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<document id="5F5A329DEF0BAC21E563C36D049305CF" ID="10.11646/zootaxa.4163.1.1" ID-DOI="10.11646/zootaxa.4163.1.1" ID-GBIF-Dataset="69b1046e-47bc-4f26-b32c-97d79da7a35d" ID-ISSN="1175-5326" ID-Zenodo-Dep="259278" ID-ZooBank="930AF345-15AD-443A-A3AE-328178ECB6C9" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1473747460051" checkinUser="plazi" docAuthor="Pamplona, Denise, Nihei, Silvio S., Couri, Márcia S. &amp; Pont, Adrian C." docDate="2016" docId="5F52879A4941FF9AA2951CD2FCA5F984" docLanguage="en" docName="zootaxa.4163.1.1.pdf" docOrigin="Zootaxa 4163 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Morellia (Morellia) quadriremis van Emden 1965" docType="treatment" docVersion="8" lastPageNumber="78" masterDocId="A36BFFE2490DFFD7A2021921FFC5FF8A" masterDocTitle="Taxonomy of Morellia Robineau-Desvoidy (Diptera: Muscidae): revision of the subgenera Morellia s. str. and Parapyrellia Townsend" masterLastPageNumber="110" masterPageNumber="1" pageNumber="77" updateTime="1698385354984" updateUser="plazi">
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<mods:title id="7547CCC94842037FC19ACB24C5E4132A">Taxonomy of Morellia Robineau-Desvoidy (Diptera: Muscidae): revision of the subgenera Morellia s. str. and Parapyrellia Townsend</mods:title>
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<mods:namePart id="EAE656494504CC2860DB3DD66AC3F92B">Pamplona, Denise</mods:namePart>
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<mods:namePart id="F87C70CA7FEF504246F23179CF40577D">Nihei, Silvio S.</mods:namePart>
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<mods:namePart id="6CAB4FD982941559D984AFB2E98A08B1">Couri, Márcia S.</mods:namePart>
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<treatment id="5F52879A4941FF9AA2951CD2FCA5F984" ID-DOI="http://doi.org/10.5281/zenodo.4547446" ID-GBIF-Taxon="124548457" ID-Zenodo-Dep="4547446" LSID="urn:lsid:plazi:treatment:5F52879A4941FF9AA2951CD2FCA5F984" httpUri="http://treatment.plazi.org/id/5F52879A4941FF9AA2951CD2FCA5F984" lastPageId="77" lastPageNumber="78" pageId="76" pageNumber="77">
<subSubSection id="9FE165074941FF9BA2951CD2FD25F987" box="[151,736,1522,1549]" pageId="76" pageNumber="77" type="nomenclature">
<paragraph id="D744368C4941FF9BA2951CD2FD25F987" blockId="76.[151,736,1522,1582]" box="[151,736,1522,1549]" pageId="76" pageNumber="77">
<heading id="8C0C81E04941FF9BA2951CD2FD25F987" bold="true" box="[151,736,1522,1549]" fontSize="11" level="1" pageId="76" pageNumber="77" reason="1">
<emphasis id="E58FEA9E4941FF9BA2951CD2FD25F987" bold="true" box="[151,736,1522,1549]" pageId="76" pageNumber="77">
34.
<taxonomicName id="10FB4D0F4941FF9BA2C11CD3FD25F987" ID-CoL="44CZX" authority="van Emden, 1965" authorityName="van Emden" authorityYear="1965" box="[195,736,1522,1549]" class="Insecta" family="Muscidae" genus="Morellia" kingdom="Animalia" order="Diptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="quadriremis" subGenus="Morellia">
<emphasis id="E58FEA9E4941FF9BA2C11CD3FEEEF986" bold="true" box="[195,299,1522,1548]" italics="true" pageId="76" pageNumber="77">Morellia</emphasis>
(
<emphasis id="E58FEA9E4941FF9BA33E1CD3FE93F986" bold="true" box="[316,342,1522,1548]" italics="true" pageId="76" pageNumber="77">M</emphasis>
.)
<emphasis id="E58FEA9E4941FF9BA36C1CD3FDC4F986" bold="true" box="[366,513,1522,1548]" italics="true" pageId="76" pageNumber="77">quadriremis</emphasis>
van
<bibRefCitation id="B36A4B7D4941FF9BA03F1CD2FD25F987" author="Emden" box="[573,736,1523,1549]" pageId="76" pageNumber="77" refString="Emden, F. I. van (1965) The Fauna of India and the adjacent countries. Diptera, 7, Muscidae, part 1. Government of India, Delhi, xiv + 647 pp." type="book" year="1965">Emden, 1965</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="9FE165074941FF9BA2951F34FE86F9A4" box="[151,323,1557,1582]" pageId="76" pageNumber="77" type="description">
<paragraph id="D744368C4941FF9BA2951F34FE86F9A4" blockId="76.[151,736,1522,1582]" box="[151,323,1557,1582]" pageId="76" pageNumber="77">
(
<figureCitation id="4FC02A094941FF9BA29D1F34FEFEF9A4" box="[159,315,1557,1582]" captionStart="FIGURES 131 143" captionStartId="75.[151,264,1879,1901]" captionTargetBox="[203,1367,193,1841]" captionTargetId="figure@75.[201,1372,193,1849]" captionTargetPageId="75" captionText="FIGURES 131 143. Males of Morellia (M.) spp. 131 134. Morellia (M.) pseudonigrisquama Shinonaga &amp; Tumrasvin. 131. Sternite 5. 132. Cercal plate. 133. Left surstylus. 134. Phallic complex, lateral view. 135 138. Morellia (M.) quadriremis van Emden. 135. Sternite 5. 136. Cercal plate. 137. Left surstylus. 138. Phallic complex, lateral view. 139 143. Morellia (M.) setosa Zielke. 139. Sternite 5. 140. Cercal plate. 141. Cercal plate, ventral view. 142. Left surstylus. 143. Phallic complex, lateral view." httpUri="https://zenodo.org/record/259303/files/figure.png" pageId="76" pageNumber="77">Figs 135138</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="9FE165074941FF9BA2951F7CFA58F9F9" box="[151,1437,1629,1651]" pageId="76" pageNumber="77" type="reference_group">
<paragraph id="D744368C4941FF9BA2951F7CFA58F9F9" blockId="76.[151,1437,1629,1651]" box="[151,1437,1629,1651]" pageId="76" pageNumber="77">
<treatmentCitationGroup id="F7EB11A24941FF9BA2951F7CFA58F9F9" box="[151,1437,1629,1651]" pageId="76" pageNumber="77">
<taxonomicName id="10FB4D0F4941FF9BA2951F7CFDA7F9F9" ID-CoL="44CZX" authority="van Emden, 1965: 105" authorityName="van Emden" authorityPageNumber="105" authorityYear="1965" box="[151,610,1629,1651]" class="Insecta" family="Muscidae" genus="Morellia" kingdom="Animalia" order="Diptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="quadriremis">
<emphasis id="E58FEA9E4941FF9BA2951F7CFEB0F9F9" bold="true" box="[151,373,1629,1651]" italics="true" pageId="76" pageNumber="77">Morellia quadriremis</emphasis>
van
<treatmentCitation id="565A109D4941FF9BA3A71F7CFDA7F9F9" author="Emden" box="[421,610,1629,1651]" page="105" pageId="76" pageNumber="77" year="1965">
<bibRefCitation id="B36A4B7D4941FF9BA3A71F7CFDF5F9F9" author="Emden" box="[421,560,1629,1651]" pageId="76" pageNumber="77" refString="Emden, F. I. van (1965) The Fauna of India and the adjacent countries. Diptera, 7, Muscidae, part 1. Government of India, Delhi, xiv + 647 pp." type="book" year="1965">Emden, 1965</bibRefCitation>
: 105
</treatmentCitation>
</taxonomicName>
. Type locality: SRI LANKA, Trincomalee Hot Wells. Holotype Ƌ, BMNH (seen).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="9FE165074941FF9AA2951F81FE67FA28" lastPageId="77" lastPageNumber="78" pageId="76" pageNumber="77" type="description">
<paragraph id="D744368C4941FF9BA2951F81FDD4F8C3" blockId="76.[151,1437,1696,2009]" pageId="76" pageNumber="77">
<emphasis id="E58FEA9E4941FF9BA2951F81FE4AF933" bold="true" box="[151,399,1696,1721]" pageId="76" pageNumber="77">Redescription. Male.</emphasis>
Colour. Black, with grey pruinosity. Frontal vitta black; face, parafacial and gena black with silver pruinosity; antenna black with grey pruinosity; palpus brown; mesonotum metallic black with a silver-dusted central vitta along presutural acrostichals, pleura grey pruinose; legs brown; haltere yellow; calypters whitish, upper calypter on outer half white and opaque; wing without dark markings; abdomen with a shifting tessellated pattern and with silver pruinosity.
</paragraph>
<paragraph id="D744368C4941FF9BA2C51E75FD70F8E6" blockId="76.[151,1437,1696,2009]" box="[199,693,1876,1901]" pageId="76" pageNumber="77">Body length: 6.0 mm; wing length: 5.0 mm.</paragraph>
<paragraph id="D744368C4941FF9AA2C51E59FD03FF7D" blockId="76.[151,1437,1696,2009]" lastBlockId="77.[151,1437,151,1550]" lastPageId="77" lastPageNumber="78" pageId="76" pageNumber="77">Head. Eyes bare but with the usual minute and scattered hairs, separated at level of anterior ocellus by 0.10 0.11 of head width; frons divergent towards lunule; ocellar triangle with 3 pairs of setae; about 13 pairs of weak frontal setae; postpedicel 1.71.9 times the length of pedicel; pedicel with 2 setae; arista plumose, with 911 cilia, bare apicad, and with some secondary plumes as far as level of 2nd to 4th cilia; vibrissa shorter than arista; 1 strong supravibrissal seta, half as long as vibrissa, and few setulae that extend to level of middle of postpedicel; 5 7 subvibrissal setae followed by 611 genal setae.</paragraph>
<paragraph id="D744368C4940FF9AA2C51825FC35FB29" blockId="77.[151,1437,151,1550]" pageId="77" pageNumber="78">Thorax- Acrostichals 0+2, the first pair weaker; dorsocentrals 0+2; intra-alars 1+1; presutural 1; prealar 1, shorter than half the length of first supra-alar; supra-alars 2; postpronotals 3; notopleuron with 2 long, subequal setae and covered with ground-setulae; postalars 2; intrapostalar 1; scutellum with 1 basal, 2 discal and 1 apical pairs of setae, and latero-ventrally with many setae, the basalmost ones downcurved; postalar wall bare; suprasquamal ridge with short pile; proepisternal depression bare; proepimeron with about 4 long setae and many setulae; prosternum broad and bare; anepisternum with a posterior row of about 7 setae, with 45 weak setae upcurved on upper-anterior corner, and covered with long setulae; katepisternals 1+2; anepimeron extensively setulose; katepimeron with 2 long setulae; anatergite bare; katatergite bare. Legs: fore femur with a posterodorsal row of setae which are shorter basad, a posteroventral row of setae on basal two-thirds, a ventral row of setae, 4 spines at apical third, and a ventral excavation at apex; fore tibia with an anterodorsal row of short and strong setae, many ventral setulae on apical half, 1 dorsal and 1 anteroventral preapical setae, and 1 ventral apical seta; fore tarsus with 2 long setae on basal segment; mid femur with a tubercle at apex of anterior to posterodorsal surfaces bearing about 8 short spines, 7 ventral setae on basal half, and an anterodorsal row of setae on basal half; mid tibia with an anterodorsal comb-like ctenidium of very short spines on basal half, 4 short and strong posterior setae on basal two-thirds, 2 posteroventral setae on middle third and 1 strong seta on apical third, 1 anterodorsal and 1 dorsal preapical setae, and 1 anteroventral, 1 posteroventral and 1 ventral apical setae; hind femur with an anteroventral row of setae, strongest on apical third, an anterodorsal row of setae, 3 dorsal setae at base, a ventral to posteroventral row of setulae on basal half, shorter but stronger in the middle, 2 posterodorsal preapical setae, a group of posterior setulae at base and few posteroventral setulae on apical fourth; hind tibia with 2 short and subequal anteroventral setae on middle third and other longer setae on apical third, an anterodorsal row of short setae on basal two-thirds of which one at middle third is longer, a dorsal row of short but strong setulae, a long and strong posterodorsal seta on apical third, 1 dorsal and 1 anterodorsal preapical setae, 1 anteroventral and 1 posteroventral apical setae, and a posterior brush-like ctenidium; hind tarsus with a golden brush ventrally. Wing with a non-uniform covering of microtrichia; dm-cu almost straight; stem-vein bare on dorsal surface, and ventrally with about 4 short setulae before humeral cross-vein and bare after humeral cross-vein; R2+3 bare on dorsal and ventral surfaces; R4+5 setulose from base to midway to r-m on dorsal surface.</paragraph>
<paragraph id="D744368C4940FF9AA2C51D90FABBFB64" blockId="77.[151,1437,151,1550]" pageId="77" pageNumber="78">
Abdomen. With a lateral tuft of setulae on syntergite 1+2; tergites 35 with a few setae laterally; tergite 5 with apical setae; sternite 1 setulose; posterior margin of sternite 5 concave and with setae, basal half bare (
<figureCitation id="4FC02A094940FF9AA7121DF4FAB7FB64" box="[1296,1394,1237,1262]" captionStart="FIGURES 131 143" captionStartId="75.[151,264,1879,1901]" captionTargetBox="[203,1367,193,1841]" captionTargetId="figure@75.[201,1372,193,1849]" captionTargetPageId="75" captionText="FIGURES 131 143. Males of Morellia (M.) spp. 131 134. Morellia (M.) pseudonigrisquama Shinonaga &amp; Tumrasvin. 131. Sternite 5. 132. Cercal plate. 133. Left surstylus. 134. Phallic complex, lateral view. 135 138. Morellia (M.) quadriremis van Emden. 135. Sternite 5. 136. Cercal plate. 137. Left surstylus. 138. Phallic complex, lateral view. 139 143. Morellia (M.) setosa Zielke. 139. Sternite 5. 140. Cercal plate. 141. Cercal plate, ventral view. 142. Left surstylus. 143. Phallic complex, lateral view." httpUri="https://zenodo.org/record/259303/files/figure.png" pageId="77" pageNumber="78">Fig. 135</figureCitation>
).
</paragraph>
<paragraph id="D744368C4940FF9AA2C51DD8FECBFAF4" blockId="77.[151,1437,151,1550]" pageId="77" pageNumber="78">
Terminalia. Cercal plate ventrally with 3 spines; surstylus with about 4 setulae basally and 2 setae apically (
<figureCitation id="4FC02A094940FF9AA29D1C3CFE84FABC" box="[159,321,1309,1334]" captionStart="FIGURES 131 143" captionStartId="75.[151,264,1879,1901]" captionTargetBox="[203,1367,193,1841]" captionTargetId="figure@75.[201,1372,193,1849]" captionTargetPageId="75" captionText="FIGURES 131 143. Males of Morellia (M.) spp. 131 134. Morellia (M.) pseudonigrisquama Shinonaga &amp; Tumrasvin. 131. Sternite 5. 132. Cercal plate. 133. Left surstylus. 134. Phallic complex, lateral view. 135 138. Morellia (M.) quadriremis van Emden. 135. Sternite 5. 136. Cercal plate. 137. Left surstylus. 138. Phallic complex, lateral view. 139 143. Morellia (M.) setosa Zielke. 139. Sternite 5. 140. Cercal plate. 141. Cercal plate, ventral view. 142. Left surstylus. 143. Phallic complex, lateral view." httpUri="https://zenodo.org/record/259303/files/figure.png" pageId="77" pageNumber="78">Figs 136137</figureCitation>
); aedeagal apodeme narrowed apically; basiphallus long; aedeagus bare; gonopod with one short seta and 4 long and strong setae on basal half, and one short seta on apical half; paramere with many setae apically (
<figureCitation id="4FC02A094940FF9AA29D1C44FEC7FAF4" box="[159,258,1381,1406]" captionStart="FIGURES 131 143" captionStartId="75.[151,264,1879,1901]" captionTargetBox="[203,1367,193,1841]" captionTargetId="figure@75.[201,1372,193,1849]" captionTargetPageId="75" captionText="FIGURES 131 143. Males of Morellia (M.) spp. 131 134. Morellia (M.) pseudonigrisquama Shinonaga &amp; Tumrasvin. 131. Sternite 5. 132. Cercal plate. 133. Left surstylus. 134. Phallic complex, lateral view. 135 138. Morellia (M.) quadriremis van Emden. 135. Sternite 5. 136. Cercal plate. 137. Left surstylus. 138. Phallic complex, lateral view. 139 143. Morellia (M.) setosa Zielke. 139. Sternite 5. 140. Cercal plate. 141. Cercal plate, ventral view. 142. Left surstylus. 143. Phallic complex, lateral view." httpUri="https://zenodo.org/record/259303/files/figure.png" pageId="77" pageNumber="78">Fig. 138</figureCitation>
).
</paragraph>
<paragraph id="D744368C4940FF9AA2C51CA8FE67FA28" blockId="77.[151,1437,151,1550]" box="[199,418,1417,1442]" pageId="77" pageNumber="78">
<emphasis id="E58FEA9E4940FF9AA2C51CA8FEE0FA28" bold="true" box="[199,293,1417,1442]" pageId="77" pageNumber="78">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="9FE165074940FF9AA2C51C8CFD00FA60" pageId="77" pageNumber="78" type="materials_examined">
<paragraph id="D744368C4940FF9AA2C51C8CFD00FA60" blockId="77.[151,1437,151,1550]" pageId="77" pageNumber="78">
<materialsCitation id="67933CD14940FF9AA2C51C8CFA5CFA4C" ID-GBIF-Occurrence="1306560157" box="[199,1433,1453,1478]" collectingDate="1890-09-07" collectionCode="BMNH" collectorName="J. W. Yerbury" country="Sri Lanka" location="Trincomalee Hot Wells" municipality="Material" pageId="77" pageNumber="78" specimenCount="1" typeStatus="holotype">
<emphasis id="E58FEA9E4940FF9AA2C51C8CFE76FA4C" bold="true" box="[199,435,1453,1478]" pageId="77" pageNumber="78">
<collectingMunicipality id="3720ACF64940FF9AA2C51C8CFEEAFA4C" box="[199,303,1453,1478]" pageId="77" pageNumber="78">Material</collectingMunicipality>
examined.
</emphasis>
<collectingCountry id="AFEC761C4940FF9AA3BF1C8FFD9FFA4C" box="[445,602,1454,1478]" name="Sri Lanka" pageId="77" pageNumber="78">SRI LANKA</collectingCountry>
:
<location id="D22460574940FF9AA0651C8CFCB0FA4C" LSID="urn:lsid:plazi:treatment:5F52879A4941FF9AA2951CD2FCA5F984:D22460574940FF9AA0651C8CFCB0FA4C" box="[615,885,1453,1478]" country="Sri Lanka" municipality="Material" name="Trincomalee Hot Wells" pageId="77" pageNumber="78">Trincomalee Hot Wells</location>
,
<date id="A345104C4940FF9AA1811C8CFC2AFA4C" box="[899,1007,1453,1478]" pageId="77" pageNumber="78" value="1890-09-07">
<collectingDate id="B301E9A44940FF9AA1811C8CFC2AFA4C" box="[899,1007,1453,1478]" pageId="77" pageNumber="78" value="1890-09-07">7.ix.1890</collectingDate>
</date>
,
<collectorName id="7A0E535A4940FF9AA1FC1C8FFB51FA4C" box="[1022,1172,1453,1478]" pageId="77" pageNumber="78">J.W. Yerbury</collectorName>
,
<typeStatus id="0840882E4940FF9AA6A11C8CFAC0FA4C" box="[1187,1285,1453,1478]" pageId="77" pageNumber="78" type="holotype">holotype</typeStatus>
Ƌ (
<collectionCode id="B1EAAE494940FF9AA7341C8FFA55FA4C" box="[1334,1424,1454,1478]" pageId="77" pageNumber="78">BMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="67933CD14940FF9AA2951CF3FD04FA60" ID-GBIF-Occurrence="1306560173" box="[151,705,1489,1514]" collectingDate="1913-10-20" collectionCode="BMNH" country="India" location="Bihar" pageId="77" pageNumber="78" specimenCount="1" stateProvince="Bihar" typeStatus="paratype">
<collectingCountry id="AFEC761C4940FF9AA2951CF3FF2EFA60" box="[151,235,1490,1514]" name="India" pageId="77" pageNumber="78">INDIA</collectingCountry>
:
<collectingRegion id="153FF86E4940FF9AA2F71CF0FEF0FA60" box="[245,309,1489,1514]" country="India" name="Bihar" pageId="77" pageNumber="78">Bihar</collectingRegion>
,
<date id="A345104C4940FF9AA3421CF3FE77FA60" box="[320,434,1490,1514]" pageId="77" pageNumber="78" value="1913-10-20">
<collectingDate id="B301E9A44940FF9AA3421CF3FE77FA60" box="[320,434,1490,1514]" pageId="77" pageNumber="78" value="1913-10-20">20.x.1913</collectingDate>
</date>
, 1 Ƌ
<typeStatus id="0840882E4940FF9AA3ED1CF3FD8AFA60" box="[495,591,1490,1514]" pageId="77" pageNumber="78" type="paratype">paratype</typeStatus>
(
<collectionCode id="B1EAAE494940FF9AA05C1CF3FD7DFA60" box="[606,696,1490,1514]" pageId="77" pageNumber="78">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="9FE165074940FF9AA2C51CD4FCA5F984" box="[199,864,1525,1550]" pageId="77" pageNumber="78" type="distribution">
<paragraph id="D744368C4940FF9AA2C51CD4FCA5F984" blockId="77.[151,1437,151,1550]" box="[199,864,1525,1550]" pageId="77" pageNumber="78">
<emphasis id="E58FEA9E4940FF9AA2C51CD4FEA5F984" bold="true" box="[199,352,1525,1550]" pageId="77" pageNumber="78">Distribution.</emphasis>
<collectingRegion id="153FF86E4940FF9AA3651CD7FE36F984" box="[359,499,1526,1550]" country="Egypt" name="Ash Sharqiyah" pageId="77" pageNumber="78">ORIENTAL</collectingRegion>
:
<collectingCountry id="AFEC761C4940FF9AA3FF1CD4FDF3F984" box="[509,566,1525,1550]" name="India" pageId="77" pageNumber="78">India</collectingCountry>
(several states),
<collectingCountry id="AFEC761C4940FF9AA0EF1CD4FC99F984" box="[749,860,1525,1550]" name="Sri Lanka" pageId="77" pageNumber="78">
<collectingRegion id="153FF86E4940FF9AA0EF1CD4FCC8F984" box="[749,781,1525,1550]" country="United Kingdom" name="Surrey" pageId="77" pageNumber="78">Sri</collectingRegion>
Lanka
</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>