177 lines
20 KiB
XML
177 lines
20 KiB
XML
<document id="391BE67530BE0305FA5E4D2343D8CA3E" ID="10.11646/zootaxa.4163.1.1" ID-DOI="10.11646/zootaxa.4163.1.1" ID-GBIF-Dataset="69b1046e-47bc-4f26-b32c-97d79da7a35d" ID-ISSN="1175-5326" ID-Zenodo-Dep="259278" ID-ZooBank="930AF345-15AD-443A-A3AE-328178ECB6C9" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1473747460051" checkinUser="plazi" docAuthor="Pamplona, Denise, Nihei, Silvio S., Couri, Márcia S. & Pont, Adrian C." docDate="2016" docId="5F52879A491EFFC4A29519B9FBE6F8E6" docLanguage="en" docName="zootaxa.4163.1.1.pdf" docOrigin="Zootaxa 4163 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Morellia (Morellia) atrisquama Malloch 1935" docType="treatment" docVersion="8" lastPageNumber="20" masterDocId="A36BFFE2490DFFD7A2021921FFC5FF8A" masterDocTitle="Taxonomy of Morellia Robineau-Desvoidy (Diptera: Muscidae): revision of the subgenera Morellia s. str. and Parapyrellia Townsend" masterLastPageNumber="110" masterPageNumber="1" pageNumber="20" updateTime="1698385354984" updateUser="plazi">
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<mods:titleInfo id="8ACC6FF170A4D63FB04F1946F9AB3EFB">
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<mods:title id="2BA62764E019D3EDC9158990E7EF2FA2">Taxonomy of Morellia Robineau-Desvoidy (Diptera: Muscidae): revision of the subgenera Morellia s. str. and Parapyrellia Townsend</mods:title>
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<mods:name id="191396252D80423A35A483891E5C890A" type="personal">
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<mods:namePart id="C4C647695975169ED18281C752A59E1F">Pamplona, Denise</mods:namePart>
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<mods:roleTerm id="0F4DCCA97F537EDB857936A6223B0244">Author</mods:roleTerm>
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<mods:namePart id="3D09D016A7ACBCCC0CD59E3FCD0B6673">Nihei, Silvio S.</mods:namePart>
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<mods:roleTerm id="B54D2B0CE25A9F2D420CC34A90D20803">Author</mods:roleTerm>
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<mods:namePart id="AF5474FCD58818B29B29E4B8F993174A">Couri, Márcia S.</mods:namePart>
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<mods:name id="ECCB47C43EACBAE7B333576484F73E6C" type="personal">
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<mods:namePart id="1ABFD425F354DFBF82096DB39076FAA4">Pont, Adrian C.</mods:namePart>
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<mods:title id="19CC95C781A91E8F5D4E8D476D1F07F3">Zootaxa</mods:title>
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<mods:part id="AFBDE91F702591887BA068A98D85E931">
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<mods:date id="1E189264B3CB7867953E96BC715D4F9A">2016</mods:date>
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<mods:detail id="DFA20137A4330EFC73412A2ABCC7BA99" type="volume">
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<mods:number id="BEA44CF3093955F784F8CCEC98918F40">4163</mods:number>
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<mods:classification id="6E06107D5658A56991FF9386E902BE46">journal article</mods:classification>
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<mods:identifier id="FC6A3E6466159CB8D28B85D05C5B3339" type="DOI">10.11646/zootaxa.4163.1.1</mods:identifier>
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<treatment id="5F52879A491EFFC4A29519B9FBE6F8E6" ID-DOI="http://doi.org/10.5281/zenodo.4547382" ID-GBIF-Taxon="124548473" ID-Zenodo-Dep="4547382" LSID="urn:lsid:plazi:treatment:5F52879A491EFFC4A29519B9FBE6F8E6" httpUri="http://treatment.plazi.org/id/5F52879A491EFFC4A29519B9FBE6F8E6" lastPageNumber="20" pageId="19" pageNumber="20">
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<subSubSection id="9FE16507491EFFC4A29519B9FD65FF38" box="[151,672,151,178]" pageId="19" pageNumber="20" type="nomenclature">
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<paragraph id="D744368C491EFFC4A29519B9FD65FF38" blockId="19.[151,672,151,211]" box="[151,672,151,178]" pageId="19" pageNumber="20">
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<heading id="8C0C81E0491EFFC4A29519B9FD65FF38" bold="true" box="[151,672,151,178]" fontSize="11" level="1" pageId="19" pageNumber="20" reason="1">
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<emphasis id="E58FEA9E491EFFC4A29519B9FD65FF38" bold="true" box="[151,672,151,178]" pageId="19" pageNumber="20">
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4.
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<taxonomicName id="10FB4D0F491EFFC4A2B619B6FD65FF38" ID-CoL="44CY7" authority="Malloch, 1935" authorityName="Malloch" authorityYear="1935" box="[180,672,151,178]" class="Insecta" family="Muscidae" genus="Morellia" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="atrisquama" subGenus="Morellia">
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<emphasis id="E58FEA9E491EFFC4A2B619B6FED9FF3B" bold="true" box="[180,284,151,177]" italics="true" pageId="19" pageNumber="20">Morellia</emphasis>
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(
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<emphasis id="E58FEA9E491EFFC4A32F19B6FE82FF3B" bold="true" box="[301,327,151,177]" italics="true" pageId="19" pageNumber="20">M</emphasis>
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||
.)
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<emphasis id="E58FEA9E491EFFC4A36219B6FE2CFF3B" bold="true" box="[352,489,151,177]" italics="true" pageId="19" pageNumber="20">atrisquama</emphasis>
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<bibRefCitation id="B36A4B7D491EFFC4A3F219B9FD65FF38" author="Malloch" box="[496,672,152,178]" pageId="19" pageNumber="20" refString="Malloch, J. R. (1935) Exotic Muscaridae (Diptera). - XL. Annals and Magazine of Natural History, Series 10, 16, 573 - 597." type="journal article" year="1935">Malloch, 1935</bibRefCitation>
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</taxonomicName>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="9FE16507491EFFC4A295199BFEE2FF59" box="[151,295,186,211]" pageId="19" pageNumber="20" type="description">
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<paragraph id="D744368C491EFFC4A295199BFEE2FF59" blockId="19.[151,672,151,211]" box="[151,295,186,211]" pageId="19" pageNumber="20">
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||
(
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<figureCitation id="4FC02A09491EFFC4A29D199BFEDBFF59" box="[159,286,186,211]" captionStart="FIGURES 19 – 20" captionStartId="20.[151,264,1953,1975]" captionTargetBox="[319,1249,207,1913]" captionTargetId="figure@20.[319,1267,193,1932]" captionTargetPageId="20" captionText="FIGURES 19 – 20. Morellia (M.) atrisquama Malloch, female. 19. Ovipositor, dorsal view. 20. Ovipositor and spermathecae, ventral view." httpUri="https://zenodo.org/record/259282/files/figure.png" pageId="19" pageNumber="20">Figs 19–20</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="9FE16507491EFFC4A2951823FA91FE92" box="[151,1364,258,280]" pageId="19" pageNumber="20" type="reference_group">
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<paragraph id="D744368C491EFFC4A2951823FA91FE92" blockId="19.[151,1364,258,280]" box="[151,1364,258,280]" pageId="19" pageNumber="20">
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<treatmentCitationGroup id="F7EB11A2491EFFC4A2951823FA91FE92" box="[151,1364,258,280]" pageId="19" pageNumber="20">
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<materialsCitation id="67933CD1491EFFC4A2951823FBEAFE92" ID-GBIF-Occurrence="1306560083" box="[151,1071,258,280]" location="Mt. Kinabalu" pageId="19" pageNumber="20" specimenCount="1" stateProvince="SABAH" typeStatus="holotype">
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<taxonomicName id="10FB4D0F491EFFC4A2951823FDF8FE92" ID-CoL="44CY7" authority="Malloch, 1935: 573" authorityName="Malloch" authorityPageNumber="573" authorityYear="1935" box="[151,573,258,280]" class="Insecta" family="Muscidae" genus="Morellia" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="atrisquama">
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<emphasis id="E58FEA9E491EFFC4A2951823FEA8FE92" bold="true" box="[151,365,258,280]" italics="true" pageId="19" pageNumber="20">Morellia atrisquama</emphasis>
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<treatmentCitation id="565A109D491EFFC4A3761823FDF8FE92" author="Malloch" box="[372,573,258,280]" page="573" pageId="19" pageNumber="20" year="1935">
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<bibRefCitation id="B36A4B7D491EFFC4A3761823FDCEFE92" author="Malloch" box="[372,523,258,280]" pageId="19" pageNumber="20" refString="Malloch, J. R. (1935) Exotic Muscaridae (Diptera). - XL. Annals and Magazine of Natural History, Series 10, 16, 573 - 597." type="journal article" year="1935">Malloch, 1935</bibRefCitation>
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: 573
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</treatmentCitation>
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||
</taxonomicName>
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.
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<typeStatus id="0840882E491EFFC4A0451822FDBCFE92" box="[583,633,259,280]" pageId="19" pageNumber="20">Type</typeStatus>
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locality:
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<collectingRegion id="153FF86E491EFFC4A0DB1823FCF4FE92" box="[729,817,258,280]" country="Malaysia" name="Sabah" pageId="19" pageNumber="20">SABAH</collectingRegion>
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,
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<location id="D2246057491EFFC4A1391822FC00FE92" LSID="urn:lsid:plazi:treatment:5F52879A491EFFC4A29519B9FBE6F8E6:D2246057491EFFC4A1391822FC00FE92" box="[827,965,258,280]" name="Mt. Kinabalu" pageId="19" pageNumber="20" stateProvince="SABAH">Mt. Kinabalu</location>
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, Kenokok
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</materialsCitation>
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. Holotype ♀, BMNH (seen).
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</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="9FE16507491EFFC4A2951864FE40F88B" pageId="19" pageNumber="20" type="description">
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<paragraph id="D744368C491EFFC4A2951864FCE7FE64" blockId="19.[151,1437,325,1901]" pageId="19" pageNumber="20">
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<emphasis id="E58FEA9E491EFFC4A2951864FE81FED4" bold="true" box="[151,324,325,350]" pageId="19" pageNumber="20">Redescription.</emphasis>
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Colour. Black, with grey pruinosity. Frontal vitta and fronto-orbital plate black; face, parafacial and gena black with silver pruinosity; antenna dark brown with grey pruinosity; palpus brown, mesonotum black with a silver-dusted central vitta in front of the acrostichals; postpronotal lobe, notopleuron and anepisternum silver pruinose; scutellum silver laterally; legs dark brown; haltere yellow; calypters brownish; wing without dark markings; abdomen black with last tergite silver pruinose.
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</paragraph>
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<paragraph id="D744368C491EFFC4A2C518D8FC8FFD9B" blockId="19.[151,1437,325,1901]" box="[199,842,505,530]" pageId="19" pageNumber="20">
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<emphasis id="E58FEA9E491EFFC4A2C518D8FEE0FD98" bold="true" box="[199,293,505,530]" pageId="19" pageNumber="20">Female.</emphasis>
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Body length. 8.0 mm; wing length: 7.0–8.0 mm.
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</paragraph>
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<paragraph id="D744368C491EFFC4A2C51B3CFDA8FC87" blockId="19.[151,1437,325,1901]" pageId="19" pageNumber="20">Head. Eyes bare, separated at level of anterior ocellus by about 0.29 of head width; frons divergent towards lunule; ocellar triangle with a pair of setae and few setulae; with 3 proclinate and 2 reclinate setae; 15 pairs of frontal setae, about 5 or 6 much shorter than the others; 5–6 fronto-orbitals; postpedicel about 2.0–2.3 times the length of pedicel; pedicel with 2 setae; arista plumose, with 29–30 cilia, bare apicad, with some secondary plumes as far as level of 9th to 11th cilia; vibrissa shorter than arista, 2–3 strong supravibrissal setae, half as long as vibrissa, and some other setulae that extend to level of basal fourth of postpedicel; 5–6 subvibrissal setae followed by 7–8 genal setae and other shorter ones.</paragraph>
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<paragraph id="D744368C491EFFC4A2C51A38FB34F98F" blockId="19.[151,1437,325,1901]" pageId="19" pageNumber="20">Thorax. Acrostichals 0+2; dorsocentrals 2+4; intra-alars 1+1; presutural 1; prealar 1, half the length the first supra-alar; supra-alars 2; postpronotals 3; notopleuron with 2 long, subequal setae and covered with groundsetulae; postalars 2; intrapostalar 1; scutellum with 1 basal, 1 latero-discal and 1 apical pairs of setae, and lateroventrally with many setae, the basalmost ones downcurved; postalar wall bare; suprasquamal ridge bare; proepisternal depression bare; proepimeron with 3 long setae (the lower one shorter) and many setulae; prosternum broad and setulose; anepisternum with a posterior row of about 7 setae, with 4 weak setae upcurved on upperanterior corner, covered with short setulae; katepisternals 1+2; anepimeron setulose medially and on subalar ridge; katepimeron bare; anatergite bare and katatergite setulose. Legs: fore femur with a posterodorsal row of setae, longer on apical half, and a complete posteroventral row of setae; fore tibia with a dorsal and an anteroventral preapical setae, and anterior surface with a ctenidium of golden setulae; mid femur without a tubercle, with 1 anterior seta on middle third, 4 ventral setae on basal half, anteroventral setulae, and 6 posterior to posterodorsal preapical setae, the 2 first weaker; mid tibia with 3 strong posteroventral setae on basal two-thirds, and a posterior row of short setae, 4 or 5 of which longer, 1 preapical seta and 2 anteroventral, 1 anterior and 2 ventral apical setae; hind femur with a complete anterodorsal row of setae and with an anteroventral row of fine setae on apical half, a weak posterior tuft at base, a row of weak setae on basal half, and few posteroventral setulae on apical fourth; hind tibia with 3 anteroventral setae on apical third, a dorsal row of setae, about 4 of which stronger, a posterodorsal row of setae, the median and basal ones longer, 1 dorsal and 1 anterodorsal preapical setae, and 1 anteroventral, 2 ventral and no posteroventral apical setae; hind tarsus with a golden brush ventrally. Wing with a uniform covering of microtrichia; dm-cu almost straight, stem-vein bare on dorsal surface, and ventrally with about 9 short setulae before the humeral cross-vein; R2+3 bare on dorsal and ventral surfaces; R4+5 with strong setulae on dorsal surface from base to r-m, and from base to two-thirds to r-m on ventral surface; 3 strong setae on basicosta.</paragraph>
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<paragraph id="D744368C491EFFC4A2C51F31FD0CF9A3" blockId="19.[151,1437,325,1901]" box="[199,713,1552,1577]" pageId="19" pageNumber="20">Abdomen. With few setae; sternite 1 setulose.</paragraph>
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<paragraph id="D744368C491EFFC4A2C51F15FC73F956" blockId="19.[151,1437,325,1901]" pageId="19" pageNumber="20">
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Ovipositor. Tergite 6 rod-like with 3 groups of very small spines; tergite 7 rod-like with 5 strong curved spines posteriorly; tergite 8 sinuous, rod-like, broadened posteriorly; enlarged spicules on intersegmental membrane between segments 6–7 and 7–8; sternite 6 small, rod-like; sternite 7 rod-like with 3 plates posteriorly, each with strong curved spines; sternite 8 very small with very small spines; epiproct large, with 4 very small spines; hypoproct with many setae; cercus flattened and not long (
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<figureCitation id="4FC02A09491EFFC4A1241FE5FC6FF956" box="[806,938,1732,1757]" captionStart="FIGURES 19 – 20" captionStartId="20.[151,264,1953,1975]" captionTargetBox="[319,1249,207,1913]" captionTargetId="figure@20.[319,1267,193,1932]" captionTargetPageId="20" captionText="FIGURES 19 – 20. Morellia (M.) atrisquama Malloch, female. 19. Ovipositor, dorsal view. 20. Ovipositor and spermathecae, ventral view." httpUri="https://zenodo.org/record/259282/files/figure.png" pageId="19" pageNumber="20">Figs 19–20</figureCitation>
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).
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</paragraph>
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<paragraph id="D744368C491EFFC4A2C51FC9FE40F88B" blockId="19.[151,1437,325,1901]" box="[199,389,1768,1793]" pageId="19" pageNumber="20">Male. Unknown.</paragraph>
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</subSubSection>
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<subSubSection id="9FE16507491EFFC4A2C51E2DFCB1F8C2" pageId="19" pageNumber="20" type="materials_examined">
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<paragraph id="D744368C491EFFC4A2C51E2DFCB1F8C2" blockId="19.[151,1437,325,1901]" pageId="19" pageNumber="20">
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||
<materialsCitation id="67933CD1491EFFC4A2C51E2DFEC4F8C2" ID-GBIF-Occurrence="1306560082" collectingDate="1929-04-22" collectionCode="BMNH" collectorName="H. M. Pendlebury" county="Material" elevation="1006" location="Kenokok" municipality="Mt. Kinabalu" pageId="19" pageNumber="20" specimenCount="1" stateProvince="SABAH" typeStatus="holotype">
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||
<emphasis id="E58FEA9E491EFFC4A2C51E2DFE76F8AF" bold="true" box="[199,435,1804,1829]" pageId="19" pageNumber="20">
|
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<collectingCounty id="3E254E00491EFFC4A2C51E2DFEEAF8AF" box="[199,303,1804,1829]" pageId="19" pageNumber="20">Material</collectingCounty>
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||
examined.
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||
</emphasis>
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||
<collectingRegion id="153FF86E491EFFC4A3BF1E2DFDE5F8AE" box="[445,544,1804,1828]" country="Malaysia" name="Sabah" pageId="19" pageNumber="20">SABAH</collectingRegion>
|
||
:
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||
<collectingMunicipality id="3720ACF6491EFFC4A02F1E2CFD0CF8AF" box="[557,713,1804,1829]" pageId="19" pageNumber="20">Mt. Kinabalu</collectingMunicipality>
|
||
,
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||
<location id="D2246057491EFFC4A0D51E2DFC84F8AF" LSID="urn:lsid:plazi:treatment:5F52879A491EFFC4A29519B9FBE6F8E6:D2246057491EFFC4A0D51E2DFC84F8AF" box="[727,833,1804,1829]" county="Material" municipality="Mt. Kinabalu" name="Kenokok" pageId="19" pageNumber="20" stateProvince="SABAH">Kenokok</location>
|
||
,
|
||
<quantity id="10039B69491EFFC4A14D1E2DFC6BF8AF" box="[847,942,1804,1829]" metricMagnitude="3" metricUnit="m" metricValue="1.0058399999999998" pageId="19" pageNumber="20" unit="ft" value="3300.0">
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<elevation id="5CD6D1BF491EFFC4A14D1E2DFC6BF8AF" box="[847,942,1804,1829]" metricMagnitude="3" metricUnit="m" metricValue="1.0058399999999998" pageId="19" pageNumber="20" unit="ft" value="3300.0">3,300 ft</elevation>
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||
</quantity>
|
||
.,
|
||
<date id="A345104C491EFFC4A1C21E2DFBFFF8AF" box="[960,1082,1804,1829]" pageId="19" pageNumber="20" value="1929-04-22">
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||
<collectingDate id="B301E9A4491EFFC4A1C21E2DFBFFF8AF" box="[960,1082,1804,1829]" pageId="19" pageNumber="20" value="1929-04-22">22.iv.1929</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="7A0E535A491EFFC4A64A1E2CFAC8F8AF" box="[1096,1293,1804,1829]" pageId="19" pageNumber="20">H.M. Pendlebury</collectorName>
|
||
,
|
||
<typeStatus id="0840882E491EFFC4A7191E2DFAB8F8AF" box="[1307,1405,1804,1829]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
|
||
Ƌ (
|
||
<collectionCode id="B1EAAE49491EFFC4A29D1E11FF3CF8C2" box="[159,249,1840,1864]" pageId="19" pageNumber="20">BMNH</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="67933CD1491EFFC4A30E1E10FCB5F8C2" ID-GBIF-Occurrence="1306560080" box="[268,880,1840,1865]" collectingDate="1929-04-22" collectingDateMax="1929-04-26" collectingDateMin="1929-04-22" collectionCode="BMNH" collectorName="H. M. Pendlebury" county="Material" elevation="1006" location="Kenokok" municipality="Mt. Kinabalu" pageId="19" pageNumber="20" specimenCount="2" specimenCount-female="2" stateProvince="SABAH" typeStatus="paratype">
|
||
same locality,
|
||
<date id="A345104C491EFFC4A3AF1E11FD92F8C3" box="[429,599,1840,1865]" pageId="19" pageNumber="20" value="1929-04-22" valueMax="1929-04-26" valueMin="1929-04-22">
|
||
<collectingDate id="B301E9A4491EFFC4A3AF1E11FD92F8C3" box="[429,599,1840,1865]" pageId="19" pageNumber="20" value="1929-04-22" valueMax="1929-04-26" valueMin="1929-04-22">22–26. iv.1929</collectingDate>
|
||
</date>
|
||
,
|
||
<specimenCount id="C1FDFD05491EFFC4A0601E11FD49F8C2" box="[610,652,1840,1864]" pageId="19" pageNumber="20" type="female">2 ♀</specimenCount>
|
||
<typeStatus id="0840882E491EFFC4A0911E10FD3BF8C3" box="[659,766,1841,1865]" pageId="19" pageNumber="20" type="paratype">paratypes</typeStatus>
|
||
(
|
||
<collectionCode id="B1EAAE49491EFFC4A10F1E11FCA2F8C2" box="[781,871,1840,1864]" pageId="19" pageNumber="20">BMNH</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9FE16507491EFFC4A2C51E75FBE6F8E6" box="[199,1059,1876,1901]" pageId="19" pageNumber="20" type="distribution">
|
||
<paragraph id="D744368C491EFFC4A2C51E75FBE6F8E6" blockId="19.[151,1437,325,1901]" box="[199,1059,1876,1901]" pageId="19" pageNumber="20">
|
||
<emphasis id="E58FEA9E491EFFC4A2C51E75FEA5F8E7" bold="true" box="[199,352,1876,1901]" pageId="19" pageNumber="20">Distribution.</emphasis>
|
||
<collectingRegion id="153FF86E491EFFC4A3651E75FE30F8E6" box="[359,501,1876,1900]" country="Egypt" name="Ash Sharqiyah" pageId="19" pageNumber="20">ORIENTAL</collectingRegion>
|
||
: known only from
|
||
<collectingRegion id="153FF86E491EFFC4A0CB1E75FCCBF8E7" box="[713,782,1876,1901]" country="Malaysia" name="Sabah" pageId="19" pageNumber="20">Sabah</collectingRegion>
|
||
on the island of Borneo.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |