasis box="[924,986,1540,1565]" italics="true" pageId="65" pageNumber="66">caesia</emphasis>
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</taxonomicName>
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are similar (that of
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<taxonomicName authorityName="Crabo & Lafontaine" authorityYear="2009" box="[234,326,1575,1601]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="mono">
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<emphasis box="[234,250,1575,1600]" italics="true" pageId="65" pageNumber="66">L</emphasis>
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.
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<emphasis box="[266,326,1575,1600]" italics="true" pageId="65" pageNumber="66">mono</emphasis>
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</taxonomicName>
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is unknown), clustering with those of
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<taxonomicName box="[771,908,1575,1601]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="lagganata">
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<emphasis box="[771,908,1575,1601]" italics="true" pageId="65" pageNumber="66">L. lagganata</emphasis>
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</taxonomicName>
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,
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<taxonomicName box="[924,1100,1575,1601]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="quadrilunata">
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<emphasis box="[924,940,1575,1600]" italics="true" pageId="65" pageNumber="66">L</emphasis>
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.
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<emphasis box="[956,1100,1575,1601]" italics="true" pageId="65" pageNumber="66">quadrilunata</emphasis>
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</taxonomicName>
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, and
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<taxonomicName authorityName="Crabo & Lafontaine" authorityYear="2009" baseAuthorityName="Barnes & Benjamin" baseAuthorityYear="1923" box="[192,292,1610,1636]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="dolosa">
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<emphasis box="[192,208,1611,1636]" italics="true" pageId="65" pageNumber="66">L</emphasis>
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.
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<emphasis box="[226,292,1610,1636]" italics="true" pageId="65" pageNumber="66">dolosa</emphasis>
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</taxonomicName>
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(
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<figureCitation box="[309,411,1610,1636]" captionStart="Figure 248" captionStartId="142.[140,212,1558,1580]" captionTargetBox="[192,1058,178,1498]" captionTargetId="figure@142.[177,1071,165,1539]" captionTargetPageId="142" captionText="Figure 248. Neighbor-joining CO1 tree of the North American Lasionycta leucocycla species-group. Symbols and codes are the same as in Fig. 247. No data is available for Lasionycta carolynae, L. illima, L. leucocycla hampa, L. macleani, L. mono, or L. uniformis handfieldi. The illustrated specimens demonstrate the variation in appearance of the species-group and are scaled to the same size." figureDoi="http://doi.org/10.5281/zenodo.3769247" httpUri="https://zenodo.org/record/3769247/files/figure.png" pageId="65" pageNumber="66">Fig. 248</figureCitation>
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). Th e sequences of
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<taxonomicName authorityName="Crabo & Lafontaine" authorityYear="2009" box="[641,736,1610,1636]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="caesia">
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<emphasis box="[641,657,1611,1636]" italics="true" pageId="65" pageNumber="66">L</emphasis>
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.
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<emphasis box="[674,736,1611,1636]" italics="true" pageId="65" pageNumber="66">caesia</emphasis>
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</taxonomicName>
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,
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<taxonomicName authorityName="Crabo & Lafontaine" authorityYear="2009" box="[754,878,1610,1636]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="brunnea">
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<emphasis box="[754,770,1611,1636]" italics="true" pageId="65" pageNumber="66">L</emphasis>
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.
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<emphasis box="[787,878,1610,1636]" italics="true" pageId="65" pageNumber="66">brunnea</emphasis>
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</taxonomicName>
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,
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<taxonomicName authority="Crabo & Lafontaine, 2009" authorityName="Crabo & Lafontaine" authorityYear="2009" box="[895,993,1610,1636]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="gelida" status="sp. n.">
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<emphasis box="[895,911,1611,1636]" italics="true" pageId="65" pageNumber="66">L</emphasis>
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.
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<emphasis box="[929,993,1610,1636]" italics="true" pageId="65" pageNumber="66">gelida</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[1003,1069,1610,1636]" pageId="65" pageNumber="66" rank="species">sp. n.</taxonomicNameLabel>
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,
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<taxonomicName authority="(Smith)" baseAuthorityName="Smith" class="Insecta" family="Noctuidae" genus="Scotogramma" kingdom="Animalia" order="Lepidoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="discolor">
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<emphasis box="[1084,1100,1611,1636]" italics="true" pageId="65" pageNumber="66">L</emphasis>
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.
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<emphasis box="[140,221,1645,1671]" italics="true" pageId="65" pageNumber="66">discolor</emphasis>
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(Smith)
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</taxonomicName>
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, and
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<taxonomicName box="[381,475,1645,1672]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="phoca">
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<emphasis box="[381,397,1646,1671]" italics="true" pageId="65" pageNumber="66">L</emphasis>
|
||
.
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<emphasis box="[413,475,1645,1671]" italics="true" pageId="65" pageNumber="66">phoca</emphasis>
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</taxonomicName>
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are fairly uniform, although this could be an artifact of small sample sizes. In contrast, many haplotypes exist for several
|
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<taxonomicName box="[879,1014,167,193]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="uniformis">
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<emphasis box="[879,895,167,192]" italics="true" pageId="66" pageNumber="67">L</emphasis>
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||
.
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<emphasis box="[910,1014,167,193]" italics="true" pageId="66" pageNumber="67">uniformis</emphasis>
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</taxonomicName>
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||
subspecies which are intermixed with those of the other species in the DNA tree (
|
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<figureCitation box="[995,1095,202,228]" captionStart="Figure 248" captionStartId="142.[140,212,1558,1580]" captionTargetBox="[192,1058,178,1498]" captionTargetId="figure@142.[177,1071,165,1539]" captionTargetPageId="142" captionText="Figure 248. Neighbor-joining CO1 tree of the North American Lasionycta leucocycla species-group. Symbols and codes are the same as in Fig. 247. No data is available for Lasionycta carolynae, L. illima, L. leucocycla hampa, L. macleani, L. mono, or L. uniformis handfieldi. The illustrated specimens demonstrate the variation in appearance of the species-group and are scaled to the same size." figureDoi="http://doi.org/10.5281/zenodo.3769247" httpUri="https://zenodo.org/record/3769247/files/figure.png" pageId="66" pageNumber="67">Fig. 248</figureCitation>
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||
). Despite this overlap, CO1 distance analysis was useful for determining the number of taxa in a region. For example, comparison of all
|
||
<taxonomicName box="[723,816,272,299]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="phoca">
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<emphasis box="[723,739,273,298]" italics="true" pageId="66" pageNumber="67">L</emphasis>
|
||
.
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<emphasis box="[754,816,272,298]" italics="true" pageId="66" pageNumber="67">phoca</emphasis>
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</taxonomicName>
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sub-group DNA samples from the Canadian Rocky Mountains identifies two dominant haplotype clusters that correlate with
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<emphasis box="[301,317,343,368]" italics="true" pageId="66" pageNumber="67">L</emphasis>
|
||
.
|
||
<emphasis box="[331,346,343,368]" italics="true" pageId="66" pageNumber="67">u</emphasis>
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.
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<taxonomicName box="[360,464,343,369]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="uniformis">
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<emphasis box="[360,464,343,369]" italics="true" pageId="66" pageNumber="67">uniformis</emphasis>
|
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</taxonomicName>
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and
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<taxonomicName authorityName="Crabo & Lafontaine" authorityYear="2009" box="[520,641,343,369]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="brunnea">
|
||
<emphasis box="[520,536,343,368]" italics="true" pageId="66" pageNumber="67">L</emphasis>
|
||
.
|
||
<emphasis box="[550,641,343,369]" italics="true" pageId="66" pageNumber="67">brunnea</emphasis>
|
||
</taxonomicName>
|
||
. Similarly, two main haplotypes from the central
|
||
<collectingCountry box="[224,376,378,404]" name="United States of America" pageId="66" pageNumber="67">United States</collectingCountry>
|
||
Rocky Mountains correspond to
|
||
<taxonomicName box="[757,869,378,404]" class="Insecta" family="Noctuidae" genus="Scotogramma" kingdom="Animalia" order="Lepidoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="discolor">
|
||
<emphasis box="[757,773,379,404]" italics="true" pageId="66" pageNumber="67">L</emphasis>
|
||
.
|
||
<emphasis box="[788,869,378,404]" italics="true" pageId="66" pageNumber="67">discolor</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="subSpecies" species="uniformis" subSpecies="fusca">
|
||
<emphasis box="[925,941,379,404]" italics="true" pageId="66" pageNumber="67">L</emphasis>
|
||
.
|
||
<emphasis italics="true" pageId="66" pageNumber="67">uniformis fusca</emphasis>
|
||
</taxonomicName>
|
||
. However, more haplotypes than identifiable species exist in the Pacific Northwest. Two of the haplotypes correspond to
|
||
<taxonomicName authorityName="Crabo & Lafontaine" authorityYear="2009" box="[579,676,448,475]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="gelida">
|
||
<emphasis box="[579,595,449,474]" italics="true" pageId="66" pageNumber="67">L</emphasis>
|
||
.
|
||
<emphasis box="[612,676,448,474]" italics="true" pageId="66" pageNumber="67">gelida</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Crabo & Lafontaine" authorityYear="2009" box="[738,834,449,475]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="caesia">
|
||
<emphasis box="[738,754,449,474]" italics="true" pageId="66" pageNumber="67">L</emphasis>
|
||
.
|
||
<emphasis box="[772,834,449,474]" italics="true" pageId="66" pageNumber="67">caesia</emphasis>
|
||
</taxonomicName>
|
||
. Th e other three major clusters are similar but variable and cannot be sorted by appearance to more than one taxon in
|
||
<taxonomicName box="[240,494,519,545]" class="Insecta" family="Noctuidae" genus="Lasionycta" kingdom="Animalia" order="Lepidoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="subSpecies" species="uniformis" subSpecies="multicolor">
|
||
<emphasis box="[240,256,519,544]" italics="true" pageId="66" pageNumber="67">L</emphasis>
|
||
.
|
||
<emphasis box="[271,494,519,545]" italics="true" pageId="66" pageNumber="67">uniformis multicolor</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |