treatments-xml/data/03/E6/87/03E687BAFFDAF300FF5EFF4C0D36C538.xml
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<document ID-DOI="10.11646/zootaxa.3219.1.1" ID-GBIF-Dataset="3fa772dc-c5b2-40c9-b28e-cd067ca9a3f6" ID-ISSN="1175-5326" ID-Zenodo-Dep="280223" checkinTime="1460447300036" checkinUser="plazi" docAuthor="Fikáček, Martin, Jia, Fenglong &amp; Prokin, Alexander" docDate="2012" docId="03E687BAFFDAF300FF5EFF4C0D36C538" docLanguage="en" docName="zt03219p053.pdf" docOrigin="Zootaxa 3219" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.3:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="3" docTitle="Pachysternum coomani Orchymont 1926" docType="treatment" docVersion="13" lastPageNumber="25" masterDocId="FFDFFFC2FFCCF318FFC9FFDB0C07C424" masterDocTitle="A review of the Asian species of the genus Pachysternum (Coleoptera: Hydrophilidae: Sphaeridiinae)" masterLastPageNumber="53" masterPageNumber="1" pageNumber="23" updateTime="1678199402054" updateUser="tatiana">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A review of the Asian species of the genus Pachysternum (Coleoptera: Hydrophilidae: Sphaeridiinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fikáček, Martin</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jia, Fenglong</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Prokin, Alexander</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2012</mods:date>
<mods:detail type="pubDate">
<mods:number>2012-03-02</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>3219</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>53</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.3219.1.1</mods:identifier>
<mods:identifier type="GBIF-Dataset">3fa772dc-c5b2-40c9-b28e-cd067ca9a3f6</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">280223</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6180434" ID-GBIF-Taxon="119509787" ID-Zenodo-Dep="6180434" LSID="urn:lsid:plazi:treatment:03E687BAFFDAF300FF5EFF4C0D36C538" httpUri="http://treatment.plazi.org/id/03E687BAFFDAF300FF5EFF4C0D36C538" lastPageId="24" lastPageNumber="25" pageId="22" pageNumber="23">
<subSubSection box="[151,661,151,177]" pageId="22" pageNumber="23" type="nomenclature">
<paragraph blockId="22.[151,661,151,210]" box="[151,661,151,177]" pageId="22" pageNumber="23">
<heading bold="true" box="[151,661,151,177]" fontSize="11" level="1" pageId="22" pageNumber="23" reason="1">
<taxonomicName ID-CoL="4BV52" authority="Orchymont, 1926" authorityName="Orchymont" authorityYear="1926" box="[151,661,151,177]" class="Insecta" family="Hydrophilidae" genus="Pachysternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="coomani">
<emphasis bold="true" box="[151,661,151,177]" pageId="22" pageNumber="23">
<emphasis bold="true" box="[151,436,151,177]" italics="true" pageId="22" pageNumber="23">Pachysternum coomani</emphasis>
Orchymont,
<number box="[603,661,151,177]" pageId="22" pageNumber="23" value="1926.0">1926</number>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,302,186,210]" pageId="22" pageNumber="23" type="description">
<paragraph blockId="22.[151,661,151,210]" box="[151,302,186,210]" pageId="22" pageNumber="23">
(
<figureCitation box="[159,253,186,210]" captionStart="FIGURES 29 32" captionStartId="8.[151,264,1975,1998]" captionTargetBox="[162,1428,577,1952]" captionTargetId="figure@8.[151,1436,561,1962]" captionTargetPageId="8" captionText="FIGURES 29 32. Male genitalia of Asian Pachysternum (a: tegmen, b: median lobe, c: apical portionof median lobe, d: sternite 9). 29: P. c a rd o n i, 30: P. coomani, 31: P. haemorrhoum; 32: P. nigrovitattum." httpUri="https://zenodo.org/record/280227/files/figure.png" pageId="22" pageNumber="23">Figs. 30</figureCitation>
,
<figureCitation box="[264,293,186,210]" captionStart="FIGURE 54" captionStartId="23.[151,250,790,813]" captionTargetBox="[177,1413,197,757]" captionTargetId="figure@23.[146,1431,186,774]" captionText="FIGURE 54. Pachysternum coomani, general habitus (a, cd: dorsal view, b: laterodorsal view). Localities of the specimens: ab: Malaysia, Kedah (male); c: same locality (female); d: Thailand, Phetchabun (male)." pageId="22" pageNumber="23">54</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="22" pageNumber="23" type="reference_group">
<paragraph blockId="22.[151,1436,251,335]" box="[151,643,251,274]" pageId="22" pageNumber="23">
<treatmentCitationGroup box="[151,643,251,274]" pageId="22" pageNumber="23">
<taxonomicName ID-CoL="4BV52" authority="Orchymont, 1926a: 221" authorityName="Orchymont" authorityPageNumber="221" authorityYear="1926" box="[151,639,251,274]" class="Insecta" family="Hydrophilidae" genus="Pachysternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="coomani">
<emphasis box="[151,389,252,274]" italics="true" pageId="22" pageNumber="23">Pachysternum coomani</emphasis>
<treatmentCitation author="Orchymont" box="[395,639,251,273]" page="221" pageId="22" pageNumber="23" year="1926">
<bibRefCitation author="Orchymont" box="[395,589,251,273]" pageId="22" pageNumber="23" pagination="201" refId="ref42865" refString="d'Orchymont, A. (1926 a) Contribution a l'etude des Hydrophilides VI. Bulletin et Annales de la Societe Entomologique de Belgique 66, 201 - 248." type="journal article" year="1926" yearSuffix="a">
Orchymont,
<number box="[523,577,251,273]" pageId="22" pageNumber="23" value="1926.0">1926</number>
a
</bibRefCitation>
:
<number box="[598,639,252,273]" pageId="22" pageNumber="23" value="221.0">221</number>
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="22.[151,1436,251,335]" pageId="22" pageNumber="23">
<treatmentCitationGroup pageId="22" pageNumber="23">
<taxonomicName ID-CoL="4BV52" authority="Orchymont" authorityName="Orchymont" authorityYear="1928" box="[151,743,282,304]" class="Insecta" family="Hydrophilidae" genus="Pachysternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="coomani">
<emphasis box="[151,389,282,304]" italics="true" pageId="22" pageNumber="23">Pachysternum coomani</emphasis>
:
<treatmentCitation box="[403,747,282,304]" pageId="22" pageNumber="23">
<bibRefCitation author="Orchymont" box="[403,743,282,304]" pageId="22" pageNumber="23" refId="ref42920" refString="d'Orchymont, A. (1928) Catalogue of Indian Insects. Part 14 - Palpicornia. Government of India, Central Publication Branch, Calcutta, 146 pp." type="book" year="1928">
Orchymont (
<number box="[532,587,283,304]" pageId="22" pageNumber="23" value="1928.0">1928</number>
:
<number box="[597,624,283,304]" pageId="22" pageNumber="23" value="83.0">83</number>
, catalogue)
</bibRefCitation>
;
</treatmentCitation>
</taxonomicName>
<treatmentCitation box="[754,1281,282,304]" pageId="22" pageNumber="23">
<bibRefCitation author="Mukhopadhyay" box="[754,1277,282,304]" pageId="22" pageNumber="23" pagination="449 - 458" refId="ref42687" refString="Mukhopadhyay, P. &amp; Sengupta, S. (2004) Insecta: Coleoptera: Polyphaga: Hydrophiloidea: Hydrophilidae. Pp. 449 - 458. In: Fauna of Manipur, State Fauna Series, volume 10. Zoological Survey of India." type="journal article" year="2004">
Mukhopadhyay &amp; Sengupta (
<number box="[1053,1109,283,304]" pageId="22" pageNumber="23" value="2004.0">2004</number>
:
<number box="[1118,1158,282,304]" pageId="22" pageNumber="23" value="453.0">453</number>
, faunistics)
</bibRefCitation>
;
</treatmentCitation>
<treatmentCitation pageId="22" pageNumber="23">
<bibRefCitation author="Hansen" pageId="22" pageNumber="23" refId="ref41882" refString="Hansen, M. (1999) World catalogue of Insects. Volume 2. Hydrophiloidea (s. str.) (Coleoptera). Apollo Books, Stenstrup, 416 pp." type="book" year="1999">
Hansen (
<number box="[1377,1432,283,304]" pageId="22" pageNumber="23" value="1999.0">1999</number>
:
<number box="[199,240,314,335]" pageId="22" pageNumber="23" value="307.0">307</number>
, catalogue)
</bibRefCitation>
.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="22" pageNumber="23" type="materials_examined">
<paragraph blockId="22.[151,1437,373,1887]" box="[151,1041,373,398]" pageId="22" pageNumber="23">
<emphasis bold="true" box="[151,304,373,398]" pageId="22" pageNumber="23">
<typeStatus box="[151,207,373,397]" pageId="22" pageNumber="23">Type</typeStatus>
locality.
</emphasis>
<collectingCountry box="[312,408,373,397]" name="Vietnam" pageId="22" pageNumber="23">Vietnam</collectingCountry>
, Hoa Binh province [coordinates ca.
<geoCoordinate box="[815,910,373,397]" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="925" value="20.833334">
<number box="[815,845,373,397]" pageId="22" pageNumber="23" value="20.0">20</number>
°
<number box="[855,885,373,397]" pageId="22" pageNumber="23" value="50.0">50</number>
'N
</geoCoordinate>
,
<geoCoordinate box="[922,1030,373,397]" direction="east" orientation="longitude" pageId="22" pageNumber="23" precision="925" value="105.333336">
<number box="[922,967,373,397]" pageId="22" pageNumber="23" value="105.0">105</number>
°
<number box="[976,1008,373,397]" pageId="22" pageNumber="23" value="20.0">20</number>
'E
</geoCoordinate>
].
</paragraph>
<paragraph blockId="22.[151,1437,373,1887]" pageId="22" pageNumber="23">
<emphasis bold="true" box="[199,494,408,433]" pageId="22" pageNumber="23">
<typeStatus box="[199,258,409,433]" pageId="22" pageNumber="23">Type</typeStatus>
material examined.
</emphasis>
<materialsCitation ID-GBIF-Occurrence="3499755330" collectionCode="IRSNB" collectorName="d'Orchym" location="Tonkin" pageId="22" pageNumber="23" specimenCount="1" specimenCount-female="1" stateProvince="Hoa Binh" typeStatus="holotype">
<typeStatus box="[501,605,409,433]" pageId="22" pageNumber="23">Holotype</typeStatus>
:
<specimenCount box="[620,716,409,433]" pageId="22" pageNumber="23" type="female">
<number box="[620,634,409,433]" pageId="22" pageNumber="23" value="1.0">1</number>
female
</specimenCount>
(
<collectionCode box="[731,820,409,433]" collectionName="Belgium, Brussels, Institut Royal des Sciences Naturelles de Belgique" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="22" pageNumber="23">IRSNB</collectionCode>
): “
<collectingRegion box="[849,959,409,433]" country="Vietnam" name="Hoa Binh" pageId="22" pageNumber="23">Hoa Binh</collectingRegion>
/
<location LSID="urn:lsid:plazi:treatment:03E687BAFFDAF300FF5EFF4C0D36C538:8E906077FFDAF30EFC1DFE420824C595" box="[980,1059,409,433]" name="Tonkin" pageId="22" pageNumber="23" stateProvince="Hoa Binh">Tonkin</location>
/
<location LSID="urn:lsid:plazi:treatment:03E687BAFFDAF300FF5EFF4C0D36C538:8E906077FFDAF30EFBF1FE4208BFC595" box="[1080,1208,409,433]" name="de Cooman" pageId="22" pageNumber="23" stateProvince="Hoa Binh">de Cooman</location>
// Coll.
<collectorName box="[1299,1418,409,433]" pageId="22" pageNumber="23">dOrchym</collectorName>
. / /
<taxonomicName box="[165,324,445,469]" class="Insecta" family="Hydrophilidae" genus="Pachysternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="51" phylum="Arthropoda" rank="genus">Pachysternum</taxonomicName>
/
<taxonomicName box="[346,442,445,469]" class="Insecta" family="Hydrophilidae" genus="Pachysternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="coomani">coomani</taxonomicName>
/ Type”.
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3712023302" box="[546,1288,445,470]" collectingDate="2009-11-07" collectingDateMax="2009-11-22" collectingDateMin="2009-11-07" collectionCode="IRSNB, NMPC, ASKC" collectorName="U. Schmidt" country="Malaysia" latitude="6.366667" location="Langkawi" longLatPrecision="929" longitude="99.675" municipality="Kedah" pageId="22" pageNumber="23" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="paratype">
<typeStatus box="[546,655,446,470]" pageId="22" pageNumber="23">Paratypes</typeStatus>
:
<specimenCount box="[670,781,445,469]" pageId="22" pageNumber="23" type="female">
<number box="[670,684,445,469]" pageId="22" pageNumber="23" value="2.0">2</number>
females
</specimenCount>
,
<number box="[792,806,445,469]" pageId="22" pageNumber="23" value="4.0">4</number>
spec. (
<collectionCode box="[885,974,445,469]" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="22" pageNumber="23">IRSNB</collectionCode>
): same data as the holotype.
</materialsCitation>
</paragraph>
<paragraph blockId="22.[151,1437,373,1887]" pageId="22" pageNumber="23">
<emphasis bold="true" box="[199,833,480,505]" pageId="22" pageNumber="23">
Additional material examined.
<materialsCitation ID-GBIF-Occurrence="3712023301" collectingDate="2009-11-07" collectingDateMax="2009-11-22" collectingDateMin="2009-11-07" collectionCode="IRSNB, NMPC, ASKC" collectorName="U. Schmidt" country="Malaysia" latitude="6.366667" location="Langkawi" longLatPrecision="929" longitude="99.675" municipality="Kedah" pageId="22" pageNumber="23" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="paratype">
<collectingCountry box="[574,728,480,505]" name="Malaysia" pageId="22" pageNumber="23">MALAYSIA</collectingCountry>
:
<collectingMunicipality box="[745,828,481,505]" pageId="22" pageNumber="23">Kedah</collectingMunicipality>
:
<specimenCount box="[844,925,481,505]" pageId="22" pageNumber="23" type="male">
<number box="[844,858,481,505]" pageId="22" pageNumber="23" value="1.0">1</number>
male
</specimenCount>
,
<number box="[941,955,481,505]" pageId="22" pageNumber="23" value="4.0">4</number>
spec. (
<collectionCode box="[1041,1125,481,505]" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="22" pageNumber="23">NMPC</collectionCode>
,
<collectionCode box="[1140,1220,481,505]" pageId="22" pageNumber="23">ASKC</collectionCode>
):
<collectingCountry box="[1242,1306,481,505]" name="Iceland" pageId="22" pageNumber="23">Pulau</collectingCountry>
<location LSID="urn:lsid:plazi:treatment:03E687BAFFDAF300FF5EFF4C0D36C538:8E906077FFDAF30EFAEDFE3A0990C5DD" box="[1316,1431,481,505]" country="Malaysia" latitude="6.366667" longLatPrecision="929" longitude="99.675" municipality="Kedah" name="Langkawi" pageId="22" pageNumber="23">Langkawi</location>
, environs of hotel
<location LSID="urn:lsid:plazi:treatment:03E687BAFFDAF300FF5EFF4C0D36C538:8E906077FFDAF30EFE92FDDE0DB5C639" box="[347,434,517,541]" country="Malaysia" latitude="6.366667" longLatPrecision="929" longitude="99.675" municipality="Kedah" name="Berjaya" pageId="22" pageNumber="23">Berjaya</location>
[=
<location LSID="urn:lsid:plazi:treatment:03E687BAFFDAF300FF5EFF4C0D36C538:8E906077FFDAF30EFE10FDDE0E51C639" box="[473,598,517,541]" country="Malaysia" latitude="6.366667" longLatPrecision="929" longitude="99.675" municipality="Kedah" name="Pantai Kok" pageId="22" pageNumber="23">Pantai Kok</location>
village, coordinates ca.
<geoCoordinate box="[870,951,517,541]" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="925" value="6.366667">
<number box="[870,884,517,541]" pageId="22" pageNumber="23" value="6.0">6</number>
°
<number box="[894,925,517,541]" pageId="22" pageNumber="23" value="22.0">22</number>
'N
</geoCoordinate>
,
<geoCoordinate box="[963,1078,517,541]" direction="east" orientation="longitude" pageId="22" pageNumber="23" precision="92" value="99.675">
<number box="[963,993,517,541]" pageId="22" pageNumber="23" value="99.0">99</number>
°
<number box="[1002,1056,517,541]" pageId="22" pageNumber="23" value="40.5">40.5</number>
'E
</geoCoordinate>
],
<date box="[1096,1247,517,541]" pageId="22" pageNumber="23" value="2009-11-07" valueMax="2009-11-22" valueMin="2009-11-07">
<collectingDate box="[1096,1247,517,541]" pageId="22" pageNumber="23" value="2009-11-07" valueMax="2009-11-22" valueMin="2009-11-07">
<number box="[1096,1156,517,541]" pageId="22" pageNumber="23" value="14.5" valueMax="22.0" valueMin="7.0">722</number>
.xi.
<number box="[1186,1247,517,541]" pageId="22" pageNumber="23" value="2009.0">2009</number>
</collectingDate>
</date>
, lgt.
<collectorName box="[1301,1433,517,541]" pageId="22" pageNumber="23">U. Schmidt</collectorName>
.
</materialsCitation>
</emphasis>
<materialsCitation ID-GBIF-Occurrence="3499755314" collectingDate="1989-11-23" collectingDateMax="1989-11-25" collectingDateMin="1989-11-23" collectionCode="FHGC" country="Myanmar" latitude="21.083334" location="Kinda" longLatPrecision="1265" longitude="96.316666" municipality="Shan" pageId="22" pageNumber="23" specimenCount="1" specimenCount-male="1">
<emphasis bold="true" box="[151,380,552,577]" pageId="22" pageNumber="23">
<collectingCountry box="[151,307,552,577]" name="Myanmar" pageId="22" pageNumber="23">MYANMAR</collectingCountry>
:
<collectingMunicipality box="[319,380,552,577]" pageId="22" pageNumber="23">Shan</collectingMunicipality>
</emphasis>
:
<specimenCount box="[395,473,553,577]" pageId="22" pageNumber="23" type="male">
<number box="[395,409,553,577]" pageId="22" pageNumber="23" value="1.0">1</number>
male
</specimenCount>
,
<number box="[485,499,553,577]" pageId="22" pageNumber="23" value="1.0">1</number>
spec. (
<collectionCode box="[578,660,553,577]" pageId="22" pageNumber="23">FHGC</collectionCode>
):
<location LSID="urn:lsid:plazi:treatment:03E687BAFFDAF300FF5EFF4C0D36C538:8E906077FFDAF30EFD6FFDF20EEDC665" box="[678,746,553,577]" country="Myanmar" latitude="21.083334" longLatPrecision="1265" longitude="96.316666" municipality="Shan" name="Kinda" pageId="22" pageNumber="23">Kinda</location>
dam,
<location LSID="urn:lsid:plazi:treatment:03E687BAFFDAF300FF5EFF4C0D36C538:8E906077FFDAF30EFCF8FDF20F8CC665" box="[817,907,553,577]" country="Myanmar" latitude="21.083334" longLatPrecision="1265" longitude="96.316666" municipality="Shan" name="Palaung" pageId="22" pageNumber="23">Palaung</location>
river mouth [coordinates ca.
<geoCoordinate box="[1237,1318,553,577]" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="925" value="21.083334">
<number box="[1237,1267,553,577]" pageId="22" pageNumber="23" value="21.0">21</number>
°
<number box="[1277,1291,553,577]" pageId="22" pageNumber="23" value="5.0">5</number>
'N
</geoCoordinate>
,
<geoCoordinate box="[1330,1424,553,577]" direction="east" orientation="longitude" pageId="22" pageNumber="23" precision="925" value="96.316666">
<number box="[1330,1362,553,577]" pageId="22" pageNumber="23" value="96.0">96</number>
°
<number box="[1372,1401,553,577]" pageId="22" pageNumber="23" value="19.0">19</number>
'E
</geoCoordinate>
],
<date box="[151,316,589,613]" pageId="22" pageNumber="23" value="1989-11-23" valueMax="1989-11-25" valueMin="1989-11-23">
<collectingDate box="[151,316,589,613]" pageId="22" pageNumber="23" value="1989-11-23" valueMax="1989-11-25" valueMin="1989-11-23">
<number box="[151,225,589,613]" pageId="22" pageNumber="23" value="24.0" valueMax="25.0" valueMin="23.0">2325</number>
.xi.
<number box="[256,316,589,613]" pageId="22" pageNumber="23" value="1989.0">1989</number>
</collectingDate>
</date>
, lgt.
<location LSID="urn:lsid:plazi:treatment:03E687BAFFDAF300FF5EFF4C0D36C538:8E906077FFDAF30EFEBFFD960DC3C641" box="[374,452,589,613]" country="Myanmar" latitude="21.083334" longLatPrecision="1265" longitude="96.316666" municipality="Shan" name="Tobias" pageId="22" pageNumber="23">Tobias</location>
.
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3499755333" collectingDate="1987-08" collectionCode="FHGC" collectorName="W. Thielen" country="Thailand" elevation="120" latitude="16.783333" location="Phetchabun" longLatPrecision="925" longitude="101.24166" municipality="Phetchabun" pageId="22" pageNumber="23" specimenCount="1" specimenCount-male="1">
<emphasis bold="true" box="[465,786,589,614]" pageId="22" pageNumber="23">
<collectingCountry box="[465,620,589,614]" name="Thailand" pageId="22" pageNumber="23">THAILAND</collectingCountry>
:
<collectingMunicipality box="[634,781,589,613]" pageId="22" pageNumber="23">Phetchabun</collectingMunicipality>
:
</emphasis>
<specimenCount box="[796,873,589,613]" pageId="22" pageNumber="23" type="male">
<number box="[796,810,589,613]" pageId="22" pageNumber="23" value="1.0">1</number>
male
</specimenCount>
(
<collectionCode box="[890,972,589,613]" pageId="22" pageNumber="23">FHGC</collectionCode>
):
<locationDeviation bearing="0.0" box="[992,1108,589,613]" distance="120000.0" pageId="22" pageNumber="23">
<quantity box="[992,1079,589,613]" metricMagnitude="5" metricUnit="m" metricValue="1.2" pageId="22" pageNumber="23" unit="km" value="120.0">120 km</quantity>
N
</locationDeviation>
<location LSID="urn:lsid:plazi:treatment:03E687BAFFDAF300FF5EFF4C0D36C538:8E906077FFDAF30EFB97FD9608E3C641" box="[1118,1252,589,613]" country="Thailand" latitude="16.783333" longLatPrecision="925" longitude="101.24166" municipality="Phetchabun" name="Phetchabun" pageId="22" pageNumber="23">Phetchabun</location>
,
<location LSID="urn:lsid:plazi:treatment:03E687BAFFDAF300FF5EFF4C0D36C538:8E906077FFDAF30EFB39FD95095CC641" box="[1264,1371,589,613]" country="Thailand" latitude="16.783333" longLatPrecision="925" longitude="101.24166" municipality="Phetchabun" name="Lom Sak" pageId="22" pageNumber="23">Lom Sak</location>
, elev.
<quantity box="[151,225,625,649]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="22" pageNumber="23" unit="m" value="120.0">
<elevation box="[151,225,625,649]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="22" pageNumber="23" unit="m" value="120.0">120 m</elevation>
</quantity>
[coordinates ca.
<geoCoordinate box="[428,520,625,649]" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="925" value="16.783333">
<number box="[428,456,625,649]" pageId="22" pageNumber="23" value="16.0">16</number>
°
<number box="[465,495,625,649]" pageId="22" pageNumber="23" value="47.0">47</number>
'N
</geoCoordinate>
<geoCoordinate box="[531,678,625,649]" direction="east" orientation="longitude" pageId="22" pageNumber="23" precision="15" value="101.24166">
<number box="[531,574,625,649]" pageId="22" pageNumber="23" value="101.0">101</number>
°
<number box="[584,613,625,649]" pageId="22" pageNumber="23" value="14.0">14</number>
'
<number box="[616,648,625,649]" pageId="22" pageNumber="23" value="30.0">30</number>
''E
</geoCoordinate>
],
<date box="[700,803,625,649]" pageId="22" pageNumber="23" value="1987-08">
<collectingDate box="[700,803,625,649]" pageId="22" pageNumber="23" value="1987-08">
viii.
<number box="[744,803,625,649]" pageId="22" pageNumber="23" value="1987.0">1987</number>
</collectingDate>
</date>
, lgt.
<collectorName box="[865,992,625,649]" pageId="22" pageNumber="23">W. Thielen</collectorName>
;
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3499755345" collectionCode="IRSNB" country="Vietnam" latitude="10.775" location="Ho Chi Minh" longLatPrecision="129" longitude="106.691666" pageId="22" pageNumber="23" specimenCount="1" stateProvince="Ho Chi Minh">
<emphasis bold="true" box="[1012,1343,625,650]" pageId="22" pageNumber="23">
<collectingCountry box="[1012,1152,625,650]" name="Vietnam" pageId="22" pageNumber="23">VIETNAM</collectingCountry>
:
<collectingRegion box="[1168,1339,625,650]" country="Vietnam" name="Ho Chi Minh City" pageId="22" pageNumber="23">Ho Chi Minh</collectingRegion>
:
</emphasis>
<specimenCount box="[1354,1436,625,649]" pageId="22" pageNumber="23" type="generic">
<number box="[1354,1368,625,649]" pageId="22" pageNumber="23" value="1.0">1</number>
spec.
</specimenCount>
(
<collectionCode box="[159,248,661,685]" collectionName="Belgium, Brussels, Institut Royal des Sciences Naturelles de Belgique" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="22" pageNumber="23">IRSNB</collectionCode>
):
<collectingRegion box="[267,345,661,685]" country="Vietnam" name="Ho Chi Minh City" pageId="22" pageNumber="23">Saigon</collectingRegion>
[=
<collectingRegion box="[384,583,661,685]" country="Vietnam" name="Ho Chi Minh City" pageId="22" pageNumber="23">Ho Chi Minh city</collectingRegion>
, coordinates ca.
<geoCoordinate box="[769,883,661,685]" degrees="10" direction="north" minutes="46.5" orientation="latitude" pageId="22" pageNumber="23" precision="92" value="10.775">
<number box="[769,796,661,685]" pageId="22" pageNumber="23" value="10.0">10</number>
°
<number box="[805,858,661,685]" pageId="22" pageNumber="23" value="46.5">46.5</number>
'N
</geoCoordinate>
<geoCoordinate box="[889,1019,661,685]" degrees="106" direction="east" minutes="41.5" orientation="longitude" pageId="22" pageNumber="23" precision="92" value="106.691666">
<number box="[889,933,661,685]" pageId="22" pageNumber="23" value="106.0">106</number>
°
<number box="[943,997,661,685]" pageId="22" pageNumber="23" value="41.5">41.5</number>
'E
</geoCoordinate>
], without date and collector.
</materialsCitation>
</paragraph>
<paragraph blockId="22.[151,1437,373,1887]" pageId="22" pageNumber="23">
<emphasis bold="true" box="[199,644,697,722]" pageId="22" pageNumber="23">
Published records.
<collectingCountry box="[428,515,697,722]" name="India" pageId="22" pageNumber="23">INDIA</collectingCountry>
: Manipur:
</emphasis>
Imphal district, Ucharthal,
<quantity box="[954,1011,697,721]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="22" pageNumber="23" unit="km" value="5.0">5 km</quantity>
E of Jiribam (
<bibRefCitation author="Mukhopadhyay &amp; Sengupta" firstAuthor="Mukhopadhyay" pageId="22" pageNumber="23" pagination="449 - 458" refId="ref42687" refString="Mukhopadhyay, P. &amp; Sengupta, S. (2004) Insecta: Coleoptera: Polyphaga: Hydrophiloidea: Hydrophilidae. Pp. 449 - 458. In: Fauna of Manipur, State Fauna Series, volume 10. Zoological Survey of India." type="journal article" year="2004">
Mukhopadhyay &amp; Sengupta
<number box="[220,281,733,757]" pageId="22" pageNumber="23" value="2004.0">2004</number>
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="22" pageNumber="23" type="diagnosis">
<paragraph blockId="22.[151,1437,373,1887]" pageId="22" pageNumber="23">
<emphasis bold="true" box="[199,463,769,794]" pageId="22" pageNumber="23">Differential diagnosis.</emphasis>
Rather easily distinguishable from all remaining species occurring in continental SE Asia by its pale reddish general coloration (
<figureCitation box="[642,725,802,826]" captionStart="FIGURE 54" captionStartId="23.[151,250,790,813]" captionTargetBox="[177,1413,197,757]" captionTargetId="figure@23.[146,1431,186,774]" captionText="FIGURE 54. Pachysternum coomani, general habitus (a, cd: dorsal view, b: laterodorsal view). Localities of the specimens: ab: Malaysia, Kedah (male); c: same locality (female); d: Thailand, Phetchabun (male)." pageId="22" pageNumber="23">
Fig.
<number box="[694,725,802,826]" pageId="22" pageNumber="23" value="54.0">54</number>
</figureCitation>
), small body size and continually arcuate outer margin of anterior tibiae (
<figureCitation box="[275,359,836,860]" captionStart="FIGURES 39 50" captionStartId="11.[151,264,1939,1962]" captionTargetBox="[230,1341,208,1896]" captionTargetId="figure@11.[201,1370,193,1918]" captionTargetPageId="11" captionText="FIGURES 39 50. Anterior tibia of Asian Pachysternum species. 39: P. apicatum (a: lectotype, b: specimen from Borneo, Sabah); 40: P. c u r v a t u m, 41: P. sandacanum; 42: P. c a rd o n i; 43: P. coomani, 44: P. haemorrhoum, 45: P. kubani, 46: P. nigrovittatum, 47: P. rugosum, 48: P. stevensi, 49: P. s u l a w e s i c u m." httpUri="https://zenodo.org/record/280228/files/figure.png" pageId="22" pageNumber="23">
Fig.
<number box="[327,359,836,860]" pageId="22" pageNumber="23" value="43.0">43</number>
</figureCitation>
); the very wide and nearly parallel-sided median lobe of the aedeagus (
<figureCitation box="[1150,1231,836,860]" captionStart="FIGURES 29 32" captionStartId="8.[151,264,1975,1998]" captionTargetBox="[162,1428,577,1952]" captionTargetId="figure@8.[151,1436,561,1962]" captionTargetPageId="8" captionText="FIGURES 29 32. Male genitalia of Asian Pachysternum (a: tegmen, b: median lobe, c: apical portionof median lobe, d: sternite 9). 29: P. c a rd o n i, 30: P. coomani, 31: P. haemorrhoum; 32: P. nigrovitattum." httpUri="https://zenodo.org/record/280227/files/figure.png" pageId="22" pageNumber="23">
Fig.
<number box="[1202,1231,836,860]" pageId="22" pageNumber="23" value="30.0">30</number>
</figureCitation>
) is also very characteristic and cannot be confused with any other species. Darker specimens with stronger interstitial microsculpture on elytra may resemble
<taxonomicName box="[425,541,903,927]" class="Insecta" family="Hydrophilidae" genus="Pachysternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="stevensi">
<emphasis box="[425,541,903,927]" italics="true" pageId="22" pageNumber="23">P. stevensi</emphasis>
</taxonomicName>
, but can be easily distinguished from it by continually arcuate outer margin of anterior tibiae (distinctly sinuate in
<taxonomicName box="[550,666,937,960]" class="Insecta" family="Hydrophilidae" genus="Pachysternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="stevensi">
<emphasis box="[550,666,937,960]" italics="true" pageId="22" pageNumber="23">P. stevensi</emphasis>
</taxonomicName>
) and wide, nearly parallel-sided median lobe. By general coloration,
<taxonomicName box="[151,416,969,993]" class="Insecta" family="Hydrophilidae" genus="Pachysternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="coomani">
<emphasis box="[151,416,969,993]" italics="true" pageId="22" pageNumber="23">Pachysternum coomani</emphasis>
</taxonomicName>
also slightly resembles Bornean specimens of
<emphasis box="[955,1085,970,993]" italics="true" pageId="22" pageNumber="23">P. a p i c a t u m</emphasis>
, from whose it also differs by continually arcuate outer margin of anterior tibiae (rather deeply sinuate in Bornean
<taxonomicName box="[1096,1227,1004,1027]" class="Insecta" family="Hydrophilidae" genus="Pachysternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="apicatum">
<emphasis box="[1096,1227,1004,1027]" italics="true" pageId="22" pageNumber="23">P. apicatum</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="description">
<paragraph blockId="22.[151,1437,373,1887]" pageId="22" pageNumber="23">
<emphasis bold="true" box="[199,348,1035,1060]" pageId="22" pageNumber="23">Description.</emphasis>
Body widely oval, shape of elytra not sexually dimorphic, female without distinctly pronounced humeral portion of elytra. Body length
<quantity box="[704,841,1071,1095]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.9" metricValueMin="2.2" pageId="22" pageNumber="23" unit="mm" value="2.55" valueMax="2.9" valueMin="2.2">2.22.9 mm</quantity>
(
<typeStatus box="[856,962,1071,1095]" pageId="22" pageNumber="23">holotype</typeStatus>
:
<quantity box="[973,1065,1071,1095]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="22" pageNumber="23" unit="mm" value="2.4">2.4 mm</quantity>
); body width
<quantity box="[1221,1359,1071,1095]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="22" pageNumber="23" unit="mm" value="1.65" valueMax="1.8" valueMin="1.5">1.51.8 mm</quantity>
(
<typeStatus pageId="22" pageNumber="23">holotype</typeStatus>
<quantity box="[207,299,1108,1132]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="22" pageNumber="23" unit="mm" value="1.6">1.6 mm</quantity>
).
</paragraph>
<paragraph blockId="22.[151,1437,373,1887]" pageId="22" pageNumber="23">
<emphasis box="[199,322,1144,1168]" italics="true" pageId="22" pageNumber="23">Coloration</emphasis>
(
<figureCitation box="[338,424,1143,1167]" captionStart="FIGURE 54" captionStartId="23.[151,250,790,813]" captionTargetBox="[177,1413,197,757]" captionTargetId="figure@23.[146,1431,186,774]" captionText="FIGURE 54. Pachysternum coomani, general habitus (a, cd: dorsal view, b: laterodorsal view). Localities of the specimens: ab: Malaysia, Kedah (male); c: same locality (female); d: Thailand, Phetchabun (male)." pageId="22" pageNumber="23">
Fig.
<number box="[392,424,1143,1167]" pageId="22" pageNumber="23" value="54.0">54</number>
</figureCitation>
). Whole dorsal surface more or less uniformly reddish brown, with slightly darker lateral portions of frons, posterolateral portions of pronotum, base of elytra and elytra at midlength. Ventral surface of head reddish brown, thoracic sclerites brown, abdominal ventrites reddish brown, legs pale reddish.
</paragraph>
<paragraph blockId="22.[151,1437,373,1887]" pageId="22" pageNumber="23">
<emphasis box="[199,439,1252,1276]" italics="true" pageId="22" pageNumber="23">External morphology</emphasis>
. Clypeus with moderately coarse but sparse punctation consisting of uniformly-sized rasp-like punctures separated from each other by ca.
<number box="[743,784,1287,1311]" pageId="22" pageNumber="23" value="2.0" valueMax="3.0" valueMin="1.0">13</number>
× puncture diameter. Punctation of frons similar to that on clypeus. Interstices on head without microsculpture. Larger punctures on pronotum rounded, shallow, much larger than small punctures; small punctures scar-like, divided from each other by ca.
<number box="[1051,1064,1359,1383]" pageId="22" pageNumber="23" value="3.0">3</number>
× puncture width. Pronotal interstices without microsculpture. Prosternum with very distinct median carina. Elytral series consisting of large, shallow punctures; interval punctation with rather fine semicircular punctures much smaller than serial ones, with sparsely intermixed setiferous punctures of the same shape and size; interstices with weak mesh-like microsculpture. Elytral series
<number box="[364,378,1503,1527]" pageId="22" pageNumber="23" value="1.0">1</number>
not impressed, lateral series only very weakly impressed. Preepisternal elevation of mesothorax without posterolateral pits. Femoral lines on metaventrite indistinctly angulate, median portion of metaventrite bearing rather fine, moderately dense setiferous punctation, interstices without microsculpture medially, with weak microsculpture consisting of partially joint ridges laterally; lateral portions with very coarse and dense, densely pubescent punctation. Anterior tibia rather wide, outer margin continually arcuate, outer series of spines not interrupted.
</paragraph>
<paragraph blockId="22.[151,1437,373,1887]" pageId="22" pageNumber="23">
<emphasis box="[199,364,1720,1744]" italics="true" pageId="22" pageNumber="23">
<collectingCountry box="[199,257,1720,1744]" name="Malta" pageId="22" pageNumber="23">Male</collectingCountry>
genitalia
</emphasis>
(
<figureCitation box="[379,460,1719,1743]" captionStart="FIGURES 29 32" captionStartId="8.[151,264,1975,1998]" captionTargetBox="[162,1428,577,1952]" captionTargetId="figure@8.[151,1436,561,1962]" captionTargetPageId="8" captionText="FIGURES 29 32. Male genitalia of Asian Pachysternum (a: tegmen, b: median lobe, c: apical portionof median lobe, d: sternite 9). 29: P. c a rd o n i, 30: P. coomani, 31: P. haemorrhoum; 32: P. nigrovitattum." httpUri="https://zenodo.org/record/280227/files/figure.png" pageId="22" pageNumber="23">
Fig.
<number box="[431,460,1719,1743]" pageId="22" pageNumber="23" value="30.0">30</number>
</figureCitation>
). Tegmen 1.0
<quantity box="[632,716,1719,1743]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="22" pageNumber="23" unit="mm" value="1.1">1.1 mm</quantity>
long, median lobe 1.0
<quantity box="[983,1067,1719,1743]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="22" pageNumber="23" unit="mm" value="1.1">1.1 mm</quantity>
long. Phallobase
<number box="[1267,1303,1719,1743]" pageId="22" pageNumber="23" value="2.5">2.5</number>
× as long as parameres, bearing small, slightly asymmetrical, distinctly detached basal manubrium. Median lobe rather wide, nearly parallel-sided at midlength, strongly narrowing apicad, apex widely rounded; gonopore subapical, rather indistinct; lateral pubescent lobes narrowly developed subapically. Sternite
<number box="[1003,1017,1828,1852]" pageId="22" pageNumber="23" value="9.0">9</number>
with moderately large V-shaped apical emargination.
</paragraph>
<caption pageId="23" pageNumber="24" startId="23.[151,250,790,813]" targetBox="[177,1413,197,757]" targetPageId="23">
<paragraph blockId="23.[151,1437,790,848]" pageId="23" pageNumber="24">
<emphasis bold="true" box="[151,289,790,813]" pageId="23" pageNumber="24">
FIGURE
<number box="[257,284,791,812]" pageId="23" pageNumber="24" value="54.0">54</number>
.
</emphasis>
<taxonomicName box="[296,535,791,813]" class="Insecta" family="Hydrophilidae" genus="Pachysternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="coomani">
<emphasis box="[296,535,791,813]" italics="true" pageId="23" pageNumber="24">Pachysternum coomani</emphasis>
</taxonomicName>
, general habitus (a, cd: dorsal view, b: laterodorsal view). Localities of the specimens: ab:
<collectingCountry box="[198,293,826,848]" name="Malaysia" pageId="23" pageNumber="24">Malaysia</collectingCountry>
, Kedah (male); c: same locality (female); d:
<collectingCountry box="[746,839,826,848]" name="Thailand" pageId="23" pageNumber="24">Thailand</collectingCountry>
, Phetchabun (male).
</paragraph>
</caption>
<caption httpUri="https://zenodo.org/record/280230/files/figure.png" pageId="23" pageNumber="24" targetBox="[311,1275,904,1723]" targetPageId="23">
<paragraph blockId="23.[151,1436,1740,1793]" pageId="23" pageNumber="24">
<emphasis bold="true" box="[151,289,1740,1763]" pageId="23" pageNumber="24">FIGURE 55.</emphasis>
Known distribution of
<taxonomicName box="[531,770,1740,1762]" class="Insecta" family="Hydrophilidae" genus="Pachysternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="coomani">
<emphasis box="[531,770,1740,1762]" italics="true" pageId="23" pageNumber="24">Pachysternum coomani</emphasis>
</taxonomicName>
(green dots: examined material; blue circles: published data, not examined).
</paragraph>
</caption>
<paragraph blockId="23.[151,1437,1867,2036]" pageId="23" pageNumber="24">
<emphasis bold="true" box="[199,315,1867,1892]" pageId="23" pageNumber="24">Variation.</emphasis>
Dorsal coloration varies slightly in the extent of the darker areas on the head, pronotum and elytra:
<typeStatus box="[151,199,1904,1928]" pageId="23" pageNumber="24">type</typeStatus>
specimens are paler (uniformly reddish) than some specimens from
<collectingCountry box="[1002,1108,1903,1927]" name="Malaysia" pageId="23" pageNumber="24">Malaysia</collectingCountry>
, in which some parts of the elytra are distinctly darker, brown (
<figureCitation box="[545,643,1939,1963]" captionStart="FIGURE 54" captionStartId="23.[151,250,790,813]" captionTargetBox="[177,1413,197,757]" captionTargetId="figure@23.[146,1431,186,774]" captionText="FIGURE 54. Pachysternum coomani, general habitus (a, cd: dorsal view, b: laterodorsal view). Localities of the specimens: ab: Malaysia, Kedah (male); c: same locality (female); d: Thailand, Phetchabun (male)." pageId="23" pageNumber="24">
Fig.
<number box="[597,628,1939,1963]" pageId="23" pageNumber="24" value="54.0">54</number>
d
</figureCitation>
). Microsculpture of elytral interstices varies from weakly developed to very distinct.
</paragraph>
</subSubSection>
<subSubSection box="[199,1437,2011,2036]" pageId="23" pageNumber="24" type="biology_ecology">
<paragraph blockId="23.[151,1437,1867,2036]" box="[199,1437,2011,2036]" pageId="23" pageNumber="24">
<emphasis bold="true" box="[199,292,2011,2036]" pageId="23" pageNumber="24">Biology.</emphasis>
The specimens from
<collectingCountry box="[528,628,2011,2035]" name="Malaysia" pageId="23" pageNumber="24">Malaysia</collectingCountry>
were collected in cow excrements; most probably a coprophilous species.
</paragraph>
</subSubSection>
<subSubSection pageId="24" pageNumber="25" type="distribution">
<paragraph blockId="24.[151,1436,151,284]" pageId="24" pageNumber="25">
<emphasis bold="true" box="[199,352,151,176]" pageId="24" pageNumber="25">Distribution.</emphasis>
Based on the examined specimens, the species seems to be widely distributed in the mainland SE Asia, reaching from northeastern
<collectingCountry box="[580,638,188,212]" name="India" pageId="24" pageNumber="25">India</collectingCountry>
(Manipur state) to southern
<collectingCountry box="[970,1065,188,212]" name="Vietnam" pageId="24" pageNumber="25">Vietnam</collectingCountry>
and Malayan peninsula, but the specimens are rather rare in collections. The species is recorded here for the first time from
<collectingCountry box="[1208,1314,223,247]" name="Malaysia" pageId="24" pageNumber="25">Malaysia</collectingCountry>
,
<collectingCountry box="[1327,1436,224,248]" name="Myanmar" pageId="24" pageNumber="25">Myanmar</collectingCountry>
and
<collectingCountry box="[198,301,260,284]" name="Thailand" pageId="24" pageNumber="25">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>