treatments-xml/data/2E/3E/87/2E3E879CB750FFC0C580A42BFBB5F833.xml

<document id="A1D4E044B67BAD72B8C8A482D57CD326" ID-DOI="10.11646/zootaxa.3368.1.9" ID-ISSN="1175-5326" ID-Zenodo-Dep="5252377" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629789103944" checkinUser="felipe" docAuthor="Chang, Cheon Young &amp; Lee, Jimin" docDate="2012" docId="2E3E879CB750FFC0C580A42BFBB5F833" docLanguage="en" docName="zt03368p210.pdf" docOrigin="Zootaxa 3368 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.3368.1.9" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Halicyclops pumilus Chang &amp; Lee 2012, sp. nov." docType="treatment" docVersion="3" lastPageNumber="209" masterDocId="D207FFE4B758FFCCC517A647FFF1FFE3" masterDocTitle="Two new species of Halicyclops (Copepoda, Cyclopoida) from the estuarine interstitial waters in South Korea *" masterLastPageNumber="210" masterPageNumber="197" pageNumber="205" updateTime="1699112272417" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="9233E635BFFEEB39C9CFDDD8563CE2DB">Two new species of Halicyclops (Copepoda, Cyclopoida) from the estuarine interstitial waters in South Korea *</mods:title>
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<mods:namePart id="BFBD55F41FA440DDEE38C9F51CFACF1D">Chang, Cheon Young</mods:namePart>
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<mods:namePart id="6A49F2D62439AD8C263754EEFC42E1AD">Lee, Jimin</mods:namePart>
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<treatment id="2E3E879CB750FFC0C580A42BFBB5F833" LSID="urn:lsid:plazi:treatment:2E3E879CB750FFC0C580A42BFBB5F833" httpUri="http://treatment.plazi.org/id/2E3E879CB750FFC0C580A42BFBB5F833" lastPageId="12" lastPageNumber="209" pageId="8" pageNumber="205">
<subSubSection id="EE8D6501B750FFC4C580A42BFE03FD66" box="[151,498,619,646]" pageId="8" pageNumber="205" type="nomenclature">
<paragraph id="A628368AB750FFC4C580A42BFE03FD66" blockId="8.[151,498,619,682]" box="[151,498,619,646]" pageId="8" pageNumber="205">
<heading id="FD6081E6B750FFC4C580A42BFE03FD66" bold="true" box="[151,498,619,646]" fontSize="11" level="1" pageId="8" pageNumber="205" reason="1">
<emphasis id="94E3EA98B750FFC4C580A42BFE03FD66" bold="true" box="[151,498,619,646]" pageId="8" pageNumber="205">
<taxonomicName id="61974D09B750FFC4C580A42BFE7CFD65" authority="Chang &amp; Lee, 2012" authorityName="Chang &amp; Lee" authorityYear="2012" box="[151,397,620,646]" class="Hexanauplia" family="Halicyclopidae" genus="Halicyclops" kingdom="Animalia" order="Cyclopoida" pageId="8" pageNumber="205" phylum="Arthropoda" rank="species" species="pumilus" status="sp. nov.">
<emphasis id="94E3EA98B750FFC4C580A42BFE7CFD65" bold="true" box="[151,397,620,646]" italics="true" pageId="8" pageNumber="205">Halicyclops pumilus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8FD057E3B750FFC4C482A42CFE03FD66" box="[405,498,619,645]" pageId="8" pageNumber="205" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="EE8D6501B750FFC4C580A4D5FEFBFD49" box="[151,266,658,682]" pageId="8" pageNumber="205" type="description">
<paragraph id="A628368AB750FFC4C580A4D5FEFBFD49" blockId="8.[151,498,619,682]" box="[151,266,658,682]" pageId="8" pageNumber="205">
(
<figureCitation id="3EAC2A0FB750FFC4C588A4D5FF14FD49" box="[159,229,658,682]" captionStart="FIGURE 5" captionStartId="9.[151,250,1970,1993]" captionTargetBox="[217,1373,192,1948]" captionTargetId="figure-43@9.[191,1396,181,1953]" captionTargetPageId="9" captionText="FIGURE 5. Halicyclops pumilus sp. nov., female. A, habitus, dorsal; B, urosome, dorsal; C, P5 exopod. Scales: A, B = 100 µm; C = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5252387" httpUri="https://zenodo.org/record/5252387/files/figure.png" pageId="8" pageNumber="205">Figs 5</figureCitation>
–7)
</paragraph>
</subSubSection>
<subSubSection id="EE8D6501B750FFC4C580A49DFBE3FC45" pageId="8" pageNumber="205" type="materials_examined">
<paragraph id="A628368AB750FFC4C580A49DFAF0FD11" blockId="8.[151,1436,729,1982]" box="[151,1281,729,755]" pageId="8" pageNumber="205">
<materialsCitation id="16FF3CD7B750FFC4C580A49DFB0CFD11" box="[151,1277,729,755]" country="South Korea" latitude="34.9844" location="Estuary of Seomjin River" longLatPrecision="1" longitude="127.77157" municipality="Gwangyang" pageId="8" pageNumber="205" specimenCount="1" typeStatus="holotype">
<emphasis id="94E3EA98B750FFC4C580A49DFEC4FD10" bold="true" box="[151,309,730,755]" inLineHeading="true" pageId="8" pageNumber="205" reason="1">
<typeStatus id="792C8828B750FFC4C580A49DFF23FD11" box="[151,210,730,754]" pageId="8" pageNumber="205">Type</typeStatus>
locality.
</emphasis>
<location id="A3486051B750FFC4C42BA49CFDAAFD11" LSID="urn:lsid:plazi:treatment:2E3E879CB750FFC0C580A42BFBB5F833:A3486051B750FFC4C42BA49CFDAAFD11" box="[316,603,730,755]" country="South Korea" latitude="34.9844" longLatPrecision="1" longitude="127.77157" municipality="Gwangyang" name="Estuary of Seomjin River" pageId="8" pageNumber="205">Estuary of Seomjin River</location>
, 
<collectingMunicipality id="464CACF0B750FFC4C771A49DFD01FD11" box="[614,752,730,754]" pageId="8" pageNumber="205">Gwangyang</collectingMunicipality>
, 
<collectingCountry id="DE80761AB750FFC4C7ECA49DFC7AFD11" box="[763,907,730,754]" name="South Korea" pageId="8" pageNumber="205">South Korea</collectingCountry>
, 
<geoCoordinate id="C3A3504DB750FFC4C681A49DFBCEFD11" box="[918,1087,729,754]" degrees="34" direction="north" minutes="59" orientation="latitude" pageId="8" pageNumber="205" precision="1" seconds="03.85" value="34.9844">34°59′03.85″N</geoCoordinate>
<geoCoordinate id="C3A3504DB750FFC4C151A49DFB0CFD11" box="[1094,1277,729,754]" degrees="127" direction="east" minutes="46" orientation="longitude" pageId="8" pageNumber="205" precision="1" seconds="17.62" value="127.77157">127°46′17.62″E</geoCoordinate>
</materialsCitation>
.
</paragraph>
<paragraph id="A628368AB750FFC4C5D0A4B8FB05FCD9" blockId="8.[151,1436,729,1982]" pageId="8" pageNumber="205">
Type material. 
<typeStatus id="792C8828B750FFC4C464A4B9FE2FFCF5" box="[371,478,766,790]" pageId="8" pageNumber="205" type="holotype">Holotype</typeStatus>
, intact female in alcohol (NIBRINV0000245093). 
<typeStatus id="792C8828B750FFC4C132A4B8FB67FCF4" box="[1061,1174,767,791]" pageId="8" pageNumber="205" type="paratype">Paratypes</typeStatus>
: two dissected females (DB20038, 20039). All collected at the type locality, 
<date id="D229104AB750FFC4C7FCA565FC58FCD9" box="[747,937,802,826]" pageId="8" pageNumber="205" value="1995-10-17">17 October 1995</date>
, leg. C.Y. Chang &amp; J.M. Lee.
</paragraph>
<paragraph id="A628368AB750FFC4C5D0A501FBE3FC45" blockId="8.[151,1436,729,1982]" pageId="8" pageNumber="205">
<emphasis id="94E3EA98B750FFC4C5D0A501FDC7FCBC" bold="true" box="[199,566,838,863]" inLineHeading="true" pageId="8" pageNumber="205" reason="1">Additional material examined.</emphasis>
<materialsCitation id="16FF3CD7B750FFC4C729A501FDF1FC61" collectingDate="1988-05-30" country="South Korea" latitude="37.4876" location="Three" longLatPrecision="1" longitude="126.621895" municipality="Jakyakdo Island" pageId="8" pageNumber="205" specimenCount="1" stateProvince="Incheon">
<location id="A3486051B750FFC4C729A501FD8EFCBD" LSID="urn:lsid:plazi:treatment:2E3E879CB750FFC0C580A42BFBB5F833:A3486051B750FFC4C729A501FD8EFCBD" box="[574,639,838,862]" country="South Korea" latitude="37.4876" longLatPrecision="1" longitude="126.621895" municipality="Jakyakdo Island" name="Three" pageId="8" pageNumber="205" stateProvince="Incheon">Three</location>
females from 
<collectingMunicipality id="464CACF0B750FFC4C631A501FC2EFCBD" box="[806,991,838,862]" pageId="8" pageNumber="205">Jakyakdo Island</collectingMunicipality>
, 
<collectingRegion id="6453F868B750FFC4C6FBA501FBB8FCBD" box="[1004,1097,838,862]" country="South Korea" name="Incheon" pageId="8" pageNumber="205">Incheon</collectingRegion>
, 
<collectingCountry id="DE80761AB750FFC4C141A501FB17FCBD" box="[1110,1254,838,862]" name="South Korea" pageId="8" pageNumber="205">South Korea</collectingCountry>
, 
<geoCoordinate id="C3A3504DB750FFC4C1E4A501FA6DFCBD" box="[1267,1436,837,862]" degrees="37" direction="north" minutes="29" orientation="latitude" pageId="8" pageNumber="205" precision="1" seconds="15.35" value="37.4876">37°29′15.35″N</geoCoordinate>
<geoCoordinate id="C3A3504DB750FFC4C580A52DFEBDFC61" box="[151,332,873,898]" degrees="126" direction="east" minutes="37" orientation="longitude" pageId="8" pageNumber="205" precision="1" seconds="18.82" value="126.621895">126°37′18.82″E</geoCoordinate>
, 
<date id="D229104AB750FFC4C44AA52DFDF1FC61" box="[349,512,874,899]" pageId="8" pageNumber="205" value="1988-05-30">
<collectingDate id="C26DE9A2B750FFC4C44AA52DFDF1FC61" box="[349,512,874,899]" pageId="8" pageNumber="205" value="1988-05-30">30 May 1988</collectingDate>
</date>
</materialsCitation>
, 
<materialsCitation id="16FF3CD7B750FFC4C707A52DFBE3FC45" collectingDate="1995-10-21" collectorName="C. Y. Chang. Four &amp; C. Y. Chang &amp; J. M. Lee." country="South Korea" latitude="34.631165" location="Sumunpo Beach" longLatPrecision="1" longitude="127.02794" municipality="Boseong" pageId="8" pageNumber="205" specimenCount="1">
<emphasis id="94E3EA98B750FFC4C707A52DFDC3FC61" box="[528,562,874,898]" italics="true" pageId="8" pageNumber="205">leg</emphasis>
<collectorName id="0B62535CB750FFC4C729A52DFCFDFC61" box="[574,780,874,898]" pageId="8" pageNumber="205">C.Y. Chang. Four</collectorName>
females from 
<location id="A3486051B750FFC4C6A9A52DFB8FFC61" LSID="urn:lsid:plazi:treatment:2E3E879CB750FFC0C580A42BFBB5F833:A3486051B750FFC4C6A9A52DFB8FFC61" box="[958,1150,874,898]" country="South Korea" latitude="34.631165" longLatPrecision="1" longitude="127.02794" municipality="Boseong" name="Sumunpo Beach" pageId="8" pageNumber="205">Sumunpo Beach</location>
, 
<collectingMunicipality id="464CACF0B750FFC4C199A52CFB02FC60" box="[1166,1267,875,899]" pageId="8" pageNumber="205">Boseong</collectingMunicipality>
, 
<collectingCountry id="DE80761AB750FFC4C014A52DFA66FC61" box="[1283,1431,874,898]" name="South Korea" pageId="8" pageNumber="205">South Korea</collectingCountry>
, 
<geoCoordinate id="C3A3504DB750FFC4C580A5C9FECEFC45" box="[151,319,909,934]" degrees="34" direction="north" minutes="37" orientation="latitude" pageId="8" pageNumber="205" precision="1" seconds="52.20" value="34.631165">34°37′52.20″N</geoCoordinate>
<geoCoordinate id="C3A3504DB750FFC4C451A5C9FE0DFC45" box="[326,508,909,934]" degrees="127" direction="east" minutes="01" orientation="longitude" pageId="8" pageNumber="205" precision="1" seconds="40.56" value="127.02794">127°01′40.56″E</geoCoordinate>
, 
<date id="D229104AB750FFC4C71FA5C9FD37FC45" box="[520,710,910,934]" pageId="8" pageNumber="205" value="1995-10-21">
<collectingDate id="C26DE9A2B750FFC4C71FA5C9FD37FC45" box="[520,710,910,934]" pageId="8" pageNumber="205" value="1995-10-21">21 October 1995</collectingDate>
</date>
, 
<emphasis id="94E3EA98B750FFC4C7C4A5C9FD04FC45" box="[723,757,910,934]" italics="true" pageId="8" pageNumber="205">leg</emphasis>
. 
<collectorName id="0B62535CB750FFC4C615A5C9FC75FC45" box="[770,900,910,934]" pageId="8" pageNumber="205">C.Y. Chang</collectorName>
&amp; 
<collectorName id="0B62535CB750FFC4C6B0A5C8FBE3FC45" box="[935,1042,911,934]" pageId="8" pageNumber="205">J.M. Lee.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="EE8D6501B750FFC4C5D0A5F5FE7EFC11" pageId="8" pageNumber="205" type="etymology">
<paragraph id="A628368AB750FFC4C5D0A5F5FE7EFC11" blockId="8.[151,1436,729,1982]" pageId="8" pageNumber="205">
<emphasis id="94E3EA98B750FFC4C5D0A5F5FEBFFC29" bold="true" box="[199,334,946,970]" inLineHeading="true" pageId="8" pageNumber="205" reason="1">Etymology.</emphasis>
The specific name comes from the Latin 
<taxonomicName id="61974D09B750FFC4C630A5F5FC71FC29" authorityName="Chang &amp; Lee" authorityYear="2012" box="[807,896,946,970]" class="Hexanauplia" family="Halicyclopidae" genus="Halicyclops" kingdom="Animalia" order="Cyclopoida" pageId="8" pageNumber="205" phylum="Arthropoda" rank="species" species="pumilus">
<emphasis id="94E3EA98B750FFC4C630A5F5FC71FC29" box="[807,896,946,970]" italics="true" pageId="8" pageNumber="205">pumilus</emphasis>
</taxonomicName>
, meaning dwarf, which refers to the diminutive habitus of the species.
</paragraph>
</subSubSection>
<subSubSection id="EE8D6501B750FFC7C5D0A5BAFEA5FAA6" lastPageId="11" lastPageNumber="208" pageId="8" pageNumber="205" type="description">
<paragraph id="A628368AB750FFC4C5D0A5BAFEBDFAD5" blockId="8.[151,1436,729,1982]" pageId="8" pageNumber="205">
<emphasis id="94E3EA98B750FFC4C5D0A5BAFEA9FBF5" bold="true" box="[199,344,1021,1046]" inLineHeading="true" pageId="8" pageNumber="205" reason="1">Description.</emphasis>
Female. Body (
<figureCitation id="3EAC2A0FB750FFC4C718A5B9FD99FBF5" box="[527,616,1022,1046]" captionStart="FIGURE 5" captionStartId="9.[151,250,1970,1993]" captionTargetBox="[217,1373,192,1948]" captionTargetId="figure-43@9.[191,1396,181,1953]" captionTargetPageId="9" captionText="FIGURE 5. Halicyclops pumilus sp. nov., female. A, habitus, dorsal; B, urosome, dorsal; C, P5 exopod. Scales: A, B = 100 µm; C = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5252387" httpUri="https://zenodo.org/record/5252387/files/figure.png" pageId="8" pageNumber="205">Fig. 5A</figureCitation>
) very small, 387.7 µm long (m = 383.6, N = 8), excluding caudal setae; maximum width at posterior end of cephalothorax, 136.9 µm; L/W about 2.83. Body a little flattened dorsoventrally. Prosome comprising cephalothorax incorporating first pedigerous somite and 3 free pedigerous somites; posterior margins of prosomites serrated; outer distal corners of prosomites weakly expanded. Cephalothorax bell-shaped, L/W about 1.06; ovoid integumental window (or depression) present; 20–24 sensilla scattered on dorsal and lateral surfaces. Rostral expansion rarely visible from dorsal view. Genital double-somite (
<figureCitation id="3EAC2A0FB750FFC4C588A291FF09FB0D" box="[159,248,1238,1262]" captionStart="FIGURE 5" captionStartId="9.[151,250,1970,1993]" captionTargetBox="[217,1373,192,1948]" captionTargetId="figure-43@9.[191,1396,181,1953]" captionTargetPageId="9" captionText="FIGURE 5. Halicyclops pumilus sp. nov., female. A, habitus, dorsal; B, urosome, dorsal; C, P5 exopod. Scales: A, B = 100 µm; C = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5252387" httpUri="https://zenodo.org/record/5252387/files/figure.png" pageId="8" pageNumber="205">Fig. 5B</figureCitation>
) a little longer than wide; lateral sides not expanded or swollen, lacking spinous processes. Mid-dorsal hyaline fringe of preanal somite strongly protruded, with 6 acute teeth along posterior margin, far extending over anal operculum.
</paragraph>
<paragraph id="A628368AB750FFC4C5D0A304FDE8F9ED" blockId="8.[151,1436,729,1982]" pageId="8" pageNumber="205">
Fu (
<figureCitation id="3EAC2A0FB750FFC4C5E2A305FEBAFAB9" box="[245,331,1346,1370]" captionStart="FIGURE 5" captionStartId="9.[151,250,1970,1993]" captionTargetBox="[217,1373,192,1948]" captionTargetId="figure-43@9.[191,1396,181,1953]" captionTargetPageId="9" captionText="FIGURE 5. Halicyclops pumilus sp. nov., female. A, habitus, dorsal; B, urosome, dorsal; C, P5 exopod. Scales: A, B = 100 µm; C = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5252387" httpUri="https://zenodo.org/record/5252387/files/figure.png" pageId="8" pageNumber="205">Fig. 5B</figureCitation>
) elongate, about 1.7–1.9 times (m = 1.78, N = 8) longer than wide; divergent posteriorly. Lateral caudal seta lying about halfway and a little dorsally. Inner caudal seta minute. Outer caudal seta 1.76 times longer than ramus. Dorsal caudal seta arising from distal end of longitudinal protuberance, 3.15 times longer than ramus, and 1.78 times longer than outer caudal seta. Small cuticular tube present outside the longitudinal protuberance, with a pore on its tip. Both inner and outer terminal caudal setae with breaking planes; proximal third bare, second third pinnate, distal third plumose.
</paragraph>
<paragraph id="A628368AB750FFC4C5D0A05DFAACF97D" blockId="8.[151,1436,729,1982]" pageId="8" pageNumber="205">
A1 (
<figureCitation id="3EAC2A0FB750FFC4C5ECA05DFEA4F9D1" box="[251,341,1562,1586]" captionStart="FIGURE 6" captionStartId="10.[151,250,1968,1991]" captionTargetBox="[171,1423,187,1943]" captionTargetId="figure-43@10.[151,1436,171,1948]" captionTargetPageId="10" captionText="FIGURE 6. Halicyclops pumilus sp. nov., female. A, A1; B, A2; C, mandible; D, maxillule; E, maxilla; F, maxilliped. Scales: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.5252391" httpUri="https://zenodo.org/record/5252391/files/figure.png" pageId="8" pageNumber="205">Fig. 6A</figureCitation>
) not reaching to halfway of cephalothorax; 6-segmented. Fourth segment 1.74 times as long as wide; last segment 2.60 times longer than wide. Second segment bearing 12 setae, including 1 plumose and 1 bipinnate setae. Third segment shortest, bearing 5 setae including 1 short spiniform seta distally. Aesthetasc each arising from fourth and last segment distally. Seta formula: 8, 12, 5, 6 (+ 1 aesthetasc), 2, 10 (+ 1 aesthetasc).
</paragraph>
<paragraph id="A628368AB750FFC4C5D0A0EDFE43F8B1" blockId="8.[151,1436,729,1982]" pageId="8" pageNumber="205">
A2 (
<figureCitation id="3EAC2A0FB750FFC4C5EEA0EDFEBEF921" box="[249,335,1706,1730]" captionStart="FIGURE 6" captionStartId="10.[151,250,1968,1991]" captionTargetBox="[171,1423,187,1943]" captionTargetId="figure-43@10.[151,1436,171,1948]" captionTargetPageId="10" captionText="FIGURE 6. Halicyclops pumilus sp. nov., female. A, A1; B, A2; C, mandible; D, maxillule; E, maxilla; F, maxilliped. Scales: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.5252391" httpUri="https://zenodo.org/record/5252391/files/figure.png" pageId="8" pageNumber="205">Fig. 6B</figureCitation>
) 3-segmented, comprising coxobasis and 2-segmented endopod. Coxobasis unornamented, armed with 2 inner distal setae, lacking outer distal seta representing exopod. First endopodal segment bearing 1 inner seta, with smooth margins lacking spinules. Second endopodal segment 3.1 times as long as wide, about 1.5 times longer than first endopodal segment; ornamented with a few spinules on caudal face proximally; armed with 5 lateral and 7 apical setae.
</paragraph>
<paragraph id="A628368AB750FFC4C5D0A119FBDBF85D" blockId="8.[151,1436,729,1982]" pageId="8" pageNumber="205">
Mandible (
<figureCitation id="3EAC2A0FB750FFC4C453A119FE6FF895" box="[324,414,1886,1910]" captionStart="FIGURE 6" captionStartId="10.[151,250,1968,1991]" captionTargetBox="[171,1423,187,1943]" captionTargetId="figure-43@10.[151,1436,171,1948]" captionTargetPageId="10" captionText="FIGURE 6. Halicyclops pumilus sp. nov., female. A, A1; B, A2; C, mandible; D, maxillule; E, maxilla; F, maxilliped. Scales: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.5252391" httpUri="https://zenodo.org/record/5252391/files/figure.png" pageId="8" pageNumber="205">Fig. 6C</figureCitation>
), with well-developed coxal gnathobase, armed with 5 smooth teeth, 4 slender spiniform setae and 1 strong pinnate seta along cutting edge, flanking 1 outer distal pinnate seta. Palp very reduced, represented by 2 naked setae on small protuberance, of which shorter one minute.
</paragraph>
<caption id="F2E86602B751FFC5C580A1F5FECAF804" ID-DOI="http://doi.org/10.5281/zenodo.5252387" ID-Zenodo-Dep="5252387" httpUri="https://zenodo.org/record/5252387/files/figure.png" pageId="9" pageNumber="206" startId="9.[151,250,1970,1993]" targetBox="[217,1373,192,1948]" targetPageId="9">
<paragraph id="A628368AB751FFC5C580A1F5FECAF804" blockId="9.[151,1436,1970,2023]" pageId="9" pageNumber="206">
<emphasis id="94E3EA98B751FFC5C580A1F5FEE5F82B" bold="true" box="[151,276,1970,1993]" pageId="9" pageNumber="206">FIGURE 5.</emphasis>
<taxonomicName id="61974D09B751FFC5C40BA1F4FE1CF82A" authority="Chang &amp; Lee, 2012" authorityName="Chang &amp; Lee" authorityYear="2012" box="[284,493,1971,1993]" class="Hexanauplia" family="Halicyclopidae" genus="Halicyclops" kingdom="Animalia" order="Cyclopoida" pageId="9" pageNumber="206" phylum="Arthropoda" rank="species" species="pumilus" status="sp. nov.">
<emphasis id="94E3EA98B751FFC5C40BA1F4FE1CF82A" box="[284,493,1971,1993]" italics="true" pageId="9" pageNumber="206">Halicyclops pumilus</emphasis>
</taxonomicName>
<emphasis id="94E3EA98B751FFC5C4E3A1F4FDB7F82B" bold="true" box="[500,582,1971,1992]" pageId="9" pageNumber="206">
<taxonomicNameLabel id="8FD057E3B751FFC5C4E3A1F4FDB7F82B" box="[500,582,1971,1992]" pageId="9" pageNumber="206" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female. A, habitus, dorsal; B, urosome, dorsal; C, P5 exopod. Scales: A, B = 100 µm; C = 20 µm.
</paragraph>
</caption>
<caption id="F2E86602B752FFC6C580A1F7FF11F809" ID-DOI="http://doi.org/10.5281/zenodo.5252391" ID-Zenodo-Dep="5252391" httpUri="https://zenodo.org/record/5252391/files/figure.png" pageId="10" pageNumber="207" startId="10.[151,250,1968,1991]" targetBox="[171,1423,187,1943]" targetPageId="10">
<paragraph id="A628368AB752FFC6C580A1F7FF11F809" blockId="10.[151,1437,1968,2026]" pageId="10" pageNumber="207">
<emphasis id="94E3EA98B752FFC6C580A1F7FEE2F824" bold="true" box="[151,275,1968,1991]" pageId="10" pageNumber="207">FIGURE 6.</emphasis>
<taxonomicName id="61974D09B752FFC6C40EA1F6FE18F824" authority="Chang &amp; Lee, 2012" authorityName="Chang &amp; Lee" authorityYear="2012" box="[281,489,1969,1991]" class="Hexanauplia" family="Halicyclopidae" genus="Halicyclops" kingdom="Animalia" order="Cyclopoida" pageId="10" pageNumber="207" phylum="Arthropoda" rank="species" species="pumilus" status="sp. nov.">
<emphasis id="94E3EA98B752FFC6C40EA1F6FE18F824" box="[281,489,1969,1991]" italics="true" pageId="10" pageNumber="207">Halicyclops pumilus</emphasis>
</taxonomicName>
<emphasis id="94E3EA98B752FFC6C4E7A1F6FDB1F825" bold="true" box="[496,576,1969,1990]" pageId="10" pageNumber="207">
<taxonomicNameLabel id="8FD057E3B752FFC6C4E7A1F6FDB1F825" box="[496,576,1969,1990]" pageId="10" pageNumber="207" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female. A, A1; B, A2; C, mandible; D, maxillule; E, maxilla; F, maxilliped. Scales: 50 µm.
</paragraph>
</caption>
<paragraph id="A628368AB753FFC7C5D0A6D0FA9DFEF8" blockId="11.[151,1437,151,728]" pageId="11" pageNumber="208">
Maxillule (
<figureCitation id="3EAC2A0FB753FFC7C453A6D0FE6EFF4C" box="[324,415,151,175]" captionStart="FIGURE 6" captionStartId="10.[151,250,1968,1991]" captionTargetBox="[171,1423,187,1943]" captionTargetId="figure-43@10.[151,1436,171,1948]" captionTargetPageId="10" captionText="FIGURE 6. Halicyclops pumilus sp. nov., female. A, A1; B, A2; C, mandible; D, maxillule; E, maxilla; F, maxilliped. Scales: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.5252391" httpUri="https://zenodo.org/record/5252391/files/figure.png" pageId="11" pageNumber="208">Fig. 6D</figureCitation>
), praecoxal arthrite bearing 4 strong spines of which innermost one ornamented with 1–2 accessory setae proximally; 7 elements with various shapes present along inner margin, including 1 strong proximalmost projection. Palp 2-segmented; coxobasis bearing 1 strong spinulose and 2 slender naked setae distally, plus 1 outer pinnate seta representing exopod; distal segment, representing endopod, with 3 long plumose setae.
</paragraph>
<paragraph id="A628368AB753FFC7C5D0A760FA88FE48" blockId="11.[151,1437,151,728]" pageId="11" pageNumber="208">
Maxilla (
<figureCitation id="3EAC2A0FB753FFC7C439A760FE73FEDC" box="[302,386,295,319]" captionStart="FIGURE 6" captionStartId="10.[151,250,1968,1991]" captionTargetBox="[171,1423,187,1943]" captionTargetId="figure-43@10.[151,1436,171,1948]" captionTargetPageId="10" captionText="FIGURE 6. Halicyclops pumilus sp. nov., female. A, A1; B, A2; C, mandible; D, maxillule; E, maxilla; F, maxilliped. Scales: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.5252391" httpUri="https://zenodo.org/record/5252391/files/figure.png" pageId="11" pageNumber="208">Fig. 6E</figureCitation>
) 4-segmented, comprising praecoxa, coxa, basis and 1-segmented endopod. Praecoxal endite with 2 setae, one of which pinnate. Coxa with 1 naked seta representing proximal endite; distal endite movable, forming 1 bisetose lobe completely fused with naked seta proximally. Basis forming 2 strong claw-like spines with 1 naked seta between them basally. Endopod carrying 5 elements of 1 naked, 2 spiniform and 2 spinulose setae.
</paragraph>
<paragraph id="A628368AB753FFC7C5D0A7F0FDDEFDBC" blockId="11.[151,1437,151,728]" pageId="11" pageNumber="208">
Maxilliped (
<figureCitation id="3EAC2A0FB753FFC7C443A7F0FE58FE2C" box="[340,425,439,463]" captionStart="FIGURE 6" captionStartId="10.[151,250,1968,1991]" captionTargetBox="[171,1423,187,1943]" captionTargetId="figure-43@10.[151,1436,171,1948]" captionTargetPageId="10" captionText="FIGURE 6. Halicyclops pumilus sp. nov., female. A, A1; B, A2; C, mandible; D, maxillule; E, maxilla; F, maxilliped. Scales: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.5252391" httpUri="https://zenodo.org/record/5252391/files/figure.png" pageId="11" pageNumber="208">Fig. 6F</figureCitation>
) 2-segmented, comprising protopod and completely defined endopod; protopod about 2 times longer than endopod. Protopod with 2 naked proximal setae and 1 distal seta ornamented with 2 setules basally each on protuberance, representing endites; 3–4 setules present on outer distal corner of protopod. Endopod bearing 5 setae in total, comprising 2 inner setae, 1 apical, 2 subapical setae, of which inner and apical setae ornamented with 1 or 2 long setules.
</paragraph>
<paragraph id="A628368AB753FFC7C5D0A42BFEE0FD48" blockId="11.[151,1437,151,728]" pageId="11" pageNumber="208">P1–P4 (Fig. 7A–D) biramous, both rami 3-segmented. Spine formula of 3,4,4,3. Seta/spine armature of P1–P4 as follows:</paragraph>
<paragraph id="A628368AB753FFC7C469A4F9FB3BFD3B" blockId="11.[151,1437,151,728]" box="[382,1226,702,728]" pageId="11" pageNumber="208">P1 coxa 0-1 basis 1-1 exp I-1; I-1; III,2,3 enp 0-1; 0-1; I,I+1,3</paragraph>
<paragraph id="A628368AB753FFC7C468A4B2FB42FCEC" blockId="11.[383,1203,757,783]" box="[383,1203,757,783]" pageId="11" pageNumber="208">P2 coxa 0-1 basis 1-0 exp I-1; I-1; IV,1,4 enp 0-1; 0-1; I,II,3</paragraph>
<paragraph id="A628368AB753FFC7C468A56BFB42FCA5" blockId="11.[383,1203,812,838]" box="[383,1203,812,838]" pageId="11" pageNumber="208">P3 coxa 0-1 basis 1-0 exp I-1; I-1; IV,1,4 enp 0-1; 0-1; I,II,3</paragraph>
<paragraph id="A628368AB753FFC7C46AA524FB45FC9E" blockId="11.[381,1204,867,893]" box="[381,1204,867,893]" pageId="11" pageNumber="208">P4 coxa 0-1 basis 1-0 exp I-1; I-1; III,1,4 enp 0-1; 0-1; I,II,2</paragraph>
<paragraph id="A628368AB753FFC7C5D0A5E6FC92FAC2" blockId="11.[151,1437,929,2034]" pageId="11" pageNumber="208">
P1 (Fig. 7A), intercoxal sclerite with hairs along distal margin of both lateral lobes; medial seta on P1 basis short but stout, not reaching to poseterior end of enp1; outer spines on exp3 become a bit (10–30%) longer distally. P2–P4, intercoxal sclerite bare along distal margin of both lateral lobes; enp2 with single inner seta. P4 (Fig. 7D), enp3 1.50 times longer than wide; both inner setae spiniform, short, nearly as long as enp3, and 23% shorter than inner apical spine; inner apical spine 1.23 times longer than enp3, and about 2 times longer than outer apical spine. P5 (
<figureCitation id="3EAC2A0FB753FFC7C5D1A212FEEFFB8E" box="[198,286,1109,1133]" captionStart="FIGURE 5" captionStartId="9.[151,250,1970,1993]" captionTargetBox="[217,1373,192,1948]" captionTargetId="figure-43@9.[191,1396,181,1953]" captionTargetPageId="9" captionText="FIGURE 5. Halicyclops pumilus sp. nov., female. A, habitus, dorsal; B, urosome, dorsal; C, P5 exopod. Scales: A, B = 100 µm; C = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.5252387" httpUri="https://zenodo.org/record/5252387/files/figure.png" pageId="11" pageNumber="208">Fig. 5C</figureCitation>
) basis and endopod completely incorporated to fifth pedigerous somite; basal seta inserted on small protuberance arising from dorsolateral corner of somite. Exopod nearly rectangular, 1.18 times longer than wide, with straight inner margin and angular inner distal corner; outer margin round, ornamented with spinule row; spines stumpy, inner apical spine longest, only 52% of exopod in length; apical seta 1.1 times longer than exopod, and 2.0 times longer than inner apical spine. P6 indistinct, represented by small genital operculum armed with 3 elements of 2 minute spinous projections and 1 long naked seta.
</paragraph>
<paragraph id="A628368AB753FFC7C580A36AFEA5FAA6" blockId="11.[151,1437,929,2034]" box="[151,340,1325,1349]" pageId="11" pageNumber="208">Male. Unknown.</paragraph>
</subSubSection>
<subSubSection id="EE8D6501B753FFC7C5D0A316FEB3FA36" pageId="11" pageNumber="208" type="biology_ecology">
<paragraph id="A628368AB753FFC7C5D0A316FEB3FA36" blockId="11.[151,1437,929,2034]" pageId="11" pageNumber="208">
<emphasis id="94E3EA98B753FFC7C5D0A316FEDAFA8A" bold="true" box="[199,299,1361,1385]" inLineHeading="true" pageId="11" pageNumber="208" reason="1">Ecology.</emphasis>
Specimens were collected by rinsing sandy sediments at two river mouths discharging into the southern coast of 
<collectingCountry id="DE80761AB753FFC7C448A331FE57FA6E" box="[351,422,1398,1421]" name="South Korea" pageId="11" pageNumber="208">Korea</collectingCountry>
, and at a small seaside ditch on the western coast of 
<collectingCountry id="DE80761AB753FFC7C6EBA331FBB2FA6E" box="[1020,1091,1398,1421]" name="South Korea" pageId="11" pageNumber="208">Korea</collectingCountry>
, where brackish shells such as Asian clams, 
<taxonomicName id="61974D09B753FFC7C421A3DEFD39FA52" authority="(Muller, 1774)" baseAuthorityName="Muller" baseAuthorityYear="1774" box="[310,712,1433,1457]" class="Choanoflagellatea" family="Acanthoecidae" genus="Corbicula" kingdom="Protozoa" order="Choanoflagellida" pageId="11" pageNumber="208" phylum="Choanozoa" rank="species" species="fluminea">
<emphasis id="94E3EA98B753FFC7C421A3DEFDE5FA52" box="[310,532,1433,1457]" italics="true" pageId="11" pageNumber="208">Corbicula fluminea</emphasis>
(Müller, 1774)
</taxonomicName>
, and purple olive clams, 
<taxonomicName id="61974D09B753FFC7C6E5A3DEFA66FA52" authority="(Deshayes, 1857)" baseAuthorityName="Deshayes" baseAuthorityYear="1857" box="[1010,1431,1433,1457]" class="Bivalvia" family="Psammobiidae" genus="Nuttallia" kingdom="Animalia" order="Cardiida" pageId="11" pageNumber="208" phylum="Mollusca" rank="species" species="japonica">
<emphasis id="94E3EA98B753FFC7C6E5A3DEFB33FA52" box="[1010,1218,1433,1457]" italics="true" pageId="11" pageNumber="208">Nuttallia japonica</emphasis>
(Deshayes, 1857)
</taxonomicName>
, were abundant.
</paragraph>
</subSubSection>
<subSubSection id="EE8D6501B753FFC0C5D0A3A6FBB5F833" lastPageId="12" lastPageNumber="209" pageId="11" pageNumber="208" type="discussion">
<paragraph id="A628368AB753FFC7C5D0A3A6FDA3F986" blockId="11.[151,1437,929,2034]" pageId="11" pageNumber="208">
<emphasis id="94E3EA98B753FFC7C5D0A3A6FECAFA1A" bold="true" box="[199,315,1505,1529]" inLineHeading="true" pageId="11" pageNumber="208" reason="1">Remarks.</emphasis>
This species belongs to the species group with the character combination of 3,4,4,3 spine formula, absence of lateral processes on genital somite, and well developed hyaline fringe of preanal somite. However, this species is characterized and easily distinguishable from its congeners by single inner seta on P2–P4 enp2 and very short spines on oval-shaped P5 exopod.
</paragraph>
<paragraph id="A628368AB753FFC7C5D0A036FF00F8FA" blockId="11.[151,1437,929,2034]" pageId="11" pageNumber="208">
The new species is strikingly similar to two interstitial species, 
<taxonomicName id="61974D09B753FFC7C690A035FB89F96A" authority="PleSa, 1961" authorityName="PleSa" authorityYear="1961" box="[903,1144,1649,1673]" class="Hexanauplia" family="Halicyclopidae" genus="Halicyclops" kingdom="Animalia" order="Cyclopoida" pageId="11" pageNumber="208" phylum="Arthropoda" rank="species" species="gauldi">
<emphasis id="94E3EA98B753FFC7C690A035FC01F96A" box="[903,1008,1649,1673]" italics="true" pageId="11" pageNumber="208">H. gauldi</emphasis>
<bibRefCitation id="C2064B7BB753FFC7C6E0A036FB89F96A" author="Plesa, C." box="[1015,1144,1649,1673]" pageId="11" pageNumber="208" pagination="1 - 13" refId="ref6939" refString="Plesa, C. (1961) New cyclopoids (Crustacea, Copepoda) of the interstitial fauna from the beaches of Ghana. Journal of the West African Science Association, 7 (1), 1 - 13." type="journal article" year="1961">PleŞa, 1961</bibRefCitation>
</taxonomicName>
from the Atlantic coast of 
<collectingCountry id="DE80761AB753FFC7C580A0D2FF12F94E" box="[151,227,1685,1709]" name="Ghana" pageId="11" pageNumber="208">Ghana</collectingCountry>
, West Africa and 
<taxonomicName id="61974D09B753FFC7C4A6A0D1FCA4F94E" authority="Lotufo &amp; Rocha, 1993" authorityName="Lotufo &amp; Rocha" authorityYear="1993" box="[433,853,1685,1709]" class="Hexanauplia" family="Halicyclopidae" genus="Halicyclops" kingdom="Animalia" order="Cyclopoida" pageId="11" pageNumber="208" phylum="Arthropoda" rank="species" species="ytororoma">
<emphasis id="94E3EA98B753FFC7C4A6A0D1FDBBF94E" box="[433,586,1686,1709]" italics="true" pageId="11" pageNumber="208">H. ytororoma</emphasis>
<bibRefCitation id="C2064B7BB753FFC7C745A0D2FCA4F94E" author="Lotufo, G. R. &amp; Rocha, C. E. F." box="[594,853,1685,1709]" pageId="11" pageNumber="208" pagination="175 - 184" refId="ref6847" refString="Lotufo, G. R. &amp; Rocha, C. E. F. (1993) Intertidal interstitial Halicyclops from the Brazilian coast (Copepoda: Cyclopoida). Hydrobiologia, 264, 175 - 184." type="journal article" year="1993">Lotufo &amp; Rocha, 1993</bibRefCitation>
</taxonomicName>
from the Atlantic coast of 
<collectingCountry id="DE80761AB753FFC7C19AA0D2FB20F94E" box="[1165,1233,1685,1709]" name="Brazil" pageId="11" pageNumber="208">Brazil</collectingCountry>
in sharing a very small body armed with single inner seta on P2–P4 enp2, elongate Fu (about 2 times longer than wide), multipronged pseudoperculum, short medial spine on basis of P1, and very stumpy spines on nearly quadrate P5 exopod.
</paragraph>
<paragraph id="A628368AB753FFC0C5D0A162FD5AFEDC" blockId="11.[151,1437,929,2034]" lastBlockId="12.[151,1436,151,320]" lastPageId="12" lastPageNumber="209" pageId="11" pageNumber="208">
Considering zoogeographical distribution and the high endemism rate of the interstitial copepods, the morphological similarities between the two marine species from the South Atlantic and the present new species from estuaries in the Northwest Pacific can be easily presumed to be the result of convergent evolution. However, in particular detail, they have a few morphological discrepancies: (1) 
<taxonomicName id="61974D09B753FFC7C689A1D5FBF6F84A" authorityName="PleSa" authorityYear="1961" box="[926,1031,1937,1961]" class="Hexanauplia" family="Halicyclopidae" genus="Halicyclops" kingdom="Animalia" order="Cyclopoida" pageId="11" pageNumber="208" phylum="Arthropoda" rank="species" species="gauldi">
<emphasis id="94E3EA98B753FFC7C689A1D5FBF6F84A" box="[926,1031,1937,1961]" italics="true" pageId="11" pageNumber="208">H. gauldi</emphasis>
</taxonomicName>
and 
<taxonomicName id="61974D09B753FFC7C12AA1D5FB22F84A" authorityName="Lotufo &amp; Rocha" authorityYear="1993" box="[1085,1235,1938,1961]" class="Hexanauplia" family="Halicyclopidae" genus="Halicyclops" kingdom="Animalia" order="Cyclopoida" pageId="11" pageNumber="208" phylum="Arthropoda" rank="species" species="ytororoma">
<emphasis id="94E3EA98B753FFC7C12AA1D5FB22F84A" box="[1085,1235,1938,1961]" italics="true" pageId="11" pageNumber="208">H. ytororoma</emphasis>
</taxonomicName>
show a spiniform inner proximalmost seta, deformed as a spatulate spine, each on P2–P3 enp3 (see 
<bibRefCitation id="C2064B7BB753FFC7C15FA1F2FAB7F82E" author="Lotufo, G. R. &amp; Rocha, C. E. F." box="[1096,1350,1973,1997]" pageId="11" pageNumber="208" pagination="175 - 184" refId="ref6847" refString="Lotufo, G. R. &amp; Rocha, C. E. F. (1993) Intertidal interstitial Halicyclops from the Brazilian coast (Copepoda: Cyclopoida). Hydrobiologia, 264, 175 - 184." type="journal article" year="1993">Lotufo &amp; Rocha 1993</bibRefCitation>
for the character state of 
<taxonomicName id="61974D09B753FFC7C44AA19DFE36F812" authorityName="PleSa" authorityYear="1961" box="[349,455,2009,2033]" class="Hexanauplia" family="Halicyclopidae" genus="Halicyclops" kingdom="Animalia" order="Cyclopoida" pageId="11" pageNumber="208" phylum="Arthropoda" rank="species" species="gauldi">
<emphasis id="94E3EA98B753FFC7C44AA19DFE36F812" box="[349,455,2009,2033]" italics="true" pageId="11" pageNumber="208">H. gauldi</emphasis>
</taxonomicName>
), while both of the setae are normally plumose in 
<taxonomicName id="61974D09B753FFC7C6EDA19DFB84F812" authorityName="Chang &amp; Lee" authorityYear="2012" box="[1018,1141,2009,2033]" class="Hexanauplia" family="Halicyclopidae" genus="Halicyclops" kingdom="Animalia" order="Cyclopoida" pageId="11" pageNumber="208" phylum="Arthropoda" rank="species" species="pumilus">
<emphasis id="94E3EA98B753FFC7C6EDA19DFB84F812" box="[1018,1141,2009,2033]" italics="true" pageId="11" pageNumber="208">H. pumilus</emphasis>
</taxonomicName>
; (2) fourth segment of A1 is much shorter in 
<taxonomicName id="61974D09B754FFC0C47DA6DEFE17FF53" authorityName="Chang &amp; Lee" authorityYear="2012" box="[362,486,152,176]" class="Hexanauplia" family="Halicyclopidae" genus="Halicyclops" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="209" phylum="Arthropoda" rank="species" species="pumilus">
<emphasis id="94E3EA98B754FFC0C47DA6DEFE17FF53" box="[362,486,152,176]" italics="true" pageId="12" pageNumber="209">H. pumilus</emphasis>
</taxonomicName>
(L/W is about 1.7–1.8, against more than 
<quantity id="616F9B6FB754FFC0C6D1A6D0FC00FF4C" box="[966,1009,151,175]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="12" pageNumber="209" unit="in" value="2.0">2 in</quantity>
<taxonomicName id="61974D09B754FFC0C6EEA6DEFB95FF53" authorityName="PleSa" authorityYear="1961" box="[1017,1124,152,176]" class="Hexanauplia" family="Halicyclopidae" genus="Halicyclops" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="209" phylum="Arthropoda" rank="species" species="gauldi">
<emphasis id="94E3EA98B754FFC0C6EEA6DEFB95FF53" box="[1017,1124,152,176]" italics="true" pageId="12" pageNumber="209">H. gauldi</emphasis>
</taxonomicName>
and 
<taxonomicName id="61974D09B754FFC0C18CA6DEFAC3FF53" authorityName="Lotufo &amp; Rocha" authorityYear="1993" box="[1179,1330,153,176]" class="Hexanauplia" family="Halicyclopidae" genus="Halicyclops" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="209" phylum="Arthropoda" rank="species" species="ytororoma">
<emphasis id="94E3EA98B754FFC0C18CA6DEFAC3FF53" box="[1179,1330,153,176]" italics="true" pageId="12" pageNumber="209">H. ytororoma</emphasis>
</taxonomicName>
); and (3) 
<taxonomicName id="61974D09B754FFC0C580A6FAFEF0FF37" authorityName="PleSa" authorityYear="1961" box="[151,257,188,212]" class="Hexanauplia" family="Halicyclopidae" genus="Halicyclops" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="209" phylum="Arthropoda" rank="species" species="gauldi">
<emphasis id="94E3EA98B754FFC0C580A6FAFEF0FF37" box="[151,257,188,212]" italics="true" pageId="12" pageNumber="209">H. gauldi</emphasis>
</taxonomicName>
has only 3 setae on P1 enp3, while 
<taxonomicName id="61974D09B754FFC0C78FA6FAFCDEFF37" authorityName="Lotufo &amp; Rocha" authorityYear="1993" box="[664,815,189,212]" class="Hexanauplia" family="Halicyclopidae" genus="Halicyclops" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="209" phylum="Arthropoda" rank="species" species="ytororoma">
<emphasis id="94E3EA98B754FFC0C78FA6FAFCDEFF37" box="[664,815,189,212]" italics="true" pageId="12" pageNumber="209">H. ytororoma</emphasis>
</taxonomicName>
and 
<taxonomicName id="61974D09B754FFC0C671A6FAFC13FF37" authorityName="Chang &amp; Lee" authorityYear="2012" box="[870,994,188,212]" class="Hexanauplia" family="Halicyclopidae" genus="Halicyclops" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="209" phylum="Arthropoda" rank="species" species="pumilus">
<emphasis id="94E3EA98B754FFC0C671A6FAFC13FF37" box="[870,994,188,212]" italics="true" pageId="12" pageNumber="209">H. pumilus</emphasis>
</taxonomicName>
have 4 setae are normal in 
<taxonomicName id="61974D09B754FFC0C00EA6FAFA64FF37" authorityName="Chang &amp; Lee" authorityYear="2012" box="[1305,1429,188,212]" class="Hexanauplia" family="Halicyclopidae" genus="Halicyclops" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="209" phylum="Arthropoda" rank="species" species="pumilus">
<emphasis id="94E3EA98B754FFC0C00EA6FAFA64FF37" box="[1305,1429,188,212]" italics="true" pageId="12" pageNumber="209">H. pumilus</emphasis>
</taxonomicName>
. Moreover, it is noteworthy to mention that the middorsal integumental depression of cephalothorax is apparently shown in 
<taxonomicName id="61974D09B754FFC0C413A742FE8EFEFF" authorityName="Chang &amp; Lee" authorityYear="2012" box="[260,383,260,284]" class="Hexanauplia" family="Halicyclopidae" genus="Halicyclops" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="209" phylum="Arthropoda" rank="species" species="pumilus">
<emphasis id="94E3EA98B754FFC0C413A742FE8EFEFF" box="[260,383,260,284]" italics="true" pageId="12" pageNumber="209">H. pumilus</emphasis>
</taxonomicName>
, while any information on it was not given nor figured in the descriptions of 
<taxonomicName id="61974D09B754FFC0C1F7A742FABBFEFF" authorityName="PleSa" authorityYear="1961" box="[1248,1354,260,284]" class="Hexanauplia" family="Halicyclopidae" genus="Halicyclops" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="209" phylum="Arthropoda" rank="species" species="gauldi">
<emphasis id="94E3EA98B754FFC0C1F7A742FABBFEFF" box="[1248,1354,260,284]" italics="true" pageId="12" pageNumber="209">H. gauldi</emphasis>
</taxonomicName>
and 
<taxonomicName id="61974D09B754FFC0C096A742FEFAFEA3" authorityName="Lotufo &amp; Rocha" authorityYear="1993" class="Hexanauplia" family="Halicyclopidae" genus="Halicyclops" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="209" phylum="Arthropoda" rank="species" species="ytororoma">
<emphasis id="94E3EA98B754FFC0C096A742FEFAFEA3" italics="true" pageId="12" pageNumber="209">H. ytororoma</emphasis>
</taxonomicName>
(
<bibRefCitation id="C2064B7BB754FFC0C40DA760FE69FEDC" author="Plesa, C." box="[282,408,295,320]" pageId="12" pageNumber="209" pagination="1 - 13" refId="ref6939" refString="Plesa, C. (1961) New cyclopoids (Crustacea, Copepoda) of the interstitial fauna from the beaches of Ghana. Journal of the West African Science Association, 7 (1), 1 - 13." type="journal article" year="1961">PleŞa 1961</bibRefCitation>
; 
<bibRefCitation id="C2064B7BB754FFC0C4B3A760FD6EFEDC" author="Lotufo, G. R. &amp; Rocha, C. E. F." box="[420,671,295,319]" pageId="12" pageNumber="209" pagination="175 - 184" refId="ref6847" refString="Lotufo, G. R. &amp; Rocha, C. E. F. (1993) Intertidal interstitial Halicyclops from the Brazilian coast (Copepoda: Cyclopoida). Hydrobiologia, 264, 175 - 184." type="journal article" year="1993">Lotufo &amp; Rocha 1993</bibRefCitation>
).
</paragraph>
<paragraph id="A628368AB754FFC0C580A1FDFBB5F833" blockId="12.[151,1092,1978,2001]" box="[151,1092,1978,2001]" pageId="12" pageNumber="209">
<emphasis id="94E3EA98B754FFC0C580A1FDFEE2F833" bold="true" box="[151,275,1978,2001]" pageId="12" pageNumber="209">FIGURE 7.</emphasis>
<taxonomicName id="61974D09B754FFC0C40EA1FDFE19F833" authority="Chang &amp; Lee, 2012" authorityName="Chang &amp; Lee" authorityYear="2012" box="[281,488,1978,2000]" class="Hexanauplia" family="Halicyclopidae" genus="Halicyclops" kingdom="Animalia" order="Cyclopoida" pageId="12" pageNumber="209" phylum="Arthropoda" rank="species" species="pumilus" status="sp. nov.">
<emphasis id="94E3EA98B754FFC0C40EA1FDFE19F833" box="[281,488,1978,2000]" italics="true" pageId="12" pageNumber="209">Halicyclops pumilus</emphasis>
</taxonomicName>
<emphasis id="94E3EA98B754FFC0C4F9A1FCFDCEF833" bold="true" box="[494,575,1979,2000]" pageId="12" pageNumber="209">
<taxonomicNameLabel id="8FD057E3B754FFC0C4F9A1FCFDCEF833" box="[494,575,1979,2000]" pageId="12" pageNumber="209" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female. A, P1; B, P2; C, P3; D, P4. Scale: 50 µm.
</paragraph>
</subSubSection>
</treatment>
</document>