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<document id="032AA99276B495D907D0C6315EB78F63" ID-CLB-Dataset="21844" ID-DOI="10.5852/ejt.2017.359" ID-GBIF-Dataset="87920495-c1bf-4bdb-b153-50e709c1d6c2" ID-ISSN="2118-9773" ID-Zenodo-Dep="3855694" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1590500868344" checkinUser="carolina" docAuthor="dAcoz, Cédric dUdekem &amp; Verheye, Marie L." docDate="2017" docId="4A5A879BFF0D68F3FDE4FE55CFFAFCFB" docLanguage="en" docName="ejt-2017-359.pdf.imf" docOrigin="European Journal of Taxonomy 359" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Subepimeria Bellan-Santini 1972" docType="treatment" docVersion="10" lastPageNumber="141" masterDocId="B663FFE3FF86687FFFF7FFE8CF17FF89" masterDocTitle="Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea)" masterLastPageNumber="553" masterPageNumber="1" pageNumber="140" updateTime="1698841458082" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="86D331B159C43A9F6A2D8E60397A23F8">Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea)</mods:title>
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<mods:namePart id="F1BC09F591FEDD24B476F57B51F89EBC">dAcoz, Cédric dUdekem</mods:namePart>
<mods:affiliation id="B0709344CFC3341CDEE2C6A5648356FF">Royal Belgian Institute of Natural Sciences, Service Heritage, Rue Vautier 29, B- 1000 Brussels, Belgium. &amp; Corresponding author: cdudekem @ naturalsciences. be</mods:affiliation>
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<mods:namePart id="3C61939B7F6C9C6807386C1186AC411A">Verheye, Marie L.</mods:namePart>
<mods:affiliation id="CB24CA8857D0FAD09C14BACF8BA96B83">Royal Belgian Institute of Natural Sciences, Operational direction Taxonomy and Phylogeny, Rue Vautier 29, B- 1000 Brussels, Belgium. &amp; Email: mverheye @ naturalsciences. be</mods:affiliation>
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Subgenus
<taxonomicName id="05F34D0EFF0D68F4FD7AFE55CB37FE5E" ID-CoL="ae1f8aee-9dc3-4f83-8867-310b6eb8ecbb" authority="Bellan-Santini, 1972" authorityName="Bellan-Santini" authorityYear="1972" box="[653,1056,445,472]" class="Malacostraca" family="Epimeriidae" genus="Subepimeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="139" pageNumber="140" phylum="Arthropoda" rank="genus">
<emphasis id="F087EA9FFF0D68F4FD7AFE55CC31FE51" bold="true" box="[653,806,445,472]" italics="true" pageId="139" pageNumber="140">Subepimeria</emphasis>
<bibRefCitation id="A6624B7CFF0D68F4FCDAFE55CB37FE5E" author="Bellan-Santini" box="[813,1056,445,471]" firstAuthor="Bellan-Santini" pageId="139" pageNumber="140" pagination="157 - 238" refId="ref132325" refString="Bellan-Santini D. 1972. Invertebres marins des XIIeme et XVeme Expeditions Antarctiques Francaises en Terre Adelie. 10. Amphipodes Gammariens. Tethys, Supplement 4: 157 - 238." type="journal article" year="1972">Bellan-Santini, 1972</bibRefCitation>
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<treatmentCitation id="4352109CFF0D68F4FF4AFDE8CD9CFD92" author="Bellan-Santini D." box="[189,651,512,539]" page="225" pageId="139" pageNumber="140" year="1972">
<taxonomicName id="05F34D0EFF0D68F4FF4AFDE8CD9CFD92" ID-CoL="ae1f8aee-9dc3-4f83-8867-310b6eb8ecbb" authority="Bellan-Santini, 1972: 225" authorityName="Bellan-Santini" authorityPageNumber="225" authorityYear="1972" box="[189,651,512,539]" class="Malacostraca" family="Epimeriidae" genus="Subepimeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="139" pageNumber="140" phylum="Arthropoda" rank="genus">
<emphasis id="F087EA9FFF0D68F4FF4AFDE8CE46FD93" box="[189,337,512,538]" italics="true" pageId="139" pageNumber="140">Subepimeria</emphasis>
<bibRefCitation id="A6624B7CFF0D68F4FEAEFDE8CD9CFD92" author="Bellan-Santini" box="[345,651,512,539]" firstAuthor="Bellan-Santini" pageId="139" pageNumber="140" pagination="157 - 238" refId="ref132325" refString="Bellan-Santini D. 1972. Invertebres marins des XIIeme et XVeme Expeditions Antarctiques Francaises en Terre Adelie. 10. Amphipodes Gammariens. Tethys, Supplement 4: 157 - 238." type="journal article" year="1972">Bellan-Santini, 1972: 225</bibRefCitation>
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.
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<taxonomicName id="05F34D0EFF0D68F4FF4AFDABCE46FDD4" ID-CoL="ae1f8aee-9dc3-4f83-8867-310b6eb8ecbb" authorityName="Bellan-Santini" authorityYear="1972" box="[189,337,579,605]" class="Malacostraca" family="Epimeriidae" genus="Subepimeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="139" pageNumber="140" phylum="Arthropoda" rank="genus">
<emphasis id="F087EA9FFF0D68F4FF4AFDABCE46FDD4" box="[189,337,579,605]" italics="true" pageId="139" pageNumber="140">Subepimeria</emphasis>
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<treatmentCitation id="4352109CFF0D68F4FE98FDACCC19FDD4" box="[367,782,579,606]" pageId="139" pageNumber="140">
<bibRefCitation id="A6624B7CFF0D68F4FE98FDACCC19FDD4" author="Karaman G. S. &amp; Barnard J. L." box="[367,782,579,606]" pageId="139" pageNumber="140" pagination="106 - 165" refId="ref140656" refString="Karaman G. S. &amp; Barnard J. L. 1979. Classificatory revisions in gammaridean Amphipoda (Crustacea), part I. Proceedings of the Biological Society of Washington 92 (1): 106 - 165. Available from http: // biodiversitylibrary. org / page / 35513813 [accessed 27 Sep. 2016]." type="journal article" year="1979">Karaman &amp; J.L. Barnard 1979: 108</bibRefCitation>
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109 (in part).
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<typeStatus id="1D48882FFF0D68F4FF4AFD6ECFEDFD29" box="[189,250,646,672]" pageId="139" pageNumber="140">Type</typeStatus>
species
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</heading>
</paragraph>
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<taxonomicName id="05F34D0EFF0D68F4FF4AFD5ECDDCFD59" ID-CoL="206dd94f-be02-43a7-bfbf-880d0998034a" authority="Bellan-Santini, 1972" authorityName="Bellan-Santini" authorityYear="1972" box="[189,715,694,720]" class="Malacostraca" family="Epimeriidae" genus="Subepimeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="139" pageNumber="140" phylum="Arthropoda" rank="species" species="geodesiae">
<emphasis id="F087EA9FFF0D68F4FF4AFD5ECED9FD59" box="[189,462,694,720]" italics="true" pageId="139" pageNumber="140">Subepimeria geodesiae</emphasis>
<bibRefCitation id="A6624B7CFF0D68F4FE22FD5ECDDCFD59" author="Bellan-Santini" box="[469,715,694,720]" firstAuthor="Bellan-Santini" pageId="139" pageNumber="140" pagination="157 - 238" refId="ref132325" refString="Bellan-Santini D. 1972. Invertebres marins des XIIeme et XVeme Expeditions Antarctiques Francaises en Terre Adelie. 10. Amphipodes Gammariens. Tethys, Supplement 4: 157 - 238." type="journal article" year="1972">Bellan-Santini, 1972</bibRefCitation>
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.
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<paragraph id="C24C368DFF0D68F4FF4AFD11CE59FC9A" blockId="139.[189,334,761,787]" box="[189,334,761,787]" pageId="139" pageNumber="140">
<heading id="990481E1FF0D68F4FF4AFD11CE59FC9A" bold="true" box="[189,334,761,787]" fontSize="11" level="3" pageId="139" pageNumber="140" reason="3">
<emphasis id="F087EA9FFF0D68F4FF4AFD11CE59FC9A" bold="true" box="[189,334,761,787]" pageId="139" pageNumber="140">Description</emphasis>
</heading>
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<paragraph id="C24C368DFF0D68F4FF4AFCC2CCCFF9E7" blockId="139.[189,1399,810,1647]" pageId="139" pageNumber="140">Body opaque, with teguments moderately calcified. Rostrum medium-sized. Eyes not conical. Pleonite 2 posteriorly produced into a small sharp tooth oriented backwards; pleonite 1 and pereionite 7 sometimes also posteriorly produced into a tooth or bump; dorsal border of pleonite 3 keeled; in lateral view that keel posteriorly terminated into a rounded bump or a squared angle. With the exception of the aforementioned mid-dorsal ornamentations, pereionites and pleosomites perfectly smooth. Coxae 13 with blunt tip, not sharply keeled. Coxa 4 with indistinct carina starting at ventral tip of coxa and terminating at posterior tip of coxa; this carina remains very close to the posteroventral border of coxa, the space in-between forming a very shallow groove; posteroventral border of coxa 4 very weakly concave, nearly straight. Coxae 56 without tooth or distinct protrusion. Mid of posterior border of epimeral plates 13 not produced into a tooth. Posteroventral tooth of epimeral plate 3 medium-sized to strong. Dorsal process of urosomite 1 produced into a triangular tooth directed upwards. Urosomite 2 without pair of small teeth pointing upwards. Lateral borders of urosomite 3 posteriorly terminated into a sharp tooth or bluntly angular. Peduncle of antenna 1 without teeth or with dentition vestigial. Mandible with pars molar process triturative. Lower lip with narrow (V-shaped) hypopharyngeal gap. Palp of maxilliped with 4 articles. Gnathopods of normal size, with carpus and propodus of medium slenderness, with palm obliquely transverse but poorly developed (gnathopods subcheliform); propodus not expanded distally; posterior border of dactylus lined by row of small oblique slender teeth. Basis of pereiopods 56 moderately broad, with posteroproximal rounded protrusion (sometimes very weak and scarcely distinct); a posterodistal tooth projecting posteriorly can be present on basis of pereiopod 5, otherwise basis posterodistally angulate or bluntly angulate. Posterior border of basis of pereiopod 7 slightly convex in proximal 0.8; at this level there is a slight angular discontinuity and the distal 0.2 can be slightly concave; posterior border terminated into a blunt angle or a blunt tooth directed in the axis of the basis. Dactylus of pereiopods 57 short. Benthic. Small species.</paragraph>
<paragraph id="C24C368DFF0D68F4FF4AF970CE43F93B" blockId="139.[189,340,1688,1714]" box="[189,340,1688,1714]" pageId="139" pageNumber="140">
<heading id="990481E1FF0D68F4FF4AF970CE43F93B" bold="true" box="[189,340,1688,1714]" fontSize="11" level="3" pageId="139" pageNumber="140" reason="3">
<emphasis id="F087EA9FFF0D68F4FF4AF970CE43F93B" bold="true" box="[189,340,1688,1714]" pageId="139" pageNumber="140">Body length</emphasis>
</heading>
</paragraph>
<paragraph id="C24C368DFF0D68F4FF4AF92FCBE3F96B" blockId="139.[189,1268,1735,1762]" box="[189,1268,1735,1762]" pageId="139" pageNumber="140">
The maximum body length recorded in
<taxonomicName id="05F34D0EFF0D68F4FD78F92FCC34F968" authorityName="Bellan-Santini" authorityYear="1972" box="[655,803,1735,1761]" class="Malacostraca" family="Epimeriidae" genus="Subepimeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="139" pageNumber="140" phylum="Arthropoda" rank="genus">
<emphasis id="F087EA9FFF0D68F4FD78F92FCC34F968" box="[655,803,1735,1761]" italics="true" pageId="139" pageNumber="140">Subepimeria</emphasis>
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species ranges between 14 and
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.
</paragraph>
</subSubSection>
<subSubSection id="8AE96506FF0D68F4FF4AF8E3CEB2F8DC" pageId="139" pageNumber="140" type="biology_ecology">
<paragraph id="C24C368DFF0D68F4FF4AF8E3CE31F8AC" blockId="139.[189,294,1803,1829]" box="[189,294,1803,1829]" pageId="139" pageNumber="140">
<heading id="990481E1FF0D68F4FF4AF8E3CE31F8AC" bold="true" box="[189,294,1803,1829]" fontSize="11" level="3" pageId="139" pageNumber="140" reason="3">
<emphasis id="F087EA9FFF0D68F4FF4AF8E3CE31F8AC" bold="true" box="[189,294,1803,1829]" pageId="139" pageNumber="140">Ecology.</emphasis>
</heading>
</paragraph>
<paragraph id="C24C368DFF0D68F4FF4AF8D3CEB2F8DC" blockId="139.[189,421,1851,1878]" box="[189,421,1851,1878]" pageId="139" pageNumber="140">
Benthic,
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52
<specimenCount id="D4F5FD04FF0D68F4FEA4F8D3CEB7F8DC" box="[339,416,1851,1878]" pageId="139" pageNumber="140" type="generic">840 m</specimenCount>
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.
</paragraph>
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<subSubSection id="8AE96506FF0A68F3FF4AFEE3CDEEFEDD" box="[189,761,267,340]" pageId="140" pageNumber="141" type="distribution">
<paragraph id="C24C368DFF0A68F3FF4AFEE3CE41FEAC" blockId="140.[189,342,267,293]" box="[189,342,267,293]" pageId="140" pageNumber="141">
<heading id="990481E1FF0A68F3FF4AFEE3CE41FEAC" bold="true" box="[189,342,267,293]" fontSize="11" level="3" pageId="140" pageNumber="141" reason="3">
<emphasis id="F087EA9FFF0A68F3FF4AFEE3CE41FEAC" bold="true" box="[189,342,267,293]" pageId="140" pageNumber="141">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="C24C368DFF0A68F3FF4AFED2CDEEFEDD" blockId="140.[189,761,314,340]" box="[189,761,314,340]" pageId="140" pageNumber="141">
Circum-Antarctic, as far north as
<collectingCountry id="BAE4761DFF0A68F3FDB0FED2CDE2FEDD" box="[583,757,314,340]" name="South Georgia and the South Sandwich Islands" pageId="140" pageNumber="141">South Georgia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="8AE96506FF0A68F3FF4AFE94CFFAFCFB" pageId="140" pageNumber="141" type="discussion">
<paragraph id="C24C368DFF0A68F3FF4AFE94CE38FE1F" blockId="140.[189,303,380,406]" box="[189,303,380,406]" pageId="140" pageNumber="141">
<heading id="990481E1FF0A68F3FF4AFE94CE38FE1F" bold="true" box="[189,303,380,406]" fontSize="11" level="3" pageId="140" pageNumber="141" reason="3">
<emphasis id="F087EA9FFF0A68F3FF4AFE94CE38FE1F" bold="true" box="[189,303,380,406]" pageId="140" pageNumber="141">Remarks</emphasis>
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Molecular data (COI, 28S) (
<bibRefCitation id="A6624B7CFF0A68F3FDEFFE43CC0EFE4F" author="Verheye" box="[536,793,427,454]" etAl="et al." firstAuthor="Verheye" pageId="140" pageNumber="141" pagination="925 - 945" refId="ref146163" refString="Verheye M., Backeljau T. &amp; d'Udekem d'Acoz C. 2016 a. Looking beneath the tip of the iceberg: diversification of the genus Epimeria on the Antarctic shelf (Crustacea, Amphipoda). In: Gutt J., David B. &amp; Isla E. (eds) High environmental variability and steep biological gradients in the waters off the northern Antarctic Peninsula. Polar Biology 39 (5): 925 - 945, online supplementary material https: // doi. org / 10.1007 / s 00300 - 016 - 1910 - 5" type="journal article" year="2016" yearSuffix="a">Verheye et al. 2016a</bibRefCitation>
) indicate that
<taxonomicName id="05F34D0EFF0A68F3FC3BFE43CB77FE4C" authorityName="Bellan-Santini" authorityYear="1972" box="[972,1120,427,453]" class="Malacostraca" family="Epimeriidae" genus="Subepimeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="140" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="F087EA9FFF0A68F3FC3BFE43CB77FE4C" box="[972,1120,427,453]" italics="true" pageId="140" pageNumber="141">Subepimeria</emphasis>
</taxonomicName>
and
<taxonomicName id="05F34D0EFF0A68F3FB55FE43CA50FE4C" authorityName="dAcoz &amp; Verheye" authorityYear="2017" box="[1186,1351,427,453]" class="Malacostraca" family="Epimeriidae" genus="Epimeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="140" pageNumber="59" phylum="Arthropoda" rank="subGenus" subGenus="Drakepimeria">
<emphasis id="F087EA9FFF0A68F3FB55FE43CA50FE4C" box="[1186,1351,427,453]" italics="true" pageId="140" pageNumber="141">Drakepimeria</emphasis>
</taxonomicName>
are sister clades (
<figureCitation id="5AC82A08FF0A68F3FE91FE27CEC4FE60" box="[358,467,463,489]" captionStart="Fig" captionStartId="551.[189,232,1824,1850]" captionTargetBox="[343,1219,266,1752]" captionTargetId="graphics@551.[343,1120,270,1752]" captionTargetPageId="551" captionText="Fig. 342. Phylogenetic tree obtained by Bayesian analysis of the concatenated COI and 28S sequences, updated from Verheye et al. (2016). Bayesian posterior probabilities and bootstrap values (from the Maximum Likelihood analysis) are indicated above the nodes of interest. Bootstrap values inferior to 50 are not indicated. Besides the species names are indicated the MOTUs codes used in Verheye et al. (2016). The Genbank accession number of the outgroup species: Gnathiphimedia sexdentata (Schellenberg, 1926) are KU870835 (COI) and KU759609 (28S)." figureDoi="http://doi.org/10.5281/zenodo.3856377" httpUri="https://zenodo.org/record/3856377/files/figure.png" pageId="140" pageNumber="141">Fig. 342</figureCitation>
). The morphology of their gnathopods and pereiopods is similar. However,
<taxonomicName id="05F34D0EFF0A68F3FF4AFE1ACE46FD85" authorityName="Bellan-Santini" authorityYear="1972" box="[189,337,498,524]" class="Malacostraca" family="Epimeriidae" genus="Subepimeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="140" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="F087EA9FFF0A68F3FF4AFE1ACE46FD85" box="[189,337,498,524]" italics="true" pageId="140" pageNumber="141">Subepimeria</emphasis>
</taxonomicName>
species are much smaller (&lt;
<quantity id="050B9B68FF0A68F3FD5CFE1BCC16FD84" box="[683,769,499,525]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="140" pageNumber="141" unit="mm" value="20.0">20 mm</quantity>
) and have only one or two (smaller) mid-dorsal teeth and no dorsolateral teeth on pereionites and pleosomites. The lateral carina of coxa 4 is also vestigial in
<taxonomicName id="05F34D0EFF0A68F3FF4AFDD2CE46FDDD" authorityName="Bellan-Santini" authorityYear="1972" box="[189,337,570,596]" class="Malacostraca" family="Epimeriidae" genus="Subepimeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="140" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="F087EA9FFF0A68F3FF4AFDD2CE46FDDD" box="[189,337,570,596]" italics="true" pageId="140" pageNumber="141">Subepimeria</emphasis>
</taxonomicName>
, whilst it is very strongly developed in
<taxonomicName id="05F34D0EFF0A68F3FCDBFDD2CCC6FDDD" authorityName="dAcoz &amp; Verheye" authorityYear="2017" box="[812,977,570,596]" class="Malacostraca" family="Epimeriidae" genus="Epimeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="140" pageNumber="59" phylum="Arthropoda" rank="subGenus" subGenus="Drakepimeria">
<emphasis id="F087EA9FFF0A68F3FCDBFDD2CCC6FDDD" box="[812,977,570,596]" italics="true" pageId="140" pageNumber="141">Drakepimeria</emphasis>
</taxonomicName>
. We believe that these conspicuous morphological differences justify the recognition of distinct subgenera for these clades.The morphological identification of species within the subgenus
<taxonomicName id="05F34D0EFF0A68F3FD27FD69CC73FD12" authorityName="Bellan-Santini" authorityYear="1972" box="[720,868,641,667]" class="Malacostraca" family="Epimeriidae" genus="Subepimeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="140" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="F087EA9FFF0A68F3FD27FD69CC73FD12" box="[720,868,641,667]" italics="true" pageId="140" pageNumber="141">Subepimeria</emphasis>
</taxonomicName>
is very difficult. Interspecific differences are based on ill-defined characters, such as the curves of coxa 4. The species are also rare, which largely prevents the study of individual and allometric differences within species. The link between immatures and adult specimens of
<taxonomicName id="05F34D0EFF0A68F3FE14FD06CC2AFC8E" authorityName="dAcoz &amp; Verheye" authorityYear="2017" box="[483,829,749,775]" class="Malacostraca" family="Epimeriidae" genus="Epimeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="140" pageNumber="141" phylum="Arthropoda" rank="species" species="iota" subGenus="Subepimeria">
<emphasis id="F087EA9FFF0A68F3FE14FD06CD47FC8E" box="[483,592,750,775]" italics="true" pageId="140" pageNumber="141">Epimeria</emphasis>
(
<emphasis id="F087EA9FFF0A68F3FD92FD05CDEEFC8E" box="[613,761,749,775]" italics="true" pageId="140" pageNumber="141">Subepimeria</emphasis>
)
<emphasis id="F087EA9FFF0A68F3FCF8FD06CC2AFC8E" box="[783,829,750,775]" italics="true" pageId="140" pageNumber="141">iota</emphasis>
</taxonomicName>
is tentative as only immature specimens were available for sequencing. As
<taxonomicName id="05F34D0EFF0A68F3FDEBFCF8CDA7FCA3" authorityName="Bellan-Santini" authorityYear="1972" box="[540,688,784,810]" class="Malacostraca" family="Epimeriidae" genus="Subepimeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="140" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="F087EA9FFF0A68F3FDEBFCF8CDA7FCA3" box="[540,688,784,810]" italics="true" pageId="140" pageNumber="141">Subepimeria</emphasis>
</taxonomicName>
species can only be distinguished by a combination of illdefined characters, we delineate their differences in a tabular format (
<tableCitation id="8F710336FF0A68F3FC2BFCDCCB26FCC7" box="[988,1073,820,846]" captionStart="Table 1" captionStartId="143.[189,258,268,294]" captionTargetPageId="143" captionText="Table 1. (continued on next page). Tentative table of character states and distribution for Subepimeria species." httpUri="http://table.plazi.org/id/968C6605FF0968F0FF4AFEE4CE0EFEC1" pageId="140" pageNumber="141" tableUuid="968C6605FF0968F0FF4AFEE4CE0EFEC1">Table 1</tableCitation>
) instead of an identification key.
</paragraph>
</subSubSection>
</treatment>
</document>