320 lines
39 KiB
XML
320 lines
39 KiB
XML
<document id="C2E0C3E5B8E389CAE81CD4AE9978C9FD" ID-DOI="10.11646/zootaxa.2319.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5317868" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629657556386" checkinUser="felipe" docAuthor="Müller, Hans-Georg & Krapp, Franz" docDate="2009" docId="03E687F82919FFD27ADC11399F1AFA76" docLanguage="en" docName="zt02319p138.pdf" docOrigin="Zootaxa 2319 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2319.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Anoplodactylus glandulifer Stock 1954" docType="treatment" docVersion="4" lastPageNumber="97" masterDocId="FFDFFF802947FFB27A4B175F9C1FFF8D" masterDocTitle="The pycnogonid fauna (Pycnogonida, Arthropoda) of the Tayrona National Park and adjoining areas on the Caribbean coast of Colombia 2319" masterLastPageNumber="138" masterPageNumber="1" pageNumber="95" updateTime="1699090509478" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="D481E155792A6C0835E67AC9C801F729">The pycnogonid fauna (Pycnogonida, Arthropoda) of the Tayrona National Park and adjoining areas on the Caribbean coast of Colombia 2319</mods:title>
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<mods:namePart id="9EF11D6240B1B9F5761AF1A168492EF9">Müller, Hans-Georg</mods:namePart>
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<mods:namePart id="44C32912BC8E00FE014D99016EDD1CFD">Krapp, Franz</mods:namePart>
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<mods:title id="DAB15804C035F80CDA246A466D17FAE5">Zootaxa</mods:title>
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<mods:date id="29F6B90338442B7E7B305FCD9AB6D7D6">2009</mods:date>
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<mods:number id="AD82F7D0B8B5A664F14BCD78475C0131">2319</mods:number>
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<taxonomicName id="4C4F4D6D2919FFEC7ADC11399E69F90D" ID-CoL="678RT" authority="Stock, 1954" authorityName="Stock" authorityYear="1954" box="[151,630,1638,1664]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" kingdom="Animalia" order="Pantopoda" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="glandulifer">
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<emphasis id="B93BEAFC2919FFEC7ADC11399E69F90D" bold="true" box="[151,630,1638,1664]" pageId="94" pageNumber="95">
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<emphasis id="B93BEAFC2919FFEC7ADC11399DFFF90D" bold="true" box="[151,480,1638,1664]" italics="true" pageId="94" pageNumber="95">Anoplodactylus glandulifer</emphasis>
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Stock, 1954
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35565652919FFEC7ADC11D69CF6F92E" box="[151,233,1673,1699]" pageId="94" pageNumber="95" type="description">
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<paragraph id="8BF036EE2919FFEC7ADC11D69CF6F92E" blockId="94.[151,630,1638,1699]" box="[151,233,1673,1699]" pageId="94" pageNumber="95">
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<figureCitation id="13742A6B2919FFEC7ADC11D69CF6F92E" box="[151,233,1673,1699]" captionStart="FIGURE 50" captionStartId="95.[151,257,1404,1428]" captionTargetBox="[203,1391,229,1382]" captionTargetId="figure-252@95.[193,1394,217,1397]" captionTargetPageId="95" captionText="FIGURE 50. Anoplodactylus glandulifer Stock, 1954, male: A ovigerous specimen, trunk lateral, B trunk dorsal, C chela, D oviger, E leg 3, F femur with two cement gland pores, G cement gland pores in lateral view, H tarsus and propodus." figureDoi="http://doi.org/10.5281/zenodo.5362510" httpUri="https://zenodo.org/record/5362510/files/figure.png" pageId="94" pageNumber="95">Fig. 50</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35565652919FFEC7ADC118898C6F8A0" pageId="94" pageNumber="95" type="reference_group">
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<paragraph id="8BF036EE2919FFEC7ADC11889984F960" blockId="94.[151,1435,1750,1838]" box="[151,1435,1750,1774]" pageId="94" pageNumber="95">
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<taxonomicName id="4C4F4D6D2919FFEC7ADC11889E63F960" ID-CoL="678RT" authority="Stock, 1954: 273" authorityName="Stock" authorityPageNumber="273" authorityYear="1954" box="[151,636,1750,1774]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" kingdom="Animalia" order="Pantopoda" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="glandulifer">
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<emphasis id="B93BEAFC2919FFEC7ADC11889DA4F963" box="[151,443,1751,1774]" italics="true" pageId="94" pageNumber="95">Anoplodactylus glandulifer</emphasis>
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Stock, 1954: 273
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</taxonomicName>
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. Stock 1958: 3. Stock 1968: 49. Arnaud 1973: 955. Stock 1974: 16. Child
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</paragraph>
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<paragraph id="8BF036EE2919FFEC7A8C11A998C6F8A0" blockId="94.[151,1435,1750,1838]" pageId="94" pageNumber="95">
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1982: 273. Child 1988: 59. Nakamura & Child 1988: 813.
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<bibRefCitation id="EFDE4B1F2919FFEC797311A99FF6F880" author="Child, C. A." box="[824,1001,1782,1805]" pageId="94" pageNumber="95" pagination="311 - 335" refId="ref76484" refString="Child, C. A. (1990) Pycnogonida of the Western Pacific Islands, VIII. Recent collections from islands of the Great Barrier Reef, Australia. Proceedings of the biological Society Washington, 103 (2), 311 - 335." type="journal article" year="1990">Child 1990: 331</bibRefCitation>
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. Müller 1990: 74.
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||
<bibRefCitation id="EFDE4B1F2919FFEC7EE611A99941F880" author="Child, C. A." box="[1197,1374,1782,1805]" pageId="94" pageNumber="95" pagination="138 - 146" refId="ref76530" refString="Child, C. A. (1991) Pycnogonida of the Western Pacific Islands. A shallow-water Guam survey, 1984. - Proceedings of the biological Society Washington 104 (1), 138 - 146." type="journal article" year="1991">Child 1991: 143</bibRefCitation>
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.
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<taxonomicName id="4C4F4D6D2919FFEC7ADC10489E63F8A0" ID-CoL="66WQZ" authority="Child, 1977: 593" authorityName="Child" authorityPageNumber="593" authorityYear="1977" box="[151,636,1814,1838]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" kingdom="Animalia" order="Pantopoda" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="multiclavus">
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<emphasis id="B93BEAFC2919FFEC7ADC10489DA1F8A3" box="[151,446,1815,1838]" italics="true" pageId="94" pageNumber="95">Anoplodactylus multiclavus</emphasis>
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Child, 1977: 593
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</taxonomicName>
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.
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<bibRefCitation id="EFDE4B1F2919FFEC78CD10499F36F8A0" author="Child, C. A." box="[646,809,1814,1837]" pageId="94" pageNumber="95" pagination="1 - 86" refId="ref76281" refString="Child, C. A. (1979) Shallow-water Pycnogonida of the Isthmus of Panama and the coasts of Middle America. Smithsonian Contributions to Zoology, 293, 1 - 86." type="journal article" year="1979">Child 1979: 58</bibRefCitation>
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. Child 1982: 372.
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<bibRefCitation id="EFDE4B1F2919FFEC79A510499880F8A0" author="Stock, J. H." box="[1006,1183,1814,1837]" pageId="94" pageNumber="95" pagination="399 - 441" refId="ref77709" refString="Stock, J. H. (1986) Pycnogonida from the Caribbean and the straits of Florida. Bulletin of marine Science, 38 (3), 399 - 441." type="journal article" year="1986">Stock 1986: 404</bibRefCitation>
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, 439.
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</paragraph>
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<paragraph id="8BF036EE2919FFEC7ADC1002983CF8FA" blockId="94.[151,1425,1885,2027]" box="[151,1059,1885,1911]" pageId="94" pageNumber="95">
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<emphasis id="B93BEAFC2919FFEC7ADC10029D1BF8FA" bold="true" box="[151,260,1885,1911]" pageId="94" pageNumber="95">Material</emphasis>
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:
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<emphasis id="B93BEAFC2919FFEC7B5810029D2FF8FA" bold="true" box="[275,304,1885,1911]" pageId="94" pageNumber="95">14</emphasis>
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.—
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<materialsCitation id="3B273CB32919FFEC7B1210029800F8FA" box="[345,1055,1885,1911]" collectingDate="1986-01-07" collectionCode="SMF" elevation="3" pageId="94" pageNumber="95" specimenCode="SMF 1525" specimenCount="1" specimenCount-male="1">
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<specimenCount id="9D49FD672919FFEC7B1210029DBBF8FA" box="[345,420,1885,1911]" pageId="94" pageNumber="95" type="male">1 male</specimenCount>
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(gravid) (
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<specimenCode id="DBE99E952919FFEC785210029E84F8FA" box="[537,667,1885,1911]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="94" pageNumber="95">SMF 1525</specimenCode>
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), coral rubble,
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<quantity id="4CB79B0B2919FFEC790010029FB2F8FA" box="[843,941,1885,1911]" metricMagnitude="0" metricUnit="m" metricValue="3.25" metricValueMax="5.0" metricValueMin="1.5" pageId="94" pageNumber="95" unit="m" value="3.25" valueMax="5.0" valueMin="1.5">
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<elevation id="0062D1DD2919FFEC790010029FB2F8FA" box="[843,941,1885,1911]" metricMagnitude="0" metricUnit="m" metricValue="3.25" metricValueMax="5.0" metricValueMin="1.5" pageId="94" pageNumber="95" unit="m" value="3.25" valueMax="5.0" valueMin="1.5">1.5–5 m</elevation>
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</quantity>
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,
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<date id="FFF1102E2919FFEC79F110029800F8FA" box="[954,1055,1885,1911]" pageId="94" pageNumber="95" value="1986-01-07">
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<collectingDate id="EFB5E9C62919FFEC79F110029800F8FA" box="[954,1055,1885,1911]" pageId="94" pageNumber="95" value="1986-01-07">7.I.1986</collectingDate>
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</date>
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BF036EE2919FFED7A8D10DB9857FF3F" blockId="94.[151,1425,1885,2027]" lastBlockId="95.[198,1096,151,178]" lastPageId="95" lastPageNumber="96" pageId="94" pageNumber="95">
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<emphasis id="B93BEAFC2919FFEC7A8D10DB9CFCF813" bold="true" box="[198,227,1924,1950]" pageId="94" pageNumber="95">21</emphasis>
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.—
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<specimenCount id="9D49FD672919FFEC7B4310DB9D7DF813" box="[264,354,1924,1950]" pageId="94" pageNumber="95" type="male">4 males</specimenCount>
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(2 ov.), 3 fem. (
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<collectionCode id="ED5EAE2B2919FFEC785410DB9E44F813" box="[543,603,1924,1950]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="94" pageNumber="95">SMF</collectionCode>
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1526), on
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<taxonomicName id="4C4F4D6D2919FFEC789110DB9F86F810" box="[730,921,1924,1949]" class="Aves" genus="Digenia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="94" pageNumber="95" phylum="Chordata" rank="species" species="simplex">
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<emphasis id="B93BEAFC2919FFEC789110DB9F86F810" box="[730,921,1924,1949]" italics="true" pageId="94" pageNumber="95">Digenia simplex</emphasis>
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</taxonomicName>
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, 5l of substratum, mesolittoral,
|
||
<date id="FFF1102E2919FFEC7F5B10DB9995F813" box="[1296,1418,1924,1950]" pageId="94" pageNumber="95" value="1985-11-08">8.XI.1985</date>
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.
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<emphasis id="B93BEAFC2919FFEC7A8D10F59CFCF849" bold="true" box="[198,227,1962,1988]" pageId="94" pageNumber="95">22</emphasis>
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.—
|
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<specimenCount id="9D49FD672919FFEC7B4710F59D48F849" box="[268,343,1962,1988]" pageId="94" pageNumber="95" type="male">1 male</specimenCount>
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(ov.) (
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||
<collectionCode id="ED5EAE2B2919FFEC7BEF10F59DF8F849" box="[420,487,1962,1988]" country="Netherlands" httpUri="http://grbio.org/cool/a65z-wv6r" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="94" pageNumber="95">ZMA</collectionCode>
|
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3358) (
|
||
<collectionCode id="ED5EAE2B2919FFEC780910F59E9AF849" box="[578,645,1962,1988]" country="Netherlands" httpUri="http://grbio.org/cool/a65z-wv6r" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="94" pageNumber="95">ZMA</collectionCode>
|
||
3358), from
|
||
<taxonomicName id="4C4F4D6D2919FFEC795410F49F8DF849" authorityName="J. V. F. Lamouroux" authorityYear="1812" box="[799,914,1963,1988]" class="Ulvophyceae" family="Halimedaceae" genus="Halimeda" kingdom="Plantae" order="Bryopsidales" pageId="94" pageNumber="95" phylum="Chlorophyta" rank="genus">
|
||
<emphasis id="B93BEAFC2919FFEC795410F49F8DF849" box="[799,914,1963,1988]" italics="true" pageId="94" pageNumber="95">Halimeda</emphasis>
|
||
</taxonomicName>
|
||
on reef top, mesolittoral,
|
||
<date id="FFF1102E2919FFEC7E8910F59957F849" box="[1218,1352,1962,1988]" pageId="94" pageNumber="95" value="1986-04-14">14.IV.1986</date>
|
||
.
|
||
<emphasis id="B93BEAFC2919FFEC7A8D108E9CFCF866" bold="true" box="[198,227,2001,2027]" pageId="94" pageNumber="95">24</emphasis>
|
||
.—1 fem. (gravid) (
|
||
<collectionCode id="ED5EAE2B2919FFEC7B83108E9E1BF866" box="[456,516,2001,2027]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="94" pageNumber="95">SMF</collectionCode>
|
||
1527),
|
||
<taxonomicName id="4C4F4D6D2919FFEC7814108E9ECEF867" box="[607,721,2001,2026]" class="Liliopsida" family="Hydrocharitaceae" genus="Thalassia" kingdom="Plantae" order="Alismatales" pageId="94" pageNumber="95" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B93BEAFC2919FFEC7814108E9ECEF867" box="[607,721,2001,2026]" italics="true" pageId="94" pageNumber="95">Thalassia</emphasis>
|
||
</taxonomicName>
|
||
with corallinaceans and
|
||
<taxonomicName id="4C4F4D6D2919FFEC79BF108E9887F867" authorityName="Splettstoesser" authorityYear="1929" box="[1012,1176,2001,2026]" class="Hydrozoa" family="Thyroscyphidae" genus="Cnidoscyphus" kingdom="Animalia" order="Leptothecata" pageId="94" pageNumber="95" phylum="Cnidaria" rank="genus">
|
||
<emphasis id="B93BEAFC2919FFEC79BF108E9887F867" box="[1012,1176,2001,2026]" italics="true" pageId="94" pageNumber="95">Cnidoscyphus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<quantity id="4CB79B0B2919FFEC7EEC108E98ECF866" box="[1191,1267,2001,2027]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="94" pageNumber="95" unit="m" value="1.5" valueMax="2.0" valueMin="1.0">1–2 m</quantity>
|
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,
|
||
<date id="FFF1102E2919FFEC7F4B108E9991F866" box="[1280,1422,2001,2027]" pageId="94" pageNumber="95" value="1985-08-09">9.VIII.1985</date>
|
||
.
|
||
<emphasis id="B93BEAFC2918FFED7A8D17C89CFCFF3C" bold="true" box="[198,227,151,177]" pageId="95" pageNumber="96">37</emphasis>
|
||
.—
|
||
<specimenCount id="9D49FD672918FFED7B4717C79D48FF3F" box="[268,343,152,178]" pageId="95" pageNumber="96" type="male">1 male</specimenCount>
|
||
(ov.) (
|
||
<collectionCode id="ED5EAE2B2918FFED7BEF17C79E1BFF3F" box="[420,516,152,178]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="95" pageNumber="96">MNHN</collectionCode>
|
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), on algae, hydroids and detritus,
|
||
<quantity id="4CB79B0B2918FFED79C517C79FA2FF3F" box="[910,957,152,178]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="95" pageNumber="96" unit="m" value="6.0">6 m</quantity>
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,
|
||
<date id="FFF1102E2918FFED798117C7985BFF3F" box="[970,1092,152,178]" pageId="95" pageNumber="96" value="1985-06-04">4.VI.1985</date>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF3066662918FFED7ADC12239D1EFA54" ID-DOI="http://doi.org/10.5281/zenodo.5362510" ID-Zenodo-Dep="5362510" httpUri="https://zenodo.org/record/5362510/files/figure.png" pageId="95" pageNumber="96" startId="95.[151,257,1404,1428]" targetBox="[203,1391,229,1382]" targetPageId="95">
|
||
<paragraph id="8BF036EE2918FFED7ADC12239D1EFA54" blockId="95.[151,1436,1404,1497]" pageId="95" pageNumber="96">
|
||
<emphasis id="B93BEAFC2918FFED7ADC12239D35FA19" bold="true" box="[151,298,1404,1428]" pageId="95" pageNumber="96">FIGURE 50.</emphasis>
|
||
<taxonomicName id="4C4F4D6D2918FFED7B7812229EF1FA19" authority="Stock, 1954" authorityName="Stock" authorityYear="1954" box="[307,750,1405,1428]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" kingdom="Animalia" order="Pantopoda" pageId="95" pageNumber="96" phylum="Arthropoda" rank="species" species="glandulifer">
|
||
<emphasis id="B93BEAFC2918FFED7B7812229E7FFA19" box="[307,608,1405,1428]" italics="true" pageId="95" pageNumber="96">Anoplodactylus glandulifer</emphasis>
|
||
Stock, 1954
|
||
</taxonomicName>
|
||
, male: A ovigerous specimen, trunk lateral, B trunk dorsal, C chela, D oviger, E leg 3, F femur with two cement gland pores, G cement gland pores in lateral view, H tarsus and propodus.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C35565652918FFD27A8D1158983EFD7B" lastPageId="96" lastPageNumber="97" pageId="95" pageNumber="96" type="description">
|
||
<paragraph id="8BF036EE2918FFED7A8D11589E53F887" blockId="95.[151,1436,1543,2001]" pageId="95" pageNumber="96">
|
||
<emphasis id="B93BEAFC2918FFED7A8D11589DDEF9AC" bold="true" box="[198,449,1543,1569]" pageId="95" pageNumber="96">Description of male</emphasis>
|
||
: Dorsal trunk outline oval, rather robust; segments 3 and 4 fused. Crurigers 1–3 bearing dorsodistally feebly developed tubercles; all crurigers with 1 or 2 short dorso-distal setae; crurigers slightly longer than their diameter, anterior three separated by 1/2 of their diameter, the third separated from the fourth by 1/3 of their diameter. Ocular process three times higher than its diameter, anteriorly inclined, with large distinctly pigmented eyes; distal part of ocular process narrowing to a lappet; abdomen of similar length to ocular process, erect or slightly inclined to the rear, 4 short apical setae. Proboscis robust, distinctly curved and apically broadly rounded.
|
||
</paragraph>
|
||
<paragraph id="8BF036EE2918FFED7A8D10499DA1F85C" blockId="95.[151,1436,1543,2001]" pageId="95" pageNumber="96">
|
||
Scape of cheliphores slender, somewhat longer than proboscis; chela slender, its fingers strongly curved; cutting edge of immovable and moveable fingers armed with 3 (usually) and 4 pointed teeth respectively. Oviger 6-articled, relatively robust; 3
|
||
<superScript id="7C3A9BA62918FFED7814103C9E72F8FC" attach="left" box="[607,621,1891,1905]" fontSize="6" pageId="95" pageNumber="96">rd</superScript>
|
||
article the longest, 1.2 times as long as 2
|
||
<superScript id="7C3A9BA62918FFED7E23103C9867F8FC" attach="none" box="[1128,1144,1891,1905]" fontSize="6" pageId="95" pageNumber="96">nd</superScript>
|
||
; 4
|
||
<superScript id="7C3A9BA62918FFED7ED2103C98B9F8FC" attach="left" box="[1177,1190,1891,1905]" fontSize="6" pageId="95" pageNumber="96">th</superScript>
|
||
article of combined length of 5
|
||
<superScript id="7C3A9BA62918FFED7B5010D29D37F816" attach="left" box="[283,296,1933,1947]" fontSize="6" pageId="95" pageNumber="96">th</superScript>
|
||
and 6
|
||
<superScript id="7C3A9BA62918FFED7B3F10D29D9EF816" attach="none" box="[372,385,1933,1947]" fontSize="6" pageId="95" pageNumber="96">th</superScript>
|
||
; articles 5 and 6 with sparsely distributed setae distally directed; distal article slender, conical, apically pointed.
|
||
</paragraph>
|
||
<paragraph id="8BF036EE2927FFD27A8D17C79E44FE4D" blockId="96.[151,1437,152,1531]" pageId="96" pageNumber="97">Legs moderately robust, feebly setose; but femur and tibiae each with a long dorso-distal seta; coxae 1 and 3 subequal in length, about as long as their diameter, 2/3 of length of coxa 2; femur longest and most robust article, with dorso-median row of 2–5 cement gland ducts shaped like a capsule in its proximal half; tibiae 1 and 2 of equal length; tarsus as long as broad, with 5 ventro-distal setae; propodus with well developed heel, distally with two robust and one slender spine, as well some setae; sole almost straight, usually with 3 small, anteriorly-curved proximal spines; on distal 2/3 of sole with long cutting lamella, with adjacent setae on both sides; main claw robust, feebly curved, extending over proximal spines of sole but not reaching to heel; auxiliaries 1/8 length of terminal claw.</paragraph>
|
||
<paragraph id="8BF036EE2927FFD27A8D16929EA3FE6A" blockId="96.[151,1437,152,1531]" box="[198,700,461,487]" pageId="96" pageNumber="97">Colouration of most specimens pale green.</paragraph>
|
||
<paragraph id="8BF036EE2927FFD27A8D16AC9D0FFD0F" blockId="96.[151,1437,152,1531]" pageId="96" pageNumber="97">
|
||
<emphasis id="B93BEAFC2927FFD27A8D16AC9D62FD80" bold="true" box="[198,381,499,525]" pageId="96" pageNumber="97">Measurements</emphasis>
|
||
: Trunk length 0.78; trunk width 0.55 (across first crurigers); length of abdomen 0.23; length of cheliphore scape 0.25; length of chela 0.14; length of articles of leg 3: coxa 1—0.18; coxa 2—0.32; coxa 3—0.19; femur 0.56; tibia 1, 0.43; tibia 2, 0.43; tarsus 0.07; propodus 0.36; main claw 0.24; auxiliary claw 0.04.
|
||
</paragraph>
|
||
<paragraph id="8BF036EE2927FFD27A8D15D1983EFD7B" blockId="96.[151,1437,152,1531]" pageId="96" pageNumber="97">
|
||
<emphasis id="B93BEAFC2927FFD27A8D15D19D3EFD25" bold="true" box="[198,289,654,680]" pageId="96" pageNumber="97">Female</emphasis>
|
||
: Apart from sex-specific characters in habitus like male. The shorter propodal cutting lamella noted by
|
||
<bibRefCitation id="EFDE4B1F2927FFD27B4D15EA9DDBFD42" author="Child, C. A." box="[262,452,693,719]" pageId="96" pageNumber="97" pagination="1 - 86" refId="ref76281" refString="Child, C. A. (1979) Shallow-water Pycnogonida of the Isthmus of Panama and the coasts of Middle America. Smithsonian Contributions to Zoology, 293, 1 - 86." type="journal article" year="1979">Child (1979: 58</bibRefCitation>
|
||
,
|
||
<figureCitation id="13742A6B2927FFD27B9A15EA9E39FD42" box="[465,550,693,719]" captionStart="FIGURE 19" captionStartId="43.[151,255,668,692]" captionTargetBox="[229,1355,196,634]" captionTargetId="figure-485@43.[223,1363,176,662]" captionTargetPageId="43" captionText="FIGURE 19. Tanystylum geminum Stock, 1954: A male oviger, B female oviger." figureDoi="http://doi.org/10.5281/zenodo.5362472" httpUri="https://zenodo.org/record/5362472/files/figure.png" pageId="96" pageNumber="97">Fig. 19</figureCitation>
|
||
) on females from
|
||
<collectingCountry id="F358767E2927FFD2794A15EA9F41FD42" box="[769,862,693,719]" name="Panama" pageId="96" pageNumber="97">Panamá</collectingCountry>
|
||
could not be found on Colombian material. The development of this lamella in females conformed exactly with that of males.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35565652927FFD27A8D145D9941FA0A" pageId="96" pageNumber="97" type="discussion">
|
||
<paragraph id="8BF036EE2927FFD27A8D145D98A5FA9E" blockId="96.[151,1437,152,1531]" pageId="96" pageNumber="97">
|
||
<emphasis id="B93BEAFC2927FFD27A8D145D9D27FC91" bold="true" box="[198,312,770,796]" pageId="96" pageNumber="97">Remarks</emphasis>
|
||
:
|
||
<taxonomicName id="4C4F4D6D2927FFD27B00145D9F5CFC91" authority="STOCK, 1954" authorityName="Stock" authorityYear="1954" box="[331,835,770,796]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" kingdom="Animalia" order="Pantopoda" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="glandulifer">
|
||
<emphasis id="B93BEAFC2927FFD27B00145D9E92FC96" box="[331,653,770,795]" italics="true" pageId="96" pageNumber="97">Anoplodactylus glandulifer</emphasis>
|
||
STOCK, 1954
|
||
</taxonomicName>
|
||
from the tropical Indo-Pacific and
|
||
<taxonomicName id="4C4F4D6D2927FFD27EBA145D9D3CFCCE" authority="Child, 1977" authorityName="Child" authorityYear="1977" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" kingdom="Animalia" order="Pantopoda" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="multiclavus">
|
||
<emphasis id="B93BEAFC2927FFD27EBA145D9983FC96" box="[1265,1436,770,795]" italics="true" pageId="96" pageNumber="97">A. multiclavus</emphasis>
|
||
Child, 1977
|
||
</taxonomicName>
|
||
from the Caribbean were regarded as very closely related species. All characters used by Child (1977b: 593) to differentiate
|
||
<taxonomicName id="4C4F4D6D2927FFD27BA7140F9E6CFCE4" authorityName="Child" authorityYear="1977" box="[492,627,848,873]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" kingdom="Animalia" order="Pantopoda" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="multiclavus">
|
||
<emphasis id="B93BEAFC2927FFD27BA7140F9E6CFCE4" box="[492,627,848,873]" italics="true" pageId="96" pageNumber="97">multiclavus</emphasis>
|
||
</taxonomicName>
|
||
from Stock’s species proved to be variable and found in
|
||
<emphasis id="B93BEAFC2927FFD27F5E140F9983FCE4" box="[1301,1436,848,873]" italics="true" pageId="96" pageNumber="97">
|
||
<taxonomicName id="4C4F4D6D2927FFD27F5E140F9988FCE4" authorityName="Stock" authorityYear="1954" box="[1301,1431,848,873]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" kingdom="Animalia" order="Pantopoda" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="glandulifer">glandulifer</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
too, so it is now considered a junior synonym of
|
||
<taxonomicName id="4C4F4D6D2927FFD278B414289F97FC1D" authorityName="Stock" authorityYear="1954" box="[767,904,887,912]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" kingdom="Animalia" order="Pantopoda" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="glandulifer">
|
||
<emphasis id="B93BEAFC2927FFD278B414289F97FC1D" box="[767,904,887,912]" italics="true" pageId="96" pageNumber="97">glandulifer</emphasis>
|
||
</taxonomicName>
|
||
. The number of cement gland ducts in the
|
||
<typeStatus id="54F4884C2927FFD27ADC14C29CE2FC3A" box="[151,253,925,951]" pageId="96" pageNumber="97" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C4F4D6D2927FFD27B6F14C29DB9FC3B" authorityName="Stock" authorityYear="1954" box="[292,422,925,950]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" kingdom="Animalia" order="Pantopoda" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="glandulifer">
|
||
<emphasis id="B93BEAFC2927FFD27B6F14C29DB9FC3B" box="[292,422,925,950]" italics="true" pageId="96" pageNumber="97">glandulifer</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F358767E2927FFD27BA514C29E7AFC3A" box="[494,613,925,951]" name="Singapore" pageId="96" pageNumber="97">Singapore</collectingCountry>
|
||
varied on different legs between 2–4. Examination of further material from
|
||
<collectingCountry id="F358767E2927FFD27A9C149B9D3CFC53" box="[215,291,964,990]" name="Kenya" pageId="96" pageNumber="97">Kenya</collectingCountry>
|
||
and
|
||
<collectingCountry id="F358767E2927FFD27B17149B9DCFFC53" box="[348,464,964,990]" name="Colombia" pageId="96" pageNumber="97">Colombia</collectingCountry>
|
||
by the first author found a variability between 2–5 ducts. Contrary to the remark in the original description of
|
||
<emphasis id="B93BEAFC2927FFD27B9814B59E45FB8E" box="[467,602,1002,1027]" italics="true" pageId="96" pageNumber="97">
|
||
<taxonomicName id="4C4F4D6D2927FFD27B9814B59E49FB8E" authorityName="Stock" authorityYear="1954" box="[467,598,1002,1027]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" kingdom="Animalia" order="Pantopoda" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="glandulifer">glandulifer</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
the anterior 3 trunk segments in the
|
||
<typeStatus id="54F4884C2927FFD27E5A14B59867FB89" box="[1041,1144,1002,1028]" pageId="96" pageNumber="97" type="holotype">holotype</typeStatus>
|
||
are clearly separated. In the recent database by Bamber & El Nagar (2007) (http://www.marinespecies.org/pycnobase/ aphia.php?p=taxlist) the two species have already been synonymized. The species is possibly related to
|
||
<taxonomicName id="4C4F4D6D2927FFD27ADC13009E41FBF5" authority="Muller, 1990" authorityName="Muller" authorityYear="1990" box="[151,606,1118,1144]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" kingdom="Animalia" order="Pantopoda" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="bourboni">
|
||
<emphasis id="B93BEAFC2927FFD27ADC13009DA0FBF5" box="[151,447,1119,1144]" italics="true" pageId="96" pageNumber="97">Anoplodactylus bourboni</emphasis>
|
||
Müller, 1990
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F358767E2927FFD278EE13019F31FBF5" box="[677,814,1118,1144]" name="Réunion" pageId="96" pageNumber="97">La Réunion</collectingCountry>
|
||
in the tropical southern Indian Ocean. The habitus in both species is similar, but the proboscis of
|
||
<taxonomicName id="4C4F4D6D2927FFD278E513DA9F20FB13" authorityName="Muller" authorityYear="1990" box="[686,831,1157,1182]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" kingdom="Animalia" order="Pantopoda" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="bourboni">
|
||
<emphasis id="B93BEAFC2927FFD278E513DA9F20FB13" box="[686,831,1157,1182]" italics="true" pageId="96" pageNumber="97">A. bourboni</emphasis>
|
||
</taxonomicName>
|
||
is significantly longer. The strongly curved chela fingers in
|
||
<taxonomicName id="4C4F4D6D2927FFD27B5913F39DBEFB48" authorityName="Muller" authorityYear="1990" box="[274,417,1196,1221]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" kingdom="Animalia" order="Pantopoda" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="bourboni">
|
||
<emphasis id="B93BEAFC2927FFD27B5913F39DBEFB48" box="[274,417,1196,1221]" italics="true" pageId="96" pageNumber="97">A. bourboni</emphasis>
|
||
</taxonomicName>
|
||
are untoothed. The cement gland ducts on femur consist of 7 dorso-median cup-like structures, differing characteristically from those found in
|
||
<taxonomicName id="4C4F4D6D2927FFD27975138C9FC0FB61" authorityName="Stock" authorityYear="1954" box="[830,991,1235,1260]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" kingdom="Animalia" order="Pantopoda" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="glandulifer">
|
||
<emphasis id="B93BEAFC2927FFD27975138C9FC0FB61" box="[830,991,1235,1260]" italics="true" pageId="96" pageNumber="97">A. glandulifer</emphasis>
|
||
</taxonomicName>
|
||
. The propodal sole in
|
||
<taxonomicName id="4C4F4D6D2927FFD27EA8138C9971FB61" authorityName="Muller" authorityYear="1990" box="[1251,1390,1235,1260]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" kingdom="Animalia" order="Pantopoda" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="bourboni">
|
||
<emphasis id="B93BEAFC2927FFD27EA8138C9971FB61" box="[1251,1390,1235,1260]" italics="true" pageId="96" pageNumber="97">A. bourboni</emphasis>
|
||
</taxonomicName>
|
||
has 7 spines, and as a consequence the distal lamella is short (cf.
|
||
<bibRefCitation id="EFDE4B1F2927FFD2792913A6982FFA9E" author="Muller, H. - G." box="[866,1072,1273,1299]" pageId="96" pageNumber="97" pagination="97 - 102" refId="ref77229" refString="Muller, H. - G. (1990 e) Pycnogonida from coral reefs at Reunion Island, Southern Indian Ocean. Zoologische Abhandlungen aus dem Staatlichen Museum fur Tierkunde Dresden, 45 (9), 97 - 102." type="journal article" year="1990">Müller 1990e: 98</bibRefCitation>
|
||
,
|
||
<figureCitation id="13742A6B2927FFD27E7713A698B2FA9E" box="[1084,1197,1273,1299]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="8.[151,255,1534,1558]" captionStartId-1="14.[151,257,1668,1692]" captionStartId-2="15.[151,255,1877,1901]" captionStartId-3="17.[151,255,1964,1988]" captionStartId-4="18.[151,256,1779,1803]" captionStartId-5="21.[151,255,1929,1953]" captionTargetBox-0="[209,1375,774,1503]" captionTargetBox-1="[208,1376,539,1654]" captionTargetBox-2="[167,1423,586,1870]" captionTargetBox-3="[270,1317,590,1908]" captionTargetBox-4="[198,1391,183,1753]" captionTargetBox-5="[162,1424,183,1911]" captionTargetId-0="figure-249@8.[203,1383,759,1527]" captionTargetId-1="figure-304@14.[204,1380,524,1662]" captionTargetId-2="figure-215@15.[155,1431,569,1871]" captionTargetId-3="figure-201@17.[226,1319,509,1985]" captionTargetId-4="figure-133@18.[190,1397,176,1773]" captionTargetId-5="figure-62@21.[162,1447,156,1953]" captionTargetPageId-0="8" captionTargetPageId-1="14" captionTargetPageId-2="15" captionTargetPageId-3="17" captionTargetPageId-4="18" captionTargetPageId-5="21" captionText-0="FIGURE 1. A, External morphology of a pycnogonid, exemplified by Ascorhynchus castelli (Dohrn, 1881) male, after Müller & Roth (1990: 164), slightly modified; abd—abdomen, ch cheliphore, l3 leg 3, lp—cruriger of leg 2, pl—palp, pr—proboscis, ov—oviger; B, proximal part of leg 3 of Anoplodactylus guachaquitae n. sp., male: cd—cement gland duct on dorsal face of femur." captionText-1="FIGURE 2. Achelia gracilis Verrill, 1900. A, male dorsal, B, female dorsal, C, male lateral, D, male cheliphore, E, female oviger, F, male oviger. Scale 200 mm for A–C, 50 mm for D–E, 100 mm for F." captionText-2="FIGURE 3. Achelia gracilis Verrill, 1900. A, male palp, B, leg 3 of male, C, leg 3 of female, D, cement gland duct, E, tarsus and propodus of male leg 3." captionText-3="FIGURE 4. Achelia sawayai Marcus, 1940. A, male dorsal, B, male dorsal, C, female dorsal, D, female dorsal, E, male lateral, F, male cheliphore, G, male palp." captionText-4="FIGURE 5. Achelia sawayai Marcus, 1940. A, male leg 3, B, cement gland duct on male leg enlarged, C, tarsus and propodus of male leg 3, D, female leg 3, E, tarsus and propodus of female leg 3, F, male oviger, G, female oviger." captionText-5="FIGURE 6. Ammothella appendiculata (Dohrn, 1881). A, male dorsal, B, male lateral, C, male cheliphore, D, female oviger, E, distal female oviger articles, F, male oviger, G, distal articles of male oviger, H, male palp, I, male leg 3, J, femur with eggs of female." figureDoi-0="http://doi.org/10.5281/zenodo.5317872" figureDoi-1="http://doi.org/10.5281/zenodo.5317874" figureDoi-2="http://doi.org/10.5281/zenodo.5317878" figureDoi-3="http://doi.org/10.5281/zenodo.5317880" figureDoi-4="http://doi.org/10.5281/zenodo.5317884" figureDoi-5="http://doi.org/10.5281/zenodo.5317888" httpUri-0="https://zenodo.org/record/5317872/files/figure.png" httpUri-1="https://zenodo.org/record/5317874/files/figure.png" httpUri-2="https://zenodo.org/record/5317878/files/figure.png" httpUri-3="https://zenodo.org/record/5317880/files/figure.png" httpUri-4="https://zenodo.org/record/5317884/files/figure.png" httpUri-5="https://zenodo.org/record/5317888/files/figure.png" pageId="96" pageNumber="97">Figs. 1–6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF036EE2927FFD27A8D127F9941FA0A" blockId="96.[151,1437,152,1531]" pageId="96" pageNumber="97">
|
||
In the research area
|
||
<taxonomicName id="4C4F4D6D2927FFD27B9A127F9E64FAB4" authorityName="Stock" authorityYear="1954" box="[465,635,1312,1337]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" kingdom="Animalia" order="Pantopoda" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="glandulifer">
|
||
<emphasis id="B93BEAFC2927FFD27B9A127F9E64FAB4" box="[465,635,1312,1337]" italics="true" pageId="96" pageNumber="97">A. glandulifer</emphasis>
|
||
</taxonomicName>
|
||
was found sporadically, without any pronounced preference of substratum or a distinct reproduction period. The vertical distribution ranges from the intertidal zone to
|
||
<specimenCount id="9D49FD672927FFD27F2C12199983FAED" box="[1383,1436,1350,1376]" pageId="96" pageNumber="97" type="generic">
|
||
<quantity id="4CB79B0B2927FFD27F2C12199987FAED" box="[1383,1432,1350,1376]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="96" pageNumber="97" unit="m" value="6.0">6 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
Data in the literature clearly indicate that the species is restricted to the uppermost shallow water region.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35565652927FFD27A8D12CC9F1AFA76" pageId="96" pageNumber="97" type="distribution">
|
||
<paragraph id="8BF036EE2927FFD27A8D12CC9F1AFA76" blockId="96.[151,1437,152,1531]" pageId="96" pageNumber="97">
|
||
<emphasis id="B93BEAFC2927FFD27A8D12CC9D40FA20" bold="true" box="[198,351,1427,1453]" pageId="96" pageNumber="97">Distribution</emphasis>
|
||
: Tropical Indo-Pacific and Caribbean. In the Caribbean the species was known form the
|
||
<collectingCountry id="F358767E2927FFD27ADC12E49D62FA58" box="[151,381,1467,1493]" name="United States Virgin Islands" pageId="96" pageNumber="97">U.S. Virgin Islands</collectingCountry>
|
||
(Child 1977b: 593), the Atlantic coast of
|
||
<collectingCountry id="F358767E2927FFD2793212E49FC8FA58" box="[889,983,1467,1493]" name="Panama" pageId="96" pageNumber="97">Panamá</collectingCountry>
|
||
and
|
||
<collectingCountry id="F358767E2927FFD27E5812E49871FA58" box="[1043,1134,1467,1493]" name="Mexico" pageId="96" pageNumber="97">Mexico</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFDE4B1F2927FFD27E3412E49926FA58" author="Child, C. A." box="[1151,1337,1467,1493]" pageId="96" pageNumber="97" pagination="1 - 86" refId="ref76281" refString="Child, C. A. (1979) Shallow-water Pycnogonida of the Isthmus of Panama and the coasts of Middle America. Smithsonian Contributions to Zoology, 293, 1 - 86." type="journal article" year="1979">Child 1979: 58</bibRefCitation>
|
||
),
|
||
<collectingCountry id="F358767E2927FFD27F1B12E49983FA58" box="[1360,1436,1467,1493]" name="Belize" pageId="96" pageNumber="97">Belize</collectingCountry>
|
||
(Child 1982: 372) and
|
||
<collectingCountry id="F358767E2927FFD27BEA12BE9E3CFA76" box="[417,547,1505,1531]" name="Saint Vincent and the Grenadines" pageId="96" pageNumber="97">St. Vincent</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFDE4B1F2927FFD2787912BE9EE7FA76" author="Stock, J. H." box="[562,760,1505,1531]" pageId="96" pageNumber="97" pagination="399 - 441" refId="ref77709" refString="Stock, J. H. (1986) Pycnogonida from the Caribbean and the straits of Florida. Bulletin of marine Science, 38 (3), 399 - 441." type="journal article" year="1986">Stock 1986: 399</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |