1112 lines
103 KiB
XML
1112 lines
103 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/subtbiol.41.76926" ID-GBIF-Dataset="51e88f30-cebb-4ab4-97a1-64f972ad1e10" ID-GBIF-Taxon="191263934" ID-Pensoft-Pub="1314-2615-41-1" ID-Pensoft-UUID="482986A3D1D15408B0FE4F5EF7587312" ID-ZooBank="73326F5CC4BE431DBE6D601A69BD7FFA" LSID="urn:lsid:zoobank.org:act:84FBEE6D-88C3-459F-B8D7-1B8A741D7393" ModsDocID="1314-2615-41-1" checkinTime="1639196417009" checkinUser="pensoft" docAuthor="Nilsai 1, Areeruk, Detcharoen 1, Matsapume, Godeiro 2, Nerivania Nunes & Jantarit 3, Sopark" docDate="2021" docId="750035DFE39158DA9033642DD6AD4619" docLanguage="en" docName="SubterBiol 41: 1-42" docOrigin="Subterranean Biology 41" docPubDate="2021-12-10" docSource="http://dx.doi.org/10.3897/subtbiol.41.76926" docTitle="Coecobrya whitteni Nilsai & Jantarit 2021, sp. nov." docType="treatment" docUuid="84FBEE6D-88C3-459F-B8D7-1B8A741D7393" docUuidSource="ZooBank" docVersion="4" id="482986A3D1D15408B0FE4F5EF7587312" lastPageNumber="1" masterDocId="482986A3D1D15408B0FE4F5EF7587312" masterDocTitle="Four new species of troglomorphic Coecobrya Yosii, 1956 (Collembola, Entomobryidae) from Thailand based on morphological and molecular evidence, with an updated key of Thai troglomorphic species" masterLastPageNumber="42" masterPageNumber="1" pageId="0" pageNumber="1" updateTime="1639202433658" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Four new species of troglomorphic Coecobrya Yosii, 1956 (Collembola, Entomobryidae) from Thailand based on morphological and molecular evidence, with an updated key of Thai troglomorphic species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Nilsai 1, Areeruk</mods:namePart>
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<mods:affiliation>Division of Biological Science, Faculty of Science, Prince of Songkla University, Hat Yai, Songkhla, 90110, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Detcharoen 1, Matsapume</mods:namePart>
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<mods:affiliation>Division of Biological Science, Faculty of Science, Prince of Songkla University, Hat Yai, Songkhla, 90110, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Godeiro 2, Nerivania Nunes</mods:namePart>
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<mods:affiliation>Division of Biological Science, Faculty of Science, Prince of Songkla University, Hat Yai, Songkhla, 90110, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jantarit 3, Sopark</mods:namePart>
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<mods:affiliation>Division of Biological Science, Faculty of Science, Prince of Songkla University, Hat Yai, Songkhla, 90110, Thailand</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Subterranean Biology</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-12-10</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>41</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>42</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/subtbiol.41.76926</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/subtbiol.41.76926</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2615-41-1</mods:identifier>
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<mods:identifier type="ZooBank">73326F5CC4BE431DBE6D601A69BD7FFA</mods:identifier>
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<mods:identifier type="Pensoft-UUID">482986A3D1D15408B0FE4F5EF7587312</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="191263934" LSID="urn:lsid:zoobank.org:act:84FBEE6D-88C3-459F-B8D7-1B8A741D7393" httpUri="http://treatment.plazi.org/id/750035DFE39158DA9033642DD6AD4619" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/84FBEE6D-88C3-459F-B8D7-1B8A741D7393" authority="Nilsai & Jantarit" authorityName="Nilsai & Jantarit" authorityYear="2021" class="Collembola" family="Entomobryidae" genus="Coecobrya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coecobrya whitteni" order="Entomobryomorpha" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="whitteni" status="sp. nov.">Coecobrya whitteni Nilsai & Jantarit</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. The distribution map of the troglomorphic Coecobrya in Thailand and the habitus of the four new species. (1) A Coecobrya whitteni sp. nov. (2) B C. troglobia sp. nov. (3) C C. ellisi sp. nov. (4) D C. phitsanulokensis sp. nov. (5) C. chumphonensis Zhang & Nilsai, 2017 (6) C. sirindhornae Jantarit, Satasook & Deharveng, 2019 (7) C. polychaeta Zhang & Nilsai, 2017 and (8) C. cavicta Nilsai & Zhang, 2017. Scale bar: 1 mm." figureDoi="10.3897/subtbiol.41.76926.figure1" httpUri="https://binary.pensoft.net/fig/621479" pageId="0" pageNumber="1">Figures 1A</figureCitation>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Coecobrya whitteni sp. nov. A Ventro-distal complex of labrum B Outer maxillary lobe C Labial palp D Ant. III organ E Chaetae of labial basis and ventral chaetotaxy of head F Mucro G Distal part of tita III and claw complex H Prelabral and labral chaetae I Mandibles." figureDoi="10.3897/subtbiol.41.76926.figure2" httpUri="https://binary.pensoft.net/fig/621480" pageId="0" pageNumber="1">, 2</figureCitation>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Coecobrya whitteni sp. nov., continued A Distal part of manubrium ventrally B Trochanteral organ C Posterior side of ventral tube and Lateral flap D Dorsal cephalic chaetotaxy E Anterior side of ventral tube F Manubrium plaque G Clypeal chaetae." figureDoi="10.3897/subtbiol.41.76926.figure3" httpUri="https://binary.pensoft.net/fig/621481" pageId="0" pageNumber="1">, 3</figureCitation>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Coecobrya whitteni sp. nov., continued A Chaetotaxy of dorsal Th. II- III B Chaetotaxy of dorsal Abd. I- III C Chaetotaxy of dorsal Abd. IV D Chaetotaxy of dorsal Abd. V." figureDoi="10.3897/subtbiol.41.76926.figure4" httpUri="https://binary.pensoft.net/fig/621482" pageId="0" pageNumber="1">, 4</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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: female on slide. Thailand, Khon Kaen province, Chum Phae district, Tham Nayn Noi (note:
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<normalizedToken originalValue="“tham”">"tham"</normalizedToken>
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=
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<normalizedToken originalValue="“cave”">"cave"</normalizedToken>
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in Thai), altitude 359 m a.m.s.l.,
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<geoCoordinate degrees="16.8292" direction="north" orientation="latitude" precision="5" value="16.8292">16.8292°N</geoCoordinate>
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,
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<geoCoordinate degrees="101.9848" direction="east" orientation="longitude" precision="5" value="101.9848">101.9848°E</geoCoordinate>
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. 3.XII.2020; S. Jantarit, A. Nilsai, K. Sarakhamhaeng and K. Jantapaso leg. (sample # THA_SJ_KKN04), dark zone of a cave, by entomological aspirator.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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: same data as holotype, 11 specimens (three female and eight subadults on slides).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Additional material</emphasis>
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: same data as holotype, 3 specimens (in ethanol)
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</paragraph>
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<paragraph pageId="0" pageNumber="1">Holotype and 11 paratypes on slides deposited in NHM-PSU.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Habitus</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. The distribution map of the troglomorphic Coecobrya in Thailand and the habitus of the four new species. (1) A Coecobrya whitteni sp. nov. (2) B C. troglobia sp. nov. (3) C C. ellisi sp. nov. (4) D C. phitsanulokensis sp. nov. (5) C. chumphonensis Zhang & Nilsai, 2017 (6) C. sirindhornae Jantarit, Satasook & Deharveng, 2019 (7) C. polychaeta Zhang & Nilsai, 2017 and (8) C. cavicta Nilsai & Zhang, 2017. Scale bar: 1 mm." figureDoi="10.3897/subtbiol.41.76926.figure1" httpUri="https://binary.pensoft.net/fig/621479" pageId="0" pageNumber="1">1A</figureCitation>
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). Medium size
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<taxonomicName class="Collembola" family="Entomobryidae" kingdom="Animalia" lsidName="" order="Entomobryomorpha" pageId="0" pageNumber="1" phylum="Arthropoda" rank="family">Entomobryidae</taxonomicName>
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. Body length 1.8-2.3 mm (holotype 1.8 mm). No scales. Eyes absent. Color: whitish in alcohol, without pigmentation. Four antennal segments. Body slender not bent nor humped at the level of Th. II. Abd. IV 3.80-5.63 times as long as Abd. III along the dorsal midline.
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</paragraph>
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<caption doi="10.3897/subtbiol.41.76926.figure1" httpUri="https://binary.pensoft.net/fig/621479" pageId="0" pageNumber="1" start="Figure 1" startId="F1">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 1.</emphasis>
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The distribution map of the troglomorphic
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<taxonomicName authorityName="Yosii" authorityYear="1956" class="Collembola" family="Entomobryidae" genus="Coecobrya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coecobrya" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Coecobrya</emphasis>
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</taxonomicName>
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in Thailand and the habitus of the four new species. (
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<emphasis bold="true" pageId="0" pageNumber="1">1</emphasis>
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)
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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<taxonomicName authorityName="Nilsai & Jantarit" authorityYear="2021" class="Collembola" family="Entomobryidae" genus="Coecobrya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coecobrya whitteni" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="whitteni">
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<emphasis italics="true" pageId="0" pageNumber="1">Coecobrya whitteni</emphasis>
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</taxonomicName>
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sp. nov. (
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<emphasis bold="true" pageId="0" pageNumber="1">2</emphasis>
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)
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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<taxonomicName class="Collembola" genus="C." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. troglobia" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="troglobia">
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<emphasis italics="true" pageId="0" pageNumber="1">C. troglobia</emphasis>
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</taxonomicName>
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sp. nov. (
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<emphasis bold="true" pageId="0" pageNumber="1">3</emphasis>
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)
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<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
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<taxonomicName class="Collembola" genus="C." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. ellisi" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ellisi">
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<emphasis italics="true" pageId="0" pageNumber="1">C. ellisi</emphasis>
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</taxonomicName>
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sp. nov. (
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<emphasis bold="true" pageId="0" pageNumber="1">4</emphasis>
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)
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<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
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<taxonomicName class="Collembola" genus="C." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. phitsanulokensis" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="phitsanulokensis">
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<emphasis italics="true" pageId="0" pageNumber="1">C. phitsanulokensis</emphasis>
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</taxonomicName>
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sp. nov. (
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<emphasis bold="true" pageId="0" pageNumber="1">5</emphasis>
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)
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<taxonomicName class="Collembola" genus="C." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. chumphonensis" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="chumphonensis">
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<emphasis italics="true" pageId="0" pageNumber="1">C. chumphonensis</emphasis>
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</taxonomicName>
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Zhang & Nilsai, 2017 (
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<emphasis bold="true" pageId="0" pageNumber="1">6</emphasis>
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)
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<taxonomicName class="Collembola" genus="C." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. sirindhornae" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sirindhornae">
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<emphasis italics="true" pageId="0" pageNumber="1">C. sirindhornae</emphasis>
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</taxonomicName>
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Jantarit, Satasook & Deharveng, 2019 (
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<emphasis bold="true" pageId="0" pageNumber="1">7</emphasis>
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)
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<taxonomicName class="Collembola" genus="C." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. polychaeta" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="polychaeta">
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<emphasis italics="true" pageId="0" pageNumber="1">C. polychaeta</emphasis>
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</taxonomicName>
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Zhang & Nilsai, 2017 and (
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<emphasis bold="true" pageId="0" pageNumber="1">8</emphasis>
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)
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<taxonomicName class="Collembola" genus="C." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. cavicta" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="cavicta">
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<emphasis italics="true" pageId="0" pageNumber="1">C. cavicta</emphasis>
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</taxonomicName>
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Nilsai & Zhang, 2017. Scale bar: 1 mm.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Pseudopores</emphasis>
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(Figs
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Coecobrya whitteni sp. nov., continued A Distal part of manubrium ventrally B Trochanteral organ C Posterior side of ventral tube and Lateral flap D Dorsal cephalic chaetotaxy E Anterior side of ventral tube F Manubrium plaque G Clypeal chaetae." figureDoi="10.3897/subtbiol.41.76926.figure3" httpUri="https://binary.pensoft.net/fig/621481" pageId="0" pageNumber="1">3F</figureCitation>
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,
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Coecobrya whitteni sp. nov., continued A Chaetotaxy of dorsal Th. II- III B Chaetotaxy of dorsal Abd. I- III C Chaetotaxy of dorsal Abd. IV D Chaetotaxy of dorsal Abd. V." figureDoi="10.3897/subtbiol.41.76926.figure4" httpUri="https://binary.pensoft.net/fig/621482" pageId="0" pageNumber="1">4A-C</figureCitation>
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). Pseudopores present as round flat disks, smaller than mac sockets, except for the coxae and manubrium where psp are as large as mac sockets, present on various parts of the body: antennae, head, tergites, coxae and manubrium. On antennae, psp located ventro-apically between the tip of antennal segments and the chaetae of the apical row, or just below the apical row of chaetae (2 psp on Ant. I, 2-3 psp on Ant. II, and 3 psp on Ant. III). On the head, 1-2 psp located externally on each peri-antennal area. On tergites, 1+1 psp close to the axis from Th. II to Abd. IV (Figs
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Coecobrya whitteni sp. nov., continued A Chaetotaxy of dorsal Th. II- III B Chaetotaxy of dorsal Abd. I- III C Chaetotaxy of dorsal Abd. IV D Chaetotaxy of dorsal Abd. V." figureDoi="10.3897/subtbiol.41.76926.figure4" httpUri="https://binary.pensoft.net/fig/621482" pageId="0" pageNumber="1">4A-C</figureCitation>
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). On coxae, 1-2 psp on coxae I, 2-3 psp on coxae II and 1-2 psp on coxae III, located close to longitudinal rows of chaetae. On manubrium, 2+2 dorso-apical ones (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Coecobrya whitteni sp. nov., continued A Distal part of manubrium ventrally B Trochanteral organ C Posterior side of ventral tube and Lateral flap D Dorsal cephalic chaetotaxy E Anterior side of ventral tube F Manubrium plaque G Clypeal chaetae." figureDoi="10.3897/subtbiol.41.76926.figure3" httpUri="https://binary.pensoft.net/fig/621481" pageId="0" pageNumber="1">3F</figureCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Clypeus and mouthparts</emphasis>
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(Figs
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Coecobrya whitteni sp. nov. A Ventro-distal complex of labrum B Outer maxillary lobe C Labial palp D Ant. III organ E Chaetae of labial basis and ventral chaetotaxy of head F Mucro G Distal part of tita III and claw complex H Prelabral and labral chaetae I Mandibles." figureDoi="10.3897/subtbiol.41.76926.figure2" httpUri="https://binary.pensoft.net/fig/621480" pageId="0" pageNumber="1">2A-C</figureCitation>
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,
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Coecobrya whitteni sp. nov. A Ventro-distal complex of labrum B Outer maxillary lobe C Labial palp D Ant. III organ E Chaetae of labial basis and ventral chaetotaxy of head F Mucro G Distal part of tita III and claw complex H Prelabral and labral chaetae I Mandibles." figureDoi="10.3897/subtbiol.41.76926.figure2" httpUri="https://binary.pensoft.net/fig/621480" pageId="0" pageNumber="1">2H, I</figureCitation>
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,
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Coecobrya whitteni sp. nov., continued A Distal part of manubrium ventrally B Trochanteral organ C Posterior side of ventral tube and Lateral flap D Dorsal cephalic chaetotaxy E Anterior side of ventral tube F Manubrium plaque G Clypeal chaetae." figureDoi="10.3897/subtbiol.41.76926.figure3" httpUri="https://binary.pensoft.net/fig/621481" pageId="0" pageNumber="1">3G</figureCitation>
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). Clypeal area with three long, smooth prefrontal; six middle chaetae (2+2 small ciliated chaetae of unequal size posteriorly and 1+1 rather long, thin, smooth chaetae anteriorly); and 1+1 long smooth lateral chaetae (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Coecobrya whitteni sp. nov., continued A Distal part of manubrium ventrally B Trochanteral organ C Posterior side of ventral tube and Lateral flap D Dorsal cephalic chaetotaxy E Anterior side of ventral tube F Manubrium plaque G Clypeal chaetae." figureDoi="10.3897/subtbiol.41.76926.figure3" httpUri="https://binary.pensoft.net/fig/621481" pageId="0" pageNumber="1">3G</figureCitation>
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). Prelabral and labral chaetae 4/5, 5, 4, all thin and smooth; the three median chaetae of the second row longer than two lateral ones (35-40 vs. 25
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<normalizedToken originalValue="µm">µm</normalizedToken>
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) (Fig.
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|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Coecobrya whitteni sp. nov. A Ventro-distal complex of labrum B Outer maxillary lobe C Labial palp D Ant. III organ E Chaetae of labial basis and ventral chaetotaxy of head F Mucro G Distal part of tita III and claw complex H Prelabral and labral chaetae I Mandibles." figureDoi="10.3897/subtbiol.41.76926.figure2" httpUri="https://binary.pensoft.net/fig/621480" pageId="0" pageNumber="1">2H</figureCitation>
|
|
). Distal border of the apical non-granulated area of the labrum with a relatively narrow median U- or V-form intrusion into the granulated area dorsally; apical edge without spines (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Coecobrya whitteni sp. nov. A Ventro-distal complex of labrum B Outer maxillary lobe C Labial palp D Ant. III organ E Chaetae of labial basis and ventral chaetotaxy of head F Mucro G Distal part of tita III and claw complex H Prelabral and labral chaetae I Mandibles." figureDoi="10.3897/subtbiol.41.76926.figure2" httpUri="https://binary.pensoft.net/fig/621480" pageId="0" pageNumber="1">2H</figureCitation>
|
|
). Ventro-distal complex of labrum well differentiated, asymmetrical, with 1+1 distal combs of 14-19 minute teeth on the right side and 14-16 strong and larger teeth on the left side (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Coecobrya whitteni sp. nov. A Ventro-distal complex of labrum B Outer maxillary lobe C Labial palp D Ant. III organ E Chaetae of labial basis and ventral chaetotaxy of head F Mucro G Distal part of tita III and claw complex H Prelabral and labral chaetae I Mandibles." figureDoi="10.3897/subtbiol.41.76926.figure2" httpUri="https://binary.pensoft.net/fig/621480" pageId="0" pageNumber="1">2A</figureCitation>
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|
), and an axial pair of long sinuous tubules. Maxillary outer lobe with one basal chaeta, one apical chaeta (basal chaeta thicker than apical one) and three smooth sublobal hairs (55-60 vs. 35-38
|
|
<normalizedToken originalValue="µm">µm</normalizedToken>
|
|
) (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Coecobrya whitteni sp. nov. A Ventro-distal complex of labrum B Outer maxillary lobe C Labial palp D Ant. III organ E Chaetae of labial basis and ventral chaetotaxy of head F Mucro G Distal part of tita III and claw complex H Prelabral and labral chaetae I Mandibles." figureDoi="10.3897/subtbiol.41.76926.figure2" httpUri="https://binary.pensoft.net/fig/621480" pageId="0" pageNumber="1">2B</figureCitation>
|
|
). Labial palp strongly modified for the genus, with 0, 5, 0, 4, 4 guards for papillae A-E, as described by
|
|
<bibRefCitation author="Fjellberg, A" journalOrPublisher="Zoologischer Anzeiger" pageId="0" pageNumber="1" pagination="309 - 330" refId="B12" refString="Fjellberg, A, 1999. The labial palp in Collembola. Zoologischer Anzeiger 237: 309 - 330" title="The labial palp in Collembola." volume="237" year="1999">Fjellberg (1999)</bibRefCitation>
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for
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<taxonomicName class="Collembola" family="Entomobryidae" kingdom="Animalia" lsidName="" order="Entomobryomorpha" pageId="0" pageNumber="1" phylum="Arthropoda" rank="family">Entomobryidae</taxonomicName>
|
|
or by
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|
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.498.9491" author="Xu, GL" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="17 - 28" refId="B36" refString="Xu, GL, Zhang, F, 2015. Two new species of Coecobrya (Collembola, Entomobryidae) from China, with an updated key to the Chinese species of the genus. ZooKeys 498: 17 - 28, DOI: https://doi.org/10.3897/zookeys.498.9491" title="Two new species of Coecobrya (Collembola, Entomobryidae) from China, with an updated key to the Chinese species of the genus." url="https://doi.org/10.3897/zookeys.498.9491" volume="498" year="2015">Xu and Zhang (2015)</bibRefCitation>
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and
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<bibRefCitation DOI="https://doi.org/10.3897/zookeys.824.31635" author="Jantarit, S" journalOrPublisher="Zookey" pageId="0" pageNumber="1" pagination="21 - 44" refId="B18" refString="Jantarit, S, Satasook, C, Deharveng, L, 2019. Coecobrya sirindhornae sp. n., the most highly troglomorphic Collembola in Southeast Asia (Collembola, Entomobryidae). Zookey 824: 21 - 44, DOI: https://doi.org/10.3897/zookeys.824.31635" title="Coecobrya sirindhornae sp. n., the most highly troglomorphic Collembola in Southeast Asia (Collembola, Entomobryidae)." url="https://doi.org/10.3897/zookeys.824.31635" volume="824" year="2019">Jantarit et al. (2019)</bibRefCitation>
|
|
for
|
|
<taxonomicName authorityName="Yosii" authorityYear="1956" class="Collembola" family="Entomobryidae" genus="Coecobrya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coecobrya" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Coecobrya</emphasis>
|
|
</taxonomicName>
|
|
. Lateral process of labial palp subcylindrical, as thick as normal chaetae, with tip beyond the apex of the labial papilla (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Coecobrya whitteni sp. nov. A Ventro-distal complex of labrum B Outer maxillary lobe C Labial palp D Ant. III organ E Chaetae of labial basis and ventral chaetotaxy of head F Mucro G Distal part of tita III and claw complex H Prelabral and labral chaetae I Mandibles." figureDoi="10.3897/subtbiol.41.76926.figure2" httpUri="https://binary.pensoft.net/fig/621480" pageId="0" pageNumber="1">2C</figureCitation>
|
|
). Mandible apex blunt and strong, asymmetrical (left with four teeth, right with five teeth) (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Coecobrya whitteni sp. nov. A Ventro-distal complex of labrum B Outer maxillary lobe C Labial palp D Ant. III organ E Chaetae of labial basis and ventral chaetotaxy of head F Mucro G Distal part of tita III and claw complex H Prelabral and labral chaetae I Mandibles." figureDoi="10.3897/subtbiol.41.76926.figure2" httpUri="https://binary.pensoft.net/fig/621480" pageId="0" pageNumber="1">2I</figureCitation>
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|
); molar plate with three strong pointed basal teeth, and 3-(5) smaller inner distal teeth, identical in both mandibles. Maxilla capitulum with a three-toothed claw and several stout ciliated lamellae; lamella 2 large and broad, lamella 3 well developed; several other lamellae present as figured in
|
|
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.824.31635" author="Jantarit, S" journalOrPublisher="Zookey" pageId="0" pageNumber="1" pagination="21 - 44" refId="B18" refString="Jantarit, S, Satasook, C, Deharveng, L, 2019. Coecobrya sirindhornae sp. n., the most highly troglomorphic Collembola in Southeast Asia (Collembola, Entomobryidae). Zookey 824: 21 - 44, DOI: https://doi.org/10.3897/zookeys.824.31635" title="Coecobrya sirindhornae sp. n., the most highly troglomorphic Collembola in Southeast Asia (Collembola, Entomobryidae)." url="https://doi.org/10.3897/zookeys.824.31635" volume="824" year="2019">Jantarit et al. (2019)</bibRefCitation>
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.
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|
</paragraph>
|
|
<caption doi="10.3897/subtbiol.41.76926.figure2" httpUri="https://binary.pensoft.net/fig/621480" pageId="0" pageNumber="1" start="Figure 2" startId="F2">
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<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="1">Figure 2.</emphasis>
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|
<taxonomicName authorityName="Nilsai & Jantarit" authorityYear="2021" class="Collembola" family="Entomobryidae" genus="Coecobrya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coecobrya whitteni" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="whitteni">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Coecobrya whitteni</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.
|
|
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
|
Ventro-distal complex of labrum
|
|
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
|
Outer maxillary lobe
|
|
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
|
|
Labial palp
|
|
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
|
|
Ant. III organ
|
|
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
|
|
Chaetae of labial basis and ventral chaetotaxy of head
|
|
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
|
|
Mucro
|
|
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
|
|
Distal part of tita III and claw complex
|
|
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
|
|
Prelabral and labral chaetae
|
|
<emphasis bold="true" pageId="0" pageNumber="1">I</emphasis>
|
|
Mandibles.
|
|
</paragraph>
|
|
</caption>
|
|
<caption doi="10.3897/subtbiol.41.76926.figure3" httpUri="https://binary.pensoft.net/fig/621481" pageId="0" pageNumber="1" start="Figure 3" startId="F3">
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="1">Figure 3.</emphasis>
|
|
<taxonomicName authorityName="Nilsai & Jantarit" authorityYear="2021" class="Collembola" family="Entomobryidae" genus="Coecobrya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coecobrya whitteni" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="whitteni">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Coecobrya whitteni</emphasis>
|
|
</taxonomicName>
|
|
sp. nov., continued
|
|
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
|
Distal part of manubrium ventrally
|
|
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
|
Trochanteral organ
|
|
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
|
|
Posterior side of ventral tube and Lateral flap
|
|
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
|
|
Dorsal cephalic chaetotaxy
|
|
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
|
|
Anterior side of ventral tube
|
|
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
|
|
Manubrium plaque
|
|
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
|
|
Clypeal chaetae.
|
|
</paragraph>
|
|
</caption>
|
|
<caption doi="10.3897/subtbiol.41.76926.figure4" httpUri="https://binary.pensoft.net/fig/621482" pageId="0" pageNumber="1" start="Figure 4" startId="F4">
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="1">Figure 4.</emphasis>
|
|
<taxonomicName authorityName="Nilsai & Jantarit" authorityYear="2021" class="Collembola" family="Entomobryidae" genus="Coecobrya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coecobrya whitteni" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="whitteni">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Coecobrya whitteni</emphasis>
|
|
</taxonomicName>
|
|
sp. nov., continued A Chaetotaxy of dorsal Th. II- III
|
|
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
|
Chaetotaxy of dorsal Abd. I- III
|
|
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
|
|
Chaetotaxy of dorsal Abd. IV
|
|
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
|
|
Chaetotaxy of dorsal Abd. V.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Antennae</emphasis>
|
|
(Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Coecobrya whitteni sp. nov. A Ventro-distal complex of labrum B Outer maxillary lobe C Labial palp D Ant. III organ E Chaetae of labial basis and ventral chaetotaxy of head F Mucro G Distal part of tita III and claw complex H Prelabral and labral chaetae I Mandibles." figureDoi="10.3897/subtbiol.41.76926.figure2" httpUri="https://binary.pensoft.net/fig/621480" pageId="0" pageNumber="1">2D</figureCitation>
|
|
). Antennae very long, approximately 3.78-5.14 times as long as cephalic diagonal. Antennal segments ratio as I: II: III: IV. 1: 2.08-2.20: 2.17-2.21: 3.75-4.40 (N = 3). Antennal segments not subdivided nor annulated. Antennal chaetal types not analysed in detail. Ant. I ventrally with many smooth spiny mic of various sizes in its basal part, many subcylindrical, hyaline sens in its middle to apical part, and many long smooth straight chaetae. The paddle-like chaetae (sensu
|
|
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4286.2.3" author="Nilsai, A" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="187 - 202" refId="B26" refString="Nilsai, A, Jantarit, S, Satasook, C, Zhang, F, 2017. Three new species of Coecobrya (Collembola: Entomobryidae) from caves in the Thai Peninsula. Zootaxa 4286 (2): 187 - 202, DOI: https://doi.org/10.11646/zootaxa.4286.2.3" title="Three new species of Coecobrya (Collembola: Entomobryidae) from caves in the Thai Peninsula." url="https://doi.org/10.11646/zootaxa.4286.2.3" volume="4286" year="2017">Nilsai et al. 2017</bibRefCitation>
|
|
) on Ant II dorsally absent. Ant. III organ with typical five sens, sens one and four subequal, hyaline; sens five acuminate, and shorter; sens two and three swollen in paddle-like chaetae shape (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Coecobrya whitteni sp. nov. A Ventro-distal complex of labrum B Outer maxillary lobe C Labial palp D Ant. III organ E Chaetae of labial basis and ventral chaetotaxy of head F Mucro G Distal part of tita III and claw complex H Prelabral and labral chaetae I Mandibles." figureDoi="10.3897/subtbiol.41.76926.figure2" httpUri="https://binary.pensoft.net/fig/621480" pageId="0" pageNumber="1">2D</figureCitation>
|
|
). Ant. IV without apical bulb. Subapical organite not distinctly knobbed, swollen, slightly enlarged apically, inserted dorsally.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Dorsal head chaetotaxy</emphasis>
|
|
(Fig.
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Coecobrya whitteni sp. nov., continued A Distal part of manubrium ventrally B Trochanteral organ C Posterior side of ventral tube and Lateral flap D Dorsal cephalic chaetotaxy E Anterior side of ventral tube F Manubrium plaque G Clypeal chaetae." figureDoi="10.3897/subtbiol.41.76926.figure3" httpUri="https://binary.pensoft.net/fig/621481" pageId="0" pageNumber="1">3D</figureCitation>
|
|
). Dorsal cephalic chaetotaxy with four antennal chaetae (An3a2 and An3 as mac, An1-2 as mes), four anterior mac (A0, A2-3 and A5), three median (M1, M2 and M4) and eight sutural (S) mac (S0, S1-7); Gr. II with four mac; A0 as mac; 5-7+5-7 scale-like structures (usually 6+6) (sensu
|
|
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.824.31635" author="Jantarit, S" journalOrPublisher="Zookey" pageId="0" pageNumber="1" pagination="21 - 44" refId="B18" refString="Jantarit, S, Satasook, C, Deharveng, L, 2019. Coecobrya sirindhornae sp. n., the most highly troglomorphic Collembola in Southeast Asia (Collembola, Entomobryidae). Zookey 824: 21 - 44, DOI: https://doi.org/10.3897/zookeys.824.31635" title="Coecobrya sirindhornae sp. n., the most highly troglomorphic Collembola in Southeast Asia (Collembola, Entomobryidae)." url="https://doi.org/10.3897/zookeys.824.31635" volume="824" year="2019">Jantarit et al. 2019</bibRefCitation>
|
|
) present below sutural mac, probably inside the integument; a pair of short cephalic trichobothria, external and close to the middle of the head (Fig.
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Coecobrya whitteni sp. nov., continued A Distal part of manubrium ventrally B Trochanteral organ C Posterior side of ventral tube and Lateral flap D Dorsal cephalic chaetotaxy E Anterior side of ventral tube F Manubrium plaque G Clypeal chaetae." figureDoi="10.3897/subtbiol.41.76926.figure3" httpUri="https://binary.pensoft.net/fig/621481" pageId="0" pageNumber="1">3D</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Ventral head chaetotaxy</emphasis>
|
|
(Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Coecobrya whitteni sp. nov. A Ventro-distal complex of labrum B Outer maxillary lobe C Labial palp D Ant. III organ E Chaetae of labial basis and ventral chaetotaxy of head F Mucro G Distal part of tita III and claw complex H Prelabral and labral chaetae I Mandibles." figureDoi="10.3897/subtbiol.41.76926.figure2" httpUri="https://binary.pensoft.net/fig/621480" pageId="0" pageNumber="1">2E</figureCitation>
|
|
). Chaetae of labial basis all smooth (mrel1l2), chaetae m e and l1 subequal, r thin and shortest, and l2 longest. The r/m length ratio: 0.13-0.14 (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Coecobrya whitteni sp. nov. A Ventro-distal complex of labrum B Outer maxillary lobe C Labial palp D Ant. III organ E Chaetae of labial basis and ventral chaetotaxy of head F Mucro G Distal part of tita III and claw complex H Prelabral and labral chaetae I Mandibles." figureDoi="10.3897/subtbiol.41.76926.figure2" httpUri="https://binary.pensoft.net/fig/621480" pageId="0" pageNumber="1">2E</figureCitation>
|
|
). Postlabial chaetae X2, X and X4 smooth, acuminate and minute chaetae, of similar size, X3 normally absent but present as mic in one individual. On each side of the cephalic groove with 9 chaetae, of which, the proximal six always long and smooth, 7th and 9th as mics, 8th always long and ciliated, one specimen with three mics distally (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Coecobrya whitteni sp. nov. A Ventro-distal complex of labrum B Outer maxillary lobe C Labial palp D Ant. III organ E Chaetae of labial basis and ventral chaetotaxy of head F Mucro G Distal part of tita III and claw complex H Prelabral and labral chaetae I Mandibles." figureDoi="10.3897/subtbiol.41.76926.figure2" httpUri="https://binary.pensoft.net/fig/621480" pageId="0" pageNumber="1">2E</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Tergite chaetotaxy</emphasis>
|
|
(Figs
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Coecobrya whitteni sp. nov., continued A Chaetotaxy of dorsal Th. II- III B Chaetotaxy of dorsal Abd. I- III C Chaetotaxy of dorsal Abd. IV D Chaetotaxy of dorsal Abd. V." figureDoi="10.3897/subtbiol.41.76926.figure4" httpUri="https://binary.pensoft.net/fig/621482" pageId="0" pageNumber="1">4A-D</figureCitation>
|
|
). Th. II with two (m1, m2) medio-medial, one (m4) medio-sublateral and 18-20 posterior mac; 1+1 ms and 2+2 sens antero-laterally (Fig.
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Coecobrya whitteni sp. nov., continued A Chaetotaxy of dorsal Th. II- III B Chaetotaxy of dorsal Abd. I- III C Chaetotaxy of dorsal Abd. IV D Chaetotaxy of dorsal Abd. V." figureDoi="10.3897/subtbiol.41.76926.figure4" httpUri="https://binary.pensoft.net/fig/621482" pageId="0" pageNumber="1">4A</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
Th. III with 23-26 mac. 2+2 sens laterally (Fig.
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Coecobrya whitteni sp. nov., continued A Chaetotaxy of dorsal Th. II- III B Chaetotaxy of dorsal Abd. I- III C Chaetotaxy of dorsal Abd. IV D Chaetotaxy of dorsal Abd. V." figureDoi="10.3897/subtbiol.41.76926.figure4" httpUri="https://binary.pensoft.net/fig/621482" pageId="0" pageNumber="1">4A</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
Abd. I with six central mac (m4i, m2-4, m2i, m4p), 1+1 ms and 1+1 sens laterally (Fig.
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Coecobrya whitteni sp. nov., continued A Chaetotaxy of dorsal Th. II- III B Chaetotaxy of dorsal Abd. I- III C Chaetotaxy of dorsal Abd. IV D Chaetotaxy of dorsal Abd. V." figureDoi="10.3897/subtbiol.41.76926.figure4" httpUri="https://binary.pensoft.net/fig/621482" pageId="0" pageNumber="1">4B</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
Abd. II with two (m3, m3e) central and one (m5) lateral mac, 2+2 tric without modified chaetae, 1+1 sens laterally and 1+1 mic near internal tric (Fig.
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Coecobrya whitteni sp. nov., continued A Chaetotaxy of dorsal Th. II- III B Chaetotaxy of dorsal Abd. I- III C Chaetotaxy of dorsal Abd. IV D Chaetotaxy of dorsal Abd. V." figureDoi="10.3897/subtbiol.41.76926.figure4" httpUri="https://binary.pensoft.net/fig/621482" pageId="0" pageNumber="1">4B</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
Abd. III with one (m3) central and three (am6, pm6, p6) lateral mac. 3+3 tric without modified chaetae, 1+1 sens laterally, 1+1 mic near m3, ms not seen (Fig.
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Coecobrya whitteni sp. nov., continued A Chaetotaxy of dorsal Th. II- III B Chaetotaxy of dorsal Abd. I- III C Chaetotaxy of dorsal Abd. IV D Chaetotaxy of dorsal Abd. V." figureDoi="10.3897/subtbiol.41.76926.figure4" httpUri="https://binary.pensoft.net/fig/621482" pageId="0" pageNumber="1">4B</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
Abd. IV with four central mac (M, A5-6, B5) and six (D3, E2-4, E2p, F3) lateral mac, 2+2 tric and about 19 long S-like chaetae and one mic in the middle below psp, without modified chaetae (Fig.
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Coecobrya whitteni sp. nov., continued A Chaetotaxy of dorsal Th. II- III B Chaetotaxy of dorsal Abd. I- III C Chaetotaxy of dorsal Abd. IV D Chaetotaxy of dorsal Abd. V." figureDoi="10.3897/subtbiol.41.76926.figure4" httpUri="https://binary.pensoft.net/fig/621482" pageId="0" pageNumber="1">4C</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
Abd. V with at least 6 obvious mac and several mes to small mac, and 3+3 sens (Fig.
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Coecobrya whitteni sp. nov., continued A Chaetotaxy of dorsal Th. II- III B Chaetotaxy of dorsal Abd. I- III C Chaetotaxy of dorsal Abd. IV D Chaetotaxy of dorsal Abd. V." figureDoi="10.3897/subtbiol.41.76926.figure4" httpUri="https://binary.pensoft.net/fig/621482" pageId="0" pageNumber="1">4D</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">Abd. VI not analysed.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
S-chaetae formula from Th. II to Abd. V: 2+ms, 2/1+ms, 2, 2, ≈ 20, 3; ps not seen, as sens on Abd. IV 1/3 as long as S-like chaetae (Figs
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Coecobrya whitteni sp. nov., continued A Chaetotaxy of dorsal Th. II- III B Chaetotaxy of dorsal Abd. I- III C Chaetotaxy of dorsal Abd. IV D Chaetotaxy of dorsal Abd. V." figureDoi="10.3897/subtbiol.41.76926.figure4" httpUri="https://binary.pensoft.net/fig/621482" pageId="0" pageNumber="1">4A-D</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Legs</emphasis>
|
|
(Figs
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Coecobrya whitteni sp. nov. A Ventro-distal complex of labrum B Outer maxillary lobe C Labial palp D Ant. III organ E Chaetae of labial basis and ventral chaetotaxy of head F Mucro G Distal part of tita III and claw complex H Prelabral and labral chaetae I Mandibles." figureDoi="10.3897/subtbiol.41.76926.figure2" httpUri="https://binary.pensoft.net/fig/621480" pageId="0" pageNumber="1">2G</figureCitation>
|
|
,
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Coecobrya whitteni sp. nov., continued A Distal part of manubrium ventrally B Trochanteral organ C Posterior side of ventral tube and Lateral flap D Dorsal cephalic chaetotaxy E Anterior side of ventral tube F Manubrium plaque G Clypeal chaetae." figureDoi="10.3897/subtbiol.41.76926.figure3" httpUri="https://binary.pensoft.net/fig/621481" pageId="0" pageNumber="1">3B</figureCitation>
|
|
). Leg long; tita of leg III slightly longer than tita of legs I and II. Legs devoid of scales, covered with ordinary ciliated chaetae of various lengths, mic not seen. Trochanteral organ with 18-21 smooth, straight, unequal spine-like chaetae (Fig.
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Coecobrya whitteni sp. nov., continued A Distal part of manubrium ventrally B Trochanteral organ C Posterior side of ventral tube and Lateral flap D Dorsal cephalic chaetotaxy E Anterior side of ventral tube F Manubrium plaque G Clypeal chaetae." figureDoi="10.3897/subtbiol.41.76926.figure3" httpUri="https://binary.pensoft.net/fig/621481" pageId="0" pageNumber="1">3B</figureCitation>
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|
). The distal whorl of tita III with 10 subequal ciliated mes, irregularly arranged, and usually claw I-II with dorso-apical clavate tenent hair (pointed in two specimens). A smooth, thin and long chaeta close to tenent hair (sensu
|
|
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.824.31635" author="Jantarit, S" journalOrPublisher="Zookey" pageId="0" pageNumber="1" pagination="21 - 44" refId="B18" refString="Jantarit, S, Satasook, C, Deharveng, L, 2019. Coecobrya sirindhornae sp. n., the most highly troglomorphic Collembola in Southeast Asia (Collembola, Entomobryidae). Zookey 824: 21 - 44, DOI: https://doi.org/10.3897/zookeys.824.31635" title="Coecobrya sirindhornae sp. n., the most highly troglomorphic Collembola in Southeast Asia (Collembola, Entomobryidae)." url="https://doi.org/10.3897/zookeys.824.31635" volume="824" year="2019">Jantarit et al. 2019</bibRefCitation>
|
|
) absent. Claw III generally with dorso-apical clavate tenent hair (8 individuals with clavate tenant hair, 4 individuals with pointed tenant hair; holotype with pointed tenent hair Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Coecobrya whitteni sp. nov. A Ventro-distal complex of labrum B Outer maxillary lobe C Labial palp D Ant. III organ E Chaetae of labial basis and ventral chaetotaxy of head F Mucro G Distal part of tita III and claw complex H Prelabral and labral chaetae I Mandibles." figureDoi="10.3897/subtbiol.41.76926.figure2" httpUri="https://binary.pensoft.net/fig/621480" pageId="0" pageNumber="1">2G</figureCitation>
|
|
). Ventro-distal smooth chaeta of tita III thick, erected, pointed, rather short. Pretarsal mic minute (2.5-3.0
|
|
<normalizedToken originalValue="µm">µm</normalizedToken>
|
|
). Claw slender and elongated. Unguis of all claw with one small inner teeth at 53 % and a pair of unequal basal teeth at about 44-49 % of inner edge from basis. Unguiculus approximately 2/3 as long as inner edge of claw, rather swollen baso-externally, pointed apically, devoid of inner tooth, with at least five minute outer teeth, often inconspicuous, at 3/4 of its length (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Coecobrya whitteni sp. nov. A Ventro-distal complex of labrum B Outer maxillary lobe C Labial palp D Ant. III organ E Chaetae of labial basis and ventral chaetotaxy of head F Mucro G Distal part of tita III and claw complex H Prelabral and labral chaetae I Mandibles." figureDoi="10.3897/subtbiol.41.76926.figure2" httpUri="https://binary.pensoft.net/fig/621480" pageId="0" pageNumber="1">2G</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Ventral tube</emphasis>
|
|
(Figs
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Coecobrya whitteni sp. nov., continued A Distal part of manubrium ventrally B Trochanteral organ C Posterior side of ventral tube and Lateral flap D Dorsal cephalic chaetotaxy E Anterior side of ventral tube F Manubrium plaque G Clypeal chaetae." figureDoi="10.3897/subtbiol.41.76926.figure3" httpUri="https://binary.pensoft.net/fig/621481" pageId="0" pageNumber="1">3C, E</figureCitation>
|
|
). Ventral tube two to three times longer than wide. Lateral flaps with 6-7+6-7 smooth chaetae except for one specimen with one ciliated chaeta present in both sides (Fig.
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Coecobrya whitteni sp. nov., continued A Distal part of manubrium ventrally B Trochanteral organ C Posterior side of ventral tube and Lateral flap D Dorsal cephalic chaetotaxy E Anterior side of ventral tube F Manubrium plaque G Clypeal chaetae." figureDoi="10.3897/subtbiol.41.76926.figure3" httpUri="https://binary.pensoft.net/fig/621481" pageId="0" pageNumber="1">3C</figureCitation>
|
|
). Anterior face with 6-8+6-8 chaetae, 2(3) of them larger than others, all ciliated, arranged roughly asymmetrically (Fig.
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Coecobrya whitteni sp. nov., continued A Distal part of manubrium ventrally B Trochanteral organ C Posterior side of ventral tube and Lateral flap D Dorsal cephalic chaetotaxy E Anterior side of ventral tube F Manubrium plaque G Clypeal chaetae." figureDoi="10.3897/subtbiol.41.76926.figure3" httpUri="https://binary.pensoft.net/fig/621481" pageId="0" pageNumber="1">3E</figureCitation>
|
|
); posterior face with at least 12 chaetae, four apical chaetae longer and larger than others and about 8-13 proximal chaetae; two straight smooth chaetae and 6-11 either small finely ciliated or mics arranged asymmetrically (Fig.
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Coecobrya whitteni sp. nov., continued A Distal part of manubrium ventrally B Trochanteral organ C Posterior side of ventral tube and Lateral flap D Dorsal cephalic chaetotaxy E Anterior side of ventral tube F Manubrium plaque G Clypeal chaetae." figureDoi="10.3897/subtbiol.41.76926.figure3" httpUri="https://binary.pensoft.net/fig/621481" pageId="0" pageNumber="1">3C</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Furcal complex</emphasis>
|
|
(Figs
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Coecobrya whitteni sp. nov. A Ventro-distal complex of labrum B Outer maxillary lobe C Labial palp D Ant. III organ E Chaetae of labial basis and ventral chaetotaxy of head F Mucro G Distal part of tita III and claw complex H Prelabral and labral chaetae I Mandibles." figureDoi="10.3897/subtbiol.41.76926.figure2" httpUri="https://binary.pensoft.net/fig/621480" pageId="0" pageNumber="1">2F</figureCitation>
|
|
,
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Coecobrya whitteni sp. nov., continued A Distal part of manubrium ventrally B Trochanteral organ C Posterior side of ventral tube and Lateral flap D Dorsal cephalic chaetotaxy E Anterior side of ventral tube F Manubrium plaque G Clypeal chaetae." figureDoi="10.3897/subtbiol.41.76926.figure3" httpUri="https://binary.pensoft.net/fig/621481" pageId="0" pageNumber="1">3A, F</figureCitation>
|
|
). Tenaculum with four large teeth of decreasing size from the basal to the distal one of each ramus, on a prominent, irregular body, with a postero-basal strong serrated chaeta bent distally. Mucrodens 1.6 times longer than manubrium. Furcula without smooth chaetae. Manubrium with a dense cover of ciliated chaetae both dorsally and ventrally. Manubrial plaque with 2+2 psp and three to six ciliated chaetae (usually 5+5) (Fig.
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Coecobrya whitteni sp. nov., continued A Distal part of manubrium ventrally B Trochanteral organ C Posterior side of ventral tube and Lateral flap D Dorsal cephalic chaetotaxy E Anterior side of ventral tube F Manubrium plaque G Clypeal chaetae." figureDoi="10.3897/subtbiol.41.76926.figure3" httpUri="https://binary.pensoft.net/fig/621481" pageId="0" pageNumber="1">3F</figureCitation>
|
|
). Distal part of manubrium ventrally with 13+13 ciliate chaetae (Fig.
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Coecobrya whitteni sp. nov., continued A Distal part of manubrium ventrally B Trochanteral organ C Posterior side of ventral tube and Lateral flap D Dorsal cephalic chaetotaxy E Anterior side of ventral tube F Manubrium plaque G Clypeal chaetae." figureDoi="10.3897/subtbiol.41.76926.figure3" httpUri="https://binary.pensoft.net/fig/621481" pageId="0" pageNumber="1">3A</figureCitation>
|
|
). Dens without spines, annulated and covered with ciliated chaetae on both sides. Distal smooth part of dens slightly longer than mucro. Mucro strong and falcate, basal spine long, reaching the tip of the mucronal tooth (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Coecobrya whitteni sp. nov. A Ventro-distal complex of labrum B Outer maxillary lobe C Labial palp D Ant. III organ E Chaetae of labial basis and ventral chaetotaxy of head F Mucro G Distal part of tita III and claw complex H Prelabral and labral chaetae I Mandibles." figureDoi="10.3897/subtbiol.41.76926.figure2" httpUri="https://binary.pensoft.net/fig/621480" pageId="0" pageNumber="1">2F</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Genital plate.</emphasis>
|
|
Female genital plate with 2+2 genital mic, male genital plate not clearly seen.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="ecology">
|
|
<paragraph pageId="0" pageNumber="1">Ecology.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<taxonomicName authorityName="Nilsai & Jantarit" authorityYear="2021" class="Collembola" family="Entomobryidae" genus="Coecobrya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coecobrya whitteni" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="whitteni">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Coecobrya whitteni</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. was found in the dark zone of a cave on the surface of the muddy ground, near the bank of a stream. It was also found in the upper levels on stalagmites, clay, gravel and rock surfaces. This species was found mainly in eutrophic habitats where piles of guano were well-presented. There was a big colony of the intermediate roundleaf bat (
|
|
<taxonomicName baseAuthorityName="Horsfield" baseAuthorityYear="1823" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" lsidName="Hipposideros larvatus" order="Chiroptera" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="larvatus">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Hipposideros larvatus</emphasis>
|
|
</taxonomicName>
|
|
) inside the cave. The air temperature of the habitat where this species was collected was 22.9-24.0 °C and the relative humidity was 85%. In the same cave, and in the same chamber, at least six different stygobitic species were also observed in the small puddles:
|
|
<taxonomicName authorityName="Botosaneanu & Stock" authorityYear="1989" class="Malacostraca" family="Bogidiellidae" genus="Aequigidiella" kingdom="Animalia" lsidName="Aequigidiella aquilifera" order="Amphipoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="aquilifera">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Aequigidiella aquilifera</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName authorityName="Kawakatsu & Mitchell" authorityYear="1989" class="Magnoliopsida" family="Asteraceae" genus="Dugesia" kingdom="Plantae" lsidName="Dugesia deharvengi" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="deharvengi">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Dugesia deharvengi</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName baseAuthorityName="Yamaguchi" baseAuthorityYear="1953" class="Clitellata" family="Haplotaxidae" genus="Heterochaetella" kingdom="Animalia" lsidName="Heterochaetella glandularis" order="Haplotaxida" pageId="0" pageNumber="1" phylum="Annelida" rank="species" species="glandularis">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Heterochaetella glandularis</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName authorityName="Spangler" authorityYear="1996" class="Insecta" family="Dytiscidae" genus="Siamoporus" kingdom="Animalia" lsidName="Siamoporus deharvengi" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="deharvengi">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Siamoporus deharvengi</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName authorityName="Magniez" authorityYear="1991" class="Malacostraca" family="Stenasellidae" genus="Stenasellus" kingdom="Animalia" lsidName="Stenasellus rigali" order="Isopoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="rigali">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Stenasellus rigali</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName authorityName="Cals & Boutin" authorityYear="1985" class="Malacostraca" family="Halosbaenidae" genus="Theosbaena" kingdom="Animalia" lsidName="Theosbaena cambodjiana" order="Thermosbaenacea" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="cambodjiana">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Theosbaena cambodjiana</emphasis>
|
|
</taxonomicName>
|
|
. The co-occurrence of these six species was first observed in 1987 (
|
|
<bibRefCitation author="Association Pyreneenne de Speleologie" journalOrPublisher="Association Pyreneenne de Speleologie, Toulouse" pageId="0" pageNumber="1" refId="B1" refString="Association Pyreneenne de Speleologie, 1988. Expeditions Thai 87-Thai 88: Rapport Speleologique et Scientifique. Association Pyreneenne de Speleologie, Toulouse" title="Expeditions Thai 87 - Thai 88: Rapport Speleologique et Scientifique." year="1988">
|
|
Association
|
|
<normalizedToken originalValue="Pyrénéenne">Pyreneenne</normalizedToken>
|
|
de
|
|
<normalizedToken originalValue="Spéléologie">Speleologie</normalizedToken>
|
|
1988
|
|
</bibRefCitation>
|
|
;
|
|
<bibRefCitation author="Deharveng, L" editor="Wilkens, H" journalOrPublisher="Amsterdam Elsevier" pageId="0" pageNumber="1" pagination="603 - 632" refId="B9" refString="Deharveng, L, Bedos, A, 2000. The cave fauna of Southeast Asia: ecology, origin, evolution. In: Wilkens, H, Culver, DC, Humphreys, WF, Eds., Ecosystem of the World 30: Subterranean Ecosystems. Amsterdam Elsevier: 603 - 632" title="The cave fauna of Southeast Asia: ecology, origin, evolution." volumeTitle="Ecosystem of the World 30: Subterranean Ecosystems." year="2000">Deharveng and Bedos 2000</bibRefCitation>
|
|
,
|
|
<bibRefCitation DOI="https://doi.org/10.1016/B978-0-12-383832-2.00032-3" author="Deharveng, L" editor="White, WB" journalOrPublisher="Academic Press, Chennai" pageId="0" pageNumber="1" pagination="238 - 250" refId="B10" refString="Deharveng, L, Bedos, A, 2012. Diversity patterns in the tropics. In: White, WB, Culver, DC, Eds., Encyclopedia of Caves. Academic Press, Chennai: 238 - 250, DOI: https://doi.org/10.1016/B978-0-12-383832-2.00032-3" title="Diversity patterns in the tropics." url="https://doi.org/10.1016/B978-0-12-383832-2.00032-3" volumeTitle="Encyclopedia of Caves." year="2012">2012</bibRefCitation>
|
|
) and from our observations they are all still present in rather large numbers of individuals. We also found a millipede (
|
|
<taxonomicName authorityName="Golovatch, Geoffroy, Mauries & VandenSpiegel" authorityYear="2011" class="Diplopoda" family="Cambalopsidae" genus="Plusioglyphiulus" kingdom="Animalia" lsidName="Plusioglyphiulus saksit" order="Spirostreptida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="saksit">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Plusioglyphiulus saksit</emphasis>
|
|
</taxonomicName>
|
|
) and a spider (
|
|
<taxonomicName authorityName="Deeleman-Reinhold" authorityYear="1995" class="Arachnida" family="Ochyroceratidae" genus="Speocera" kingdom="Animalia" lsidName="Speocera deharvengi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="deharvengi">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Speocera deharvengi</emphasis>
|
|
</taxonomicName>
|
|
) in the same habitat where this new species was found. Most of these observed species only known as endemics to this cave. The discovery of this new species in Tham Nayn Noi emphasizes the importance of this cave in harboring the endemic subterranean fauna of the area.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="etymology">
|
|
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<taxonomicName authorityName="Nilsai & Jantarit" authorityYear="2021" class="Collembola" family="Entomobryidae" genus="Coecobrya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coecobrya whitteni" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="whitteni">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Coecobrya whitteni</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. is named in honor of the late Tony Whitten in appreciation of his enormous contributions to the discovery and conservation of karst and cave invertebrates across Asia.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="remarks">
|
|
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
Among the described troglomorphic
|
|
<taxonomicName authorityName="Yosii" authorityYear="1956" class="Collembola" family="Entomobryidae" genus="Coecobrya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coecobrya" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Coecobrya</emphasis>
|
|
</taxonomicName>
|
|
species in Thailand,
|
|
<taxonomicName authorityName="Nilsai & Jantarit" authorityYear="2021" class="Collembola" family="Entomobryidae" genus="Coecobrya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coecobrya whitteni" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="whitteni">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Coecobrya whitteni</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. is near to
|
|
<taxonomicName class="Collembola" genus="C." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. cavicta" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="cavicta">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">C. cavicta</emphasis>
|
|
</taxonomicName>
|
|
Nilsai & Zhang, 2017 from Satun province, southern Thailand in the body length, dorsal head chaetotaxy, labial chaetae, number of sublobal hairs on maxillary outer lobe, number of mac on Abd.III and number of chaetae on ventrodistal part of manubrium. However, it differs from
|
|
<taxonomicName class="Collembola" genus="C." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. cavicta" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="cavicta">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">C. cavicta</emphasis>
|
|
</taxonomicName>
|
|
by the combination of various characters i.e. longer length of antennae, number of mac of dorsal tergits on Th. II (21-23 vs. 36-37), Th. III (23-26 vs. 35), Abd. II (2 vs. 3) and Abd. IV (11 vs. 18-20), number of inner teeth of claw (3 vs. 2) and number of smooth chaetae of trochanteral organe (18-21 vs. 15-16) (see Table
|
|
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Comparison of troglomorphic Coecobrya in Thailand: C. chumphonensis Zhang & Nilsai, 2017, C. cavicta Nilsai & Zhang, 2017, C. polychaeta Zhang & Nilsai, 2017, C. sirindhornae Jantarit, Satasook & Deharveng, 2019, C. nupa Christiansen & Bellinger, 1992. C. whitteni Nilsai and Jantarit sp. nov., C. troglobia Jantarit and Nilsai sp. nov., C. ellisi Jantarit and Nilsai sp. nov., C. phitsanulokensis Jantarit and Nilsai sp. nov. Key: c = ciliated chaetae, s = smooth chaetae,? = not given in literature description. Characters indicated in parentheses are rarely observed." httpUri="http://table.plazi.org/id/5378ECD9BC57B445F8A5FA2E10CEA7D7" pageId="0" pageNumber="1" tableUuid="5378ECD9BC57B445F8A5FA2E10CEA7D7">1</tableCitation>
|
|
). In fact,
|
|
<taxonomicName authorityName="Nilsai & Jantarit" authorityYear="2021" class="Collembola" family="Entomobryidae" genus="Coecobrya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coecobrya whitteni" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="whitteni">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Coecobrya whitteni</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. is most similar to
|
|
<taxonomicName class="Collembola" genus="C." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. troglobia" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="troglobia">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">C. troglobia</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. which is described in this work since the caves where these two species are found are only 3.4 km apart in a straight line, separated by limestone mountains. The altitudes of the two caves differs by about 200 m. The two species are similar in body size, antennal length, number of sublobal hair on the maxillary outer lobe, number of chaetae on Gr. II, labial chaetae, number of mac on Abd. I-IV, number of inner teeth of the claw, tenent hair, similar number in ventral tube chaetae both anterior and posterior side as well as a lateral flap, number of chaetae on trochanteral organ (Table
|
|
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Comparison of troglomorphic Coecobrya in Thailand: C. chumphonensis Zhang & Nilsai, 2017, C. cavicta Nilsai & Zhang, 2017, C. polychaeta Zhang & Nilsai, 2017, C. sirindhornae Jantarit, Satasook & Deharveng, 2019, C. nupa Christiansen & Bellinger, 1992. C. whitteni Nilsai and Jantarit sp. nov., C. troglobia Jantarit and Nilsai sp. nov., C. ellisi Jantarit and Nilsai sp. nov., C. phitsanulokensis Jantarit and Nilsai sp. nov. Key: c = ciliated chaetae, s = smooth chaetae,? = not given in literature description. Characters indicated in parentheses are rarely observed." httpUri="http://table.plazi.org/id/5378ECD9BC57B445F8A5FA2E10CEA7D7" pageId="0" pageNumber="1" tableUuid="5378ECD9BC57B445F8A5FA2E10CEA7D7">1</tableCitation>
|
|
). However,
|
|
<taxonomicName authorityName="Nilsai & Jantarit" authorityYear="2021" class="Collembola" family="Entomobryidae" genus="Coecobrya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coecobrya whitteni" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="whitteni">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Coecobrya whitteni</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. differs from
|
|
<taxonomicName class="Collembola" genus="C." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. troglobia" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="troglobia">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">C. troglobia</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. by the number of An dorsal mac on the head (4 vs. 5) and number of chaetae on Th. II (medio-medial mac = 2 vs. 3, medio-sublateral mac = 1 vs. 4) (Table
|
|
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Comparison of troglomorphic Coecobrya in Thailand: C. chumphonensis Zhang & Nilsai, 2017, C. cavicta Nilsai & Zhang, 2017, C. polychaeta Zhang & Nilsai, 2017, C. sirindhornae Jantarit, Satasook & Deharveng, 2019, C. nupa Christiansen & Bellinger, 1992. C. whitteni Nilsai and Jantarit sp. nov., C. troglobia Jantarit and Nilsai sp. nov., C. ellisi Jantarit and Nilsai sp. nov., C. phitsanulokensis Jantarit and Nilsai sp. nov. Key: c = ciliated chaetae, s = smooth chaetae,? = not given in literature description. Characters indicated in parentheses are rarely observed." httpUri="http://table.plazi.org/id/5378ECD9BC57B445F8A5FA2E10CEA7D7" pageId="0" pageNumber="1" tableUuid="5378ECD9BC57B445F8A5FA2E10CEA7D7">1</tableCitation>
|
|
).
|
|
<taxonomicName authorityName="Nilsai & Jantarit" authorityYear="2021" class="Collembola" family="Entomobryidae" genus="Coecobrya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coecobrya whitteni" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="whitteni">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Coecobrya whitteni</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. has in fact already been mentioned by
|
|
<bibRefCitation author="Deharveng, L" editor="Wilkens, H" journalOrPublisher="Amsterdam Elsevier" pageId="0" pageNumber="1" pagination="603 - 632" refId="B9" refString="Deharveng, L, Bedos, A, 2000. The cave fauna of Southeast Asia: ecology, origin, evolution. In: Wilkens, H, Culver, DC, Humphreys, WF, Eds., Ecosystem of the World 30: Subterranean Ecosystems. Amsterdam Elsevier: 603 - 632" title="The cave fauna of Southeast Asia: ecology, origin, evolution." volumeTitle="Ecosystem of the World 30: Subterranean Ecosystems." year="2000">Deharveng and Bedos (2000)</bibRefCitation>
|
|
as an undescribed cf.
|
|
<taxonomicName authorityName="Yosii" authorityYear="1956" class="Collembola" family="Entomobryidae" genus="Coecobrya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coecobrya" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Coecobrya</emphasis>
|
|
</taxonomicName>
|
|
species (Fig. 31.8A page 625) from oligotrophic habitats. (Note that in this reference the cave is misnamed as Tham Kubio).
|
|
</paragraph>
|
|
<caption ID-Table-UUID="5378ECD9BC57B445F8A5FA2E10CEA7D7" httpUri="http://table.plazi.org/id/5378ECD9BC57B445F8A5FA2E10CEA7D7" pageId="0" pageNumber="1" start="Table 1" startId="T1">
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="1">Table 1.</emphasis>
|
|
Comparison of troglomorphic
|
|
<taxonomicName authorityName="Yosii" authorityYear="1956" class="Collembola" family="Entomobryidae" genus="Coecobrya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coecobrya" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Coecobrya</emphasis>
|
|
</taxonomicName>
|
|
in Thailand:
|
|
<taxonomicName class="Collembola" genus="C." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. chumphonensis" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="chumphonensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">C. chumphonensis</emphasis>
|
|
</taxonomicName>
|
|
Zhang & Nilsai, 2017,
|
|
<taxonomicName class="Collembola" genus="C." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. cavicta" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="cavicta">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">C. cavicta</emphasis>
|
|
</taxonomicName>
|
|
Nilsai & Zhang, 2017,
|
|
<taxonomicName class="Collembola" genus="C." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. polychaeta" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="polychaeta">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">C. polychaeta</emphasis>
|
|
</taxonomicName>
|
|
Zhang & Nilsai, 2017,
|
|
<taxonomicName class="Collembola" genus="C." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. sirindhornae" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sirindhornae">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">C. sirindhornae</emphasis>
|
|
</taxonomicName>
|
|
Jantarit, Satasook & Deharveng, 2019,
|
|
<taxonomicName class="Collembola" genus="C." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. nupa" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nupa">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">C. nupa</emphasis>
|
|
</taxonomicName>
|
|
Christiansen & Bellinger, 1992.
|
|
<taxonomicName class="Collembola" genus="C." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. whitteni" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="whitteni">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">C. whitteni</emphasis>
|
|
</taxonomicName>
|
|
Nilsai and Jantarit sp. nov.,
|
|
<taxonomicName class="Collembola" genus="C." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. troglobia" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="troglobia">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">C. troglobia</emphasis>
|
|
</taxonomicName>
|
|
Jantarit and Nilsai sp. nov.,
|
|
<taxonomicName class="Collembola" genus="C." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. ellisi" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ellisi">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">C. ellisi</emphasis>
|
|
</taxonomicName>
|
|
Jantarit and Nilsai sp. nov.,
|
|
<taxonomicName class="Collembola" genus="C." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. phitsanulokensis" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="phitsanulokensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">C. phitsanulokensis</emphasis>
|
|
</taxonomicName>
|
|
Jantarit and Nilsai sp. nov. Key: c = ciliated chaetae, s = smooth chaetae,? = not given in literature description. Characters indicated in parentheses are rarely observed.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<table pageId="0" pageNumber="1">
|
|
<tr pageId="0" pageNumber="1">
|
|
<th colspan="2" pageId="0" pageNumber="1" rowspan="1">Characters/species</th>
|
|
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
|
<taxonomicName class="Collembola" genus="C." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. chumphonensis" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="chumphonensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">C. chumphonensis</emphasis>
|
|
</taxonomicName>
|
|
</th>
|
|
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
|
<taxonomicName class="Collembola" genus="C." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. cavicta" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="cavicta">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">C. cavicta</emphasis>
|
|
</taxonomicName>
|
|
</th>
|
|
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
|
<taxonomicName class="Collembola" genus="C." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. polychaeta" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="polychaeta">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">C. polychaeta</emphasis>
|
|
</taxonomicName>
|
|
</th>
|
|
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
|
<taxonomicName class="Collembola" genus="C." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. sirindhornae" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sirindhornae">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">C. sirindhornae</emphasis>
|
|
</taxonomicName>
|
|
</th>
|
|
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
|
<taxonomicName class="Collembola" genus="C." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. nupa" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nupa">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">C. nupa</emphasis>
|
|
</taxonomicName>
|
|
</th>
|
|
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
|
<taxonomicName class="Collembola" genus="C." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. whitteni" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="whitteni">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">C. whitteni</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.
|
|
</th>
|
|
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
|
<taxonomicName class="Collembola" genus="C." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. troglobia" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="troglobia">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">C. troglobia</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.
|
|
</th>
|
|
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
|
<taxonomicName class="Collembola" genus="C." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. ellisi" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ellisi">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">C. ellisi</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.
|
|
</th>
|
|
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
|
<taxonomicName class="Collembola" genus="C." higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="C. phitsanulokensis" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="phitsanulokensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">C. phitsanulokensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.
|
|
</th>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="2" pageId="0" pageNumber="1" rowspan="1">Body length</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">up to 2.82 mm</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.72 mm</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">up to 2.58 mm</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">up to 2.6 mm</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.0 mm</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.8-2.3 mm</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.0-2.4 mm</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.6-2.1 mm</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.8-2.3 mm</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="2" pageId="0" pageNumber="1" rowspan="1">Ant./head ratio</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3.70-4.48</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.67</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">5.91-7.12</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">8.0-12.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">6.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3.78-5.14</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">4.0-6.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3.5-6.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3.1-4.2</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="2" pageId="0" pageNumber="1" rowspan="1">Long smooth straight chaetae on Ant. I</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">absent</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">present</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">absent</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">present</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">?</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">present</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">present</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">present</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">present</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="2" pageId="0" pageNumber="1" rowspan="1">No. of paddle-like chaetae on Ant. II</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2-4</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">?</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">10-12</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">?</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">not seen</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">not seen</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">not seen</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">not seen</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="11" pageId="0" pageNumber="1" rowspan="1">Number of chaetae on clypeus</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="1" isEmpty="true" pageId="0" pageNumber="1" rowspan="1">-</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Prefrontal area</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3s</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">?</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3s</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3s</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">?</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3s</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3s</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3s</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3s</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="1" isEmpty="true" pageId="0" pageNumber="1" rowspan="1">-</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Facial area</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">8s</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">?</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2s; 7-10c</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2s; 7-10c</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">?</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2s; 6c</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2s; 4c</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2s; 5c</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2s; 4-7c</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="11" pageId="0" pageNumber="1" rowspan="1">Number of chaetae on dorsal head</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="1" isEmpty="true" pageId="0" pageNumber="1" rowspan="1">-</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">An area</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">4</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">4</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">?</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">4</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">5</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">5</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="1" isEmpty="true" pageId="0" pageNumber="1" rowspan="1">-</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">M series</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">4</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">?</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="1" isEmpty="true" pageId="0" pageNumber="1" rowspan="1">-</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Group II (Gr. II)</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">6(7)</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">4</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">?</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">4</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">4</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">4(5)</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">4(5)</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="2" pageId="0" pageNumber="1" rowspan="1">A0</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">mac</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">mac</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">mic</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">mes</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">?</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">mac</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">mac</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">mac</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">mac</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="2" pageId="0" pageNumber="1" rowspan="1">Number of sublobal hairs on maxillary outer lobe</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">4</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">4</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">4</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">4</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="2" pageId="0" pageNumber="1" rowspan="1">Lateral process of labial palp</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">short</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">short</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">long</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">long</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">short</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">long</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">long</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">long</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">long</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="2" pageId="0" pageNumber="1" rowspan="1">Labial chaetae shape</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">mRel1l2</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">mrel1l2</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">M1m2rel1l2</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">m1m2rel1l2</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">M1m2rel1l2</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">mrel1l2</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">mrel1l2</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">m(M)1m2rel1l2</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">m1m2rel1l2</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="2" pageId="0" pageNumber="1" rowspan="1">Postlabial chaetae X</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">minute</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">minute</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">normal</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">normal</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">?</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">minute</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">minute</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">normal</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">minute</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="2" pageId="0" pageNumber="1" rowspan="1">Number of chaetae along cephalic groove</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">4s; 5-7c</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3-4s; 7-8c</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">6-7s; 5-10c</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">7-9s; 3-7c</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">?</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">5-6s; 1-2c1-2mics</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">6s; 1c; 2mics</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2-5s; 1-5c; 2mics</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">4-7s; 1-2c; 1-2(3-4)mics</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="11" pageId="0" pageNumber="1" rowspan="1">Chaetotaxy of Th. II</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="1" isEmpty="true" pageId="0" pageNumber="1" rowspan="1">-</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Number of medio-medial mac</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">4-6</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">7(6)</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="1" isEmpty="true" pageId="0" pageNumber="1" rowspan="1">-</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Number of medio-sublateral mac</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">4</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">4</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="1" isEmpty="true" pageId="0" pageNumber="1" rowspan="1">-</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Number of posterior mac</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">25-32</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">30-31</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">29-40</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">15-18</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">15?</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">18-20</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">15-18</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">28-31</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">31-33</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="2" pageId="0" pageNumber="1" rowspan="1">Number of mac on Th. III</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">32-35</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">35</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">35-43</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">32-35</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">15?</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">23-26</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">24-27</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">29-36</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">32-33</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="2" pageId="0" pageNumber="1" rowspan="1">Number of mac on Abd. I</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">6-7</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">6-7</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">8-9</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">6</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">4</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">6</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">6</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">7</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">7</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="2" pageId="0" pageNumber="1" rowspan="1">Number of central mac on Abd. II</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">4(3)</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3(4)</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="11" pageId="0" pageNumber="1" rowspan="1">Chaetotaxy of Abd. III</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="1" isEmpty="true" pageId="0" pageNumber="1" rowspan="1">-</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Number of central mac</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="1" isEmpty="true" pageId="0" pageNumber="1" rowspan="1">-</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Number of lateral mac</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2?</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="1" isEmpty="true" pageId="0" pageNumber="1" rowspan="1">-</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">ms</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">not seen</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">not seen</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">present</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">not seen</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">?</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">not seen</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">not seen</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">present</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">present</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="11" pageId="0" pageNumber="1" rowspan="1">Chaetotaxy of Abd. IV</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="1" isEmpty="true" pageId="0" pageNumber="1" rowspan="1">-</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Number of central mac</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">7</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">7-9</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">6</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">6</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">4</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">5</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">5</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">8</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">9</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="1" isEmpty="true" pageId="0" pageNumber="1" rowspan="1">-</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Number of lateral mac</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">10-12</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">11</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">9</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">8</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">?</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">6</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">6</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">9</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">8</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="11" pageId="0" pageNumber="1" rowspan="1">Ventral tube chaetae</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="1" isEmpty="true" pageId="0" pageNumber="1" rowspan="1">-</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">anterior face</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">9-12c</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">?</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">12c</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">10c</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">?</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">6-8c</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">7-8c</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">6-7c</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">6-7c</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="1" isEmpty="true" pageId="0" pageNumber="1" rowspan="1">-</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">posterior face</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">13</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">?</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">20-31</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">20-30</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">7</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">8-13</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">14-18</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">12-13</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">12-15</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="1" isEmpty="true" pageId="0" pageNumber="1" rowspan="1">-</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">lateral flap</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">7(10)</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">?</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">9-12</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">7-8s (5c)</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">6</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">6-7s (1c)</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">7-9s (1-2c)</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">6-7s; 1-3c</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">7-8s</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="2" pageId="0" pageNumber="1" rowspan="1">Tenent hair shape</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">usually pointed</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">pointed</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">pointed</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">pointed</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">pointed</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">clavate/pointed</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">clavate/pointed</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">pointed</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">pointed</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="2" pageId="0" pageNumber="1" rowspan="1">Number of ungual inner teeth</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="2" pageId="0" pageNumber="1" rowspan="1">Unguiculus outer edge shape</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">serrate</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">serrate</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">serrate</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2-3 teeth</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">?</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">serrate</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">serrate</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">?</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">serrate</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="2" pageId="0" pageNumber="1" rowspan="1">Smooth chaetae on trochanteral organ</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">12-22</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">15-16</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">15-25</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">12-18</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">16</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">18-21</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">19-23</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">17-23</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">19</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="2" pageId="0" pageNumber="1" rowspan="1">Chaetae on manubrial plaque</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">4-7</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">4</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">4-10</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">5</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3-6c</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">5c</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">5-6c</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">5-6c</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="2" pageId="0" pageNumber="1" rowspan="1">Chaetae on ventrodistal part of manubrium</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">11-15c</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">13c</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">15-25c</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">8-10c</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">?</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">13c</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">?</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">10-12c</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">13+13c</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="2" pageId="0" pageNumber="1" rowspan="1">Mucronal spine</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">nearly reaching mucronal apex</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">nearly reaching mucronal apex</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">nearly reaching mucronal apex</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">nearly reaching mucronal apex</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">beyond mucronal apex</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">nearly reaching mucronal apex</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">nearly reaching mucronal apex</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">nearly reaching mucronal apex</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">nearly reaching mucronal apex</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="1">
|
|
<td colspan="2" pageId="0" pageNumber="1" rowspan="1">Locality</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Chumphon, Thailand</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Satun, Thailand</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Satun, Thailand</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Satun, Thailand</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Maui, Hawaii</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Khon Kaen, Thailand</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Khon Kaen, Thailand</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Phetchabun, Thailand</td>
|
|
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Phitsanulok, Thailand</td>
|
|
</tr>
|
|
</table>
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |