303 lines
37 KiB
XML
303 lines
37 KiB
XML
<document id="3FECB08C7CE2EA8858B23A836873FB2C" ID-DOI="10.11646/zootaxa.3700.4.1" ID-GBIF-Dataset="e3577bac-9620-4b98-904b-935228c918b9" ID-ISSN="1175-5326" ID-Zenodo-Dep="216107" ID-ZooBank="153B3E22-7C34-474D-9FF8-7B381BE5CDA6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460664072544" checkinUser="plazi" docAuthor="Linkem, Charles W. & Brown, Rafe M." docDate="2013" docId="DE2187C6FFC6FF93FF6AFA946C85457B" docLanguage="en" docName="zt03700p533.pdf" docOrigin="Zootaxa 3700 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Parvoscincus agtorum Linkem & Brown, 2013, sp. nov." docType="treatment" docVersion="9" lastPageNumber="516" masterDocId="2218FFBEFFCBFF9CFFFDFFA2686A4643" masterDocTitle="Systematic revision of the Parvoscincus decipiens (Boulenger, 1894) complex of Philippine forest skinks (Squamata: Scincidae: Lygosominae) with descriptions of seven new species" masterLastPageNumber="533" masterPageNumber="501" pageNumber="514" updateTime="1698347816056" updateUser="plazi">
|
||
<mods:mods id="B72A1891DDB7A24C2C9C4AA8D2CA770C" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="FD8DF99576AC147017E5FDBA8FD8F891">
|
||
<mods:title id="3574839DBD4EBFE9EDD53C68C2921846">Systematic revision of the Parvoscincus decipiens (Boulenger, 1894) complex of Philippine forest skinks (Squamata: Scincidae: Lygosominae) with descriptions of seven new species</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="825D1AC853F02EFB64749EE86CDC9EEA" type="personal">
|
||
<mods:role id="7693ED2E6CBC3600B2A756CF80C8DC2A">
|
||
<mods:roleTerm id="D9B107FD22DA989EC0CC60E12E5F1B47">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="A3372C15A52A57BD07FEB9C03F9A03A8">Linkem, Charles W.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="7899A1B63C4B1BE4B4BC03A12665B897" type="personal">
|
||
<mods:role id="BA7EF1FBEC0148D3E1A3E6803ACCF89A">
|
||
<mods:roleTerm id="3124BC43AE2D486CC761776489C65228">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="B416D4BA0BE884508F2F1EB27F27C0B3">Brown, Rafe M.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="BDA8789B276F56C50D5AC8B5A7B1AA3D">text</mods:typeOfResource>
|
||
<mods:relatedItem id="B6ECABF31FF7F4BD09BDA6B7C25942FC" type="host">
|
||
<mods:titleInfo id="ED0FC3E816A1F41BE4366A6F5A21D1B8">
|
||
<mods:title id="8D55B7D502069A93535FB08D2C382CEA">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="B92212C6EC3992B5B43242B2B02C0DD4">
|
||
<mods:date id="BFC246142D1EFE0AE157F41D32658403">2013</mods:date>
|
||
<mods:detail id="CFE2AD39B1761C4FD1361FCE133CE8FE" type="volume">
|
||
<mods:number id="FEF3CA196512E9354A1C39FEE756311D">3700</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="58DB205564EE233CD4107A1D68295BB6" type="issue">
|
||
<mods:number id="C6B59D66B4402777B0155DB05DC3122C">4</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="FF84AA9ACDB7EF6904FFA7D7ED3B3A89" unit="page">
|
||
<mods:start id="621BDD2A906A4745A2248C4201745FA7">501</mods:start>
|
||
<mods:end id="8B9AB804B63C042CB077E7D8D6075620">533</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="BA421A0A37BD9B496AEFC474067D727E">journal article</mods:classification>
|
||
<mods:identifier id="9086C2CFBFB5FBE57F31E856E63DFA62" type="DOI">10.11646/zootaxa.3700.4.1</mods:identifier>
|
||
<mods:identifier id="9896CF2396FBF7C707F0DD4A4FD037A6" type="GBIF-Dataset">e3577bac-9620-4b98-904b-935228c918b9</mods:identifier>
|
||
<mods:identifier id="FCC62DF23F3BD67C2A78AE918C50B9A7" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="459BD53D49C4E967414698146E3DDE3C" type="Zenodo-Dep">216107</mods:identifier>
|
||
<mods:identifier id="9255EE46D58E17AC25BF04723D02B108" type="ZooBank">153B3E22-7C34-474D-9FF8-7B381BE5CDA6</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="DE2187C6FFC6FF93FF6AFA946C85457B" ID-DOI="http://doi.org/10.5281/zenodo.5617916" ID-GBIF-Taxon="119574986" ID-Zenodo-Dep="5617916" LSID="urn:lsid:plazi:treatment:DE2187C6FFC6FF93FF6AFA946C85457B" httpUri="http://treatment.plazi.org/id/DE2187C6FFC6FF93FF6AFA946C85457B" lastPageId="15" lastPageNumber="516" pageId="13" pageNumber="514">
|
||
<subSubSection id="1E92655BFFC6FF91FF6AFA946B6F43F4" pageId="13" pageNumber="514" type="nomenclature">
|
||
<paragraph id="563736D0FFC6FF91FF6AFA946A614313" blockId="13.[151,523,1334,1394]" box="[151,523,1334,1360]" pageId="13" pageNumber="514">
|
||
<heading id="0D7F81BCFFC6FF91FF6AFA946A614313" bold="true" box="[151,523,1334,1360]" fontSize="11" level="1" pageId="13" pageNumber="514" reason="1">
|
||
<emphasis id="64FCEAC2FFC6FF91FF6AFA946A614313" bold="true" box="[151,523,1334,1360]" pageId="13" pageNumber="514">
|
||
<taxonomicName id="91884D53FFC6FF91FF6AFA9469CF4313" box="[151,421,1334,1360]" class="Reptilia" family="Scincidae" genus="Parvoscincus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="514" phylum="Chordata" rank="species" species="agtorum" status="sp. nov.">
|
||
<emphasis id="64FCEAC2FFC6FF91FF6AFA9469CF4313" bold="true" box="[151,421,1334,1360]" italics="true" pageId="13" pageNumber="514">Parvoscincus agtorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7FCF57B9FFC6FF91FE50FA946A614313" box="[429,523,1334,1360]" pageId="13" pageNumber="514" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="563736D0FFC6FF91FF6AFAFB69F64332" blockId="13.[151,523,1334,1394]" box="[151,412,1369,1394]" pageId="13" pageNumber="514">
|
||
<figureCitation id="CEB32A55FFC6FF91FF6AFAFB688C4331" box="[151,230,1369,1394]" captionStart="FIGURE 2" captionStartId="3.[151,250,1527,1549]" captionTargetBox="[151,1429,193,1505]" captionTargetId="figure@3.[151,1436,193,1506]" captionTargetPageId="3" captionText="FIGURE 2. Partitioned Bayesian majority rule consensus estimate of molecular phylogeny from the ND 2 data of Parvoscincus. Populations within the P. decipiens complex are genetically unique and are diagnosed as new species based on morphological differences. Black circles label nodes with posterior probabilities above 0.95." httpUri="https://zenodo.org/record/216109/files/figure.png" pageId="13" pageNumber="514">Figs. 2</figureCitation>
|
||
; 4J, K, L; 5G, H
|
||
</paragraph>
|
||
<paragraph id="563736D0FFC6FF91FF6AFA036B6F43F4" blockId="13.[151,773,1441,1463]" box="[151,773,1441,1463]" pageId="13" pageNumber="514">
|
||
<taxonomicName id="91884D53FFC6FF91FF6AFA036AAD43F4" authority="Brown & Alcala, 1980: 186" authorityName="Brown & Alcala" authorityPageNumber="186" authorityYear="1980" box="[151,711,1441,1463]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="514" phylum="Chordata" rank="species" species="decipiens">
|
||
<emphasis id="64FCEAC2FFC6FF91FF6AFA0369F543F4" box="[151,415,1441,1463]" italics="true" pageId="13" pageNumber="514">Sphenomorphus decipiens</emphasis>
|
||
: Brown & Alcala, 1980: 186
|
||
</taxonomicName>
|
||
(part)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1E92655BFFC6FF91FF6AFA466A0B402B" pageId="13" pageNumber="514" type="materials_examined">
|
||
<paragraph id="563736D0FFC6FF91FF6AFA466AE64006" blockId="13.[151,1436,1508,2037]" pageId="13" pageNumber="514">
|
||
<emphasis id="64FCEAC2FFC6FF91FF6AFA46696143BE" bold="true" box="[151,267,1508,1533]" pageId="13" pageNumber="514">
|
||
<typeStatus id="89338872FFC6FF91FF6AFA46696C43BE" box="[151,262,1508,1533]" pageId="13" pageNumber="514" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<collectingCountry id="2E9F7640FFC6FF91FEEEFA47693A43BF" box="[275,336,1509,1532]" name="Panama" pageId="13" pageNumber="514">PNM</collectingCountry>
|
||
9782 (formerly KU 323310: RMB Field No. 10732); Female;
|
||
<collectingCountry id="2E9F7640FFC6FF91FBE1FA466CF743BE" box="[1052,1181,1508,1533]" name="Philippines" pageId="13" pageNumber="514">Philippines</collectingCountry>
|
||
, Luzon Island, Aurora Province, Municipality of San Luis, Barangay Lipimental, N: 15.65366º, E: 121.50734º, elevation
|
||
<quantity id="91709B35FFC6FF91FB1FF9A56D43405C" box="[1250,1321,1543,1568]" metricMagnitude="2" metricUnit="m" metricValue="5.4" pageId="13" pageNumber="514" unit="m" value="540.0">540 m</quantity>
|
||
above sea level, collected
|
||
<date id="22361010FFC6FF91FEBAF98E69CA4007" box="[327,416,1580,1604]" pageId="13" pageNumber="514" value="1923-06">23 June</date>
|
||
, 2009 by C. D. Siler.
|
||
</paragraph>
|
||
<paragraph id="563736D0FFC6FF91FF3AF9ED6A0B402B" blockId="13.[151,1436,1508,2037]" box="[199,609,1615,1640]" pageId="13" pageNumber="514">
|
||
<emphasis id="64FCEAC2FFC6FF91FF3AF9ED6900402B" bold="true" box="[199,362,1615,1640]" pageId="13" pageNumber="514">Genetic Data.</emphasis>
|
||
GenBank
|
||
<accessionNumber id="49DBAB33FFC6FF91FE1FF9ED6A34402B" box="[482,606,1615,1640]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KF425343" pageId="13" pageNumber="514">KF425343</accessionNumber>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1E92655BFFC6FF92FF3AF9D669434696" lastPageId="14" lastPageNumber="515" pageId="13" pageNumber="514" type="diagnosis">
|
||
<paragraph id="563736D0FFC6FF91FF3AF9D66BDB40BB" blockId="13.[151,1436,1508,2037]" pageId="13" pageNumber="514">
|
||
<emphasis id="64FCEAC2FFC6FF91FF3AF9D6692B40CE" bold="true" box="[199,321,1652,1677]" pageId="13" pageNumber="514">Diagnosis.</emphasis>
|
||
<taxonomicName id="91884D53FFC6FF91FEB5F9D76A2F40CF" box="[328,581,1653,1676]" class="Reptilia" family="Scincidae" genus="Parvoscincus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="514" phylum="Chordata" rank="species" species="agtorum" status="sp. nov.">
|
||
<emphasis id="64FCEAC2FFC6FF91FEB5F9D76A2F40CF" box="[328,581,1653,1676]" italics="true" pageId="13" pageNumber="514">Parvoscincus agtorum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="64FCEAC2FFC6FF91FDB1F9D66ACC40CE" bold="true" box="[588,678,1652,1677]" pageId="13" pageNumber="514">
|
||
<taxonomicNameLabel id="7FCF57B9FFC6FF91FDB1F9D66ACC40CE" box="[588,678,1652,1677]" pageId="13" pageNumber="514" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be identified by the following combination of characters: (1) A small body size (SVL at maturity
|
||
<quantity id="91709B35FFC6FF91FDE6F93A6AEF40F3" box="[539,645,1688,1712]" metricMagnitude="-2" metricUnit="m" metricValue="4.49" pageId="13" pageNumber="514" unit="mm" value="44.9">44.9 mm</quantity>
|
||
); (2) MBSR = 39; (3) PV = 71; (4) dorsal scales non-striated without apical pits; (5) apical pits on forelimbs and hind limbs; (6) four enlarged supraoculars; (7) anterior and posterior loreals undivided laterally; (8) two preoculars; (9) and 17 Toe IV SDL.
|
||
</paragraph>
|
||
<paragraph id="563736D0FFC6FF92FF3AF8A769434696" blockId="13.[151,1436,1508,2037]" lastBlockId="14.[151,1436,151,2024]" lastPageId="14" lastPageNumber="515" pageId="13" pageNumber="514">
|
||
<taxonomicName id="91884D53FFC6FF91FF3AF8A769AF415F" box="[199,453,1797,1820]" class="Reptilia" family="Scincidae" genus="Parvoscincus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="514" phylum="Chordata" rank="species" species="agtorum" status="sp. nov.">
|
||
<emphasis id="64FCEAC2FFC6FF91FF3AF8A769AF415F" box="[199,453,1797,1820]" italics="true" pageId="13" pageNumber="514">Parvoscincus agtorum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="64FCEAC2FFC6FF91FE30F8A66A4D415E" bold="true" box="[461,551,1796,1821]" pageId="13" pageNumber="514">
|
||
<taxonomicNameLabel id="7FCF57B9FFC6FF91FE30F8A66A4D415E" box="[461,551,1796,1821]" pageId="13" pageNumber="514" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is sister to
|
||
<taxonomicName id="91884D53FFC6FF91FD51F8A76B0F415E" box="[684,869,1796,1821]" class="Reptilia" family="Scincidae" genus="Parvoscincus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="514" phylum="Chordata" rank="species" species="arvindiesmosi" status="sp. nov.">
|
||
<emphasis id="64FCEAC2FFC6FF91FD51F8A76B0F415E" box="[684,869,1796,1821]" italics="true" pageId="13" pageNumber="514">P. arvindiesmosi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="64FCEAC2FFC6FF91FC90F8A66BA2415E" bold="true" box="[877,968,1796,1821]" pageId="13" pageNumber="514">
|
||
<taxonomicNameLabel id="7FCF57B9FFC6FF91FC90F8A66BA2415E" box="[877,968,1796,1821]" pageId="13" pageNumber="514" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, and closely related to
|
||
<taxonomicName id="91884D53FFC6FF91FB2CF8A76DFF415F" box="[1233,1429,1797,1820]" class="Reptilia" family="Scincidae" genus="Parvoscincus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="514" phylum="Chordata" rank="species" species="jimmymcguirei" status="sp. nov.">
|
||
<emphasis id="64FCEAC2FFC6FF91FB2CF8A76DFF415F" box="[1233,1429,1797,1820]" italics="true" pageId="13" pageNumber="514">P. jimmymcguirei</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="64FCEAC2FFC6FF91FF6AF885689F4103" bold="true" box="[151,245,1831,1856]" pageId="13" pageNumber="514">
|
||
<taxonomicNameLabel id="7FCF57B9FFC6FF91FF6AF885689F4103" box="[151,245,1831,1856]" pageId="13" pageNumber="514" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="91884D53FFC6FF91FEFAF88B69FB4103" box="[263,401,1831,1856]" class="Reptilia" family="Scincidae" genus="Parvoscincus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="514" phylum="Chordata" rank="species" species="abstrusus" status="sp. nov.">
|
||
<emphasis id="64FCEAC2FFC6FF91FEFAF88B69FB4103" box="[263,401,1831,1856]" italics="true" pageId="13" pageNumber="514">P. abstrusus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="64FCEAC2FFC6FF91FE60F88569904103" bold="true" box="[413,506,1831,1856]" pageId="13" pageNumber="514">
|
||
<taxonomicNameLabel id="7FCF57B9FFC6FF91FE60F88569904103" box="[413,506,1831,1856]" pageId="13" pageNumber="514" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, and
|
||
<taxonomicName id="91884D53FFC6FF91FDBCF88B6AA04103" box="[577,714,1831,1856]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="514" phylum="Chordata" rank="species" sensu="stricto" species="decipiens">
|
||
<emphasis id="64FCEAC2FFC6FF91FDBCF88B6AA04103" box="[577,714,1831,1856]" italics="true" pageId="13" pageNumber="514">P. decipiens</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7FCF57B9FFC6FF91FD28F88A6B0F4103" box="[725,869,1831,1856]" pageId="13" pageNumber="514" sensu="stricto">sensu stricto</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="CEB32A55FFC6FF91FC85F8856BA84103" box="[888,962,1831,1856]" captionStart="FIGURE 2" captionStartId="3.[151,250,1527,1549]" captionTargetBox="[151,1429,193,1505]" captionTargetId="figure@3.[151,1436,193,1506]" captionTargetPageId="3" captionText="FIGURE 2. Partitioned Bayesian majority rule consensus estimate of molecular phylogeny from the ND 2 data of Parvoscincus. Populations within the P. decipiens complex are genetically unique and are diagnosed as new species based on morphological differences. Black circles label nodes with posterior probabilities above 0.95." httpUri="https://zenodo.org/record/216109/files/figure.png" pageId="13" pageNumber="514">Fig. 2</figureCitation>
|
||
).
|
||
<taxonomicName id="91884D53FFC6FF91FC27F88B6CB14103" box="[986,1243,1833,1856]" class="Reptilia" family="Scincidae" genus="Parvoscincus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="514" phylum="Chordata" rank="species" species="agtorum" status="sp. nov.">
|
||
<emphasis id="64FCEAC2FFC6FF91FC27F88B6CB14103" box="[986,1243,1833,1856]" italics="true" pageId="13" pageNumber="514">Parvoscincus agtorum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="64FCEAC2FFC6FF91FB1BF8856D2E4103" bold="true" box="[1254,1348,1831,1856]" pageId="13" pageNumber="514">
|
||
<taxonomicNameLabel id="7FCF57B9FFC6FF91FB1BF8856D2E4103" box="[1254,1348,1831,1856]" pageId="13" pageNumber="514" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished from all these species by the lack of dorsal apical pits (vs. present); greater number of midbody scale rows (39 vs. <37); presence of two preoculars (vs. three).
|
||
<taxonomicName id="91884D53FFC6FF91FCC6F8D36BD241CB" box="[827,952,1905,1928]" class="Reptilia" family="Scincidae" genus="Parvoscincus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="514" phylum="Chordata" rank="species" species="agtorum" status="sp. nov.">
|
||
<emphasis id="64FCEAC2FFC6FF91FCC6F8D36BD241CB" box="[827,952,1905,1928]" italics="true" pageId="13" pageNumber="514">P. agtorum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="64FCEAC2FFC6FF91FC3CF8CD6C7641CB" bold="true" box="[961,1052,1903,1928]" pageId="13" pageNumber="514">
|
||
<taxonomicNameLabel id="7FCF57B9FFC6FF91FC3CF8CD6C7641CB" box="[961,1052,1903,1928]" pageId="13" pageNumber="514" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be further diagnosed from
|
||
<taxonomicName id="91884D53FFC6FF91FA7AF8D3695441EF" class="Reptilia" family="Scincidae" genus="Parvoscincus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="514" phylum="Chordata" rank="species" species="jimmymcguirei" status="sp. nov.">
|
||
<emphasis id="64FCEAC2FFC6FF91FA7AF8D3695441EF" italics="true" pageId="13" pageNumber="514">P. jimmymcguirei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="64FCEAC2FFC6FF91FEB9F83669F741EE" bold="true" box="[324,413,1940,1965]" pageId="13" pageNumber="514">
|
||
<taxonomicNameLabel id="7FCF57B9FFC6FF91FEB9F83669F741EE" box="[324,413,1940,1965]" pageId="13" pageNumber="514" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by the presence of two preoculars (vs. three preoculars); slightly larger body size (44.91 vs. <
|
||
<quantity id="91709B35FFC6FF91FF4DF81A69434193" box="[176,297,1976,2000]" metricMagnitude="-2" metricUnit="m" metricValue="4.412" pageId="13" pageNumber="514" unit="mm" value="44.12">44.12 mm</quantity>
|
||
); dorsal color of
|
||
<emphasis id="64FCEAC2FFC6FF91FE0FF81B6A064193" box="[498,620,1977,2000]" italics="true" pageId="13" pageNumber="514">P. a g t o r u m</emphasis>
|
||
<emphasis id="64FCEAC2FFC6FF91FD84F81A6ABC4192" bold="true" box="[633,726,1976,2001]" pageId="13" pageNumber="514">sp. nov.</emphasis>
|
||
is much darker than
|
||
<taxonomicName id="91884D53FFC6FF91FC32F81B6CFE4193" box="[975,1172,1977,2000]" class="Reptilia" family="Scincidae" genus="Parvoscincus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="514" phylum="Chordata" rank="species" species="jimmymcguirei" status="sp. nov.">
|
||
<emphasis id="64FCEAC2FFC6FF91FC32F81B6CFE4193" box="[975,1172,1977,2000]" italics="true" pageId="13" pageNumber="514">P. jimmymcguirei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="64FCEAC2FFC6FF91FB63F81A6C904192" bold="true" box="[1182,1274,1976,2001]" pageId="13" pageNumber="514">
|
||
<taxonomicNameLabel id="7FCF57B9FFC6FF91FB63F81A6C904192" box="[1182,1274,1976,2001]" pageId="13" pageNumber="514" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and lacks the series of bright spots along the dorsal edge of the dorsolateral band.
|
||
<taxonomicName id="91884D53FFC6FF91FC57F87F6CC341B7" box="[938,1193,2013,2036]" class="Reptilia" family="Scincidae" genus="Parvoscincus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="514" phylum="Chordata" rank="species" species="agtorum" status="sp. nov.">
|
||
<emphasis id="64FCEAC2FFC6FF91FC57F87F6CC341B7" box="[938,1193,2013,2036]" italics="true" pageId="13" pageNumber="514">Parvoscincus agtorum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="64FCEAC2FFC6FF91FB4CF87E6D6741B6" bold="true" box="[1201,1293,2012,2037]" pageId="13" pageNumber="514">
|
||
<taxonomicNameLabel id="7FCF57B9FFC6FF91FB4CF87E6D6741B6" box="[1201,1293,2012,2037]" pageId="13" pageNumber="514" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has a bright white line from the eye to ear (absent in
|
||
<taxonomicName id="91884D53FFC5FF92FD96FF3B6B4546F3" box="[619,815,153,176]" class="Reptilia" family="Scincidae" genus="Parvoscincus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="515" phylum="Chordata" rank="species" species="jimmymcguirei" status="sp. nov.">
|
||
<emphasis id="64FCEAC2FFC5FF92FD96FF3B6B4546F3" box="[619,815,153,176]" italics="true" pageId="14" pageNumber="515">P. jimmymcguirei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="64FCEAC2FFC5FF92FCCAFF356BF946F3" bold="true" box="[823,915,151,176]" pageId="14" pageNumber="515">
|
||
<taxonomicNameLabel id="7FCF57B9FFC5FF92FCCAFF356BF946F3" box="[823,915,151,176]" pageId="14" pageNumber="515" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
) and an incomplete gular collar (vs. speckled gular region)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1E92655BFFC5FF93FF3AFF436CF94793" lastPageId="15" lastPageNumber="516" pageId="14" pageNumber="515" type="description">
|
||
<paragraph id="563736D0FFC5FF92FF3AFF436B454726" blockId="14.[151,1436,151,2024]" pageId="14" pageNumber="515">
|
||
<taxonomicName id="91884D53FFC5FF92FF3AFF4369AE46BB" box="[199,452,225,248]" class="Reptilia" family="Scincidae" genus="Parvoscincus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="515" phylum="Chordata" rank="species" species="agtorum" status="sp. nov.">
|
||
<emphasis id="64FCEAC2FFC5FF92FF3AFF4369AE46BB" box="[199,452,225,248]" italics="true" pageId="14" pageNumber="515">Parvoscincus agtorum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="64FCEAC2FFC5FF92FE36FF7D6A4E46BB" bold="true" box="[459,548,223,248]" pageId="14" pageNumber="515">
|
||
<taxonomicNameLabel id="7FCF57B9FFC5FF92FE36FF7D6A4E46BB" box="[459,548,223,248]" pageId="14" pageNumber="515" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished from
|
||
<taxonomicName id="91884D53FFC5FF92FCA9FF436C6746BB" box="[852,1037,223,248]" class="Reptilia" family="Scincidae" genus="Parvoscincus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="515" phylum="Chordata" rank="species" species="arvindiesmosi" status="sp. nov.">
|
||
<emphasis id="64FCEAC2FFC5FF92FCA9FF436C6746BB" box="[852,1037,223,248]" italics="true" pageId="14" pageNumber="515">P. arvindiesmosi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="64FCEAC2FFC5FF92FBE9FF7D6C0446BB" bold="true" box="[1044,1134,223,248]" pageId="14" pageNumber="515">
|
||
<taxonomicNameLabel id="7FCF57B9FFC5FF92FBE9FF7D6C0446BB" box="[1044,1134,223,248]" pageId="14" pageNumber="515" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by the presence of a broad snout (
|
||
<collectingCountry id="2E9F7640FFC5FF92FF1FFEA6697E475F" box="[226,276,260,284]" name="India" pageId="14" pageNumber="515">IND</collectingCountry>
|
||
/RostL> 0.50 vs. <0.49); larger body size (SVL 44.91 vs. <
|
||
<quantity id="91709B35FFC5FF92FC3EFEA66C53475F" box="[963,1081,260,284]" metricMagnitude="-2" metricUnit="m" metricValue="4.314" pageId="14" pageNumber="515" unit="mm" value="43.14">43.14 mm</quantity>
|
||
); the presence of an incomplete gular collar and brown speckling on throat (vs. white throat); dorsal body color dark brown (vs. dorsal color lighttan with a broad dorsolateral band extending onto dorsum).
|
||
</paragraph>
|
||
<paragraph id="563736D0FFC5FF92FF3AFED36A03447F" blockId="14.[151,1436,151,2024]" pageId="14" pageNumber="515">
|
||
<taxonomicName id="91884D53FFC5FF92FF3AFED369AD47CB" box="[199,455,369,392]" class="Reptilia" family="Scincidae" genus="Parvoscincus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="515" phylum="Chordata" rank="species" species="agtorum" status="sp. nov.">
|
||
<emphasis id="64FCEAC2FFC5FF92FF3AFED369AD47CB" box="[199,455,369,392]" italics="true" pageId="14" pageNumber="515">Parvoscincus agtorum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="64FCEAC2FFC5FF92FE2DFECD6A4747CB" bold="true" box="[464,557,367,392]" pageId="14" pageNumber="515">
|
||
<taxonomicNameLabel id="7FCF57B9FFC5FF92FE2DFECD6A4747CB" box="[464,557,367,392]" pageId="14" pageNumber="515" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished from
|
||
<taxonomicName id="91884D53FFC5FF92FC91FED36B9F47CB" box="[876,1013,367,392]" class="Reptilia" family="Scincidae" genus="Parvoscincus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="515" phylum="Chordata" rank="species" species="abstrusus" status="sp. nov.">
|
||
<emphasis id="64FCEAC2FFC5FF92FC91FED36B9F47CB" box="[876,1013,367,392]" italics="true" pageId="14" pageNumber="515">P. abstrusus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="64FCEAC2FFC5FF92FC02FECD6C3147CB" bold="true" box="[1023,1115,367,392]" pageId="14" pageNumber="515">
|
||
<taxonomicNameLabel id="7FCF57B9FFC5FF92FC02FECD6C3147CB" box="[1023,1115,367,392]" pageId="14" pageNumber="515" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by having a single anterior loreal (vs. laterally divided anterior loreal); the lack of dorsal apical pits (vs. apical pits weak to missing); higher number of paravertebral scales (71 vs. <69); by having a cream throat with brown speckles and a brown incomplete gular collar (vs. throat black in males and white in females); by having a white line from eye to ear that becomes spots posterior to the ear (vs. presence of a white line from eye to forelimb); and by the presence of strong vertebral spots (vs. weak vertebral spots).
|
||
</paragraph>
|
||
<paragraph id="563736D0FFC5FF92FF3AFDEB6BDB44B3" blockId="14.[151,1436,151,2024]" pageId="14" pageNumber="515">
|
||
<taxonomicName id="91884D53FFC5FF92FF3AFDEB69AE4423" box="[199,452,585,608]" class="Reptilia" family="Scincidae" genus="Parvoscincus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="515" phylum="Chordata" rank="species" species="agtorum" status="sp. nov.">
|
||
<emphasis id="64FCEAC2FFC5FF92FF3AFDEB69AE4423" box="[199,452,585,608]" italics="true" pageId="14" pageNumber="515">Parvoscincus agtorum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="64FCEAC2FFC5FF92FE36FDE56A4F4423" bold="true" box="[459,549,583,608]" pageId="14" pageNumber="515">
|
||
<taxonomicNameLabel id="7FCF57B9FFC5FF92FE36FDE56A4F4423" box="[459,549,583,608]" pageId="14" pageNumber="515" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished from
|
||
<taxonomicName id="91884D53FFC5FF92FCABFDEB6BB64423" box="[854,988,583,608]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="515" phylum="Chordata" rank="species" species="decipiens">
|
||
<emphasis id="64FCEAC2FFC5FF92FCABFDEB6BB64423" box="[854,988,583,608]" italics="true" pageId="14" pageNumber="515">P. decipiens</emphasis>
|
||
</taxonomicName>
|
||
by being larger (SVL 44.91 vs. <
|
||
<quantity id="91709B35FFC5FF92FAA0FDE568AD44C7" metricMagnitude="-2" metricUnit="m" metricValue="4.061" pageId="14" pageNumber="515" unit="mm" value="40.61">40.61 mm</quantity>
|
||
); having more paravertebral scales (71 vs. <63); more midbody scale rows (39 vs. 32–34); the presence of forelimb apical pores (vs. weak or missing apical pores); by having a cream colored throat with brown speckling and a brown incomplete collar (vs. throat white with a few brown streaks); by having brown labials (vs. white labials); and by having a dark brown dorsum (vs. light brown dorsum).
|
||
</paragraph>
|
||
<paragraph id="563736D0FFC5FF92FF3AFD5E69054279" blockId="14.[151,1436,151,2024]" pageId="14" pageNumber="515">
|
||
<emphasis id="64FCEAC2FFC5FF92FF3AFD5E69B54556" bold="true" box="[199,479,764,789]" pageId="14" pageNumber="515">
|
||
Description of
|
||
<typeStatus id="89338872FFC5FF92FE84FD5E69B54556" box="[377,479,764,789]" pageId="14" pageNumber="515" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
. A small-sized
|
||
<taxonomicName id="91884D53FFC5FF92FD69FD5F6B414557" box="[660,811,765,788]" class="Reptilia" family="Scincidae" genus="Parvoscincus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="515" phylum="Chordata" rank="genus">
|
||
<emphasis id="64FCEAC2FFC5FF92FD69FD5F6B414557" box="[660,811,765,788]" italics="true" pageId="14" pageNumber="515">Parvoscincus</emphasis>
|
||
</taxonomicName>
|
||
, SVL
|
||
<quantity id="91709B35FFC5FF92FC8AFD5E6BB44557" box="[887,990,764,788]" metricMagnitude="-2" metricUnit="m" metricValue="4.49" pageId="14" pageNumber="515" unit="mm" value="44.9">44.9 mm</quantity>
|
||
, with clawed, pentadactyl limbs. Snout pointed in lateral profile; rostral wide forming an oval dorsal margin with the nasals and frontonasal scale; frontonasal wider than long, in contact with nasals, rostral, anterior loreals, and prefrontal scales; prefrontals in broad medial contact left overlapping right, in contact with anterior and posterior loreals, frontal, frontonasal, and 1st supraciliary; frontal slightly longer than wide, in contact with two supraoculars, posterior apex rounded; four enlarged supraoculars, 1st largest, 2nd widest; frontoparietals fused, in contact with three supraoculars; interparietal arrowhead-shaped with parietal eye in posterior third; parietals in narrow overlap, right overlapping left, in contact with fourth supraocular, postsupraocular, primary and secondary temporal; nuchals same size as dorsals, not obliquely enlarged.
|
||
</paragraph>
|
||
<paragraph id="563736D0FFC5FF92FF3AFBE76BAC435B" blockId="14.[151,1436,151,2024]" pageId="14" pageNumber="515">Nasal pierced in center by large naris, surrounded anteriorly by rostral, dorsally by frontonasal, posteriorly by anterior loreal, and ventrally by 1st supralabial; single anterior loreal, posterior loreal wider than anterior; preoculars two; seven supralabials, 5th widest and under center of eye; supraciliaries 12, anterior four and posterior two larger than rest of series; 15 ciliaries; lower eyelid scaly and transparent, lacking non-scaled “window;” suboculars nine, largest anteriorly; primary temporals three, secondary temporals two, lower overlapping upper; ear large (EarD [1.66]/EyeD [2.34] = 0.71), round, and moderately sunk.</paragraph>
|
||
<paragraph id="563736D0FFC5FF92FF3AFA8169E243EB" blockId="14.[151,1436,151,2024]" pageId="14" pageNumber="515">Infralabials seven, decreasing in size posteriorly in series; mental large, forming a straight suture with a single large postmental and first infralabials; postmental contacts anterior two infralabials; chin scales increasing in number posteriorly (one, three, five) and then blending into size and shape of gular scales; gular scales slightly smaller than ventrals.</paragraph>
|
||
<paragraph id="563736D0FFC5FF92FF3AFA116BB0407B" blockId="14.[151,1436,151,2024]" pageId="14" pageNumber="515">Body non-elongate (AGD [25.3]/SVL [44.9] = 0.56), cylindrical, with 39 equal-sized midbody scales, limbs overlapping when adpressed; lateral body scales with two or three rows of apical pits; paravertebral scales 71, imbricate, without apical pits. Regenerated tail short, shorter than body (TL [37.98]/SVL [44.9] = 0.84) triangular at base, thicker dorsally than ventrally; subcaudal scales nondifferentiated.</paragraph>
|
||
<paragraph id="563736D0FFC5FF92FF3AF9E169E84153" blockId="14.[151,1436,151,2024]" pageId="14" pageNumber="515">Forelimbs smaller than hind limbs (FLL [3.64]/HLL [5.07] = 0.72), pentadactyl; dorsal forelimb scales smaller than body scale, ventral forelimb scales much smaller than ventral scales, dorsal and ventral forelimb scales imbricate with multiple rows of apical pits; multiple rows of dorsal scales on digits. Relative digit length with lamellae (L/R) in parentheses IV(11/12)> III(10/11)> II(9/8)> V(7/7)> I(5/5). Palmar scales irregular, raised, forming ventral protrusions from palmar surface; large set of four scales on distolateral edge of Digit V to the wrist, largest scale at wrist.</paragraph>
|
||
<paragraph id="563736D0FFC5FF92FF3AF8B9690841AB" blockId="14.[151,1436,151,2024]" pageId="14" pageNumber="515">
|
||
<collectingCountry id="2E9F7640FFC5FF92FF3AF8B968954177" box="[199,255,1819,1844]" name="India" pageId="14" pageNumber="515">Hind</collectingCountry>
|
||
limbs small (HLL [5.07]/SVL [44.9] = 0.11), pentadactyl; dorsal and ventral hind limb scales smaller than body scales; dorsal scales bearing apical pits, ventral scales with single row of apical pits; multiple scale rows on dorsal side of digits. Lamellae slightly keeled. Relative digit length with lamellae (L/R) in parentheses: IV(17/ 17)> III(13/13)> V(10/10)> II(9/9)> I(6/6). Plantar scales irregular, slightly raised; four large, ventrally pointed scales along ankle/plantar margin; ventrally raised scales along distolateral edge of Digit V to ankle, increasing in size toward ankle.
|
||
</paragraph>
|
||
<paragraph id="563736D0FFC4FF93FF3AFF356B1B4696" blockId="15.[151,1437,151,824]" pageId="15" pageNumber="516">Precloacal region with series of enlarged scales between pelvic region and cloaca, more elongate than ventral scales; medial precloacal scales larger, overlapping lateral scales.</paragraph>
|
||
<paragraph id="563736D0FFC4FF93FF3AFF7D6CF94793" blockId="15.[151,1437,151,824]" pageId="15" pageNumber="516">
|
||
<emphasis id="64FCEAC2FFC4FF93FF3AFF7D6A8846BB" bold="true" box="[199,738,223,248]" pageId="15" pageNumber="516">
|
||
Coloration of
|
||
<typeStatus id="89338872FFC4FF93FE90FF7D69B946BB" box="[365,467,223,248]" pageId="15" pageNumber="516" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="CEB32A55FFC4FF93FE1EFF7D6A5846BB" box="[483,562,223,248]" captionStart="FIGURE 4" captionStartId="8.[151,250,1891,1913]" captionTargetBox="[189,1352,195,1861]" captionTargetId="figure@8.[189,1362,193,1865]" captionTargetPageId="8" captionText="FIGURE 4. Photographs of heads from ventral, lateral, and dorsal perspective to show color pattern variation among species. Images A, B, C Parvoscincus decipiens; D, E, F Parvoscincus jimmymcguirei sp. nov. holotype; G, H, I, Parvoscincus arvindiesmosi sp. nov. holotype; J, K, L Parvoscincus agtorum sp. nov. holotype; M, N, O, Parvoscincus abstrusus sp. nov. holotype (Male); P, Q, R, Parvoscincus abstrusus sp. nov. paratype (Female); S, T, U, Parvoscincus banahaoensis sp. nov. holotype; V, W, X, Parvoscincus palaliensis sp. nov. holotype; Y, Z, AA, Parvoscincus aurorus sp. nov. holotype." httpUri="https://zenodo.org/record/216111/files/figure.png" pageId="15" pageNumber="516">Figs. 4</figureCitation>
|
||
J, K, L; 5G, H)
|
||
</emphasis>
|
||
. Dorsal ground color brown with dark brown square vertebral marks. Dorsolateral band dark brown, originating at posterior apex of eye; blending into flank coloration by midbody; becoming a dark set of blotches over hind limb to base of tail. Bordered ventrally by white line one scale wide from eye to ear, transitioning to white spots posterior to ear. Flanks brown with lots of dorsolateral streaking. Ventrum cream colored, mental region with a few brown flecks, gular region with brown incomplete collar. Brown flecks near vent and base of tail. Dorsal aspect of hind limbs and forelimbs dark brown with white spotting on posterior portion of limb. Ventral limbs same color as ventrum, hands and feet light brown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1E92655BFFC4FF93FF3AFE7E6A604423" pageId="15" pageNumber="516" type="distribution">
|
||
<paragraph id="563736D0FFC4FF93FF3AFE7E6997445B" blockId="15.[151,1437,151,824]" pageId="15" pageNumber="516">
|
||
<emphasis id="64FCEAC2FFC4FF93FF3AFE7E69D247B6" bold="true" box="[199,440,476,501]" pageId="15" pageNumber="516">
|
||
Distribution (
|
||
<figureCitation id="CEB32A55FFC4FF93FE97FE7E69C547B6" box="[362,431,476,501]" captionStart="FIGURE 1" captionStartId="2.[151,250,1962,1984]" captionTargetBox="[311,1275,507,1947]" captionTargetId="figure@2.[311,1275,507,1949]" captionTargetPageId="2" captionText="FIGURE 1. Map of Luzon Island showing the collection localities for the new species and P. decipiens." httpUri="https://zenodo.org/record/216108/files/figure.png" pageId="15" pageNumber="516">Fig. 1</figureCitation>
|
||
)
|
||
</emphasis>
|
||
. The new species is known only from one locality in Aurora Province, Municipality of San Luis, Barangay Lipimental.
|
||
</paragraph>
|
||
<paragraph id="563736D0FFC4FF93FF3AFD866A604423" blockId="15.[151,1437,151,824]" pageId="15" pageNumber="516">
|
||
<emphasis id="64FCEAC2FFC4FF93FF3AFD8669EE447E" bold="true" box="[199,388,548,573]" pageId="15" pageNumber="516">Natural History</emphasis>
|
||
. Found in leaf litter in secondary growth forest away from water. The one female specimen collected has two oviductal eggs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1E92655BFFC4FF93FF3AFDCE6C85457B" pageId="15" pageNumber="516" type="etymology">
|
||
<paragraph id="563736D0FFC4FF93FF3AFDCE6C85457B" blockId="15.[151,1437,151,824]" pageId="15" pageNumber="516">
|
||
<emphasis id="64FCEAC2FFC4FF93FF3AFDCE692C44C6" bold="true" box="[199,326,620,645]" pageId="15" pageNumber="516">Etymology</emphasis>
|
||
. The specific epithet honors the Agta tribal group of Eastern Luzon that live in the Aurora region where this new species is found. These people are diminutive in stature, have an established cultural history of interaction with reptiles, including predation on humans by reticulated pythons (
|
||
<taxonomicName id="91884D53FFC4FF93FB99FD166DFE448E" box="[1124,1428,692,717]" class="Reptilia" family="Pythonidae" genus="Broghammerus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="516" phylum="Chordata" rank="species" species="reticulatus">
|
||
<emphasis id="64FCEAC2FFC4FF93FB99FD166DFE448E" box="[1124,1428,692,717]" italics="true" pageId="15" pageNumber="516">Broghammerus reticulatus</emphasis>
|
||
</taxonomicName>
|
||
; Headland & Greene 2011). The Agta people play a prominent role in helping to conserve the forests of the Sierra Madre Mountain Range and are increasingly threatened by mining and logging industries, which destructively extract resources from their ancestral tribal domains (van der Ploeg
|
||
<emphasis id="64FCEAC2FFC4FF93FC72FC836BAA457B" box="[911,960,799,824]" italics="true" pageId="15" pageNumber="516">et al</emphasis>
|
||
. 2011; Minter
|
||
<emphasis id="64FCEAC2FFC4FF93FB9AFC836CF2457B" box="[1127,1176,799,824]" italics="true" pageId="15" pageNumber="516">et al</emphasis>
|
||
. 2012).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |