346 lines
50 KiB
XML
346 lines
50 KiB
XML
<document id="988B9AC248085026A04453083E8DD31D" ID-DOI="10.11646/zootaxa.3750.5.7" ID-GBIF-Dataset="d35e62c7-975b-4dff-892e-e8e5c926820e" ID-ISSN="1175-5326" ID-Zenodo-Dep="217852" ID-ZooBank="22C6D62A-D6CA-4BBE-AD0D-29174C2A2769" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460711129622" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2013" docId="7505414FFF927370FF29FC35FBE04426" docLanguage="en" docName="zt03750p568.pdf" docOrigin="Zootaxa 3750 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Alloraphes myrmecophilus Franz" docType="treatment" docVersion="6" lastPageNumber="567" masterDocId="893C3937FF837362FFBEFFC2FFB14304" masterDocTitle="Revision of the Neotropical genus Alloraphes Franz (Coleoptera: Staphylinidae: Scydmaeninae)" masterLastPageNumber="568" masterPageNumber="549" pageNumber="566" updateTime="1698357997849" updateUser="plazi">
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<mods:titleInfo id="7AC650093577D30728965FBC5414BC94">
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<mods:title id="9B00DF4EFF52FE7E29A1B8BF82613811">Revision of the Neotropical genus Alloraphes Franz (Coleoptera: Staphylinidae: Scydmaeninae)</mods:title>
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<mods:name id="4446D6C954ECD79421DBF85872010C2D" type="personal">
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<mods:roleTerm id="0CE586025DEA8E09611D426BDA38BC43">Author</mods:roleTerm>
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<mods:namePart id="E808592CE14BD21B40BD606965B6B608">Jałoszyński, Paweł</mods:namePart>
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<mods:typeOfResource id="5817942F20243642571F5C40745B5BDE">text</mods:typeOfResource>
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<mods:title id="4B635940E357CE0969B7D1518DFBDD98">Zootaxa</mods:title>
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<mods:part id="745DD4F7D67C78772DF96ECC2050827B">
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<mods:date id="93157F19818815D9FD9166E6F42B6DBD">2013</mods:date>
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<mods:detail id="5CA2669AB9C87EA8EB7D3B11C6AE28C8" type="volume">
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<mods:number id="B47F833ED7545A4A88EEEC620A52082D">3750</mods:number>
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<mods:number id="5D7F444B655DF8D0D3B05B6D86968431">5</mods:number>
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<mods:classification id="A334BE0A42A3D1A7F1F2B887A0F602E3">journal article</mods:classification>
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<mods:identifier id="25D85571D773CE3E40C1498508953387" type="DOI">10.11646/zootaxa.3750.5.7</mods:identifier>
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<mods:identifier id="98B647FAD3DE4416BC5FDA96EE688DDC" type="GBIF-Dataset">d35e62c7-975b-4dff-892e-e8e5c926820e</mods:identifier>
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<mods:identifier id="7366064927380B2A3695CEBE11C19E3B" type="ISSN">1175-5326</mods:identifier>
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<treatment id="7505414FFF927370FF29FC35FBE04426" ID-DOI="http://doi.org/10.5281/zenodo.6154537" ID-GBIF-Taxon="119585163" ID-Zenodo-Dep="6154537" LSID="urn:lsid:plazi:treatment:7505414FFF927370FF29FC35FBE04426" httpUri="http://treatment.plazi.org/id/7505414FFF927370FF29FC35FBE04426" lastPageId="18" lastPageNumber="567" pageId="17" pageNumber="566">
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<subSubSection id="B5B6A3D2FF927373FF29FC35FD7C477C" pageId="17" pageNumber="566" type="nomenclature">
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<paragraph id="FD13F059FF927373FF29FC35FD164716" blockId="17.[151,679,1015,1075]" box="[151,679,1015,1042]" pageId="17" pageNumber="566">
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<heading id="A65B4735FF927373FF29FC35FD164716" bold="true" box="[151,679,1015,1042]" fontSize="11" level="1" pageId="17" pageNumber="566" reason="1">
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<emphasis id="CFD82C4BFF927373FF29FC35FD164716" bold="true" box="[151,679,1015,1042]" pageId="17" pageNumber="566">
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<taxonomicName id="3AAC8BDAFF927373FF29FC35FD9F4716" ID-CoL="BYZG" authority="Franz" authorityName="Franz" box="[151,558,1015,1042]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="566" phylum="Arthropoda" rank="species" species="myrmecophilus">
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<emphasis id="CFD82C4BFF927373FF29FC35FE6D4715" bold="true" box="[151,476,1015,1041]" italics="true" pageId="17" pageNumber="566">Alloraphes myrmecophilus</emphasis>
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Franz
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</taxonomicName>
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stat. rest.
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="FD13F059FF927373FF29FBD8FD994737" blockId="17.[151,679,1015,1075]" box="[151,552,1050,1075]" pageId="17" pageNumber="566">
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(
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<figureCitation id="6597ECDCFF927373FF21FBD8FF4A4737" box="[159,251,1050,1075]" captionStart="FIGURES 1 – 12" captionStartId="3.[151,264,1879,1902]" captionTargetBox="[217,1363,203,1830]" captionTargetId="figure@3.[209,1368,195,1845]" captionTargetPageId="3" captionText="FIGURES 1 – 12. Dorsal (1 – 9, 11 – 12) and laterodorsal (10) habitus of Alloraphes Franz. Alloraphes jamaicae Franz, holotype male (1); A. chiapasensis Franz, holotype male (2); A. yucatani Franz, holotype male (3); A. lenkoi Franz, paratype male (4) and paratype female (5); A. peruanus Franz, holotype male (6) and paratype female (7; but see the Remarks section in the redescription); A. dentatus sp. n., holotype male (8); A. peckorum sp. n., holotype male (9 – 10); A. myrmecophilus Franz, stat. rest., paratype male (11);? Alloraphes magnus Franz, holotype female (12)." httpUri="https://zenodo.org/record/217853/files/figure.png" pageId="17" pageNumber="566">Figs. 11</figureCitation>
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,
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<figureCitation id="6597ECDCFF927373FEB9FBD8FE944737" box="[263,293,1050,1075]" captionStart="FIGURES 22 – 26" captionStartId="7.[151,264,1642,1665]" captionTargetBox="[222,1331,193,1600]" captionTargetId="figure@7.[200,1342,193,1608]" captionTargetPageId="7" captionText="FIGURES 22 – 26. Alloraphes jamaicae Franz, holotype male (22 – 24); A. myrmecophilus Franz stat. rest., paratype male (25);? Alloraphes magnus Franz, holotype female (26). Head in ventral view (22); prothorax in ventral view (23); pterothorax in ventral view (24); body venter (25); and mesothorax in ventral view (26). Abbreviations: aest 2, mesanepisternum; aest 3, metanepisternum; ai, asetose impression; amc, anterior marginal carina of mesocoxal cavity; ar, anterior ridge; bst, basisternal part of prosternum; bst, basistipes; cd, cardo; cx, coxa; el, elytron; epm 3, metepimeron; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; lss, lateral suture of submentum; mc, median carina of mesoventrite; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mesostipes; msvp, mesoventral intercoxal process; mtap, metaventral anterior process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; pl, posterior lobe of mesocoxal projection; pre, prepectus; ppf, palpifer; psp, prosternal process; submentum; tm, temple; v 3, metaventrite" httpUri="https://zenodo.org/record/217857/files/figure.png" pageId="17" pageNumber="566">25</figureCitation>
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,
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<figureCitation id="6597ECDCFF927373FE8FFBD9FECB4737" box="[305,378,1051,1075]" captionStart="FIGURES 27 – 43" captionStartId="8.[151,264,1654,1677]" captionTargetBox="[156,1426,184,1626]" captionTargetId="figure@8.[151,1436,167,1633]" captionTargetPageId="8" captionText="FIGURES 27 – 43. Aedeagus (27 – 34, 36 – 43) and female genitalia (35) of Alloraphes Franz in ventral (27, 29, 31, 33, 35, 36, 38, 40, 42) and lateral (28, 30, 32, 34, 37, 39, 41, 43) views. Alloraphes jamaicae Franz, holotype (27 – 28); A. chiapasensis Franz, holotype, parameres missing (29 – 30); A. yucatani Franz, holotype (31 – 32); A. lenkoi Franz, paratypes (33 – 35); A. peruanus Franz, holotype (36 – 37); A. dentatus sp. n., holotype (38 – 39); A. peckorum sp. n., holotype (40 – 41); A. myrmecophilus Franz, stat. rest., paratype (42 – 43). Abbreviations: ap, apical projection; bpa, basal pumping apparatus; ov, egg; sp, spermatheca." httpUri="https://zenodo.org/record/217858/files/figure.png" pageId="17" pageNumber="566">42–43</figureCitation>
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,
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<figureCitation id="6597ECDCFF927373FE3BFBD8FE7F4737" box="[389,462,1050,1075]" captionStart="FIGURES 44 – 60" captionStartId="10.[151,264,1803,1826]" captionTargetBox="[162,1411,295,1754]" captionTargetId="figure@10.[151,1436,283,1765]" captionTargetPageId="10" captionText="FIGURES 44 – 60. Aedeagus (44 – 51, 53 – 60) and female genitalia (52) of Alloraphes Franz in ventral (44, 46, 48, 50, 52, 53, 55, 57, 59) and lateral (45, 47, 49, 51, 54, 56, 58, 60) views. Alloraphes jamaicae Franz, holotype (44 – 45); A. chiapasensis Franz, holotype, paramere detached (46 – 47); A. yucatani Franz, holotype (48 – 49); A. lenkoi Franz, paratypes (50 – 52); A. peruanus Franz, holotype (53 – 54); A. dentatus sp. n., holotype (55 – 56); A. peckorum sp. n., holotype (57 – 58); A. myrmecophilus Franz, stat. rest., paratype (59 – 60). Abbreviations: ap, apical projection; bpa, basal pumping apparatus; ov, egg; sp, spermatheca." httpUri="https://zenodo.org/record/217859/files/figure.png" pageId="17" pageNumber="566">59–60</figureCitation>
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,
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<figureCitation id="6597ECDCFF927373FE67FBD8FE464737" box="[473,503,1050,1075]" captionStart="FIGURES 61 – 67" captionStartId="11.[151,264,1809,1832]" captionTargetBox="[161,1423,953,1744]" captionTargetId="figure@11.[151,1436,939,1757]" captionTargetPageId="11" captionText="FIGURES 61 – 67. Original sets of labels pinned under historical type specimens. Alloraphes jamaicae Franz, holotype (61); A. chiapasensis Franz, holotype (62); A. yucatani Franz, holotype (63); A. lenkoi Franz, paratype (64); A. peruanus Franz, holotype (65); A. myrmecophilus Franz, stat. rest., paratype (66);? Alloraphes magnus Franz, holotype (67)." httpUri="https://zenodo.org/record/217860/files/figure.png" pageId="17" pageNumber="566">66</figureCitation>
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,
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<figureCitation id="6597ECDCFF927373FDBDFBD8FD914737" box="[515,544,1050,1075]" captionStart="FIGURE 68" captionStartId="12.[151,250,1290,1312]" captionTargetBox="[151,1418,167,1268]" captionTargetId="figure@12.[151,1436,167,1268]" captionTargetPageId="12" captionText="FIGURE 68. Distribution of Alloraphes: a, A. jamaicae Franz; b, A. chiapasensis Franz; c, A. yucatani Franz; d, A. lenkoi Franz; e, A. peruanus Franz; f, A. dentatus sp. n.; g, A. peckorum sp. n.; h, A. myrmecophilus Franz stat. rest.; i, A. bolivarensis Franz; j,? Alloraphes magnus Franz." httpUri="https://zenodo.org/record/217861/files/figure.png" pageId="17" pageNumber="566">68</figureCitation>
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)
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</paragraph>
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<paragraph id="FD13F059FF927373FF29FBA0FD7C477C" blockId="17.[151,779,1122,1175]" box="[151,717,1122,1144]" pageId="17" pageNumber="566">
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<taxonomicName id="3AAC8BDAFF927373FF29FBA0FDD6477C" authority="Franz, 1980a: 214" authorityName="Franz" authorityPageNumber="214" authorityYear="1980" box="[151,615,1122,1144]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="566" phylum="Arthropoda" rank="species" species="myrmecophilus">
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<emphasis id="CFD82C4BFF927373FF29FBA0FE16477C" box="[151,423,1122,1144]" italics="true" pageId="17" pageNumber="566">Alloraphes myrmecophilus</emphasis>
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Franz, 1980a: 214
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</taxonomicName>
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, Fig. 201.
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</paragraph>
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</subSubSection>
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<subSubSection id="B5B6A3D2FF927373FF29FB43FCBA4793" box="[151,779,1153,1175]" pageId="17" pageNumber="566" type="description">
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<paragraph id="FD13F059FF927373FF29FB43FCBA4793" blockId="17.[151,779,1122,1175]" box="[151,779,1153,1175]" pageId="17" pageNumber="566">
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<taxonomicName id="3AAC8BDAFF927373FF29FB43FDFC4793" authority="Franz" authorityName="Franz" box="[151,589,1153,1175]" class="Insecta" family="Staphylinidae" genus="Parastenichnaphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="566" phylum="Arthropoda" rank="species" species="myrmecophilus">
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<emphasis id="CFD82C4BFF927373FF29FB43FE4B4793" box="[151,506,1153,1175]" italics="true" pageId="17" pageNumber="566">Parastenichnaphes myrmecophilus</emphasis>
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(Franz)
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</taxonomicName>
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; Franz, 1989: 278.
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</paragraph>
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</subSubSection>
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<subSubSection id="B5B6A3D2FF927373FF29FB06FC2746D3" pageId="17" pageNumber="566" type="materials_examined">
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<paragraph id="FD13F059FF927373FF29FB06FB464620" blockId="17.[151,1436,1220,2037]" pageId="17" pageNumber="566">
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<emphasis id="CFD82C4BFF927373FF29FB06FDE347D9" bold="true" box="[151,594,1220,1245]" pageId="17" pageNumber="566">
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<typeStatus id="22174EFBFF927373FF29FB06FF6347D9" box="[151,210,1220,1245]" pageId="17" pageNumber="566">Type</typeStatus>
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material.
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<typeStatus id="22174EFBFF927373FEF0FB06FE7747D9" box="[334,454,1220,1245]" pageId="17" pageNumber="566" type="paratype">Paratypes</typeStatus>
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(15 exx:
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<specimenCount id="EBAA3BD0FF927373FD88FB06FDE347D9" box="[566,594,1220,1245]" pageId="17" pageNumber="566" type="generic">12</specimenCount>
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</emphasis>
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♂,
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<emphasis id="CFD82C4BFF927373FDC1FB06FD3C47D9" bold="true" box="[639,653,1220,1245]" pageId="17" pageNumber="566">3</emphasis>
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♀
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<emphasis id="CFD82C4BFF927373FD15FB06FC1D47D9" bold="true" box="[683,940,1220,1245]" pageId="17" pageNumber="566">
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):
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<collectingCountry id="85BBB0C9FF927373FD78FB06FCBE47D9" box="[710,783,1220,1245]" name="Brazil" pageId="17" pageNumber="566">Brazil</collectingCountry>
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(São Paulo):
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</emphasis>
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all with two labels (
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<figureCitation id="6597ECDCFF927373FB26FB06FB5C47D9" box="[1176,1261,1220,1245]" captionStart="FIGURES 61 – 67" captionStartId="11.[151,264,1809,1832]" captionTargetBox="[161,1423,953,1744]" captionTargetId="figure@11.[151,1436,939,1757]" captionTargetPageId="11" captionText="FIGURES 61 – 67. Original sets of labels pinned under historical type specimens. Alloraphes jamaicae Franz, holotype (61); A. chiapasensis Franz, holotype (62); A. yucatani Franz, holotype (63); A. lenkoi Franz, paratype (64); A. peruanus Franz, holotype (65); A. myrmecophilus Franz, stat. rest., paratype (66);? Alloraphes magnus Franz, holotype (67)." httpUri="https://zenodo.org/record/217860/files/figure.png" pageId="17" pageNumber="566">Fig. 66</figureCitation>
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): "Barueri, Sao Paulo /
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<collectingCountry id="85BBB0C9FF927373FF50FB25FE854604" box="[238,308,1255,1280]" name="Brazil" pageId="17" pageNumber="566">Brasil</collectingCountry>
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, lg. Lenko" with numbers 22, 23 or 27 on reverse sides [white, printed; reverse handwritten in blue ink]; "
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<taxonomicName id="3AAC8BDAFF927373FF60FACEFEE84620" box="[222,345,1292,1316]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="566" phylum="Arthropoda" rank="genus">
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<emphasis id="CFD82C4BFF927373FF60FACEFEE84620" box="[222,345,1292,1316]" italics="true" pageId="17" pageNumber="566">Alloraphes</emphasis>
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</taxonomicName>
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/
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<taxonomicName id="3AAC8BDAFF927373FED1FACEFDAB4620" box="[367,538,1292,1316]" class="Insecta" family="Staphylinidae" genus="Parastenichnaphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="566" phylum="Arthropoda" rank="species" species="myrmecophilus">
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<emphasis id="CFD82C4BFF927373FED1FACEFDAB4620" box="[367,538,1292,1316]" italics="true" pageId="17" pageNumber="566">myrmecophilus</emphasis>
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</taxonomicName>
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m. /
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<typeStatus id="22174EFBFF927373FDEDFACEFD454620" box="[595,756,1292,1316]" pageId="17" pageNumber="566" type="paratype">PARATYPUS</typeStatus>
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" [yellow, handwritten and printed] (NHMW).
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</paragraph>
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<paragraph id="FD13F059FF927373FF79FAEDFC2746D3" blockId="17.[151,1436,1220,2037]" pageId="17" pageNumber="566">
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<emphasis id="CFD82C4BFF927373FF79FAEDFE13464C" bold="true" box="[199,418,1327,1352]" pageId="17" pageNumber="566">Revised diagnosis.</emphasis>
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Male and female: frons shallowly impressed between supraantennal tubercles and sharply demarcated from convex vertex by broadly V-shaped and narrow transverse groove between eyes; pronotum with nearly evenly arcuate ante-basal transverse groove indistinctly deepened in middle; elytra unmodified. Male: aedeagus in ventral view strikingly stout, with distinctly asymmetrical apex of median lobe, without apical projections; each slender paramere bearing single long and thin seta.
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</paragraph>
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</subSubSection>
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<subSubSection id="B5B6A3D2FF927370FF79FA26FBEC4138" lastPageId="18" lastPageNumber="567" pageId="17" pageNumber="566" type="description">
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<paragraph id="FD13F059FF927373FF79FA26FE6A451B" blockId="17.[151,1436,1220,2037]" pageId="17" pageNumber="566">
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<emphasis id="CFD82C4BFF927373FF79FA26FEC546F9" bold="true" box="[199,372,1508,1533]" pageId="17" pageNumber="566">Redescription.</emphasis>
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Body of male (
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<figureCitation id="6597ECDCFF927373FDFFFA26FD2946F8" box="[577,664,1508,1533]" captionStart="FIGURES 1 – 12" captionStartId="3.[151,264,1879,1902]" captionTargetBox="[217,1363,203,1830]" captionTargetId="figure@3.[209,1368,195,1845]" captionTargetPageId="3" captionText="FIGURES 1 – 12. Dorsal (1 – 9, 11 – 12) and laterodorsal (10) habitus of Alloraphes Franz. Alloraphes jamaicae Franz, holotype male (1); A. chiapasensis Franz, holotype male (2); A. yucatani Franz, holotype male (3); A. lenkoi Franz, paratype male (4) and paratype female (5); A. peruanus Franz, holotype male (6) and paratype female (7; but see the Remarks section in the redescription); A. dentatus sp. n., holotype male (8); A. peckorum sp. n., holotype male (9 – 10); A. myrmecophilus Franz, stat. rest., paratype male (11);? Alloraphes magnus Franz, holotype female (12)." httpUri="https://zenodo.org/record/217853/files/figure.png" pageId="17" pageNumber="566">Fig. 11</figureCitation>
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) moderately strongly convex, brown, covered with light brown vestiture; BL
|
||
<quantity id="3A545DBCFF927373FE8FF9C5FE66451B" box="[305,471,1543,1568]" metricMagnitude="-4" metricUnit="m" metricValue="9.299999999999999" metricValueMax="9.6" metricValueMin="9.0" pageId="17" pageNumber="566" unit="mm" value="0.9299999999999999" valueMax="0.96" valueMin="0.9">0.90–0.96 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="FD13F059FF927373FF79F9EEFED14444" blockId="17.[151,1436,1220,2037]" pageId="17" pageNumber="566">
|
||
Head broadest at large, moderately strongly convex and moderately coarsely faceted eyes, HL
|
||
<quantity id="3A545DBCFF927373FB46F9EEFA2D4540" box="[1272,1436,1580,1604]" metricMagnitude="-4" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="17" pageNumber="566" unit="mm" value="0.115" valueMax="0.13" valueMin="0.1">0.10–0.13 mm</quantity>
|
||
(mean
|
||
<quantity id="3A545DBCFF927373FF59F98DFEE2456C" box="[231,339,1615,1640]" metricMagnitude="-4" metricUnit="m" metricValue="6.8999999999999995" pageId="17" pageNumber="566" unit="mm" value="0.69">0.69 mm</quantity>
|
||
), HW 0.14–015 mm (mean
|
||
<quantity id="3A545DBCFF927373FD23F98DFCB8456C" box="[669,777,1615,1640]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="17" pageNumber="566" unit="mm" value="0.15">0.15 mm</quantity>
|
||
); tempora absent, eyes adjacent to occipital constriction; vertex distinctly convex, sharply demarcated from frons by narrow and long broadly V-shaped transverse groove (tip directed anteriorly); frons between weakly raised supra-antennal tubercles slightly impressed. Punctures on frons and vertex inconspicuous, fine; setae short, barely discernible. Antennae slender, AnL 25–
|
||
<quantity id="3A545DBCFF927373FB58F97EFAFC45D0" box="[1254,1357,1724,1749]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="17" pageNumber="566" unit="mm" value="0.28">0.28 mm</quantity>
|
||
(mean
|
||
<quantity id="3A545DBCFF927373FF29F922FF4F45FC" box="[151,254,1760,1784]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="17" pageNumber="566" unit="mm" value="0.27">0.27 mm</quantity>
|
||
); antennomere I about 1.5x as long as broad; II about twice as long as broad; III–V about as long as broad; VI–VII each slightly transverse; VIII–X each distinctly transverse; XI only about 1.3x as long as broad and about 1.8x as long as X.
|
||
</paragraph>
|
||
<paragraph id="FD13F059FF927370FF79F88FFAF943D1" blockId="17.[151,1436,1220,2037]" lastBlockId="18.[151,1436,151,1040]" lastPageId="18" lastPageNumber="567" pageId="17" pageNumber="566">
|
||
Pronotum about as long as broad, broadest in anterior third; PL
|
||
<quantity id="3A545DBCFF927373FC32F88EFB9E4460" box="[908,1071,1868,1893]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" metricValueMax="2.0" metricValueMin="1.6" pageId="17" pageNumber="566" unit="mm" value="0.18" valueMax="0.2" valueMin="0.16">0.16–0.20 mm</quantity>
|
||
(mean
|
||
<quantity id="3A545DBCFF927373FB3FF88EFB584460" box="[1153,1257,1868,1892]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="17" pageNumber="566" unit="mm" value="0.18">0.18 mm</quantity>
|
||
), PW
|
||
<quantity id="3A545DBCFF927373FA92F88EFF72448C" metricMagnitude="-4" metricUnit="m" metricValue="1.75" metricValueMax="1.9" metricValueMin="1.6" pageId="17" pageNumber="566" unit="mm" value="0.175" valueMax="0.19" valueMin="0.16">0.16–0.19 mm</quantity>
|
||
(mean
|
||
<quantity id="3A545DBCFF927373FE9FF8B2FE3F448C" box="[289,398,1904,1928]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="17" pageNumber="566" unit="mm" value="0.18">0.18 mm</quantity>
|
||
). Anterior margin and lateral margins in anterior half strongly rounded; sides distinctly narrowing caudad, pronotum distinctly constricted near posterior fourth; posterior margin slightly arcuate; antebasal transverse groove distinct and weakly, evenly arcuate, indistinctly deepened in middle; lateral impressions small and shallow, indistinct. Punctures on pronotal disc inconspicuous, fine and sparse; setae sparse, short and nearly recumbent, sides of pronotum covered sparsely with short and relatively thin bristles directed latero-caudad (indiscernible in dry-mounted specimens but visible in transparent mounts under a compound microscope).
|
||
</paragraph>
|
||
<paragraph id="FD13F059FF917370FF79FF1DFB494261" blockId="18.[151,1436,151,1040]" pageId="18" pageNumber="567">
|
||
Elytra more convex than pronotum, oval and unmodified, broadest between middle and anterior third; EL
|
||
<quantity id="3A545DBCFF917370FF29FEC6FE884218" box="[151,313,260,284]" metricMagnitude="-4" metricUnit="m" metricValue="4.05" metricValueMax="4.3" metricValueMin="3.8" pageId="18" pageNumber="567" unit="mm" value="0.405" valueMax="0.43" valueMin="0.38">0.38–0.43 mm</quantity>
|
||
(mean
|
||
<quantity id="3A545DBCFF917370FE32FEC6FE454218" box="[396,500,260,284]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="18" pageNumber="567" unit="mm" value="0.4">0.40 mm</quantity>
|
||
), EW
|
||
<quantity id="3A545DBCFF917370FD86FEC6FD6A4218" box="[568,731,260,284]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" metricValueMax="3.0" metricValueMin="2.8" pageId="18" pageNumber="567" unit="mm" value="0.29" valueMax="0.3" valueMin="0.28">0.28–0.30 mm</quantity>
|
||
(mean
|
||
<quantity id="3A545DBCFF917370FC93FEC6FC244218" box="[813,917,260,284]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="18" pageNumber="567" unit="mm" value="0.28">0.28 mm</quantity>
|
||
), EI 1.36–1.45 (mean 1.42); sub-humeral lines barely discernible at elytral bases, short and indistinct. Punctures on elytra only slightly more distinct than those on pronotum, small and shallow; setae short, sparse and nearly recumbent.
|
||
<collectingCountry id="85BBB0C9FF917370FC03FE8EFC444261" box="[957,1013,332,357]" name="India" pageId="18" pageNumber="567">Hind</collectingCountry>
|
||
wings well developed.
|
||
</paragraph>
|
||
<paragraph id="FD13F059FF917370FF79FEB2FD21428C" blockId="18.[151,1436,151,1040]" box="[199,656,367,392]" pageId="18" pageNumber="567">Legs long and slender; all tibiae straight.</paragraph>
|
||
<paragraph id="FD13F059FF917370FF79FE56FC4042F0" blockId="18.[151,1436,151,1040]" pageId="18" pageNumber="567">
|
||
Aedeagus (
|
||
<figureCitation id="6597ECDCFF917370FEF6FE56FE6542A8" box="[328,468,404,429]" captionStart="FIGURES 27 – 43" captionStartId="8.[151,264,1654,1677]" captionTargetBox="[156,1426,184,1626]" captionTargetId="figure@8.[151,1436,167,1633]" captionTargetPageId="8" captionText="FIGURES 27 – 43. Aedeagus (27 – 34, 36 – 43) and female genitalia (35) of Alloraphes Franz in ventral (27, 29, 31, 33, 35, 36, 38, 40, 42) and lateral (28, 30, 32, 34, 37, 39, 41, 43) views. Alloraphes jamaicae Franz, holotype (27 – 28); A. chiapasensis Franz, holotype, parameres missing (29 – 30); A. yucatani Franz, holotype (31 – 32); A. lenkoi Franz, paratypes (33 – 35); A. peruanus Franz, holotype (36 – 37); A. dentatus sp. n., holotype (38 – 39); A. peckorum sp. n., holotype (40 – 41); A. myrmecophilus Franz, stat. rest., paratype (42 – 43). Abbreviations: ap, apical projection; bpa, basal pumping apparatus; ov, egg; sp, spermatheca." httpUri="https://zenodo.org/record/217858/files/figure.png" pageId="18" pageNumber="567">Figs. 42–43</figureCitation>
|
||
,
|
||
<figureCitation id="6597ECDCFF917370FE5DFE56FD9B42A9" box="[483,554,404,429]" captionStart="FIGURES 44 – 60" captionStartId="10.[151,264,1803,1826]" captionTargetBox="[162,1411,295,1754]" captionTargetId="figure@10.[151,1436,283,1765]" captionTargetPageId="10" captionText="FIGURES 44 – 60. Aedeagus (44 – 51, 53 – 60) and female genitalia (52) of Alloraphes Franz in ventral (44, 46, 48, 50, 52, 53, 55, 57, 59) and lateral (45, 47, 49, 51, 54, 56, 58, 60) views. Alloraphes jamaicae Franz, holotype (44 – 45); A. chiapasensis Franz, holotype, paramere detached (46 – 47); A. yucatani Franz, holotype (48 – 49); A. lenkoi Franz, paratypes (50 – 52); A. peruanus Franz, holotype (53 – 54); A. dentatus sp. n., holotype (55 – 56); A. peckorum sp. n., holotype (57 – 58); A. myrmecophilus Franz, stat. rest., paratype (59 – 60). Abbreviations: ap, apical projection; bpa, basal pumping apparatus; ov, egg; sp, spermatheca." httpUri="https://zenodo.org/record/217859/files/figure.png" pageId="18" pageNumber="567">59–60</figureCitation>
|
||
) stout and lightly sclerotized; AeL
|
||
<quantity id="3A545DBCFF917370FC73FE56FB8942A8" box="[973,1080,404,428]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="18" pageNumber="567" unit="mm" value="0.13">0.13 mm</quantity>
|
||
; in ventral view apical part of median lobe distinctly asymmetrical and with deeply and irregularly emarginated apical margin; apical projections missing; parameres slender and each with only one thin and long apical seta.
|
||
</paragraph>
|
||
<paragraph id="FD13F059FF917370FF79FE3DFBEC4138" blockId="18.[151,1436,151,1040]" pageId="18" pageNumber="567">
|
||
Female. Externally indiscernible from male. BL
|
||
<quantity id="3A545DBCFF917370FD53FDC2FC224113" box="[749,915,512,536]" metricMagnitude="-4" metricUnit="m" metricValue="7.35" metricValueMax="7.4" metricValueMin="7.3" pageId="18" pageNumber="567" unit="mm" value="0.735" valueMax="0.74" valueMin="0.73">0.73–0.74 mm</quantity>
|
||
; HL
|
||
<quantity id="3A545DBCFF917370FC73FDC2FBC24113" box="[973,1139,512,536]" metricMagnitude="-4" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="18" pageNumber="567" unit="mm" value="0.135" valueMax="0.14" valueMin="0.13">0.13–0.14 mm</quantity>
|
||
, HW
|
||
<quantity id="3A545DBCFF917370FB08FE3DFAED4113" box="[1206,1372,511,536]" metricMagnitude="-4" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="18" pageNumber="567" unit="mm" value="0.145" valueMax="0.15" valueMin="0.14">0.14–0.15 mm</quantity>
|
||
, AnL
|
||
<quantity id="3A545DBCFF917370FF29FDE6FF4F4138" box="[151,254,548,573]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="18" pageNumber="567" unit="mm" value="0.25">0.25 mm</quantity>
|
||
; PL
|
||
<quantity id="3A545DBCFF917370FE8FFDE6FE264138" box="[305,407,548,572]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="18" pageNumber="567" unit="mm" value="0.18">0.18 mm</quantity>
|
||
, PW
|
||
<quantity id="3A545DBCFF917370FE6DFDE6FD8B4138" box="[467,570,548,572]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="18" pageNumber="567" unit="mm" value="0.18">0.18 mm</quantity>
|
||
; EL
|
||
<quantity id="3A545DBCFF917370FDD1FDE6FD644138" box="[623,725,548,572]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="18" pageNumber="567" unit="mm" value="0.43">0.43 mm</quantity>
|
||
, EW
|
||
<quantity id="3A545DBCFF917370FCADFDE6FC094138" box="[787,952,548,572]" metricMagnitude="-4" metricUnit="m" metricValue="2.9499999999999997" metricValueMax="3.0" metricValueMin="2.9" pageId="18" pageNumber="567" unit="mm" value="0.295" valueMax="0.3" valueMin="0.29">0.29–0.30 mm</quantity>
|
||
, EI 1.42–1.48.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B5B6A3D2FF917370FF79FD85FD274164" box="[199,662,583,608]" pageId="18" pageNumber="567" type="distribution">
|
||
<paragraph id="FD13F059FF917370FF79FD85FD274164" blockId="18.[151,1436,151,1040]" box="[199,662,583,608]" pageId="18" pageNumber="567">
|
||
<emphasis id="CFD82C4BFF917370FF79FD85FED14164" bold="true" box="[199,352,583,608]" pageId="18" pageNumber="567">Distribution.</emphasis>
|
||
Southern
|
||
<collectingCountry id="85BBB0C9FF917370FE6DFD85FDA64164" box="[467,535,583,608]" name="Brazil" pageId="18" pageNumber="567">Brazil</collectingCountry>
|
||
(
|
||
<figureCitation id="6597ECDCFF917370FD99FD85FDC84164" box="[551,633,583,608]" captionStart="FIGURE 68" captionStartId="12.[151,250,1290,1312]" captionTargetBox="[151,1418,167,1268]" captionTargetId="figure@12.[151,1436,167,1268]" captionTargetPageId="12" captionText="FIGURE 68. Distribution of Alloraphes: a, A. jamaicae Franz; b, A. chiapasensis Franz; c, A. yucatani Franz; d, A. lenkoi Franz; e, A. peruanus Franz; f, A. dentatus sp. n.; g, A. peckorum sp. n.; h, A. myrmecophilus Franz stat. rest.; i, A. bolivarensis Franz; j,? Alloraphes magnus Franz." httpUri="https://zenodo.org/record/217861/files/figure.png" pageId="18" pageNumber="567">Fig. 68</figureCitation>
|
||
h).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B5B6A3D2FF917370FF79FDAEFE344779" pageId="18" pageNumber="567" type="discussion">
|
||
<paragraph id="FD13F059FF917370FF79FDAEFB9540CC" blockId="18.[151,1436,151,1040]" pageId="18" pageNumber="567">
|
||
<emphasis id="CFD82C4BFF917370FF79FDAEFE8A4181" bold="true" box="[199,315,620,645]" pageId="18" pageNumber="567">Remarks.</emphasis>
|
||
This remarkably small and slender species has strikingly stout aedeagus that differs from typically elongate and slender genitalia of all remaining congeners. However, details of morphology studied in a transparent mount (
|
||
<figureCitation id="6597ECDCFF917370FF4EFD76FEF541C9" box="[240,324,692,717]" captionStart="FIGURES 22 – 26" captionStartId="7.[151,264,1642,1665]" captionTargetBox="[222,1331,193,1600]" captionTargetId="figure@7.[200,1342,193,1608]" captionTargetPageId="7" captionText="FIGURES 22 – 26. Alloraphes jamaicae Franz, holotype male (22 – 24); A. myrmecophilus Franz stat. rest., paratype male (25);? Alloraphes magnus Franz, holotype female (26). Head in ventral view (22); prothorax in ventral view (23); pterothorax in ventral view (24); body venter (25); and mesothorax in ventral view (26). Abbreviations: aest 2, mesanepisternum; aest 3, metanepisternum; ai, asetose impression; amc, anterior marginal carina of mesocoxal cavity; ar, anterior ridge; bst, basisternal part of prosternum; bst, basistipes; cd, cardo; cx, coxa; el, elytron; epm 3, metepimeron; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; lss, lateral suture of submentum; mc, median carina of mesoventrite; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mesostipes; msvp, mesoventral intercoxal process; mtap, metaventral anterior process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; pl, posterior lobe of mesocoxal projection; pre, prepectus; ppf, palpifer; psp, prosternal process; submentum; tm, temple; v 3, metaventrite" httpUri="https://zenodo.org/record/217857/files/figure.png" pageId="18" pageNumber="567">Fig. 25</figureCitation>
|
||
) do not differ from those of the
|
||
<typeStatus id="22174EFBFF917370FD7EFD77FD4141C9" box="[704,752,693,717]" pageId="18" pageNumber="567">type</typeStatus>
|
||
species of
|
||
<taxonomicName id="3AAC8BDAFF917370FCCAFD76FC5E41C8" box="[884,1007,692,716]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="567" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CFD82C4BFF917370FCCAFD76FC5E41C8" box="[884,1007,692,716]" italics="true" pageId="18" pageNumber="567">Alloraphes</emphasis>
|
||
</taxonomicName>
|
||
, and the aedeagus differs only in the general shape, and not in important characters, as the free parameres, basal foramen distant from base of median lobe, basally located lentiform sclerotization and the asymmetrical apex of median lobe. Franz (1980a) originally placed this species in
|
||
<taxonomicName id="3AAC8BDAFF917370FE2EFCE2FDBA403C" box="[400,523,800,824]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="567" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CFD82C4BFF917370FE2EFCE2FDBA403C" box="[400,523,800,824]" italics="true" pageId="18" pageNumber="567">Alloraphes</emphasis>
|
||
</taxonomicName>
|
||
, but later (Franz 1989) transferred it to
|
||
<taxonomicName id="3AAC8BDAFF917370FC6AFCE2FB1A403C" box="[980,1195,800,824]" class="Insecta" family="Staphylinidae" genus="Parastenichnaphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="567" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CFD82C4BFF917370FC6AFCE2FB1A403C" box="[980,1195,800,824]" italics="true" pageId="18" pageNumber="567">Parastenichnaphes</emphasis>
|
||
</taxonomicName>
|
||
. It is unclear why he did so, as his
|
||
<taxonomicName id="3AAC8BDAFF917370FEFCFC86FDA94058" box="[322,536,836,860]" class="Insecta" family="Staphylinidae" genus="Parastenichnaphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="567" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CFD82C4BFF917370FEFCFC86FDA94058" box="[322,536,836,860]" italics="true" pageId="18" pageNumber="567">Parastenichnaphes</emphasis>
|
||
</taxonomicName>
|
||
was defined on the basis of the aedeagus without parameres, in contrast to
|
||
<taxonomicName id="3AAC8BDAFF917370FF29FCAAFEA34084" box="[151,274,872,896]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="567" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CFD82C4BFF917370FF29FCAAFEA34084" box="[151,274,872,896]" italics="true" pageId="18" pageNumber="567">Alloraphes</emphasis>
|
||
</taxonomicName>
|
||
having the parameres (the identification key in Franz (1989)). In the original description Franz (1980a) illustrated the aedeagus of
|
||
<taxonomicName id="3AAC8BDAFF917370FE77FC4EFD2440A0" box="[457,661,908,932]" class="Insecta" family="Staphylinidae" genus="Parastenichnaphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="567" phylum="Arthropoda" rank="species" species="myrmecophilus">
|
||
<emphasis id="CFD82C4BFF917370FE77FC4EFD2440A0" box="[457,661,908,932]" italics="true" pageId="18" pageNumber="567">A. myrmecophilus</emphasis>
|
||
</taxonomicName>
|
||
with distinct parameres, and this observation was confirmed during the present study. Therefore, the original status of
|
||
<taxonomicName id="3AAC8BDAFF917370FD77FC72FC2240CC" box="[713,915,944,968]" class="Insecta" family="Staphylinidae" genus="Parastenichnaphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="567" phylum="Arthropoda" rank="species" species="myrmecophilus">
|
||
<emphasis id="CFD82C4BFF917370FD77FC72FC2240CC" box="[713,915,944,968]" italics="true" pageId="18" pageNumber="567">A. myrmecophilus</emphasis>
|
||
</taxonomicName>
|
||
is restituted.
|
||
</paragraph>
|
||
<paragraph id="FD13F059FF917370FF79FC17FF5A470B" blockId="18.[151,1436,151,1040]" pageId="18" pageNumber="567">
|
||
Franz (1980a) gives additional collecting data for the
|
||
<typeStatus id="22174EFBFF917370FC9EFC17FCE140E9" box="[800,848,981,1005]" pageId="18" pageNumber="567">type</typeStatus>
|
||
series of
|
||
<taxonomicName id="3AAC8BDAFF917370FC02FC16FB3740E8" box="[956,1158,980,1004]" class="Insecta" family="Staphylinidae" genus="Parastenichnaphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="567" phylum="Arthropoda" rank="species" species="myrmecophilus">
|
||
<emphasis id="CFD82C4BFF917370FC02FC16FB3740E8" box="[956,1158,980,1004]" italics="true" pageId="18" pageNumber="567">A. myrmecophilus</emphasis>
|
||
</taxonomicName>
|
||
: in nests of
|
||
<taxonomicName id="3AAC8BDAFF917370FAB0FC16FF544714" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="567" phylum="Arthropoda" rank="species" species="rufipes">
|
||
<emphasis id="CFD82C4BFF917370FAB0FC16FF544714" italics="true" pageId="18" pageNumber="567">Camponotus rufipes</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="FD13F059FF917370FF29FBA6FE344779" blockId="18.[151,389,1124,1149]" box="[151,389,1124,1149]" pageId="18" pageNumber="567">
|
||
<emphasis id="CFD82C4BFF917370FF29FBA6FE344779" bold="true" box="[151,389,1124,1149]" pageId="18" pageNumber="567">Species inquirendae</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B5B6A3D2FF917370FF29FB69FBE04426" pageId="18" pageNumber="567" type="description">
|
||
<paragraph id="FD13F059FF917370FF29FB69FE4447C1" blockId="18.[151,501,1195,1221]" box="[151,501,1195,1221]" pageId="18" pageNumber="567">
|
||
<taxonomicName id="3AAC8BDAFF917370FF29FB69FE4447C1" authority="Franz" authorityName="Franz" box="[151,501,1195,1221]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="567" phylum="Arthropoda" rank="species" species="bolivarensis">
|
||
<emphasis id="CFD82C4BFF917370FF29FB69FE4447C1" bold="true" box="[151,501,1195,1221]" pageId="18" pageNumber="567">
|
||
<emphasis id="CFD82C4BFF917370FF29FB69FE1747C0" bold="true" box="[151,422,1195,1220]" italics="true" pageId="18" pageNumber="567">Alloraphes bolivarensis</emphasis>
|
||
Franz
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="FD13F059FF917370FF29FB36FD39460E" blockId="18.[151,648,1268,1290]" box="[151,648,1268,1290]" pageId="18" pageNumber="567">
|
||
<taxonomicName id="3AAC8BDAFF917370FF29FB36FD9F460E" authority="Franz, 1988: 80" authorityName="Franz" authorityPageNumber="80" authorityYear="1988" box="[151,558,1268,1290]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="567" phylum="Arthropoda" rank="species" species="bolivarensis">
|
||
<emphasis id="CFD82C4BFF917370FF29FB36FE36460E" box="[151,391,1268,1290]" italics="true" pageId="18" pageNumber="567">Alloraphes bolivarensis</emphasis>
|
||
Franz, 1988: 80
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="6597ECDCFF917370FD87FB36FD35460E" box="[569,644,1268,1290]" captionStart="FIGURES 22 – 26" captionStartId="7.[151,264,1642,1665]" captionTargetBox="[222,1331,193,1600]" captionTargetId="figure@7.[200,1342,193,1608]" captionTargetPageId="7" captionText="FIGURES 22 – 26. Alloraphes jamaicae Franz, holotype male (22 – 24); A. myrmecophilus Franz stat. rest., paratype male (25);? Alloraphes magnus Franz, holotype female (26). Head in ventral view (22); prothorax in ventral view (23); pterothorax in ventral view (24); body venter (25); and mesothorax in ventral view (26). Abbreviations: aest 2, mesanepisternum; aest 3, metanepisternum; ai, asetose impression; amc, anterior marginal carina of mesocoxal cavity; ar, anterior ridge; bst, basisternal part of prosternum; bst, basistipes; cd, cardo; cx, coxa; el, elytron; epm 3, metepimeron; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; lss, lateral suture of submentum; mc, median carina of mesoventrite; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mesostipes; msvp, mesoventral intercoxal process; mtap, metaventral anterior process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; pl, posterior lobe of mesocoxal projection; pre, prepectus; ppf, palpifer; psp, prosternal process; submentum; tm, temple; v 3, metaventrite" httpUri="https://zenodo.org/record/217857/files/figure.png" pageId="18" pageNumber="567">Fig. 24</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="FD13F059FF917370FF29FAF4FCA9454E" blockId="18.[151,1437,1334,1827]" pageId="18" pageNumber="567">
|
||
This species is known from a single male collected in Ciudad Bolívar,
|
||
<collectingCountry id="85BBB0C9FF917370FC78FAF4FB88464B" box="[966,1081,1334,1359]" name="Venezuela" pageId="18" pageNumber="567">Venezuela</collectingCountry>
|
||
(
|
||
<figureCitation id="6597ECDCFF917370FBF5FAF4FB11464B" box="[1099,1184,1334,1359]" captionStart="FIGURE 68" captionStartId="12.[151,250,1290,1312]" captionTargetBox="[151,1418,167,1268]" captionTargetId="figure@12.[151,1436,167,1268]" captionTargetPageId="12" captionText="FIGURE 68. Distribution of Alloraphes: a, A. jamaicae Franz; b, A. chiapasensis Franz; c, A. yucatani Franz; d, A. lenkoi Franz; e, A. peruanus Franz; f, A. dentatus sp. n.; g, A. peckorum sp. n.; h, A. myrmecophilus Franz stat. rest.; i, A. bolivarensis Franz; j,? Alloraphes magnus Franz." httpUri="https://zenodo.org/record/217861/files/figure.png" pageId="18" pageNumber="567">
|
||
Fig.
|
||
<date id="8912D699FF917370FB3FFAF4FB11464B" box="[1153,1184,1334,1359]" pageId="18" pageNumber="567" value="1968-01">68</date>
|
||
</figureCitation>
|
||
i). According to Franz (1988), the
|
||
<typeStatus id="22174EFBFF917370FEA6FA98FECB4677" box="[280,378,1370,1395]" pageId="18" pageNumber="567" type="holotype">holotype</typeStatus>
|
||
was deposited in his collection (currently housed at NHMW). The entire Franz Collection was surveyed by the author in 2004, and
|
||
<taxonomicName id="3AAC8BDAFF917370FD8AFABDFD6D4692" box="[564,732,1406,1430]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="567" phylum="Arthropoda" rank="species" species="bolivarensis">
|
||
<emphasis id="CFD82C4BFF917370FD8AFABDFD6D4692" box="[564,732,1406,1430]" italics="true" pageId="18" pageNumber="567">A. bolivarensis</emphasis>
|
||
</taxonomicName>
|
||
was not found; repeated search by the NHMW curator Harald Schillhammer in 2013 was also unsuccessful (email from H. Schillhammer dated
|
||
<date id="8912D699FF917370FB94FA61FB0D46BF" box="[1066,1212,1443,1467]" pageId="18" pageNumber="567" value="2013-09-09">9 Sept. 2013</date>
|
||
). However, cases of labeling
|
||
<typeStatus id="22174EFBFF917370FF47FA05FE9846DB" box="[249,297,1479,1503]" pageId="18" pageNumber="567">type</typeStatus>
|
||
specimens with a different name than the subsequently published one are not infrequent in the Franz Collection (Jałoszyński, pers. obs.) and it is still possible that the
|
||
<typeStatus id="22174EFBFF917370FCD3FA28FC7E4507" box="[877,975,1514,1539]" pageId="18" pageNumber="567" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="3AAC8BDAFF917370FC4DFA29FB2B4507" box="[1011,1178,1515,1539]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="567" phylum="Arthropoda" rank="species" species="bolivarensis">
|
||
<emphasis id="CFD82C4BFF917370FC4DFA29FB2B4507" box="[1011,1178,1515,1539]" italics="true" pageId="18" pageNumber="567">A. bolivarensis</emphasis>
|
||
</taxonomicName>
|
||
can be found in future. Since the description and illustrations presented by Franz (1988) are insufficiently detailed to unambiguously identify this species, it must remain a
|
||
<emphasis id="CFD82C4BFF917370FD81F9F1FCA0454F" box="[575,785,1587,1611]" italics="true" pageId="18" pageNumber="567">species inquirenda</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="FD13F059FF917370FF79F995FBE04426" blockId="18.[151,1437,1334,1827]" pageId="18" pageNumber="567">
|
||
Franz (1988) mentioned that the elytral apices of
|
||
<taxonomicName id="3AAC8BDAFF917370FC9CF995FC7E456A" box="[802,975,1622,1646]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="567" phylum="Arthropoda" rank="species" species="bolivarensis">
|
||
<emphasis id="CFD82C4BFF917370FC9CF995FC7E456A" box="[802,975,1622,1646]" italics="true" pageId="18" pageNumber="567">A. bolivarensis</emphasis>
|
||
</taxonomicName>
|
||
are protruding posteriorly, which may resemble modifications seen in
|
||
<taxonomicName id="3AAC8BDAFF917370FE40F9B9FD344596" box="[510,645,1659,1682]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="567" phylum="Arthropoda" rank="species" species="jamaicae">
|
||
<emphasis id="CFD82C4BFF917370FE40F9B9FD344596" box="[510,645,1659,1682]" italics="true" pageId="18" pageNumber="567">A. jamaicae</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="3AAC8BDAFF917370FD12F9B9FC9C4596" box="[684,813,1659,1682]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="567" phylum="Arthropoda" rank="species" species="yucatani">
|
||
<emphasis id="CFD82C4BFF917370FD12F9B9FC9C4596" box="[684,813,1659,1682]" italics="true" pageId="18" pageNumber="567">A. yucatani</emphasis>
|
||
</taxonomicName>
|
||
. The aedeagus illustrated in latero-ventral view (Franz 1988:
|
||
<figureCitation id="6597ECDCFF917370FF61F95CFE8145B3" box="[223,304,1694,1719]" captionStart="FIGURES 22 – 26" captionStartId="7.[151,264,1642,1665]" captionTargetBox="[222,1331,193,1600]" captionTargetId="figure@7.[200,1342,193,1608]" captionTargetPageId="7" captionText="FIGURES 22 – 26. Alloraphes jamaicae Franz, holotype male (22 – 24); A. myrmecophilus Franz stat. rest., paratype male (25);? Alloraphes magnus Franz, holotype female (26). Head in ventral view (22); prothorax in ventral view (23); pterothorax in ventral view (24); body venter (25); and mesothorax in ventral view (26). Abbreviations: aest 2, mesanepisternum; aest 3, metanepisternum; ai, asetose impression; amc, anterior marginal carina of mesocoxal cavity; ar, anterior ridge; bst, basisternal part of prosternum; bst, basistipes; cd, cardo; cx, coxa; el, elytron; epm 3, metepimeron; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; lss, lateral suture of submentum; mc, median carina of mesoventrite; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mesostipes; msvp, mesoventral intercoxal process; mtap, metaventral anterior process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; pl, posterior lobe of mesocoxal projection; pre, prepectus; ppf, palpifer; psp, prosternal process; submentum; tm, temple; v 3, metaventrite" httpUri="https://zenodo.org/record/217857/files/figure.png" pageId="18" pageNumber="567">Fig. 24</figureCitation>
|
||
) and without any internal details has unusually elongate and slender apical part of the median lobe, dissimilar to those of
|
||
<taxonomicName id="3AAC8BDAFF917370FE2AF901FDAD45DE" box="[404,540,1731,1754]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="567" phylum="Arthropoda" rank="species" species="jamaicae">
|
||
<emphasis id="CFD82C4BFF917370FE2AF901FDAD45DE" box="[404,540,1731,1754]" italics="true" pageId="18" pageNumber="567">A. jamaicae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="3AAC8BDAFF917370FDE6F901FD6B45DE" box="[600,730,1731,1754]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="567" phylum="Arthropoda" rank="species" species="yucatani">
|
||
<emphasis id="CFD82C4BFF917370FDE6F901FD6B45DE" box="[600,730,1731,1754]" italics="true" pageId="18" pageNumber="567">A. yucatani</emphasis>
|
||
</taxonomicName>
|
||
, but somehow resembling the aedeagus of
|
||
<taxonomicName id="3AAC8BDAFF917370FB6EF901FAEE45DF" box="[1232,1375,1731,1755]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="567" phylum="Arthropoda" rank="species" species="peckorum">
|
||
<emphasis id="CFD82C4BFF917370FB6EF901FAEE45DF" box="[1232,1375,1731,1755]" italics="true" pageId="18" pageNumber="567">A. peckorum</emphasis>
|
||
</taxonomicName>
|
||
. The parameres of
|
||
<taxonomicName id="3AAC8BDAFF917370FE87F925FE5445FA" box="[313,485,1766,1790]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="567" phylum="Arthropoda" rank="species" species="bolivarensis">
|
||
<emphasis id="CFD82C4BFF917370FE87F925FE5445FA" box="[313,485,1766,1790]" italics="true" pageId="18" pageNumber="567">A. bolivarensis</emphasis>
|
||
</taxonomicName>
|
||
were illustrated as slender, barely broadened distally and much shorter than the median lobe; if this character is correct than no other
|
||
<taxonomicName id="3AAC8BDAFF917370FD53F8C9FCD94427" box="[749,872,1803,1827]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="567" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CFD82C4BFF917370FD53F8C9FCD94427" box="[749,872,1803,1827]" italics="true" pageId="18" pageNumber="567">Alloraphes</emphasis>
|
||
</taxonomicName>
|
||
has such parameres.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |