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<document id="C869CFE8624F4C446B9AF2383525F91C" ID-DOI="10.17161/jom.i110.17001" ID-ISSN="2325-4467" ID-ZooBank="urn:lsid:zoobank.org:pub:53C3B699-96AD-4692-9AB3-05058738EADB" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722456082791" checkinUser="felipe" docAuthor="Engel, Michael S." docDate="2022" docId="926F87F592020D0D15AD1EB8B1DEFD80" docLanguage="en" docName="JMelittology.2022.110.1-51.pdf" docOrigin="Journal of Melittology 2022 (110)" docSource="http://dx.doi.org/10.17161/jom.i110.17001" docStyle="DocumentStyle:51747547F162FCF7F4DBBB3D518F67EC.3:JMelittology.2013-.journal_article.2cover" docStyleId="51747547F162FCF7F4DBBB3D518F67EC" docStyleName="JMelittology.2013-.journal_article.2cover" docStyleVersion="3" docTitle="Scaptotrigona ederi Engel 2022, new species" docType="treatment" docUuid="3004F475-B4C8-48ED-8C9A-350A4EAA18D2" docUuidSource="ZooBank" docVersion="1" lastPageNumber="12" masterDocId="6E56FF8D92050D00141C1A19B220FFD9" masterDocTitle="Notes on South American stingless bees of the genus Scaptotrigona (Hymenoptera: Apidae), Part II: Subgroup A of the postica species group" masterLastPageNumber="51" masterPageNumber="1" pageNumber="6" updateTime="1722456085954" updateUser="felipe" zenodo-license-document="CC-BY-NC-ND-4.0">
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<mods:title id="CF19F3690AC72564480EB7D491CEAF9F">Notes on South American stingless bees of the genus Scaptotrigona (Hymenoptera: Apidae), Part II: Subgroup A of the postica species group</mods:title>
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<mods:name id="6F08D3EED4D7B0C49872C59F551A8883" type="personal">
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<mods:namePart id="B5FC4850754D873EAF9C7BB6E563C397">Engel, Michael S.</mods:namePart>
<mods:affiliation id="A1D8E2C67C80703606E96E1C43535D92">Division of Entomology, Natural History Museum and Department of Ecology &amp; Evolutionary Biology, 1501 Crestline Drive - Suite 140, University of Kansas, Lawrence, Kansas 66045 - 4415, USA (msengel @ ku. edu). &amp; Division of Invertebrate Zoölogy, American Museum of Natural History, Central Park West at 79 Street, New York, New York 10024 - 5192, USA.</mods:affiliation>
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<mods:title id="29945FED6CF840C44B574C5751599457">Journal of Melittology</mods:title>
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<mods:part id="B0585CFFD356804F2CF71300006971E6">
<mods:date id="9F0C7E89112F25FAAEF4CAEC62E4EE8E">2022</mods:date>
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<mods:number id="2334CD1708DDDE58B5169B6B2E13E8F1">2022-03-14</mods:number>
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<mods:detail id="8D753101F6F13A368C5C554CA9D0077B" type="volume">
<mods:number id="1B58EE397FE9F64E2A22485DF03E5789">2022</mods:number>
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<mods:number id="4E2EA0AAFD8B26908D9E93B0F138A719">110</mods:number>
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<mods:classification id="55BD96077DBEAB837BC79D993E378968">journal article</mods:classification>
<mods:identifier id="1F9B09587F857E575DE28F1C17ABB79E" type="DOI">10.17161/jom.i110.17001</mods:identifier>
<mods:identifier id="3A684E52E6FEBEA689CD81D40B33E253" type="ISSN">2325-4467</mods:identifier>
<mods:identifier id="5413EF00A30D0A4496C1FF9DB4663FF4" type="ZooBank">urn:lsid:zoobank.org:pub:53C3B699-96AD-4692-9AB3-05058738EADB</mods:identifier>
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<treatment id="926F87F592020D0D15AD1EB8B1DEFD80" LSID="urn:lsid:zoobank.org:act:3004F475-B4C8-48ED-8C9A-350A4EAA18D2" httpUri="http://treatment.plazi.org/id/926F87F592020D0D15AD1EB8B1DEFD80" lastPageId="13" lastPageNumber="12" pageId="7" pageNumber="6">
<subSubSection id="52DC656892020D0715AD1EB8B61FFB00" pageId="7" pageNumber="6" type="nomenclature">
<paragraph id="1A7936E392020D0715AD1EB8B15FFB60" blockId="7.[433,895,1185,1209]" box="[433,895,1185,1209]" pageId="7" pageNumber="6">
<heading id="4131818F92020D0715AD1EB8B15FFB60" box="[433,895,1185,1209]" centered="true" fontSize="10" level="2" pageId="7" pageNumber="6" reason="2">
<taxonomicName id="DDC64D6092020D0715AD1EB8B0C7FB60" authority="Engel" authorityName="Engel" authorityYear="2022" box="[433,743,1185,1209]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="ederi" status="sp. nov.">
<emphasis id="28B2EAF192020D0715AD1EB8B0BBFB60" bold="true" box="[433,667,1185,1209]" italics="true" pageId="7" pageNumber="6">Scaptotrigona ederi</emphasis>
Engel
</taxonomicName>
,
<taxonomicNameLabel id="3381578A92020D0716EE1EBBB15FFB60" box="[754,895,1185,1209]" pageId="7" pageNumber="6" rank="species">new species</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="1A7936E392020D0714EC1EDAB61FFB00" blockId="7.[240,1087,1219,1273]" box="[240,1087,1219,1241]" pageId="7" pageNumber="6">
ZooBank:
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="52DC656892020D0716491EF8B0FBFB20" box="[597,731,1249,1273]" pageId="7" pageNumber="6" type="description">
<paragraph id="1A7936E392020D0716491EF8B0FBFB20" blockId="7.[240,1087,1219,1273]" box="[597,731,1249,1273]" pageId="7" pageNumber="6">
(
<figureCitation id="82FD2A6692020D0716421EF8B0F2FB20" box="[606,722,1249,1273]" captionStart-0="Figures 1–2" captionStart-1="Figures 3–6" captionStart-2="Figures 7–11" captionStartId-0="8.[160,241,1212,1234]" captionStartId-1="10.[161,242,1457,1479]" captionStartId-2="12.[160,241,1357,1379]" captionTargetBox-0="[175,1153,237,1199]" captionTargetBox-1="[165,1151,227,1434]" captionTargetBox-2="[160,1156,223,1346]" captionTargetId-0="figure-201@8.[160,1168,224,1206]" captionTargetId-1="figure-91@10.[160,1168,224,1451]" captionTargetId-2="figure-180@12.[160,1168,223,1351]" captionTargetPageId-0="8" captionTargetPageId-1="10" captionTargetPageId-2="12" captionText-0="Figures 1–2. Workers of Scaptotrigona ederi, new species. 1. Lateral habitus of individual on the darker end of the color spectrum. 2. Lateral habitus of individual on the lighter end of the spectrum." captionText-1="Figures 3–6. Workers of Scaptotrigona ederi, new species. 3. Facial view of individual on the lighter end of the color spectrum. 4. Facial view, darker worker. 5. Dorsal habitus, darker worker. 6. Dorsal habitus, lighter worker." captionText-2="Figures 7–11. Workers of Scaptotrigona ederi, new species. 7. Dorsal view of mesosoma, lighter worker. 8. Dorsal view of mesosoma, darker worker. 9. Mesoscutellum. 10. Prolateral surface of metatibia, metatarsus, and metapretarsus. 11. Apical view of metasoma." pageId="7" pageNumber="6">Figs. 1–11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="52DC656892020D0714CC1F38B1E2F900" pageId="7" pageNumber="6" type="diagnosis">
<paragraph id="1A7936E392020D0714CC1F38B1E2F900" blockId="7.[160,1168,1313,1754]" pageId="7" pageNumber="6">
DIAGNOSIS: This is a comparatively abundant and distinctive species in the western Andean region of
<collectingCountry id="62D1767392020D0715611F58B3D0FA80" box="[381,496,1345,1369]" name="Colombia" pageId="7" pageNumber="6">Colombia</collectingCountry>
around Cali and currently known to range in elevation from
<quantity id="DD3E9B0692020D0714FE1F78B34FFAA0" box="[226,367,1377,1401]" metricMagnitude="3" metricUnit="m" metricValue="1.1199999999999999" metricValueMax="1.5" metricValueMin="0.74" pageId="7" pageNumber="6" unit="m" value="1120.0" valueMax="1500.0" valueMin="740.0">740–1500 m</quantity>
, making it one of the highest occurrences for
<taxonomicName id="DDC64D6092020D07178A1F7BB60CFAA0" box="[918,1068,1378,1401]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="28B2EAF192020D07178A1F7BB60CFAA0" box="[918,1068,1378,1401]" italics="true" pageId="7" pageNumber="6">Scaptotrigona</emphasis>
</taxonomicName>
(second only to
<taxonomicName id="DDC64D6092020D0714E71F9BB3B6FA40" authorityName="Engel" authorityYear="2022" box="[251,406,1409,1433]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="magdalenae">
<emphasis id="28B2EAF192020D0714E71F9BB3B6FA40" box="[251,406,1409,1433]" italics="true" pageId="7" pageNumber="6">S. magdalenae</emphasis>
</taxonomicName>
,
<emphasis id="28B2EAF192020D0715B81F98B02CFA40" box="[420,524,1409,1433]" italics="true" pageId="7" pageNumber="6">vide infra</emphasis>
). Reports of
<taxonomicName id="DDC64D6092020D0716B01F9BB0DDFA40" authorityName="Engel" authorityYear="2022" box="[684,765,1409,1433]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF192020D0716B01F9BB0DDFA40" box="[684,765,1409,1433]" italics="true" pageId="7" pageNumber="6">S. ederi</emphasis>
</taxonomicName>
(as a
<emphasis id="28B2EAF192020D07175E1F9BB1C1FA40" box="[834,993,1409,1433]" italics="true" pageId="7" pageNumber="6">nomen nudum</emphasis>
) from the East of
<collectingCountry id="62D1767392020D0714A31FB8B312FA60" box="[191,306,1441,1465]" name="Colombia" pageId="7" pageNumber="6">Colombia</collectingCountry>
and elsewhere have generally been based on misidentifications. Among species in this subgroup,
<taxonomicName id="DDC64D6092020D0715CC1FDBB001FA00" authorityName="Engel" authorityYear="2022" box="[464,545,1473,1497]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF192020D0715CC1FDBB001FA00" box="[464,545,1473,1497]" italics="true" pageId="7" pageNumber="6">S. ederi</emphasis>
</taxonomicName>
is most superficially similar to those species of sub- group B in relation to its generally dark coloration, although it still falls within sub- group A at the darker end of the spectrum. The species is one of a few that lacks a defined genal tooth projecting upward from the lower margin of the lateral concavity in the preoccipital carina. Among those without such a defined genal tooth, the dark coloration, black or at least dark fuscous bristles of the vertex and mesoscutal antero- lateral corners, and nearly entirely black bristles of the metabasitarsus (and frequently the metatibia), serve to distinguish this species most readily. In addition, the promi- nent bristles of metasomal terga
<collectionCode id="7CD7AE2692020D0716031CDBB01AF900" box="[543,570,1730,1753]" pageId="7" pageNumber="6">III</collectionCode>
–
<collectionCode id="7CD7AE2692020D0716541CDBB07BF900" box="[584,603,1730,1753]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="7" pageNumber="6" type="Museum">V</collectionCode>
are yellow to fulvous in color.
</paragraph>
</subSubSection>
<caption id="4EB9666B920D0D0814BC1EA5B32BFAD2" pageId="8" pageNumber="7" startId="8.[160,241,1212,1234]" targetBox="[175,1153,237,1199]" targetPageId="8" targetType="figure">
<paragraph id="1A7936E3920D0D0814BC1EA5B32BFAD2" blockId="8.[160,1168,1212,1291]" pageId="8" pageNumber="7">
<emphasis id="28B2EAF1920D0D0814BC1EA5B304FB08" bold="true" box="[160,292,1212,1234]" pageId="8" pageNumber="7">Figures 1–2.</emphasis>
Workers of
<taxonomicName id="DDC64D60920D0D0815AA1EA4B055FB08" authorityName="Engel" authorityYear="2022" box="[438,629,1212,1234]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="ederi" status="sp. nov.">
<emphasis id="28B2EAF1920D0D0815AA1EA4B055FB08" box="[438,629,1212,1234]" italics="true" pageId="8" pageNumber="7">Scaptotrigona ederi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="3381578A920D0D08169F1EA4B127FB0B" box="[643,775,1212,1234]" pageId="8" pageNumber="7" rank="species">new species</taxonomicNameLabel>
.
<emphasis id="28B2EAF1920D0D0817071EA5B10DFB08" bold="true" box="[795,813,1212,1233]" pageId="8" pageNumber="7">1.</emphasis>
Lateral habitus of individual on the darker end of the color spectrum.
<emphasis id="28B2EAF1920D0D0816201EC0B06EFB37" bold="true" box="[572,590,1241,1262]" pageId="8" pageNumber="7">2.</emphasis>
Lateral habitus of individual on the lighter end of the spectrum.
</paragraph>
</caption>
<subSubSection id="52DC6568920D0D0C14CC1F2FB30FFA3C" lastPageId="12" lastPageNumber="11" pageId="8" pageNumber="7" type="description">
<paragraph id="1A7936E3920D0D0814CC1F2FB194F937" blockId="8.[160,1175,1334,1774]" pageId="8" pageNumber="7">
DESCRIPTION: ⚲: Total body length approximately
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, forewing length (to base of humeral sclerite)
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. Head wider than long, width
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, length
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; compound eye length
<quantity id="DD3E9B06920D0D0816BB1F6FB16AFA54" box="[679,842,1398,1422]" metricMagnitude="-3" metricUnit="m" metricValue="1.375" metricValueMax="1.39" metricValueMin="1.36" pageId="8" pageNumber="7" unit="mm" value="1.375" valueMax="1.39" valueMin="1.36">1.36–1.39 mm</quantity>
; upper interorbital distance
<quantity id="DD3E9B06920D0D0814BC1F8FB363FA74" box="[160,323,1430,1454]" metricMagnitude="-3" metricUnit="m" metricValue="1.58" metricValueMax="1.61" metricValueMin="1.55" pageId="8" pageNumber="7" unit="mm" value="1.58" valueMax="1.61" valueMin="1.55">1.55–1.61 mm</quantity>
, lower interorbital distance
<quantity id="DD3E9B06920D0D08168D1F8FB116FA74" box="[657,822,1430,1454]" metricMagnitude="-3" metricUnit="m" metricValue="1.4849999999999999" metricValueMax="1.52" metricValueMin="1.45" pageId="8" pageNumber="7" unit="mm" value="1.4849999999999999" valueMax="1.52" valueMin="1.45">1.45–1.52 mm</quantity>
. Scape length
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, about as long as torulocellar distance, torulocellar distance
<quantity id="DD3E9B06920D0D0817661FAFB600FA14" box="[890,1056,1462,1486]" metricMagnitude="-4" metricUnit="m" metricValue="8.95" metricValueMax="9.1" metricValueMin="8.8" pageId="8" pageNumber="7" unit="mm" value="0.895" valueMax="0.91" valueMin="0.88">0.88–0.91 mm</quantity>
; flagello- mere I longer than wide, longer than flagellomere II, remaining flagellomeres about as long as wide to slightly longer than wide except apical flagellomere longer than wide. Clypeus broader than long, approximately 1.7–1.75× as wide as long, length
<quantity id="DD3E9B06920D0D0810391C0FB2F1F994" metricMagnitude="-4" metricUnit="m" metricValue="6.25" metricValueMax="6.4" metricValueMin="6.1" pageId="8" pageNumber="7" unit="mm" value="0.625" valueMax="0.64" valueMin="0.61">0.61–0.64 mm</quantity>
, width
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. Malar area long, length approximately 1.55–1.7× flagellar diameter or 0.75–0.82× basal mandibular width. Preoccipital ridge carinate, weakly lamellate dorsally, with bordering medial indentation shallower than lateral inden- tations, laterally concavity interrupting carina comparatively shallow, lower margin of concavity with carina forming rounded angle, sometimes slightly acute, without sharply acute lamellate projection (genal tooth) extending upward.
</paragraph>
<paragraph id="1A7936E3920C0D0914CD1AFBB0B3FA80" blockId="9.[161,1175,225,1785]" pageId="9" pageNumber="8">
Integument of labiomaxillary complex yellow orange except prementum typically brown; labrum yellow orange to dark orange; mandible yellow orange to ferruginous; clypeus largely yellow orange to orange except typically with dark brown mark at medioapical margin and apicolateral corners from apical margin to epistomal sulcus (sometimes absent or faint, and sometimes these all merged medioapically), often with mediolongitudinal brown mark from base to about midlength, epistomal sulcus well delineated by brown to black; supraclypeal area brown to dark brown; malar space often dark brown, sometimes light brown to orange; face below tangent of antennal toruli orange to light yellow brown, typically slightly darker toward lateral borders of antennal toruli and often extending along inner orbit as narrow strip to point slightly above upper tangent of antennal toruli; scape yellow orange except typically brown dorsoapically, sometimes brown extending along entire dorsal surface; pedicel and flagellum typically dark brown above, with narrow underside of flagellum orange to light brown; remainder of face dark brown to black, sometimes with small areas of light brown immediately above antennal toruli; posterior of head dark brown; upper gena dark brown, remainder of gena orange, although if malar space dark brown, then such dark brown extends onto gena ventroapically; postgena orange to yellow orange. Mesosoma typically dark brown to black, with ferruginous apex to mesoscutellum, except sometimes with areas of reddish orange to reddish brown on mesepisternum, typically as small areas posteriorly and ventrally, and sometimes ventrally on propo- deum, and mesoscutellum ranging from wholly orange to wholly black, with degrees of intermixing of colors between, V-shaped incision in mesoscutellum typically black; tegula wholly yellow orange and semi-translucent, sometimes with slight brownish darkening anteriorly; legs typically yellow orange on coxae, trochanters, basal por- tions of tibiae, distitarsomeres, and inferior and superior areas of metabasitarsus, with remainder of tibiae, femora, and tarsi dark brown to black, except sometimes meso- and metacoxae light brown. Wing membranes hyaline clear to lightly parchment col- ored; veins yellow orange. Metasoma generally dark brown to black except anterior-facing surface of tergum I typically reddish brown to light yellow brown, dorsal-facing disc of tergum I often medially lighter reddish brown to yellowish brown and with exceptionally narrow apical marginal zone of reddish brown to yellow and terga II–
<collectionCode id="7CD7AE26920C0D0910681EBBB6B0FB60" box="[1140,1168,1186,1209]" pageId="9" pageNumber="8">IV</collectionCode>
with variably developed apical marginal zones of light brown to more frequently yel- low, sometimes quite broad on terga II and
<collectionCode id="7CD7AE26920C0D0916811EFBB09AFB20" box="[669,698,1250,1273]" pageId="9" pageNumber="8">III</collectionCode>
; tergum II sometimes with light reddish brown laterally and on disc and blending gradually to yellow apical marginal zone; yellow apical marginal zones of terga
<collectionCode id="7CD7AE26920C0D0916751F3BB0A4FAE0" box="[617,644,1314,1337]" pageId="9" pageNumber="8">III</collectionCode>
and
<collectionCode id="7CD7AE26920C0D0916DE1F3BB0FEFAE0" box="[706,734,1314,1337]" pageId="9" pageNumber="8">IV</collectionCode>
typically obscured by dense tomen- tum; sterna reddish brown to dark brown.
</paragraph>
<paragraph id="1A7936E3920C0D0B14CD1F7BB63FFE80" blockId="9.[161,1175,225,1785]" lastBlockId="11.[160,1168,225,1788]" lastPageId="11" lastPageNumber="10" pageId="9" pageNumber="8">
Integument mostly smooth and shining between punctures, punctures of head difficult to see, those of mesosoma and metasoma clear; clypeus with small, shallow, nearly contiguous punctures, in some small areas punctures separated by up to 0.75× a puncture width, punctures somewhat faint; supraclypeal area with small, shallow, nearly contiguous punctures; lower face with small punctures separated less than a puncture width, slightly fainter and more spaced than on clypeus, and slightly sparser along orbital border and in malar space; upper face with small punctures nearly con- tiguous to contiguous medially, such punctures becoming shallower toward ocelli and giving surface a roughened appearance, laterally punctures smaller and more distinct, separated by a puncture width or less; ocellocular area with punctures of upper lat- eral face becoming denser and blending posteriorly to slightly larger punctures like those of medial face; vertex and posterior of head punctures with shallow, coarse, contiguous punctures giving surface a roughened appearance, such punctures blend- ing to minute, shallow, punctures separated by less than a puncture width on gena; postgena nearly impunctate and smooth. Mesoscutum and mesoscutellum with small contiguous punctures, punctures coarser and more irregular in shape along borders; pleura with punctures as on mesoscutum, those punctures ventrally a bit more sepa- rated, posteriorly punctures more irregularly shaped and elongate; metepisternum with small, nearly contiguous punctures; lateral surface of propodeum as on metepi- sternum; basal area of propodeum tessellate to microalveolate. Metasomal terga with minute contiguous punctures blending to imbricate integument in apical marginal zones, except anterior-facing surface of tergum I impunctate and smooth; punctures laterally blending to imbricate integument, punctures weaker on tergum
<collectionCode id="7CD7AE26920E0D0B17E41B3BB62CFEE0" box="[1016,1036,290,313]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="11" pageNumber="10" type="Museum">V</collectionCode>
, tergum
<collectionCode id="7CD7AE26920E0D0B10681B3BB6B0FEE0" box="[1140,1168,290,313]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="11" pageNumber="10">VI</collectionCode>
largely impunctate and faintly imbricate; sterna finely and faintly imbricate.
</paragraph>
<caption id="4EB9666B920F0D0A14BD1FA8B07EF9D8" pageId="10" pageNumber="9" startId="10.[161,242,1457,1479]" targetBox="[165,1151,227,1434]" targetPageId="10" targetType="figure">
<paragraph id="1A7936E3920F0D0A14BD1FA8B07EF9D8" blockId="10.[161,1168,1457,1537]" pageId="10" pageNumber="9">
<emphasis id="28B2EAF1920F0D0A14BD1FA8B305FA1F" bold="true" box="[161,293,1457,1479]" pageId="10" pageNumber="9">Figures 3–6.</emphasis>
Workers of
<taxonomicName id="DDC64D60920F0D0A15AA1FABB054FA1F" authorityName="Engel" authorityYear="2022" box="[438,628,1457,1479]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="ederi" status="sp. nov.">
<emphasis id="28B2EAF1920F0D0A15AA1FABB054FA1F" box="[438,628,1457,1479]" italics="true" pageId="10" pageNumber="9">Scaptotrigona ederi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="3381578A920F0D0A169E1FABB126FA1E" box="[642,774,1457,1479]" pageId="10" pageNumber="9" rank="species">new species</taxonomicNameLabel>
.
<emphasis id="28B2EAF1920F0D0A17061FA8B10CFA1F" bold="true" box="[794,812,1457,1478]" pageId="10" pageNumber="9">3.</emphasis>
Facial view of individual on the lighter end of the color spectrum.
<emphasis id="28B2EAF1920F0D0A163A1FD7B018FA3A" bold="true" box="[550,568,1486,1507]" pageId="10" pageNumber="9">4.</emphasis>
Facial view, darker worker.
<emphasis id="28B2EAF1920F0D0A179D1FD7B1B3FA3A" bold="true" box="[897,915,1486,1507]" pageId="10" pageNumber="9">5.</emphasis>
Dorsal habitus, darker worker.
<emphasis id="28B2EAF1920F0D0A151C1FF2B332F9D9" bold="true" box="[256,274,1515,1536]" pageId="10" pageNumber="9">6.</emphasis>
Dorsal habitus, lighter worker.
</paragraph>
</caption>
<paragraph id="1A7936E3920E0D0B14CC1B78B126F900" blockId="11.[160,1168,225,1788]" pageId="11" pageNumber="10">
Fine pubescence generally consisting of sparse to scatered, minute to short, ap- pressed or decumbent yellow setae, such setae often simple but sometimes with min- ute branches or plumose, as well as areas of short, white, plumose setae such as dense on pronotal lobe, largely obscuring integument on metepisternum and lateral surface of propodeum; fine setae sparse on clypeus, more numerous on face and progressively longer on upper face and vertex; scape with sparse, minute, appressed setae and two erect to suberect, simple bristles at base (frequently obscured or even sometimes bro- ken off); terga I–
<collectionCode id="7CD7AE26920E0D0B1542185BB35AFD80" box="[350,378,578,601]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="11" pageNumber="10">VI</collectionCode>
with plumose yellow tomentum, on tergum I such setae are diffuse, more suberect, and not obscuring integument, frequently worn such that in some spec- imens they are completely absent to only scatered in irregular places (most easily seen in profile), tergum II with such setae diffuse to sometimes lacking laterally (most easily seen in profile), but dense in apical marginal zone, terga
<collectionCode id="7CD7AE26920E0D0B174D18DBB14CFD00" box="[849,876,706,729]" pageId="11" pageNumber="10">III</collectionCode>
–
<collectionCode id="7CD7AE26920E0D0B176618DBB1ADFD00" box="[890,909,706,729]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="11" pageNumber="10" type="Museum">V</collectionCode>
with such tomentum dense and obscuring integument, tomentum paler and sparser on tergum
<collectionCode id="7CD7AE26920E0D0B100018FBB61BFD20" box="[1052,1083,738,761]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="11" pageNumber="10">VI</collectionCode>
; sterna II–
<collectionCode id="7CD7AE26920E0D0B14A3191BB2F2FCC0" box="[191,210,770,793]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="11" pageNumber="10" type="Museum">V</collectionCode>
with scopa composed of elongate, erect, simple setae with characteristically wavy apices, such scopal setae yellowish. Bristles (thick, typically erect, often simple or with minute pectinate branches apically) typically sparse on body except in certain areas as noted. Labrum with sparse, erect, simple, yellow to fulvous bristles, those medially longest; vertex with numerous erect, simple, fulvous to sometimes fuscous bristles, longest medially and extending across vertex to just posterior to upper orbit; prono- tal dorsolateral margin near and anterior to pronotal lobe with bristles, sometimes arched apically, yellow to fulvous; mesoscutum anterior and lateral borders with erect bristles, those on lateral border short, bristles largely fulvous to sometimes fuscous along anterior margin, typically fuscous bristles intermixed with fulvous bristles in an- terolateral corner but sometimes intermixed along entire anterior margin, bristles on lateral border typically fuscous to black; tegula with short, suberect bristles anteriorly, such bristles typically yellow or fulvous, more uncommonly fuscous; mesoscutellum with abundant, elongate, erect bristles posteriorly, such bristles minutely pectinate apically, mesoscutellar bristles typically light fulvous, sometimes more darkly ful- vous; mesepisternum with scatered, subdecumbent fulvous bristles, longer ventrally, along rounded anterior border with preëpisternum a line of abundant, suberect to subdecumbent bristles extending ventrally from pronotal lobe, such bristles fulvous to dark fulvous; lateral surface of propodeum with scatered, long, erect, yellow to fulvous bristles emerging amid dense, appressed, white, plumose setae. Legs with abundant bristles, those on basal podites typically yellow to fulvous, those apically on tibiae and tarsi typically black, sometimes with black bristles apically on femora, particularly metafemur, distitarsi typically with fulvous bristles dominant; metatibia with elongate simple black corbicular bristles and margins with long, arched, largely simple, black bristles; penicillum black; rastellum fulvous to fuscous; metabasitarsus with scatered erect black bristles on prolateral surface and numerous black bristles on margins intermixed with fulvous bristles, those of inferior margin most numerous; a few black bristles on distitarsomeres. Metasoma with numerous, prominent, suberect to subdecumbent, yellow to fulvous bristles on terga
<collectionCode id="7CD7AE26920E0D0B17341C9BB163F940" box="[808,835,1666,1689]" pageId="11" pageNumber="10">III</collectionCode>
–
<collectionCode id="7CD7AE26920E0D0B174C1C9BB143F940" box="[848,867,1666,1689]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="11" pageNumber="10" type="Museum">V</collectionCode>
arising amid tomentum, such bristles sparse to absent on tergum II, and more scatered on tergum
<collectionCode id="7CD7AE26920E0D0B10031CBBB61BF960" box="[1055,1083,1698,1721]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="11" pageNumber="10">VI</collectionCode>
except longer and more numerous laterally and on margin.
</paragraph>
<paragraph id="1A7936E3920E0D0B14CC1CFBB30FF920" blockId="11.[160,1168,225,1788]" box="[208,303,1762,1788]" pageId="11" pageNumber="10">
<specimenCount id="0CC0FD6A920E0D0B14CC1CFBB2C4F925" box="[208,228,1762,1788]" count="1" pageId="11" pageNumber="10" type="female">♀</specimenCount>
:
<emphasis id="28B2EAF1920E0D0B14ED1CFBB308F920" box="[241,296,1762,1785]" italics="true" pageId="11" pageNumber="10">Latet</emphasis>
.
</paragraph>
<caption id="4EB9666B92090D0C14BC1F54B19CFA45" pageId="12" pageNumber="11" startId="12.[160,241,1357,1379]" targetBox="[160,1156,223,1346]" targetPageId="12" targetType="figure">
<paragraph id="1A7936E392090D0C14BC1F54B19CFA45" blockId="12.[160,1168,1357,1437]" pageId="12" pageNumber="11">
<emphasis id="28B2EAF192090D0C14BC1F54B30EFABB" bold="true" box="[160,302,1357,1379]" pageId="12" pageNumber="11">Figures 7–11.</emphasis>
Workers of
<taxonomicName id="DDC64D6092090D0C15A41F54B054FABB" authorityName="Engel" authorityYear="2022" box="[440,628,1357,1378]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="ederi" status="sp. nov.">
<emphasis id="28B2EAF192090D0C15A41F54B054FABB" box="[440,628,1357,1378]" italics="true" pageId="12" pageNumber="11">Scaptotrigona ederi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="3381578A92090D0C169C1F54B122FABA" box="[640,770,1357,1379]" pageId="12" pageNumber="11" rank="species">new species</taxonomicNameLabel>
.
<emphasis id="28B2EAF192090D0C170E1F54B104FABB" bold="true" box="[786,804,1357,1378]" pageId="12" pageNumber="11">7.</emphasis>
Dorsal view of mesosoma, lighter worker.
<emphasis id="28B2EAF192090D0C151C1F73B332FAA6" bold="true" box="[256,274,1386,1407]" pageId="12" pageNumber="11">8.</emphasis>
Dorsal view of mesosoma, darker worker.
<emphasis id="28B2EAF192090D0C16F81F73B0D6FAA6" bold="true" box="[740,758,1386,1407]" pageId="12" pageNumber="11">9.</emphasis>
Mesoscutellum.
<emphasis id="28B2EAF192090D0C17AD1F73B1EFFAA6" bold="true" box="[945,975,1386,1407]" pageId="12" pageNumber="11">10.</emphasis>
Prolateral surface of metatibia, metatarsus, and metapretarsus.
<emphasis id="28B2EAF192090D0C16981F9FB082FA42" bold="true" box="[644,674,1414,1435]" pageId="12" pageNumber="11">11.</emphasis>
Apical view of metasoma.
</paragraph>
</caption>
<paragraph id="1A7936E392090D0C14CC1FD7B30FFA3C" blockId="12.[160,1168,1486,1797]" box="[208,303,1486,1512]" pageId="12" pageNumber="11">
<specimenCount id="0CC0FD6A92090D0C14CC1FD7B2C4FA31" box="[208,228,1486,1512]" count="1" pageId="12" pageNumber="11" type="male">♂</specimenCount>
:
<emphasis id="28B2EAF192090D0C14ED1FD7B308FA3C" box="[241,296,1486,1509]" italics="true" pageId="12" pageNumber="11">Latet</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="52DC656892090D0D14CC1FF4B65FFE80" lastPageId="13" lastPageNumber="12" pageId="12" pageNumber="11" type="materials_examined">
<paragraph id="1A7936E392090D0C14CC1FF4B396F9FC" blockId="12.[160,1168,1486,1797]" pageId="12" pageNumber="11">
<materialsCitation id="AAAE3CBE92090D0C14CC1FF4B396F9FC" collectingDate="1977-02-04" collectionCode="SEMC" collectorName="M. D. Breed &amp; C. D. Michener" country="Colombia" location="Dagua" pageId="12" pageNumber="11" specimenCount="1" typeStatus="holotype">
<typeStatus id="C57D884192090D0C14CC1FF4B368F9DC" box="[208,328,1517,1541]" pageId="12" pageNumber="11" type="holotype">HOLOTYPE</typeStatus>
: ⚲,
<collectingCountry id="62D1767392090D0C15731FF4B3C5F9DC" box="[367,485,1517,1541]" name="Colombia" pageId="12" pageNumber="11">Colombia</collectingCountry>
: Prov. Valle,
<quantity id="DD3E9B0692090D0C16631FF4B09AF9DC" box="[639,698,1517,1541]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="11" unit="km" value="5.0">5 km</quantity>
SE of
<location id="1F19603892090D0C171A1FF7B176F9DF" LSID="urn:lsid:plazi:treatment:926F87F592020D0D15AD1EB8B1DEFD80:1F19603892090D0C171A1FF7B176F9DF" box="[774,854,1518,1542]" country="Colombia" name="Dagua" pageId="12" pageNumber="11">Dagua</location>
,
<date id="6E78102392090D0C177F1FF4B1C1F9DC" box="[867,993,1517,1541]" pageId="12" pageNumber="11" value="1977-02-04">
<collectingDate id="7E3CE9CB92090D0C177F1FF4B1C1F9DC" box="[867,993,1517,1541]" pageId="12" pageNumber="11" value="1977-02-04">4 Feb 1977</collectingDate>
</date>
,
<collectorName id="B733533592090D0C17F11FF7B654F9DC" box="[1005,1140,1517,1541]" pageId="12" pageNumber="11">M.D. Breed</collectorName>
&amp;
<collectorName id="B733533592090D0C14BC1C14B36CF9FC" box="[160,332,1549,1573]" pageId="12" pageNumber="11">C.D. Michener</collectorName>
(
<collectionCode id="7CD7AE2692090D0C15461C14B38AF9FC" box="[346,426,1549,1573]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="12" pageNumber="11" type="Museum">SEMC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="1A7936E392090D0D14CC1C34B65FFE80" blockId="12.[160,1168,1486,1797]" lastBlockId="13.[160,1168,225,601]" lastPageId="13" lastPageNumber="12" pageId="12" pageNumber="11">
<materialsCitation id="AAAE3CBE92090D0C14CC1C34B3F4F9BC" collectingDate="1977-02-04" collectionCode="SEMC" collectorName="M. D. Breed &amp; C. D. Michener" country="Colombia" location="Prov. Valle" pageId="12" pageNumber="11" specimenCount="1" typeStatus="paratype">
<typeStatus id="C57D884192090D0C14CC1C34B36EF99C" box="[208,334,1581,1605]" pageId="12" pageNumber="11" type="paratype">PARATYPES</typeStatus>
: 14⚲⚲,
<collectingCountry id="62D1767392090D0C15871C34B031F99C" box="[411,529,1581,1605]" name="Colombia" pageId="12" pageNumber="11">Colombia</collectingCountry>
:
<location id="1F19603892090D0C16071C37B0BDF99C" LSID="urn:lsid:plazi:treatment:926F87F592020D0D15AD1EB8B1DEFD80:1F19603892090D0C16071C37B0BDF99C" box="[539,669,1581,1605]" country="Colombia" name="Prov. Valle" pageId="12" pageNumber="11">Prov. Valle</location>
,
<locationDeviation id="0BCB2FC192090D0C16B41C34B159F99F" box="[680,889,1581,1606]" location="Dagua" pageId="12" pageNumber="11">
<quantity id="DD3E9B0692090D0C16B41C34B0C1F99C" box="[680,737,1581,1605]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="11" unit="km" value="5.0">5 km</quantity>
SE of Dagua
</locationDeviation>
,
<date id="6E78102392090D0C17981C34B620F99C" box="[900,1024,1581,1605]" pageId="12" pageNumber="11" value="1977-02-04">
<collectingDate id="7E3CE9CB92090D0C17981C34B620F99C" box="[900,1024,1581,1605]" pageId="12" pageNumber="11" value="1977-02-04">4 Feb 1977</collectingDate>
</date>
,
<collectorName id="B733533592090D0C10161C37B6B0F99C" box="[1034,1168,1581,1605]" pageId="12" pageNumber="11">M.D. Breed</collectorName>
&amp;
<collectorName id="B733533592090D0C14A21C54B34CF9BC" box="[190,364,1613,1637]" pageId="12" pageNumber="11">C.D. Michener</collectorName>
(
<collectionCode id="7CD7AE2692090D0C15611C54B3ECF9BC" box="[381,460,1613,1637]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="12" pageNumber="11" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
; 2⚲⚲,
<materialsCitation id="AAAE3CBE92090D0C16001C54B3B4F95C" collectingDate="1976-10-06" collectionCode="SEMC" collectorName="Breed" country="Colombia" location="Valle" municipality="Loboguerrera" pageId="12" pageNumber="11" specimenCount="1" typeStatus="paratype">
<collectingCountry id="62D1767392090D0C16001C54B0B2F9BC" box="[540,658,1613,1637]" name="Colombia" pageId="12" pageNumber="11">Colombia</collectingCountry>
:
<location id="1F19603892090D0C16831C54B0FCF9BC" LSID="urn:lsid:plazi:treatment:926F87F592020D0D15AD1EB8B1DEFD80:1F19603892090D0C16831C54B0FCF9BC" box="[671,732,1613,1637]" country="Colombia" municipality="Loboguerrera" name="Valle" pageId="12" pageNumber="11">Valle</location>
, near
<collectingMunicipality id="FA1DAC9992090D0C173A1C54B1EAF9BC" box="[806,970,1613,1637]" pageId="12" pageNumber="11">Loboguerrera</collectingMunicipality>
,
<date id="6E78102392090D0C17C41C54B679F9BC" box="[984,1113,1613,1637]" pageId="12" pageNumber="11" value="1976-10-06">
<collectingDate id="7E3CE9CB92090D0C17C41C54B679F9BC" box="[984,1113,1613,1637]" pageId="12" pageNumber="11" value="1976-10-06">6 Oct 1976</collectingDate>
</date>
, Mi- chael
<collectorName id="B733533592090D0C14FB1C74B30CF95C" box="[231,300,1645,1669]" pageId="12" pageNumber="11">Breed</collectorName>
(
<collectionCode id="7CD7AE2692090D0C15201C74B3ABF95C" box="[316,395,1645,1669]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="12" pageNumber="11" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
; 14⚲⚲,
<materialsCitation id="AAAE3CBE92090D0C15F51C74B078F97C" collectingDate="1977-02-04" collectionCode="SEMC" collectorName="Breed &amp; C. D. Michener" country="Colombia" elevation="740" location="Prov. Valle" municipality="Loboguerrera" pageId="12" pageNumber="11" specimenCount="1" typeStatus="paratype">
<collectingCountry id="62D1767392090D0C15F51C74B07FF95C" box="[489,607,1645,1669]" name="Colombia" pageId="12" pageNumber="11">Colombia</collectingCountry>
:
<location id="1F19603892090D0C16701C77B0D1F95C" LSID="urn:lsid:plazi:treatment:926F87F592020D0D15AD1EB8B1DEFD80:1F19603892090D0C16701C77B0D1F95C" box="[620,753,1645,1669]" country="Colombia" municipality="Loboguerrera" name="Prov. Valle" pageId="12" pageNumber="11">Prov. Valle</location>
,
<collectingMunicipality id="FA1DAC9992090D0C16E31C74B183F95C" box="[767,931,1645,1669]" pageId="12" pageNumber="11">Loboguerrera</collectingMunicipality>
,
<quantity id="DD3E9B0692090D0C17AD1C74B1DCF95C" box="[945,1020,1645,1669]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="12" pageNumber="11" unit="m" value="740.0">
<elevation id="91EBD1D092090D0C17AD1C74B1DCF95C" box="[945,1020,1645,1669]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="12" pageNumber="11" unit="m" value="740.0">740 m</elevation>
</quantity>
,
<date id="6E78102392090D0C10161C74B6ACF95C" box="[1034,1164,1645,1669]" pageId="12" pageNumber="11" value="1977-02-04">
<collectingDate id="7E3CE9CB92090D0C10161C74B6ACF95C" box="[1034,1164,1645,1669]" pageId="12" pageNumber="11" value="1977-02-04">4 Feb 1977</collectingDate>
</date>
, M.D.
<collectorName id="B733533592090D0C14FE1C94B307F97C" box="[226,295,1677,1701]" pageId="12" pageNumber="11">Breed</collectorName>
&amp;
<collectorName id="B733533592090D0C15541C94B3D3F97C" box="[328,499,1677,1701]" pageId="12" pageNumber="11">C.D. Michener</collectorName>
(
<collectionCode id="7CD7AE2692090D0C161D1C94B070F97C" box="[513,592,1677,1701]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="12" pageNumber="11" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
; 7⚲⚲,
<materialsCitation id="AAAE3CBE92090D0C16861C94B084F91C" collectingDate="1977-02-16" collectionCode="SEMC" collectorName="Breed &amp; C. D. Michener" country="Colombia" elevation="1000" location="Prov. Valle" municipality="Cali" pageId="12" pageNumber="11" specimenCount="1" typeStatus="paratype">
<collectingCountry id="62D1767392090D0C16861C94B130F97C" box="[666,784,1677,1701]" name="Colombia" pageId="12" pageNumber="11">Colombia</collectingCountry>
:
<location id="1F19603892090D0C17061C97B1BDF97C" LSID="urn:lsid:plazi:treatment:926F87F592020D0D15AD1EB8B1DEFD80:1F19603892090D0C17061C97B1BDF97C" box="[794,925,1677,1701]" country="Colombia" municipality="Cali" name="Prov. Valle" pageId="12" pageNumber="11">Prov. Valle</location>
,
<collectingMunicipality id="FA1DAC9992090D0C17B41C94B1FAF97C" box="[936,986,1677,1701]" pageId="12" pageNumber="11">Cali</collectingMunicipality>
,
<quantity id="DD3E9B0692090D0C17F81C94B61AF97C" box="[996,1082,1677,1701]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="11" unit="m" value="1000.0">
<elevation id="91EBD1D092090D0C17F81C94B61AF97C" box="[996,1082,1677,1701]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="11" unit="m" value="1000.0">1000 m</elevation>
</quantity>
,
<date id="6E78102392090D0C10591C94B2F8F91C" pageId="12" pageNumber="11" value="1977-02-16">
<collectingDate id="7E3CE9CB92090D0C10591C94B2F8F91C" pageId="12" pageNumber="11" value="1977-02-16">16 Feb 1977</collectingDate>
</date>
, M.D.
<collectorName id="B733533592090D0C153B1CB4B34CF91C" box="[295,364,1709,1733]" pageId="12" pageNumber="11">Breed</collectorName>
&amp;
<collectorName id="B733533592090D0C158C1CB4B01DF91C" box="[400,573,1709,1733]" pageId="12" pageNumber="11">C.D. Michener</collectorName>
(
<collectionCode id="7CD7AE2692090D0C16511CB4B0BCF91C" box="[589,668,1709,1733]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="12" pageNumber="11" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
; 2⚲⚲,
<materialsCitation id="AAAE3CBE92090D0C16F51CB4B1BDF93C" collectingDate="1976-09-23" collectionCode="SEMC" collectorName="Bell &amp; Breed &amp; Michener" country="Colombia" elevation="1500" location="Valle" municipality="Lago Calima" pageId="12" pageNumber="11" specimenCount="1" typeStatus="paratype">
<collectingCountry id="62D1767392090D0C16F51CB4B17FF91C" box="[745,863,1709,1733]" name="Colombia" pageId="12" pageNumber="11">Colombia</collectingCountry>
:
<location id="1F19603892090D0C17771CB4B188F91C" LSID="urn:lsid:plazi:treatment:926F87F592020D0D15AD1EB8B1DEFD80:1F19603892090D0C17771CB4B188F91C" box="[875,936,1709,1733]" country="Colombia" municipality="Lago Calima" name="Valle" pageId="12" pageNumber="11">Valle</location>
,
<collectingMunicipality id="FA1DAC9992090D0C17A81CB7B66EF91C" box="[948,1102,1709,1734]" pageId="12" pageNumber="11">Lago Calima</collectingMunicipality>
,
<quantity id="DD3E9B0692090D0C10471CB4B29AF93C" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="11" unit="m" value="1500.0">
<elevation id="91EBD1D092090D0C10471CB4B29AF93C" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="11" unit="m" value="1500.0">1500 m</elevation>
</quantity>
, IX-23-76 [
<date id="6E78102392090D0C152E1CD4B033F93C" box="[306,531,1741,1765]" pageId="12" pageNumber="11" value="1976-09-23">
<collectingDate id="7E3CE9CB92090D0C152E1CD4B033F93C" box="[306,531,1741,1765]" pageId="12" pageNumber="11" value="1976-09-23">23 September 1976</collectingDate>
</date>
],
<collectorName id="B733533592090D0C163F1CD4B072F93C" box="[547,594,1741,1765]" pageId="12" pageNumber="11">Bell</collectorName>
,
<collectorName id="B733533592090D0C16401CD4B083F93C" box="[604,675,1741,1765]" pageId="12" pageNumber="11">Breed</collectorName>
, &amp;
<collectorName id="B733533592090D0C16D41CD4B118F93C" box="[712,824,1741,1765]" pageId="12" pageNumber="11">Michener</collectorName>
(
<collectionCode id="7CD7AE2692090D0C175A1CD4B1B5F93C" box="[838,917,1741,1765]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="12" pageNumber="11" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
; 1⚲,
<materialsCitation id="AAAE3CBE92090D0D17D31CD4B356FF20" collectingDate="1976-09-27" collectionCode="SEMC" collectorName="Bell &amp; Breed &amp; Michener" country="Colombia" elevation="1300" lastPageId="13" lastPageNumber="12" location="Valle" municipality="Buenaventura Rd." pageId="12" pageNumber="11" specimenCount="1" typeStatus="paratype">
<collectingCountry id="62D1767392090D0C17D31CD4B665F93C" box="[975,1093,1741,1765]" name="Colombia" pageId="12" pageNumber="11">Colombia</collectingCountry>
:
<location id="1F19603892090D0C10531CD4B6ACF93C" LSID="urn:lsid:plazi:treatment:926F87F592020D0D15AD1EB8B1DEFD80:1F19603892090D0C10531CD4B6ACF93C" box="[1103,1164,1741,1765]" country="Colombia" municipality="Buenaventura Rd." name="Valle" pageId="12" pageNumber="11">Valle</location>
, km 33, old
<collectingMunicipality id="FA1DAC9992090D0C15371CF7B025F8DC" box="[299,517,1773,1797]" pageId="12" pageNumber="11">Buenaventura Rd.</collectingMunicipality>
,
<quantity id="DD3E9B0692090D0C16081CF4B04DF8DC" box="[532,621,1773,1797]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="12" pageNumber="11" unit="m" value="1300.0">
<elevation id="91EBD1D092090D0C16081CF4B04DF8DC" box="[532,621,1773,1797]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="12" pageNumber="11" unit="m" value="1300.0">1300 m</elevation>
</quantity>
, IX-27-76 [
<date id="6E78102392090D0C16F11CF4B1F5F8DC" box="[749,981,1773,1797]" pageId="12" pageNumber="11" value="1976-09-27">
<collectingDate id="7E3CE9CB92090D0C16F11CF4B1F5F8DC" box="[749,981,1773,1797]" pageId="12" pageNumber="11" value="1976-09-27">27 September 1976</collectingDate>
</date>
],
<collectorName id="B733533592090D0C17F51CF4B638F8DC" box="[1001,1048,1773,1797]" pageId="12" pageNumber="11">Bell</collectorName>
,
<collectorName id="B733533592090D0C103A1CF4B64DF8DC" box="[1062,1133,1773,1797]" pageId="12" pageNumber="11">Breed</collectorName>
, &amp;
<collectorName id="B733533592080D0D14BC1AF8B330FF20" box="[160,272,225,249]" pageId="13" pageNumber="12">Michener</collectorName>
(
<collectionCode id="7CD7AE2692080D0D15031AF8B34EFF20" box="[287,366,225,249]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="13" pageNumber="12" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
; 21⚲⚲,
<materialsCitation id="AAAE3CBE92080D0D15DB1AF8B044FEC0" collectingDate="1976-09-22" collectionCode="SEMC" collectorName="Valle &amp; El Carmen &amp; Bell &amp; Breed &amp; Michener" country="Colombia" elevation="1370" location="Colombia" pageId="13" pageNumber="12" specimenCount="1" typeStatus="paratype">
<collectingCountry id="62D1767392080D0D15DB1AF8B01DFF20" box="[455,573,225,249]" name="Colombia" pageId="13" pageNumber="12">Colombia</collectingCountry>
:
<collectorName id="B733533592080D0D16541AF8B0A5FF20" box="[584,645,225,249]" pageId="13" pageNumber="12">Valle</collectorName>
,
<collectorName id="B733533592080D0D168D1AF8B131FF20" box="[657,785,225,249]" pageId="13" pageNumber="12">El Carmen</collectorName>
,
<quantity id="DD3E9B0692080D0D17001AF8B153FF20" box="[796,883,225,249]" metricMagnitude="3" metricUnit="m" metricValue="1.3699999999999999" pageId="13" pageNumber="12" unit="m" value="1370.0">
<elevation id="91EBD1D092080D0D17001AF8B153FF20" box="[796,883,225,249]" metricMagnitude="3" metricUnit="m" metricValue="1.3699999999999999" pageId="13" pageNumber="12" unit="m" value="1370.0">1370 m</elevation>
</quantity>
, IX-22-76 [
<date id="6E78102392080D0D17F21AF8B2FAFEC0" pageId="13" pageNumber="12" value="1976-09-22">
<collectingDate id="7E3CE9CB92080D0D17F21AF8B2FAFEC0" pageId="13" pageNumber="12" value="1976-09-22">22 September 1976</collectingDate>
</date>
],
<collectorName id="B733533592080D0D14F71B18B33AFEC0" box="[235,282,257,281]" pageId="13" pageNumber="12">Bell</collectorName>
,
<collectorName id="B733533592080D0D15381B18B34BFEC0" box="[292,363,257,281]" pageId="13" pageNumber="12">Breed</collectorName>
, &amp;
<collectorName id="B733533592080D0D158C1B18B020FEC0" box="[400,512,257,281]" pageId="13" pageNumber="12">Michener</collectorName>
(
<collectionCode id="7CD7AE2692080D0D16111B18B07CFEC0" box="[525,604,257,281]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="13" pageNumber="12" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
; 20⚲⚲,
<materialsCitation id="AAAE3CBE92080D0D16AF1B18B1A9FEE0" collectingDate="1977-02-14" collectionCode="SEMC" collectorName="M. D. Breed &amp; C. D. Michener" country="Colombia" elevation="1200" location="Prov. Valle" pageId="13" pageNumber="12" specimenCount="1" typeStatus="paratype">
<collectingCountry id="62D1767392080D0D16AF1B18B109FEC0" box="[691,809,257,281]" name="Colombia" pageId="13" pageNumber="12">Colombia</collectingCountry>
:
<location id="1F19603892080D0D172E1B1BB194FEC0" LSID="urn:lsid:plazi:treatment:926F87F592020D0D15AD1EB8B1DEFD80:1F19603892080D0D172E1B1BB194FEC0" box="[818,948,257,281]" country="Colombia" name="Prov. Valle" pageId="13" pageNumber="12">Prov. Valle</location>
,
<quantity id="DD3E9B0692080D0D17A21B18B625FEC0" box="[958,1029,257,281]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="12" unit="km" value="10.0">10 km</quantity>
SW Jamun- di,
<date id="6E78102392080D0D14DB1B38B375FEE0" box="[199,341,289,313]" pageId="13" pageNumber="12" value="1977-02-14">
<collectingDate id="7E3CE9CB92080D0D14DB1B38B375FEE0" box="[199,341,289,313]" pageId="13" pageNumber="12" value="1977-02-14">14 Feb 1977</collectingDate>
</date>
,
<quantity id="DD3E9B0692080D0D157E1B38B399FEE0" box="[354,441,289,313]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="13" pageNumber="12" unit="m" value="1200.0">
<elevation id="91EBD1D092080D0D157E1B38B399FEE0" box="[354,441,289,313]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="13" pageNumber="12" unit="m" value="1200.0">1200 m</elevation>
</quantity>
,
<collectorName id="B733533592080D0D15DA1B3BB06FFEE0" box="[454,591,289,313]" pageId="13" pageNumber="12">M.D. Breed</collectorName>
&amp;
<collectorName id="B733533592080D0D16681B38B101FEE0" box="[628,801,289,313]" pageId="13" pageNumber="12">C.D. Michener</collectorName>
(
<collectionCode id="7CD7AE2692080D0D172D1B38B1A0FEE0" box="[817,896,289,313]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="13" pageNumber="12" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
; 5⚲⚲,
<materialsCitation id="AAAE3CBE92080D0D17D31B38B65CFE80" collectingDate="1977-02-14" collectionCode="SEMC" collectorName="M. D. Breed &amp; C. D. Michener" country="Colombia" elevation="1100" location="Prov. Valle" pageId="13" pageNumber="12" specimenCount="1" typeStatus="paratype">
<collectingCountry id="62D1767392080D0D17D31B38B665FEE0" box="[975,1093,289,313]" name="Colombia" pageId="13" pageNumber="12">Colombia</collectingCountry>
:
<location id="1F19603892080D0D104D1B3BB2FDFE80" LSID="urn:lsid:plazi:treatment:926F87F592020D0D15AD1EB8B1DEFD80:1F19603892080D0D104D1B3BB2FDFE80" country="Colombia" name="Prov. Valle" pageId="13" pageNumber="12">Prov. Valle</location>
,
<locationDeviation id="0BCB2FC192080D0D14F41B5BB3E0FE80" box="[232,448,321,345]" location="Jamundi" pageId="13" pageNumber="12">
<quantity id="DD3E9B0692080D0D14F41B5BB302FE80" box="[232,290,321,345]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="13" pageNumber="12" unit="km" value="7.0">7 km</quantity>
SW Jamundi
</locationDeviation>
,
<date id="6E78102392080D0D15D71B58B076FE80" box="[459,598,321,345]" pageId="13" pageNumber="12" value="1977-02-14">
<collectingDate id="7E3CE9CB92080D0D15D71B58B076FE80" box="[459,598,321,345]" pageId="13" pageNumber="12" value="1977-02-14">14 Feb 1977</collectingDate>
</date>
,
<quantity id="DD3E9B0692080D0D167C1B58B096FE80" box="[608,694,321,345]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="13" pageNumber="12" unit="m" value="1100.0">
<elevation id="91EBD1D092080D0D167C1B58B096FE80" box="[608,694,321,345]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="13" pageNumber="12" unit="m" value="1100.0">1100 m</elevation>
</quantity>
,
<collectorName id="B733533592080D0D16DD1B5BB168FE80" box="[705,840,321,345]" pageId="13" pageNumber="12">M.D. Breed</collectorName>
&amp;
<collectorName id="B733533592080D0D17761B58B635FE80" box="[874,1045,321,345]" pageId="13" pageNumber="12">C.D. Michener</collectorName>
(
<collectionCode id="7CD7AE2692080D0D10381B58B654FE80" box="[1060,1140,321,345]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="13" pageNumber="12" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="52DC656892080D0D14CC1B78B1DEFD80" pageId="13" pageNumber="12" type="etymology">
<paragraph id="1A7936E392080D0D14CC1B78B1DEFD80" blockId="13.[160,1168,225,601]" pageId="13" pageNumber="12">
ETYMOLOGY: This species has been frequently referenced in the literature but has never been described formally. In order to maintain some continuity the original name is maintained. The specific epithet was first proposed by Herbert F. Schwarz (1883– 1960) for an unnamed member of the Eder family. Henry J. (1870–1949) and Walter Eder (
<emphasis id="28B2EAF192080D0D14F41BF8B30CFE20" box="[232,300,481,505]" italics="true" pageId="13" pageNumber="12">floruit</emphasis>
1927), son and grandson of the pioneering sugar industrialist James Mar- tin Eder (1838–1921), provided logistical assistance to Schwarz during his 1935 collect- ing visit to Cali (
<bibRefCitation id="7E574B1292080D0D157E1838B02DFDE0" author="Schwarz, H. P." box="[354,525,545,569]" pageId="13" pageNumber="12" pagination="1 - 546" refId="ref22362" refString="Schwarz, H. P. 1948. Stingless bees (Meliponidae) of the Western Hemisphere. Lestrimelita and the following subgenera of Trigona: Trigona, Paratrigona, Schwarziana, Parapartamona, Cephalotrigona, Oxytrigona, Scaura, and Mourella. Bulletin of the American Museum of Natural History 90: i - xviii, 1 - 546, + 8 pls." type="book chapter" year="1948">Schwarz, 1948</bibRefCitation>
). Herein, I take the presumption of proposing that the specific epithet honors the patriarch of the family in
<collectingCountry id="62D1767392080D0D17141858B15EFD80" box="[776,894,577,601]" name="Colombia" pageId="13" pageNumber="12">Colombia</collectingCountry>
, J.M. Eder.
</paragraph>
</subSubSection>
</treatment>
</document>
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