475 lines
67 KiB
XML
475 lines
67 KiB
XML
<document id="6429F976CE19DAED15F06561272D0B3B" ID-DOI="10.11646/zootaxa.3949.1.3" ID-GBIF-Dataset="c5c2caed-3746-4779-b176-8c56cea9c188" ID-ISSN="1175-5326" ID-Zenodo-Dep="233514" ID-ZooBank="694D37C6-F98D-49B5-BDC2-FCB07767BCBB" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461045164769" checkinUser="plazi" docAuthor="González, Fabio Arturo & Medina, Claudia Alejandra" docDate="2015" docId="A55717192B57944A6F80FC0CFC0DFD6D" docLanguage="en" docName="zt03949p090.pdf" docOrigin="Zootaxa 3949 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ontherus felicitae Gonzalez & Medina, new species" docType="treatment" docVersion="8" masterDocId="596E6F612B56944C6F17FFDBFFFCFFA0" masterDocTitle="The genus Ontherus Erichson 1847 (Coleoptera: Scarabaeidae: Scarabaeinae): description of a new species, and notes on the genus in Colombia" updateTime="1698576529699" updateUser="plazi">
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<mods:titleInfo id="13F89E0C230270460FDEEDC9561E31F1">
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<mods:title id="A547F3D2AA6DD7A21CBD2A6FFF1AE97E">The genus Ontherus Erichson 1847 (Coleoptera: Scarabaeidae: Scarabaeinae): description of a new species, and notes on the genus in Colombia</mods:title>
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<mods:namePart id="D506DC034CE87D6A9701B7D83540485B">González, Fabio Arturo</mods:namePart>
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<mods:roleTerm id="49697E6B9FC116CF6C9C2CD74DD93F3A">Author</mods:roleTerm>
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<mods:namePart id="B6B447B447EA654D2828DE8FE9AD8FBC">Medina, Claudia Alejandra</mods:namePart>
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<mods:typeOfResource id="7819EB871AEB7BCEAC68ECD8BECA0F55">text</mods:typeOfResource>
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<mods:title id="5D4A36A303E18065C1884F4040320350">Zootaxa</mods:title>
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<mods:part id="CC0D0C12909A1465AB1F4D9756BDB136">
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<mods:date id="9CF97D9F8958FCBD4B95065FA141503E">2015</mods:date>
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<mods:detail id="BE480D81BFB58390F90EA1B072079562" type="volume">
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<mods:number id="42CA2FC4709F36BF62281159388FFAC9">3949</mods:number>
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<mods:classification id="100902B8FD6040F288C990BD7344FC0C">journal volume</mods:classification>
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<mods:identifier id="9AF9A56AAF901734648305661C51AC55" type="DOI">10.11646/zootaxa.3949.1.3</mods:identifier>
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<mods:identifier id="D31FEB6395E25CA33ADBBB9B28095CD5" type="ISSN">1175-5326</mods:identifier>
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<treatment id="A55717192B57944A6F80FC0CFC0DFD6D" ID-DOI="http://doi.org/10.5281/zenodo.5663997" ID-GBIF-Taxon="119610393" ID-Zenodo-Dep="5663997" LSID="urn:lsid:plazi:treatment:A55717192B57944A6F80FC0CFC0DFD6D" httpUri="http://treatment.plazi.org/id/A55717192B57944A6F80FC0CFC0DFD6D" lastPageId="6" pageId="1">
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<subSubSection id="65E4F5842B57944D6F80FC0CFDF2FBB3" pageId="1" type="nomenclature">
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<paragraph id="2D41A60F2B57944D6F80FC0CFCF5FC52" blockId="1.[151,777,983,1043]" box="[151,777,983,1010]" pageId="1">
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<heading id="760911632B57944D6F80FC0CFCF5FC52" bold="true" box="[151,777,983,1010]" fontSize="11" level="1" pageId="1" reason="1">
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<emphasis id="1F8A7A1D2B57944D6F80FC0CFCF5FC52" bold="true" box="[151,777,983,1010]" pageId="1">
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<taxonomicName id="EAFEDD8C2B57944D6F80FC0CFD8EFC52" ID-CoL="7ZQ3V" authority="Gonzalez & Medina" authorityName="Gonzalez & Medina" box="[151,626,983,1010]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="felicitae" status="sp. nov.">
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<emphasis id="1F8A7A1D2B57944D6F80FC0CFE93FC51" bold="true" box="[151,367,983,1009]" italics="true" pageId="1">Ontherus felicitae</emphasis>
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González & Medina
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</taxonomicName>
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<taxonomicNameLabel id="04B9C7662B57944D6D6EFC03FCF5FC52" box="[633,777,984,1010]" pageId="1" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="2D41A60F2B57944D6F80FC21FDF2FBB3" blockId="1.[151,777,983,1043]" box="[151,526,1018,1043]" pageId="1">
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(
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<figureCitation id="B5C5BA8A2B57944D6F89FC21FEDAFBB3" box="[158,294,1018,1043]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1863,1886]" captionTargetBox="[316,1265,214,1828]" captionTargetId="figure@2.[304,1284,193,1842]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Figures 1 – 3 Ontherus felicitae n. sp. Figure 1, dorsal view (holotype); Figure 2, frontal view, head and pronotum (holotype); Figure 3, elytral sculpture (holotype). Figures 4 – 6 male of Ontherus compressicornis Luederwaldt 1931. Figure 4, dorsal view; Figure 5, frontal view head and pronotum; Figure 6, elytral sculpture." httpUri="https://zenodo.org/record/233515/files/figure.png" pageId="1">Figures 1–3</figureCitation>
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,
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<figureCitation id="B5C5BA8A2B57944D6E24FC20FEA3FBB3" box="[307,351,1019,1043]" captionStart="FIGURES 7 – 9" captionStartId="3.[151,264,881,904]" captionTargetBox="[265,1325,184,863]" captionTargetId="figure@3.[252,1334,173,876]" captionTargetPageId="3" captionText="FIGURES 7 – 9. Figures 7 – 8 Ontherus felicitae n. sp. Figure 7, female dorsal view (paratype); Figure 8, 8 th elytral stria, female (paratype). Figure 9, Ontherus compressicornis Luederwaldt 1931, 8 th elytral stria, female. Black arrow pseudoepipleuron, white arrow 8 th elytral stria." httpUri="https://zenodo.org/record/233516/files/figure.png" pageId="1">7–8</figureCitation>
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,
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<figureCitation id="B5C5BA8A2B57944D6E7CFC20FE48FBB3" box="[363,436,1019,1043]" captionStart="FIGURES 10 – 18" captionStartId="4.[151,264,1915,1938]" captionTargetBox="[241,1344,208,1881]" captionTargetId="figure@4.[229,1358,193,1894]" captionTargetPageId="4" captionText="FIGURES 10 – 18. Figures 10 – 14 male genitalia Ontherus felicitae n. sp. Figure 10, lateral view aedeagus; Figure 11, basal sclerite; Figure 12, plate-shape sclerite; Figure 13, accessory sclerite; Figure 14, elongate sclerite. Figures 15 – 18 male genitalia Ontherus compressicornis Luederwaldt 1931. Figure 15, lateral view aedeagus; Figure 16, basal sclerite; Figure 17, plate-shape sclerite; Figure 18, elongate sclerite." httpUri="https://zenodo.org/record/233517/files/figure.png" pageId="1">10–14</figureCitation>
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,
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<figureCitation id="B5C5BA8A2B57944D6EA8FC20FDFAFBB3" box="[447,518,1019,1043]" captionStart="FIGURES 19 – 24" captionStartId="5.[151,264,1274,1297]" captionTargetBox="[192,1393,203,1243]" captionTargetId="figure@5.[180,1406,193,1253]" captionTargetPageId="5" captionText="FIGURES 19 – 24. Cephalic horn size variation and plate-shape sclerite in Ontherus felicitae n. sp. Figures 19 – 20 welldeveloped male (holotype). Figure 19, lateral view head and pronotum; Figure 20, plate-shape sclerite. Figures 21 – 22 moderately developed male (paratype). Figure 21, lateral view, head and pronotum; Figure 22, plate-shape sclerite. Figures 23 – 24 underdeveloped male (paratype). Figure 23, lateral view, head and pronotum; Figure 24, plate-shape sclerite." httpUri="https://zenodo.org/record/233518/files/figure.png" pageId="1">19–24</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="65E4F5842B57944D6F80FBE4FA88F99E" pageId="1" type="materials_examined">
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<paragraph id="2D41A60F2B57944D6F80FBE4FDCCFB60" blockId="1.[151,1437,1087,2014]" pageId="1">
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<typeStatus id="F24518AD2B57944D6F80FBE4FEFFFBF8" box="[151,259,1087,1112]" pageId="1" type="holotype">Holotype</typeStatus>
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. ♂,
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<emphasis id="1F8A7A1D2B57944D6E51FBE4FD7DFBF8" bold="true" box="[326,641,1087,1112]" pageId="1">
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<collectingCountry id="55E9E69F2B57944D6E51FBE4FE15FBF8" box="[326,489,1087,1112]" name="Colombia" pageId="1">COLOMBIA</collectingCountry>
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: Antioquia:
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</emphasis>
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Jardín, Reserva Natural Mesenia, Secondary Forest,
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,
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,
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,
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<date id="594080CF2B57944D6E84FBB9FE18FBDB" box="[403,484,1122,1147]" pageId="1" value="2012-02">ii.2012</date>
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, Edwards Felicity, T.Exc.H. #1 (IAvH-E-147505)/ [handwritten] Mesenia, 2012,
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<collectingCountry id="55E9E69F2B57944D6F80FB5EFEF4FB3E" box="[151,264,1157,1182]" name="Colombia" pageId="1">Colombia</collectingCountry>
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, Forest. Near, Trap #1/ [handwritten, yellow label] Mesenia, # 126/ [red label]
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<taxonomicName id="EAFEDD8C2B57944D6BDBFB5EFA61FB3D" box="[1228,1437,1157,1181]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="felicitae">
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<emphasis id="1F8A7A1D2B57944D6BDBFB5EFA61FB3D" box="[1228,1437,1157,1181]" italics="true" pageId="1">Ontherus felicitae</emphasis>
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</taxonomicName>
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<typeStatus id="F24518AD2B57944D6F80FB7CFF03FB60" box="[151,255,1191,1216]" pageId="1" type="holotype">Holotype</typeStatus>
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<bibRefCitation id="496FDBFE2B57944D6E11FB7CFDD0FB60" author="Gonzalez" box="[262,556,1191,1216]" pageId="1" refString="Medina, C. A. & Gonzalez, A. (2014) Escarabajos coprofagos de la subfamilia Scarabaeinae. In: Pizano, C. & Garcia, H. (Eds.), El bosque seco tropical en Colombia. Instituto de Investigacion de Recursos Biologicos Alexander von Humboldt (IAvH) Bogota, pp. 195 - 213." type="book chapter" year="2014">González & Medina 2014</bibRefCitation>
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.
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</paragraph>
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<typeStatus id="F24518AD2B57944D6FD0FB10FEC4FB43" box="[199,312,1227,1251]" pageId="1" type="paratype">Paratypes</typeStatus>
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.
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<collectingCountry id="55E9E69F2B57944D6E50FB11FE16FB43" box="[327,490,1226,1251]" name="Colombia" pageId="1">COLOMBIA</collectingCountry>
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: Antioquia:
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</emphasis>
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Jardín, PNN Tatamá, Reserva Natural Mesenia, Forest,
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,
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, Edwards Felicity, T.Exc.H. #1 (IAvH-E-147509), Jardín, PNN Tatamá, Reserva Natural Mesenia, Forest,
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, 1♀,
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, Edwards Felicity, T.Exc.H. #2 (IAvH-E- 147511), Jardín, Reserva Natural Mesenia, Forest,
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,
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<specimenCount id="3BF86D862B57944D6B5EFAE8FB91FAEB" box="[1097,1133,1331,1355]" pageId="1" type="generic">1♂</specimenCount>
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,
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,
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, 1♀,
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<date id="594080CF2B57944D6F80FAACFF15FA30" box="[151,233,1399,1424]" pageId="1" value="2012-02">ii.2012</date>
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, Edwards Felicity, T.Exc.H. #2a (IAvH-E-147513), Jardín, Reserva Natural Mesenia, Forest,
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<geoCoordinate id="48CAC0C82B57944D6A37FAACFA6BFA30" box="[1312,1431,1399,1424]" direction="north" orientation="latitude" pageId="1" precision="15" value="5.4772224">5°28'38''N</geoCoordinate>
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,
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,
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,
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<date id="594080CF2B57944D6ED9FA41FDE3FA13" box="[462,543,1434,1459]" pageId="1" value="2012-02">ii.2012</date>
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, Edwards Felicity, T.Exc.H. #3 (IAvH-E-147510), Jardín, Reserva Natural Mesenia, Secondary Forest,
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<geoCoordinate id="48CAC0C82B57944D6EC1FA66FDB0FA76" box="[470,588,1469,1494]" direction="north" orientation="latitude" pageId="1" precision="15" value="5.488333">5°29'18''N</geoCoordinate>
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,
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<geoCoordinate id="48CAC0C82B57944D6D4EFA66FD1DFA76" box="[601,737,1469,1494]" direction="west" orientation="longitude" pageId="1" precision="15" value="-75.894165">75°53'39''W</geoCoordinate>
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,
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<quantity id="EA060BEA2B57944D6DFAFA66FCC2FA76" box="[749,830,1469,1494]" metricMagnitude="3" metricUnit="m" metricValue="2.357" pageId="1" unit="m" value="2357.0">2357m</quantity>
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,
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<specimenCount id="3BF86D862B57944D6C5EFA65FC93FA76" box="[841,879,1470,1494]" pageId="1" type="generic">2♂</specimenCount>
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,
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<date id="594080CF2B57944D6C6CFA66FC30FA76" box="[891,972,1469,1494]" pageId="1" value="2012-02">ii.2012</date>
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, Edwards Felicity, T.Exc.H. #2 (IAvH-E- 147514, IAvH-E-147515), Jardín, Reserva Natural Mesenia, Secondary Forest,
|
||
<geoCoordinate id="48CAC0C82B57944D6B3DFA04FB5DFA58" box="[1066,1185,1503,1528]" direction="north" orientation="latitude" pageId="1" precision="15" value="5.4980555">5°29'53''N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="48CAC0C82B57944D6BA7FA04FAC5FA58" box="[1200,1337,1503,1528]" direction="west" orientation="longitude" pageId="1" precision="15" value="-75.88306">75°52'59''W</geoCoordinate>
|
||
,
|
||
<quantity id="EA060BEA2B57944D6A50FA3BFA64FA58" box="[1351,1432,1504,1528]" metricMagnitude="3" metricUnit="m" metricValue="2.232" pageId="1" unit="m" value="2232.0">2232m</quantity>
|
||
,
|
||
<specimenCount id="3BF86D862B57944D6F80F9D8FF47F9BB" box="[151,187,1539,1563]" pageId="1" type="generic">2♂</specimenCount>
|
||
,
|
||
<date id="594080CF2B57944D6FDBF9D9FEDFF9BB" box="[204,291,1538,1563]" pageId="1" value="2012-06">vi.2012</date>
|
||
, Edwards Felicity, T.Exc.H. (IAvH-E-147506, IAvH-E-147508), Jardín, Reserva Natural Mesenia, Secondary Forest,
|
||
<geoCoordinate id="48CAC0C82B57944D6E7FF9FEFE23F99E" box="[360,479,1573,1598]" direction="north" orientation="latitude" pageId="1" precision="15" value="5.4905553">5°29'26''N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="48CAC0C82B57944D6EFCF9FEFD8FF99E" box="[491,627,1573,1598]" direction="west" orientation="longitude" pageId="1" precision="15" value="-75.8875">75°53'15''W</geoCoordinate>
|
||
,
|
||
<quantity id="EA060BEA2B57944D6D68F9FDFD2CF99E" box="[639,720,1574,1598]" metricMagnitude="3" metricUnit="m" metricValue="2.334" pageId="1" unit="m" value="2334.0">2334m</quantity>
|
||
,
|
||
<specimenCount id="3BF86D862B57944D6DCDF9FDFD03F99E" box="[730,767,1574,1598]" pageId="1" type="generic">1♂</specimenCount>
|
||
,
|
||
<date id="594080CF2B57944D6C1BF9FEFC9FF99E" box="[780,867,1573,1598]" pageId="1" value="2012-06">vi.2012</date>
|
||
, Edwards Felicity, T.Exc.H. (IAvH-E-147512).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="65E4F5842B57944F6FD0F99CFAE6FA85" lastPageId="3" pageId="1" type="diagnosis">
|
||
<paragraph id="2D41A60F2B57944D6FD0F99CFF14F87E" blockId="1.[151,1437,1087,2014]" pageId="1">
|
||
<emphasis id="1F8A7A1D2B57944D6FD0F99CFEBDF9C0" bold="true" box="[199,321,1607,1632]" pageId="1">Diagnosis.</emphasis>
|
||
the species belongs to
|
||
<emphasis id="1F8A7A1D2B57944D6D5DF993FD3FF9FF" box="[586,707,1608,1631]" italics="true" pageId="1">Mexicanus</emphasis>
|
||
species group as defined by
|
||
<bibRefCitation id="496FDBFE2B57944D6B18F99CFB4CF9C0" author="Genier" box="[1039,1200,1607,1632]" pageId="1" refString="Genier, F. (1996) A revision of the Neotropical genus Ontherus Erichson (Coleoptera: Scarabaeidae, Scarabaeinae). Memoirs of the Entomological Society of Canada, 170, 1 - 169. http: // dx. doi. org / 10.4039 / entm 128170 fv" type="journal article" year="1996">Génier (1996)</bibRefCitation>
|
||
, since
|
||
<taxonomicName id="EAFEDD8C2B57944D6BE8F993FA81F9C0" box="[1279,1405,1608,1632]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="felicitae" status="sp. nov.">
|
||
<emphasis id="1F8A7A1D2B57944D6BE8F993FAEFF9FF" box="[1279,1299,1608,1631]" italics="true" pageId="1">O</emphasis>
|
||
.
|
||
<emphasis id="1F8A7A1D2B57944D6A34F993FA81F9C0" box="[1315,1405,1608,1632]" italics="true" pageId="1">felicitae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="04B9C7662B57944D6A92F99CFF44F923" pageId="1" rank="species">
|
||
<emphasis id="1F8A7A1D2B57944D6A92F99CFF44F923" bold="true" pageId="1">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
has the anterior suture of median lobe of metasternum straight; pronotum with carina (
|
||
<figureCitation id="B5C5BA8A2B57944D6B8FF9B1FACCF923" box="[1176,1328,1642,1667]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1863,1886]" captionTargetBox="[316,1265,214,1828]" captionTargetId="figure@2.[304,1284,193,1842]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Figures 1 – 3 Ontherus felicitae n. sp. Figure 1, dorsal view (holotype); Figure 2, frontal view, head and pronotum (holotype); Figure 3, elytral sculpture (holotype). Figures 4 – 6 male of Ontherus compressicornis Luederwaldt 1931. Figure 4, dorsal view; Figure 5, frontal view head and pronotum; Figure 6, elytral sculpture." httpUri="https://zenodo.org/record/233515/files/figure.png" pageId="1">Figs. 1 and 2</figureCitation>
|
||
); head of large males with horn compressed laterally; parameres apically expanded (
|
||
<figureCitation id="B5C5BA8A2B57944D6CC1F956FBD4F906" box="[982,1064,1677,1702]" captionStart="FIGURES 10 – 18" captionStartId="4.[151,264,1915,1938]" captionTargetBox="[241,1344,208,1881]" captionTargetId="figure@4.[229,1358,193,1894]" captionTargetPageId="4" captionText="FIGURES 10 – 18. Figures 10 – 14 male genitalia Ontherus felicitae n. sp. Figure 10, lateral view aedeagus; Figure 11, basal sclerite; Figure 12, plate-shape sclerite; Figure 13, accessory sclerite; Figure 14, elongate sclerite. Figures 15 – 18 male genitalia Ontherus compressicornis Luederwaldt 1931. Figure 15, lateral view aedeagus; Figure 16, basal sclerite; Figure 17, plate-shape sclerite; Figure 18, elongate sclerite." httpUri="https://zenodo.org/record/233517/files/figure.png" pageId="1">Fig. 10</figureCitation>
|
||
), and females with 2 denticles on the head (
|
||
<figureCitation id="B5C5BA8A2B57944D6E13F974FEB4F968" box="[260,328,1711,1736]" captionStart="FIGURES 7 – 9" captionStartId="3.[151,264,881,904]" captionTargetBox="[265,1325,184,863]" captionTargetId="figure@3.[252,1334,173,876]" captionTargetPageId="3" captionText="FIGURES 7 – 9. Figures 7 – 8 Ontherus felicitae n. sp. Figure 7, female dorsal view (paratype); Figure 8, 8 th elytral stria, female (paratype). Figure 9, Ontherus compressicornis Luederwaldt 1931, 8 th elytral stria, female. Black arrow pseudoepipleuron, white arrow 8 th elytral stria." httpUri="https://zenodo.org/record/233516/files/figure.png" pageId="1">Fig. 7</figureCitation>
|
||
). The new species is closely related to
|
||
<taxonomicName id="EAFEDD8C2B57944D6DECF96BFB5AF968" authority="Luederwaldt 1931" authorityName="Luederwaldt" authorityYear="1931" box="[763,1190,1711,1736]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="compressicornis">
|
||
<emphasis id="1F8A7A1D2B57944D6DECF96BFCF3F967" box="[763,783,1712,1735]" italics="true" pageId="1">O</emphasis>
|
||
.
|
||
<emphasis id="1F8A7A1D2B57944D6C0AF96BFC28F967" box="[797,980,1712,1735]" italics="true" pageId="1">compressicornis</emphasis>
|
||
Luederwaldt 1931
|
||
</taxonomicName>
|
||
due to: 1. large males with cephalic horn compressed laterally; 2. large males with pronotal carina with inner portion lower than lateral portion, without tubercles (
|
||
<figureCitation id="B5C5BA8A2B57944D6ED0F92EFDA6F8AE" box="[455,602,1781,1806]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1863,1886]" captionTargetBox="[316,1265,214,1828]" captionTargetId="figure@2.[304,1284,193,1842]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Figures 1 – 3 Ontherus felicitae n. sp. Figure 1, dorsal view (holotype); Figure 2, frontal view, head and pronotum (holotype); Figure 3, elytral sculpture (holotype). Figures 4 – 6 male of Ontherus compressicornis Luederwaldt 1931. Figure 4, dorsal view; Figure 5, frontal view head and pronotum; Figure 6, elytral sculpture." httpUri="https://zenodo.org/record/233515/files/figure.png" pageId="1">Figs. 2 and 5</figureCitation>
|
||
) and 3. strial punctures, simples (not ocellate), and larger, punctures more than twice as wide as connecting stria (
|
||
<figureCitation id="B5C5BA8A2B57944D6D42F8CCFD10F890" box="[597,748,1815,1840]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1863,1886]" captionTargetBox="[316,1265,214,1828]" captionTargetId="figure@2.[304,1284,193,1842]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Figures 1 – 3 Ontherus felicitae n. sp. Figure 1, dorsal view (holotype); Figure 2, frontal view, head and pronotum (holotype); Figure 3, elytral sculpture (holotype). Figures 4 – 6 male of Ontherus compressicornis Luederwaldt 1931. Figure 4, dorsal view; Figure 5, frontal view head and pronotum; Figure 6, elytral sculpture." httpUri="https://zenodo.org/record/233515/files/figure.png" pageId="1">Figs. 3 and 6</figureCitation>
|
||
). But
|
||
<taxonomicName id="EAFEDD8C2B57944D6C24F8C3FC4DF890" box="[819,945,1816,1840]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="felicitae" status="sp. nov.">
|
||
<emphasis id="1F8A7A1D2B57944D6C24F8C3FCBBF88F" box="[819,839,1816,1839]" italics="true" pageId="1">O</emphasis>
|
||
.
|
||
<emphasis id="1F8A7A1D2B57944D6C40F8C3FC4DF890" box="[855,945,1816,1840]" italics="true" pageId="1">felicitae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1F8A7A1D2B57944D6CAEF8CCFC05F890" bold="true" box="[953,1017,1815,1840]" pageId="1">
|
||
<taxonomicNameLabel id="04B9C7662B57944D6CAEF8CCFC05F890" box="[953,1017,1815,1840]" pageId="1" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be separated by: 1. the minutely punctate pronotal disc (
|
||
<figureCitation id="B5C5BA8A2B57944D6EB7F8E1FDCBF8F3" box="[416,567,1850,1875]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1863,1886]" captionTargetBox="[316,1265,214,1828]" captionTargetId="figure@2.[304,1284,193,1842]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Figures 1 – 3 Ontherus felicitae n. sp. Figure 1, dorsal view (holotype); Figure 2, frontal view, head and pronotum (holotype); Figure 3, elytral sculpture (holotype). Figures 4 – 6 male of Ontherus compressicornis Luederwaldt 1931. Figure 4, dorsal view; Figure 5, frontal view head and pronotum; Figure 6, elytral sculpture." httpUri="https://zenodo.org/record/233515/files/figure.png" pageId="1">Figs. 1 and 2</figureCitation>
|
||
), punctures not visible at
|
||
<date id="594080CF2B57944D6C4BF8E0FC87F8F3" box="[860,891,1851,1875]" pageId="1">8x</date>
|
||
; 2. elytral striae punctures almost three times as wide as connecting stria (
|
||
<figureCitation id="B5C5BA8A2B57944D6ECFF886FDDAF8D6" box="[472,550,1885,1910]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1863,1886]" captionTargetBox="[316,1265,214,1828]" captionTargetId="figure@2.[304,1284,193,1842]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Figures 1 – 3 Ontherus felicitae n. sp. Figure 1, dorsal view (holotype); Figure 2, frontal view, head and pronotum (holotype); Figure 3, elytral sculpture (holotype). Figures 4 – 6 male of Ontherus compressicornis Luederwaldt 1931. Figure 4, dorsal view; Figure 5, frontal view head and pronotum; Figure 6, elytral sculpture." httpUri="https://zenodo.org/record/233515/files/figure.png" pageId="1">Fig. 3</figureCitation>
|
||
); 3. elytral striae deeper apically; 4. eighth elytral stria with punctures longitudinally elongated and irregularly shaped (not rounded) (
|
||
<figureCitation id="B5C5BA8A2B57944D6C4BF8A4FC5DF838" box="[860,929,1919,1944]" captionStart="FIGURES 7 – 9" captionStartId="3.[151,264,881,904]" captionTargetBox="[265,1325,184,863]" captionTargetId="figure@3.[252,1334,173,876]" captionTargetPageId="3" captionText="FIGURES 7 – 9. Figures 7 – 8 Ontherus felicitae n. sp. Figure 7, female dorsal view (paratype); Figure 8, 8 th elytral stria, female (paratype). Figure 9, Ontherus compressicornis Luederwaldt 1931, 8 th elytral stria, female. Black arrow pseudoepipleuron, white arrow 8 th elytral stria." httpUri="https://zenodo.org/record/233516/files/figure.png" pageId="1">Fig. 8</figureCitation>
|
||
); 5. plate-shape sclerite of the internal sac of the aedeagus elongated, straight and slightly bent (
|
||
<figureCitation id="B5C5BA8A2B57944D6DCBF879FCCEF81B" box="[732,818,1954,1979]" captionStart="FIGURES 10 – 18" captionStartId="4.[151,264,1915,1938]" captionTargetBox="[241,1344,208,1881]" captionTargetId="figure@4.[229,1358,193,1894]" captionTargetPageId="4" captionText="FIGURES 10 – 18. Figures 10 – 14 male genitalia Ontherus felicitae n. sp. Figure 10, lateral view aedeagus; Figure 11, basal sclerite; Figure 12, plate-shape sclerite; Figure 13, accessory sclerite; Figure 14, elongate sclerite. Figures 15 – 18 male genitalia Ontherus compressicornis Luederwaldt 1931. Figure 15, lateral view aedeagus; Figure 16, basal sclerite; Figure 17, plate-shape sclerite; Figure 18, elongate sclerite." httpUri="https://zenodo.org/record/233517/files/figure.png" pageId="1">Fig. 12</figureCitation>
|
||
); 6. internal sac of the aedeagus with small accessory sclerite
|
||
</paragraph>
|
||
<caption id="7981F6872B54944E6F80F89CFBC0F83A" httpUri="https://zenodo.org/record/233515/files/figure.png" pageId="2" targetBox="[316,1265,214,1828]" targetPageId="2">
|
||
<paragraph id="2D41A60F2B54944E6F80F89CFBC0F83A" blockId="2.[151,1436,1863,1946]" pageId="2">
|
||
<emphasis id="1F8A7A1D2B54944E6F80F89CFEC2F8FC" bold="true" box="[151,318,1863,1886]" pageId="2">FIGURES 1–6.</emphasis>
|
||
Figures 1–3
|
||
<taxonomicName id="EAFEDD8C2B54944E6ED8F89CFD76F8FD" box="[463,650,1863,1885]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="felicitae" status="sp. nov.">
|
||
<emphasis id="1F8A7A1D2B54944E6ED8F89CFD76F8FD" box="[463,650,1863,1885]" italics="true" pageId="2">Ontherus felicitae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1F8A7A1D2B54944E6D83F893FD2DF8FD" bold="true" box="[660,721,1864,1885]" pageId="2">
|
||
<taxonomicNameLabel id="04B9C7662B54944E6D83F893FD2DF8FD" box="[660,721,1864,1885]" pageId="2" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Figure 1, dorsal view (holotype); Figure 2, frontal view, head and pronotum (holotype); Figure 3, elytral sculpture (holotype). Figures 4–6 male of
|
||
<taxonomicName id="EAFEDD8C2B54944E6CDBF8BDFA65F8DC" authority="Luederwaldt 1931" authorityName="Luederwaldt" authorityYear="1931" box="[972,1433,1894,1916]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="compressicornis">
|
||
<emphasis id="1F8A7A1D2B54944E6CDBF8BDFB2AF8DC" box="[972,1238,1894,1916]" italics="true" pageId="2">Ontherus compressicornis</emphasis>
|
||
Luederwaldt 1931
|
||
</taxonomicName>
|
||
. Figure 4, dorsal view; Figure 5, frontal view head and pronotum; Figure 6, elytral sculpture.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="7981F6872B55944F6F80FCAAFE4BFC65" httpUri="https://zenodo.org/record/233516/files/figure.png" pageId="3" targetBox="[265,1325,184,863]" targetPageId="3">
|
||
<paragraph id="2D41A60F2B55944F6F80FCAAFE4BFC65" blockId="3.[151,1436,881,965]" pageId="3">
|
||
<emphasis id="1F8A7A1D2B55944F6F80FCAAFEC6FC27" bold="true" box="[151,314,881,904]" pageId="3">FIGURES 7–9.</emphasis>
|
||
Figures 7–8
|
||
<taxonomicName id="EAFEDD8C2B55944F6EAAFCA9FD8FFC28" box="[445,627,882,904]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="felicitae" status="sp. nov.">
|
||
<emphasis id="1F8A7A1D2B55944F6EAAFCA9FD8FFC28" box="[445,627,882,904]" italics="true" pageId="3">Ontherus felicitae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1F8A7A1D2B55944F6D6FFCA9FD4DFC27" bold="true" box="[632,689,882,903]" pageId="3">
|
||
<taxonomicNameLabel id="04B9C7662B55944F6D6FFCA9FD4DFC27" box="[632,689,882,903]" pageId="3" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Figure 7, female dorsal view (paratype); Figure 8, 8th elytral stria, female (paratype). Figure 9,
|
||
<taxonomicName id="EAFEDD8C2B55944F6E61FC4BFCB5FC06" authority="Luederwaldt 1931" authorityName="Luederwaldt" authorityYear="1931" box="[374,841,912,934]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="compressicornis">
|
||
<emphasis id="1F8A7A1D2B55944F6E61FC4BFD7EFC06" box="[374,642,912,934]" italics="true" pageId="3">Ontherus compressicornis</emphasis>
|
||
Luederwaldt 1931
|
||
</taxonomicName>
|
||
, 8th elytral stria, female. Black arrow pseudoepipleuron, white arrow 8th elytral stria.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="2D41A60F2B55944F6F80FC37FF06FB35" blockId="3.[151,1437,1004,2037]" pageId="3">
|
||
between the elongate and plate-shape sclerites (
|
||
<figureCitation id="B5C5BA8A2B55944F6DC3FC37FCD2FBA5" box="[724,814,1004,1029]" captionStart="FIGURES 10 – 18" captionStartId="4.[151,264,1915,1938]" captionTargetBox="[241,1344,208,1881]" captionTargetId="figure@4.[229,1358,193,1894]" captionTargetPageId="4" captionText="FIGURES 10 – 18. Figures 10 – 14 male genitalia Ontherus felicitae n. sp. Figure 10, lateral view aedeagus; Figure 11, basal sclerite; Figure 12, plate-shape sclerite; Figure 13, accessory sclerite; Figure 14, elongate sclerite. Figures 15 – 18 male genitalia Ontherus compressicornis Luederwaldt 1931. Figure 15, lateral view aedeagus; Figure 16, basal sclerite; Figure 17, plate-shape sclerite; Figure 18, elongate sclerite." httpUri="https://zenodo.org/record/233517/files/figure.png" pageId="3">Fig. 13</figureCitation>
|
||
). In
|
||
<taxonomicName id="EAFEDD8C2B55944F6C7AFC36FBB0FBA4" box="[877,1100,1005,1028]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="compressicornis">
|
||
<emphasis id="1F8A7A1D2B55944F6C7AFC36FC7DFBA4" box="[877,897,1005,1028]" italics="true" pageId="3">O</emphasis>
|
||
.
|
||
<emphasis id="1F8A7A1D2B55944F6C81FC36FBB0FBA4" box="[918,1100,1005,1028]" italics="true" pageId="3">compressicornis</emphasis>
|
||
</taxonomicName>
|
||
: 1. the pronotal disc larger punctures visible at
|
||
<date id="594080CF2B55944F6E60FBCAFE6FFB89" box="[375,403,1041,1065]" pageId="3">8x</date>
|
||
(
|
||
<figureCitation id="B5C5BA8A2B55944F6EB5FBCBFDCAFB89" box="[418,566,1040,1065]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1863,1886]" captionTargetBox="[316,1265,214,1828]" captionTargetId="figure@2.[304,1284,193,1842]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Figures 1 – 3 Ontherus felicitae n. sp. Figure 1, dorsal view (holotype); Figure 2, frontal view, head and pronotum (holotype); Figure 3, elytral sculpture (holotype). Figures 4 – 6 male of Ontherus compressicornis Luederwaldt 1931. Figure 4, dorsal view; Figure 5, frontal view head and pronotum; Figure 6, elytral sculpture." httpUri="https://zenodo.org/record/233515/files/figure.png" pageId="3">Figs. 4 and 5</figureCitation>
|
||
); 2. elytral striae punctures almost five times as wide as connecting stria (
|
||
<figureCitation id="B5C5BA8A2B55944F6A78FBCBFF5BFBED" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1863,1886]" captionTargetBox="[316,1265,214,1828]" captionTargetId="figure@2.[304,1284,193,1842]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Figures 1 – 3 Ontherus felicitae n. sp. Figure 1, dorsal view (holotype); Figure 2, frontal view, head and pronotum (holotype); Figure 3, elytral sculpture (holotype). Figures 4 – 6 male of Ontherus compressicornis Luederwaldt 1931. Figure 4, dorsal view; Figure 5, frontal view head and pronotum; Figure 6, elytral sculpture." httpUri="https://zenodo.org/record/233515/files/figure.png" pageId="3">Fig. 6</figureCitation>
|
||
); 3. elytral striae slightly deeper apically; 4. eighth elytral stria with punctures rounded (
|
||
<figureCitation id="B5C5BA8A2B55944F6BB7FBEFFB1BFBED" box="[1184,1255,1076,1101]" captionStart="FIGURES 7 – 9" captionStartId="3.[151,264,881,904]" captionTargetBox="[265,1325,184,863]" captionTargetId="figure@3.[252,1334,173,876]" captionTargetPageId="3" captionText="FIGURES 7 – 9. Figures 7 – 8 Ontherus felicitae n. sp. Figure 7, female dorsal view (paratype); Figure 8, 8 th elytral stria, female (paratype). Figure 9, Ontherus compressicornis Luederwaldt 1931, 8 th elytral stria, female. Black arrow pseudoepipleuron, white arrow 8 th elytral stria." httpUri="https://zenodo.org/record/233516/files/figure.png" pageId="3">Fig. 9</figureCitation>
|
||
); 5. plate-shape sclerite of the internal sac of the aedeagus sinuated (
|
||
<figureCitation id="B5C5BA8A2B55944F6DEEFB83FCB3FBD1" box="[761,847,1112,1137]" captionStart="FIGURES 10 – 18" captionStartId="4.[151,264,1915,1938]" captionTargetBox="[241,1344,208,1881]" captionTargetId="figure@4.[229,1358,193,1894]" captionTargetPageId="4" captionText="FIGURES 10 – 18. Figures 10 – 14 male genitalia Ontherus felicitae n. sp. Figure 10, lateral view aedeagus; Figure 11, basal sclerite; Figure 12, plate-shape sclerite; Figure 13, accessory sclerite; Figure 14, elongate sclerite. Figures 15 – 18 male genitalia Ontherus compressicornis Luederwaldt 1931. Figure 15, lateral view aedeagus; Figure 16, basal sclerite; Figure 17, plate-shape sclerite; Figure 18, elongate sclerite." httpUri="https://zenodo.org/record/233517/files/figure.png" pageId="3">Fig. 17</figureCitation>
|
||
); 6. internal sac of the aedeagus without accessory sclerites.
|
||
</paragraph>
|
||
<paragraph id="2D41A60F2B55944F6FD0FB7BFAE6FA85" blockId="3.[151,1437,1004,2037]" pageId="3">
|
||
The females of
|
||
<taxonomicName id="EAFEDD8C2B55944F6E6FFB7AFE09FB19" box="[376,501,1185,1209]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="felicitae" status="sp. nov.">
|
||
<emphasis id="1F8A7A1D2B55944F6E6FFB7AFE70FB18" box="[376,396,1185,1208]" italics="true" pageId="3">O</emphasis>
|
||
.
|
||
<emphasis id="1F8A7A1D2B55944F6E8CFB7AFE09FB19" box="[411,501,1185,1209]" italics="true" pageId="3">felicitae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="04B9C7662B55944F6EEAFB7BFDC1FB19" box="[509,573,1184,1209]" pageId="3" rank="species">
|
||
<emphasis id="1F8A7A1D2B55944F6EEAFB7BFDC1FB19" bold="true" box="[509,573,1184,1209]" pageId="3">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="EAFEDD8C2B55944F6D63FB7AFCB1FB18" box="[628,845,1185,1208]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="compressicornis">
|
||
<emphasis id="1F8A7A1D2B55944F6D63FB7AFD74FB18" box="[628,648,1185,1208]" italics="true" pageId="3">O</emphasis>
|
||
.
|
||
<emphasis id="1F8A7A1D2B55944F6D80FB7AFCB1FB18" box="[663,845,1185,1208]" italics="true" pageId="3">compressicornis</emphasis>
|
||
</taxonomicName>
|
||
can be separated by:
|
||
<taxonomicName id="EAFEDD8C2B55944F6B53FB7AFB3DFB19" box="[1092,1217,1185,1209]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="felicitae" status="sp. nov.">
|
||
<emphasis id="1F8A7A1D2B55944F6B53FB7AFBA4FB18" box="[1092,1112,1185,1208]" italics="true" pageId="3">O</emphasis>
|
||
.
|
||
<emphasis id="1F8A7A1D2B55944F6B71FB7AFB3DFB19" box="[1126,1217,1185,1209]" italics="true" pageId="3">felicitae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1F8A7A1D2B55944F6BDFFB7BFAF4FB19" bold="true" box="[1224,1288,1184,1209]" pageId="3">
|
||
<taxonomicNameLabel id="04B9C7662B55944F6BDFFB7BFAF4FB19" box="[1224,1288,1184,1209]" pageId="3" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
elytral striae punctures almost three times as wide as connecting stria, and eighth elytral stria with punctures longitudinally elongated, irregularly shaped (not rounded) (
|
||
<figureCitation id="B5C5BA8A2B55944F6D84FB33FD24FAA1" box="[659,728,1256,1281]" captionStart="FIGURES 7 – 9" captionStartId="3.[151,264,881,904]" captionTargetBox="[265,1325,184,863]" captionTargetId="figure@3.[252,1334,173,876]" captionTargetPageId="3" captionText="FIGURES 7 – 9. Figures 7 – 8 Ontherus felicitae n. sp. Figure 7, female dorsal view (paratype); Figure 8, 8 th elytral stria, female (paratype). Figure 9, Ontherus compressicornis Luederwaldt 1931, 8 th elytral stria, female. Black arrow pseudoepipleuron, white arrow 8 th elytral stria." httpUri="https://zenodo.org/record/233516/files/figure.png" pageId="3">Fig. 8</figureCitation>
|
||
). In
|
||
<taxonomicName id="EAFEDD8C2B55944F6C18FB32FC15FAA0" box="[783,1001,1257,1280]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="compressicornis">
|
||
<emphasis id="1F8A7A1D2B55944F6C18FB32FCDFFAA0" box="[783,803,1257,1280]" italics="true" pageId="3">O</emphasis>
|
||
.
|
||
<emphasis id="1F8A7A1D2B55944F6C24FB32FC15FAA0" box="[819,1001,1257,1280]" italics="true" pageId="3">compressicornis</emphasis>
|
||
</taxonomicName>
|
||
the elytral striae punctures are almost five times as wide as connecting stria, and the punctures on the eighth elytral stria are rounded (
|
||
<figureCitation id="B5C5BA8A2B55944F6BD0FAD7FAF7FA85" box="[1223,1291,1292,1317]" captionStart="FIGURES 7 – 9" captionStartId="3.[151,264,881,904]" captionTargetBox="[265,1325,184,863]" captionTargetId="figure@3.[252,1334,173,876]" captionTargetPageId="3" captionText="FIGURES 7 – 9. Figures 7 – 8 Ontherus felicitae n. sp. Figure 7, female dorsal view (paratype); Figure 8, 8 th elytral stria, female (paratype). Figure 9, Ontherus compressicornis Luederwaldt 1931, 8 th elytral stria, female. Black arrow pseudoepipleuron, white arrow 8 th elytral stria." httpUri="https://zenodo.org/record/233516/files/figure.png" pageId="3">Fig. 9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="65E4F5842B55944A6FD0FAEBFBACFDC0" lastPageId="6" pageId="3" type="description">
|
||
<paragraph id="2D41A60F2B5594496FD0FAEBFB4CF972" blockId="3.[151,1437,1004,2037]" lastBlockId="5.[151,1437,1433,2034]" lastPageId="5" pageId="3">
|
||
<emphasis id="1F8A7A1D2B55944F6FD0FAEBFEA5FAE9" bold="true" box="[199,345,1328,1353]" pageId="3">Description.</emphasis>
|
||
<typeStatus id="F24518AD2B55944F6E76FAEBFE3FFAE9" box="[353,451,1328,1353]" pageId="3" type="holotype">holotype</typeStatus>
|
||
male: size, length, from pronotal anterior margin to elytral apex,
|
||
<quantity id="EA060BEA2B55944F6BACFAEAFACEFAE8" box="[1211,1330,1329,1353]" metricMagnitude="-2" metricUnit="m" metricValue="1.217" pageId="3" unit="mm" value="12.17">12.17 mm</quantity>
|
||
; humeral width,
|
||
<quantity id="EA060BEA2B55944F6FFDFA8EFEABFACC" box="[234,343,1365,1389]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="3" unit="mm" value="7.3">7.30 mm</quantity>
|
||
. Body black with weakly reddish reflection ventrally, easier to see on meso-metafemur and metasternum, and weakly bluish reflection dorsally, easier to see on elytra.
|
||
<emphasis id="1F8A7A1D2B55944F6B19FAA3FBB2FA31" bold="true" box="[1038,1102,1400,1425]" pageId="3">Head</emphasis>
|
||
with a backwardly directed, laterally gradually compressed horn (
|
||
<figureCitation id="B5C5BA8A2B55944F6D2EFA47FD70FA15" box="[569,652,1436,1461]" captionStart="FIGURES 19 – 24" captionStartId="5.[151,264,1274,1297]" captionTargetBox="[192,1393,203,1243]" captionTargetId="figure@5.[180,1406,193,1253]" captionTargetPageId="5" captionText="FIGURES 19 – 24. Cephalic horn size variation and plate-shape sclerite in Ontherus felicitae n. sp. Figures 19 – 20 welldeveloped male (holotype). Figure 19, lateral view head and pronotum; Figure 20, plate-shape sclerite. Figures 21 – 22 moderately developed male (paratype). Figure 21, lateral view, head and pronotum; Figure 22, plate-shape sclerite. Figures 23 – 24 underdeveloped male (paratype). Figure 23, lateral view, head and pronotum; Figure 24, plate-shape sclerite." httpUri="https://zenodo.org/record/233518/files/figure.png" pageId="3">Fig. 19</figureCitation>
|
||
), truncate apically. Clypeus and genae transversely wrinkled; anterior edge of clypeus feebly bisinuate, edge between the sinuation and the clypeogenal junction rounded; clypeogenal junction feebly notched; clypeofrontal suture feebly noticeable; vertex with a deep depressions, on the internal side of each eye (
|
||
<figureCitation id="B5C5BA8A2B55944F6E3EF9D3FE97F981" box="[297,363,1544,1569]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1863,1886]" captionTargetBox="[316,1265,214,1828]" captionTargetId="figure@2.[304,1284,193,1842]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Figures 1 – 3 Ontherus felicitae n. sp. Figure 1, dorsal view (holotype); Figure 2, frontal view, head and pronotum (holotype); Figure 3, elytral sculpture (holotype). Figures 4 – 6 male of Ontherus compressicornis Luederwaldt 1931. Figure 4, dorsal view; Figure 5, frontal view head and pronotum; Figure 6, elytral sculpture." httpUri="https://zenodo.org/record/233515/files/figure.png" pageId="3">Fig. 1</figureCitation>
|
||
).
|
||
<emphasis id="1F8A7A1D2B55944F6E92F9D3FE02F981" bold="true" box="[389,510,1544,1569]" pageId="3">Pronotum</emphasis>
|
||
smooth with simple punctures (not ocellate); punctures in the antero-lateral areas denser and larger than in the disc, the disc is smooth, the punctures are small and not easily visible (
|
||
<figureCitation id="B5C5BA8A2B55944F6BEEF9F7FA73F9E5" box="[1273,1423,1580,1605]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1863,1886]" captionTargetBox="[316,1265,214,1828]" captionTargetId="figure@2.[304,1284,193,1842]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Figures 1 – 3 Ontherus felicitae n. sp. Figure 1, dorsal view (holotype); Figure 2, frontal view, head and pronotum (holotype); Figure 3, elytral sculpture (holotype). Figures 4 – 6 male of Ontherus compressicornis Luederwaldt 1931. Figure 4, dorsal view; Figure 5, frontal view head and pronotum; Figure 6, elytral sculpture." httpUri="https://zenodo.org/record/233515/files/figure.png" pageId="3">Figs. 1 and 2</figureCitation>
|
||
). Punctures separated by twice the diameter in the anterolateral areas and by more than twice the length of the diameter in the disc; disc with two oblique raised carinae (
|
||
<figureCitation id="B5C5BA8A2B55944F6C2FF9AFFC82F92D" box="[824,894,1652,1677]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1863,1886]" captionTargetBox="[316,1265,214,1828]" captionTargetId="figure@2.[304,1284,193,1842]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Figures 1 – 3 Ontherus felicitae n. sp. Figure 1, dorsal view (holotype); Figure 2, frontal view, head and pronotum (holotype); Figure 3, elytral sculpture (holotype). Figures 4 – 6 male of Ontherus compressicornis Luederwaldt 1931. Figure 4, dorsal view; Figure 5, frontal view head and pronotum; Figure 6, elytral sculpture." httpUri="https://zenodo.org/record/233515/files/figure.png" pageId="3">Fig. 2</figureCitation>
|
||
); carina with irregular border (small denticles), except lateral and inner edges that are smooth and rounded; pronotal anterior margin wider and sinuated behind the eyes; anterior, lateral and posterior area of hypomeron with setae, setae denser anteriorly, without punctures or setae submedially;
|
||
<emphasis id="1F8A7A1D2B55944F6E66F93BFE03F959" bold="true" box="[369,511,1760,1785]" pageId="3">Prosternum</emphasis>
|
||
with dense punctures, each puncture with a long seta.
|
||
<emphasis id="1F8A7A1D2B55944F6B62F93BFAE4F959" bold="true" box="[1141,1304,1760,1785]" pageId="3">Mesosternum</emphasis>
|
||
with dense punctures, each puncture separated by less than one diameter, and setae shorter than those of the prosternum, except along the mesocoxa where they are subequal in size.
|
||
<emphasis id="1F8A7A1D2B55944F6C92F8F3FBDBF8E1" bold="true" box="[901,1063,1832,1857]" pageId="3">Metasternum</emphasis>
|
||
smooth and glabrous on disc, shagreened and with few setae on lateral lobes; lateral lobes punctures larger posteriorly and denser anteriorly. Median lobe punctures larger and denser than discal punctures; median lobe of metasternum anteriorly with atrophied carina medially, mesometasternal suture almost straight.
|
||
<emphasis id="1F8A7A1D2B55944F6C87F84FFC18F80D" bold="true" box="[912,996,1940,1965]" pageId="3">Elytral</emphasis>
|
||
striae deeper apically; strial punctures simple, separated by one diameter on the disc and by less than one diameter apically; 1–7th strial punctures rounded, eighth stria with punctures longitudinally elongated, irregularly shaped (not rounded); strial punctures almost three times as wide as the connecting stria (
|
||
<figureCitation id="B5C5BA8A2B5394496DC3FA42FCEBFA11" box="[724,791,1433,1458]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1863,1886]" captionTargetBox="[316,1265,214,1828]" captionTargetId="figure@2.[304,1284,193,1842]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Figures 1 – 3 Ontherus felicitae n. sp. Figure 1, dorsal view (holotype); Figure 2, frontal view, head and pronotum (holotype); Figure 3, elytral sculpture (holotype). Figures 4 – 6 male of Ontherus compressicornis Luederwaldt 1931. Figure 4, dorsal view; Figure 5, frontal view head and pronotum; Figure 6, elytral sculpture." httpUri="https://zenodo.org/record/233515/files/figure.png" pageId="5">Fig. 3</figureCitation>
|
||
), except at the apex of the sixth and seventh striae where the punctures are slightly larger.
|
||
<emphasis id="1F8A7A1D2B5394496D10FA66FD7AFA76" bold="true" box="[519,646,1469,1494]" pageId="5">Interstriae</emphasis>
|
||
smooth, with simple punctures (
|
||
<figureCitation id="B5C5BA8A2B5394496CE1FA66FBC6FA76" box="[1014,1082,1469,1494]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1863,1886]" captionTargetBox="[316,1265,214,1828]" captionTargetId="figure@2.[304,1284,193,1842]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Figures 1 – 3 Ontherus felicitae n. sp. Figure 1, dorsal view (holotype); Figure 2, frontal view, head and pronotum (holotype); Figure 3, elytral sculpture (holotype). Figures 4 – 6 male of Ontherus compressicornis Luederwaldt 1931. Figure 4, dorsal view; Figure 5, frontal view head and pronotum; Figure 6, elytral sculpture." httpUri="https://zenodo.org/record/233515/files/figure.png" pageId="5">Fig. 3</figureCitation>
|
||
); punctures small and disperse, smaller than the line of the striae and separated by 2–3 times their diameter.
|
||
<emphasis id="1F8A7A1D2B5394496B52FA3AFAC2FA5A" bold="true" box="[1093,1342,1505,1530]" pageId="5">Abdominal sternites</emphasis>
|
||
weakly shagreened; sternite 1–5 reduced in width medially, sternite 6 as wide as 2–5 combined medially.
|
||
<emphasis id="1F8A7A1D2B5394496A3EF9DEFA60F9BE" bold="true" box="[1321,1436,1541,1566]" pageId="5">Pygidium</emphasis>
|
||
approximately 1.5 times wider than long; pygidium smooth between punctures, punctures fine and slightly larger than the punctures on the middle of the sixth abdominal sternite.
|
||
<emphasis id="1F8A7A1D2B5394496CB0F996FC23F9C6" bold="true" box="[935,991,1613,1638]" pageId="5">Legs</emphasis>
|
||
, anterior tibiae tridentate, basal tooth smallest, basal and medial teeth laterally projected, apical tooth projecting forward; anterior femur weakly shagreened, with large setiferous punctures on posterior half; surface of middle and posterior femur weakly shagreened, with small punctures separated by 2–3 times their diameter; hind leg unmodified.
|
||
</paragraph>
|
||
<caption id="7981F6872B5294486F80F8A0FDFBF84D" httpUri="https://zenodo.org/record/233517/files/figure.png" pageId="4" targetBox="[241,1344,208,1881]" targetPageId="4">
|
||
<paragraph id="2D41A60F2B5294486F80F8A0FDFBF84D" blockId="4.[151,1436,1915,2029]" pageId="4">
|
||
<emphasis id="1F8A7A1D2B5294486F80F8A0FEAAF831" bold="true" box="[151,342,1915,1938]" pageId="4">FIGURES 10–18.</emphasis>
|
||
Figures 10–14 male genitalia
|
||
<taxonomicName id="EAFEDD8C2B5294486D84F8A7FCB7F832" box="[659,843,1916,1938]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="felicitae" status="sp. nov.">
|
||
<emphasis id="1F8A7A1D2B5294486D84F8A7FCB7F832" box="[659,843,1916,1938]" italics="true" pageId="4">Ontherus felicitae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1F8A7A1D2B5294486C44F8A7FC73F831" bold="true" box="[851,911,1916,1937]" pageId="4">
|
||
<taxonomicNameLabel id="04B9C7662B5294486C44F8A7FC73F831" box="[851,911,1916,1937]" pageId="4" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Figure 10, lateral view aedeagus; Figure 11, basal sclerite; Figure 12, plate-shape sclerite; Figure 13, accessory sclerite; Figure 14, elongate sclerite. Figures 15–18 male genitalia
|
||
<taxonomicName id="EAFEDD8C2B5294486F80F862FD9DF86F" authority="Luederwaldt 1931" authorityName="Luederwaldt" authorityYear="1931" box="[151,609,1977,1999]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="compressicornis">
|
||
<emphasis id="1F8A7A1D2B5294486F80F862FE5CF86F" box="[151,416,1977,1999]" italics="true" pageId="4">Ontherus compressicornis</emphasis>
|
||
Luederwaldt 1931
|
||
</taxonomicName>
|
||
. Figure 15, lateral view aedeagus; Figure 16, basal sclerite; Figure 17, plate-shape sclerite; Figure 18, elongate sclerite.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="7981F6872B5394496F80FB21FAF8FACC" httpUri="https://zenodo.org/record/233518/files/figure.png" pageId="5" targetBox="[192,1393,203,1243]" targetPageId="5">
|
||
<paragraph id="2D41A60F2B5394496F80FB21FAF8FACC" blockId="5.[151,1436,1274,1388]" pageId="5">
|
||
<emphasis id="1F8A7A1D2B5394496F80FB21FEA4FAAF" bold="true" box="[151,344,1274,1297]" pageId="5">FIGURES 19–24.</emphasis>
|
||
Cephalic horn size variation and plate-shape sclerite in
|
||
<taxonomicName id="EAFEDD8C2B5394496CAEFB21FB88FAB0" box="[953,1140,1274,1296]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="felicitae" status="sp. nov.">
|
||
<emphasis id="1F8A7A1D2B5394496CAEFB21FB88FAB0" box="[953,1140,1274,1296]" italics="true" pageId="5">Ontherus felicitae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1F8A7A1D2B5394496B69FB20FB47FAB0" bold="true" box="[1150,1211,1275,1296]" pageId="5">
|
||
<taxonomicNameLabel id="04B9C7662B5394496B69FB20FB47FAB0" box="[1150,1211,1275,1296]" pageId="5" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Figures 19–20 welldeveloped male (holotype). Figure 19, lateral view head and pronotum; Figure 20, plate-shape sclerite. Figures 21–22 moderately developed male (paratype). Figure 21, lateral view, head and pronotum; Figure 22, plate-shape sclerite. Figures 23– 24 underdeveloped male (paratype). Figure 23, lateral view, head and pronotum; Figure 24, plate-shape sclerite.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="2D41A60F2B5394496FD0F906FCC6F86E" blockId="5.[151,1437,1433,2034]" pageId="5">
|
||
<emphasis id="1F8A7A1D2B5394496FD0F906FE8FF956" bold="true" box="[199,371,1757,1782]" pageId="5">Male genitalia</emphasis>
|
||
:
|
||
<emphasis id="1F8A7A1D2B5394496E95F906FE18F956" bold="true" box="[386,484,1757,1782]" pageId="5">aedeagu</emphasis>
|
||
s (
|
||
<figureCitation id="B5C5BA8A2B5394496D17F906FDAEF956" box="[512,594,1757,1782]" captionStart="FIGURES 10 – 18" captionStartId="4.[151,264,1915,1938]" captionTargetBox="[241,1344,208,1881]" captionTargetId="figure@4.[229,1358,193,1894]" captionTargetPageId="4" captionText="FIGURES 10 – 18. Figures 10 – 14 male genitalia Ontherus felicitae n. sp. Figure 10, lateral view aedeagus; Figure 11, basal sclerite; Figure 12, plate-shape sclerite; Figure 13, accessory sclerite; Figure 14, elongate sclerite. Figures 15 – 18 male genitalia Ontherus compressicornis Luederwaldt 1931. Figure 15, lateral view aedeagus; Figure 16, basal sclerite; Figure 17, plate-shape sclerite; Figure 18, elongate sclerite." httpUri="https://zenodo.org/record/233517/files/figure.png" pageId="5">Fig. 10</figureCitation>
|
||
) with phallobase elongated, nearly twice as long as parameres. Parameres sinuated in lateral view, apex turned up. Internal sac well developed with four sclerites in the apical area.
|
||
<emphasis id="1F8A7A1D2B5394496A4CF8DAFF10F89E" bold="true" pageId="5">Basal sclerite</emphasis>
|
||
(
|
||
<figureCitation id="B5C5BA8A2B5394496E16F8FEFEA9F89E" box="[257,341,1829,1854]" captionStart="FIGURES 10 – 18" captionStartId="4.[151,264,1915,1938]" captionTargetBox="[241,1344,208,1881]" captionTargetId="figure@4.[229,1358,193,1894]" captionTargetPageId="4" captionText="FIGURES 10 – 18. Figures 10 – 14 male genitalia Ontherus felicitae n. sp. Figure 10, lateral view aedeagus; Figure 11, basal sclerite; Figure 12, plate-shape sclerite; Figure 13, accessory sclerite; Figure 14, elongate sclerite. Figures 15 – 18 male genitalia Ontherus compressicornis Luederwaldt 1931. Figure 15, lateral view aedeagus; Figure 16, basal sclerite; Figure 17, plate-shape sclerite; Figure 18, elongate sclerite." httpUri="https://zenodo.org/record/233517/files/figure.png" pageId="5">Fig. 11</figureCitation>
|
||
) robust, well sclerotized, with a thick base and two lateral arms, one shorter, and the other elongated and bent apically.
|
||
<emphasis id="1F8A7A1D2B5394496EC0F892FD41F8C2" bold="true" box="[471,701,1865,1890]" pageId="5">Plate-shape sclerite</emphasis>
|
||
(
|
||
<figureCitation id="B5C5BA8A2B5394496DDAF892FCDDF8C1" box="[717,801,1865,1890]" captionStart="FIGURES 10 – 18" captionStartId="4.[151,264,1915,1938]" captionTargetBox="[241,1344,208,1881]" captionTargetId="figure@4.[229,1358,193,1894]" captionTargetPageId="4" captionText="FIGURES 10 – 18. Figures 10 – 14 male genitalia Ontherus felicitae n. sp. Figure 10, lateral view aedeagus; Figure 11, basal sclerite; Figure 12, plate-shape sclerite; Figure 13, accessory sclerite; Figure 14, elongate sclerite. Figures 15 – 18 male genitalia Ontherus compressicornis Luederwaldt 1931. Figure 15, lateral view aedeagus; Figure 16, basal sclerite; Figure 17, plate-shape sclerite; Figure 18, elongate sclerite." httpUri="https://zenodo.org/record/233517/files/figure.png" pageId="5">Fig. 12</figureCitation>
|
||
), called sigmoid sclerite in
|
||
<bibRefCitation id="496FDBFE2B5394496B41F892FB0AF8C2" author="Genier" box="[1110,1270,1865,1890]" pageId="5" refString="Genier, F. (1996) A revision of the Neotropical genus Ontherus Erichson (Coleoptera: Scarabaeidae, Scarabaeinae). Memoirs of the Entomological Society of Canada, 170, 1 - 169. http: // dx. doi. org / 10.4039 / entm 128170 fv" type="journal article" year="1996">Génier (1996)</bibRefCitation>
|
||
, is slender and elongated, with the apices bent laterally.
|
||
<emphasis id="1F8A7A1D2B5394496D66F8B6FCB4F826" bold="true" box="[625,840,1901,1926]" pageId="5">Accessory sclerite</emphasis>
|
||
(
|
||
<figureCitation id="B5C5BA8A2B5394496C4CF8B6FC52F826" box="[859,942,1901,1926]" captionStart="FIGURES 10 – 18" captionStartId="4.[151,264,1915,1938]" captionTargetBox="[241,1344,208,1881]" captionTargetId="figure@4.[229,1358,193,1894]" captionTargetPageId="4" captionText="FIGURES 10 – 18. Figures 10 – 14 male genitalia Ontherus felicitae n. sp. Figure 10, lateral view aedeagus; Figure 11, basal sclerite; Figure 12, plate-shape sclerite; Figure 13, accessory sclerite; Figure 14, elongate sclerite. Figures 15 – 18 male genitalia Ontherus compressicornis Luederwaldt 1931. Figure 15, lateral view aedeagus; Figure 16, basal sclerite; Figure 17, plate-shape sclerite; Figure 18, elongate sclerite." httpUri="https://zenodo.org/record/233517/files/figure.png" pageId="5">Fig. 13</figureCitation>
|
||
) small and oval shaped.
|
||
<emphasis id="1F8A7A1D2B5394496BC4F8B6FA67F826" bold="true" box="[1235,1435,1901,1926]" pageId="5">Elongate sclerite</emphasis>
|
||
(
|
||
<figureCitation id="B5C5BA8A2B5394496F88F84AFF0EF809" box="[159,242,1937,1962]" captionStart="FIGURES 10 – 18" captionStartId="4.[151,264,1915,1938]" captionTargetBox="[241,1344,208,1881]" captionTargetId="figure@4.[229,1358,193,1894]" captionTargetPageId="4" captionText="FIGURES 10 – 18. Figures 10 – 14 male genitalia Ontherus felicitae n. sp. Figure 10, lateral view aedeagus; Figure 11, basal sclerite; Figure 12, plate-shape sclerite; Figure 13, accessory sclerite; Figure 14, elongate sclerite. Figures 15 – 18 male genitalia Ontherus compressicornis Luederwaldt 1931. Figure 15, lateral view aedeagus; Figure 16, basal sclerite; Figure 17, plate-shape sclerite; Figure 18, elongate sclerite." httpUri="https://zenodo.org/record/233517/files/figure.png" pageId="5">Fig. 14</figureCitation>
|
||
) with two separated filaments. Medial area of the internal sac without sclerites. Submedial area without defined raspules, but covered by homogeneous small scales.
|
||
</paragraph>
|
||
<paragraph id="2D41A60F2B53944A6FD0F802FD0AFEE0" blockId="5.[151,1437,1433,2034]" lastBlockId="6.[151,1437,151,717]" lastPageId="6" pageId="5">
|
||
<emphasis id="1F8A7A1D2B5394496FD0F802FEC7F852" bold="true" box="[199,315,2009,2034]" pageId="5">Variation.</emphasis>
|
||
The
|
||
<typeStatus id="F24518AD2B5394496E60F801FE1FF852" box="[375,483,2010,2034]" pageId="5" type="paratype">paratypes</typeStatus>
|
||
vary in sexual features and in the body development of the males. Female, head with two dentincles, without horn (
|
||
<figureCitation id="B5C5BA8A2B50944A6EE1FF4CFDC2FF10" box="[502,574,151,176]" captionStart="FIGURES 7 – 9" captionStartId="3.[151,264,881,904]" captionTargetBox="[265,1325,184,863]" captionTargetId="figure@3.[252,1334,173,876]" captionTargetPageId="3" captionText="FIGURES 7 – 9. Figures 7 – 8 Ontherus felicitae n. sp. Figure 7, female dorsal view (paratype); Figure 8, 8 th elytral stria, female (paratype). Figure 9, Ontherus compressicornis Luederwaldt 1931, 8 th elytral stria, female. Black arrow pseudoepipleuron, white arrow 8 th elytral stria." httpUri="https://zenodo.org/record/233516/files/figure.png" pageId="6">Fig. 7</figureCitation>
|
||
); clypeus and genae more wrinkled than developed males, vertex without depressions. Pronotal punctures nearly twice as large as in developed males, mainly in the antero-lateral areas; anterior third of pronotum with two straight transversal swelling, as long as the distance between the eyes (
|
||
<figureCitation id="B5C5BA8A2B50944A6A5EFF04FA72FF58" box="[1353,1422,223,248]" captionStart="FIGURES 7 – 9" captionStartId="3.[151,264,881,904]" captionTargetBox="[265,1325,184,863]" captionTargetId="figure@3.[252,1334,173,876]" captionTargetPageId="3" captionText="FIGURES 7 – 9. Figures 7 – 8 Ontherus felicitae n. sp. Figure 7, female dorsal view (paratype); Figure 8, 8 th elytral stria, female (paratype). Figure 9, Ontherus compressicornis Luederwaldt 1931, 8 th elytral stria, female. Black arrow pseudoepipleuron, white arrow 8 th elytral stria." httpUri="https://zenodo.org/record/233516/files/figure.png" pageId="6">Fig. 7</figureCitation>
|
||
). Sixth abdominal sternite as wide as 1–5 combined medially. Pygidium approximately three times wider than long. Anterior tibiae quadridentate, the basal tooth smallest.
|
||
</paragraph>
|
||
<paragraph id="2D41A60F2B50944A6FD0FE97FDB0FE70" blockId="6.[151,1437,151,717]" pageId="6">
|
||
Small males, head with small conical or subquadrate horn (
|
||
<figureCitation id="B5C5BA8A2B50944A6C49FE97FBF3FEC4" box="[862,1039,332,357]" captionStart="FIGURES 19 – 24" captionStartId="5.[151,264,1274,1297]" captionTargetBox="[192,1393,203,1243]" captionTargetId="figure@5.[180,1406,193,1253]" captionTargetPageId="5" captionText="FIGURES 19 – 24. Cephalic horn size variation and plate-shape sclerite in Ontherus felicitae n. sp. Figures 19 – 20 welldeveloped male (holotype). Figure 19, lateral view head and pronotum; Figure 20, plate-shape sclerite. Figures 21 – 22 moderately developed male (paratype). Figure 21, lateral view, head and pronotum; Figure 22, plate-shape sclerite. Figures 23 – 24 underdeveloped male (paratype). Figure 23, lateral view, head and pronotum; Figure 24, plate-shape sclerite." httpUri="https://zenodo.org/record/233518/files/figure.png" pageId="6">Figs. 21 and 23</figureCitation>
|
||
); clypeus and genae more wrinkled than developed males; vertex with shallower depressions. Pronotal punctures nearly twice as large as in developed males, principally in the anterolateral areas; anterior third of pronotum with two straight transversal swelling, as longs as the distance between the eyes.
|
||
</paragraph>
|
||
<paragraph id="2D41A60F2B50944A6FD0FE06FC99FDB8" blockId="6.[151,1437,151,717]" pageId="6">In small males and females the lateral swelling of pronotum, may be the same size or larger than inner swelling, depending of the development grade of the individual.</paragraph>
|
||
<paragraph id="2D41A60F2B50944A6FD0FDFFFBACFDC0" blockId="6.[151,1437,151,717]" pageId="6">
|
||
The greater variation found in the genitalia was observed in the plate sclerite, mainly in the size of the sclerite correlated with the size of the cephalic horn and the body of the beetle (
|
||
<figureCitation id="B5C5BA8A2B50944A6CADFD9CFBB8FDC0" box="[954,1092,583,608]" captionStart="FIGURES 19 – 24" captionStartId="5.[151,264,1274,1297]" captionTargetBox="[192,1393,203,1243]" captionTargetId="figure@5.[180,1406,193,1253]" captionTargetPageId="5" captionText="FIGURES 19 – 24. Cephalic horn size variation and plate-shape sclerite in Ontherus felicitae n. sp. Figures 19 – 20 welldeveloped male (holotype). Figure 19, lateral view head and pronotum; Figure 20, plate-shape sclerite. Figures 21 – 22 moderately developed male (paratype). Figure 21, lateral view, head and pronotum; Figure 22, plate-shape sclerite. Figures 23 – 24 underdeveloped male (paratype). Figure 23, lateral view, head and pronotum; Figure 24, plate-shape sclerite." httpUri="https://zenodo.org/record/233518/files/figure.png" pageId="6">Figs. 19–24</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="65E4F5842B50944A6FD0FDB7FACDFD25" box="[199,1329,620,645]" pageId="6" type="etymology">
|
||
<paragraph id="2D41A60F2B50944A6FD0FDB7FACDFD25" blockId="6.[151,1437,151,717]" box="[199,1329,620,645]" pageId="6">
|
||
<emphasis id="1F8A7A1D2B50944A6FD0FDB7FEBAFD25" bold="true" box="[199,326,620,645]" pageId="6">Etymology</emphasis>
|
||
. A patronymic after Felicity Edwards who collected all known specimens of this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="65E4F5842B50944A6FD0FD54FC0DFD6D" pageId="6" type="discussion">
|
||
<paragraph id="2D41A60F2B50944A6FD0FD54FC0DFD6D" blockId="6.[151,1437,151,717]" pageId="6">
|
||
<emphasis id="1F8A7A1D2B50944A6FD0FD54FEC8FD08" bold="true" box="[199,308,655,680]" pageId="6">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="EAFEDD8C2B50944A6E55FD4BFE43FD08" box="[322,447,656,680]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="felicitae" status="sp. nov.">
|
||
<emphasis id="1F8A7A1D2B50944A6E55FD4BFEAAFD07" box="[322,342,656,679]" italics="true" pageId="6">O</emphasis>
|
||
.
|
||
<emphasis id="1F8A7A1D2B50944A6E72FD4BFE43FD08" box="[357,447,656,680]" italics="true" pageId="6">felicitae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1F8A7A1D2B50944A6ED0FD54FDFAFD08" bold="true" box="[455,518,655,680]" pageId="6">
|
||
<taxonomicNameLabel id="04B9C7662B50944A6ED0FD54FDFAFD08" box="[455,518,655,680]" pageId="6" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
will key out to couplet eleven and to
|
||
<taxonomicName id="EAFEDD8C2B50944A6CA3FD4BFB71FD07" box="[948,1165,656,679]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="compressicornis">
|
||
<emphasis id="1F8A7A1D2B50944A6CA3FD4BFC34FD07" box="[948,968,656,679]" italics="true" pageId="6">O</emphasis>
|
||
.
|
||
<emphasis id="1F8A7A1D2B50944A6CC0FD4BFB71FD07" box="[983,1165,656,679]" italics="true" pageId="6">compressicornis</emphasis>
|
||
</taxonomicName>
|
||
in Génier’s key (1996). However it can be separated by the six characters described in the diagnosis.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |