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<document id="554E242C963222FF934CC69EC8728822" ID-CLB-Dataset="62482" ID-DOI="10.5281/zenodo.6603807" ID-GBIF-Dataset="916bf600-5bc0-438d-9c6c-7fff4c5a388f" ID-ISBN="978-84-941892-3-4" ID-Zenodo-Dep="6603807" IM.illustrations_requiresApprovalFor="plazi" IM.metadata_requiresApprovalFor="plazi" IM.taxonomicNames_requiresApprovalFor="plazi" checkinTime="1654101401716" checkinUser="carolina" docAuthor="Don E. Wilson, Thomas E. Lacher, Jr &amp; Russell A. Mittermeier" docDate="2016" docId="0131878A073EFF90FA95F71D6A0F43B6" docLanguage="en" docName="hbmw_6_Geomyidae_0234.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 6 Lagomorphs and Rodents I, Barcelona: Lynx Edicions" docTitle="Cratogeomys merriami" docType="treatment" docVersion="9" lastPageNumber="252" masterDocId="FD08FFF2072CFF83FFF3FFF96B0F4602" masterDocTitle="Geomyidae" masterLastPageNumber="269" masterPageNumber="234" pageNumber="251" updateTime="1699338600100" updateUser="ExternalLinkService">
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<mods:title id="B59AB381CE0E572D2F8A123DEC76C801">Geomyidae</mods:title>
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<mods:namePart id="E16C6D5687AD513E4A0B3E5C4FCFFAC4">Don E. Wilson</mods:namePart>
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<mods:namePart id="0A44D2153D36FE802DD51A3E3A7A54D7">Thomas E. Lacher, Jr</mods:namePart>
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<mods:namePart id="600D6E633B7D14DB0A4E426FE1508634">Russell A. Mittermeier</mods:namePart>
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<mods:title id="651734D622A2BB93AFE64FE216936268">Handbook of the Mammals of the World Volume 6 Lagomorphs and Rodents I</mods:title>
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<treatment id="0131878A073EFF90FA95F71D6A0F43B6" ID-DOI="http://doi.org/10.5281/zenodo.6603747" ID-GBIF-Taxon="195708558" ID-Zenodo-Dep="6603747" LSID="urn:lsid:plazi:treatment:0131878A073EFF90FA95F71D6A0F43B6" httpUri="http://treatment.plazi.org/id/0131878A073EFF90FA95F71D6A0F43B6" lastPageId="19" lastPageNumber="252" pageId="18" pageNumber="251">
<subSubSection id="C1826517073EFF91FA95F71D6E914F10" box="[1382,1438,2276,2322]" pageId="18" pageNumber="251" type="multiple">
<paragraph id="8927369C073EFF91FA95F71D6E914F10" blockId="18.[1379,2463,2276,2363]" box="[1382,1438,2276,2322]" pageId="18" pageNumber="251">
<heading id="D26F81F0073EFF91FA95F71D6E914F10" box="[1382,1438,2276,2322]" pageId="18" pageNumber="251">
<figureCitation id="11A32A19073EFF91FA95F71D6E914F10" box="[1382,1438,2276,2322]" captionStart="Plate 13: Geomyidae" captionStartId="14.[104,134,3318,3339]" captionTargetBox="[12,2731,15,3644]" captionTargetPageId="13" captionText="25. Michoacan Pocket Gopher (Zygogeomys trichopus), 26. Giant Pocket Gopher (Orthogeomys grandis), 27. Hispid Pocket Gopher (Heterogeomys hispidus), 28. Big Pocket Gopher (Heterogeomys lanius), 29. Chiriqui Pocket Gopher (Heterogeomys cavator), 30. Darien Pocket Gopher (Heterogeomys dariensis), 31. Cherries Pocket Gopher (Heterogeomys cherrier), 32. Variable Pocket Gopher (Heterogeomys heterodus), 33. Underwoods Pocket Gopher (Heterogeomys underwood), 34. Bullers Pocket Gopher (Pappogeomys bulleri), 35. Yellow-faced Pocket Gopher (Cratogeomys castanops), 36. Goldmans Pocket Gopher (Cratogeomys goldman), 37. Oriental Basin Pocket Gopher (Cratogeomys fulvescens), 38. Merriams Pocket Gopher (Cratogeomys merriami), 39. Cofre de Perote Pocket Gopher (Cratogeomys perotensis), 40. Volcan de Toluca Pocket Gopher (Cratogeomys planiceps), 41. Smoky Pocket Gopher (Cratogeomys fumosus)" figureDoi="http://doi.org/10.5281/zenodo.6603903" httpUri="https://zenodo.org/record/6603903/files/figure.png" pageId="18" pageNumber="251">38.</figureCitation>
</heading>
</paragraph>
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<subSubSection id="C1826517073EFF91FA5CF71D6CC24F10" box="[1455,1997,2276,2322]" pageId="18" pageNumber="251" type="vernacular_names">
<paragraph id="8927369C073EFF91FA5CF71D6CC24F10" blockId="18.[1379,2463,2276,2363]" box="[1455,1997,2276,2322]" pageId="18" pageNumber="251">
<heading id="D26F81F0073EFF91FA5CF71D6CC24F10" box="[1455,1997,2276,2322]" pageId="18" pageNumber="251">
<vernacularName id="079B46B2073EFF91FA5CF71D6CC24F10" box="[1455,1997,2276,2322]" pageId="18" pageNumber="251">Merriams Pocket Gopher</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C1826517073EFF91F7E5F71D62914F10" box="[2070,2462,2276,2322]" pageId="18" pageNumber="251" type="nomenclature">
<paragraph id="8927369C073EFF91F7E5F71D62914F10" blockId="18.[1379,2463,2276,2363]" box="[2070,2462,2276,2322]" pageId="18" pageNumber="251">
<heading id="D26F81F0073EFF91F7E5F71D62914F10" box="[2070,2462,2276,2322]" pageId="18" pageNumber="251">
<taxonomicName id="4E984D1F073EFF91F7E5F71D62914F10" ID-CoL="Z9MM" baseAuthorityName="Thomas" baseAuthorityYear="1893" box="[2070,2462,2276,2322]" class="Mammalia" family="Geomyidae" genus="Cratogeomys" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="251" phylum="Chordata" rank="species" species="merriami">
<emphasis id="BBECEA8E073EFF91F7E5F71D62914F10" box="[2070,2462,2276,2322]" italics="true" pageId="18" pageNumber="251">Cratogeomys merriami</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C1826517073EFF91FA97F6DD63EA4F3B" box="[1380,2277,2340,2361]" pageId="18" pageNumber="251" type="vernacular_names">
<paragraph id="8927369C073EFF91FA97F6DD63EA4F3B" blockId="18.[1379,2463,2276,2363]" box="[1380,2277,2340,2361]" pageId="18" pageNumber="251">
<heading id="D26F81F0073EFF91FA97F6DD63EA4F3B" box="[1380,2277,2340,2361]" pageId="18" pageNumber="251">
<emphasis id="BBECEA8E073EFF91FA97F6DD6EBF4F3B" bold="true" box="[1380,1456,2340,2361]" pageId="18" pageNumber="251">French:</emphasis>
<vernacularName id="079B46B2073EFF91FA4AF6DD6D794F3B" box="[1465,1654,2340,2361]" pageId="18" pageNumber="251">Gaufre de Merriam</vernacularName>
/
<emphasis id="BBECEA8E073EFF91F97FF6DD6DE94F3B" bold="true" box="[1676,1766,2340,2361]" pageId="18" pageNumber="251">German:</emphasis>
<vernacularName id="079B46B2073EFF91F903F6DD6CC74F3B" box="[1776,1992,2340,2361]" pageId="18" pageNumber="251">Merriam-Taschenratte</vernacularName>
/
<emphasis id="BBECEA8E073EFF91F82DF6DD63364F3B" bold="true" box="[2014,2105,2340,2361]" pageId="18" pageNumber="251">Spanish:</emphasis>
<vernacularName id="079B46B2073EFF91F7CCF6DD63EA4F3B" box="[2111,2277,2340,2361]" pageId="18" pageNumber="251">Tuza de Merriam</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C1826517073EFF91F83BF69661194F8A" box="[1992,2582,2415,2440]" pageId="18" pageNumber="251" type="reference_group">
<paragraph id="8927369C073EFF91F83BF69661194F8A" blockId="18.[1992,2586,2415,2833]" box="[1992,2582,2415,2440]" pageId="18" pageNumber="251">
<emphasis id="BBECEA8E073EFF91F83BF696636D4F8A" bold="true" box="[1992,2146,2415,2440]" pageId="18" pageNumber="251">Taxonomy.</emphasis>
<taxonomicName id="4E984D1F073EFF91F783F696611E4F8A" ID-CoL="3FQC5" authority="Thomas, 1893" authorityName="Thomas" authorityYear="1893" box="[2160,2577,2415,2440]" class="Mammalia" family="Geomyidae" genus="Geomys" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="251" phylum="Chordata" rank="species" species="merriami">Geomys merriami Thomas, 1893</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection id="C1826517073EFF91F83AF67461194FD4" pageId="18" pageNumber="251" type="materials_examined">
<paragraph id="8927369C073EFF91F83AF67461194FD4" blockId="18.[1992,2586,2415,2833]" pageId="18" pageNumber="251">
<materialsCitation id="39F03CC1073EFF91F83AF67461194FD4" ID-GBIF-Occurrence="3799834323" pageId="18" pageNumber="251">“S. Mexico.” Restricted by C. H. Merriam in 1895 to “probably the Valley of Mexico.”</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C1826517073EFF91F83FF6196DE84DD4" pageId="18" pageNumber="251" type="discussion">
<paragraph id="8927369C073EFF91F83FF6196DE84DD4" blockId="18.[1992,2586,2415,2833]" lastBlockId="18.[1380,2586,2840,3462]" pageId="18" pageNumber="251">
<taxonomicName id="4E984D1F073EFF91F83FF61963674FFF" authorityName="Merriam" authorityYear="1895" box="[1996,2152,2528,2557]" class="Mammalia" family="Geomyidae" genus="Cratogeomys" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="251" phylum="Chordata" rank="genus">Cratogeomys</taxonomicName>
was regarded as a subgenus of
<taxonomicName id="4E984D1F073EFF91F7CCF5FD63E74C27" authorityName="Merriam" authorityYear="1895" box="[2111,2280,2564,2597]" class="Mammalia" family="Geomyidae" genus="Pappogeomys" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="251" phylum="Chordata" rank="genus">Pappogeomys</taxonomicName>
by R. J. Russell in 1968 but was returned to generic status by R. L. Honeycutt and S. L. Willams in 1982. Prior to systematic revision of C.
<taxonomicName id="4E984D1F073EFF91F7D1F55863994CC0" authorityName="Thomas" authorityYear="1893" box="[2082,2198,2721,2754]" class="Mammalia" family="Geomyidae" genus="Geomys" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="251" phylum="Chordata" rank="species" species="merriami">merriami</taxonomicName>
by M. S. Hafner and colleagues in 2005, C.
<taxonomicName id="4E984D1F073EFF91F718F53062604CE8" authorityName="Merriam" authorityYear="1895" box="[2283,2415,2761,2794]" class="Mammalia" family="Geomyidae" genus="Cratogeomys" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="251" phylum="Chordata" rank="species" species="fulvescens">fulvescens</taxonomicName>
and
<taxonomicName id="4E984D1F073EFF91F630F53063114D13" authorityName="Merriam" authorityYear="1895" class="Mammalia" family="Geomyidae" genus="Cratogeomys" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="268" phylum="Chordata" rank="species" species="perotensis">C. perotensis</taxonomicName>
were considered subspecies of C.
<taxonomicName id="4E984D1F073EFF91FA7CF4E16D0C4D3B" authorityName="Thomas" authorityYear="1893" box="[1423,1539,2840,2873]" class="Mammalia" family="Geomyidae" genus="Geomys" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="251" phylum="Chordata" rank="species" species="merriami">merriami</taxonomicName>
. Molecular studies show C.
<taxonomicName id="4E984D1F073EFF91F86EF4E1631F4D3B" authorityName="Thomas" authorityYear="1893" box="[1949,2064,2840,2873]" class="Mammalia" family="Geomyidae" genus="Geomys" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="251" phylum="Chordata" rank="species" species="merriami">merriami</taxonomicName>
to be closely related to C.
<taxonomicName id="4E984D1F073EFF91F666F4E161164D3B" authorityName="Merriam" authorityYear="1895" box="[2453,2585,2840,2873]" class="Mammalia" family="Geomyidae" genus="Cratogeomys" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="251" phylum="Chordata" rank="species" species="fulvescens">fulvescens</taxonomicName>
and
<taxonomicName id="4E984D1F073EFF91FA5AF4BE6D454D62" authorityName="Merriam" authorityYear="1895" box="[1449,1610,2887,2912]" class="Mammalia" family="Geomyidae" genus="Cratogeomys" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="268" phylum="Chordata" rank="species" species="perotensis">C. perotensis</taxonomicName>
, and these three species together with
<taxonomicName id="4E984D1F073EFF91F77AF4BE62264D62" authorityName="Merriam" authorityYear="1895" box="[2185,2345,2887,2912]" class="Mammalia" family="Geomyidae" genus="Cratogeomys" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="267" phylum="Chordata" rank="species" species="goldmani">C. goldmani</taxonomicName>
and
<taxonomicName id="4E984D1F073EFF91F684F4BE61154D62" baseAuthorityName="Baird" baseAuthorityYear="1852" box="[2423,2586,2887,2912]" class="Mammalia" family="Geomyidae" genus="Cratogeomys" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="267" phylum="Chordata" rank="species" species="castanops">C. castanops</taxonomicName>
comprise the
<taxonomicName id="4E984D1F073EFF91F9DCF4936DDE4D85" baseAuthorityName="Baird" baseAuthorityYear="1852" box="[1583,1745,2922,2951]" class="Mammalia" family="Geomyidae" genus="Cratogeomys" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="267" phylum="Chordata" rank="species" species="castanops">C. castanops</taxonomicName>
species group. J. L. Patton in 2005 recognized six subspecies of C.
<taxonomicName id="4E984D1F073EFF91FA1FF4776D6F4DAD" authorityName="Thomas" authorityYear="1893" box="[1516,1632,2958,2991]" class="Mammalia" family="Geomyidae" genus="Geomys" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="251" phylum="Chordata" rank="species" species="merriami">merriami</taxonomicName>
, but M. S. Hafner and colleagues in 2005 found no subspecies-level differentiation. Monotypic.
</paragraph>
</subSubSection>
<subSubSection id="C1826517073EFF91FA97F4256CF64A4E" pageId="18" pageNumber="251" type="distribution">
<caption id="DDE76614073EFF91FA97F4256CF64A4E" ID-DOI="http://doi.org/10.5281/zenodo.6603889" ID-Zenodo-Dep="6603889" httpUri="https://zenodo.org/record/6603889/files/figure.png" inLine="true" pageId="18" pageNumber="251" targetBox="[1377,1968,2415,2829]" targetPageId="18">
<paragraph id="8927369C073EFF91FA97F4256CF64A4E" blockId="18.[1380,2586,2840,3462]" pageId="18" pageNumber="251">
<emphasis id="BBECEA8E073EFF91FA97F4256D1C4DFF" bold="true" box="[1380,1555,3036,3069]" pageId="18" pageNumber="251">Distribution.</emphasis>
Mexican endemic, patchily distributed in the S part of the Valley of Mexico (Sierra de Las Cruces, Sierra de Ajusco, Popocatéptl and Iztaccthutl Mts, and from Lerma at the E Toluca Valley E into W Puebla).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="C1826517073EFF90FA97F3AF69C24761" lastPageId="19" lastPageNumber="252" pageId="18" pageNumber="251" type="description">
<paragraph id="8927369C073EFF90FA97F3AF69C24761" blockId="18.[1380,2586,2840,3462]" lastBlockId="19.[182,1390,283,1460]" lastPageId="19" lastPageNumber="252" pageId="18" pageNumber="251">
<emphasis id="BBECEA8E073EFF91FA97F3AF6D6B4A71" bold="true" box="[1380,1636,3158,3187]" pageId="18" pageNumber="251">Descriptive notes.</emphasis>
Head-body 230-285 mm (males) and 200-255 mm (females), tail 90-125 mm (males) and 80-120 mm (females); weight 580-700 g (males) and 450-670 g (females). Male Merriams Pocket Gophers are usually larger than females, but sexual dimorphism is not as pronounced as it is in many other geomyid species. Merriams Pocket Gopheris large for the genus. It has a fusiform body shape typical of all pocket gophers, and it possesses fur-lined cheek pouches that open external to the mouth. Dorsal coloris highly variable, from dull chestnut-brown to slate black. Ventral color is similar to dorsal color but paler. Anterior surface of each upper incisor has a single medial groove slightly displaced to the inner side of the tooth. Chromosomal complement has 2n = 36 and FN = 68.
</paragraph>
</subSubSection>
<subSubSection id="C1826517073FFF90FF4AFE93693D442A" pageId="19" pageNumber="252" type="biology_ecology">
<paragraph id="8927369C073FFF90FF4AFE93693D442A" blockId="19.[182,1390,283,1460]" pageId="19" pageNumber="252">
<emphasis id="BBECEA8E073FFF90FF4AFE936A274789" bold="true" box="[185,296,362,395]" pageId="19" pageNumber="252">Habitat.</emphasis>
Wide variety of habitats, most characterized by arid conditions, sandy soils, and xeric-adapted vegetation. In parts ofits distribution, Merriams Pocket
<taxonomicName id="4E984D1F073FFF90FAF3FE6C6E6247B0" box="[1280,1389,405,434]" class="Reptilia" family="Testudinidae" genus="Gopher" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="252" phylum="Chordata" rank="genus">Gopher</taxonomicName>
can be found in oak, pine, and fir woodlands and grassy mountain meadows, and it is known from near timberline on volcanic peaks east of Mexico City. It is known from elevations of 1800-4000 m.
</paragraph>
</subSubSection>
<subSubSection id="C1826517073FFF90FF4AFDD66E394566" pageId="19" pageNumber="252" type="food_feeding">
<paragraph id="8927369C073FFF90FF4AFDD66E394566" blockId="19.[182,1390,283,1460]" pageId="19" pageNumber="252">
<emphasis id="BBECEA8E073FFF90FF4AFDD66ACE4452" bold="true" box="[185,449,559,592]" pageId="19" pageNumber="252">Food and Feeding.</emphasis>
There is no specific information available for this species, but all pocket gophers that have been studied eat predominately underground roots and tubers and a limited amount of surface vegetation. Merriams Pocket
<taxonomicName id="4E984D1F073FFF90FB9EFD876FD7449D" box="[1133,1240,638,671]" class="Reptilia" family="Testudinidae" genus="Gopher" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="252" phylum="Chordata" rank="genus">Gopher</taxonomicName>
readily invades cultivated fields and is considered an agricultural pest wherever it comes in contact with humans. As in all other pocket gophers, the burrow system is a series of shallow feeding tunnels radiating spoke-like from a deeper, central network that contains one or more nest chambers and several smaller chambers for storage of food or fecal pellets. Burrow systems of Merriams Pocket
<taxonomicName id="4E984D1F073FFF90FD40FCBA68114566" box="[691,798,835,868]" class="Reptilia" family="Testudinidae" genus="Gopher" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="252" phylum="Chordata" rank="genus">Gopher</taxonomicName>
can encompass 400 m* of surface area.
</paragraph>
</subSubSection>
<subSubSection id="C1826517073FFF90FF4BFC936FBB45D8" pageId="19" pageNumber="252" type="breeding">
<paragraph id="8927369C073FFF90FF4BFC936FBB45D8" blockId="19.[182,1390,283,1460]" pageId="19" pageNumber="252">
<emphasis id="BBECEA8E073FFF90FF4BFC936A314589" bold="true" box="[184,318,874,907]" pageId="19" pageNumber="252">Breeding.</emphasis>
Captures of pregnant female Merriams Pocket Gophers suggest that they breed throughout the year, with peak reproductive activities in October—March. Females may have multiple litters per year, with litters usually of 1-3 young.
</paragraph>
</subSubSection>
<subSubSection id="C1826517073FFF90FF45FC196E65427A" pageId="19" pageNumber="252" type="activity">
<paragraph id="8927369C073FFF90FF45FC196E65427A" blockId="19.[182,1390,283,1460]" pageId="19" pageNumber="252">
<emphasis id="BBECEA8E073FFF90FF45FC196AAE4203" bold="true" box="[182,417,992,1025]" pageId="19" pageNumber="252">Activity patterns.</emphasis>
Merriams Pocket Gophers in the Valley of Mexico were active at any hour of the day, with periods of peak activity around dawn and dusk. Digging activity was generally higher in the dry season (especially December—January) and lower at the beginning of the wet season (June). It does not hibernate and is active year-round.
</paragraph>
</subSubSection>
<subSubSection id="C1826517073FFF90FF4BFB876FC04317" pageId="19" pageNumber="252" type="biology_ecology">
<paragraph id="8927369C073FFF90FF4BFB876FC04317" blockId="19.[182,1390,283,1460]" pageId="19" pageNumber="252">
<emphasis id="BBECEA8E073FFF90FF4BFB876887429D" bold="true" box="[184,904,1150,1183]" pageId="19" pageNumber="252">Movements, Home range and Social organization.</emphasis>
There is no specific information available for this species, but Merriams Pocket Gopheris likely solitary and aggressively territorial. Individuals probably leave their burrow systems only rarely, meaning that their home range is defined by size and extent of their burrow system.
</paragraph>
</subSubSection>
<subSubSection id="C1826517073FFF90FF4BFAE56FDD433F" box="[184,1234,1308,1341]" pageId="19" pageNumber="252" type="conservation">
<paragraph id="8927369C073FFF90FF4BFAE56FDD433F" blockId="19.[182,1390,283,1460]" box="[184,1234,1308,1341]" pageId="19" pageNumber="252">
<emphasis id="BBECEA8E073FFF90FF4BFAE5691A433F" bold="true" box="[184,533,1308,1341]" pageId="19" pageNumber="252">Status and Conservation.</emphasis>
Classified as Least Concern on The IUCN Red Last.
</paragraph>
</subSubSection>
<subSubSection id="C1826517073FFF90FF4AFAB56A0F43B6" pageId="19" pageNumber="252" type="bibRefCitation_list">
<paragraph id="8927369C073FFF90FF4AFAB56A0F43B6" blockId="19.[182,1390,283,1460]" pageId="19" pageNumber="252">
<emphasis id="BBECEA8E073FFF90FF4AFAB56A5E4367" bold="true" box="[185,337,1356,1381]" pageId="19" pageNumber="252">Bibliography.</emphasis>
Alvarez-Castafieda, Castro-Arellano, Lacher &amp; Vazquez (2008f), Ceballos (2014), Hafner et al. (2005), Honeycutt &amp; Williams (1982), Merriam (1895), Patton (2005b), Russell (1968b), Whisson &amp; Villa-Cornejo (1994).
</paragraph>
</subSubSection>
</treatment>
</document>