369 lines
39 KiB
XML
369 lines
39 KiB
XML
<document ID-DOI="10.11646/phytotaxa.577.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="7517526" checkinTime="1673303962232" checkinUser="plazi" docAuthor="Castillón, Eduardo Estrada, Rebman, Jon P., González, Carlos & Quintanilla, José Ángel Villarreal" docDate="2022" docId="536B878DFFB3FFB9FF2C0AEFFA34F7E3" docLanguage="en" docName="phytotaxa.577.1.1.pdf" docOrigin="Phytotaxa 577 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.577.1.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.5:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="5" docTitle="Astragalus tijuanensis A. E. Estrada, Rebman, C. Gonzalez & Villarreal 2022, sp. nov." docType="treatment" docVersion="1" lastPageNumber="3" masterDocId="AF52FFF5FFB2FFBBFFA40C7BFFB1FFCF" masterDocTitle="Astragalus tijuanensis and Astragalus brauntonii var. lativexillum, a new species and a new variety in the genus Astragalus (Fabaceae) from extreme northwestern Baja California, Mexico" masterLastPageNumber="13" masterPageNumber="1" pageNumber="2" updateTime="1673552936955" updateUser="jonas" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Astragalus tijuanensis and Astragalus brauntonii var. lativexillum, a new species and a new variety in the genus Astragalus (Fabaceae) from extreme northwestern Baja California, Mexico</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Castillón, Eduardo Estrada</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0003-1061-9862</mods:nameIdentifier>
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<mods:affiliation>Facultad de Ciencias Forestales, Universidad Autónoma de Nuevo León, C. P. 67700, Linares, Nuevo León, México. aeduardoestradac @ prodigy. net; https: // orcid. org / 0000 - 0003 - 1061 - 9862</mods:affiliation>
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<mods:nameIdentifier type="email">aeduardoestradac@prodigy.net</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rebman, Jon P.</mods:namePart>
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<mods:affiliation>San Diego Natural History Museum, P. O. Box 121390, San Diego, California, 92112.</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>González, Carlos</mods:namePart>
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<mods:affiliation>Jaime Nuno 2227, Col. Hidalgo, C. P. 22880, Ensenada, Baja California, México.</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Quintanilla, José Ángel Villarreal</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Phytotaxa</mods:title>
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</mods:titleInfo>
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||
<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-12-29</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>577</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>13</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.11646/phytotaxa.577.1.1</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/phytotaxa.577.1.1</mods:identifier>
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<mods:identifier type="ISSN">1179-3163</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7517526</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:536B878DFFB3FFB9FF2C0AEFFA34F7E3" httpUri="http://treatment.plazi.org/id/536B878DFFB3FFB9FF2C0AEFFA34F7E3" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
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<paragraph blockId="1.[136,1260,1684,1711]" box="[136,1260,1684,1711]" pageId="1" pageNumber="2">
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<heading box="[136,1260,1684,1711]" fontSize="11" level="2" pageId="1" pageNumber="2" reason="3">
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<taxonomicName authority="A. E. Estrada, Rebman, C. Gonzalez & Villarreal" authorityName="A. E. Estrada, Rebman, C. Gonzalez & Villarreal" authorityYear="2022" box="[136,954,1684,1711]" class="Magnoliopsida" family="Fabaceae" genus="Astragalus" kingdom="Plantae" order="Fabales" pageId="1" pageNumber="2" phylum="Tracheophyta" rank="species" species="tijuanensis" status="sp. nov.">
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<emphasis bold="true" box="[136,394,1684,1711]" italics="true" pageId="1" pageNumber="2">Astragalus tijuanensis</emphasis>
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A. E. Estrada, Rebman, C. González & Villarreal
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</taxonomicName>
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||
,
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||
<emphasis box="[964,1049,1685,1710]" italics="true" pageId="1" pageNumber="2">
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<taxonomicNameLabel box="[964,1049,1685,1710]" pageId="1" pageNumber="2" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
(
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<figureCitation box="[1064,1130,1684,1710]" captionStart="FIGURE 1" captionStartId="3.[136,229,1922,1944]" captionTargetBox="[177,1436,190,1898]" captionTargetId="figure-19@3.[151,1436,190,1898]" captionTargetPageId="3" captionText="FIGURE 1. Astragalus tijuanensis, A) Whole plant showing leaves, inflorescences, flowers and fruits. B) Stipules. C) Bracts. D) inflorescence enlarged to see detail of the flowers. E) Calyx. F) Banner: front view (left) and side view (right). G) Wing. H) Keel. I) Stamens. J) Ovary. K) Fruit. L) Seed." figureDoi="http://doi.org/10.5281/zenodo.7517528" httpUri="https://zenodo.org/record/7517528/files/figure.png" pageId="1" pageNumber="2">Fig. 1</figureCitation>
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and
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<figureCitation box="[1185,1251,1684,1710]" captionStart="FIGURE 2" captionStartId="4.[136,229,1962,1984]" captionTargetBox="[138,1449,190,1938]" captionTargetId="figure-28@4.[138,1449,190,1938]" captionTargetPageId="4" captionText="FIGURE 2. Astragalus tijuanensis, part of the plant showing its leaves, inflorescences, flowers, and fruits." figureDoi="http://doi.org/10.5281/zenodo.7517530" httpUri="https://zenodo.org/record/7517530/files/figure.png" pageId="1" pageNumber="2">Fig. 2</figureCitation>
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)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="2" type="diagnosis">
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<paragraph blockId="1.[136,1452,1720,2071]" pageId="1" pageNumber="2">
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Morphologically similar to
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<taxonomicName authority="A. Gray" authorityName="A. Gray" authorityYear="1868" box="[499,813,1720,1746]" class="Magnoliopsida" family="Fabaceae" genus="Astragalus" kingdom="Plantae" order="Fabales" pageId="1" pageNumber="2" phylum="Tracheophyta" rank="species" species="palmeri">
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<emphasis box="[499,714,1720,1746]" italics="true" pageId="1" pageNumber="2">Astragalus palmeri</emphasis>
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A. Gray
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</taxonomicName>
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, but differing in
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<typeStatus box="[1001,1049,1721,1747]" pageId="1" pageNumber="2">type</typeStatus>
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and stipule size, number of flowers per raceme, petal colors, curvature and size of the banner, number of ovules, and diameter of the pod and the absence of pubescence.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="2" type="materials_examined">
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<paragraph blockId="1.[136,1452,1720,2071]" pageId="1" pageNumber="2">
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<materialsCitation collectingDate="2022-06-05" collectionCode="SD, MEXU" collectorName="C. Gonzalez" country="Mexico" county="Type" elevation="171" latitude="32.3364" location="Tijuana" longLatPrecision="1" longitude="-116.96239" municipality="Municipio Tijuana" pageId="1" pageNumber="2" specimenCount="1" stateProvince="Baja California" typeStatus="holotype">
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<collectingCounty box="[189,248,1828,1854]" pageId="1" pageNumber="2">Type</collectingCounty>
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:—
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<collectingCountry box="[281,395,1828,1854]" name="Mexico" pageId="1" pageNumber="2">MEXICO</collectingCountry>
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.
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<collectingRegion box="[405,578,1828,1854]" country="Mexico" name="Baja California" pageId="1" pageNumber="2">Baja California</collectingRegion>
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||
(
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||
<collectingMunicipality box="[592,808,1828,1854]" pageId="1" pageNumber="2">
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Municipio
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<location LSID="urn:lsid:plazi:treatment:536B878DFFB3FFB9FF2C0AEFFA34F7E3:DE1D6040FFB3FFBAFD690B5FFC99F8F1" box="[717,808,1828,1854]" country="Mexico" county="Type" latitude="32.3364" longLatPrecision="1" longitude="-116.96239" municipality="Municipio Tijuana" name="Tijuana" pageId="1" pageNumber="2" stateProvince="Baja California">Tijuana</location>
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</collectingMunicipality>
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||
): along
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||
<location LSID="urn:lsid:plazi:treatment:536B878DFFB3FFB9FF2C0AEFFA34F7E3:DE1D6040FFB3FFBAFCDB0B5FFC45F8F1" box="[895,1012,1828,1854]" country="Mexico" county="Type" latitude="32.3364" longLatPrecision="1" longitude="-116.96239" municipality="Municipio Tijuana" name="Boulevard" pageId="1" pageNumber="2" stateProvince="Baja California">Boulevard</location>
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2000
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<locationDeviation box="[1082,1419,1828,1854]" firstLocation="Rosarito" pageId="1" pageNumber="2" secondLocation="Tijuana">between Rosarito and Tijuana</locationDeviation>
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, in the vicinity of the south side of the metropolitan area of the city of
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<location LSID="urn:lsid:plazi:treatment:536B878DFFB3FFB9FF2C0AEFFA34F7E3:DE1D6040FFB3FFBAFCD90B33FC64F8AD" box="[893,981,1864,1890]" country="Mexico" county="Type" latitude="32.3364" longLatPrecision="1" longitude="-116.96239" municipality="Municipio Tijuana" name="Tijuana" pageId="1" pageNumber="2" stateProvince="Baja California">Tijuana</location>
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,
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<geoCoordinate box="[992,1181,1864,1891]" degrees="32" direction="north" minutes="20" orientation="latitude" pageId="1" pageNumber="2" precision="1" seconds="11.04" value="32.3364">32° 20’11.04” N</geoCoordinate>
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,
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<geoCoordinate box="[1192,1405,1864,1891]" degrees="116" direction="west" minutes="57" orientation="longitude" pageId="1" pageNumber="2" precision="1" seconds="44.593" value="-116.96239">116°57’44.593” W</geoCoordinate>
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, alt.
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<quantity box="[136,209,1900,1927]" metricMagnitude="2" metricUnit="m" metricValue="1.71" pageId="1" pageNumber="2" unit="m" value="171.0">
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||
<elevation box="[136,209,1900,1927]" metricMagnitude="2" metricUnit="m" metricValue="1.71" pageId="1" pageNumber="2" unit="m" value="171.0">171 m</elevation>
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</quantity>
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||
,
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<date box="[221,359,1900,1927]" pageId="1" pageNumber="2" value="2022-06-05">
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<collectingDate box="[221,359,1900,1927]" pageId="1" pageNumber="2" value="2022-06-05">5 June 2022</collectingDate>
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</date>
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||
,
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||
<emphasis box="[370,556,1900,1926]" italics="true" pageId="1" pageNumber="2">
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<collectorName box="[370,507,1900,1926]" pageId="1" pageNumber="2">C. González</collectorName>
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104
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</emphasis>
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||
(
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<typeStatus box="[571,677,1900,1926]" pageId="1" pageNumber="2">Holotype</typeStatus>
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<collectionCode box="[684,728,1900,1926]" country="USA" lsid="urn:lsid:biocol.org:col:15659" name="San Diego Natural History Museum" pageId="1" pageNumber="2" type="Herbarium">SD</collectionCode>
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!; isotype
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<collectionCode box="[831,924,1900,1926]" country="Mexico" lsid="urn:lsid:biocol.org:col:15658" name="Universidad Nacional Autónoma de México" pageId="1" pageNumber="2" type="Herbarium">MEXU</collectionCode>
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!)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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||
<subSubSection lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2" type="description">
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<paragraph blockId="1.[136,1452,1720,2071]" lastBlockId="2.[136,1452,158,2093]" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
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<emphasis bold="true" box="[189,331,1936,1962]" pageId="1" pageNumber="2">Herbaceous</emphasis>
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, caulescent, perennial with deep single buried root, without superficial crown of a woody taproot, 13 ×
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<quantity box="[193,268,1972,1999]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="2" unit="cm" value="0.7">0.7 cm</quantity>
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long.
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<emphasis bold="true" box="[338,410,1972,1998]" pageId="1" pageNumber="2">Stems</emphasis>
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three to several, up to
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<quantity box="[663,731,1972,1999]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="2" unit="cm" value="30.0">30 cm</quantity>
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long,
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<quantity box="[801,915,1972,1999]" metricMagnitude="-3" metricUnit="m" metricValue="5.25" metricValueMax="7.5" metricValueMin="3.0" pageId="1" pageNumber="2" unit="mm" value="5.25" valueMax="7.5" valueMin="3.0">3–7.5 mm</quantity>
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diameter, erect, arising from the base, the lower part of the stems,
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<quantity box="[337,438,2008,2035]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="1" pageNumber="2" unit="cm" value="8.5" valueMax="10.0" valueMin="7.0">7–10 cm</quantity>
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, gray color, with stipules and lower leaves persistent, the leaves dried throughout, without leaflets, the upper part of the stem green, striate, minutely and densely strigulose, the trichomes
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<quantity box="[1248,1386,2044,2071]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="1" pageNumber="2" unit="mm" value="0.2" valueMax="0.3" valueMin="0.1">0.1–0.3 mm</quantity>
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long, appressed, white.
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<emphasis bold="true" box="[340,435,158,184]" pageId="2" pageNumber="3">Stipules</emphasis>
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free or semi–amplexicaul, clasping ¼ to ½ of the stem´s circumference, 2–3 ×
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<quantity box="[1350,1447,158,185]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="2" pageNumber="3" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
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, persistent, almost always wider than longer, triangular to triangular–ovate, white to light brown, sparsely strigulose or glabrate, with only few trichomes on the edges.
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<emphasis bold="true" box="[734,816,230,256]" pageId="2" pageNumber="3">Leaves</emphasis>
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odd–pinnate,
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<quantity box="[983,1111,230,257]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" metricValueMax="8.5" metricValueMin="4.5" pageId="2" pageNumber="3" unit="cm" value="6.5" valueMax="8.5" valueMin="4.5">4.5–8.5 cm</quantity>
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long, petiole (3–)11–12(–17) mm long, leaflets (17)21–29(33) per leaf, opposite, rarely some of them alternate, (2–)8–10 ×
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<quantity box="[1224,1342,266,293]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="2" pageNumber="3" unit="mm" value="0.85" valueMax="1.0" valueMin="0.7">0.7–1 mm</quantity>
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||
, linear to linear–oblong, petiolule
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<quantity box="[412,499,302,329]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="3" unit="mm" value="0.2">0.2 mm</quantity>
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long, separated from each other by
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||
<quantity box="[906,1002,302,329]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="2" pageNumber="3" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
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||
, glabrate and light–green adaxially, and dark–green and sparsely strigulose abaxially as stem pubescence, flattened or slightly concave, acute at both ends.
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||
<emphasis bold="true" box="[136,305,374,400]" pageId="2" pageNumber="3">Inflorescences</emphasis>
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in axillary open racemes,
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||
<quantity box="[619,727,374,401]" metricMagnitude="-2" metricUnit="m" metricValue="4.75" metricValueMax="5.0" metricValueMin="4.5" pageId="2" pageNumber="3" unit="cm" value="4.75" valueMax="5.0" valueMin="4.5">4.5–5 cm</quantity>
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||
long, peduncles erect or incurved–ascending, when young (in flower stage),
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<quantity box="[298,381,410,437]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="2" pageNumber="3" unit="cm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 cm</quantity>
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||
long, elongating with age (5–)7–11(–14) cm long (in fruit), exceeding the length of the leaves, sparsely strigulose,
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||
<quantity box="[359,495,446,473]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="2" pageNumber="3" unit="mm" value="0.15" valueMax="0.2" valueMin="0.1">0.1–0.2 mm</quantity>
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long, the trichomes white; floriferous axis
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||
<quantity box="[985,1090,446,473]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="2" pageNumber="3" unit="cm" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 cm</quantity>
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||
long, elongating
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<quantity box="[1289,1393,446,473]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" metricValueMax="5.0" metricValueMin="2.2" pageId="2" pageNumber="3" unit="cm" value="3.6" valueMax="5.0" valueMin="2.2">2.2–5 cm</quantity>
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||
long in fruit, with 13–19 flowers per raceme.
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||
<emphasis bold="true" box="[592,670,482,508]" pageId="2" pageNumber="3">Bracts</emphasis>
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||
0.6–1.2 ×
|
||
<quantity box="[791,905,482,509]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="2" pageNumber="3" unit="mm" value="0.85" valueMax="1.0" valueMin="0.7">0.7–1 mm</quantity>
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||
long, ovate to triangular, white to green, opaque to somewhat translucent, sometimes with slight purple tones in the upper half, subglabrate to sparsely strigulose, sometimes the midvein is evident, pedicel ascending or spreading, arched outward,
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||
<quantity box="[1068,1182,554,581]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="2" pageNumber="3" unit="mm" value="1.15" valueMax="1.3" valueMin="1.0">1–1.3 mm</quantity>
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||
long, slightly thickened in fruit, caducous; bracteoles absent.
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||
<emphasis bold="true" box="[559,654,590,616]" pageId="2" pageNumber="3">Flowers</emphasis>
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||
<quantity box="[663,785,590,617]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="2" pageNumber="3" unit="mm" value="12.5" valueMax="13.0" valueMin="12.0">12–13 mm</quantity>
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||
long, calyx (3.5–)3.9–4(–4.2) ×
|
||
<quantity box="[1158,1297,590,617]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" metricValueMax="2.7" metricValueMin="2.5" pageId="2" pageNumber="3" unit="mm" value="2.6" valueMax="2.7" valueMin="2.5">2.5–2.7 mm</quantity>
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||
, campanulate to wide–campanulate, the tube
|
||
<quantity box="[488,624,626,653]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="3.3" metricValueMin="3.2" pageId="2" pageNumber="3" unit="mm" value="3.25" valueMax="3.3" valueMin="3.2">3.2–3.3 mm</quantity>
|
||
long, inequilaterous or slightly oblique sparsely strigulose, the trichomes white and purple mixed, occasionally subglabrous with few trichomes, the teeth
|
||
<quantity box="[1089,1229,662,689]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="2" pageNumber="3" unit="mm" value="0.8" valueMax="0.9" valueMin="0.7">0.7–0.9 mm</quantity>
|
||
long, triangular to triangular–subulate, separated by obtuse or rounded sinuses; petals white with purple tones, the banner sessile, 12.5–13 ×
|
||
<quantity box="[161,279,734,761]" metricMagnitude="-3" metricUnit="m" metricValue="6.65" metricValueMax="7.0" metricValueMin="6.3" pageId="2" pageNumber="3" unit="mm" value="6.65" valueMax="7.0" valueMin="6.3">6.3–7 mm</quantity>
|
||
, obovate, elliptic obovate to obovate–rhombic, gradually attenuated at the base, the notch
|
||
<quantity box="[1315,1452,734,761]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="2" pageNumber="3" unit="mm" value="0.45" valueMax="0.5" valueMin="0.4">0.4–0.5 mm</quantity>
|
||
deep, folded at the middle, its margins then folded backward, slightly bent backward from the keel, recurved through 25°–30° from the vertical; wings 11.9–12.2 ×
|
||
<quantity box="[652,769,806,833]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" metricValueMax="2.3" metricValueMin="2.0" pageId="2" pageNumber="3" unit="mm" value="2.15" valueMax="2.3" valueMin="2.0">2–2.3 mm</quantity>
|
||
, the claw
|
||
<quantity box="[882,996,806,833]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" metricValueMax="5.2" metricValueMin="5.0" pageId="2" pageNumber="3" unit="mm" value="5.1" valueMax="5.2" valueMin="5.0">5–5.2 mm</quantity>
|
||
long, the blade
|
||
<quantity box="[1176,1290,806,833]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" metricValueMax="7.0" metricValueMin="6.8" pageId="2" pageNumber="3" unit="mm" value="6.9" valueMax="7.0" valueMin="6.8">6.8–7 mm</quantity>
|
||
long, straight, narrowly obliquely oblong–obovate, rounded to truncate at apex, basally auriculate; keel 9.4–9.8 ×
|
||
<quantity box="[1266,1404,842,869]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="2" pageNumber="3" unit="mm" value="2.65" valueMax="2.8" valueMin="2.5">2.5–2.8 mm</quantity>
|
||
, the claw
|
||
<quantity box="[195,308,878,905]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" metricValueMax="5.0" metricValueMin="4.8" pageId="2" pageNumber="3" unit="mm" value="4.9" valueMax="5.0" valueMin="4.8">4.8–5 mm</quantity>
|
||
long, the blade
|
||
<quantity box="[485,598,878,905]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" metricValueMax="5.0" metricValueMin="4.6" pageId="2" pageNumber="3" unit="mm" value="4.8" valueMax="5.0" valueMin="4.6">4.6–5 mm</quantity>
|
||
long, half obovate, incurved, apex rounded, auriculate; Stamens 10–merous, diadelphous, 9 fused by their filaments into a white striped sheath
|
||
<quantity box="[919,1036,914,941]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" metricValueMax="5.2" metricValueMin="5.0" pageId="2" pageNumber="3" unit="mm" value="5.1" valueMax="5.2" valueMin="5.0">5–5.2 mm</quantity>
|
||
long, the free part
|
||
<quantity box="[1267,1384,914,941]" metricMagnitude="-3" metricUnit="m" metricValue="4.15" metricValueMax="4.3" metricValueMin="4.0" pageId="2" pageNumber="3" unit="mm" value="4.15" valueMax="4.3" valueMin="4.0">4–4.3 mm</quantity>
|
||
long, enfolding the ovary, the vexilar stamen
|
||
<quantity box="[574,708,950,977]" metricMagnitude="-3" metricUnit="m" metricValue="8.95" metricValueMax="9.3" metricValueMin="8.6" pageId="2" pageNumber="3" unit="mm" value="8.95" valueMax="9.3" valueMin="8.6">8.6–9.3 mm</quantity>
|
||
long, standing free; anthers yellow, round, 0.9–1 ×
|
||
<quantity box="[1278,1366,950,977]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="3" unit="mm" value="0.2">0.2 mm</quantity>
|
||
.
|
||
<emphasis bold="true" box="[1375,1451,950,976]" pageId="2" pageNumber="3">Ovary</emphasis>
|
||
unilocular, 32–38 ovulate, sessile, linear–oblong, green, glabrous, 4.2–4.8 ×
|
||
<quantity box="[1007,1095,986,1013]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="3" unit="mm" value="0.8">0.8 mm</quantity>
|
||
, the style linear, glabrous, style
|
||
<quantity box="[136,252,1022,1049]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="2" pageNumber="3" unit="mm" value="4.25" valueMax="4.5" valueMin="4.0">4–4.5 mm</quantity>
|
||
long, glabrate, incurved, stigma terminal, minute, glabrous.
|
||
<emphasis bold="true" box="[945,992,1022,1048]" pageId="2" pageNumber="3">Pod</emphasis>
|
||
spreading or slightly ascending, sessile, (17–)19–20(–24) ×
|
||
<quantity box="[354,447,1058,1085]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="2" pageNumber="3" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">6–8 mm</quantity>
|
||
, incurved, semi–lunate in profile, at first subcylindrical to elliptic, sub–fleshy, acute in both ends, and at maturity strongly compressed laterally or low convex on the lateral faces, somewhat plump (not inflated like a bladder), ventral suture concave, keeled, dorsal suture almost straight to low convex, base acute, apiculate and rigid at the apex, the beak
|
||
<quantity box="[432,546,1166,1193]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="2" pageNumber="3" unit="mm" value="0.85" valueMax="1.0" valueMin="0.7">0.7–1 mm</quantity>
|
||
long, with the sub–persistent style, the valves coriaceous, stramineous, brown to light–brown, glabrous, noticeably transversely reticulate and wrinkled, unilocular, septum absent; dehiscence apical; seeds
|
||
<quantity box="[204,339,1238,1265]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="2" pageNumber="3" unit="mm" value="2.05" valueMax="2.2" valueMin="1.9">1.9–2.2 mm</quantity>
|
||
long, mitten–shaped, black, opaque.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="2" pageNumber="3" type="distribution">
|
||
<paragraph blockId="2.[136,1452,158,2093]" pageId="2" pageNumber="3">
|
||
<emphasis bold="true" box="[189,524,1274,1300]" pageId="2" pageNumber="3">Distribution and habitat:—</emphasis>
|
||
As far as we know,
|
||
<taxonomicName authorityName="A. E. Estrada, Rebman, C. Gonzalez & Villarreal" authorityYear="2022" box="[761,1014,1274,1300]" class="Magnoliopsida" family="Fabaceae" genus="Astragalus" kingdom="Plantae" order="Fabales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="tijuanensis">
|
||
<emphasis box="[761,1014,1274,1300]" italics="true" pageId="2" pageNumber="3">Astragalus tijuanensis</emphasis>
|
||
</taxonomicName>
|
||
is only known from the
|
||
<typeStatus box="[1309,1357,1275,1301]" pageId="2" pageNumber="3">type</typeStatus>
|
||
locality on the south side of the metropolitan area of Tijuana,
|
||
<collectingRegion box="[760,935,1310,1336]" country="Mexico" name="Baja California" pageId="2" pageNumber="3">Baja California</collectingRegion>
|
||
to the east of Rosarito (
|
||
<figureCitation box="[1217,1288,1310,1336]" captionStart="FIGURE 3" captionStartId="5.[136,229,894,916]" captionTargetBox="[340,1247,190,870]" captionTargetId="figure-19@5.[340,1247,190,870]" captionTargetPageId="5" captionText="FIGURE 3. Map showing the distribution of Astragalus tijuanensis (white star) and A. brauntonii var. lativexillum (black dot) in Tijuana, Baja California, Mexico." figureDoi="http://doi.org/10.5281/zenodo.7517532" httpUri="https://zenodo.org/record/7517532/files/figure.png" pageId="2" pageNumber="3">Fig. 3</figureCitation>
|
||
). This species was found in a recently burned, disturbed, chaparral area on open gravelly or clayish hillsides, low slopes or flats, on clayish substrate associated with
|
||
<taxonomicName authority="Engl. (1883: 339)" authorityName="Engl." authorityPageNumber="339" authorityYear="1883" box="[568,988,1382,1408]" class="Magnoliopsida" family="Anacardiaceae" genus="Malosma" kingdom="Plantae" order="Sapindales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="laurina">
|
||
<emphasis box="[568,766,1382,1408]" italics="true" pageId="2" pageNumber="3">Malosma laurina</emphasis>
|
||
Engl. (1883: 339)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Nutt. (1842: 259)" authorityName="Nutt." authorityPageNumber="259" authorityYear="1842" box="[1006,1445,1382,1409]" class="Magnoliopsida" family="Ericaceae" genus="Xylococcus" kingdom="Plantae" order="Ericales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="bicolor">
|
||
<emphasis box="[1006,1227,1382,1408]" italics="true" pageId="2" pageNumber="3">Xylococcus bicolor</emphasis>
|
||
Nutt. (1842: 259)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Greene (1906: 208)" authorityName="Greene" authorityPageNumber="208" authorityYear="1906" box="[136,694,1418,1444]" class="Magnoliopsida" family="Malvaceae" genus="Malacothamnus" kingdom="Plantae" order="Malvales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="fasciculatus">
|
||
<emphasis box="[136,460,1418,1444]" italics="true" pageId="2" pageNumber="3">Malacothamnus fasciculatus</emphasis>
|
||
<bibRefCitation author="Greene, E. L." box="[469,694,1418,1444]" pageId="2" pageNumber="3" pagination="208" refId="ref7334" refString="Greene, E. L. (1906) Malacothamnus fasciculatus. Leaflets of Botanical Observation and Criticism 1: 208" type="journal article" year="1906">Greene (1906: 208)</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Hook. & Arn. (1832: 139)" authorityName="Hook. & Arn." authorityPageNumber="139" authorityYear="1832" box="[710,1313,1418,1444]" class="Magnoliopsida" family="Rosaceae" genus="Adenostoma" kingdom="Plantae" order="Rosales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="fasciculatum">
|
||
<emphasis box="[710,1001,1418,1444]" italics="true" pageId="2" pageNumber="3">Adenostoma fasciculatum</emphasis>
|
||
Hook. & Arn. (1832: 139)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Greene (1897: 424)" authorityName="Greene" authorityPageNumber="424" authorityYear="1897" class="Magnoliopsida" family="Asteraceae" genus="Deinandra" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="fasciculata">
|
||
<emphasis italics="true" pageId="2" pageNumber="3">Deinandra fasciculata</emphasis>
|
||
<bibRefCitation author="Greene, E. L." box="[265,483,1454,1480]" pageId="2" pageNumber="3" pagination="424" refId="ref7299" refString="Greene, E. L. (1897) Deinandra fasciculata Greene. In: Flora Franciscana: An Attempt to Classify and Describe the Vascular Plants of Middle California. pp. 424." type="book chapter" year="1897">Greene (1897: 424)</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="(D. D. Keck) B. G. Baldwin (1999: 468)" authorityName="B. G. Baldwin" authorityPageNumber="468" authorityYear="1999" baseAuthorityName="D. D. Keck" box="[496,1172,1454,1480]" class="Magnoliopsida" family="Asteraceae" genus="Deinandra" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="conjugens">
|
||
<emphasis box="[496,737,1454,1480]" italics="true" pageId="2" pageNumber="3">Deinandra conjugens</emphasis>
|
||
(D.D. Keck) B.G.
|
||
<bibRefCitation author="Baldwin, B. G." box="[933,1172,1454,1480]" pageId="2" pageNumber="3" pagination="468" refId="ref6780" refString="Baldwin, B. G. (1999) New combinations and new genera in the North American tarweeds (Compositae: Madiinae). Novon 9 (4): 468. https: // doi. org / 10.2307 / 3392143" type="journal article" year="1999">Baldwin (1999: 468)</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="(Lindl.) M. Roem. (1847: 105)" authorityName="M. Roem." authorityPageNumber="105" authorityYear="1847" baseAuthorityName="Lindl." class="Magnoliopsida" family="Rosaceae" genus="Heteromeles" kingdom="Plantae" order="Rosales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="arbutifolia">
|
||
<emphasis box="[1185,1452,1454,1480]" italics="true" pageId="2" pageNumber="3">Heteromeles arbutifolia</emphasis>
|
||
(Lindl.) M. Roem. (1847: 105)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Gentry (1946: 181)" authorityName="Gentry" authorityPageNumber="181" authorityYear="1946" box="[505,919,1490,1516]" class="Liliopsida" family="Asparagaceae" genus="Nolina" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="interrata">
|
||
<emphasis box="[505,690,1490,1516]" italics="true" pageId="2" pageNumber="3">Nolina interrata</emphasis>
|
||
<bibRefCitation author="Gentry, H. S." box="[698,919,1490,1516]" pageId="2" pageNumber="3" pagination="181" refId="ref7126" refString="Gentry, H. S. (1946) A new Nolina from southern California. Madrono 8: 181, fig. 1, tab. 19." type="journal article" year="1946">Gentry (1946: 181)</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="(DC.) Torr. & A. Gray (1842: 193)" authorityName="Torr. & A. Gray" authorityPageNumber="193" authorityYear="1842" baseAuthorityName="DC." class="Magnoliopsida" family="Asteraceae" genus="Gutierrezia" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="californica">
|
||
<emphasis box="[935,1213,1490,1517]" italics="true" pageId="2" pageNumber="3">Gutierrezia californica (</emphasis>
|
||
DC.) Torr. & A.Gray (1842: 193)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="S. Stokes (1910: 351)" authorityName="S. Stokes" authorityPageNumber="351" authorityYear="1910" box="[283,988,1526,1553]" class="Magnoliopsida" family="Polygonaceae" genus="Eriogonum" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="variety" species="fasciculatum" variety="foliolosum">
|
||
<emphasis box="[283,560,1526,1552]" italics="true" pageId="2" pageNumber="3">Eriogonum fasciculatum</emphasis>
|
||
var.
|
||
<emphasis box="[617,735,1526,1552]" italics="true" pageId="2" pageNumber="3">foliolosum</emphasis>
|
||
S.
|
||
<bibRefCitation author="Stokes, S." box="[773,988,1526,1553]" pageId="2" pageNumber="3" pagination="351" refId="ref7821" refString="Stokes, S. (1910) Eriogonum fasciculatum var. foliolosum. Bulletin of the New York Botanical Garden 6: 351." type="journal article" year="1910">Stokes (1910: 351)</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Nutt (1838: 267 - 268)" authorityName="Nutt" authorityPageNumber="267 - 268" authorityYear="1838" class="Magnoliopsida" family="Rhamnaceae" genus="Ceanothus" kingdom="Plantae" order="Rosales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="verrucosus">
|
||
<emphasis box="[1003,1254,1526,1552]" italics="true" pageId="2" pageNumber="3">Ceanothus verrucosus</emphasis>
|
||
Nutt (1838: 267- 268)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="S. Watson (1876: 126)" authorityName="S. Watson" authorityPageNumber="126" authorityYear="1876" box="[201,691,1562,1589]" class="Magnoliopsida" family="Rhamnaceae" genus="Adolphia" kingdom="Plantae" order="Rosales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="californica">
|
||
<emphasis box="[201,434,1562,1588]" italics="true" pageId="2" pageNumber="3">Adolphia californica</emphasis>
|
||
S.
|
||
<bibRefCitation author="Watson, S." box="[469,691,1562,1589]" pageId="2" pageNumber="3" pagination="126" refId="ref7872" refString="Watson, S. (1876) Description of new speciesof plants, chiefly Californian, with revisoon of certain genera. Proceedings of the American Academy of Arts and Sciences 11: 126. https: // doi. org / 10.2307 / 20021459" type="journal article" year="1876">Watson (1876: 126)</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName authority="Graham (1828: 2837)" authorityName="Graham" authorityPageNumber="2837" authorityYear="1828" box="[760,1206,1562,1588]" class="Magnoliopsida" family="Solanaceae" genus="Nicotiana" kingdom="Plantae" order="Solanales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="glauca">
|
||
<emphasis box="[760,953,1562,1588]" italics="true" pageId="2" pageNumber="3">Nicotiana glauca</emphasis>
|
||
<bibRefCitation author="Graham, R." box="[960,1206,1562,1588]" pageId="2" pageNumber="3" pagination="175" refId="ref7155" refString="Graham, R. (1828) Description of new and rare plants. The Edinburgh new philosophical journal 55: t. 175." type="journal article" year="1828">Graham (1828: 2837)</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="2" pageNumber="3" type="etymology">
|
||
<paragraph blockId="2.[136,1452,158,2093]" pageId="2" pageNumber="3">
|
||
<emphasis bold="true" box="[189,356,1598,1624]" pageId="2" pageNumber="3">Etymology:—</emphasis>
|
||
The name of this new species refers to the
|
||
<typeStatus box="[831,879,1599,1625]" pageId="2" pageNumber="3">type</typeStatus>
|
||
locality in the municipality of Tijuana, and to date, it is the only place known where this species grows in a small population.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="2" pageNumber="3" type="biology_ecology">
|
||
<paragraph blockId="2.[136,1452,158,2093]" pageId="2" pageNumber="3">
|
||
<emphasis bold="true" box="[189,351,1670,1696]" pageId="2" pageNumber="3">Phenology:—</emphasis>
|
||
According to the presence of reproductive structures on the
|
||
<typeStatus box="[1017,1065,1671,1697]" pageId="2" pageNumber="3">type</typeStatus>
|
||
specimen, flowering likely started in early May and was continuing in early June while fruiting probably started in early June and will likely continue until the end of July or early August.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="2" pageNumber="3" type="conservation">
|
||
<paragraph blockId="2.[136,1452,158,2093]" pageId="2" pageNumber="3">
|
||
<emphasis bold="true" box="[189,467,1778,1804]" pageId="2" pageNumber="3">Conservation status:—</emphasis>
|
||
This new species was discovered while surveying and documenting populations of a cross-border rare plant species called
|
||
<taxonomicName authorityName="B. G. Baldwin" authorityYear="1999" baseAuthorityName="D. D. Keck" box="[574,819,1814,1840]" class="Magnoliopsida" family="Asteraceae" genus="Deinandra" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="conjugens">
|
||
<emphasis box="[574,819,1814,1840]" italics="true" pageId="2" pageNumber="3">Deinandra conjugens</emphasis>
|
||
</taxonomicName>
|
||
as part of a binational collaborative project. This new taxon was only encountered in a single population at the
|
||
<typeStatus box="[796,844,1851,1877]" pageId="2" pageNumber="3">type</typeStatus>
|
||
locality. Even though we documented other regional populations of
|
||
<taxonomicName authorityName="B. G. Baldwin" authorityYear="1999" baseAuthorityName="D. D. Keck" box="[312,558,1886,1912]" class="Magnoliopsida" family="Asteraceae" genus="Deinandra" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="conjugens">
|
||
<emphasis box="[312,558,1886,1912]" italics="true" pageId="2" pageNumber="3">Deinandra conjugens</emphasis>
|
||
</taxonomicName>
|
||
in the area, this new
|
||
<taxonomicName box="[820,941,1886,1912]" class="Magnoliopsida" family="Fabaceae" genus="Astragalus" kingdom="Plantae" order="Fabales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="genus">
|
||
<emphasis box="[820,941,1886,1912]" italics="true" pageId="2" pageNumber="3">Astragalus</emphasis>
|
||
</taxonomicName>
|
||
was not found elsewhere. The only known population is found within the Tijuana metropolitan area, which is being strongly affected by human activity (personal observation). According to GeoCAT, the extent of its distribution and the number of occurrences, as recommended by the IUCN Red List guidelines (
|
||
<bibRefCitation author="IUCN" box="[522,660,1994,2020]" pageId="2" pageNumber="3" refId="ref7414" refString="IUCN. (2017) Guidelines for Using the IUCN Red List Categories and Criteria. Version 14. Prepared by the Standards and Petition subcommittee. Available from: http: // cmsdocs. s 3. amazonaws. com / RedListGuidelines. pdf (accessed June 2022)" type="book" year="2017">IUCN 2017</bibRefCitation>
|
||
),
|
||
<taxonomicName authorityName="A. E. Estrada, Rebman, C. Gonzalez & Villarreal" authorityYear="2022" box="[680,930,1994,2020]" class="Magnoliopsida" family="Fabaceae" genus="Astragalus" kingdom="Plantae" order="Fabales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="tijuanensis">
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<emphasis box="[680,930,1994,2020]" italics="true" pageId="2" pageNumber="3">Astragalus tijuanensis</emphasis>
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</taxonomicName>
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should be placed in the Critically Endangered (CR) category, criteria B2ab(iii). Under the current conditions of impact due to fires, the presence of livestock, and the development of human settlements in the area of distribution of this new species, it faces a high risk of extinction.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |