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<document id="296B842DAB651A7B51901E36051F90F1" ID-CLB-Dataset="24286" ID-DOI="10.11646/zootaxa.4728.2.2" ID-GBIF-Dataset="a1c40b73-0c11-4863-a188-412c33523e02" ID-ISSN="1175-5326" ID-Zenodo-Dep="3621117" ID-ZooBank="AE4F8089-7D2E-4ABA-AA09-CD504705788A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1579677041620" checkinUser="plazi" docAuthor="Wagner, Rüdiger &amp; Withers, Phil" docDate="2020" docId="784D878B601F21068FF100F4FD358CBD" docLanguage="en" docName="zootaxa.4728.2.2.pdf" docOrigin="Zootaxa 4728 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tonnoiriella filistylis Vaillant &amp; Moubayed 1987" docType="treatment" docVersion="7" lastPageNumber="202" masterDocId="8474FFF3600C21158F66006DFFB28F20" masterDocTitle="The West-Palearctic species of the genus Tonnoiriella Vaillant, 1971 (Diptera: Psychodidae, Psychodinae)" masterLastPageNumber="210" masterPageNumber="183" pageNumber="202" updateTime="1698778572713" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="3A924DD96196614F0594CF9A5C3CF58F">The West-Palearctic species of the genus Tonnoiriella Vaillant, 1971 (Diptera: Psychodidae, Psychodinae)</mods:title>
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<mods:namePart id="A4DDCD7FA7127ED521FDBCF9DE97D0A3">Wagner, Rüdiger</mods:namePart>
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<mods:namePart id="C0D3CDE1CDE62D11CCB86A80084F1089">Withers, Phil</mods:namePart>
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<mods:title id="F91C6F2243C8CF518E13DAFB3DE04B15">Zootaxa</mods:title>
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<mods:date id="CEFD220F7C960F27AF26DC68A10132A7">2020</mods:date>
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<mods:number id="9D18BC9906E59D776C7E3AA9371A204D">4728</mods:number>
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<mods:identifier id="A9396C327B4D9C165EA256803A3EDDE3" type="DOI">10.11646/zootaxa.4728.2.2</mods:identifier>
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<treatment id="784D878B601F21068FF100F4FD358CBD" ID-DOI="http://doi.org/10.5281/zenodo.5918607" ID-GBIF-Taxon="161827854" ID-Zenodo-Dep="5918607" LSID="urn:lsid:plazi:treatment:784D878B601F21068FF100F4FD358CBD" httpUri="http://treatment.plazi.org/id/784D878B601F21068FF100F4FD358CBD" lastPageNumber="202" pageId="19" pageNumber="202">
<subSubSection id="B8FE6516601F21068FF100F4FD408F93" box="[151,754,153,180]" pageId="19" pageNumber="202" type="nomenclature">
<paragraph id="F05B369D601F21068FF100F4FD408F93" blockId="19.[151,754,153,215]" box="[151,754,153,180]" pageId="19" pageNumber="202">
<heading id="AB1381F1601F21068FF100F4FD408F93" bold="true" box="[151,754,153,180]" fontSize="11" level="1" pageId="19" pageNumber="202" reason="1">
<taxonomicName id="37E44D1E601F21068FF100F4FD408F93" ID-CoL="7CHTJ" authority="Vaillant &amp; Moubayed, 1987" authorityName="Vaillant &amp; Moubayed" authorityYear="1987" box="[151,754,153,180]" class="Insecta" family="Psychodidae" genus="Tonnoiriella" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="202" phylum="Arthropoda" rank="species" species="filistylis">
<emphasis id="C290EA8F601F21068FF100F4FD408F93" bold="true" box="[151,754,153,180]" pageId="19" pageNumber="202">
<emphasis id="C290EA8F601F21068FF100F4FE208F94" bold="true" box="[151,402,153,180]" italics="true" pageId="19" pageNumber="202">Tonnoiriella filistylis</emphasis>
<bibRefCitation id="94754B6C601F21068EFF00F4FD408F93" author="Vaillant, F. &amp; Moubayed, J." box="[409,754,153,180]" pageId="19" pageNumber="202" pagination="121 - 127" refId="ref20807" refString="Vaillant, F. &amp; Moubayed, J. (1987) Cinq especes nouvelles de Dipteres Psychodidae Psychodinae du Liban. Annales de Limnologie, 23 (2), 121 - 127. https: // doi. org / 10.1051 / limn / 1987008" type="journal article" year="1987">Vaillant &amp; Moubayed, 1987</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B8FE6516601F21068FF100D0FE9C8FF7" box="[151,302,189,215]" pageId="19" pageNumber="202" type="description">
<paragraph id="F05B369D601F21068FF100D0FE9C8FF7" blockId="19.[151,754,153,215]" box="[151,302,189,215]" pageId="19" pageNumber="202">
(
<figureCitation id="68DF2A18601F21068FF900D0FE948FF7" box="[159,294,189,215]" captionStart="FIGURES 32–42" captionStartId="17.[151,264,1698,1723]" captionTargetBox="[175,1372,209,1639]" captionTargetId="figure@17.[151,1436,181,1672]" captionTargetPageId="17" captionText="FIGURES 32–42: 32–34 T. mollis (holotype), 32 styles and hypandrium, 33 epandrial processes, lateral view, 34 epandrium, epandrial processes and proctiger; 35–37 T. mollis (paratype), 35 hypandrium, 36 epandrial processes, lateral view, 37 aedeagus dorsal view; 38–40 T. anchoriformis 38 hypandrium, gonopods, aedeagus ventral view, 39 gonostylus, 40 epandrial processes with tenacula; 41–42 T. filistylis 41 apical part of gonostylus, 42 genitalia ventral view, (figs 41 and 42 from Vaillant &amp; Moubayed 1987)." figureDoi="http://doi.org/10.5281/zenodo.3621127" httpUri="https://zenodo.org/record/3621127/files/figure.png" pageId="19" pageNumber="202">Figs. 41, 42</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B8FE6516601F21068FF10091FBBE8E35" box="[151,1036,252,277]" pageId="19" pageNumber="202" type="reference_group">
<paragraph id="F05B369D601F21068FF10091FBBE8E35" blockId="19.[151,1036,252,277]" box="[151,1036,252,277]" pageId="19" pageNumber="202">
<taxonomicName id="37E44D1E601F21068FF10091FD3D8E34" ID-CoL="7CHTJ" authority="Vaillant &amp; Moubayed, 1987" authorityName="Vaillant &amp; Moubayed" authorityYear="1987" box="[151,655,252,277]" class="Insecta" family="Psychodidae" genus="Tonnoiriella" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="202" phylum="Arthropoda" rank="species" species="filistylis">
<emphasis id="C290EA8F601F21068FF10091FED88E34" box="[151,362,252,276]" italics="true" pageId="19" pageNumber="202">Tonnoiriella filistylis</emphasis>
<bibRefCitation id="94754B6C601F21068E160091FD3D8E34" author="Vaillant, F. &amp; Moubayed, J." box="[368,655,252,277]" pageId="19" pageNumber="202" pagination="121 - 127" refId="ref20807" refString="Vaillant, F. &amp; Moubayed, J. (1987) Cinq especes nouvelles de Dipteres Psychodidae Psychodinae du Liban. Annales de Limnologie, 23 (2), 121 - 127. https: // doi. org / 10.1051 / limn / 1987008" type="journal article" year="1987">Vaillant &amp; Moubayed, 1987</bibRefCitation>
</taxonomicName>
: Annales de Limnol 23 (2): 124–125.
</paragraph>
</subSubSection>
<subSubSection id="B8FE6516601F21068FF1012FFA8C8EA2" pageId="19" pageNumber="202" type="materials_examined">
<paragraph id="F05B369D601F21068FF1012FFA8C8EA2" blockId="19.[151,1437,322,925]" pageId="19" pageNumber="202">
<emphasis id="C290EA8F601F21068FF1012FFF4D8E7D" bold="true" box="[151,255,322,349]" pageId="19" pageNumber="202">Material</emphasis>
:
<materialsCitation id="408C3CC0601F21068E6B012EFA8C8EA2" ID-GBIF-Occurrence="2556505432" collectingDate="1982-05-17" collectorName="Moubayed" country="Lebanon" elevation="650" location="Hermel" municipality="Orontes river" pageId="19" pageNumber="202" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="2F5F883F601F21068E6B012EFECA8E7D" box="[269,376,323,349]" pageId="19" pageNumber="202" type="holotype">Holotype</typeStatus>
,
<specimenCount id="E6E2FD14601F21068EE4012EFE178E7D" box="[386,421,323,349]" pageId="19" pageNumber="202" type="male">1♂</specimenCount>
collected near
<location id="F53B6046601F21068D2A012EFD118E7D" LSID="urn:lsid:plazi:treatment:784D878B601F21068FF100F4FD358CBD:F53B6046601F21068D2A012EFD118E7D" box="[588,675,323,349]" country="Lebanon" municipality="Orontes river" name="Hermel" pageId="19" pageNumber="202">Hermel</location>
,
<collectingCountry id="88F3760D601F21068DCB012EFCA08E7D" box="[685,786,323,349]" name="Lebanon" pageId="19" pageNumber="202">Lebanon</collectingCountry>
, collected near the spring of a small affluent of the
<collectingMunicipality id="103FACE7601F21068A22012EFF7E8EA1" pageId="19" pageNumber="202">Orontes river</collectingMunicipality>
at
<elevation id="7BC9D1AE601F21068F88010BFEC38EA1" box="[238,369,358,385]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="19" pageNumber="202" unit="m" value="650.0">
<quantity id="371C9B78601F21068F88010BFE878EA1" box="[238,309,358,385]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="19" pageNumber="202" unit="m" value="650.0">650 m</quantity>
a.s.l.
</elevation>
(
<date id="845A105D601F21068EE6010AFDA58EA1" box="[384,535,359,385]" pageId="19" pageNumber="202" value="1982-05-17">
<collectingDate id="941EE9B5601F21068EE6010AFDA58EA1" box="[384,535,359,385]" pageId="19" pageNumber="202" value="1982-05-17">17 May 1982</collectingDate>
</date>
) leg.
<collectorName id="5D11534B601F21068D30010AFD7C8EA1" box="[598,718,359,385]" pageId="19" pageNumber="202">Moubayed</collectorName>
(specimen not in the collection Vaillant, probably lost).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="B8FE6516601F21068FA101E7FE698DE5" pageId="19" pageNumber="202" type="description">
<paragraph id="F05B369D601F21068FA101E7FEA98ECD" blockId="19.[151,1437,322,925]" pageId="19" pageNumber="202">
<emphasis id="C290EA8F601F21068FA101E7FEE38E85" bold="true" box="[199,337,394,421]" pageId="19" pageNumber="202">Description</emphasis>
: Head with eyebridge of 3 facet rows. Antenna with scape and pedicel broader than long, only 3 flagellomeres existent; relative length: 24-23-19-20-22-? Palpus with only 3 segments remaining, relative length: 34-40-38-?.
</paragraph>
<paragraph id="F05B369D601F21068FA1019AFCB38D31" blockId="19.[151,1437,322,925]" box="[199,769,502,529]" pageId="19" pageNumber="202">
Wing length
<quantity id="371C9B78601F21068E3E019AFE038D31" box="[344,433,503,529]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="19" pageNumber="202" unit="mm" value="2.1">2.1 mm</quantity>
; relation length by width 2.95
</paragraph>
<paragraph id="F05B369D601F21068FA10276FC618D5D" blockId="19.[151,1437,322,925]" pageId="19" pageNumber="202">
Male terminalia (
<figureCitation id="68DF2A18601F21068EEB0276FE538D15" box="[397,481,539,565]" captionStart="FIGURES 32–42" captionStartId="17.[151,264,1698,1723]" captionTargetBox="[175,1372,209,1639]" captionTargetId="figure@17.[151,1436,181,1672]" captionTargetPageId="17" captionText="FIGURES 32–42: 32–34 T. mollis (holotype), 32 styles and hypandrium, 33 epandrial processes, lateral view, 34 epandrium, epandrial processes and proctiger; 35–37 T. mollis (paratype), 35 hypandrium, 36 epandrial processes, lateral view, 37 aedeagus dorsal view; 38–40 T. anchoriformis 38 hypandrium, gonopods, aedeagus ventral view, 39 gonostylus, 40 epandrial processes with tenacula; 41–42 T. filistylis 41 apical part of gonostylus, 42 genitalia ventral view, (figs 41 and 42 from Vaillant &amp; Moubayed 1987)." figureDoi="http://doi.org/10.5281/zenodo.3621127" httpUri="https://zenodo.org/record/3621127/files/figure.png" pageId="19" pageNumber="202">Fig. 42</figureCitation>
) with hypandrium linguiform prolonged in the middle. Gonocoxites approximately 1.8x longer than wide; gonostyli longer than gonocoxites, almost straight, apical section suddenly thinner, flattened and with of 4 setae bases in line that are in contact to each other (
<figureCitation id="68DF2A18601F21068C14020EFC778D5D" box="[882,965,611,637]" captionStart="FIGURES 32–42" captionStartId="17.[151,264,1698,1723]" captionTargetBox="[175,1372,209,1639]" captionTargetId="figure@17.[151,1436,181,1672]" captionTargetPageId="17" captionText="FIGURES 32–42: 32–34 T. mollis (holotype), 32 styles and hypandrium, 33 epandrial processes, lateral view, 34 epandrium, epandrial processes and proctiger; 35–37 T. mollis (paratype), 35 hypandrium, 36 epandrial processes, lateral view, 37 aedeagus dorsal view; 38–40 T. anchoriformis 38 hypandrium, gonopods, aedeagus ventral view, 39 gonostylus, 40 epandrial processes with tenacula; 41–42 T. filistylis 41 apical part of gonostylus, 42 genitalia ventral view, (figs 41 and 42 from Vaillant &amp; Moubayed 1987)." figureDoi="http://doi.org/10.5281/zenodo.3621127" httpUri="https://zenodo.org/record/3621127/files/figure.png" pageId="19" pageNumber="202">Fig. 41</figureCitation>
).
</paragraph>
<paragraph id="F05B369D601F21068FA102EAFE698DE5" blockId="19.[151,1437,322,925]" pageId="19" pageNumber="202">Epandrium rhomboid, apically converging; epandrial processes basally swollen, apically with 3 feathered tenacula, one long and 2 short.</paragraph>
</subSubSection>
<subSubSection id="B8FE6516601F21068FA102A3FE408C2D" pageId="19" pageNumber="202" type="distribution">
<paragraph id="F05B369D601F21068FA102A3FE408C2D" blockId="19.[151,1437,322,925]" pageId="19" pageNumber="202">
<emphasis id="C290EA8F601F21068FA102A3FEEB8DC9" bold="true" box="[199,345,718,745]" pageId="19" pageNumber="202">Distribution</emphasis>
:
<collectingCountry id="88F3760D601F21068E0102A2FE7E8DC9" box="[359,460,719,745]" name="Lebanon" pageId="19" pageNumber="202">Lebanon</collectingCountry>
, known only from the
<typeStatus id="2F5F883F601F21068DA002A2FD448DC9" box="[710,758,719,745]" pageId="19" pageNumber="202">type</typeStatus>
locality Hermel, collected on the bank of a small tributary of the Orontes river in
<date id="845A105D601F21068E1F029EFE5C8C2D" box="[377,494,755,781]" pageId="19" pageNumber="202" value="1982-05">May 1982</date>
.
</paragraph>
</subSubSection>
<subSubSection id="B8FE6516601F21068FA1037AFD358CBD" pageId="19" pageNumber="202" type="discussion">
<paragraph id="F05B369D601F21068FA1037AFD358CBD" blockId="19.[151,1437,322,925]" pageId="19" pageNumber="202">
<emphasis id="C290EA8F601F21068FA1037AFE868C11" bold="true" box="[199,308,791,817]" pageId="19" pageNumber="202">Remarks</emphasis>
: The species is well characterized by the medially prolonged hypandrium, the elongate gonostylus with seta bases in line and the short basally swollen epandrial processes with 3 tenacula, 1 long, 2 short. The whereabouts of the sole
<typeStatus id="2F5F883F601F21068E000332FE7A8C59" box="[358,456,863,889]" pageId="19" pageNumber="202" type="holotype">holotype</typeStatus>
specimen is unknown. It is not in the Vaillant collection in the Lausanne Museum, nor is it in the personal collection of Moubayed.
</paragraph>
</subSubSection>
</treatment>
</document>
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