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<document id="EF967429697F07F0A2BABFCA1C833013" ID-CLB-Dataset="24286" ID-DOI="10.11646/zootaxa.4728.2.2" ID-GBIF-Dataset="a1c40b73-0c11-4863-a188-412c33523e02" ID-ISSN="1175-5326" ID-Zenodo-Dep="3621117" ID-ZooBank="AE4F8089-7D2E-4ABA-AA09-CD504705788A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1579677041620" checkinUser="plazi" docAuthor="Wagner, Rüdiger &amp; Withers, Phil" docDate="2020" docId="784D878B6007211E8FF100F4FE758BA0" docLanguage="en" docName="zootaxa.4728.2.2.pdf" docOrigin="Zootaxa 4728 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tonnoiriella ikariae Wagner &amp; Withers 2020, sp. nov." docType="treatment" docVersion="7" lastPageNumber="194" masterDocId="8474FFF3600C21158F66006DFFB28F20" masterDocTitle="The West-Palearctic species of the genus Tonnoiriella Vaillant, 1971 (Diptera: Psychodidae, Psychodinae)" masterLastPageNumber="210" masterPageNumber="183" pageNumber="194" updateTime="1698778572713" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="1B41CD0955777ABE24F7E6E39F761699">The West-Palearctic species of the genus Tonnoiriella Vaillant, 1971 (Diptera: Psychodidae, Psychodinae)</mods:title>
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<mods:namePart id="6C39C35C7ED8CF99E7D9417FEAA2C200">Wagner, Rüdiger</mods:namePart>
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<treatment id="784D878B6007211E8FF100F4FE758BA0" ID-DOI="http://doi.org/10.5281/zenodo.5918591" ID-GBIF-Taxon="161827869" ID-Zenodo-Dep="5918591" LSID="urn:lsid:plazi:treatment:784D878B6007211E8FF100F4FE758BA0" httpUri="http://treatment.plazi.org/id/784D878B6007211E8FF100F4FE758BA0" lastPageNumber="194" pageId="11" pageNumber="194">
<subSubSection id="B8FE65166007211E8FF100F4FE5F8F94" box="[151,493,153,180]" pageId="11" pageNumber="194" type="nomenclature">
<paragraph id="F05B369D6007211E8FF100F4FE5F8F94" blockId="11.[151,493,153,216]" box="[151,493,153,180]" pageId="11" pageNumber="194">
<heading id="AB1381F16007211E8FF100F4FE5F8F94" bold="true" box="[151,493,153,180]" fontSize="11" level="1" pageId="11" pageNumber="194" reason="1">
<emphasis id="C290EA8F6007211E8FF100F4FE5F8F94" bold="true" box="[151,493,153,180]" pageId="11" pageNumber="194">
<taxonomicName id="37E44D1E6007211E8FF100F4FE3A8F94" ID-CoL="9HX42" authority="Wagner &amp; Withers, 2020" authorityName="Wagner &amp; Withers" authorityYear="2020" box="[151,392,153,180]" class="Insecta" family="Psychodidae" genus="Tonnoiriella" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="194" phylum="Arthropoda" rank="species" species="ikariae" status="sp. nov.">
<emphasis id="C290EA8F6007211E8FF100F4FE3A8F94" bold="true" box="[151,392,153,180]" italics="true" pageId="11" pageNumber="194">Tonnoiriella ikariae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D9A357F46007211E8EE900F7FE5F8F94" box="[399,493,154,180]" pageId="11" pageNumber="194" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B8FE65166007211E8FF100D0FF488FF8" box="[151,250,189,216]" pageId="11" pageNumber="194" type="description">
<paragraph id="F05B369D6007211E8FF100D0FF488FF8" blockId="11.[151,493,153,216]" box="[151,250,189,216]" pageId="11" pageNumber="194">
(
<figureCitation id="68DF2A186007211E8FC600D0FF438FF8" box="[160,241,189,216]" captionStart="FIGURES 1525" captionStartId="10.[151,264,1910,1935]" captionTargetBox="[213,1374,200,1864]" captionTargetId="figure@10.[196,1391,181,1885]" captionTargetPageId="10" captionText="FIGURES 1525: 15 T. sieberti genitalia ventral view; 16 T. ikariae genitalia ventral view; 1721 T. italiae, genitalia ventral view of specimen from northern Italy (17), Sicily (18), aedeagus of specimen from N Italy (19), Sicily, Villa doso (20) and Sicily Castelbuono (21); 22 T. turcica genitalia ventral view; 23 T. rhodesica genitalia ventral view; 24 T syriensis genitalia ventral view, 25 T. syriensis epandrium and epandrial processes oblique view." figureDoi="http://doi.org/10.5281/zenodo.3621123" httpUri="https://zenodo.org/record/3621123/files/figure.png" pageId="11" pageNumber="194">Fig. 16</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B8FE65166007211E8FF10168FDA78E63" pageId="11" pageNumber="194" type="materials_examined">
<paragraph id="F05B369D6007211E8FF10168FDA78E63" blockId="11.[151,1437,261,1152]" pageId="11" pageNumber="194">
<emphasis id="C290EA8F6007211E8FF10168FF4D8E00" bold="true" box="[151,255,261,288]" pageId="11" pageNumber="194">Material</emphasis>
:
<materialsCitation id="408C3CC06007211E8E680168FDA38E63" ID-GBIF-Occurrence="2556505437" collectingDate="1979-05-30" collectionCode="RW" collectorName="Malicky" country="Greece" elevation="180" latitude="37.633335" location="Ikaria island" longLatPrecision="1180" longitude="26.216667" pageId="11" pageNumber="194" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<specimenCount id="E6E2FD146007211E8E680168FE808E3F" box="[270,306,261,287]" pageId="11" pageNumber="194" type="male">1♂</specimenCount>
,
<typeStatus id="2F5F883F6007211E8E580168FE118E3F" box="[318,419,261,287]" pageId="11" pageNumber="194" type="holotype">holotype</typeStatus>
,
<collectingCountry id="88F3760D6007211E8EC80168FDB28E3F" box="[430,512,261,287]" name="Greece" pageId="11" pageNumber="194">Greece</collectingCountry>
,
<location id="F53B60466007211E8D6C0168FD2A8E3F" LSID="urn:lsid:plazi:treatment:784D878B6007211E8FF100F4FE758BA0:F53B60466007211E8D6C0168FD2A8E3F" box="[522,664,261,287]" country="Greece" latitude="37.633335" longLatPrecision="1180" longitude="26.216667" name="Ikaria island" pageId="11" pageNumber="194">Ikaria island</location>
,
<locationDeviation id="E1E92FBF6007211E8DC50168FC698E00" box="[675,987,261,288]" location="East of Karavostomou" pageId="11" pageNumber="194">
<quantity id="371C9B786007211E8DC50168FD698E3F" box="[675,731,261,287]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="194" unit="km" value="2.0">2 km</quantity>
East of Karavostomou
</locationDeviation>
(
<geoCoordinate id="95D0505A6007211E8C8C0168FBFE8E00" box="[1002,1100,261,288]" degrees="37" direction="north" minutes="38" orientation="latitude" pageId="11" pageNumber="194" precision="925" value="37.633335">37°38N</geoCoordinate>
/
<geoCoordinate id="95D0505A6007211E8B310168FB068E00" box="[1111,1204,261,288]" degrees="26" direction="east" minutes="13" orientation="longitude" pageId="11" pageNumber="194" precision="925" value="26.216667">26°13E</geoCoordinate>
), spring,
<quantity id="371C9B786007211E8A7D0168FAD08E00" box="[1307,1378,261,288]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="11" pageNumber="194" unit="m" value="180.0">
<elevation id="7BC9D1AE6007211E8A7D0168FAD08E00" box="[1307,1378,261,288]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="11" pageNumber="194" unit="m" value="180.0">180 m</elevation>
</quantity>
a.s.l,
<date id="845A105D6007211E8FF10144FE828E63" box="[151,304,297,324]" pageId="11" pageNumber="194" value="1979-05-30">
<collectingDate id="941EE9B56007211E8FF10144FE828E63" box="[151,304,297,324]" pageId="11" pageNumber="194" value="1979-05-30">30 May 1979</collectingDate>
</date>
, leg.
<collectorName id="5D11534B6007211E8E0D0144FE7A8E63" box="[363,456,297,323]" pageId="11" pageNumber="194">Malicky</collectorName>
(
<collectionCode id="96F5AE586007211E8EB10144FDBB8E63" box="[471,521,297,323]" pageId="11" pageNumber="194">RW</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B8FE65166007211E8FA10120FB6E8CEB" pageId="11" pageNumber="194" type="description">
<paragraph id="F05B369D6007211E8FA10120FC658ED8" blockId="11.[151,1437,261,1152]" pageId="11" pageNumber="194">
<emphasis id="C290EA8F6007211E8FA10120FEE98E48" bold="true" box="[199,347,333,360]" pageId="11" pageNumber="194">Description:</emphasis>
Head with an eyebridge of 3 facet rows; distance between eyes 4 facet diameters; several postocular bristles behind eyes. Antenna with scape, pedicel and 14 flagellomeres: absolute length 0.038-0.05-0.042-0.044- 0.042-0.042-0.044-0.044-0.046-0.046-0.036-0.036-0.03-0.022-
<quantity id="371C9B786007211E8C3D01F8FBB98E90" box="[859,1035,405,432]" metricMagnitude="-5" metricUnit="m" metricValue="3.2" metricValueMax="4.3999999999999995" metricValueMin="2.0" pageId="11" pageNumber="194" unit="mm" value="0.032" valueMax="0.044" valueMin="0.02">0.02-0.044 mm</quantity>
; relative length 19-25-21-22-21-21- 22-22-23-23-
<date id="845A105D6007211E8E5201D4FE098EF4" box="[308,443,441,468]" pageId="11" pageNumber="194" value="2018-11-18" valueMax="2018-11-15" valueMin="2018-11-18">18-18-15-11</date>
-10-12. Flagellomeres 511 with a pair of digitiform curved ascoids. Palpus segments, absolute length: 0.05-0.062-
<quantity id="371C9B786007211E8EB701B0FD228ED8" box="[465,656,477,504]" metricMagnitude="-4" metricUnit="m" metricValue="1.06" metricValueMax="1.38" metricValueMin="0.74" pageId="11" pageNumber="194" unit="mm" value="0.106" valueMax="0.138" valueMin="0.074">0.074-0.138 mm</quantity>
; relative lenght 25-31-37-69.
</paragraph>
<paragraph id="F05B369D6007211E8FA1026CFC708D3C" blockId="11.[151,1437,261,1152]" box="[199,962,513,540]" pageId="11" pageNumber="194">
Wing 2.70 times longer than wide; length
<quantity id="371C9B786007211E8DF9026CFCB78D3C" box="[671,773,513,540]" metricMagnitude="-3" metricUnit="m" metricValue="1.82" pageId="11" pageNumber="194" unit="mm" value="1.82">1.82 mm</quantity>
, width
<quantity id="371C9B786007211E8C31026CFC0C8D3C" box="[855,958,513,540]" metricMagnitude="-4" metricUnit="m" metricValue="6.7" pageId="11" pageNumber="194" unit="mm" value="0.67">0.67 mm</quantity>
.
</paragraph>
<paragraph id="F05B369D6007211E8FA10248FBBF8DA7" blockId="11.[151,1437,261,1152]" pageId="11" pageNumber="194">
Male terminalia (
<figureCitation id="68DF2A186007211E8EE10248FE6A8D60" box="[391,472,549,576]" captionStart="FIGURES 1525" captionStartId="10.[151,264,1910,1935]" captionTargetBox="[213,1374,200,1864]" captionTargetId="figure@10.[196,1391,181,1885]" captionTargetPageId="10" captionText="FIGURES 1525: 15 T. sieberti genitalia ventral view; 16 T. ikariae genitalia ventral view; 1721 T. italiae, genitalia ventral view of specimen from northern Italy (17), Sicily (18), aedeagus of specimen from N Italy (19), Sicily, Villa doso (20) and Sicily Castelbuono (21); 22 T. turcica genitalia ventral view; 23 T. rhodesica genitalia ventral view; 24 T syriensis genitalia ventral view, 25 T. syriensis epandrium and epandrial processes oblique view." figureDoi="http://doi.org/10.5281/zenodo.3621123" httpUri="https://zenodo.org/record/3621123/files/figure.png" pageId="11" pageNumber="194">Fig. 16</figureCitation>
) with hypandrium expanded along 1/3 of its ventral length with 2 lateral and 1 median widenings. Gonocoxite approximately
<date id="845A105D6007211E8D2A0224FDDA8D43" box="[588,616,585,611]" pageId="11" pageNumber="194">2x</date>
longer than its median width; inner side with a basal group of setae, just before middle 2 long setae; gonostylus longer than gonocoxite, almost straight.
</paragraph>
<paragraph id="F05B369D6007211E8FA102FFFE868C37" blockId="11.[151,1437,261,1152]" pageId="11" pageNumber="194">Parameres with a broad ventral bridge, lobes just reach the bases of the long setae on gonocoxite, ventrally in the midst with a racket-shaped brownish spot of aedeagus sheath; above a pair of processes as long as the more ventral processes that do not meet in the middle. Further dorsal large basally round condyles with parallel inner edges form the keel.</paragraph>
<paragraph id="F05B369D6007211E8FA1034CFC5F8C88" blockId="11.[151,1437,261,1152]" pageId="11" pageNumber="194">Basiphallus longer than wide, basally broader, apically bilobed; ventral lobe bifid, ventral part connected to the distiphallus, longer part slightly bent with blunt tip; distiphallus longer than gonocoxites angular, with both ends acute, the broader end with a broad projection, the smaller end double-headed. The end of the dorsal distiphallus is across anchor-shaped sclerite clearly thinner than the its ventral counterpart.</paragraph>
<paragraph id="F05B369D6007211E8FA103DCFB6E8CEB" blockId="11.[151,1437,261,1152]" box="[199,1244,945,972]" pageId="11" pageNumber="194">Epandrium rhomboid, epandrial processes almost straight, apically with 8 feathered tenacula.</paragraph>
</subSubSection>
<subSubSection id="B8FE65166007211E8FA103B8FDC58CCF" box="[199,631,981,1008]" pageId="11" pageNumber="194" type="distribution">
<paragraph id="F05B369D6007211E8FA103B8FDC58CCF" blockId="11.[151,1437,261,1152]" box="[199,631,981,1008]" pageId="11" pageNumber="194">
<emphasis id="C290EA8F6007211E8FA103B8FEEB8CD0" bold="true" box="[199,345,981,1008]" pageId="11" pageNumber="194">Distribution</emphasis>
:
<collectingCountry id="88F3760D6007211E8E0E03B8FE088CCF" box="[360,442,981,1007]" name="Greece" pageId="11" pageNumber="194">Greece</collectingCountry>
, island of Ikaria.
</paragraph>
</subSubSection>
<subSubSection id="B8FE65166007211E8FA10397FB6E8B33" box="[199,1244,1017,1044]" pageId="11" pageNumber="194" type="etymology">
<paragraph id="F05B369D6007211E8FA10397FB6E8B33" blockId="11.[151,1437,261,1152]" box="[199,1244,1017,1044]" pageId="11" pageNumber="194">
<emphasis id="C290EA8F6007211E8FA10397FEE08B34" bold="true" box="[199,338,1018,1044]" pageId="11" pageNumber="194">Etymology:</emphasis>
The epithet is derived from the island of Ikaria, where the species was collected.
</paragraph>
</subSubSection>
<subSubSection id="B8FE65166007211E8FA10473FE758BA0" pageId="11" pageNumber="194" type="discussion">
<paragraph id="F05B369D6007211E8FA10473FE758BA0" blockId="11.[151,1437,261,1152]" pageId="11" pageNumber="194">
<emphasis id="C290EA8F6007211E8FA10473FE868B18" bold="true" box="[199,308,1054,1080]" pageId="11" pageNumber="194">Remarks</emphasis>
: Another species of the
<taxonomicName id="37E44D1E6007211E8D210473FD0A8B17" authorityName="Wagner" authorityYear="1993" box="[583,696,1053,1079]" class="Insecta" family="Psychodidae" genus="Tonnoiriella" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="194" phylum="Arthropoda" rank="species" species="sieberti">
<emphasis id="C290EA8F6007211E8D210473FD0A8B17" box="[583,696,1053,1079]" italics="true" pageId="11" pageNumber="194">T. sieberti</emphasis>
</taxonomicName>
-group. The middle part of the hypandrium is less prominent than in
<taxonomicName id="37E44D1E6007211E8FD2042FFEAF8B7B" authorityName="Wagner" authorityYear="1986" box="[180,285,1090,1115]" class="Insecta" family="Psychodidae" genus="Tonnoiriella" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="194" phylum="Arthropoda" rank="species" species="turcica">
<emphasis id="C290EA8F6007211E8FD2042FFEAF8B7B" box="[180,285,1090,1115]" italics="true" pageId="11" pageNumber="194">T. turcica</emphasis>
</taxonomicName>
and
<taxonomicName id="37E44D1E6007211E8E34042FFE738B7B" authorityName="Wagner" authorityYear="1993" box="[338,449,1089,1115]" class="Insecta" family="Psychodidae" genus="Tonnoiriella" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="194" phylum="Arthropoda" rank="species" species="sieberti">
<emphasis id="C290EA8F6007211E8E34042FFE738B7B" box="[338,449,1089,1115]" italics="true" pageId="11" pageNumber="194">T. sieberti</emphasis>
</taxonomicName>
. The shape of the hypandrium, and the long and angular distiphallus distinguish the new species from its congeners.
</paragraph>
</subSubSection>
</treatment>
</document>