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<mods:title id="C3942B542478E501D5FAA6236B0FDC85">Revision of the West-Palaearctic species of the tribe Cerioidini (Diptera, Syrphidae)</mods:title>
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<taxonomicName id="72A7E9BAFFA059327AAAFE767443FDAD" ID-CoL="69M4C" authority="Stackelberg, 1928" authorityName="Stackelberg" authorityYear="1928" box="[151,608,511,538]" class="Insecta" family="Syrphidae" genus="Ceriana" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="caesarea">
<emphasis id="87D34E2BFFA059327AAAFE767443FDAD" bold="true" box="[151,608,511,538]" pageId="15" pageNumber="166">
<emphasis id="87D34E2BFFA059327AAAFE767749FDAE" bold="true" box="[151,362,511,537]" italics="true" pageId="15" pageNumber="166">Ceriana caesarea</emphasis>
(
<bibRefCitation id="D136EFC8FFA059327B44FD89747BFDAD" author="Stackelberg" box="[377,600,512,538]" pageId="15" pageNumber="166" refString="Stackelberg, A. A. 1928. Descriptio species novae generis Cerioides Rd. (Dipt.). Izvestiia Stavropol entomologi Obshchestva., 5, 1 - 2." type="journal article" year="1928">Stackelberg, 1928</bibRefCitation>
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<figureCitation id="2D9C8EBCFFA059327AAAFDAB76D9FD8C" box="[151,250,546,571]" captionStart="FIGURES 1 6" captionStartId="2.[151,264,1946,1968]" captionTargetBox="[211,1367,195,1907]" captionTargetId="figure@2.[211,1370,195,1907]" captionTargetPageId="2" captionText="FIGURES 1 6. Ceriana adults, overall appearance, dorsal view. 1. Ceriana brunettii male, Tajikistan. 2. Ceriana brunettii female, Uzbekistan. 3. Ceriana caesarea male, Turkmenistan. 4. Ceriana caesarea female, Turkmenistan. 5. Ceriana glaebosa male paratype 6 Ceriana glaebosa female paratype. Scale line = 5 mm." httpUri="https://zenodo.org/record/272627/files/figure.png" pageId="15" pageNumber="166">Figs 3, 4</figureCitation>
,
<figureCitation id="2D9C8EBCFFA059327B3BFDAA776DFD8C" box="[262,334,547,571]" captionStart="FIGURES 19 24" captionStartId="5.[151,264,1488,1510]" captionTargetBox="[151,1436,193,1466]" captionTargetId="figure@5.[151,1436,193,1467]" captionTargetPageId="5" captionText="FIGURES 19 24. Ceriana adults, overall appearance, lateral view. 19 Ceriana brunettii male, Tajikistan 20 Ceriana brunettii female, Uzbekistan 21 Ceriana caesarea male, Turkmenistan 22 Ceriana caesarea female, Turkmenistan 23 Ceriana glaebosa male, paratype 24 abdomen Ceriana glaebosa female, paratype. Scale line = 5 mm." httpUri="https://zenodo.org/record/167928/files/figure.png" pageId="15" pageNumber="166">21, 22</figureCitation>
,
<figureCitation id="2D9C8EBCFFA059327B67FDAA775AFD8C" box="[346,377,547,571]" captionStart="FIGURES 37 48" captionStartId="9.[151,264,1866,1888]" captionTargetBox="[178,1354,204,1778]" captionTargetId="figure@9.[153,1433,193,1844]" captionTargetPageId="9" captionText="FIGURES 37 48. Head, frontal view. 37 Ceriana brunettii male, Tajikistan 38 Ceriana caesarea male, Turkmenistan 39 Ceriana conopsoides male, Greece 40 Ceriana conopsoides female, Russia 41 Ceriana glaebosa male paratype 42 Ceriana media male paratype, Afghanistan 43 Ceriana naja male, Tajikistan 44 Ceriana sartorum male, Turkmenistan 45 Ceriana sartorum female, Tajikistan 46 Ceriana skevingtoni male paratype, Azerbaijan Talish 47 Ceriana vespiformis male, Greece 48 Ceriana vespiformis female, Rhodos. Scale line = 2 mm." httpUri="https://zenodo.org/record/167931/files/figure.png" pageId="15" pageNumber="166">38</figureCitation>
,
<figureCitation id="2D9C8EBCFFA059327BB9FDAB7780FD8C" box="[388,419,546,571]" captionStart="FIGURES 49 55. 49 52" captionStartId="11.[151,264,1746,1768]" captionTargetBox="[335,1213,1140,1715]" captionTargetId="figure@11.[313,1273,1134,1724]" captionTargetPageId="11" captionText="FIGURES 49 55. 49 52 Procoxa of Cerioidini species, posterior view: 49 Ceriana conopsoides, male, Russia 50 Ceriana vespiformis, male, Montenegro 51 Primocerioides regale, female, Cyprus 52 Sphiximorpha subsessilis, male, The Netherlands. 53 Basisternum of Ceriana conopsoides, frontal view, male, Russia 54 Mesotrochanter of Ceriana caesarea, ventral view, male, Turkmenistan 55 Basisternum of Sphiximorpha subsessilis, frontal view, male, Greece. Scale line = 0.5 mm (a: 49, 50; b: 51, 52; c: 53 55)." httpUri="https://zenodo.org/record/167932/files/figure.png" pageId="15" pageNumber="166">54</figureCitation>
,
<figureCitation id="2D9C8EBCFFA059327B93FDAB77D4FD8C" box="[430,503,546,571]" captionStart="FIGURES 59 63" captionStartId="16.[151,264,1807,1829]" captionTargetBox="[349,1235,841,1779]" captionTargetId="figure@16.[313,1273,810,1785]" captionTargetPageId="16" captionText="FIGURES 59 63. Male genitalia of Ceriana caesarea, Turkmenistan. 59 Epandrium and hypandrium, lateral view 60 Aedeagus, lateral view 61 Epandrium, dorsal view 62 Epandrium, ventral view 63 Hypandrium, ventral view. Scale line = 0.5 mm (a: 59, 61 63; b: 60). Legends: a epandrium b ventral rim c cercus d apical part of dorsal lobe of surstylus e medial part of dorsal lobe f basal part of dorsal lobe g ventral lobe h membranous part of ventral lobe i surstylar apodeme j apex of hypandrium k superior lobe of hypandrium l lingula m ventro-medial margin of hypandrium n base of hypandrium o stem of aedeagus p head of aedeagus q apex of aedeagus r ventro-medial projection of aedeagus s baso-ventral projection of aedeagus." httpUri="https://zenodo.org/record/167934/files/figure.png" pageId="15" pageNumber="166">5963</figureCitation>
,
<figureCitation id="2D9C8EBCFFA05932783FFDAB7402FD8C" box="[514,545,546,571]" captionStart="FIGURES 64 70" captionStartId="17.[151,264,912,934]" captionTargetBox="[348,1226,227,862]" captionTargetId="figure@17.[313,1273,195,891]" captionTargetPageId="17" captionText="FIGURES 64 70. Metatrochanter of Ceriana species, ventral view. 64 Ceriana brunettii, male, Azerbaijan. 65 Ceriana caesarea male, Turkmenistan 66 Ceriana conopsoides male, Estonia 67 Ceriana glaebosa paratype, male 68 Ceriana media paratype, male 69 Ceriana naja male, Tajikistan 70 Ceriana sartorum male, Kyrgyzstan. Scale line = 0.5 mm (a: 65; b: 67; c: 64, 66, 68 70)." httpUri="https://zenodo.org/record/167935/files/figure.png" pageId="15" pageNumber="166">65</figureCitation>
,
<figureCitation id="2D9C8EBCFFA059327811FDAA7469FD8C" box="[556,586,547,571]" captionStart="FIGURES 83 86" captionStartId="24.[151,264,565,587]" captionTargetBox="[332,1248,219,508]" captionTargetId="figure@24.[313,1273,193,544]" captionTargetPageId="24" captionText="FIGURES 83 86. Mesofemur of Ceriana species, male, ventro-lateral view. 83 Ceriana caesarea, Turkmenistan 84 Ceriana conopsoides, Russia 85 Ceriana media sp. n., paratype 86 Ceriana skevingtoni sp. n., holotype. Scale line = 1 mm." httpUri="https://zenodo.org/record/167938/files/figure.png" pageId="15" pageNumber="166">83</figureCitation>
,
<figureCitation id="2D9C8EBCFFA05932786BFDAA7457FD8C" box="[598,628,547,571]" captionStart="FIGURES 90 93" captionStartId="26.[151,264,860,882]" captionTargetBox="[342,1165,453,779]" captionTargetId="figure@26.[313,1273,411,839]" captionTargetPageId="26" captionText="FIGURES 90 93. Metafemur of Ceriana species, male 90 Ceriana caesarea (with tibia), Uzbekistan 91 Ceriana naja, Tajikistan 92 Ceriana sartorum, Kyrgyzstan 93 Ceriana vespiformis, Montenegro. Scale line = 1 mm." httpUri="https://zenodo.org/record/167940/files/figure.png" pageId="15" pageNumber="166">90</figureCitation>
,
<figureCitation id="2D9C8EBCFFA0593278BDFDAB7489FD8C" box="[640,682,546,571]" captionStart="FIGURES 166 168" captionStartId="49.[151,264,1952,1974]" captionTargetBox="[153,1420,901,1934]" captionTargetId="figure@49.[153,1433,901,1935]" captionTargetPageId="49" captionText="FIGURES 166 168. Distribution maps of the West-Palaearctic Cerioidini. 166 Ceriana brunettii (yellow circle), Ceriana caesarea (blue square) and Ceriana media (red triangle) 167 Ceriana naja (blue circle), Ceriana sartorum (yellow triangle) and Ceriana skevingtoni (red square) 168 Ceriana conopsoides (purple circle)." httpUri="https://zenodo.org/record/167957/files/figure.png" pageId="15" pageNumber="166">166</figureCitation>
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<taxonomicName id="72A7E9BAFFA059327AAAFDE37412FD37" ID-CoL="8BDQK" authority="Stackelberg, 1928: 1" authorityName="Stackelberg" authorityPageNumber="1" authorityYear="1928" box="[151,561,618,640]" class="Insecta" family="Syrphidae" genus="Cerioides" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="caesarea">
<emphasis id="87D34E2BFFA059327AAAFDE37778FD37" box="[151,347,618,640]" italics="true" pageId="15" pageNumber="166">Cerioides caesarea</emphasis>
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: 1
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locality:
<collectingCountry id="CDB0D2A9FFA0593278F2FDE37578FD37" box="[719,859,618,640]" name="Turkmenistan" pageId="15" pageNumber="166">Turkmenistan</collectingCountry>
[HT
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<taxonomicName id="72A7E9BAFFA059327AAAFD0077C5FD28" authority="Sack (1932)" authorityName="Sack" authorityYear="1932" box="[151,486,649,671]" class="Insecta" family="Syrphidae" genus="Cerioides" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="caesarea">
<emphasis id="87D34E2BFFA059327AAAFD007779FD28" box="[151,346,649,671]" italics="true" pageId="15" pageNumber="166">Cerioides caesarea</emphasis>
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<bibRefCitation id="D136EFC8FFA059327B5AFD0077C5FD28" author="Sack" box="[359,486,649,671]" pageId="15" pageNumber="166" refString="Sack, P. (1932) Syrphidae. In: Lindner, E. (Ed.), Die Fliegen der Palaearktischen Region. 31, E. Scheizerbartsche Verlagsbuchhandlung. Stuttgart, 451 pp." type="book" year="1932">Sack (1932)</bibRefCitation>
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<taxonomicName id="72A7E9BAFFA059327AAAFD207428FD09" ID-CoL="69M4C" authority="Violovitsh (1974)" authorityName="Violovitsh" authorityYear="1974" box="[151,523,680,702]" class="Insecta" family="Syrphidae" genus="Ceriana" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="caesarea" status="comb. nov.">
<emphasis id="87D34E2BFFA059327AAAFD207768FD09" box="[151,331,681,702]" italics="true" pageId="15" pageNumber="166">Ceriana caesarea</emphasis>
:
<bibRefCitation id="D136EFC8FFA059327B65FD217428FD09" author="Violovitsh" box="[344,523,680,702]" pageId="15" pageNumber="166" refString="Violovitsh, N. A. (1974) Brief survey of palaearctic species of the genus Ceriana Rafinesque, 1815 (Diptera, Syrphidae). Izvestiȳa Sirbirskogo Otdeleniya Akademii Nauk (Biologicheskoi), 1, 81 - 88. [in Russian, with English summary]." type="journal article" year="1974">Violovitsh (1974)</bibRefCitation>
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<taxonomicNameLabel id="9CE0F350FFA05932782CFD2174A7FD09" box="[529,644,680,702]" pageId="15" pageNumber="166">comb. nov.</taxonomicNameLabel>
,
<bibRefCitation id="D136EFC8FFA0593278B0FD217574FD09" author="Krivosheina" box="[653,855,680,702]" pageId="15" pageNumber="166" refString="Krivosheina, M. G. (2001) To the biology of the xylophilous flies of the genus Ceriana Rafinesque, 1815 (Diptera: Syrphidae) with the redescriptions of the larvae and puparia of C. caesarea Stackelberg and C. naja Violovitsh. International Journal of Dipterology, 12, 57 - 64." type="journal article" year="2001">Krivosheina (2001)</bibRefCitation>
.
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<emphasis id="87D34E2BFFA059327AAAFD63771EFCB4" bold="true" box="[151,317,746,771]" pageId="15" pageNumber="166">Redescription</emphasis>
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<emphasis id="87D34E2BFFA059327B71FD637783FCB4" bold="true" box="[332,416,746,771]" pageId="15" pageNumber="166">MALE</emphasis>
(
<figureCitation id="2D9C8EBCFFA059327B92FD6377D4FCB4" box="[431,503,746,771]" captionStart="FIGURES 1 6" captionStartId="2.[151,264,1946,1968]" captionTargetBox="[211,1367,195,1907]" captionTargetId="figure@2.[211,1370,195,1907]" captionTargetPageId="2" captionText="FIGURES 1 6. Ceriana adults, overall appearance, dorsal view. 1. Ceriana brunettii male, Tajikistan. 2. Ceriana brunettii female, Uzbekistan. 3. Ceriana caesarea male, Turkmenistan. 4. Ceriana caesarea female, Turkmenistan. 5. Ceriana glaebosa male paratype 6 Ceriana glaebosa female paratype. Scale line = 5 mm." httpUri="https://zenodo.org/record/272627/files/figure.png" pageId="15" pageNumber="166">Figs 3</figureCitation>
,
<figureCitation id="2D9C8EBCFFA059327838FD627407FCB4" box="[517,548,747,771]" captionStart="FIGURES 19 24" captionStartId="5.[151,264,1488,1510]" captionTargetBox="[151,1436,193,1466]" captionTargetId="figure@5.[151,1436,193,1467]" captionTargetPageId="5" captionText="FIGURES 19 24. Ceriana adults, overall appearance, lateral view. 19 Ceriana brunettii male, Tajikistan 20 Ceriana brunettii female, Uzbekistan 21 Ceriana caesarea male, Turkmenistan 22 Ceriana caesarea female, Turkmenistan 23 Ceriana glaebosa male, paratype 24 abdomen Ceriana glaebosa female, paratype. Scale line = 5 mm." httpUri="https://zenodo.org/record/167928/files/figure.png" pageId="15" pageNumber="166">21</figureCitation>
). Body length: 12.0
<quantity id="725F3FDCFFA05932792FFD62755AFCB5" box="[786,889,747,771]" metricMagnitude="-2" metricUnit="m" metricValue="1.3199999999999998" pageId="15" pageNumber="166" unit="mm" value="13.2">13.2 mm</quantity>
; wing length: 9.0
<quantity id="725F3FDCFFA059327E76FD627287FCB5" box="[1099,1188,747,771]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="15" pageNumber="166" unit="mm" value="9.2">9.2 mm</quantity>
.
<emphasis id="87D34E2BFFA059327E92FD6372CCFCB4" bold="true" box="[1199,1263,746,771]" pageId="15" pageNumber="166">Head</emphasis>
(
<figureCitation id="2D9C8EBCFFA059327EC3FD637371FCB4" box="[1278,1362,746,771]" captionStart="FIGURES 37 48" captionStartId="9.[151,264,1866,1888]" captionTargetBox="[178,1354,204,1778]" captionTargetId="figure@9.[153,1433,193,1844]" captionTargetPageId="9" captionText="FIGURES 37 48. Head, frontal view. 37 Ceriana brunettii male, Tajikistan 38 Ceriana caesarea male, Turkmenistan 39 Ceriana conopsoides male, Greece 40 Ceriana conopsoides female, Russia 41 Ceriana glaebosa male paratype 42 Ceriana media male paratype, Afghanistan 43 Ceriana naja male, Tajikistan 44 Ceriana sartorum male, Turkmenistan 45 Ceriana sartorum female, Tajikistan 46 Ceriana skevingtoni male paratype, Azerbaijan Talish 47 Ceriana vespiformis male, Greece 48 Ceriana vespiformis female, Rhodos. Scale line = 2 mm." httpUri="https://zenodo.org/record/167931/files/figure.png" pageId="15" pageNumber="166">Fig. 38</figureCitation>
). Face slightly protruding antero-ventrally with low and hardly visible tubercle and slightly concave below antennae. Head 2.52.6 times wider than face just below the antennae; eye contiguity 0.400.48 times as long as length of frons; angle of eyes at eye contiguity 115125o. Face yellow with broad black vitta between eye margin and mouth edge. Frons yellow; vertical triangle and dorsal surface of head capsule yellow; ocellar triangle between ocelli black. Frontal prominence 2.83.2 times longer than wide; relative length of pedicel is as 1.1: 1: 0.921.0. Frontal prominence brown to dark-brown, basally yellow to black with dark-yellow baso-ventral part; antennae and arista black.
<emphasis id="87D34E2BFFA059327ADEFC60771FFBB5" bold="true" box="[227,316,1001,1026]" pageId="15" pageNumber="166">Thorax</emphasis>
. Scutum black with yellow maculae on postpronotum, notopleuron and posterior to notopleuron; a yellow vitta from postalar callus to suture at both sides, sometimes very short and narrow; two oval maculae medially anterior to scutellum. Pleuron black with three yellow maculae, one each on anepisternum, dorsal 1/3 of katepisternum and on the anteroventral corner of anterior anepimeron. Metasternum with short pile and a low, rounded elevation medially on antero-ventral part. Scutellum entirely yellow.
<emphasis id="87D34E2BFFA059327E0AFBF0724CFB25" bold="true" box="[1079,1135,1145,1170]" pageId="15" pageNumber="166">Legs</emphasis>
. Coxae black; trochanter brown-yellow; apico-ventral part of metafemur dark-brown to black; basal 3/4 to entire metatibia yellow. Clear patch of yellow setulae present on antero-basal part of profemur; metatrochanter without setulae (
<figureCitation id="2D9C8EBCFFA059327F08FB4873ACFB6D" box="[1333,1423,1217,1242]" captionStart="FIGURES 64 70" captionStartId="17.[151,264,912,934]" captionTargetBox="[348,1226,227,862]" captionTargetId="figure@17.[313,1273,195,891]" captionTargetPageId="17" captionText="FIGURES 64 70. Metatrochanter of Ceriana species, ventral view. 64 Ceriana brunettii, male, Azerbaijan. 65 Ceriana caesarea male, Turkmenistan 66 Ceriana conopsoides male, Estonia 67 Ceriana glaebosa paratype, male 68 Ceriana media paratype, male 69 Ceriana naja male, Tajikistan 70 Ceriana sartorum male, Kyrgyzstan. Scale line = 0.5 mm (a: 65; b: 67; c: 64, 66, 68 70)." httpUri="https://zenodo.org/record/167935/files/figure.png" pageId="15" pageNumber="166">Fig. 65</figureCitation>
). Mesotrochanter dorsally with rounded appendix; mesofemur with small and narrow flattened area antero-basally (
<figureCitation id="2D9C8EBCFFA059327AA2FA8076D1FA95" box="[159,242,1289,1314]" captionStart="FIGURES 83 86" captionStartId="24.[151,264,565,587]" captionTargetBox="[332,1248,219,508]" captionTargetId="figure@24.[313,1273,193,544]" captionTargetPageId="24" captionText="FIGURES 83 86. Mesofemur of Ceriana species, male, ventro-lateral view. 83 Ceriana caesarea, Turkmenistan 84 Ceriana conopsoides, Russia 85 Ceriana media sp. n., paratype 86 Ceriana skevingtoni sp. n., holotype. Scale line = 1 mm." httpUri="https://zenodo.org/record/167938/files/figure.png" pageId="15" pageNumber="166">Fig. 83</figureCitation>
); metatrochanter a with high keel and a broad rim laterally (
<figureCitation id="2D9C8EBCFFA0593279AFFA8075C5FA95" box="[914,998,1289,1314]" captionStart="FIGURES 49 55. 49 52" captionStartId="11.[151,264,1746,1768]" captionTargetBox="[335,1213,1140,1715]" captionTargetId="figure@11.[313,1273,1134,1724]" captionTargetPageId="11" captionText="FIGURES 49 55. 49 52 Procoxa of Cerioidini species, posterior view: 49 Ceriana conopsoides, male, Russia 50 Ceriana vespiformis, male, Montenegro 51 Primocerioides regale, female, Cyprus 52 Sphiximorpha subsessilis, male, The Netherlands. 53 Basisternum of Ceriana conopsoides, frontal view, male, Russia 54 Mesotrochanter of Ceriana caesarea, ventral view, male, Turkmenistan 55 Basisternum of Sphiximorpha subsessilis, frontal view, male, Greece. Scale line = 0.5 mm (a: 49, 50; b: 51, 52; c: 53 55)." httpUri="https://zenodo.org/record/167932/files/figure.png" pageId="15" pageNumber="166">Fig. 54</figureCitation>
); metatibia without appendix on apicoventral part; metafemur (
<figureCitation id="2D9C8EBCFFA059327B88FAA4742AFAF1" box="[437,521,1325,1350]" captionStart="FIGURES 90 93" captionStartId="26.[151,264,860,882]" captionTargetBox="[342,1165,453,779]" captionTargetId="figure@26.[313,1273,411,839]" captionTargetPageId="26" captionText="FIGURES 90 93. Metafemur of Ceriana species, male 90 Ceriana caesarea (with tibia), Uzbekistan 91 Ceriana naja, Tajikistan 92 Ceriana sartorum, Kyrgyzstan 93 Ceriana vespiformis, Montenegro. Scale line = 1 mm." httpUri="https://zenodo.org/record/167940/files/figure.png" pageId="15" pageNumber="166">Fig. 90</figureCitation>
) elongate and slightly curved.
<emphasis id="87D34E2BFFA059327953FAA4758CFAF1" bold="true" box="[878,943,1325,1350]" pageId="15" pageNumber="166">Wing</emphasis>
. Hyaline with dark-brown maculae, one at cross-vein r-m and apically at vein R2+3. Loop in vein R4+5 very deep without appendix. Cross-vein r-m slightly curved. Membrane entirely but sparsely covered in microtrichia except alula which is only narrowly microtrichose along the anterior margin. Alula broad, 2.52.6 times longer than wide.
<emphasis id="87D34E2BFFA0593279E7FA15726DFA02" bold="true" box="[986,1102,1436,1461]" pageId="15" pageNumber="166">Abdomen</emphasis>
. Black and yellow coloured. Length of tergite I: II: III: IV is as 1: 2.02.3: 1.51.8: 2.12.2. Width of yellow fascia medially: length of tergite of respectively tergite II, III and IV as 1: 3.43.7, 1: 3.34.1 and 1: 2.62.8.Tergite II wider than long, anterior part most narrow; length of tergite II: width of tergite II at respectively anterior: posterior as 1: 1.01.1: 1.1.Tergite I predominantly yellow with large postero-medial black maculae; fascia on tergite II concave medially, straight towards lateral margin; fascia on tergite III straight throughout; fascia on tergite IV medially concave, straight only narrowed laterally and reaching lateral margin. Tergite IV slightly emarginated. Tergite I with low medial elevation; tergites III and IV medially without longitudinal elevation. Tergite III and IV with grey pollinosity as in
<figureCitation id="2D9C8EBCFFA059327AAAF93576FFF962" box="[151,220,1724,1749]" captionStart="FIGURES 1 6" captionStartId="2.[151,264,1946,1968]" captionTargetBox="[211,1367,195,1907]" captionTargetId="figure@2.[211,1370,195,1907]" captionTargetPageId="2" captionText="FIGURES 1 6. Ceriana adults, overall appearance, dorsal view. 1. Ceriana brunettii male, Tajikistan. 2. Ceriana brunettii female, Uzbekistan. 3. Ceriana caesarea male, Turkmenistan. 4. Ceriana caesarea female, Turkmenistan. 5. Ceriana glaebosa male paratype 6 Ceriana glaebosa female paratype. Scale line = 5 mm." httpUri="https://zenodo.org/record/272627/files/figure.png" pageId="15" pageNumber="166">Fig. 3</figureCitation>
. Sternites IIII with yellow posterior fascia.
<emphasis id="87D34E2BFFA0593278E4F9357564F962" bold="true" box="[729,839,1724,1749]" pageId="15" pageNumber="166">Genitalia</emphasis>
. Epandrium with broad ventral rim (
<figureCitation id="2D9C8EBCFFA059327ED6F9357353F962" box="[1259,1392,1724,1749]" captionStart="FIGURES 59 63" captionStartId="16.[151,264,1807,1829]" captionTargetBox="[349,1235,841,1779]" captionTargetId="figure@16.[313,1273,810,1785]" captionTargetPageId="16" captionText="FIGURES 59 63. Male genitalia of Ceriana caesarea, Turkmenistan. 59 Epandrium and hypandrium, lateral view 60 Aedeagus, lateral view 61 Epandrium, dorsal view 62 Epandrium, ventral view 63 Hypandrium, ventral view. Scale line = 0.5 mm (a: 59, 61 63; b: 60). Legends: a epandrium b ventral rim c cercus d apical part of dorsal lobe of surstylus e medial part of dorsal lobe f basal part of dorsal lobe g ventral lobe h membranous part of ventral lobe i surstylar apodeme j apex of hypandrium k superior lobe of hypandrium l lingula m ventro-medial margin of hypandrium n base of hypandrium o stem of aedeagus p head of aedeagus q apex of aedeagus r ventro-medial projection of aedeagus s baso-ventral projection of aedeagus." httpUri="https://zenodo.org/record/167934/files/figure.png" pageId="15" pageNumber="166">Figs 59, 61</figureCitation>
); in dorsal view, cerci broadly sickle shaped, with pile about as long as width of cerci (
<figureCitation id="2D9C8EBCFFA059327E61F9697291F94E" box="[1116,1202,1760,1785]" captionStart="FIGURES 59 63" captionStartId="16.[151,264,1807,1829]" captionTargetBox="[349,1235,841,1779]" captionTargetId="figure@16.[313,1273,810,1785]" captionTargetPageId="16" captionText="FIGURES 59 63. Male genitalia of Ceriana caesarea, Turkmenistan. 59 Epandrium and hypandrium, lateral view 60 Aedeagus, lateral view 61 Epandrium, dorsal view 62 Epandrium, ventral view 63 Hypandrium, ventral view. Scale line = 0.5 mm (a: 59, 61 63; b: 60). Legends: a epandrium b ventral rim c cercus d apical part of dorsal lobe of surstylus e medial part of dorsal lobe f basal part of dorsal lobe g ventral lobe h membranous part of ventral lobe i surstylar apodeme j apex of hypandrium k superior lobe of hypandrium l lingula m ventro-medial margin of hypandrium n base of hypandrium o stem of aedeagus p head of aedeagus q apex of aedeagus r ventro-medial projection of aedeagus s baso-ventral projection of aedeagus." httpUri="https://zenodo.org/record/167934/files/figure.png" pageId="15" pageNumber="166">Fig. 61</figureCitation>
); surstylus bi-lobed, dorsal lobe short broadly elongate medially narrowed, ventral lobe broadly circular with membranous basal part (
<figureCitation id="2D9C8EBCFFA059327AA2F8A176D7F8F6" box="[159,244,1832,1857]" captionStart="FIGURES 59 63" captionStartId="16.[151,264,1807,1829]" captionTargetBox="[349,1235,841,1779]" captionTargetId="figure@16.[313,1273,810,1785]" captionTargetPageId="16" captionText="FIGURES 59 63. Male genitalia of Ceriana caesarea, Turkmenistan. 59 Epandrium and hypandrium, lateral view 60 Aedeagus, lateral view 61 Epandrium, dorsal view 62 Epandrium, ventral view 63 Hypandrium, ventral view. Scale line = 0.5 mm (a: 59, 61 63; b: 60). Legends: a epandrium b ventral rim c cercus d apical part of dorsal lobe of surstylus e medial part of dorsal lobe f basal part of dorsal lobe g ventral lobe h membranous part of ventral lobe i surstylar apodeme j apex of hypandrium k superior lobe of hypandrium l lingula m ventro-medial margin of hypandrium n base of hypandrium o stem of aedeagus p head of aedeagus q apex of aedeagus r ventro-medial projection of aedeagus s baso-ventral projection of aedeagus." httpUri="https://zenodo.org/record/167934/files/figure.png" pageId="15" pageNumber="166">Fig. 59</figureCitation>
); surstylar apodeme separated in two triangular shaped sclerotized parts (
<figureCitation id="2D9C8EBCFFA059327E11F8A172A3F8F6" box="[1068,1152,1832,1857]" captionStart="FIGURES 59 63" captionStartId="16.[151,264,1807,1829]" captionTargetBox="[349,1235,841,1779]" captionTargetId="figure@16.[313,1273,810,1785]" captionTargetPageId="16" captionText="FIGURES 59 63. Male genitalia of Ceriana caesarea, Turkmenistan. 59 Epandrium and hypandrium, lateral view 60 Aedeagus, lateral view 61 Epandrium, dorsal view 62 Epandrium, ventral view 63 Hypandrium, ventral view. Scale line = 0.5 mm (a: 59, 61 63; b: 60). Legends: a epandrium b ventral rim c cercus d apical part of dorsal lobe of surstylus e medial part of dorsal lobe f basal part of dorsal lobe g ventral lobe h membranous part of ventral lobe i surstylar apodeme j apex of hypandrium k superior lobe of hypandrium l lingula m ventro-medial margin of hypandrium n base of hypandrium o stem of aedeagus p head of aedeagus q apex of aedeagus r ventro-medial projection of aedeagus s baso-ventral projection of aedeagus." httpUri="https://zenodo.org/record/167934/files/figure.png" pageId="15" pageNumber="166">Fig. 62</figureCitation>
); hypandrium with broad base and narrower apex; with short, broad and rounded lingula (
<figureCitation id="2D9C8EBCFFA05932795BF8C575CBF8D2" box="[870,1000,1868,1893]" captionStart="FIGURES 59 63" captionStartId="16.[151,264,1807,1829]" captionTargetBox="[349,1235,841,1779]" captionTargetId="figure@16.[313,1273,810,1785]" captionTargetPageId="16" captionText="FIGURES 59 63. Male genitalia of Ceriana caesarea, Turkmenistan. 59 Epandrium and hypandrium, lateral view 60 Aedeagus, lateral view 61 Epandrium, dorsal view 62 Epandrium, ventral view 63 Hypandrium, ventral view. Scale line = 0.5 mm (a: 59, 61 63; b: 60). Legends: a epandrium b ventral rim c cercus d apical part of dorsal lobe of surstylus e medial part of dorsal lobe f basal part of dorsal lobe g ventral lobe h membranous part of ventral lobe i surstylar apodeme j apex of hypandrium k superior lobe of hypandrium l lingula m ventro-medial margin of hypandrium n base of hypandrium o stem of aedeagus p head of aedeagus q apex of aedeagus r ventro-medial projection of aedeagus s baso-ventral projection of aedeagus." httpUri="https://zenodo.org/record/167934/files/figure.png" pageId="15" pageNumber="166">Figs 59, 63</figureCitation>
); in lateral view, superior lobe boxingglove shaped, articulating with hypandrium (
<figureCitation id="2D9C8EBCFFA059327896F8F97526F83E" box="[683,773,1904,1929]" captionStart="FIGURES 59 63" captionStartId="16.[151,264,1807,1829]" captionTargetBox="[349,1235,841,1779]" captionTargetId="figure@16.[313,1273,810,1785]" captionTargetPageId="16" captionText="FIGURES 59 63. Male genitalia of Ceriana caesarea, Turkmenistan. 59 Epandrium and hypandrium, lateral view 60 Aedeagus, lateral view 61 Epandrium, dorsal view 62 Epandrium, ventral view 63 Hypandrium, ventral view. Scale line = 0.5 mm (a: 59, 61 63; b: 60). Legends: a epandrium b ventral rim c cercus d apical part of dorsal lobe of surstylus e medial part of dorsal lobe f basal part of dorsal lobe g ventral lobe h membranous part of ventral lobe i surstylar apodeme j apex of hypandrium k superior lobe of hypandrium l lingula m ventro-medial margin of hypandrium n base of hypandrium o stem of aedeagus p head of aedeagus q apex of aedeagus r ventro-medial projection of aedeagus s baso-ventral projection of aedeagus." httpUri="https://zenodo.org/record/167934/files/figure.png" pageId="15" pageNumber="166">Fig. 59</figureCitation>
); aedeagus with elongated and pointed projection basoventrally, with a narrowly elongate projection with pointed apex, originating from medial part, ventro-medially, apex bird of prey-head shaped (
<figureCitation id="2D9C8EBCFFA059327BC3F8317470F866" box="[510,595,1976,2001]" captionStart="FIGURES 59 63" captionStartId="16.[151,264,1807,1829]" captionTargetBox="[349,1235,841,1779]" captionTargetId="figure@16.[313,1273,810,1785]" captionTargetPageId="16" captionText="FIGURES 59 63. Male genitalia of Ceriana caesarea, Turkmenistan. 59 Epandrium and hypandrium, lateral view 60 Aedeagus, lateral view 61 Epandrium, dorsal view 62 Epandrium, ventral view 63 Hypandrium, ventral view. Scale line = 0.5 mm (a: 59, 61 63; b: 60). Legends: a epandrium b ventral rim c cercus d apical part of dorsal lobe of surstylus e medial part of dorsal lobe f basal part of dorsal lobe g ventral lobe h membranous part of ventral lobe i surstylar apodeme j apex of hypandrium k superior lobe of hypandrium l lingula m ventro-medial margin of hypandrium n base of hypandrium o stem of aedeagus p head of aedeagus q apex of aedeagus r ventro-medial projection of aedeagus s baso-ventral projection of aedeagus." httpUri="https://zenodo.org/record/167934/files/figure.png" pageId="15" pageNumber="166">Fig. 60</figureCitation>
).
<emphasis id="87D34E2BFFA05932785AF83174FCF866" bold="true" box="[615,735,1976,2001]" pageId="15" pageNumber="166">FEMALE</emphasis>
(
<figureCitation id="2D9C8EBCFFA0593278D2F831751AF867" box="[751,825,1976,2001]" captionStart="FIGURES 1 6" captionStartId="2.[151,264,1946,1968]" captionTargetBox="[211,1367,195,1907]" captionTargetId="figure@2.[211,1370,195,1907]" captionTargetPageId="2" captionText="FIGURES 1 6. Ceriana adults, overall appearance, dorsal view. 1. Ceriana brunettii male, Tajikistan. 2. Ceriana brunettii female, Uzbekistan. 3. Ceriana caesarea male, Turkmenistan. 4. Ceriana caesarea female, Turkmenistan. 5. Ceriana glaebosa male paratype 6 Ceriana glaebosa female paratype. Scale line = 5 mm." httpUri="https://zenodo.org/record/272627/files/figure.png" pageId="15" pageNumber="166">Figs 4</figureCitation>
,
<figureCitation id="2D9C8EBCFFA059327978F8317546F867" box="[837,869,1976,2000]" captionStart="FIGURES 19 24" captionStartId="5.[151,264,1488,1510]" captionTargetBox="[151,1436,193,1466]" captionTargetId="figure@5.[151,1436,193,1467]" captionTargetPageId="5" captionText="FIGURES 19 24. Ceriana adults, overall appearance, lateral view. 19 Ceriana brunettii male, Tajikistan 20 Ceriana brunettii female, Uzbekistan 21 Ceriana caesarea male, Turkmenistan 22 Ceriana caesarea female, Turkmenistan 23 Ceriana glaebosa male, paratype 24 abdomen Ceriana glaebosa female, paratype. Scale line = 5 mm." httpUri="https://zenodo.org/record/167928/files/figure.png" pageId="15" pageNumber="166">22</figureCitation>
). Body length:
<quantity id="725F3FDCFFA059327E29F8317298F867" box="[1044,1211,1976,2001]" metricMagnitude="-2" metricUnit="m" metricValue="1.4649999999999999" metricValueMax="1.52" metricValueMin="1.41" pageId="15" pageNumber="166" unit="mm" value="14.649999999999999" valueMax="15.2" valueMin="14.1">14.115.2 mm</quantity>
; wing length:
<quantity id="725F3FDCFFA059327F60F83176DDF843" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" metricValueMax="1.09" metricValueMin="1.05" pageId="15" pageNumber="166" unit="mm" value="10.7" valueMax="10.9" valueMin="10.5">10.5 10.9 mm</quantity>
. Similar to male, except for normal sexual dimorphism and the following characters.
<emphasis id="87D34E2BFFA059327E85F85572D4F842" bold="true" box="[1208,1271,2012,2037]" pageId="15" pageNumber="166">Head</emphasis>
. Head 2.12.2 times wider than face just below the antennae. Frontal prominence 2.63.2 times longer than wide; relative length of pedicel is as 1.11.2: 1: 1.01.1. Face with very narrow medial brown vittae from below frontal prominence towards tubercle. Frons and dorsal surface of head capsule entirely yellow; ocellar triangle and vertex black; small yellow triangular macula anterior of ocellar triangle.
<emphasis id="87D34E2BFFBF592D7938FE8D751EFEAA" bold="true" box="[773,829,260,285]" pageId="16" pageNumber="167">Legs</emphasis>
. Metatrochanter and femora without black spicules.
<emphasis id="87D34E2BFFBF592D7AAAFEAE76FBFEF7" bold="true" box="[151,216,295,320]" pageId="16" pageNumber="167">Wing</emphasis>
. With sub anterior black fascia on cells sc, r1 and anterior half of r2+3. Alula broad, 2.5 times longer than wide.
<emphasis id="87D34E2BFFBF592D7AAAFEC7772FFED0" bold="true" box="[151,268,334,359]" pageId="16" pageNumber="167">Abdomen</emphasis>
. Length of tergite I: II: III: IV: V is as 1: 2.02.1: 1.51.7: 1.82.0: 0.91.1. Width of yellow fascia medially: length of tergite of respectively tergite II, III and IV as 1: 2.93.1, 1: 2.53.1 and 1: 1:2.02.5. Tergite II wider than long, anterior part most narrow; length of tergite II: width of tergite II at respectively anterior: posterior as 1: 1.11.2: 1.31.4.
</paragraph>
<paragraph id="B5189239FFBF592D7AFAFE5775F2FD88" blockId="16.[151,1437,151,755]" pageId="16" pageNumber="167">
<emphasis id="87D34E2BFFBF592D7AFAFE577716FE40" bold="true" box="[199,309,478,503]" pageId="16" pageNumber="167">Variation</emphasis>
.
<bibRefCitation id="D136EFC8FFBF592D7B7BFE577402FE40" author="Krivosheina" box="[326,545,478,503]" pageId="16" pageNumber="167" refString="Krivosheina, M. G. (2001) To the biology of the xylophilous flies of the genus Ceriana Rafinesque, 1815 (Diptera: Syrphidae) with the redescriptions of the larvae and puparia of C. caesarea Stackelberg and C. naja Violovitsh. International Journal of Dipterology, 12, 57 - 64." type="journal article" year="2001">Krivosheina (2001)</bibRefCitation>
discussed the intraspecific variability of
<taxonomicName id="72A7E9BAFFBF592D79CEFE56725BFE40" box="[1011,1144,479,503]" class="Insecta" family="Syrphidae" genus="Ceriana" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="167" phylum="Arthropoda" rank="species" species="caesarea">
<emphasis id="87D34E2BFFBF592D79CEFE56725BFE40" box="[1011,1144,479,503]" italics="true" pageId="16" pageNumber="167">C. caesarea</emphasis>
</taxonomicName>
and, after examination of the male genitalia of specimens with different colouration concluded that they belonged to the same taxon. All the specimens examined in the present study belong to the yellow phenotype.
</paragraph>
</subSubSection>
<subSubSection id="FDBDC1B2FFBF592D7AFAFDC37579FDD4" box="[199,858,586,611]" pageId="16" pageNumber="167" type="materials_examined">
<paragraph id="B5189239FFBF592D7AFAFDC37579FDD4" blockId="16.[151,1437,151,755]" box="[199,858,586,611]" pageId="16" pageNumber="167">
<emphasis id="87D34E2BFFBF592D7AFAFDC3778AFDD4" bold="true" box="[199,425,586,611]" pageId="16" pageNumber="167">Material examined</emphasis>
. 12 Ƌ, 5 ♀ (NHM, ISEA, NBC, ZIN).
</paragraph>
</subSubSection>
<subSubSection id="FDBDC1B2FFBF592D7AFAFDE775ABFD30" box="[199,904,622,647]" pageId="16" pageNumber="167" type="distribution">
<paragraph id="B5189239FFBF592D7AFAFDE775ABFD30" blockId="16.[151,1437,151,755]" box="[199,904,622,647]" pageId="16" pageNumber="167">
<emphasis id="87D34E2BFFBF592D7AFAFDE7777AFD30" bold="true" box="[199,345,622,647]" pageId="16" pageNumber="167">Distribution</emphasis>
(
<figureCitation id="2D9C8EBCFFBF592D7B55FDE777E8FD30" box="[360,459,622,647]" captionStart="FIGURES 166 168" captionStartId="49.[151,264,1952,1974]" captionTargetBox="[153,1420,901,1934]" captionTargetId="figure@49.[153,1433,901,1935]" captionTargetPageId="49" captionText="FIGURES 166 168. Distribution maps of the West-Palaearctic Cerioidini. 166 Ceriana brunettii (yellow circle), Ceriana caesarea (blue square) and Ceriana media (red triangle) 167 Ceriana naja (blue circle), Ceriana sartorum (yellow triangle) and Ceriana skevingtoni (red square) 168 Ceriana conopsoides (purple circle)." httpUri="https://zenodo.org/record/167957/files/figure.png" pageId="16" pageNumber="167">Fig. 166</figureCitation>
).
<collectingCountry id="CDB0D2A9FFBF592D7BE3FDE7746CFD30" box="[478,591,622,647]" name="Tajikistan" pageId="16" pageNumber="167">Tajikistan</collectingCountry>
,
<collectingCountry id="CDB0D2A9FFBF592D7867FDE774DBFD30" box="[602,760,622,647]" name="Turkmenistan" pageId="16" pageNumber="167">Turkmenistan</collectingCountry>
,
<collectingCountry id="CDB0D2A9FFBF592D793EFDE775A6FD30" box="[771,901,622,647]" name="Uzbekistan" pageId="16" pageNumber="167">Uzbekistan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="FDBDC1B2FFBF592D7AFAFD1B7312FD44" pageId="16" pageNumber="167" type="biology_ecology">
<paragraph id="B5189239FFBF592D7AFAFD1B7312FD44" blockId="16.[151,1437,151,755]" pageId="16" pageNumber="167">
<emphasis id="87D34E2BFFBF592D7AFAFD1B773CFD1C" bold="true" box="[199,287,658,683]" pageId="16" pageNumber="167">Biology</emphasis>
. The immature stages of this species are found under the bark and in water-filled holes of
<taxonomicName id="72A7E9BAFFBF592D7F7DFD1B7734FD78" class="Magnoliopsida" family="Salicaceae" genus="Populus" kingdom="Plantae" order="Malpighiales" pageId="16" pageNumber="167" phylum="Tracheophyta" rank="species" species="diversifolia">
<emphasis id="87D34E2BFFBF592D7F7DFD1B7734FD78" italics="true" pageId="16" pageNumber="167">Populus diversifolia</emphasis>
</taxonomicName>
; the puparium is described (
<bibRefCitation id="D136EFC8FFBF592D7846FD3F7572FD78" author="Krivosheina" box="[635,849,694,719]" pageId="16" pageNumber="167" refString="Krivosheina, M. G. (2001) To the biology of the xylophilous flies of the genus Ceriana Rafinesque, 1815 (Diptera: Syrphidae) with the redescriptions of the larvae and puparia of C. caesarea Stackelberg and C. naja Violovitsh. International Journal of Dipterology, 12, 57 - 64." type="journal article" year="2001">Krivosheina 2001</bibRefCitation>
). One female was found in riparian forests and woodlands. Adults fly from early April to late June. Adults have been observed visiting
<taxonomicName id="72A7E9BAFFBF592D7E4FFD5572ECFD44" box="[1138,1231,732,755]" class="Magnoliopsida" family="Tamaricaceae" genus="Tamarix" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" order="Caryophyllales" pageId="16" pageNumber="167" phylum="Tracheophyta" rank="genus">
<emphasis id="87D34E2BFFBF592D7E4FFD5572ECFD44" box="[1138,1231,732,755]" italics="true" pageId="16" pageNumber="167">Tamarix</emphasis>
</taxonomicName>
flowers.
</paragraph>
</subSubSection>
</treatment>
</document>