155 lines
18 KiB
XML
155 lines
18 KiB
XML
<document id="74ED88EC2527B9A1427CC29460648F9B" ID-DOI="10.5281/zenodo.178251" ID-GBIF-Dataset="905dea16-dd69-45f9-9fe2-5b1aa6af93b6" ID-ISSN="1175-5326" ID-Zenodo-Dep="178251" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459886785444" checkinUser="plazi" docAuthor="Hippa, Heikki & Vilkamaa, Pekka" docDate="2007" docId="03DB45439C7CFFC143DDFC3FFBD5FEAB" docLanguage="en" docName="zt01556p050.pdf" docOrigin="Zootaxa 1556" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Keilbachia foveolata Hippa & Vilkamaa, 2007, sp. n." docType="treatment" docVersion="9" lastPageNumber="43" masterDocId="FFE23D3B9C70FFCC434AFFE0FFC7FFA9" masterDocTitle="The flagria group of Keilbachia Mohrig (Diptera, Sciaridae) in a biodiversity hot spot: nine new sympatric species from Kambaiti, Myanmar" masterLastPageNumber="50" masterPageNumber="31" pageNumber="43" updateTime="1698227854892" updateUser="plazi">
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<mods:mods id="BD73722CEAC26D9DAA1340C2D1447457" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="BBA00AEF9C24FE7A627F2DC2C31F550A">
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<mods:title id="6CF675364C6096FDC4AE7743A69D59A3">The flagria group of Keilbachia Mohrig (Diptera, Sciaridae) in a biodiversity hot spot: nine new sympatric species from Kambaiti, Myanmar</mods:title>
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</mods:titleInfo>
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<mods:name id="B69997B8BEA43C3D7DC098A7A9EB7CA5" type="personal">
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<mods:roleTerm id="1A6D4A9499409FB7F427068B2426AB2D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6C3E30FE73AC6D9A9F596823405DE912">Hippa, Heikki</mods:namePart>
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<mods:name id="471E8A47251001FD26F0DF360CDB3E51" type="personal">
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<mods:roleTerm id="4806509894B6D388D6B856D313B4F647">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5E2B1E9C9EC769E311D661B5BB0CFD55">Vilkamaa, Pekka</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="ED1AB2A64CEE3FB05C1ADEDF34AE7A09">text</mods:typeOfResource>
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<mods:titleInfo id="FA8363004D80698F87E7E917B372A737">
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<mods:title id="1CA1AC00FF5D336F940FB25DD8958715">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="AB88267C0FB8E34537E8856CEF72DEA0">
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<mods:date id="025E15FA009D8AC3E69B697294CFBB15">2007</mods:date>
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<mods:detail id="05C974D81F41C0A0ED40A3FBA5A6BE69" type="volume">
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<mods:number id="BEFA3869335A0166E1DDB13D01B9DC70">1556</mods:number>
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<mods:start id="174C65D31F6A908DB517807B40E02788">31</mods:start>
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<mods:end id="8E1D0DBEC77175A0D65F18FDBAF746DA">50</mods:end>
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<mods:classification id="C5E69C39C54D83A5CA80B1202315D23E">journal article</mods:classification>
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<mods:identifier id="910E049389B778B760706E6E1BA97CF3" type="DOI">10.5281/zenodo.178251</mods:identifier>
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<mods:identifier id="8BD43BC72F0422BFAD84EB402B8D5BB9" type="GBIF-Dataset">905dea16-dd69-45f9-9fe2-5b1aa6af93b6</mods:identifier>
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<mods:identifier id="8FC4627707842B68242E54244D397A1B" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="3C3D01C229DA04DA451B5474D7319E52" type="Zenodo-Dep">178251</mods:identifier>
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<treatment id="03DB45439C7CFFC143DDFC3FFBD5FEAB" ID-DOI="http://doi.org/10.5281/zenodo.6241302" ID-GBIF-Taxon="119371644" ID-Zenodo-Dep="6241302" LSID="urn:lsid:plazi:treatment:03DB45439C7CFFC143DDFC3FFBD5FEAB" httpUri="http://treatment.plazi.org/id/03DB45439C7CFFC143DDFC3FFBD5FEAB" lastPageId="13" lastPageNumber="43" pageId="12" pageNumber="43">
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<subSubSection id="C368A7DE9C7CFFC043DDFC3FFE1FFBB5" pageId="12" pageNumber="43" type="nomenclature">
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<paragraph id="8BCDF4559C7CFFC043DDFC3FFE1FFC50" blockId="12.[151,472,991,1052]" box="[151,472,991,1017]" pageId="12" pageNumber="43">
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<heading id="D08543399C7CFFC043DDFC3FFE1FFC50" bold="true" box="[151,472,991,1017]" fontSize="11" level="1" pageId="12" pageNumber="43" reason="1">
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<emphasis id="B90628479C7CFFC043DDFC3FFE1FFC50" bold="true" box="[151,472,991,1017]" pageId="12" pageNumber="43">
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<taxonomicName id="4C728FD69C7CFFC043DDFC3FFE49FC50" ID-CoL="3R4LN" box="[151,398,991,1017]" class="Insecta" family="Sciaridae" genus="Keilbachia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="foveolata" status="sp. nov.">
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<emphasis id="B90628479C7CFFC043DDFC3FFE49FC50" bold="true" box="[151,398,991,1017]" italics="true" pageId="12" pageNumber="43">Keilbachia foveolata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A235953C9C7CFFC042DCFC3FFE1FFC50" box="[406,472,991,1017]" pageId="12" pageNumber="43" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BCDF4559C7CFFC043DDFBE2FE1FFBB5" blockId="12.[151,472,991,1052]" box="[151,472,1026,1052]" pageId="12" pageNumber="43">
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(
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<figureCitation id="1349E8D09C7CFFC043D5FBE2FF22FBB5" box="[159,229,1026,1052]" captionStart="FIGURE 1" captionStartId="3.[151,255,1939,1963]" captionTargetBox="[232,1367,193,1884]" captionTargetId="figure@3.[198,1385,154,1913]" captionTargetPageId="3" captionText="FIGURE 1. Maxillary palpus. A, Keilbachia acamptochaeta sp. n. (paratype). B, K. flabellata sp. n. (holotype). C, K. gigas sp. n. (holotype). D, K. fasciata sp. n. (holotype). E, K. foveolata sp. n. (holotype). F, K. bicuspis sp. n. (holotype). G, K. rima sp. n. (holotype). H, K. filigera sp. n. (holotype). I, K. oligonema sp. n. (holotype). Scale 0.10 mm." httpUri="https://zenodo.org/record/178252/files/figure.png" pageId="12" pageNumber="43">Figs 1</figureCitation>
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E, 3F, 4B, 8B, C, D)
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</paragraph>
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</subSubSection>
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<subSubSection id="C368A7DE9C7CFFC043DDFBB2FE5EF8CD" pageId="12" pageNumber="43" type="materials_examined">
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<paragraph id="8BCDF4559C7CFFC043DDFBB2FF0BFB3D" blockId="12.[151,1437,1106,2012]" pageId="12" pageNumber="43">
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<emphasis id="B90628479C7CFFC043DDFBB2FEAAFBC5" bold="true" box="[151,365,1106,1132]" pageId="12" pageNumber="43">Material studied.</emphasis>
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<emphasis id="B90628479C7CFFC0423FFBB3FDE6FBC5" box="[373,545,1107,1132]" italics="true" pageId="12" pageNumber="43">
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<typeStatus id="54C94AF79C7CFFC0423FFBB3FE27FBC5" box="[373,480,1107,1132]" pageId="12" pageNumber="43" type="holotype">Holotype</typeStatus>
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male
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</emphasis>
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(in
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<collectionCode id="ED636C909C7CFFC0411BFBB2FD6EFBC5" box="[593,681,1106,1132]" pageId="12" pageNumber="43">NRMS</collectionCode>
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). N.E.
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<collectingCountry id="F365B4C59C7CFFC041B0FBB2FC8DFBC5" box="[762,842,1106,1132]" name="Myanmar" pageId="12" pageNumber="43">Burma</collectingCountry>
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(
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<collectingCountry id="F365B4C59C7CFFC04013FBB2FC16FBC5" box="[857,977,1106,1132]" name="Myanmar" pageId="12" pageNumber="43">Myanmar</collectingCountry>
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), Kambaiti,
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<quantity id="4C8A59B09C7CFFC0472BFBB2FB7DFBC5" box="[1121,1210,1106,1132]" metricMagnitude="3" metricUnit="m" metricValue="2.1336" pageId="12" pageNumber="43" unit="ft" value="7000.0">7000 ft</quantity>
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.,
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<date id="FFCCD2959C7CFFC04781FBB2FAF3FBC5" box="[1227,1332,1106,1132]" pageId="12" pageNumber="43" value="1934-04-04">4.4.1934</date>
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, R. Malaise.
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</paragraph>
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<paragraph id="8BCDF4559C7CFFC0438CFB42FBC2F895" blockId="12.[151,1437,1106,2012]" pageId="12" pageNumber="43">
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Male.
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<emphasis id="B90628479C7CFFC0425EFB42FEADFB15" bold="true" box="[276,362,1186,1212]" pageId="12" pageNumber="43">Colour</emphasis>
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. The single specimen is strongly faded, almost unicolorous pale yellowish-brown.
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<emphasis id="B90628479C7CFFC04618FB42FA52FB15" bold="true" box="[1362,1429,1186,1212]" pageId="12" pageNumber="43">Head</emphasis>
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. Eye bridge,
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<figureCitation id="1349E8D09C7CFFC04261FB2AFEB4FB4D" box="[299,371,1226,1252]" captionStart="FIGURE 8" captionStartId="13.[151,255,1962,1986]" captionTargetBox="[151,1388,351,1882]" captionTargetId="figure@13.[151,1436,317,1935]" captionTargetPageId="13" captionText="FIGURE 8. Eye bridge (A, B), frontal view, part of hypopygium (C, E) and gonostylus (D, F), ventral view. A, E, F, Keilbachia gigas sp. n. (holotype). B, C, D, K. foveolata sp. n. (B, D, E holotype, C non-type). Scale 0.10 mm." httpUri="https://zenodo.org/record/178259/files/figure.png" pageId="12" pageNumber="43">Fig. 8</figureCitation>
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B, 3 facets wide, medially with a gap of two facets width. Face (prefrons) with 7 setae. Clypeus non-setose. Maxillary palpus (
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<figureCitation id="1349E8D09C7CFFC0412EFB12FD61FAA5" box="[612,678,1266,1292]" captionStart="FIGURE 1" captionStartId="3.[151,255,1939,1963]" captionTargetBox="[232,1367,193,1884]" captionTargetId="figure@3.[198,1385,154,1913]" captionTargetPageId="3" captionText="FIGURE 1. Maxillary palpus. A, Keilbachia acamptochaeta sp. n. (paratype). B, K. flabellata sp. n. (holotype). C, K. gigas sp. n. (holotype). D, K. fasciata sp. n. (holotype). E, K. foveolata sp. n. (holotype). F, K. bicuspis sp. n. (holotype). G, K. rima sp. n. (holotype). H, K. filigera sp. n. (holotype). I, K. oligonema sp. n. (holotype). Scale 0.10 mm." httpUri="https://zenodo.org/record/178252/files/figure.png" pageId="12" pageNumber="43">Fig. 1</figureCitation>
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E) with 2 palpomeres. Palpomere 1 with 1 seta, with many hyaline sensilla scattered on the dorsal surface, without sensory pit. Palpomere 2 with 4 setae. Antennal flagellomeres rather inflated in KOH and not drawn, flagellomere 4, in a non-inflated condition, apparently twice as long as broad.
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<emphasis id="B90628479C7CFFC042FBFA8AFDCAFA2D" bold="true" box="[433,525,1386,1412]" pageId="12" pageNumber="43">Thorax</emphasis>
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. Anterior pronotum with 1 seta. Episternum 1 with 5 setae.
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<emphasis id="B90628479C7CFFC04791FA8AFAD8FA2D" bold="true" box="[1243,1311,1386,1412]" pageId="12" pageNumber="43">Wing</emphasis>
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(
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<figureCitation id="1349E8D09C7CFFC0467AFA8AFABDFA2D" box="[1328,1402,1386,1412]" captionStart="FIGURE 4" captionStartId="6.[151,255,1974,1998]" captionTargetBox="[302,1325,197,1973]" captionTargetId="figure@6.[271,1328,160,2005]" captionTargetPageId="6" captionText="FIGURE 4. Wing, dorsal view. A, Keilbachia flabellata sp. n. (holotype). B, K. foveolata sp. n. (holotype). C, K. gigas sp. n. (holotype). D, K. rima sp. n. (paratype). Scale 1.0 mm." httpUri="https://zenodo.org/record/178255/files/figure.png" pageId="12" pageNumber="43">Fig. 4</figureCitation>
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B). Length
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<quantity id="4C8A59B09C7CFFC043BBFA72FE9AFA05" box="[241,349,1426,1452]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" pageId="12" pageNumber="43" unit="mm" value="2.75">2.75 mm</quantity>
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. Width/length 0.41. R1/R 0.98. c/
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<geoCoordinate id="EE4692929C7CFFC041A3FA72FCFAFA05" box="[745,829,1426,1452]" direction="west" orientation="longitude" pageId="12" pageNumber="43" precision="555" value="-0.64">w 0.64</geoCoordinate>
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. r-m and bM nearly equally long, r-m/bM 1.08, r-m with 1 seta very close to Rs, bM non-setose.
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<emphasis id="B90628479C7CFFC041E6FA5AFD20FA7D" bold="true" box="[684,743,1466,1492]" pageId="12" pageNumber="43">Legs</emphasis>
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. Apex of fore tibia,
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<figureCitation id="1349E8D09C7CFFC04091FA5AFBD8FA7D" box="[987,1055,1466,1492]" captionStart="FIGURE 3" captionStartId="5.[151,255,1905,1929]" captionTargetBox="[188,1393,195,1864]" captionTargetId="figure@5.[163,1428,158,1890]" captionTargetPageId="5" captionText="FIGURE 3. Apical part of fore tibia, prolateral view. A, Keilbachia oligonema sp. n. (Burma). B, K. fasciata sp. n. (paratype). C, K. rima sp. n. (holotype). D, K. acamptochaeta sp. n. (paratype). E, K. flabellata sp. n. (holotype). F, K. foveolata sp. n. (holotype). G, K. filigera sp. n. (holotype). H, K. bicuspis sp. n. (holotype). I, K. gigas sp. n. (holotype). Scale 0.10 mm." httpUri="https://zenodo.org/record/178254/files/figure.png" pageId="12" pageNumber="43">
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Fig.
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<specimenCount id="9D743FDC9C7CFFC0475BFA5AFBD8FA7D" box="[1041,1055,1466,1492]" pageId="12" pageNumber="43" type="generic">3</specimenCount>
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</figureCitation>
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F. Length of fore basitarsomere/ length of fore tibia ca. 0.72. Length of fore tibia/length of fore femur ca. 1.60. Length of hind tibia/length of scutum+scutellum ca 2.26.
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<emphasis id="B90628479C7CFFC04297F9EAFD90F98D" bold="true" box="[477,599,1546,1572]" pageId="12" pageNumber="43">Abdomen</emphasis>
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. Sternite 8 visible in lateral view and the setae difficult to count, with ca. 20 setae, in two transverse rows on posterior half.
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<emphasis id="B90628479C7CFFC04068F9D2FC0CF9E5" bold="true" box="[802,971,1586,1612]" pageId="12" pageNumber="43">Hypopygium.</emphasis>
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<figureCitation id="1349E8D09C7CFFC0409FF9D2FBDCF9E5" box="[981,1051,1586,1612]" captionStart="FIGURE 8" captionStartId="13.[151,255,1962,1986]" captionTargetBox="[151,1388,351,1882]" captionTargetId="figure@13.[151,1436,317,1935]" captionTargetPageId="13" captionText="FIGURE 8. Eye bridge (A, B), frontal view, part of hypopygium (C, E) and gonostylus (D, F), ventral view. A, E, F, Keilbachia gigas sp. n. (holotype). B, C, D, K. foveolata sp. n. (B, D, E holotype, C non-type). Scale 0.10 mm." httpUri="https://zenodo.org/record/178259/files/figure.png" pageId="12" pageNumber="43">Fig. 8</figureCitation>
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C, D: Gonostylus with 2–3 long curved mesial megasetae arising from a common basal body; whether the number of these megasetae really is two on the left side in the
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<typeStatus id="54C94AF79C7CFFC04284F962FDF3F935" box="[462,564,1666,1692]" pageId="12" pageNumber="43" type="holotype">holotype</typeStatus>
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is uncertain because the number of sockets is impossible to see. Between the curved megasetae and the apical tooth with two widely-separated megasetae. Near the base of the apical tooth with two or three strong setae diverging from other nearby setae, at least one of which has a curvature near base. Posteriorly to the ventral intercoxal area with a small pit, not like the other species. Tegmen very pale in this specimen and the details difficult to see: it seems rather simple.
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</paragraph>
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<paragraph id="8BCDF4559C7CFFC0438CF8AAFE5EF8CD" blockId="12.[151,1437,1106,2012]" box="[198,409,1866,1892]" pageId="12" pageNumber="43">Female unknown.</paragraph>
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</subSubSection>
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<subSubSection id="C368A7DE9C7CFFC1438CF892FBD5FEAB" lastPageId="13" lastPageNumber="44" pageId="12" pageNumber="43" type="discussion">
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<paragraph id="8BCDF4559C7CFFC1438CF892FBD5FEAB" blockId="12.[151,1437,1106,2012]" lastBlockId="13.[151,1436,152,258]" lastPageId="13" lastPageNumber="44" pageId="12" pageNumber="43">
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<emphasis id="B90628479C7CFFC0438CF892FE95F825" bold="true" box="[198,338,1906,1932]" pageId="12" pageNumber="43">Discussion.</emphasis>
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<taxonomicName id="4C728FD69C7CFFC04217F893FD90F825" box="[349,599,1907,1932]" class="Insecta" family="Sciaridae" genus="Keilbachia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="foveolata">
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<emphasis id="B90628479C7CFFC04217F893FD90F825" box="[349,599,1907,1932]" italics="true" pageId="12" pageNumber="43">Keilbachia foveolata</emphasis>
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</taxonomicName>
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is similar to
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<taxonomicName id="4C728FD69C7CFFC04048F893FCA7F825" box="[770,864,1907,1932]" class="Insecta" family="Sciaridae" genus="Keilbachia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="rima">
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<emphasis id="B90628479C7CFFC04048F893FCA7F825" box="[770,864,1907,1932]" italics="true" pageId="12" pageNumber="43">K. rima</emphasis>
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</taxonomicName>
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. It is distinguished by having two palpomeres instead of three and by lacking a megaseta at the base of the apical tooth of the gonostylus. In
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<taxonomicName id="4C728FD69C7CFFC04641F87BFA5BF81D" box="[1291,1436,1947,1972]" class="Insecta" family="Sciaridae" genus="Keilbachia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="foveolata">
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<emphasis id="B90628479C7CFFC04641F87BFA5BF81D" box="[1291,1436,1947,1972]" italics="true" pageId="12" pageNumber="43">K. foveolata</emphasis>
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</taxonomicName>
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there are two or three strong setae ventrally near the base of the apical tooth of the gonostylus, which differ distinctly from the other nearby setae; in
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<taxonomicName id="4C728FD69C7DFFC14137FF78FD10FF18" box="[637,727,152,178]" class="Insecta" family="Sciaridae" genus="Keilbachia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="44" phylum="Arthropoda" rank="species" species="rima">
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<emphasis id="B90628479C7DFFC14137FF78FD57FF18" box="[637,656,152,177]" italics="true" pageId="13" pageNumber="44">K</emphasis>
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.
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<emphasis id="B90628479C7DFFC141EAFF78FD10FF18" box="[672,727,152,177]" italics="true" pageId="13" pageNumber="44">rima</emphasis>
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</taxonomicName>
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such setae are lacking.
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<taxonomicName id="4C728FD69C7DFFC140BAFF78FB47FF18" box="[1008,1152,152,177]" class="Insecta" family="Sciaridae" genus="Keilbachia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="44" phylum="Arthropoda" rank="species" species="foveolata">
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<emphasis id="B90628479C7DFFC140BAFF78FB47FF18" box="[1008,1152,152,177]" italics="true" pageId="13" pageNumber="44">K. foveolata</emphasis>
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</taxonomicName>
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differs from
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<taxonomicName id="4C728FD69C7DFFC14656FF78FAB1FF18" box="[1308,1398,152,177]" class="Insecta" family="Sciaridae" genus="Keilbachia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="44" phylum="Arthropoda" rank="species" species="rima">
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||
<emphasis id="B90628479C7DFFC14656FF78FAB1FF18" box="[1308,1398,152,177]" italics="true" pageId="13" pageNumber="44">K. rima</emphasis>
|
||
</taxonomicName>
|
||
by having a small sclerotized pit medially in the membrane on the intercoxal area of hypopygium. In this respect,
|
||
<taxonomicName id="4C728FD69C7DFFC143DDFF08FEE1FEA8" box="[151,294,232,257]" class="Insecta" family="Sciaridae" genus="Keilbachia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="44" phylum="Arthropoda" rank="species" species="foveolata">
|
||
<emphasis id="B90628479C7DFFC143DDFF08FEE1FEA8" box="[151,294,232,257]" italics="true" pageId="13" pageNumber="44">K. foveolata</emphasis>
|
||
</taxonomicName>
|
||
resembles
|
||
<emphasis id="B90628479C7DFFC142E0FF08FDAEFEA8" box="[426,617,232,257]" italics="true" pageId="13" pageNumber="44">
|
||
<taxonomicName id="4C728FD69C7DFFC142E0FF08FDECFEA8" box="[426,555,232,257]" class="Insecta" family="Sciaridae" genus="Keilbachia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="44" phylum="Arthropoda" rank="genus">Keilbachia</taxonomicName>
|
||
sp. 3
|
||
</emphasis>
|
||
, but the latter has three palpomeres.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |