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<document id="C16BEC99931BF569EB8ADEC006C3045B" ID-CLB-Dataset="42305" ID-DOI="10.11646/zootaxa.3878.5.7" ID-GBIF-Dataset="cb3f8689-9f3e-451a-93d0-d0e88d4e50ff" ID-ISSN="1175-5326" ID-Zenodo-Dep="227753" ID-ZooBank="C69D72C1-2E03-46E4-9C3E-3E36AE82563E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460952962255" checkinUser="plazi" docAuthor="Sharifian, Sana, Kamrani, Ehsan &amp; Sharifian, Salim" docDate="2014" docId="017DF3129A36FFF0BDAA44EE3B381423" docLanguage="en" docName="zt03878p490.pdf" docOrigin="Zootaxa 3878 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sodhiana iranica Sharifian, Kamrani &amp; Sharifian, 2014, n. sp." docType="treatment" docVersion="9" lastPageNumber="489" masterDocId="FD448B6A9A37FFF4BD3D44793F3F1736" masterDocTitle="A new species of freshwater crab of the genus Sodhiana Yeo &amp; Ng, 2012 (Decapoda: Brachyura: Gecarcinucidae) from southern Iran" masterLastPageNumber="490" masterPageNumber="485" pageNumber="486" updateTime="1698585728926" updateUser="ExternalLinkService">
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<mods:title id="DD3C91E5EC7BAC55AB941397A3DA986D">A new species of freshwater crab of the genus Sodhiana Yeo &amp; Ng, 2012 (Decapoda: Brachyura: Gecarcinucidae) from southern Iran</mods:title>
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<mods:namePart id="E17EDE3BB08ABB5DF3697FF7AEC09209">Sharifian, Sana</mods:namePart>
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<mods:namePart id="6CA9750AE5B6B4DEF5268D862BEC7F2B">Kamrani, Ehsan</mods:namePart>
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<mods:namePart id="9114ECF0D55DCAB3FA4DC820C4FBC32A">Sharifian, Salim</mods:namePart>
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<mods:date id="06406593E8A78F8E3214B161E06D0553">2014</mods:date>
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<treatment id="017DF3129A36FFF0BDAA44EE3B381423" ID-DOI="http://doi.org/10.5281/zenodo.6138478" ID-GBIF-Taxon="119591917" ID-Zenodo-Dep="6138478" LSID="urn:lsid:plazi:treatment:017DF3129A36FFF0BDAA44EE3B381423" httpUri="http://treatment.plazi.org/id/017DF3129A36FFF0BDAA44EE3B381423" lastPageId="4" lastPageNumber="489" pageId="1" pageNumber="486">
<subSubSection id="C1CE118F9A36FFF5BDAA44EE3E2E17E5" pageId="1" pageNumber="486" type="nomenclature">
<paragraph id="896B42049A36FFF5BDAA44EE3E8D1784" blockId="1.[151,434,151,211]" box="[151,434,151,178]" pageId="1" pageNumber="486">
<heading id="D223F5689A36FFF5BDAA44EE3E8D1784" bold="true" box="[151,434,151,178]" fontSize="11" level="1" pageId="1" pageNumber="486" reason="1">
<emphasis id="BBA09E169A36FFF5BDAA44EE3E8D1784" bold="true" box="[151,434,151,178]" pageId="1" pageNumber="486">
<taxonomicName id="4ED439879A36FFF5BDAA44EE3E571787" ID-CoL="4XYTS" box="[151,360,151,177]" class="Malacostraca" family="Gecarcinucidae" genus="Sodhiana" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="486" phylum="Arthropoda" rank="species" species="iranica" status="sp. nov.">
<emphasis id="BBA09E169A36FFF5BDAA44EE3E571787" bold="true" box="[151,360,151,177]" italics="true" pageId="1" pageNumber="486">Sodhiana iranica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A093236D9A36FFF5BC5244E13E8D1784" box="[367,434,152,178]" pageId="1" pageNumber="486" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="896B42049A36FFF5BDAA44C33E2E17E5" blockId="1.[151,434,151,211]" box="[151,273,186,211]" pageId="1" pageNumber="486">
(
<figureCitation id="11EF5E819A36FFF5BDA244C33E3617E5" box="[159,265,186,211]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="1.[151,250,1945,1967]" captionStartId-1="2.[151,250,1373,1395]" captionStartId-2="3.[151,250,1376,1398]" captionTargetBox-0="[345,1239,605,1913]" captionTargetBox-1="[340,1246,193,1351]" captionTargetBox-2="[167,1418,201,1333]" captionTargetId-0="figure@1.[345,1240,603,1914]" captionTargetId-1="figure@2.[340,1247,193,1351]" captionTargetId-2="figure@3.[151,1436,193,1355]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="FIGURE 1. Sodhiana iranica n. sp., holotype male (30.2 × 23.1 mm) (ZRC 2014.0240), Bastak, Iran. A, overall dorsal view; B, frontal view of carapace; C, ventral view showing anterior thoracic sternum and abdomen." captionText-1="FIGURE 2. Sodhiana iranica n. sp., A, C, D, F, G, holotype male (30.2 × 23.1 mm) (ZRC 2014.0240), Bastak, Iran; B, H, paratype male (29.3 × 23.0 mm) (ZRC 2014.0241), Bastak, Iran; E, I, paratype female (29.0 × 23.1 mm) (ZRC 2014.0241), Bastak, Iran. A, B, dorsal views of carapaces; C, left third maxilliped; D, male abdomen (somites 2 6 and telson); E, female thoracic sternum showing vulvae; F, H, I, major chelae; G, minor chela." captionText-2="FIGURE 3. Sodhiana iranica n. sp., holotype male (30.2 × 23.1 mm) (ZRC 2014.0240), Bastak, Iran. A, ventral view of left G 1; B, ventral view of distal part of left G 1; C, dorsal view of distal part of left G 1; D, left G 2. Scales = 1.0 mm." httpUri-0="https://zenodo.org/record/227754/files/figure.png" httpUri-1="https://zenodo.org/record/227755/files/figure.png" httpUri-2="https://zenodo.org/record/227756/files/figure.png" pageId="1" pageNumber="486">Figs. 13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C1CE118F9A36FFF5BDAA44843AB91500" pageId="1" pageNumber="486" type="materials_examined">
<paragraph id="896B42049A36FFF5BDAA44843C6616B7" blockId="1.[151,1436,253,566]" pageId="1" pageNumber="486">
<emphasis id="BBA09E169A36FFF5BDAA44843EBE1620" bold="true" box="[151,385,253,278]" pageId="1" pageNumber="486">Material examined.</emphasis>
<typeStatus id="566FFCA69A36FFF5BCB544843ECA1620" box="[392,501,253,278]" pageId="1" pageNumber="486" type="holotype">Holotype</typeStatus>
: male (30.2 ×
<quantity id="4E2CEFE19A36FFF5BFAA44873DC31620" box="[663,764,254,278]" metricMagnitude="-2" metricUnit="m" metricValue="2.31" pageId="1" pageNumber="486" unit="mm" value="23.1">23.1 mm</quantity>
) (
<collectionCode id="EFC5DAC19A36FFF5BE2E44873C741620" box="[787,843,254,278]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="486">ZRC</collectionCode>
2014.0240), Bastak,
<collectingCountry id="F1C302949A36FFF5B90044873B521623" box="[1085,1133,254,277]" name="Iran" pageId="1" pageNumber="486">Iran</collectingCountry>
,
<geoCoordinate id="ECE024C39A36FFF5B94544873BE61620" box="[1144,1241,254,278]" direction="north" orientation="latitude" pageId="1" pageNumber="486" precision="925" value="27.216667">27°13N</geoCoordinate>
<geoCoordinate id="ECE024C39A36FFF5B9DC44843A7F1620" box="[1249,1344,253,278]" direction="east" orientation="longitude" pageId="1" pageNumber="486" precision="925" value="54.666668">54°40E</geoCoordinate>
, coll. S. Sharifian,
<date id="FD6A64C49A36FFF5BC2C45583EF4160F" box="[273,459,289,314]" pageId="1" pageNumber="486" value="2014-03-12">12 March 2014</date>
.
<typeStatus id="566FFCA69A36FFF5BCE6455B3D72160C" box="[475,589,290,314]" pageId="1" pageNumber="486" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9FD2898D9A36FFF5BF6045583D92160C" box="[605,685,289,314]" pageId="1" pageNumber="486" type="male">1 male</specimenCount>
(29.3 × 23.0 mm),
<specimenCount id="9FD2898D9A36FFF5BE9945583B29160C" box="[932,1046,289,314]" pageId="1" pageNumber="486" type="female">3 females</specimenCount>
(largest 29.0 ×
<quantity id="4E2CEFE19A36FFF5B9E045583A77160F" box="[1245,1352,289,313]" metricMagnitude="-2" metricUnit="m" metricValue="2.31" pageId="1" pageNumber="486" unit="mm" value="23.1">23.1 mm</quantity>
) (
<collectionCode id="EFC5DAC19A36FFF5B85945583AA3160F" box="[1380,1436,289,313]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="486">ZRC</collectionCode>
2014.0241), same data as
<typeStatus id="566FFCA69A36FFF5BC85453C3D211668" box="[440,542,325,350]" pageId="1" pageNumber="486" type="holotype">holotype</typeStatus>
;
<specimenCount id="9FD2898D9A36FFF5BF14453C3D401668" box="[553,639,325,350]" pageId="1" pageNumber="486" type="male">5 males</specimenCount>
(largest 30.3 ×
<quantity id="4E2CEFE19A36FFF5BE13453C3CA91668" box="[814,918,325,350]" metricMagnitude="-2" metricUnit="m" metricValue="2.61" pageId="1" pageNumber="486" unit="mm" value="26.1">26.1 mm</quantity>
),
<specimenCount id="9FD2898D9A36FFF5BE95453F3B2B1668" box="[936,1044,325,350]" pageId="1" pageNumber="486" type="female">4 females</specimenCount>
(largest 33.8 ×
<quantity id="4E2CEFE19A36FFF5B9FE453F3A171668" box="[1219,1320,326,350]" metricMagnitude="-2" metricUnit="m" metricValue="2.94" pageId="1" pageNumber="486" unit="mm" value="29.4">29.4 mm</quantity>
) (
<collectionCode id="EFC5DAC19A36FFF5B803453F3AAF1668" box="[1342,1424,326,350]" pageId="1" pageNumber="486">ZCUH</collectionCode>
), Bastak,
<collectingCountry id="F1C302949A36FFF5BDCC45133E1E16B7" box="[241,289,362,385]" name="Iran" pageId="1" pageNumber="486">Iran</collectingCountry>
,
<geoCoordinate id="ECE024C39A36FFF5BC1145103EB316B7" box="[300,396,361,385]" direction="north" orientation="latitude" pageId="1" pageNumber="486" precision="925" value="27.216667">27°13N</geoCoordinate>
<geoCoordinate id="ECE024C39A36FFF5BCA945103ECB16B4" box="[404,500,361,386]" direction="east" orientation="longitude" pageId="1" pageNumber="486" precision="925" value="54.666668">54°40E</geoCoordinate>
, coll. S. Sharifian,
<date id="FD6A64C49A36FFF5BFF745103C6A16B7" box="[714,853,361,386]" pageId="1" pageNumber="486" value="2014-05-01">1 May 2014</date>
.
</paragraph>
<paragraph id="896B42049A36FFF5BDFA45F43AB91500" blockId="1.[151,1436,253,566]" pageId="1" pageNumber="486">
<emphasis id="BBA09E169A36FFF5BDFA45F43C3A1690" bold="true" box="[199,773,397,422]" pageId="1" pageNumber="486">Comparative material [photographs examined].</emphasis>
<taxonomicName id="4ED439879A36FFF5BE3145F43BA31690" authority="Alcock, 1909" authorityName="Alcock" authorityYear="1909" box="[780,1180,397,422]" class="Malacostraca" family="Gecarcinucidae" genus="Sodhiana" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="486" phylum="Arthropoda" rank="species" species="blanfordi">
<emphasis id="BBA09E169A36FFF5BE3145F43CDA1693" box="[780,997,397,421]" italics="true" pageId="1" pageNumber="486">Sodhiana blanfordi</emphasis>
(
<bibRefCitation id="ED453FF59A36FFF5BEC845F43BAC1690" author="Alcock" box="[1013,1171,397,422]" pageId="1" pageNumber="486" refString="Alcock, A. (1909) Diagnoses of new species and varieties of freshwater crabs. Records of the Indian Museum, 4 (3), 375 - 381." type="journal article" year="1909">Alcock, 1909</bibRefCitation>
)
</taxonomicName>
:
<typeStatus id="566FFCA69A36FFF5B99545F73A3B1690" box="[1192,1284,398,422]" pageId="1" pageNumber="486" type="syntype">Syntype</typeStatus>
male (46.4 × 35.0 mm) (NHM 1905.10.13.5), Pishin, Baluchistan,
<collectingCountry id="F1C302949A36FFF5BE3A45C83C5616FC" box="[775,873,433,458]" name="Pakistan" pageId="1" pageNumber="486">Pakistan</collectingCountry>
;
<typeStatus id="566FFCA69A36FFF5BE4445CB3CE116FC" box="[889,990,434,458]" pageId="1" pageNumber="486" type="syntype">syntypes</typeStatus>
,
<specimenCount id="9FD2898D9A36FFF5BED345C83B7716FC" box="[1006,1096,433,458]" pageId="1" pageNumber="486" type="male">4 males</specimenCount>
(largest 43.2 ×
<quantity id="4E2CEFE19A36FFF5B83745C83A4816FF" box="[1290,1399,433,457]" metricMagnitude="-2" metricUnit="m" metricValue="3.32" pageId="1" pageNumber="486" unit="mm" value="33.2">33.2 mm</quantity>
),
<specimenCount id="9FD2898D9A36FFF5B8B345C83FD116D8" pageId="1" pageNumber="486" type="female">5 females</specimenCount>
(largest 36.9 ×
<quantity id="4E2CEFE19A36FFF5BC9D45AF3D3A16D8" box="[416,517,470,494]" metricMagnitude="-2" metricUnit="m" metricValue="2.88" pageId="1" pageNumber="486" unit="mm" value="28.8">28.8 mm</quantity>
) (ZSI 4043/4), Pishin, Baluchistan,
<collectingCountry id="F1C302949A36FFF5BEA645AC3CC216D8" box="[923,1021,469,494]" name="Pakistan" pageId="1" pageNumber="486">Pakistan</collectingCountry>
, coll. W.T. Blanford, no date;
<specimenCount id="9FD2898D9A36FFF5B86D45AF3AA316D8" box="[1360,1436,469,494]" pageId="1" pageNumber="486" type="male">1 male</specimenCount>
(41.8 ×
<quantity id="4E2CEFE19A36FFF5BDCD45803E671527" box="[240,344,505,529]" metricMagnitude="-2" metricUnit="m" metricValue="3.18" pageId="1" pageNumber="486" unit="mm" value="31.8">31.8 mm</quantity>
),
<specimenCount id="9FD2898D9A36FFF5BC5145803EF21524" box="[364,461,505,530]" pageId="1" pageNumber="486" type="female">1 female</specimenCount>
(37.3 ×
<quantity id="4E2CEFE19A36FFF5BF1045803DAC1527" box="[557,659,505,529]" metricMagnitude="-2" metricUnit="m" metricValue="2.88" pageId="1" pageNumber="486" unit="mm" value="28.8">28.8 mm</quantity>
) (
<collectionCode id="EFC5DAC19A36FFF5BF9745803DD91527" box="[682,742,505,529]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="486">ZRC</collectionCode>
), Punjgoor, Beluchistan,
<collectingCountry id="F1C302949A36FFF5B93D45803B5D1524" box="[1024,1122,505,530]" name="Pakistan" pageId="1" pageNumber="486">Pakistan</collectingCountry>
, coll. Q. Kazmi,
<date id="FD6A64C49A36FFF5B81C45803FEC1500" pageId="1" pageNumber="486" value="2003-01-19">19 January 2003</date>
;
<specimenCount id="9FD2898D9A36FFF5BDE046673E0B1500" box="[221,308,541,566]" pageId="1" pageNumber="486" type="male">2 males</specimenCount>
(larger 37.1 × 28.0 mm) (
<collectionCode id="EFC5DAC19A36FFF5BF6A46673DAC1500" box="[599,659,542,566]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="486">ZRC</collectionCode>
), Punjgoor, Beluchistan,
<collectingCountry id="F1C302949A36FFF5BE9146643B311500" box="[940,1038,541,566]" name="Pakistan" pageId="1" pageNumber="486">Pakistan</collectingCountry>
, coll. Q. Kazmi,
<date id="FD6A64C49A36FFF5B9F646673ABD1500" box="[1227,1410,542,566]" pageId="1" pageNumber="486" value="2003-08-19">19 August 2003</date>
.
</paragraph>
</subSubSection>
<caption id="DDAB128C9A36FFF5BDAA43E03B7710F8" httpUri="https://zenodo.org/record/227754/files/figure.png" pageId="1" pageNumber="486" targetBox="[345,1239,605,1913]" targetPageId="1">
<paragraph id="896B42049A36FFF5BDAA43E03B7710F8" blockId="1.[151,1436,1945,1998]" pageId="1" pageNumber="486">
<emphasis id="BBA09E169A36FFF5BDAA43E03E2C1098" bold="true" box="[151,275,1945,1967]" pageId="1" pageNumber="486">FIGURE 1.</emphasis>
<taxonomicName id="4ED439879A36FFF5BC2743E03EF61099" box="[282,457,1945,1967]" class="Malacostraca" family="Gecarcinucidae" genus="Sodhiana" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="486" phylum="Arthropoda" rank="species" species="iranica" status="sp. nov.">
<emphasis id="BBA09E169A36FFF5BC2743E03EF61099" box="[282,457,1945,1967]" italics="true" pageId="1" pageNumber="486">Sodhiana iranica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A093236D9A36FFF5BCED43E03D351098" box="[464,522,1945,1966]" pageId="1" pageNumber="486" rank="species">
<emphasis id="BBA09E169A36FFF5BCED43E03D351098" bold="true" box="[464,522,1945,1966]" pageId="1" pageNumber="486">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype male (30.2 × 23.1 mm) (ZRC 2014.0240), Bastak, Iran. A, overall dorsal view; B, frontal view of carapace; C, ventral view showing anterior thoracic sternum and abdomen.
</paragraph>
</caption>
<caption id="DDAB128C9A35FFF6BDAA41243C5412F9" httpUri="https://zenodo.org/record/227755/files/figure.png" pageId="2" pageNumber="487" targetBox="[340,1246,193,1351]" targetPageId="2">
<paragraph id="896B42049A35FFF6BDAA41243C5412F9" blockId="2.[151,1436,1373,1487]" pageId="2" pageNumber="487">
<emphasis id="BBA09E169A35FFF6BDAA41243E2B1244" bold="true" box="[151,276,1373,1395]" pageId="2" pageNumber="487">FIGURE 2.</emphasis>
<taxonomicName id="4ED439879A35FFF6BC2041243EF21245" box="[285,461,1373,1395]" class="Malacostraca" family="Gecarcinucidae" genus="Sodhiana" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="487" phylum="Arthropoda" rank="species" species="iranica" status="sp. nov.">
<emphasis id="BBA09E169A35FFF6BC2041243EF21245" box="[285,461,1373,1395]" italics="true" pageId="2" pageNumber="487">Sodhiana iranica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A093236D9A35FFF6BCEB41243D2E1244" box="[470,529,1373,1394]" pageId="2" pageNumber="487" rank="species">
<emphasis id="BBA09E169A35FFF6BCEB41243D2E1244" bold="true" box="[470,529,1373,1394]" pageId="2" pageNumber="487">n. sp.</emphasis>
</taxonomicNameLabel>
, A, C, D, F, G, holotype male (30.2 × 23.1 mm) (ZRC 2014.0240), Bastak, Iran; B, H, paratype male (29.3 × 23.0 mm) (ZRC 2014.0241), Bastak, Iran; E, I, paratype female (29.0 × 23.1 mm) (ZRC 2014.0241), Bastak, Iran. A, B, dorsal views of carapaces; C, left third maxilliped; D, male abdomen (somites 26 and telson); E, female thoracic sternum showing vulvae; F, H, I, major chelae; G, minor chela.
</paragraph>
</caption>
<subSubSection id="C1CE118F9A35FFF7BDFA418F3AB612CF" lastPageId="3" lastPageNumber="488" pageId="2" pageNumber="487" type="description">
<paragraph id="896B42049A35FFF6BDFA418F3AAA1019" blockId="2.[151,1437,1526,2019]" pageId="2" pageNumber="487">
<emphasis id="BBA09E169A35FFF6BDFA418F3E661139" bold="true" box="[199,345,1526,1551]" pageId="2" pageNumber="487">Description.</emphasis>
Carapace broader than long; dorsal surfaces smooth; epigastric cristae low, formed by low transverse striae, not cristate, slightly anterior to postorbital cristae; postorbital cristae distinct, sharp, low, defined by small rounded granules, just reaching anterolateral margin at epibranchial tooth. External orbital tooth broadly triangular, outer margin lined with small rounded granules, about 3 times length of inner margin, margin near epibranchial tooth more angular, subdentiform. Epibranchial tooth distinct, triangular, just posterior of postorbital cristae, separated from margin by distinct V-shaped cleft (
<figureCitation id="11EF5E819A35FFF6BE6342D33C9011F5" box="[862,943,1706,1731]" captionStart="FIGURE 1" captionStartId="1.[151,250,1945,1967]" captionTargetBox="[345,1239,605,1913]" captionTargetId="figure@1.[345,1240,603,1914]" captionTargetPageId="1" captionText="FIGURE 1. Sodhiana iranica n. sp., holotype male (30.2 × 23.1 mm) (ZRC 2014.0240), Bastak, Iran. A, overall dorsal view; B, frontal view of carapace; C, ventral view showing anterior thoracic sternum and abdomen." httpUri="https://zenodo.org/record/227754/files/figure.png" pageId="2" pageNumber="487">Figs. 1</figureCitation>
A, B, 2A, B). Third maxilliped ischium subrectangular, smooth, with submedian depression but no obvious longitudinal median sulcus; merus rectangular; exopod slender, reaching to midlength of merus, flagellum elongate (
<figureCitation id="11EF5E819A35FFF6BE98428B3CD6103D" box="[933,1001,1778,1803]" captionStart="FIGURE 2" captionStartId="2.[151,250,1373,1395]" captionTargetBox="[340,1246,193,1351]" captionTargetId="figure@2.[340,1247,193,1351]" captionTargetPageId="2" captionText="FIGURE 2. Sodhiana iranica n. sp., A, C, D, F, G, holotype male (30.2 × 23.1 mm) (ZRC 2014.0240), Bastak, Iran; B, H, paratype male (29.3 × 23.0 mm) (ZRC 2014.0241), Bastak, Iran; E, I, paratype female (29.0 × 23.1 mm) (ZRC 2014.0241), Bastak, Iran. A, B, dorsal views of carapaces; C, left third maxilliped; D, male abdomen (somites 2 6 and telson); E, female thoracic sternum showing vulvae; F, H, I, major chelae; G, minor chela." httpUri="https://zenodo.org/record/227755/files/figure.png" pageId="2" pageNumber="487">Fig. 2</figureCitation>
C). Cutting edge of propodus of larger chela with large proximal tooth formed by fusion of several smaller teeth; tip of finger subspatulate (
<figureCitation id="11EF5E819A35FFF6B9C1436F3A011019" box="[1276,1342,1814,1839]" captionStart="FIGURE 2" captionStartId="2.[151,250,1373,1395]" captionTargetBox="[340,1246,193,1351]" captionTargetId="figure@2.[340,1247,193,1351]" captionTargetPageId="2" captionText="FIGURE 2. Sodhiana iranica n. sp., A, C, D, F, G, holotype male (30.2 × 23.1 mm) (ZRC 2014.0240), Bastak, Iran; B, H, paratype male (29.3 × 23.0 mm) (ZRC 2014.0241), Bastak, Iran; E, I, paratype female (29.0 × 23.1 mm) (ZRC 2014.0241), Bastak, Iran. A, B, dorsal views of carapaces; C, left third maxilliped; D, male abdomen (somites 2 6 and telson); E, female thoracic sternum showing vulvae; F, H, I, major chelae; G, minor chela." httpUri="https://zenodo.org/record/227755/files/figure.png" pageId="2" pageNumber="487">Fig. 2</figureCitation>
F, H, I).
</paragraph>
<paragraph id="896B42049A35FFF6BDFA43433CDE10D5" blockId="2.[151,1437,1526,2019]" pageId="2" pageNumber="487">
Male thoracic sternum smooth; male abdomen narrowly triangular, somite 6 trapezoidal with very slightly concave lateral margins, longer than broad; longer than telson (
<figureCitation id="11EF5E819A35FFF6BE5B43273C8D1041" box="[870,946,1886,1911]" captionStart="FIGURE 1" captionStartId="1.[151,250,1945,1967]" captionTargetBox="[345,1239,605,1913]" captionTargetId="figure@1.[345,1240,603,1914]" captionTargetPageId="1" captionText="FIGURE 1. Sodhiana iranica n. sp., holotype male (30.2 × 23.1 mm) (ZRC 2014.0240), Bastak, Iran. A, overall dorsal view; B, frontal view of carapace; C, ventral view showing anterior thoracic sternum and abdomen." httpUri="https://zenodo.org/record/227754/files/figure.png" pageId="2" pageNumber="487">Figs. 1</figureCitation>
C, 2D). G1 with terminal segment straight, subconical, tip directed upwards, slightly inwards (toward median part of thoracic sternum), about one-fifth length of subterminal segment. G2 distal segment long, about half length of basal segment (
<figureCitation id="11EF5E819A35FFF6B9B243DF3BE61089" box="[1167,1241,1958,1983]" captionStart="FIGURE 3" captionStartId="3.[151,250,1376,1398]" captionTargetBox="[167,1418,201,1333]" captionTargetId="figure@3.[151,1436,193,1355]" captionTargetPageId="3" captionText="FIGURE 3. Sodhiana iranica n. sp., holotype male (30.2 × 23.1 mm) (ZRC 2014.0240), Bastak, Iran. A, ventral view of left G 1; B, ventral view of distal part of left G 1; C, dorsal view of distal part of left G 1; D, left G 2. Scales = 1.0 mm." httpUri="https://zenodo.org/record/227756/files/figure.png" pageId="2" pageNumber="487">Fig. 3</figureCitation>
). Female vulvae rounded, on anterior half of sternite 6, with large opercular cover (
<figureCitation id="11EF5E819A35FFF6BE4343B33CFE10D5" box="[894,961,1994,2019]" captionStart="FIGURE 2" captionStartId="2.[151,250,1373,1395]" captionTargetBox="[340,1246,193,1351]" captionTargetId="figure@2.[340,1247,193,1351]" captionTargetPageId="2" captionText="FIGURE 2. Sodhiana iranica n. sp., A, C, D, F, G, holotype male (30.2 × 23.1 mm) (ZRC 2014.0240), Bastak, Iran; B, H, paratype male (29.3 × 23.0 mm) (ZRC 2014.0241), Bastak, Iran; E, I, paratype female (29.0 × 23.1 mm) (ZRC 2014.0241), Bastak, Iran. A, B, dorsal views of carapaces; C, left third maxilliped; D, male abdomen (somites 2 6 and telson); E, female thoracic sternum showing vulvae; F, H, I, major chelae; G, minor chela." httpUri="https://zenodo.org/record/227755/files/figure.png" pageId="2" pageNumber="487">Fig. 2</figureCitation>
E).
</paragraph>
<caption id="DDAB128C9A34FFF7BDAA41193A3212A3" httpUri="https://zenodo.org/record/227756/files/figure.png" pageId="3" pageNumber="488" targetBox="[167,1418,201,1333]" targetPageId="3">
<paragraph id="896B42049A34FFF7BDAA41193A3212A3" blockId="3.[151,1436,1376,1429]" pageId="3" pageNumber="488">
<emphasis id="BBA09E169A34FFF7BDAA41193E2C1243" bold="true" box="[151,275,1376,1398]" pageId="3" pageNumber="488">FIGURE 3.</emphasis>
<taxonomicName id="4ED439879A34FFF7BC2641193EF51240" box="[283,458,1376,1398]" class="Malacostraca" family="Gecarcinucidae" genus="Sodhiana" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="488" phylum="Arthropoda" rank="species" species="iranica" status="sp. nov.">
<emphasis id="BBA09E169A34FFF7BC2641193EF51240" box="[283,458,1376,1398]" italics="true" pageId="3" pageNumber="488">Sodhiana iranica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A093236D9A34FFF7BCEF41193D331243" box="[466,524,1376,1397]" pageId="3" pageNumber="488" rank="species">
<emphasis id="BBA09E169A34FFF7BCEF41193D331243" bold="true" box="[466,524,1376,1397]" pageId="3" pageNumber="488">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype male (30.2 × 23.1 mm) (ZRC 2014.0240), Bastak, Iran. A, ventral view of left G1; B, ventral view of distal part of left G1; C, dorsal view of distal part of left G1; D, left G2. Scales = 1.0 mm.
</paragraph>
</caption>
<paragraph id="896B42049A34FFF7BDFA41C53AB612CF" blockId="3.[151,1437,1468,2033]" pageId="3" pageNumber="488">
<emphasis id="BBA09E169A34FFF7BDFA41C53E1512E3" bold="true" box="[199,298,1468,1493]" pageId="3" pageNumber="488">Habitat.</emphasis>
The species inhabits a freshwater spring located in a semi-mountainous area in
<collectingCountry id="F1C302949A34FFF7B98841C43BDD12E2" box="[1205,1250,1469,1492]" name="Iran" pageId="3" pageNumber="488">Iran</collectingCountry>
(Eellod region), covered by dense stands of common reeds and salt cedar trees in the periphery, with mats of algae on the bottom.
</paragraph>
</subSubSection>
<subSubSection id="C1CE118F9A34FFF7BDFA427D3B7E112B" box="[199,1089,1540,1565]" pageId="3" pageNumber="488" type="distribution">
<paragraph id="896B42049A34FFF7BDFA427D3B7E112B" blockId="3.[151,1437,1468,2033]" box="[199,1089,1540,1565]" pageId="3" pageNumber="488">
<emphasis id="BBA09E169A34FFF7BDFA427D3E5F112B" bold="true" box="[199,352,1540,1565]" pageId="3" pageNumber="488">Distribution.</emphasis>
So far only known from Bastak,
<collectingCountry id="F1C302949A34FFF7BFE8427C3C3D112A" box="[725,770,1541,1564]" name="Iran" pageId="3" pageNumber="488">Iran</collectingCountry>
(but see “Remarks” below).
</paragraph>
</subSubSection>
<subSubSection id="C1CE118F9A34FFF7BDFA42513CCF1177" box="[199,1008,1576,1601]" pageId="3" pageNumber="488" type="etymology">
<paragraph id="896B42049A34FFF7BDFA42513CCF1177" blockId="3.[151,1437,1468,2033]" box="[199,1008,1576,1601]" pageId="3" pageNumber="488">
<emphasis id="BBA09E169A34FFF7BDFA42513E711177" bold="true" box="[199,334,1576,1601]" pageId="3" pageNumber="488">Etymology.</emphasis>
The specific epithet
<taxonomicName id="4ED439879A34FFF7BF0542503DB61176" box="[568,649,1577,1600]" class="Malacostraca" family="Gecarcinucidae" genus="Sodhiana" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="488" phylum="Arthropoda" rank="species" species="iranica">
<emphasis id="BBA09E169A34FFF7BF0542503DB61176" box="[568,649,1577,1600]" italics="true" pageId="3" pageNumber="488">iranica</emphasis>
</taxonomicName>
is named after the
<typeStatus id="566FFCA69A34FFF7BE6242503CB01177" box="[863,911,1577,1601]" pageId="3" pageNumber="488">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C1CE118F9A34FFF7BDFA42353AB811C3" pageId="3" pageNumber="488" type="discussion">
<paragraph id="896B42049A34FFF7BDFA42353AB811C3" blockId="3.[151,1437,1468,2033]" pageId="3" pageNumber="488">
<emphasis id="BBA09E169A34FFF7BDFA42353E041153" bold="true" box="[199,315,1612,1637]" pageId="3" pageNumber="488">Remarks.</emphasis>
<taxonomicName id="4ED439879A34FFF7BC7F42353DF21153" authority="Alcock, 1909" authorityName="Alcock" authorityYear="1909" box="[322,717,1612,1637]" class="Malacostraca" family="Gecarcinucidae" genus="Sodhiana" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="488" phylum="Arthropoda" rank="species" species="blanfordi">
<emphasis id="BBA09E169A34FFF7BC7F42353D241152" box="[322,539,1612,1636]" italics="true" pageId="3" pageNumber="488">Sodhiana blanfordi</emphasis>
(
<bibRefCitation id="ED453FF59A34FFF7BF1742353DFA1153" author="Alcock" box="[554,709,1612,1637]" pageId="3" pageNumber="488" refString="Alcock, A. (1909) Diagnoses of new species and varieties of freshwater crabs. Records of the Indian Museum, 4 (3), 375 - 381." type="journal article" year="1909">Alcock, 1909</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4ED439879A34FFF7BE3842343C4D1152" box="[773,882,1613,1636]" class="Malacostraca" family="Gecarcinucidae" genus="Sodhiana" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="488" phylum="Arthropoda" rank="species" species="iranica" status="sp. nov.">
<emphasis id="BBA09E169A34FFF7BE3842343C4D1152" box="[773,882,1613,1636]" italics="true" pageId="3" pageNumber="488">S. iranica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A093236D9A34FFF7BE4742353C851153" box="[890,954,1612,1637]" pageId="3" pageNumber="488" rank="species">
<emphasis id="BBA09E169A34FFF7BE4742353C851153" bold="true" box="[890,954,1612,1637]" pageId="3" pageNumber="488">n. sp.</emphasis>
</taxonomicNameLabel>
are superficially similar in the form of the carapace and ambulatory legs. The most significant difference is in the form of the G1. The terminal segment of the G1 of
<taxonomicName id="4ED439879A34FFF7BDDD42EC3E71119A" box="[224,334,1685,1708]" class="Malacostraca" family="Gecarcinucidae" genus="Sodhiana" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="488" phylum="Arthropoda" rank="species" species="iranica" status="sp. nov.">
<emphasis id="BBA09E169A34FFF7BDDD42EC3E71119A" box="[224,334,1685,1708]" italics="true" pageId="3" pageNumber="488">S. iranica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A093236D9A34FFF7BC6B42ED3EAA119B" box="[342,405,1684,1709]" pageId="3" pageNumber="488" rank="species">
<emphasis id="BBA09E169A34FFF7BC6B42ED3EAA119B" bold="true" box="[342,405,1684,1709]" pageId="3" pageNumber="488">n. sp.</emphasis>
</taxonomicNameLabel>
is gently bent inwards, with the outer margin more curved and the tip directed towards the median part of the sternoabdominal cavity (
<figureCitation id="11EF5E819A34FFF7BFBD42C13DFE11E6" box="[640,705,1720,1745]" captionStart="FIGURE 3" captionStartId="3.[151,250,1376,1398]" captionTargetBox="[167,1418,201,1333]" captionTargetId="figure@3.[151,1436,193,1355]" captionTargetPageId="3" captionText="FIGURE 3. Sodhiana iranica n. sp., holotype male (30.2 × 23.1 mm) (ZRC 2014.0240), Bastak, Iran. A, ventral view of left G 1; B, ventral view of distal part of left G 1; C, dorsal view of distal part of left G 1; D, left G 2. Scales = 1.0 mm." httpUri="https://zenodo.org/record/227756/files/figure.png" pageId="3" pageNumber="488">Fig. 3</figureCitation>
AC). In
<taxonomicName id="4ED439879A34FFF7BE1042C13C8F11E6" box="[813,944,1720,1744]" class="Malacostraca" family="Gecarcinucidae" genus="Sodhiana" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="488" phylum="Arthropoda" rank="species" species="blanfordi">
<emphasis id="BBA09E169A34FFF7BE1042C13C8F11E6" box="[813,944,1720,1744]" italics="true" pageId="3" pageNumber="488">S. blanfordi</emphasis>
</taxonomicName>
, the terminal segment is evenly conical and straight (
<figureCitation id="11EF5E819A34FFF7BDC742A53E0311C3" box="[250,316,1756,1781]" captionStart="FIGURE 4" captionStartId="4.[151,250,1712,1734]" captionTargetBox="[340,1246,844,1690]" captionTargetId="figure@4.[340,1247,844,1691]" captionTargetPageId="4" captionText="FIGURE 4. Sodhiana blanfordi. A, C, E G, syntype male (43.2 × 33.2 mm) (ZSI 4043 / 4), Pakistan; B, D, syntype male (46.4 × 35.0 mm) (NHM 1905.10.13.5), Pakistan. A, B, dorsal views of carapaces; C, D, frontal views of carapaces; E, ventral view showing anterior thoracic sternum and abdomen; F, dorsal view of right G 1; G, ventral view of right G 1." httpUri="https://zenodo.org/record/227757/files/figure.png" pageId="3" pageNumber="488">Fig. 4</figureCitation>
F, G). These differences are consistent for the specimens of different sizes of both species studied.
</paragraph>
</subSubSection>
<subSubSection id="C1CE118F9A34FFF0BDFA43793D2F1676" lastPageId="4" lastPageNumber="489" pageId="3" pageNumber="488" type="reference_group">
<paragraph id="896B42049A34FFF0BDFA43793D2F1676" blockId="3.[151,1437,1468,2033]" lastBlockId="4.[151,1437,151,789]" lastPageId="4" lastPageNumber="489" pageId="3" pageNumber="488">
<taxonomicName id="4ED439879A34FFF7BDFA43793EB5102E" ID-CoL="4XYTS" box="[199,394,1792,1816]" class="Malacostraca" family="Gecarcinucidae" genus="Sodhiana" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="488" phylum="Arthropoda" rank="species" species="iranica" status="sp. nov.">
<emphasis id="BBA09E169A34FFF7BDFA43793EB5102E" box="[199,394,1792,1816]" italics="true" pageId="3" pageNumber="488">Sodhiana iranica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A093236D9A34FFF7BCAF43793EED102F" box="[402,466,1792,1817]" pageId="3" pageNumber="488" rank="species">
<emphasis id="BBA09E169A34FFF7BCAF43793EED102F" bold="true" box="[402,466,1792,1817]" pageId="3" pageNumber="488">n. sp.</emphasis>
</taxonomicNameLabel>
is easily separated from
<taxonomicName id="4ED439879A34FFF7BFD243793CB8102E" ID-CoL="6XPDM" box="[751,903,1792,1816]" class="Malacostraca" family="Gecarcinucidae" genus="Sartoriana" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="488" phylum="Arthropoda" rank="species" species="rokitanskyi">
<emphasis id="BBA09E169A34FFF7BFD243793DC21021" box="[751,765,1792,1815]" italics="true" pageId="3" pageNumber="488">S</emphasis>
.
<emphasis id="BBA09E169A34FFF7BE3143793CB8102E" box="[780,903,1792,1816]" italics="true" pageId="3" pageNumber="488">rokitanskyi</emphasis>
</taxonomicName>
by having the epigastric cristae lower and less pronounced, and coming close or almost in contact with the postorbital cristae (
<figureCitation id="11EF5E819A34FFF7B937435D3B6C100B" box="[1034,1107,1828,1853]" captionStart="FIGURE 1" captionStartId="1.[151,250,1945,1967]" captionTargetBox="[345,1239,605,1913]" captionTargetId="figure@1.[345,1240,603,1914]" captionTargetPageId="1" captionText="FIGURE 1. Sodhiana iranica n. sp., holotype male (30.2 × 23.1 mm) (ZRC 2014.0240), Bastak, Iran. A, overall dorsal view; B, frontal view of carapace; C, ventral view showing anterior thoracic sternum and abdomen." httpUri="https://zenodo.org/record/227754/files/figure.png" pageId="3" pageNumber="488">Figs. 1</figureCitation>
A, 2A, B), whereas it is more prominent in
<taxonomicName id="4ED439879A34FFF7BC1343313EF91056" ID-CoL="6XPDM" box="[302,454,1864,1888]" class="Malacostraca" family="Gecarcinucidae" genus="Sartoriana" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="488" phylum="Arthropoda" rank="species" species="rokitanskyi">
<emphasis id="BBA09E169A34FFF7BC1343313E031069" box="[302,316,1864,1887]" italics="true" pageId="3" pageNumber="488">S</emphasis>
.
<emphasis id="BBA09E169A34FFF7BC7643313EF91056" box="[331,454,1864,1888]" italics="true" pageId="3" pageNumber="488">rokitanskyi</emphasis>
</taxonomicName>
(
<bibRefCitation id="ED453FF59A34FFF7BCE843313DAD1056" author="Pretzmann" box="[469,658,1864,1888]" pageId="3" pageNumber="488" refString="Pretzmann, G. (1971) Ergebnisse einiger Sammelreisen nach Vorderasien I. Teil: Zwei neue Unterarten von Susswasserkrabben. Annalen des Naturhistorischen Museums in Wien, 75, 473 - 475. [plates 1 - 6.]" type="journal article" year="1971">Pretzmann 1971</bibRefCitation>
: pl. 5 fig. 11;
<bibRefCitation id="ED453FF59A34FFF7BE1343313B2C1056" author="Kamrani" box="[814,1043,1864,1889]" pageId="3" pageNumber="488" refString="Kamrani, E., Ng, P. K. L., Mirzadeh, M. &amp; Nakhodai, S. (2009) The marsh crab, Sodhiana rokitanskyi (Pretzmann, 1971) (Decapoda, Brachyura, Gecarcinucidae) from southern Iran. Zootaxa, 2305, 24 - 32." type="journal article" year="2009">
Kamrani
<emphasis id="BBA09E169A34FFF7BEAA43303CEF1056" box="[919,976,1864,1888]" italics="true" pageId="3" pageNumber="488">et al.</emphasis>
2009
</bibRefCitation>
: fig. 1A;
<bibRefCitation id="ED453FF59A34FFF7B94043303A0D1056" author="Yeo" box="[1149,1330,1864,1889]" pageId="3" pageNumber="488" refString="Yeo, D. C. J. &amp; Ng, P. K. L. (2012) Sodhiana, A new genus of freshwater crab from south Asia (Decapoda: Brachyura: Gecarcinucidae). Raffles Bulletin of Zoology, 25, 279 - 284." type="journal article" year="2012">Yeo &amp; Ng 2012</bibRefCitation>
: fig. 2B). The G1 of
<taxonomicName id="4ED439879A34FFF7BC2443143EB710B2" ID-CoL="4XYTS" box="[281,392,1901,1924]" class="Malacostraca" family="Gecarcinucidae" genus="Sodhiana" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="488" phylum="Arthropoda" rank="species" species="iranica" status="sp. nov.">
<emphasis id="BBA09E169A34FFF7BC2443143EB710B2" box="[281,392,1901,1924]" italics="true" pageId="3" pageNumber="488">S. iranica</emphasis>
</taxonomicName>
<emphasis id="BBA09E169A34FFF7BCAF43153EEC10B3" bold="true" box="[402,467,1900,1925]" pageId="3" pageNumber="488">
<taxonomicNameLabel id="A093236D9A34FFF7BCAF43153EEC10B3" box="[402,467,1900,1925]" pageId="3" pageNumber="488" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is relatively more slender with the terminal segment proportionately shorter and tip gently bent inwards (
<figureCitation id="11EF5E819A34FFF7BCBF43E93EFD109E" box="[386,450,1936,1961]" captionStart="FIGURE 3" captionStartId="3.[151,250,1376,1398]" captionTargetBox="[167,1418,201,1333]" captionTargetId="figure@3.[151,1436,193,1355]" captionTargetPageId="3" captionText="FIGURE 3. Sodhiana iranica n. sp., holotype male (30.2 × 23.1 mm) (ZRC 2014.0240), Bastak, Iran. A, ventral view of left G 1; B, ventral view of distal part of left G 1; C, dorsal view of distal part of left G 1; D, left G 2. Scales = 1.0 mm." httpUri="https://zenodo.org/record/227756/files/figure.png" pageId="3" pageNumber="488">Fig. 3</figureCitation>
AC), in contrast to
<taxonomicName id="4ED439879A34FFF7BF9543E93C00109E" ID-CoL="6XPDM" box="[680,831,1936,1960]" class="Malacostraca" family="Gecarcinucidae" genus="Sartoriana" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="488" phylum="Arthropoda" rank="species" species="rokitanskyi">
<emphasis id="BBA09E169A34FFF7BF9543E93D891091" box="[680,694,1936,1959]" italics="true" pageId="3" pageNumber="488">S</emphasis>
.
<emphasis id="BBA09E169A34FFF7BFFE43E93C00109E" box="[707,831,1936,1960]" italics="true" pageId="3" pageNumber="488">rokitanskyi</emphasis>
</taxonomicName>
, where the G1 is proportionately stouter and the tip of the G1 terminal segment is gently bent outwards (
<bibRefCitation id="ED453FF59A34FFF7BFF643CC3CB610FB" author="Pretzmann" box="[715,905,1973,1997]" pageId="3" pageNumber="488" refString="Pretzmann, G. (1971) Ergebnisse einiger Sammelreisen nach Vorderasien I. Teil: Zwei neue Unterarten von Susswasserkrabben. Annalen des Naturhistorischen Museums in Wien, 75, 473 - 475. [plates 1 - 6.]" type="journal article" year="1971">Pretzmann 1971</bibRefCitation>
: pl. 6 fig. 16;
<bibRefCitation id="ED453FF59A34FFF7B91543CD3A3010FB" author="Kamrani" box="[1064,1295,1972,1997]" pageId="3" pageNumber="488" refString="Kamrani, E., Ng, P. K. L., Mirzadeh, M. &amp; Nakhodai, S. (2009) The marsh crab, Sodhiana rokitanskyi (Pretzmann, 1971) (Decapoda, Brachyura, Gecarcinucidae) from southern Iran. Zootaxa, 2305, 24 - 32." type="journal article" year="2009">
Kamrani
<emphasis id="BBA09E169A34FFF7B9AF43CC3BF510FA" box="[1170,1226,1972,1996]" italics="true" pageId="3" pageNumber="488">et al.</emphasis>
2009
</bibRefCitation>
: fig. 4A, B).
<taxonomicName id="4ED439879A34FFF7BDAA43A13E6310C6" ID-CoL="4XYTS" box="[151,348,2008,2032]" class="Malacostraca" family="Gecarcinucidae" genus="Sodhiana" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="488" phylum="Arthropoda" rank="species" species="iranica" status="sp. nov.">
<emphasis id="BBA09E169A34FFF7BDAA43A13E6310C6" box="[151,348,2008,2032]" italics="true" pageId="3" pageNumber="488">Sodhiana iranica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A093236D9A34FFF7BC5B43A13E9710C7" box="[358,424,2008,2033]" pageId="3" pageNumber="488" rank="species">
<emphasis id="BBA09E169A34FFF7BC5B43A13E9710C7" bold="true" box="[358,424,2008,2033]" pageId="3" pageNumber="488">n. sp.</emphasis>
</taxonomicNameLabel>
can be separated from the poorly known
<taxonomicName id="4ED439879A34FFF7BEAF43A13A2210C7" ID-CoL="6XPDL" authority="Pretzmann, 1963" authorityName="Pretzmann" authorityYear="1963" box="[914,1309,2008,2033]" class="Malacostraca" family="Gecarcinucidae" genus="Sartoriana" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="488" phylum="Arthropoda" rank="species" species="afghaniensis">
<emphasis id="BBA09E169A34FFF7BEAF43A13B0110C6" box="[914,1086,2008,2032]" italics="true" pageId="3" pageNumber="488">S. afghaniensis</emphasis>
(
<bibRefCitation id="ED453FF59A34FFF7B96C43A13A2A10C7" author="Pretzmann" box="[1105,1301,2008,2033]" pageId="3" pageNumber="488" refString="Pretzmann, G. (1963) Weiterer Bericht uber die mediterranen und vorderasiatischen Potamoniden. Annalen des Naturhistorischen Museums in Wien, 66, 373 - 380. [patels 1 - 3.]" type="journal article" year="1963">Pretzmann, 1963</bibRefCitation>
)
</taxonomicName>
by its less convex dorsal carapace surface and proportionately longer G1 terminal segment that has the tip directed inwards (versus straighter G1 with a relatively shorter terminal segment in
<taxonomicName id="4ED439879A33FFF0BEBE44C53B1217E2" ID-CoL="6XPDL" box="[899,1069,188,212]" class="Malacostraca" family="Gecarcinucidae" genus="Sartoriana" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="489" phylum="Arthropoda" rank="species" species="afghaniensis">
<emphasis id="BBA09E169A33FFF0BEBE44C53B1217E2" box="[899,1069,188,212]" italics="true" pageId="4" pageNumber="489">S. afghaniensis</emphasis>
</taxonomicName>
) (cf.
<bibRefCitation id="ED453FF59A33FFF0B95644C43A1917E3" author="Pretzmann" box="[1131,1318,188,213]" pageId="4" pageNumber="489" refString="Pretzmann, G. (1963) Weiterer Bericht uber die mediterranen und vorderasiatischen Potamoniden. Annalen des Naturhistorischen Museums in Wien, 66, 373 - 380. [patels 1 - 3.]" type="journal article" year="1963">Pretzmann 1963</bibRefCitation>
: pl. 2 figs. 5, 6;
<bibRefCitation id="ED453FF59A33FFF0BDE844993EAC17CE" author="Yeo" box="[213,403,224,248]" pageId="4" pageNumber="489" refString="Yeo, D. C. J. &amp; Ng, P. K. L. (2012) Sodhiana, A new genus of freshwater crab from south Asia (Decapoda: Brachyura: Gecarcinucidae). Raffles Bulletin of Zoology, 25, 279 - 284." type="journal article" year="2012">Yeo &amp; Ng 2012</bibRefCitation>
: fig. 2A). The carapace of the
<typeStatus id="566FFCA69A33FFF0BFC044A63C6017CE" box="[765,863,223,248]" pageId="4" pageNumber="489" type="holotype">holotype</typeStatus>
and only specimen of
<taxonomicName id="4ED439879A33FFF0B95144993A2717CE" ID-CoL="6XPDL" box="[1132,1304,224,248]" class="Malacostraca" family="Gecarcinucidae" genus="Sartoriana" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="489" phylum="Arthropoda" rank="species" species="afghaniensis">
<emphasis id="BBA09E169A33FFF0B95144993A2717CE" box="[1132,1304,224,248]" italics="true" pageId="4" pageNumber="489">S. afghaniensis</emphasis>
</taxonomicName>
is partially crushed but fresh material from near the
<typeStatus id="566FFCA69A33FFF0BF51457C3DA3162B" box="[620,668,261,285]" pageId="4" pageNumber="489">type</typeStatus>
locality shows that adults have a distinctly convex dorsal carapace surface (P.K.L. Ng, unpublished).
</paragraph>
</subSubSection>
<subSubSection id="C1CE118F9A33FFF0BDFA45353B381423" pageId="4" pageNumber="489" type="materials_examined">
<paragraph id="896B42049A33FFF0BDFA45353B0C15B2" blockId="4.[151,1437,151,789]" pageId="4" pageNumber="489">
<bibRefCitation id="ED453FF59A33FFF0BDFA45353E871652" box="[199,440,332,357]" pageId="4" pageNumber="489" refString="Kamrani, E., Ng, P. K. L., Mirzadeh, M. &amp; Nakhodai, S. (2009) The marsh crab, Sodhiana rokitanskyi (Pretzmann, 1971) (Decapoda, Brachyura, Gecarcinucidae) from southern Iran. Zootaxa, 2305, 24 - 32." type="journal article">
Kamrani
<emphasis id="BBA09E169A33FFF0BC1245343E581652" box="[303,359,332,356]" italics="true" pageId="4" pageNumber="489">et al.</emphasis>
(2009)
</bibRefCitation>
discussed the differences between
<taxonomicName id="4ED439879A33FFF0BE0345353CEB1652" box="[830,980,332,356]" class="Malacostraca" family="Gecarcinucidae" genus="Sartoriana" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="489" phylum="Arthropoda" rank="species" species="rokitanskyi">
<emphasis id="BBA09E169A33FFF0BE0345353CEB1652" box="[830,980,332,356]" italics="true" pageId="4" pageNumber="489">S. rokitanskyi</emphasis>
</taxonomicName>
and
<taxonomicName id="4ED439879A33FFF0B93445353BB41652" box="[1033,1163,332,356]" class="Malacostraca" family="Gecarcinucidae" genus="Sodhiana" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="489" phylum="Arthropoda" rank="species" species="blanfordi">
<emphasis id="BBA09E169A33FFF0B93445353BB41652" box="[1033,1163,332,356]" italics="true" pageId="4" pageNumber="489">S. blanfordi</emphasis>
</taxonomicName>
, and noted that the form of the posterior margin of the epistome and structure of the proximal tooth on the propodal finger of the larger chela were different. These characters may be affected by size. Although the specimens of
<taxonomicName id="4ED439879A33FFF0B9A445ED3A1D169A" box="[1177,1314,404,428]" class="Malacostraca" family="Gecarcinucidae" genus="Sodhiana" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="489" phylum="Arthropoda" rank="species" species="blanfordi">
<emphasis id="BBA09E169A33FFF0B9A445ED3A1D169A" box="[1177,1314,404,428]" italics="true" pageId="4" pageNumber="489">S. blanfordi</emphasis>
</taxonomicName>
examined (through the courtesy of P.K.L. Ng) had the proximal tooth on the pollex of the larger chela composed of several smaller teeth,
<bibRefCitation id="ED453FF59A33FFF0BC0845A53D4316C3" author="Alcock" box="[309,636,476,501]" pageId="4" pageNumber="489" refString="Alcock, A. (1910) Brachyura I. Fasc. II. The Indian Freshwater Crabs- Potamonidae. Catalogue of the Indian Decapod Crustacea in the collection of the Indian Museum. Calcutta, 135 pp. [plates 1 - 14.]" type="book chapter" year="1910">Alcock (1910: pl. 4 fig. 16A)</bibRefCitation>
figured a large male that had a large complete tooth. This suggests that the large tooth is merely the consequence of complete fusion of several smaller teeth, certainly the case in the present material of
<taxonomicName id="4ED439879A33FFF0BC52465D3EE3150A" box="[367,476,548,572]" class="Malacostraca" family="Gecarcinucidae" genus="Sodhiana" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="489" phylum="Arthropoda" rank="species" species="iranica">
<emphasis id="BBA09E169A33FFF0BC52465D3EE3150A" box="[367,476,548,572]" italics="true" pageId="4" pageNumber="489">S. iranica</emphasis>
</taxonomicName>
(
<figureCitation id="11EF5E819A33FFF0BCD6465D3D12150A" box="[491,557,548,573]" captionStart="FIGURE 2" captionStartId="2.[151,250,1373,1395]" captionTargetBox="[340,1246,193,1351]" captionTargetId="figure@2.[340,1247,193,1351]" captionTargetPageId="2" captionText="FIGURE 2. Sodhiana iranica n. sp., A, C, D, F, G, holotype male (30.2 × 23.1 mm) (ZRC 2014.0240), Bastak, Iran; B, H, paratype male (29.3 × 23.0 mm) (ZRC 2014.0241), Bastak, Iran; E, I, paratype female (29.0 × 23.1 mm) (ZRC 2014.0241), Bastak, Iran. A, B, dorsal views of carapaces; C, left third maxilliped; D, male abdomen (somites 2 6 and telson); E, female thoracic sternum showing vulvae; F, H, I, major chelae; G, minor chela." httpUri="https://zenodo.org/record/227755/files/figure.png" pageId="4" pageNumber="489">Fig. 2</figureCitation>
F, H, I). The same appears to be true for the form of the median triangular lobe on the posterior margin of the epistome. Smaller specimens have the lateral margins gently concave (cf.
<bibRefCitation id="ED453FF59A33FFF0B81D463E3FC815B2" author="Kamrani" pageId="4" pageNumber="489" refString="Kamrani, E., Ng, P. K. L., Mirzadeh, M. &amp; Nakhodai, S. (2009) The marsh crab, Sodhiana rokitanskyi (Pretzmann, 1971) (Decapoda, Brachyura, Gecarcinucidae) from southern Iran. Zootaxa, 2305, 24 - 32." type="journal article" year="2009">
Kamrani
<emphasis id="BBA09E169A33FFF0B8B546303F8B15B2" italics="true" pageId="4" pageNumber="489">et al.</emphasis>
2009
</bibRefCitation>
: fig. 2B) but larger ones have almost straight lateral margins (
<figureCitation id="11EF5E819A33FFF0BE9646153CD115B2" box="[939,1006,620,645]" captionStart="FIGURE 4" captionStartId="4.[151,250,1712,1734]" captionTargetBox="[340,1246,844,1690]" captionTargetId="figure@4.[340,1247,844,1691]" captionTargetPageId="4" captionText="FIGURE 4. Sodhiana blanfordi. A, C, E G, syntype male (43.2 × 33.2 mm) (ZSI 4043 / 4), Pakistan; B, D, syntype male (46.4 × 35.0 mm) (NHM 1905.10.13.5), Pakistan. A, B, dorsal views of carapaces; C, D, frontal views of carapaces; E, ventral view showing anterior thoracic sternum and abdomen; F, dorsal view of right G 1; G, ventral view of right G 1." httpUri="https://zenodo.org/record/227757/files/figure.png" pageId="4" pageNumber="489">Fig. 4</figureCitation>
C, D).
</paragraph>
<paragraph id="896B42049A33FFF0BDFA46E93B381423" blockId="4.[151,1437,151,789]" pageId="4" pageNumber="489">
It is noteworthy that the locality of the new species in Bastak,
<collectingCountry id="F1C302949A33FFF0BEA546E93CF71591" box="[920,968,656,679]" name="Iran" pageId="4" pageNumber="489">Iran</collectingCountry>
, is over
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to the west of the
<typeStatus id="566FFCA69A33FFF0B85146E93AA3159E" box="[1388,1436,656,680]" pageId="4" pageNumber="489">type</typeStatus>
locality of
<taxonomicName id="4ED439879A33FFF0BC2E46CD3EA715FA" box="[275,408,692,716]" class="Malacostraca" family="Gecarcinucidae" genus="Sodhiana" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="489" phylum="Arthropoda" rank="species" species="blanfordi">
<emphasis id="BBA09E169A33FFF0BC2E46CD3EA715FA" box="[275,408,692,716]" italics="true" pageId="4" pageNumber="489">S. blanfordi</emphasis>
</taxonomicName>
(Pishin, Baluchistan) in western
<collectingCountry id="F1C302949A33FFF0BE2E46CD3C4A15FB" box="[787,885,692,717]" name="Pakistan" pageId="4" pageNumber="489">Pakistan</collectingCountry>
, but separated by various high mountain ranges. The
<typeStatus id="566FFCA69A33FFF0BDF446A13FC615C6" box="[201,249,728,752]" pageId="4" pageNumber="489">type</typeStatus>
locality of
<taxonomicName id="4ED439879A33FFF0BC4A46A13D3015C6" box="[375,527,728,752]" class="Malacostraca" family="Gecarcinucidae" genus="Sartoriana" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="489" phylum="Arthropoda" rank="species" species="rokitanskyi">
<emphasis id="BBA09E169A33FFF0BC4A46A13D3015C6" box="[375,527,728,752]" italics="true" pageId="4" pageNumber="489">S. rokitanskyi</emphasis>
</taxonomicName>
occurs in near a hot spring in Geno, southern
<collectingCountry id="F1C302949A33FFF0B93046A13B0515D9" box="[1037,1082,728,751]" name="Iran" pageId="4" pageNumber="489">Iran</collectingCountry>
(
<bibRefCitation id="ED453FF59A33FFF0B97546AE3A1315C6" author="Kamrani" box="[1096,1324,727,752]" pageId="4" pageNumber="489" refString="Kamrani, E., Ng, P. K. L., Mirzadeh, M. &amp; Nakhodai, S. (2009) The marsh crab, Sodhiana rokitanskyi (Pretzmann, 1971) (Decapoda, Brachyura, Gecarcinucidae) from southern Iran. Zootaxa, 2305, 24 - 32." type="journal article" year="2009">
Kamrani
<emphasis id="BBA09E169A33FFF0B98F46A03BD515C6" box="[1202,1258,728,752]" italics="true" pageId="4" pageNumber="489">et al.</emphasis>
2009
</bibRefCitation>
), this area is some
<quantity id="4E2CEFE19A33FFF0BDCE46853E771423" box="[243,328,764,789]" metricMagnitude="5" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="489" unit="km" value="200.0">200 km</quantity>
to the east of Bastak,
<collectingCountry id="F1C302949A33FFF0BF0246843D501422" box="[575,623,765,788]" name="Iran" pageId="4" pageNumber="489">Iran</collectingCountry>
, the
<typeStatus id="566FFCA69A33FFF0BF9946843DEB1423" box="[676,724,765,789]" pageId="4" pageNumber="489">type</typeStatus>
locality of
<taxonomicName id="4ED439879A33FFF0BE6946853CFE1422" box="[852,961,764,788]" class="Malacostraca" family="Gecarcinucidae" genus="Sodhiana" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="489" phylum="Arthropoda" rank="species" species="iranica" status="sp. nov.">
<emphasis id="BBA09E169A33FFF0BE6946853CFE1422" box="[852,961,764,788]" italics="true" pageId="4" pageNumber="489">S. iranica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A093236D9A33FFF0BEF546853B381423" box="[968,1031,764,789]" pageId="4" pageNumber="489" rank="species">
<emphasis id="BBA09E169A33FFF0BEF546853B381423" bold="true" box="[968,1031,764,789]" pageId="4" pageNumber="489">n. sp.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>