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<document id="7B6E697D150B63CA67F21CDE7562F829" ID-CLB-Dataset="35667" ID-DOI="http://dx.doi.org/10.3897/zookeys.599.8168" ID-GBIF-Dataset="644955ba-354e-4fe4-a132-54aa98d8d11b" ID-PMC="PMC4926696" ID-Pensoft-Pub="1313-2970-599-1" ID-PubMed="27408595" ID-ZooBank="8BEE967F7C6946928210A440AD8E2018" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1313-2970-599-1" ModsDocOrigin="ZooKeys 599" ModsDocTitle="Taxonomic revision of the rock-dwelling door snail genus Montenegrina Boettger, 1877 (Mollusca, Gastropoda, Clausiliidae)" checkinTime="1466146683286" checkinUser="pensoft" docAuthor="Feher, Zoltan &amp; Szekeres, Miklos" docDate="2016" docId="666597C7554955B0F099E0BCC6E314B5" docLanguage="en" docName="ZooKeys 599: 1-137" docOrigin="ZooKeys 599" docSource="http://dx.doi.org/10.3897/zookeys.599.8168" docTitle="Montenegrina soosi Eross &amp; Szekeres 2006" docType="treatment" docVersion="7" lastPageNumber="93" masterDocId="FFFFFFBDDF41B16FB84A7D17FFFD0B15" masterDocTitle="Taxonomic revision of the rock-dwelling door snail genus Montenegrina Boettger, 1877 (Mollusca, Gastropoda, Clausiliidae)" masterLastPageNumber="137" masterPageNumber="1" pageNumber="93" updateTime="1701376832386" updateUser="ExternalLinkService">
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<mods:title id="F153A90F45823D845FC85FD8565435D1">Taxonomic revision of the rock-dwelling door snail genus Montenegrina Boettger, 1877 (Mollusca, Gastropoda, Clausiliidae)</mods:title>
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<mods:namePart id="D8D2C85F86694E8A15C7978BF83CA7B8">Feher, Zoltan</mods:namePart>
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<mods:namePart id="058116161612396451E9880133D8CD7A">Szekeres, Miklos</mods:namePart>
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<mods:date id="D151B6D1665D87C77D10361CCA64D97D">2016</mods:date>
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<mods:classification id="3226ACA9B3E418ED757E49AEDEA04191">journal article</mods:classification>
<mods:identifier id="BB9EC616FF962FB5519BD410EC623E50" type="DOI">http://dx.doi.org/10.3897/zookeys.599.8168</mods:identifier>
<mods:identifier id="F9C9C25CE9BDA7C0F6BF2901B3E5AFA5" type="Pensoft-Pub">1313-2970-599-1</mods:identifier>
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<treatment id="666597C7554955B0F099E0BCC6E314B5" ID-GBIF-Taxon="127862034" LSID="urn:lsid:plazi:treatment:666597C7554955B0F099E0BCC6E314B5" httpUri="http://treatment.plazi.org/id/666597C7554955B0F099E0BCC6E314B5" lastPageNumber="93" pageId="92" pageNumber="93">
<subSubSection id="C87585F0751FF2BA9FEEDB256FBCC83C" pageId="92" pageNumber="93" type="multiple">
<paragraph id="CC9AD3C850E63375DE644CF9FD6AC7B5" pageId="92" pageNumber="93">Taxon classification Animalia Stylommatophora Clausiliidae</paragraph>
</subSubSection>
<subSubSection id="5B233AB8D57C3F131CC68D7AB56D4E6C" pageId="92" pageNumber="93" type="nomenclature">
<paragraph id="0D1B44BAF6E60ECF9F5B6A3C5C936AC9" pageId="92" pageNumber="93">
<taxonomicName id="BAB770EEEC352049535E03E3C0D17FA7" ID-CoL="73Q83" authority="Eross &amp; Szekeres, 2006" authorityName="Eross &amp; Szekeres" authorityYear="2006" class="Gastropoda" family="Clausiliidae" genus="Montenegrina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Montenegrina soosi" order="Stylommatophora" pageId="92" pageNumber="93" phylum="Mollusca" rank="species" species="soosi">
Montenegrina soosi
<normalizedToken id="860279CC10835BCE241E4275D82158ED" originalValue="Erőss">Eross</normalizedToken>
&amp; Szekeres, 2006
</taxonomicName>
Fig. 28H
</paragraph>
</subSubSection>
<subSubSection id="5978B21038DE03F0D5BDEFEE7730237E" pageId="92" pageNumber="93" type="reference_group">
<paragraph id="1469E37CD05E83244ECE52397F09177E" pageId="92" pageNumber="93">
<taxonomicName id="A679B998F4D0EBEBD3DCC6919DCD8B59" class="Gastropoda" family="Clausiliidae" genus="Montenegrina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Montenegrina janinensis" order="Stylommatophora" pageId="92" pageNumber="93" phylum="Mollusca" rank="subSpecies" species="janinensis" subSpecies="soosi">Montenegrina janinensis soosi</taxonomicName>
<normalizedToken id="AB93CB9FC91D2177F08C64D7CCB42912" originalValue="Erőss">Eross</normalizedToken>
&amp; Szekeres, 2006 in
<bibRefCitation id="B9E75A23D9A6E77CBECB0934CA6749AF" author="Eross, Z" journalOrPublisher="Annalen des Naturhistorischen Museums in Wien" pageId="113" pageNumber="114" pagination="181 - 208" title="New taxa of Montenegrina O. Boettger, 1877 (Gastropoda: Pulmonata: Clausiliidae) from Albania and the neighbouring regions." volume="107 B" year="2006">
<normalizedToken id="73C545F9F5EAD25207B36F54EC8485CA" originalValue="Erőss">Eross</normalizedToken>
et al. 2006
</bibRefCitation>
: 197, fig. 18. -
<bibRefCitation id="2F22A3FB3A4B4226B87B7CE5879D5ACF" author="Nordsieck, H" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="114" pageNumber="115" pagination="71 - 87" title="Revisory notes on the genus Montenegrina O. Boettger 1877 and description of new taxa (Gastropoda: Stylommatophora: Clausiliidae)." url="10.1127/arch.moll/0003-9284/138/071-087" volume="138" year="2009">Nordsieck 2009</bibRefCitation>
: 75.
</paragraph>
</subSubSection>
<subSubSection id="DC5F2BF41628223F27660662A1D6AF9C" pageId="92" pageNumber="93" type="diagnosis">
<paragraph id="BCAC71D11E2EBC1E98EE09CE3F8EAB80" pageId="92" pageNumber="93">Diagnosis.</paragraph>
<paragraph id="5DC9DDB7CC42E00B4AD40450BCBAE231" pageId="92" pageNumber="93">Shell small, light yellowish corneous. All whorls smooth. Neck inflexed, densely striate-costate. Basal and peripheral crests well recognizable. Peristome attached, ovoid to angular, with simple margin. Lamellae superior and spiralis long overlap. In front view lamella inferior moderately emerged, medium-bent subcolumellaris visible. Lunella broad, dorsolateral-lateral to lateral, mostly not fused to the basalis. Subclaustralis residual, sulcalis strong. Plica superior fused to the principalis. Anterior plica superior long, mostly separate from the lunella complex. Clausilium plate not visible through the aperture.</paragraph>
</subSubSection>
<subSubSection id="478C6183039C36D18650C2298F6C3A9A" pageId="92" pageNumber="93" type="description">
<paragraph id="55E560D6B8F5FC392C3B7DF8DACBDA30" pageId="92" pageNumber="93">Dimensions</paragraph>
<paragraph id="D271F4D73F809C89DF4D9E35ED5DDDCA" pageId="92" pageNumber="93">(in mm). Hs: 11.4-14.6 (holotype 14.6), Ws: 2.7-3.2 (holotype 3.2).</paragraph>
</subSubSection>
<subSubSection id="5AA2E0A617C8D9E2796C05DE27A624DD" pageId="92" pageNumber="93" type="type locality">
<paragraph id="DF7BCB845D535EA7B9162D5A0818B435" pageId="92" pageNumber="93">Type locality.</paragraph>
<paragraph id="4BDF95B33021E1C853D8789359617526" pageId="92" pageNumber="93">
Albania,
<normalizedToken id="2663C5B1269282C0A182FC66DFDF3AEF" originalValue="Kukës">Kukes</normalizedToken>
District, between Kolesjan and Resk, 1 km N of the Ploshtan junction, along the
<normalizedToken id="477C158DB30D48A60E476BA004CD3E3A" originalValue="Kukës">Kukes</normalizedToken>
to Peshkopi road, 750 m,
<geoCoordinate id="7209C30D9F67535234431C4B1F8989B8" direction="north" orientation="latitude" precision="5" value="41.9598">41.9598°N</geoCoordinate>
,
<geoCoordinate id="BE1E9C5B5EEC62A8E5586AB9BCCE8847" direction="east" orientation="longitude" precision="5" value="20.3938">20.3938°E</geoCoordinate>
.
</paragraph>
</subSubSection>
<subSubSection id="841C0609979A3D411E913C46242B11D1" pageId="92" pageNumber="93" type="type material">
<paragraph id="C0055F3D019FA87BFE5159B0B52D219E" pageId="92" pageNumber="93">Type material.</paragraph>
<paragraph id="EE3F327FA9604E22D13D0D4A82E1A0A6" pageId="92" pageNumber="93">Type locality, leg. ZE, ZF, JK, DM, 25.vi.2003, holotype (HNHM 94868), paratypes (NHMUK 20050224, HNC 63188, HNHM 94869/23, NHMW 103285, RMNH 100317, SMF 328089).</paragraph>
</subSubSection>
<subSubSection id="6DF9DA894F89D714D50F3BE8EAA82431" pageId="92" pageNumber="93" type="materials_examined">
<paragraph id="80453101B28E63A75013D7A254ABCE93" pageId="92" pageNumber="93">Other material.</paragraph>
<paragraph id="E872EA4206F8B9AC5326AE5FFE662181" pageId="92" pageNumber="93">
Type locality, leg. TD, ZE, ZF, DM, 9.x.2005 (HNHM 96825); 1.8 km S of the Kolesjan junction,
<geoCoordinate id="D528AD88026EE392A02B0D9599F8AD2D" direction="north" orientation="latitude" precision="5" value="41.9613">41.9613°N</geoCoordinate>
,
<geoCoordinate id="263164785502378DABA44A2801F9F26E" direction="east" orientation="longitude" precision="5" value="20.3955">20.3955°E</geoCoordinate>
, leg. AR, NR, PR, ix.2013 (NHMW 110430/MN/0112); 7.4 km S of the Kolesjan junction,
<geoCoordinate id="BD81D67CE11C7ABBA224326DC2D0054A" direction="north" orientation="latitude" precision="5" value="41.926">41.9260°N</geoCoordinate>
,
<geoCoordinate id="A45684243A7E1629FB665814DA3D4154" direction="east" orientation="longitude" precision="5" value="20.3872">20.3872°E</geoCoordinate>
, leg. AR, NR, PR, ix.2013 (NHMW 110430/MN/0113); at the mouth of the
<normalizedToken id="C2AC997AA4FFF03A2E0D3E2F92FE453D" originalValue="Përroi">Perroi</normalizedToken>
i
<normalizedToken id="175B379305464C0C1DE011A9A4D0B123" originalValue="Bushtricës">Bushtrices</normalizedToken>
, 300 m,
<geoCoordinate id="F037E8B9DCF8A50324878DBAFDD1E903" direction="north" orientation="latitude" precision="5" value="41.9352">41.9352°N</geoCoordinate>
,
<geoCoordinate id="8EE42A787CFDD1B9D312AB5E7B49E0B8" direction="east" orientation="longitude" precision="5" value="20.3632">20.3632°E</geoCoordinate>
, leg. TD, ZE, ZF, DM, 9.x.2005 (HNHM 99637); same locality, leg. ZE, ZF, JG, 2.vii.2015 (NHMW 110430/MN/0116); 1.3 km S of the mouth of the
<normalizedToken id="C9F92899DDC41FA9F4D361E90051B54A" originalValue="Përroi">Perroi</normalizedToken>
i
<normalizedToken id="D5C77105ADCE73A498F55263B77D8527" originalValue="Bushtricës">Bushtrices</normalizedToken>
, 400 m,
<geoCoordinate id="1BCA8074E70B102380258AFD2E3E7132" direction="north" orientation="latitude" precision="5" value="41.9254">41.9254°N</geoCoordinate>
,
<geoCoordinate id="4C142335CAEA598F38AFB26B8835D679" direction="east" orientation="longitude" precision="5" value="20.3569">20.3569°E</geoCoordinate>
, leg. ZE, ZF, JG, 2.vii.2015 (NHMW 110430/MN/0115); bridge of the Lumi i Drinit te Zi (between
<normalizedToken id="602E07CE257C3E97F453A79F95D6019F" originalValue="Skavicë">Skavice</normalizedToken>
and Tej-Drinit), 320 m,
<geoCoordinate id="BE52DD0B0FBA4FC62FBB5F5DE7F1A041" direction="north" orientation="latitude" precision="5" value="41.9237">41.9237°N</geoCoordinate>
,
<geoCoordinate id="3AAF8F0B8DD96C05BFD3EAA8BD2528BA" direction="east" orientation="longitude" precision="5" value="20.3539">20.3539°E</geoCoordinate>
, leg. TD, ZE, ZF, DM, 9.x.2005 (HNHM 99644); bridge of the Lumi i Drinit te Zi (between
<normalizedToken id="3AB47FEA84BD8811B71865F65848AACF" originalValue="Draj-Reç">Draj-Rec</normalizedToken>
and
<normalizedToken id="007D497B3B526A7A13B4CA79EC60DB50" originalValue="Zall-Reç">Zall-Rec</normalizedToken>
), 310 m,
<geoCoordinate id="B0BD3FD14B9C1EEDC873B7B13577A6D8" direction="north" orientation="latitude" precision="5" value="41.888">41.8880°N</geoCoordinate>
,
<geoCoordinate id="7FD388E499DC0660949825176FD7A110" direction="east" orientation="longitude" precision="5" value="20.3165">20.3165°E</geoCoordinate>
, leg. TD, ZE, ZF, DM, 9.x.2005 (HNHM 99640); same locality, leg. ZE, ZF, JG, 2.vii.2015 (NHMW 110430/MN/0114); Tej-Drinit, 4.5 km from the bridge of the Lumi i Drinit te Zi, 620 m,
<geoCoordinate id="D70477DCAD589E5CB75A338A732AC583" direction="north" orientation="latitude" precision="5" value="41.9423">41.9423°N</geoCoordinate>
,
<geoCoordinate id="39A6383091E511BE8B9CD7DB65476CA8" direction="east" orientation="longitude" precision="5" value="20.3593">20.3593°E</geoCoordinate>
, leg. TD, ZE, ZF, DM, 9.x.2005 (HNHM 99641); 2.5 km N of
<normalizedToken id="DC64DFB3A68B98F02223930F33E491E4" originalValue="Bushtricë">Bushtrice</normalizedToken>
, gorge at the Ura e
<normalizedToken id="44A6786F09E14E14D73ECAC809AB4EA4" originalValue="Lapavës">Lapaves</normalizedToken>
, 600 m,
<geoCoordinate id="756B1AC7947D100543D92D3DBF4C4B5C" direction="north" orientation="latitude" precision="5" value="41.8949">41.8949°N</geoCoordinate>
,
<geoCoordinate id="9714D02C8118CCFFB6F5FCBE7C431974" direction="east" orientation="longitude" precision="5" value="20.4169">20.4169°E</geoCoordinate>
, leg. ZE, ZF, JK, DM, 25.vi.2003 (HNHM 94870); same locality, leg. LD, ZE, ZF, AH, DM, 26.vi.2007 (HNHM 99642); 1 km S of Resk along the
<normalizedToken id="39210A811F68C8C8BD2C8F6584DDC896" originalValue="Kukës">Kukes</normalizedToken>
to Peshkopi road, 690 m,
<geoCoordinate id="AAB3EA84515E8F7057334EEE7F1F31B8" direction="north" orientation="latitude" precision="5" value="41.926">41.9260°N</geoCoordinate>
,
<geoCoordinate id="5485FC2E292B73AD0030FCBEAD097057" direction="east" orientation="longitude" precision="5" value="20.3871">20.3871°E</geoCoordinate>
, leg. LD, ZE, ZF, AH, DM, 26.vi.2007 (HNHM 99643);
<normalizedToken id="D12CBFB1DD08BBE6D9D7850EB77980BF" originalValue="Arrën">Arren</normalizedToken>
, 1150 m,
<geoCoordinate id="8F5BE820BDE8472688FD74967902D4CE" direction="north" orientation="latitude" precision="5" value="41.9242">41.9242°N</geoCoordinate>
,
<geoCoordinate id="1D9B38EE85BBA1E947472896DDDDAC47" direction="east" orientation="longitude" precision="5" value="20.2813">20.2813°E</geoCoordinate>
, leg. TD, ZE, ZF, DM, 9.x.2005 (HNHM 99639); Gurri i
<normalizedToken id="25F827734B9955EB7085ED47653277A7" originalValue="Shkallës">Shkalles</normalizedToken>
, E of
<normalizedToken id="E693BF4B8CC7C5938B78960B6FA88A4E" originalValue="Arrën">Arren</normalizedToken>
, 1300 m,
<geoCoordinate id="79D693914B82A319F0955B1403874962" direction="north" orientation="latitude" precision="5" value="41.9188">41.9188°N</geoCoordinate>
,
<geoCoordinate id="53EB9AD204ABF554869D3E470CA78E80" direction="east" orientation="longitude" precision="5" value="20.2693">20.2693°E</geoCoordinate>
, leg. TD, ZE, ZF, DM, 9.x.2005 (HNHM 99638).
</paragraph>
</subSubSection>
<subSubSection id="1F4FD63ED2926788ED0964D6A368F79C" pageId="92" pageNumber="93" type="distribution">
<paragraph id="B5C9794F17DDE526DEB1B6495D2A3E0B" pageId="92" pageNumber="93">Distribution.</paragraph>
<paragraph id="493193A9CE199A891A236474D654C5D0" pageId="92" pageNumber="93">
This taxon is distributed in the catchment area of the Drin i Zi (Black Drin) between the
<normalizedToken id="2272A5EE6CD160687AAF1626F4EB3B97" originalValue="Mirditë">Mirdite</normalizedToken>
and Mt. Gjalica e
<normalizedToken id="17998E0F1D19D04DB4AC0BC25D55EF2C" originalValue="Lumës">Lumes</normalizedToken>
(Fig. 32).
</paragraph>
<caption id="FEB21A3B3E317A6F9ACA79644C6E9076" pageId="92" pageNumber="93">
<paragraph id="CA14D0CFBC4F0E7FC1A44EB80025C9FC" pageId="92" pageNumber="93">
Figure 32. Distribution of
<taxonomicName id="EDECBBC2A938A6829D98BC9517730EE9" class="Gastropoda" family="Clausiliidae" genus="Montenegrina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Montenegrina soosi" order="Stylommatophora" pageId="92" pageNumber="93" phylum="Mollusca" rank="species" species="soosi">Montenegrina soosi</taxonomicName>
and
<taxonomicName id="940C3F6EA97EA994E77DD4B5E058C680" class="Gastropoda" family="Clausiliidae" genus="Montenegrina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Montenegrina sporadica" order="Stylommatophora" pageId="92" pageNumber="93" phylum="Mollusca" rank="species" species="sporadica">Montenegrina sporadica</taxonomicName>
.
<taxonomicName id="3588B5BB061B2342279958381190CD92" class="Gastropoda" family="Clausiliidae" genus="Montenegrina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Montenegrina soosi" order="Stylommatophora" pageId="92" pageNumber="93" phylum="Mollusca" rank="species" species="soosi">Montenegrina soosi</taxonomicName>
(circle);
<taxonomicName id="2C58B32FC358C2C6109911B25EE2A8E5" class="Gastropoda" family="Clausiliidae" genus="Montenegrina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Montenegrina sporadica" order="Stylommatophora" pageId="92" pageNumber="93" phylum="Mollusca" rank="subSpecies" species="sporadica" subSpecies="sporadica">Montenegrina sporadica sporadica</taxonomicName>
(diamond);
<taxonomicName id="E4738F518D88B6B85A32DADF02E79693" class="Gastropoda" family="Clausiliidae" genus="Montenegrina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Montenegrina sporadica" order="Stylommatophora" pageId="92" pageNumber="93" phylum="Mollusca" rank="subSpecies" species="sporadica" subSpecies="tropojana">Montenegrina sporadica tropojana</taxonomicName>
ssp. n. (empty circle with dot).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="680D5094CF196EC5C2FE8536E7D89EAF" pageId="92" pageNumber="93" type="remarks">
<paragraph id="B23071B2C1D201C40DCC36A96F3D25EA" pageId="92" pageNumber="93">Remarks.</paragraph>
<paragraph id="F57D5FDB428579AB5FAB5153821961DF" pageId="92" pageNumber="93">
As its shell does not reveal a clear affiliation to any other taxa,
<taxonomicName id="E0FE9D676EAF09EB2EE125C650656B63" class="Gastropoda" family="Clausiliidae" genus="Montenegrina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Montenegrina soosi" order="Stylommatophora" pageId="92" pageNumber="93" phylum="Mollusca" rank="species" species="soosi">Montenegrina soosi</taxonomicName>
is tentatively regarded as a distinct species. In the
<normalizedToken id="114F45E7F489ECF1C65C521ED5B9DACA" originalValue="Lapavë">Lapave</normalizedToken>
Gorge it occurs syntopically with the nominotypical form of
<taxonomicName id="CB251F75E0FEAD37C5D20332AC42A5D5" class="Gastropoda" family="Clausiliidae" genus="Montenegrina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Montenegrina skipetarica" order="Stylommatophora" pageId="92" pageNumber="93" phylum="Mollusca" rank="species" species="skipetarica">Montenegrina skipetarica</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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