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<document id="39DDC8F1B62FD08696920370441326B6" ID-CLB-Dataset="22261" ID-DOI="10.11646/zootaxa.4772.3.1" ID-GBIF-Dataset="d807f185-5391-4c8c-84c6-2550370b3cd3" ID-ISSN="1175-5326" ID-Zenodo-Dep="3819654" ID-ZooBank="F546A223-59A0-4DA1-9102-AD7BC482105F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589175628250" checkinUser="plazi" docAuthor="DellAngelo, Bruno, Landau, Bernard M., Sosso, Maurizio &amp; Taviani, Marco" docDate="2020" docId="03D21D39FFB83952FF7BFA8EFC8BDF47" docLanguage="en" docName="zootaxa.4772.3.1.pdf" docOrigin="Zootaxa 4772 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Callochiton Gray 1847" docType="treatment" docVersion="7" lastPageNumber="412" masterDocId="FFEB6541FFB23959FFECFFB3FFF0DE03" masterDocTitle="Late Pleistocene Red Sea Mollusca: 1. Polyplacophora" masterLastPageNumber="449" masterPageNumber="401" pageNumber="411" updateTime="1698839092336" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="AD6E65AD172CF9AC2057528A8F06EFE5">Late Pleistocene Red Sea Mollusca: 1. Polyplacophora</mods:title>
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<mods:namePart id="31804E8A58CDE40F89C8D614BB45BC4C">DellAngelo, Bruno</mods:namePart>
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<mods:namePart id="AC708199CBB5FE277BB0C3C2A8B38E29">Landau, Bernard M.</mods:namePart>
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<mods:namePart id="83917A46AE1DBABF8851468EB64E2E6E">Sosso, Maurizio</mods:namePart>
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<mods:namePart id="CD099E9B7E6692D794D18EA1D275CA45">Taviani, Marco</mods:namePart>
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<treatment id="03D21D39FFB83952FF7BFA8EFC8BDF47" ID-DOI="http://doi.org/10.5281/zenodo.3844075" ID-GBIF-Taxon="164035823" ID-Zenodo-Dep="3844075" LSID="urn:lsid:plazi:treatment:03D21D39FFB83952FF7BFA8EFC8BDF47" httpUri="http://treatment.plazi.org/id/03D21D39FFB83952FF7BFA8EFC8BDF47" lastPageId="11" lastPageNumber="412" pageId="10" pageNumber="411">
<subSubSection id="C361FFA4FFB83953FF7BFA8EFEA0DB54" box="[151,336,1341,1368]" pageId="10" pageNumber="411" type="nomenclature">
<paragraph id="8BC4AC2FFFB83953FF7BFA8EFEA0DB54" blockId="10.[151,336,1341,1404]" box="[151,336,1341,1368]" pageId="10" pageNumber="411">
<heading id="D08C1B43FFB83953FF7BFA8EFEA0DB54" bold="true" box="[151,336,1341,1368]" fontSize="11" level="1" pageId="10" pageNumber="411" reason="1">
<emphasis id="B90F703DFFB83953FF7BFA8EFEA0DB54" bold="true" box="[151,336,1341,1368]" pageId="10" pageNumber="411">
<taxonomicName id="4C7BD7ACFFB83953FF7BFA8EFED5DB5B" authorityName="Gray" authorityYear="1847" box="[151,293,1341,1368]" class="Polyplacophora" family="Callochitonidae" genus="Callochiton" kingdom="Animalia" order="Chitonida" pageId="10" pageNumber="411" phylum="Mollusca" rank="genus">
<emphasis id="B90F703DFFB83953FF7BFA8EFED5DB5B" bold="true" box="[151,293,1341,1368]" italics="true" pageId="10" pageNumber="411">Callochiton</emphasis>
</taxonomicName>
sp.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C361FFA4FFB83953FF7BFAD2FF1CDB7F" box="[151,236,1377,1404]" pageId="10" pageNumber="411" type="description">
<paragraph id="8BC4AC2FFFB83953FF7BFAD2FF1CDB7F" blockId="10.[151,336,1341,1404]" box="[151,236,1377,1404]" pageId="10" pageNumber="411">
(
<figureCitation id="1340B0AAFFB83953FF4CFAD2FF13DB7F" box="[160,227,1377,1404]" captionStart="FIGURE 6" captionStartId="11.[151,250,610,635]" captionTargetBox="[151,1436,345,585]" captionTargetId="figure@11.[151,1436,345,585]" captionTargetPageId="11" captionText="FIGURE 6. Callochiton sp., Egypt, Hurghada (St. 13), Late Pleistocene (last interglacial MIS5e). AD. MZB 50529, intermediate valve, width 4 mm, dorsal view, detail of tegmentum surface of central area, ventral and frontal views. Scale bar: 100 µm (B)." figureDoi="http://doi.org/10.5281/zenodo.3825468" httpUri="https://zenodo.org/record/3825468/files/figure.png" pageId="10" pageNumber="411">Fig. 6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C361FFA4FFB83953FF7BFA1BFAA4DBC0" box="[151,1364,1448,1475]" pageId="10" pageNumber="411" type="materials_examined">
<paragraph id="8BC4AC2FFFB83953FF7BFA1BFAA4DBC0" blockId="10.[151,1437,1448,2015]" box="[151,1364,1448,1475]" pageId="10" pageNumber="411">
<emphasis id="B90F703DFFB83953FF7BFA1BFE71DBC0" bold="true" box="[151,385,1448,1475]" pageId="10" pageNumber="411">Material examined.</emphasis>
<materialsCitation id="3B13A672FFB83953FE6BFA1AFAA0DBC0" ID-GBIF-Occurrence="2611558309" box="[391,1360,1448,1475]" collectionCode="MZB" country="Egypt" location="Hurghada" pageId="10" pageNumber="411" specimenCount="1">
<collectingCountry id="F36CECBFFFB83953FE6BFA1AFE3DDBC0" box="[391,461,1449,1475]" name="Egypt" pageId="10" pageNumber="411">Egypt</collectingCountry>
,
<location id="8EA4FAF4FFB83953FE34FA1AFDBADBC0" LSID="urn:lsid:plazi:treatment:03D21D39FFB83952FF7BFA8EFC8BDF47:8EA4FAF4FFB83953FE34FA1AFDBADBC0" box="[472,586,1449,1475]" country="Egypt" name="Hurghada" pageId="10" pageNumber="411">Hurghada</location>
: St. 13: 1 intermediate valve, width
<quantity id="4C8301CAFFB83953FC33FA1AFBD2DBC0" box="[991,1058,1449,1475]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="411" unit="mm" value="4.0">4 mm</quantity>
,
<figureCitation id="1340B0AAFFB83953FBC1FA1AFB81DBC0" box="[1069,1137,1449,1475]" captionStart="FIGURE 6" captionStartId="11.[151,250,610,635]" captionTargetBox="[151,1436,345,585]" captionTargetId="figure@11.[151,1436,345,585]" captionTargetPageId="11" captionText="FIGURE 6. Callochiton sp., Egypt, Hurghada (St. 13), Late Pleistocene (last interglacial MIS5e). AD. MZB 50529, intermediate valve, width 4 mm, dorsal view, detail of tegmentum surface of central area, ventral and frontal views. Scale bar: 100 µm (B)." figureDoi="http://doi.org/10.5281/zenodo.3825468" httpUri="https://zenodo.org/record/3825468/files/figure.png" pageId="10" pageNumber="411">Figs 6</figureCitation>
AD (
<collectionCode id="ED6A34EAFFB83953FB54FA1AFB06DBC0" box="[1208,1270,1449,1475]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="10" pageNumber="411" type="Museum">MZB</collectionCode>
50529)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C361FFA4FFB83953FF2BFA7FFE74D898" pageId="10" pageNumber="411" type="description">
<paragraph id="8BC4AC2FFFB83953FF2BFA7FFC30D808" blockId="10.[151,1437,1448,2015]" pageId="10" pageNumber="411">
<emphasis id="B90F703DFFB83953FF2BFA7FFEA9DBE4" bold="true" box="[199,345,1484,1511]" pageId="10" pageNumber="411">Description.</emphasis>
Intermediate valves trapezoidal, rounded in anterior profile, elevated (height/width ratio 0.61), anterior margin almost straight in jugal area, lateral areas weakly raised.
</paragraph>
<paragraph id="8BC4AC2FFFB83953FF2BF9A6FDDED850" blockId="10.[151,1437,1448,2015]" pageId="10" pageNumber="411">Tegmentum surface appears rough, sculptured with fine granules, fusing into continuous longitudinal lines in central area, radially in lateral areas.</paragraph>
<paragraph id="8BC4AC2FFFB83953FF2BF9EEFE74D898" blockId="10.[151,1437,1448,2015]" pageId="10" pageNumber="411">Articulamentum with short, wide apophyses connected at jugum by lamina, insertion plates short, with two slits, slit rays visible.</paragraph>
</subSubSection>
<subSubSection id="C361FFA4FFB83952FF2BF916FBF0DF1C" lastPageId="11" lastPageNumber="412" pageId="10" pageNumber="411" type="discussion">
<paragraph id="8BC4AC2FFFB83953FF2BF916FCADD977" blockId="10.[151,1437,1448,2015]" pageId="10" pageNumber="411">
<emphasis id="B90F703DFFB83953FF2BF916FECBD8BC" bold="true" box="[199,315,1701,1727]" pageId="10" pageNumber="411">Remarks.</emphasis>
This intermediate valve is characterised by the lack of longitudinal grooves on the pleural area (as in
<taxonomicName id="4C7BD7ACFFB83953FF7BF97AFEE4D8E0" authorityName="Ferreira" authorityYear="1983" box="[151,276,1737,1763]" class="Polyplacophora" family="Callochitonidae" genus="Callochiton" kingdom="Animalia" order="Chitonida" pageId="10" pageNumber="411" phylum="Mollusca" rank="species" species="vanninii">
<emphasis id="B90F703DFFB83953FF7BF97AFEE4D8E0" box="[151,276,1737,1763]" italics="true" pageId="10" pageNumber="411">C. vanninii</emphasis>
</taxonomicName>
), the rounded anterior profile and the strong elevation (H/W = 0.61), higher than that seen in intermediate valves of
<taxonomicName id="4C7BD7ACFFB83953FEDCF95EFE5ED904" authorityName="Ferreira" authorityYear="1983" box="[304,430,1773,1799]" class="Polyplacophora" family="Callochitonidae" genus="Callochiton" kingdom="Animalia" order="Chitonida" pageId="10" pageNumber="411" phylum="Mollusca" rank="species" species="vanninii">
<emphasis id="B90F703DFFB83953FEDCF95EFE5ED904" box="[304,430,1773,1799]" italics="true" pageId="10" pageNumber="411">C. vanninii</emphasis>
</taxonomicName>
(H/W = 0.300.48). These characters do not agree with those of the other named Indian Ocean
<taxonomicName id="4C7BD7ACFFB83953FF0BF8A2FE9BD928" authorityName="Gray" authorityYear="1847" box="[231,363,1809,1835]" class="Polyplacophora" family="Callochitonidae" genus="Callochiton" kingdom="Animalia" order="Chitonida" pageId="10" pageNumber="411" phylum="Mollusca" rank="genus">
<emphasis id="B90F703DFFB83953FF0BF8A2FE9BD928" box="[231,363,1809,1835]" italics="true" pageId="10" pageNumber="411">Callochiton</emphasis>
</taxonomicName>
species,
<taxonomicName id="4C7BD7ACFFB83953FE39F8A2FC51D928" authority="Kaas &amp; Van Belle, 1985 b" authorityName="Kaas &amp; Van Belle" authorityYear="1985" box="[469,929,1808,1835]" class="Polyplacophora" family="Callochitonidae" genus="Callochiton" kingdom="Animalia" order="Chitonida" pageId="10" pageNumber="411" phylum="Mollusca" rank="species" species="clausadeae">
<emphasis id="B90F703DFFB83953FE39F8A2FD85D928" box="[469,629,1809,1835]" italics="true" pageId="10" pageNumber="411">C. clausadeae</emphasis>
<bibRefCitation id="EFEAD1DEFFB83953FD92F8A2FC51D928" author="Kaas, P. &amp; Van Belle, R. A." box="[638,929,1808,1835]" pageId="10" pageNumber="411" refId="ref32880" refString="Kaas, P. &amp; Van Belle, R. A. (1985 b) Monograph of Living Chitons (Mollusca: Polyplacophora). Vol. 2. Suborder Ischnochi- tonina, Ischnochitonidae: Schizoplacinae, Callochitoninae &amp; Lepidochitoninae. E. J. Brill, W. Backhuys, Leiden, 198 pp." type="book" year="1985">Kaas &amp; Van Belle, 1985b</bibRefCitation>
</taxonomicName>
(from
<collectingCountry id="F36CECBFFFB83953FC1FF8A2FB8CD928" box="[1011,1148,1809,1835]" name="Madagascar" pageId="10" pageNumber="411">Madagascar</collectingCountry>
,
<collectingCountry id="F36CECBFFFB83953FB65F8A2FB09D928" box="[1161,1273,1809,1835]" name="Mauritius" pageId="10" pageNumber="411">Mauritius</collectingCountry>
,
<collectingCountry id="F36CECBFFFB83953FAE9F8A2FA97D928" box="[1285,1383,1809,1835]" name="Réunion" pageId="10" pageNumber="411">Réunion</collectingCountry>
, and the
<collectingCountry id="F36CECBFFFB83953FF2CF886FE86D953" box="[192,374,1845,1872]" name="Maldives" pageId="10" pageNumber="411">Maldive Islands</collectingCountry>
),
<taxonomicName id="4C7BD7ACFFB83953FE66F886FCFCD94C" authority="Kaas &amp; Van Belle, 1998" authorityName="Kaas &amp; Van Belle" authorityYear="1998" box="[394,780,1845,1871]" class="Polyplacophora" family="Callochitonidae" genus="Callochiton" kingdom="Animalia" order="Chitonida" pageId="10" pageNumber="411" phylum="Mollusca" rank="species" species="levatus">
<emphasis id="B90F703DFFB83953FE66F886FE09D94C" box="[394,505,1845,1871]" italics="true" pageId="10" pageNumber="411">C. levatus</emphasis>
<bibRefCitation id="EFEAD1DEFFB83953FE13F886FCFCD94C" author="Kaas, P. &amp; Van Belle, R. A." box="[511,780,1845,1871]" pageId="10" pageNumber="411" refId="ref33036" refString="Kaas, P. &amp; Van Belle, R. A. (1998) Catalogue of living chitons (Mollusca, Polyplacophora). Second, revised edition. Backuys Publishers, Leiden, 204 pp." type="book" year="1998">Kaas &amp; Van Belle, 1998</bibRefCitation>
</taxonomicName>
(from
<collectingCountry id="F36CECBFFFB83953FCB4F886FC11D94C" box="[856,993,1845,1871]" name="Madagascar" pageId="10" pageNumber="411">Madagascar</collectingCountry>
,
<collectingCountry id="F36CECBFFFB83953FC00F886FBBED94C" box="[1004,1102,1845,1871]" name="Réunion" pageId="10" pageNumber="411">Réunion</collectingCountry>
,
<collectingCountry id="F36CECBFFFB83953FBB4F886FB35D94C" box="[1112,1221,1845,1871]" name="Mauritius" pageId="10" pageNumber="411">Mauritius</collectingCountry>
and Chagos Archipelago) and
<taxonomicName id="4C7BD7ACFFB83953FECEF8EAFD61D970" authority="Schwabe, 2003" authorityName="Schwabe" authorityYear="2003" box="[290,657,1881,1907]" class="Polyplacophora" family="Callochitonidae" genus="Callochiton" kingdom="Animalia" order="Chitonida" pageId="10" pageNumber="411" phylum="Mollusca" rank="species" species="christamariae">
<emphasis id="B90F703DFFB83953FECEF8EAFE2FD970" box="[290,479,1881,1907]" italics="true" pageId="10" pageNumber="411">C. christamariae</emphasis>
<bibRefCitation id="EFEAD1DEFFB83953FE0AF8EAFD61D970" author="Schwabe, E." box="[486,657,1881,1907]" pageId="10" pageNumber="411" pagination="19 - 27" refId="ref35129" refString="Schwabe, E. (2003) Taxonomic notes on chitons. 3. Notes on the genus Callochiton Gray, 1847 (Mollusca: Polyplacophora: Callochitonidae) from the Indian Ocean. Malakologische Abhandlungen, 21, 19 - 27." type="journal article" year="2003">Schwabe, 2003</bibRefCitation>
</taxonomicName>
(from
<collectingCountry id="F36CECBFFFB83953FD33F8EAFCA1D977" box="[735,849,1881,1908]" name="Mauritius" pageId="10" pageNumber="411">Mauritius</collectingCountry>
).
</paragraph>
<paragraph id="8BC4AC2FFFB83952FF2BF8CEFBF0DF1C" blockId="10.[151,1437,1448,2015]" lastBlockId="11.[151,1436,153,324]" lastPageId="11" lastPageNumber="412" pageId="10" pageNumber="411">
Two fossil species of
<taxonomicName id="4C7BD7ACFFB83953FE54F8CEFDCCD994" authorityName="Gray" authorityYear="1847" box="[440,572,1917,1943]" class="Polyplacophora" family="Callochitonidae" genus="Callochiton" kingdom="Animalia" order="Chitonida" pageId="10" pageNumber="411" phylum="Mollusca" rank="genus">
<emphasis id="B90F703DFFB83953FE54F8CEFDCCD994" box="[440,572,1917,1943]" italics="true" pageId="10" pageNumber="411">Callochiton</emphasis>
</taxonomicName>
have a similar dorsal elevation.
<taxonomicName id="4C7BD7ACFFB83953FC4AF8CEFB7ED994" authorityName="Dell'Angelo &amp; Palazzi" authorityYear="1994" box="[934,1166,1917,1943]" class="Polyplacophora" family="Callochitonidae" genus="Callochiton" kingdom="Animalia" order="Chitonida" pageId="10" pageNumber="411" phylum="Mollusca" rank="species" species="calcatus">
<emphasis id="B90F703DFFB83953FC4AF8CEFB7ED994" box="[934,1166,1917,1943]" italics="true" pageId="10" pageNumber="411">Callochiton calcatus</emphasis>
</taxonomicName>
<bibRefCitation id="EFEAD1DEFFB83953FB79F8CEFF3FD9B8" author="Dell'Angelo, B. &amp; Palazzi, S." pageId="10" pageNumber="411" pagination="15 - 23" refId="ref31351" refString="Dell'Angelo, B. &amp; Palazzi, S. (1994) Callochiton calcatus n. sp. con note su Callochiton septemvalvis (Montagu, 1803). La Conchiglia, 26 (273), 15 - 23." type="journal article" year="1994">DellAngelo &amp; Palazzi, 1994</bibRefCitation>
(H/W = 0.650.72), known from the Recent Mediterranean and the submerged Pleistocene of the Tyrrhenian Sea geographic area between Capraia and Capo Corso,
<collectingCountry id="F36CECBFFFB83953FCE2F876FCB1D9DC" box="[782,833,1989,2015]" name="Italy" pageId="10" pageNumber="411">Italy</collectingCountry>
(
<bibRefCitation id="EFEAD1DEFFB83953FCBEF876FB7ED9DC" author="Dell'Angelo, B. &amp; Giusti, F." box="[850,1166,1989,2015]" pageId="10" pageNumber="411" pagination="51 - 58" refId="ref31080" refString="Dell'Angelo, B. &amp; Giusti, F. (1997) I Polyplacophora di una tafocenosi profonda. La Conchiglia, 29 (283), 51 - 58." type="journal article" year="1997">DellAngelo &amp; Giusti 1997</bibRefCitation>
), differs from
<emphasis id="B90F703DFFB83952FADEF876FF4BDEB0" italics="true" lastPageId="11" lastPageNumber="412" pageId="10" pageNumber="411">Callochi- ton</emphasis>
sp. mainly by the presence of longitudinal grooves on the pleural areas and the more angulated intermediate valves.
<taxonomicName id="4C7BD7ACFFB93952FF1BFF0EFE2DDED4" authorityName="DellAngelo, Landau, Van Dingenen &amp; Ceulemans, 201" authorityYear="2018" box="[247,477,189,215]" class="Polyplacophora" family="Callochitonidae" genus="Callochiton" kingdom="Animalia" order="Chitonida" pageId="11" pageNumber="412" phylum="Mollusca" rank="species" species="pouweri">
<emphasis id="B90F703DFFB93952FF1BFF0EFE2DDED4" box="[247,477,189,215]" italics="true" pageId="11" pageNumber="412">Callochiton pouweri</emphasis>
</taxonomicName>
<bibRefCitation id="EFEAD1DEFFB93952FE0FFF0EFB9ADED4" author="Dell'Angelo, B. &amp; Landau, B. &amp; Van Dingenen, F. &amp; Ceulemans, F." box="[483,1130,189,215]" pageId="11" pageNumber="412" pagination="1 - 62" refId="ref31168" refString="Dell'Angelo, B., Landau, B., Van Dingenen, F. &amp; Ceulemans, F. (2018 b) The upper Miocene chitons of northwest France (Mollusca: Polyplacophora). Zootaxa, 4447 (1), 1 - 62. https: // doi. org / 10.11646 / zootaxa. 4447.1.1" type="journal article" year="2018">DellAngelo, Landau, Van Dingenen &amp; Ceulemans, 2018b</bibRefCitation>
(H/W = 0.580.64), known from the upper Miocene of Anjou,
<collectingCountry id="F36CECBFFFB93952FDDEFF52FD71DEF8" box="[562,641,225,251]" name="France" pageId="11" pageNumber="412">France</collectingCountry>
, differs from
<taxonomicName id="4C7BD7ACFFB93952FCCFFF52FC57DEF8" authorityName="Gray" authorityYear="1847" box="[803,935,225,251]" class="Polyplacophora" family="Callochitonidae" genus="Callochiton" kingdom="Animalia" order="Chitonida" pageId="11" pageNumber="412" phylum="Mollusca" rank="genus">
<emphasis id="B90F703DFFB93952FCCFFF52FC57DEF8" box="[803,935,225,251]" italics="true" pageId="11" pageNumber="412">Callochiton</emphasis>
</taxonomicName>
sp. mainly by the presence of longitudinal grooves on the pleural areas and the valves being carinated in anterior profile.
</paragraph>
</subSubSection>
<subSubSection id="C361FFA4FFB93952FF2BFE9BFC8BDF47" box="[199,891,296,324]" pageId="11" pageNumber="412" type="distribution">
<paragraph id="8BC4AC2FFFB93952FF2BFE9BFC8BDF47" blockId="11.[151,1436,153,324]" box="[199,891,296,324]" pageId="11" pageNumber="412">
<emphasis id="B90F703DFFB93952FF2BFE9BFE91DF40" bold="true" box="[199,353,296,323]" pageId="11" pageNumber="412">Distribution.</emphasis>
Late Pleistocene:
<collectingCountry id="F36CECBFFFB93952FDC3FE9AFD82DF40" box="[559,626,297,323]" name="Egypt" pageId="11" pageNumber="412">Egypt</collectingCountry>
(Hurghada: this study).
</paragraph>
</subSubSection>
</treatment>
</document>